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Balanites aegyptiaca Balanitaceae - World Agroforestry Centre

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<strong>Balanites</strong> <strong>aegyptiaca</strong><br />

heglig berries ( in the sudan), egyptian myrobalan, desert date (dried fruit)<br />

LOCAL NAMES<br />

Amharic (kudkuda,jemo,bedeno); Arabic (zachun,zaccone,heglig<br />

(tree),zacon,kuge,lalob (fruit)); Bemba (katikayengele,mubambwangoma);<br />

Bengali (hin); English (soap berry tree,simple-thorned torchwood,simple<br />

thorned torch tree,Jericho balsam,lalob tree,heglig,Egyptian<br />

myrobalan,desert date,torch wood); French (myrobalou d’Egypte,dattier<br />

sauvage,dattier du desert,myrobalau d' Egypte); Hindi<br />

(engua,ingudi,betu,hingan,hingn,hingot,hongot,hingota); Lozi (mwalabwe);<br />

Luganda (musongole); Mandinka (sumpo); Nyanja (nkuyu); Sanskrit<br />

(ingudi); Swahili (mjunju,mwambangoma); Tamil (nanjunda); Tigrigna<br />

(indrur,mekie); Tongan (mulyanzovu,mwalabwe); Trade name (desert date<br />

(dried fruit),heglig berries ( in the sudan),egyptian myrobalan)<br />

BOTANIC DESCRIPTION<br />

<strong>Balanites</strong> <strong>aegyptiaca</strong> is a multibranched, spiny shrub or tree up to l0 m<br />

high. Crown rounded, dense (but still seen through) with long stout<br />

branchlets. Trunk and bark grey, deeply fissured longitudinally.<br />

Leaves compound and spirally arranged on the shoots, dark green with 2<br />

firm coriaceous leaflets; dimensions and shapes varying widely. Petiole<br />

canaliculate, from 5 mm to 20 mm with a short rachis. Most accounts<br />

indicate a maximum length of 8 mm for Uganda. Margin of each leaflet<br />

entire; lamina generally up to 6 cm long, 4 cm broad, although apparently<br />

smaller (1-3 x 0.3-1.5 cm) in the Sahara and in Palestine.<br />

Inflorescence a sessile or shortly pedunculate fascicle of a few flowers.<br />

Flower buds ovoid and tomentose. Individual flowers hermaphroditic,<br />

pentamerous an actinomorphic, 8-14 mm in diameter and generally<br />

greenish-yellow. Pedicels densely greyish, pubescent and rarely reaching<br />

10 mm in length, although 15 mm is reported for Zambia and Zimbabwe.<br />

The usual length is about 8 mm.<br />

Fruit ellipsoid, up to 4 cm long, green. Ripe fruit brown or pale brown with<br />

a brittle coat enclosing a brown or brown-green sticky pulp and a hard<br />

stone seed.<br />

The name <strong>Balanites</strong> (from the Greek for acorn, referring to the fruit) was<br />

given in 1813 by Alire Delile and replaced Agialid (derived from the Arabic<br />

name for the tree, 'heglig').<br />

BIOLOGY<br />

Flowering behaviour varies. There is no definite time for flowering in the<br />

Sahel, although flowering most likely takes place in the dry season.<br />

Flowering in Nigeria varies between November and April with ripe fruits<br />

becoming available in December and January and occasionally later, from<br />

March to July. Elsewhere, fruiting and foliage production occur at the<br />

height of the dry season. Pollination is presumably by insects as flowers<br />

are scented, and flower structure facilitates insect activity. The 1st fruiting<br />

is at 5-8 years, yields increasing until 25 years of age for the tree.<br />

The fruit apparently takes at least 1 year to mature and ripen. Birds and<br />

mammals eat the fleshy and edible fruit, discarding, regurgitating or<br />

evacuating the stone.<br />

<strong>Agroforestry</strong> Database 4.0 (Orwa et al.2009)<br />

(L.) Del.<br />

<strong>Balanitaceae</strong><br />

Detail of leaves, thorns and flowers. (USAID)<br />

Tree in Machakos, Kenya. (Fernandes<br />

E.C.M)<br />

Detail of unripe fruits. (Fernandes E.C.M)<br />

Page 1 of 5


<strong>Balanites</strong> <strong>aegyptiaca</strong><br />

heglig berries ( in the sudan), egyptian myrobalan, desert date (dried fruit)<br />

(L.) Del.<br />

<strong>Balanitaceae</strong><br />

ECOLOGY<br />

B. <strong>aegyptiaca</strong> has wide ecological distribution; however, it reaches its maximum development as an individual tree on<br />

low-lying, level alluvial sites with deep sandy loam and uninterrupted access to water such as valley floors, riverbanks or<br />

the foot of rocky slopes. It is intolerant to shade after the seedling stage and therefore prefers open woodland or<br />

savannah for natural regeneration.<br />

BIOPHYSICAL LIMITS<br />

Altitude: 0-2 000 m, Mean annual temperature: 20 -30 deg. C, Mean annual rainfall: 250-1200 mm<br />

Soil type: The soils in its range tend to be deep sands, sandy clay loams, sandy loams or clays.<br />

DOCUMENTED SPECIES DISTRIBUTION<br />

Native:<br />

Exotic:<br />

Algeria, Angola, Benin, Burkina Faso, Burundi, Cameroon, Chad, Cote d'Ivoire, Democratic<br />

Republic of Congo, Djibouti, Egypt, Eritrea, Ethiopia, Gambia, Ghana, Guinea, India, Israel, Kenya,<br />

Libyan Arab Jamahiriya, Morocco, Myanmar, Nigeria, Saudi Arabia, Senegal, Somalia, Sudan,<br />

Tanzania, Uganda, Yemen, Republic of, Zambia, Zimbabwe<br />

Cape Verde, Dominican Republic, Puerto Rico<br />

Native range<br />

Exotic range<br />

The map above shows countries where the species has been planted. It does neither<br />

suggest that the species can be planted in every ecological zone within that country,<br />

nor that the species can not be planted in other countries than those depicted. Since<br />

some tree species are invasive, you need to follow biosafety procedures that apply to<br />

your planting site.<br />

<strong>Agroforestry</strong> Database 4.0 (Orwa et al.2009)<br />

Page 2 of 5


The map above shows countries where the species has been planted. It does neither<br />

suggest that the species can be planted in every ecological zone within that country,<br />

nor that the species can not be planted in other countries than those depicted. Since<br />

<strong>Balanites</strong> <strong>aegyptiaca</strong><br />

(L.) Del.<br />

some tree species are invasive, you need to follow biosafety procedures that apply to<br />

your planting site.<br />

<strong>Balanitaceae</strong><br />

heglig berries ( in the sudan), egyptian myrobalan, desert date (dried fruit)<br />

PRODUCTS<br />

Food: The fleshy pulp of both unripe and ripe fruit is edible and eaten dried or fresh. The fruit is processed into a drink<br />

and sweetmeats in Ghana, an alcoholic liquor in Nigeria, a soup ingredient in Sudan. Young leaves and tender shoots<br />

are used as a vegetable, which is boiled, pounded, then fried or fat added to prepare it. The flowers are a<br />

supplementary food in West Africa and an ingredient of ‘dawa dawa’ flavouring in Nigeria. Flowers are sucked to obtain<br />

nectar.<br />

Fodder: The fresh and dried leaves, fruit and sprouts are all eaten by livestock. As shown in an experiment in Burkina<br />

Faso, B. <strong>aegyptiaca</strong> contributed up to 38% of the dry-matter intake of goats in the dry season. Kernel meal, the residue<br />

remaining after oil extraction, is widely used in Senegal, Sudan and Uganda as a stock feed. The tree is lopped for<br />

fodder in India (Maharashta, Madhya Pradesh, Tamil Nadu and Rajasthan).<br />

Fuel: The wood is good firewood; it produces considerable heat and very little smoke, making it particularly suitable for<br />

indoor use. It produces high-quality charcoal, and it has been suggested that the nutshell is suitable for industrial<br />

activated charcoal. The calorific value is estimated at 4600 kcal/kg.<br />

Fibre: A strong fibre is obtained from the bark.<br />

Timber: The wood is pale yellow or yellowish-brown. Heartwood and sapwood are not clearly differentiated. The wood<br />

is hard, durable, worked easily and made into yokes, wooden spoons, pestles, mortars, handles, stools and combs. It<br />

shows no serious seasoning defects and no tendency towards surface checking or splitting. The wood saws cleanly and<br />

easily, planes without difficulty to a smooth finish and is easy to chisel. It glues firmly and takes a clear varnish. The<br />

timber has traditionally been a minor product. The usually small log size and the prevalence of stem fluting makes<br />

sawmill processing difficult.<br />

Gum or resin: A greenish-yellow to orange-red resin is produced from the stems. It is sucked and chewed when fresh.<br />

It is used as a glue for sticking feathers onto arrow shafts and spearheads and in the repair of handle cracks and arrows.<br />

Lipids: The kernels produce edible oil used for cooking. The oil remains stable when heated and has a high smoking<br />

point, and therefore its free fatty acid content is low. Its scent and taste are acceptable.<br />

Alcohol: The fruit of B. <strong>aegyptiaca</strong> may be used to brew an alcoholic drink.<br />

Poison: An emulsion made from the fruit or bark is lethal to the freshwater snails that are the host of miracidia and<br />

cercaria stages of bilharzia and to a water flea that acts as a host to the guinea worm. A fish poison can be obtained<br />

from the fruit, root and the bark. The active agent of the poison is saponin. The compound is toxic to fish but does not<br />

affect mammals and rapidly becomes inert, so that fish retrieved are edible. However, in the Fada region of Cote<br />

d’Ivoire, the poison is reported to damage the sight of fishermen after they have used it for 5-6 years.<br />

Medicine: Decoction of root is used to treat malaria. Roots boiled in soup are used against oedema and stomach pains.<br />

Roots are used as an emetic; bark infusion is used to treat heartburn. Wood gum mixed with maize meal porridge is<br />

used to treat chest pains. The bark is used to deworm cattle in Rajasthan.<br />

Other products: The seeds are used for rosary beads, necklaces and in the game of warri played in Sudan.<br />

SERVICES<br />

Shade or shelter: The usually evergreen behaviour potentially makes B. <strong>aegyptiaca</strong> an attractive element to introduce<br />

into shelterbelts, although because of its slow growth, it is not suitable as a principal species.<br />

Boundary or barrier or support: As a thorny tree, B. <strong>aegyptiaca</strong> is useful for fencing. Boundary and amenity plantings<br />

are widespread in Africa. Cut branches are used to make livestock enclosures.<br />

<strong>Agroforestry</strong> Database 4.0 (Orwa et al.2009)<br />

Page 3 of 5


<strong>Balanites</strong> <strong>aegyptiaca</strong><br />

heglig berries ( in the sudan), egyptian myrobalan, desert date (dried fruit)<br />

(L.) Del.<br />

<strong>Balanitaceae</strong><br />

TREE MANAGEMENT<br />

Coppices and pollards well and can regenerate after lopping and heavy browsing. Where fruit is the principal interest,<br />

pollarding and coppicing for obtaining fodder are seldom employed.<br />

GERMPLASM MANAGEMENT<br />

Seed storage behaviour is orthodox; viability can be maintained for 2 years in air-dry storage at cool temperatures or<br />

for several years in hermetic storage at 3 deg. C with 6-10% mc. One kilogram of cleaned, extracted seeds, air-dried to<br />

15% mc, contains 500-1500 seeds.<br />

PESTS AND DISEASES<br />

B. <strong>aegyptiaca</strong> suffers from repeated locust and beetle attack and a high degree of parasitic infestation (in Gountoure,<br />

Burkina Faso, 50% of the population had leaf galls, bugs or scales). Two microfungi, Phorma balanitis and Septoria<br />

balanitis, are the only reported dependent fungi.<br />

<strong>Agroforestry</strong> Database 4.0 (Orwa et al.2009)<br />

Page 4 of 5


<strong>Balanites</strong> <strong>aegyptiaca</strong><br />

heglig berries ( in the sudan), egyptian myrobalan, desert date (dried fruit)<br />

(L.) Del.<br />

<strong>Balanitaceae</strong><br />

FURTHER READNG<br />

Albrecht J. ed. 1993. Tree seed hand book of Kenya. GTZ Forestry Seed Center Muguga, Nairobi, Kenya.<br />

Anon. 1986. The useful plants of India. Publications & Information Directorate, CSIR, New Delhi, India.<br />

Baumer M. 1983. Notes on trees and shrubs in arid and semi-arid regions. Rome FAO. Forestry Division.<br />

Beentje HJ. 1994. Kenya trees, shrubs and lianas. National Museums of Kenya.<br />

Bein E. 1996. Useful trees and shrubs in Eritrea. Regional Soil Conservation Unit (RSCU), Nairobi, Kenya.<br />

Bekele-Tesemma A, Birnie A, Tengnas B. 1993. Useful trees and shrubs for Ethiopia. Regional Soil Conservation Unit<br />

(RSCU), Swedish International Development Authority (SIDA).<br />

Birnie A. 1997. What tree is that? A beginner's guide to 40 trees in Kenya. Jacaranda designs Ltd.<br />

Booth FEM, Wickens GE. 1988. Non-timber uses of selected arid zone trees and shrubs in Africa. FAO Conservation<br />

Guide. No. 19. Rome.<br />

Coates-Palgrave K. 1988. Trees of southern Africa. C.S. Struik Publishers Cape Town.<br />

Dale IR, Greenway PJ. 1961. Kenya trees and shrubs. Buchanan’s Kenya Estates Ltd.<br />

Eggeling. 1940. Indigenous trees of Uganda. Govt. of Uganda.<br />

Hall JB, Walker HD. 1991. <strong>Balanites</strong> <strong>aegyptiaca</strong>: a monograph. School of Agriculture and Forest Sciences Publication<br />

Number: 3. University of Wales, Bangor.<br />

Hines DA, Eckman K. 1993. Indigenous multipurpose trees for Tanzania: uses and economic benefits to the people.<br />

Cultural survival Canada and Development Services Foundation of Tanzania.<br />

Hocking D. 1993. Trees for Drylands. Oxford & IBH Publishing Co. New Delhi.<br />

Hong TD, Linington S, Ellis RH. 1996. Seed storage behaviour: a compendium. Handbooks for Genebanks: No. 4.<br />

IPGRI.<br />

IBPGR-Kew. 1984. Forage and browse plants for arid and semi-arid Africa. Rome. IBPGR.<br />

ICRAF. 1992. A selection of useful trees and shrubs for Kenya: Notes on their identification, propagation and<br />

management for use by farming and pastoral communities. ICRAF.<br />

Katende AB et al. 1995. Useful trees and shrubs for Uganda. Identification, Propagation and Management for<br />

Agricultural and Pastoral Communities. Regional Soil Conservation Unit (RSCU), Swedish International Development<br />

Authority (SIDA).<br />

Kitilit JK. 2003. Evaluation of Acacia seyal and <strong>Balanites</strong> <strong>aegyptiaca</strong> as sources of nutrients for ruminants. 78p.<br />

Mbuya LP et al. 1994. Useful trees and shrubs for Tanzania: Identification, Propagation and Management for<br />

Agricultural and Pastoral Communities. Regional Soil Conservation Unit (RSCU), Swedish International Development<br />

Authority (SIDA).<br />

Noad T, Birnie A. 1989. Trees of Kenya. General Printers, Nairobi.<br />

Sahni KC. 1968. Important trees of the northern Sudan. United Nations and FAO.<br />

Storrs AEG. 1995. Know your trees: some common trees found in Zambia. Regional Soil Conservation Unit (RSCU).<br />

Vogt K. 1995. A field guide to the identification, propagation and uses of common trees and shrubs of dryland Sudan.<br />

SOS Sahel International (UK).<br />

Weber F, Stoney C. 1986. Reforestation in arid lands. VITA Arlington, Virginia.<br />

SUGGESTED CITATION<br />

Orwa C, Mutua A , Kindt R , Jamnadass R, Simons A. 2009. Agroforestree Database:a tree reference and selection guide<br />

version 4.0 (http://www.worldagroforestry.org/af/treedb/)<br />

<strong>Agroforestry</strong> Database 4.0 (Orwa et al.2009)<br />

Page 5 of 5

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