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Ver. 19.2
April 17th, 2024

041.0000 Family Polypodiaceae J. & C.Presl; J. & C. Presl, Delic. Prag. 159 (1822)
67 genera, ca. 1700 spp. Phylogeny: Wei & Zhang 2022
Species listed in database: 1718; hybrids: 47
= Platyceriaceae Ching; Acta Phytotax. Sin. 16(3): 18 (1978)
= Drynariaceae Ching; Acta Phytotax. Sin. 16: 1-37 (1978)
= Grammitidaceae Newman; Hist. Brit. Ferns 7 (1840) (Grammitaceae auct., missp.)
= Gymnogrammitidaceae Ching; Acta Phytotax. Sin. 11: 12 (1966)
= Loxogrammaceae Ching ex Pic.Serm.; Webbia 29: 11 (1975)
= Platyceriaceae Ching; Sunyatsenia 5 (1940)
= Pleurosoriopsidaceae Sa.Kurata & Ikebe ex Ching; Acta Phytotax. Sin. 16(4): 17 (1978)
041.2001 Genus Pyrrosia Mirb.; Hist. Nat. Gen. Pl. 4: 70 (1803)
Distribution: Tropical Asia including Pacific. Cyclophorus, Drymoglossum and Saxiglossum are frequently treated as separate genera. Subgenera according to Vasques et al. (2017)
Species listed in database: 65; hybrids: 1
= Apalophlebia C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 137 (1851)
= Asteroglossum J.Sm.ms.; ex Moore, Ind. 179 (1860)
= Candollea Mirb.; Lam. & Mirb., Hist. Nat. Vég. Cl. Fam. 5: 86 (1802)
= Cyclophorus Desv.; Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 300 (1811)
= Digramma Kunze; Bot. Zeitung (Berlin) 209 (1848)
= Drymoglossum C.Presl; Tent. Pterid. 227 (1836)
= Galeoglossa C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 132 (1851)
= Gyrosorium C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 139 (1851)
= Neoniphopsis Nakai; Bot. Mag. (Tokyo) 42: 217 (1928)
= Nesoris Raf.; Fl. Tellur. 4: 104 (1838)
= Niphobolus Kaulf.; Enum. Filic. 124 (1824) (nom. superfl.)
= Niphopsis J.Sm.; Cat. Cult. Ferns 6 (1857)
= Oetosis Kuntze; Revis. Gen. Pl. 2: 817 (1891)
= Polycampium C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 135 (1851)
= Polypodium subgen.Niphidium (Lellinger) Proctor; R. A. Howard, Fl. Lesser Antilles 2: 343 (1977)
= Pteropsis Desv.; Mém. Soc. Linn. Paris 6: 218 (1827)
= Saxiglossum Ching; Contr. Inst. Bot. Nat. Acad. Peiping 2: 1 (1933)
= Scytopteris C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 133 (1851)
= Solenopteris Wall.; Tidsskr. f. Naturvid. 1: 259 (1822)
= Sphaerostichum C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 134 (1851)
species :
Pyrrosia abbreviata (Zoll. & Moritzi) Tagawa; Acta Phytotax. Geobot. 25: 180 (1973)
Distribution: Sumatra; Java; Lesser Sunda Isl. (Lombok, Bali, Sumbawa, Flores)
42 JAW LSI SUM
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Cyclophorus abbreviatus (Zoll. & Moritzi) C.Chr.; Index Filic. 197 (1905)
= Niphobolus abbreviatus Zoll.; Natuur- Geneesk. Arch. Ned.-Indië 1: 397 (1844)
= Polypodium abbreviatum (Zoll. & Moritzi) Mett.; Fil. Hort. Bot. Lips. 3 (1856)
= Pyrrosia abbreviata (Zoll. & Moritzi) Hovenkamp; Blumea 30(1): 208 (1984)
= Sphaerostichum abbreviatum (Zoll. & Moritzi) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 135 (1851)
species :
Pyrrosia adnascens (Sw.) Ching; Bull. Chin. Bot. Soc. 1: 45 (1935)
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Yunnan); Nepal; Bhutan; India (Arunachal Pradesh, Sikkim, Darjeeling, S-India, NE-India); Bangladesh; Thailand; Laos; Cambodia; Vietnam; Sumatra; Borneo (Sabah, Sarawak, Kalimantan); Vanuatu [I]
36 CHI 40 BAN BHU IND NEP 41 CBD LAO THA VIE 42 BOR SUM 60 van
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Cyclophorus adnascens (Sw.) Desv.; Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 300 (1811)
= Cyclophorus adnascens f. dichotomus Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 192 (1922)
= Cyclophorus adnascens f. pernudus Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 10 (1912)
= Cyclophorus adnascens var. minor Alderw.; Malayan Ferns 681 (1909)
= Craspedaria pertusa (Roxb. ex Hook.) Link; Fil. Spec. 118 (1841)
= Cyclophorus pustulosus Christ; Notul. Syst. (Paris) 1: 187 (1910)
= Niphobolus adnascens (Sw.) Kaulf.; Enum. Filic. 124 (1824) (pro parte)
= Niphobolus pertusus (Roxb. ex Hook.) Spreng.; Syst. Veg. ed. 16. 4(1): 44 (1827)
= Polypodium adnascens Sw.; Syn. 25: 222, t. 2, f. 2 (1806)
= Polypodium pertusum Roxb. ex Hook.; Exot. Fl. 2: t. 162 (1825)
= Pyrrosia adnascens f. calcicola K.H.Shing; J. Jap. Bot. 72(1): 28 (1997)
species :
Pyrrosia albicans (Blume) Ching; Bull. Chin. Bot. Soc. 1: 72 (1935)
Distribution: Thailand; Laos; Sumatra; Java; Lesser Sunda Isl. (Bali, Lombok, Sumbawa, Flores)
41 LAO THA 42 JAW LSI SUM
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Cyclophorus albicans (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 131 (1851)
= Cyclophorus blumeanus (Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 130 (1851)
= Cyclophorus elaphoglossoides Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 139 (1920)
= Cyclophorus flocciger (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 131 (1851)
= Cyclophorus lancifolius Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 23: 8 (1916)
= Cyclophorus nigropunctatus Rosenst.; Meded. Rijks Herb. no. 31: 4 (1917)
= Niphobolus albicans Blume; Enum. Pl. Javae 2: 107 (1828)
= Niphobolus blumeanus Kunze; Bot. Zeitung (Berlin) 120 (1848)
= Niphobolus flocciger Blume; Enum. Pl. Javae 2: 107 (1828)
= Niphobolus flocciger var. loriformis Kunze; (1848)
= Polypodium acrostichoides Willd.; Sp. Pl. ed. 4, 5: 156 (1810) (non G. Forst. (1786))
= Polypodium albicans (Blume) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 127, n. 257 (1857)
= Polypodium floccigerum (Blume) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 129, n. 262 (1857)
= Pyrrosia floccigera (Blume) Ching; Bull. Chin. Bot. Soc. 1: 71 (1935)
species :
Pyrrosia angustata (Sw.) Ching; Bull. Chin. Bot. Soc. 1: 49 (1935)
Distribution: S-Myanmar; peninsular Thailand; peninsular Malaysia (Penang, Perak, Selangor, Negeri Sembilan, Melaka, Terengganu, Pahang, Johor); Singapore; Sumatra; Borneo (Sabah); ?Java; Moluccas; ?Society Isl.; ?Gambier Isl. (?Mangareva); ?Austral Isl. (?Rurutu)
41 MYA THA 42 BOR MLY MOL SNG SUM
Remarks: subgen. 5 Niphopsis
Conservation Status: vulnerable (Singapore)
Synonyms: hidden
= Cyclophorus angustatus (Sw.) Desv.; Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 300 (1811)
= Cyclophorus micraster Copel.; Univ. Calif. Publ. Bot. 12: 405 (1931)
= Drynaria angustata (Sw.) Fée; Mém. Foug. 5 (Gen. Filic.): 270 (1852)
= Drynaria macrosora Fée; Mém. Foug. 5. (Gen. Filic.) 270 (1852)
= Niphobolus angustatus (Sw.) Spreng.; Syst. Veg. ed. 16, 4(1): 44 (1827)
= Niphobolus sphaerocephalus (Blume) Hook. & Grev.; Icon. Filic. t. 94 (1829)
= Niphopsis angustatus (Sw.) J.Sm.; Cat. Cult. Ferns 6 (1857)
= Phymatodes sphaerocephala (Blume) C.Presl; Tent. Pterid. 196 (1836)
= Pleopeltis angustata (Sw.) C.Presl; Tent. Pterid. 193 (1836)
= Pleopeltis sphaerocephala Blume; Fl. Javae, Fil. 64 (1828)
= Polypodium angustatum Sw.; Syn. Fil. 27, 224 (1806)
= Polypodium coriaceum Roxb.; Calcutta J. Nat. Hist. 4: 481, t. 28 (1844)
= Polypodium sphaerocephalum (Blume) Wall.; Numer. List no. 272 (1829), nomen
= Pyrrosia micraster (Copel.) Tagawa; J. Jap. Bot. 22: 163 (1949)
species :
Pyrrosia angustissima (Giesenh. ex Diels) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 26: 171 (1975)
Distribution: China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangxi, Shaanxi, Shanxi, Zhejiang); Japan; Taiwan; SW-Thailand
36 CHI 38 JAP TAI 41 THA
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Cyclophorus cavalerianus (Christ) C.Chr.; Index Filic. 198 (1905)
= Cyclophorus sasakii Hayata; Icon. Pl. Formosan. 6: 158, f. 61 (1916)
= Cyclophorus taenioides C.Chr.; Index Filic. 201 (1905)
= Niphobolus angustissimus Giesenh. ex Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 326 (1899), non Polypodium angustissimum Fée (1873)
= Niphobolus cavalerianus Christ; Bull. Acad. Int. Géogr. Bot., sér. 3, 13: 107 (1904)
= Pyrrosia angustissima (Giesenh. ex Diels) C.M.Kuo; Taiwania 30: 68 (1985)
= Saxiglossum angustissimum (Giesenh. ex Diels) Ching; Acta Phytotax. Sin. 10: 301 (1965)
= Saxiglossum sasakii (Hayata) Tagawa; J. Jap. Bot. 12: 753 (1936)
= Saxiglossum taeniodes (C.Chr.) Ching; Contr. Inst. Bot. Nat. Acad. Peiping 2: 1 (1933)
= Saxiglossum taeniodes var. sasakii (Hayata) Masam. ex Satomi; J. Geobot. 9: 3 (1960)
species :
Pyrrosia assimilis (Baker) Ching; Bull. Chin. Bot. Soc. 1 49 (1935)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Henan, Hunan, Jiangxi, Sichuan, Xinjiang, Yunnan, Zhejiang); Taiwan
36 CHI 38 TAI
Remarks: subgen. 3 Neoniphopsis. Not in India
Synonyms: hidden
= Cyclophorus assimilis (Baker) C.Chr.; Index Filic. 198 (1905)
= Cyclophorus assimilis f. lobatus Y.C.Wu, K.K.Wong & Pong; Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ. 3: 336, pl. 154 (1932)
= Cyclophorus assimilis var. mollifrons Hand.-Mazz.; Symb. Sin. 6: 46 (1929)
= Niphobolus assimilis (Baker) Diels; Bot. Jahrb. Syst. 29: 207 (1900)
= Polypodium assimilis Baker; J. Bot. 13: 1875: 201 (1875)
= Pyrrosia assimilis f. lobata (Y.C.Wu, K.K.Wong & Pong) Ching; Bull. Chin. Bot. Soc. 1: 50 (1935)
= Pyrrosia assimilis var. longissima Ching; Lingnan Sci. J. 33 (1945)
= Pyrrosia dimorpha X.H.Guo & S.B.Zhou; Bull. Bot. Res., Harbin 25: 389 (2005), not (Copeland) Parris (1980)
species :
Pyrrosia asterosora (Baker) Hovenkamp; Blumea 30(1): 208 (1984)
Distribution: Sumatra
42 SUM
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Cyclophorus asterosorus (Baker) C.Chr.; Index Filic. 198 (1905)
= Polypodium asterosorum Baker; J. Bot. 18: 214 (1880)
species :
Pyrrosia avaratra Rakotondr. & Hovenkamp; Novon 22(1): 75 (2012)
Distribution: Madagascar
29 MDG
Remarks: subgen. 6 Solis
species :
Pyrrosia beddomei (C.B.Clarke) Hovenkamp ex Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 3: 255 (2020)
Distribution: S-India
40 IND
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Drymoglossum beddomei C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 576 (1880)
= Drymoglossum heterophyllum var. beddomei (C.B.Clarke) C.Chr.; Dansk Bot. Ark. 6: 85 (1929)
= Drymoglossum piloselloides var. beddomei (C.B.Clarke) Bedd.; Handb. Ferns Brit. India: 413 (1880)
species :
Pyrrosia bonii (Christ) Ching; Bull. Chin. Bot. Soc. 1: 67 (1935)
Distribution: China (Guangxi, Guizhou); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Cyclophorus bonii (Christ) C.Chr.; Index Filic. 198 (1905)
= Niphobolus bonii Christ ex Giesenh.; Farngat. Niphobolus: 120 (1901)
= Pyrrosia shennongensis K.H.Shing; J. Jap. Bot. 72(2): 73 (1997)
= Pyrrosia subtruncata Ching; Bull. Chin. Bot. Soc. 1: 67 (1935)
species :
Pyrrosia boothii (Hook.) Ching; Bull. Chin. Bot. Soc. 1 66 (1935)
Distribution: China (Yunnan); Tibet; Bhutan; India (Arunachal Pradesh, Sikkim); Myanmar
36 CHI TIB 40 BHU IND 41 MYA
Remarks: subgen. 3 Neoniphopsis
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cyclophorus boothii (Hook.) C.Chr.; Index Filic. 198 (1905)
= Cyclophorus subvelutinus (Christ) C.Chr.; Index Filic. 201 (1905)
= Niphobolus boothii (Hook.) Bedd.; Handb. Ferns Brit. India, t. 258 (1868)
= Niphobolus subvelutinus Christ; Annuaire Conserv. Jard. Bot. Genève 7-8: 331 (1904)
= Polypodium boothii Hook.; Sp. Fil. 5: 53 (1863)
= Pyrrosia fengiana Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 73 (1941)
= Pyrrosia subvelutina (Christ) Ching; Bull. Chin. Bot. Soc. 1: 50 (1935)
species :
Pyrrosia calvata (Baker) Ching; Bull. Chin. Bot. Soc. 1: 62 (1935)
Distribution: China (Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Shaanxi, Sichuan, Yunnan, Zhejiang)
36 CHI
Synonyms: hidden
= Cyclophorus calvatus (Baker) C.Chr.; Index Filic. 198 (1905)
= Cyclophorus esquirolii H.Lév.; Flore du Kouy-tcheou 478 (1915)
= Niphobolus calvatus (Baker) Diels; Bot. Jahrb. Syst. 29: 207 (1900)
= Polypodium calvatum Baker; J. Bot. 17: 304 (1879)
= Pyrrosia pseudocalvata Ching; Boufford & K. H. Shing, J. Arnold Arbor. 64(1): 38 (1983)
species :
Pyrrosia ceylanica (Giesenh.) Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 2: 134 (1960)
Distribution: S-India (Karnataka, Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
Remarks: subgen. 4 Niphobolus
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cyclophorus ceylanicus (Giesenh.) C.Chr.; Index Filic. 198 (1905)
= Niphobolus ceylanicus Giesenh.; Farngatt. Niphobolus 216, f. 19 (1901)
species :
Pyrrosia christii (Giesenh.) Ching; Bull. Chin. Bot. Soc. 1: 58 (1935)
Distribution: Borneo (Mt. Kinabalu, E-Kalimantan); Brunei
42 BOR BRN
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Cyclophorus christii (Giesenh.) C.Chr.; Index Filic. 198 (1905)
= Niphobolus christii Giesenh.; Farngatt. Niphobolus 140 (1901)
species :
Pyrrosia confluens (R.Br.) Ching; Bull. Chin. Bot. Soc. 1: 49 (1935)
Name: Robber fern
Distribution: Australia (Queensland, NE-New South Wales); Lord Howe Isl.; Norfolk Isl.; New Caledonia; Vanuatu; Society Isl. (Tahiti); Austral Isl. (Tubuai)
50 AUS LHI NFK 60 NWC VAN 61 AUI SCI
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Cyclophorus confluens (R.Br.) C.Chr.; Index Filic. 198 (1905)
= Cyclophorus scytopteris (Fée) E.Fourn.; Ann. Sci. Nat., Bot., sér. V, 18: 286 (1873)
= Cyclophorus spicatus Domin; Biblioth. Bot. 20: 189 (1913)
= Drymoglossum cunninghamii T.Moore; Index Filic. (T. Moore) 31 (1857)
= Niphobolus acrostichoides Kaulf.; Enum. Filic. 127 (1824)
= Niphobolus confluens (R.Br.) Spreng.; Syst. Veg. ed. 16, 4(1): 44 (1827)
= Niphobolus scytopteris Fée; Mém. Foug. 5 (Gen. Filic.): 262 (1852)
= Niphobolus spicatus Domin; Biblioth. Bot. 20: 189 (1913), pro syn.
= Pleopeltis glabra (Mett.) E.Fourn.; Bull. Soc. Bot. France 16: 424 (1869)
= Polypodium confluens R.Br.; Prodr. Fl. Nov. Holland. 146 (1810)
= Polypodium confluens var. lobatum F.M.Bailey; Proc. Roy. Soc. Queensland 1: 15 (1885)
= Polypodium glabrum Mett.; Abh. Senckenberg. Naturf. Ges. 2: 123, n. 245, t. 3, f. 6-8 (1857) (non Burm. (1768))
= Polypodium rupestre Hook.; Sp. Fil. 5: 46 (1863), nom. illeg.
= Polypodium spicatum (Domin) F.M.Bailey; Queensl. Dept. Agric. Stock, Bot. Bull. (1913)
= Scytopteris acrostichoides (Kaulf.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 134 (1851)
variant :
var. dielsii (C.Chr.) Hovenkamp; Blumea 30(1): 208 (1984)
Name: Silvery Felt fern
Distribution: Australia (NE-Queensland, Atherton Tableland)
50 AUS
Synonyms: hidden
= Cyclophorus dielsii C.Chr.; Dansk Bot. Ark. 9(3) 57, t. 6(2) (1937)
= Cyclophorus intermedius Goy; Queensl. Nat. 10: 48, t. 6 (1937)
= Pyrrosia intermedia (Goy) K.H.Shing; Amer. Fern J. 73(3): 78 (1983)
= Polypodium serpens var. grande F.M.Bailey; Queensl. Dept. Agric. Stock, Bot. Bull. (1913)
= Pyrrosia dielsii (C.Chr.) Tindale; Contr. New South Wales Natl. Herb. Flora Ser. 208-211: 32 (1961)
species :
Pyrrosia costata (Wall. ex C.Presl) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 22: 100 (1967)
Distribution: China (Yunnan); Tibet; Nepal; Bhutan; N-India (Uttarakhand, Darjeeling, Sikkim, Meghalaya, Garo, Khasia Hills, Bihar, Orissa, Tamil Nadu, Kerala, Arunachal Pradesh); Bangladesh; Myanmar (Kachin, etc.); N-Thailand; Vietnam
36 CHI TIB 40 BAN BHU IND NEP 41 MYA THA VIE
Remarks: subgen. 1 Pyrrosia
Synonyms: hidden
= Apalophlebia costata (Wall. ex C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 138 (1851)
= Cyclophorus beddomeanus (Giesenh.) C.Chr.; Index Filic. 198 (1905)
= Niphobolus beddomeanus f. fallax Giesenh.
= Niphobolus beddomeanus Giesenh.; Farngatt. Niph. 101 (1901), nom. superfl.
= Niphobolus costatus Wall. ex C.Presl; Tent. Pterid.: 200 (1836)
= Niphobolus costatus C.Presl ex Bedd.; Handb. Ferns Brit. India, t. 120 (1865)
= Polypodium costatum (Wall. ex C.Presl) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 131 (1856), nom. illeg.
= Pyrrosia beddomeana (Giesenh.) Ching; Bull. Chin. Bot. Soc. 1: 68 (1935), nom. superfl.
species :
Pyrrosia davidii (Baker) Ching; Acta Phytotax. Sin. 10: 301 (1965)
Distribution: China (Gansu, Guizhou, Hebei, Henan, Hubei, Hunan, Liaoning, Nei Mongol, Shaanxi, Shandong, Shanxi, Sichuan, Yunnan); Tibet; South Korea (Cheju Isl.)
36 CHI TIB 38 KOR
Remarks: subgen. 3 Neoniphopsis. Kept separate from P. porosa by Flora of China
Synonyms: hidden
= Cyclophorus davidii (Baker) H.Lév.; Flore du Kouy-tcheou 478 (1915)
= Cyclophorus davidii (Baker) Hand.-Mazz.; Symb. Sin. 6: 46 (1929), isonym
= Cyclophorus gralla (Giesenh.) C.Chr.; Index Filic. 199 (1905)
= Cyclophorus pekinensis C.Chr.; Index Filic. 200 (1905)
= Cyclophorus pekinensis f. minor C.Chr.; Acta Hort. Gothob. 1: 105 (1924)
= Cyclophorus subfissus Hayata; Icon. Pl. Formosan. 6: 264 (1916)
= Cyclophorus transmorrisonensis Hayata; Icon. Pl. Formosan. 4: 256, f. 180 (1914)
= Niphobolus davidii (Baker) Giesenh. ex Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 325 (1899), not Franch. (1887)
= Niphobolus gralla Giesenh.; Farngat. Niphobolus: 128 (1901)
= Niphobolus transmorrisonensis Hayata; Icon. Pl. Formosan. 4: 257 (1914), nom. altern.
= Polypodium davidii Baker; Ann. Bot. (Oxford) 5: 572 (1891) (non Franch. (1887))
= Pyrrosia gralla (Giesenh.) Ching; Bull. Chin. Bot. Soc. 1: 50 (1935)
= Pyrrosia nudicaulis Ching; Bull. Chin. Bot. Soc. 1: 52 (1935)
= Pyrrosia pekinensis (C.Chr.) Ching; Bull. Chin. Bot. Soc. 1: 51 (1935)
= Pyrrosia subfissa (Hayata) Ching; Bull. Chin. Bot. Soc. 1: 72 (1935)
= Pyrrosia transmorrisonensis (Hayata) Ching; Bull. Chin. Bot. Soc. 1: 72 (1935)
species :
Pyrrosia distichocarpa (Mett.) K.H.Shing; Amer. Fern J. 73(3): 78 (1983)
Distribution: Sumatra
42 SUM
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Cyclophorus distichocarpus (Mett.) C.Chr.; Index Filic. 198 (1905)
= Cyclophorus grandis Ridl.; J. Malayan Branch Roy. Asiat. Soc. 1: 112 (1923)
= Cyclophorus winkleri Rosenst.; Repert. Spec. Nov. Regni Veg. 7: 149 (1909)
= Niphobolus distichocarpus (Mett.) Giesenh. ex Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 325 (1899)
= Polypodium distichocarpum Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 231 (1866)
= Pyrrosia winkleri (Rosenst.) Tagawa; Acta Phytotax. Geobot. 25: 180 (1973)
species :
Pyrrosia drakeana (Franch.) Ching; Bull. Chin. Bot. Soc. 1: 65 (1935)
Distribution: China (Gansu, Guangxi, Guizhou, Henan, Hubei, Shaanxi, Sichuan, Yunnan); Tibet; India (Arunachal Pradesh)
36 CHI TIB 40 IND
Remarks: subgen. 3 Neoniphopsis
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cyclophorus drakeanus (Franch.) C.Chr.; Index Filic. 198 (1905)
= Niphobolus drakeanus (Franch.) Diels; Bot. Jahrb. Syst. 29: 206 (1900)
= Niphobolus drakeanus f. elongatus Christ ex Diels; Bot. Jahrb. Syst. 29: 207 (1900)
= Polypodium drakeanum Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 7: 165 (1883)
= Pyrrosia pseudodrakeana K.H.Shing; Vasc. Pl. Hengduan Mount. 1: 170 (1993), without latin descr.
species :
Pyrrosia eberhardtii (Christ) Ching; Bull. Chin. Bot. Soc. 1: 59 (1935)
Distribution: China (Guangdong, Hainan); Thailand; Vietnam
36 CHI 41 THA VIE
Remarks: frequently included in P. heteractis but kept separate by Flora of China
Synonyms: hidden
= Cyclophorus eberhardtii Christ; J. Bot. (Morot) 21: 237, 270 (1908)
= Pyrrosia oblonga Ching; Bull. Chin. Bot. Soc. 1: 58 (1935)
species :
Pyrrosia elaeagnifolia (Bory) Hovenkamp; Blumea 30(1): 208 (1984), as ´eleagnifolia´
Name: Leather-leaf fern
Distribution: New Zealand (Three Kings Isl., North Isl., South Isl., Stewart Isl.); Kermadec Isl.; Chatham Isl.
51 CTM KER NZL
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Polypodium elaeagnifolium Bory; Dup., Voy. Bot. 1: 259, t. 31, f. 1 (1828), as ´eleagnifolium´
= Polypodium rupestre var. sinuatum Colenso; Trans. & Proc. New Zealand Inst. 17: 257 (1885)
species :
Pyrrosia ensata Ching & K.H.Shing; J. Jap. Bot. 72(1): 28 (1997)
Distribution: China (Yunnan); Tibet
36 CHI TIB
Remarks: subgen. 5 Niphopsis
species :
Pyrrosia fallax (Alderw.) M.G.Price; Kalikasan 3: 177 (1974)
Distribution: Moluccas (Ambon); New Guinea (Papua New Guinea); Bismarck Arch. (New Hanover); Solomon Isl. (Bougainville, Kolombangara, Makira, Malaita)
42 MOL NWG 43 BIS SOL
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Drymoglossum fallax Alderw.; Philipp. J. Sci., Bot. 11: 111, t. 6 (1916)
= Drymoglossum schlechteri Hieron. & Brause; Bot. Jahrb. Syst. 66: 177 (1920)
species :
Pyrrosia flocculosa (D.Don) Ching; Bull. Chin. Bot. Soc. 66 (1935)
Distribution: China (Yunnan, Guangxi); Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Siliguri, Meghalaya, Khasia Hills, Jaintla Hills, Shillong, Arunachal Pradesh); Bangladesh (Silhet); Myanmar (Kachin, etc.); N-Thailand; Vietnam
36 CHI 40 BAN BHU IND NEP 41 MYA THA VIE
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Apalophlebia flocculosa (D.Don) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 139 (1851)
= Cyclophorus annamensis (Christ) C.Chr.; Index Filic. 198 (1905)
= Cyclophorus flocculosus (D.Don) C.Chr.; Index Filic. 199 (1905)
= Cyclophorus rhomboidalis Bonap.; Notes Ptéridol. 7: 126, 191 (1918)
= Niphobolus annamensis Christ; J. Bot. (Morot) 19: 74 (1905)
= Niphobolus detergibilis (Hook.) Bedd.; Ferns Brit. India: t. 162 (1866)
= Niphobolus flocculosus (D.Don) Bedd.; Handb. Ferns Brit. India, t. 162 (1867)
= Niphobolus flocculosus (D.Don) Spreng.; Syst. Veg. ed. 16, 4(1): 45 (1827)
= Polypodium detergibile Hook.; Sp. Fil. 5: 49 (1863)
= Polypodium flocculosum D.Don; Prodr. Fl. Nepal. 1 (1825)
= Polypodium tomentosum Roxb.; Calcutta J. Nat. Hist. 4: 483 (1844), nom. illeg.
= Pyrrosia annamensis (Christ) Li Bing Zhang, X.M.Zhou & C.W.Chen; Phytotaxa 309(1): 90 (2017)
= Pyrrosia mollis f. rhomboidalis (Bonap.) C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 206 (1939)
= Pyrrosia rhomboidalis (Bonap.) Ching; Bull. Chin. Bot. Soc. 1: 72 (1935)
species :
Pyrrosia foveolata (Alston) C.V.Morton; Contr. U.S. Nat. Herb. 38: 234 (1973)
Distribution: Moluccas; New Guinea (Irian Jaya, Papua New Guinea, Goodenough Isl.); Bismarck Arch. (New Ireland)
42 MOL NWG 43 BIS
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Cyclophorus brassii Copel.; Univ. Calif. Publ. Bot. 18: 226 (1942)
= Cyclophorus foveolatus Alston; J. Bot. 78: 226 (1940) (Oct.)
= Pyrrosia brassii (Copel.) Pic.Serm.; Webbia 31(1): 249 (1977)
variant :
var. lauterbachii (Christ) Hovenkamp; Blumea 30(1): 208 (1984)
Distribution: New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.
42 NWG 43 BIS
Synonyms: hidden
= Cyclophorus dispar Christ; Nova Guinea 8: 155 (1909)
= Cyclophorus lauterbachii (Christ) C.Chr.; Index Filic. 199 (1905)
= Niphobolus lauterbachii Christ; K. Schum. & Lauterb., Fl. Schutzgeb. Südsee 142 (1901)
= Pyrrosia dispar (Christ) K.H.Shing; Amer. Fern J. 73(3): 78 (1983)
= Pyrrosia lauterbachii (Christ) Ching; Bull. Chin. Bot. Soc. 1: 49 (1935)
species :
Pyrrosia gardneri (Mett.) Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 2: 134 (1960)
Distribution: Sri Lanka
40 SRL
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Cyclophorus gardneri (Mett.) C.Chr.; Index Filic. 199 (1905)
= Niphobolus gardneri (Mett.) Kunze ex J.Sm.; Cat. Cult. Ferns: 12 (1857)
= Polypodium gardneri Mett.; Abh. Senckenberg. Naturf. Ges. 2: 129 (1856)
species :
Pyrrosia glabra (Desv.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 3: 258-259 (2020), et Indian J. Forestry 43(1): 82 (2021)
Distribution: China (Yunnan, Hainan, Sichuan); Nepal; Bhutan; India (Darjeeling, Sikkim, Arunachal Pradesh, Manipur, Meghalaya, S-India); Bangladesh; Myanmar; Laos; Philippines
36 CHI 40 BAN BHU IND NEP 41 LAO MYA 42 PHI
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Cyclophorus glaber Desv.; Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 301 (1811)
= Cyclophorus nudus (Giesenh.) C.Chr.; Index Filic. 200 (1905)
= Cyclophorus varius (Kaulf.) Gaudich.; Voy. Uranie [Freycinet] 364 (1827)
= Niphobolus adnascens var. varius (Kaulf.) Keyserl.; Polyp. Herb. Bunge 37 (1873)
= Niphobolus glaber (Desv.) Kaulf.; Enum. Filic. 127 (1824) (non Cyclophorus Desv. (1811))
= Niphobolus nudus Giesenh.; Farngatt. Niph. 149 (1901)
= Niphobolus varius Kaulf.; Enum. Filic. 125 (1824)
= Polypodium varium (Kaulf.) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 126 (1856), nom. illeg., non L. (1753)
= Pyrrosia nuda (Giesenh.) Ching; Bull. Chin. Bot. Soc. 1: 70 (1935)
= Pyrrosia varia (Kaulf.) Farw.; Amer. Midl. Naturalist 12: 302 (1931)
species :
Pyrrosia hastata (Thunb. ex Houtt.) Ching; Bull. Chin. Bot. Soc. 1: 48 (1935)
Distribution: China (Anhui); North Korea; South Korea (incl. Cheju Isl.); Japan
36 CHI 38 JAP KOR NKO
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Acrostichum hastatum Thunb. ex Houtt.; Natuurl. Hist. 14: 68, pl. 95, f. 2 (1783)
= Cyclophorus hastatus (Thunb. ex Houtt.) C.Chr.; Index Filic. 199 (1905)
= Niphobolus hastatus (Thunb. ex Houtt.) Kunze; Bot. Zeitung (Berlin) 6: 505 (1848)
= Niphobolus tricuspis (Sw.) J.Sm.; Hist. Fil. 98 (1875)
= Polycampium hastatum (Thunb. ex Houtt.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 137 (1851)
= Polypodium tricuspe Sw.; J. Bot. (Schrader) 1800(2): 21 (1801)
= Pyrrosia tricuspis (Sw.) Tagawa; J. Jap. Bot. 32: 354 (1957)
species :
Pyrrosia heteractis (Mett.) Ching; Bull. Chin. Bot. Soc. 1: 57 (1935)
Distribution: China (Guangxi, Hainan, Sichuan, Yunnan); Tibet; Nepal; Bhutan; N-India (Arunacha. Pradesh, Sikkim, Darjeeling, NE-India); Myanmar (Kachin, etc.); Thailand; Laos; Cambodia; Vietnam
36 CHI TIB 40 BHU IND NEP 41 CBD LAO MYA THA VIE
Remarks: subgen. 5 Niphopsis. P. eberhardtii is frequently included here, but kept separate by Flora of China, see there
Synonyms: hidden
= Cyclophorus heteractis (Mett.) C.Chr.; Index Filic. 99 (1905)
= Cyclophorus heteractis var. minor C.Chr.; Contr. U.S. Natl. Herb. 26: 335 (1931)
= Niphobolus heteractis (Mett.) J.Sm.; Ferns Brit. For. ed. II: 296 (1877)
= Polypodium heteractis Mett.; Kuhn, Linnaea 36: 140 (1869)
= Pyrrosia fuohaiensis Ching & K.H.Shing; J. Jap. Bot. 72(2): 76, f. 1(7-9) (1997)
= Pyrrosia heteractis var. minor (C.Chr.) Ching; Bull. Chin. Bot. Soc. 1: 58 (1935)
= Pyrrosia latifolia Ching & S.K.Wu; Fl. Xizangica 1: 340 (1983)
= Pyrrosia lingua var. heteractis (Mett.) Hovenkamp; Blumea 30(1): 208 (1984)
species :
Pyrrosia heterophylla (L.) M.G.Price; Kalikasan 3: 177 (1974)
Distribution: Seychelles; S-India; Sri Lanka
29 SEY 40 IND SRL
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Acrostichum heterophyllum L.; Sp. Pl. 2: 1067 (1753)
= Drymoglossum ellipticum (Willd.) T.Moore; Index Filic. (T. Moore) 31 (1857)
= Drymoglossum heterophyllum (L.) C.Chr.; Index Filic. 246 (1905), isonym
= Drymoglossum heterophyllum (L.) Trimen; J. Linn. Soc., Bot. 24: 152 (1887)
= Elaphoglossum heterophyllum (L.) Keyserl.; (1873)
= Lemmaphyllum heterophyllum (L.) Luerss.; (1882)
= Notholaena heterophylla (L.) Kaulf.; (1824)
= Oetosis heterophylla (L.) Kuntze; (1891)
= Pteris ceilanica Wikstr.; Vet. Akad. Handl. 1825: 443 (1826)
= Pteris elliptica Willd.; Sp. Pl. ed. 4, 5: 356 (1810) (non Poir. (1804))
= Pteropsis elliptica (Willd.) Desv.; Mem. Soc. Linn. Paris 6, 2: 218 (1827)
= Pteropsis heterophylla (L.) Desv.; (1827)
= Pteropsis nummularia Desv.; Mem. Soc. Linn. Paris 6, 2: 218 (1827)
= Taenitis heterophylla (L.) R.Br.; (1810) ex Mett. (1856)
species :
Pyrrosia kinabaluensis Hovenkamp; Blumea 30(1): 207 (1984)
Distribution: Borneo (Mt. Kinabalu, Sabah, Sarawak, Kalimantan)
42 BOR
Remarks: subgen. 5 Niphopsis
species :
Pyrrosia laevis (J.Sm. ex Bedd.) Ching; Bull. Chin. Bot. Soc. 1: 52 (1935)
Distribution: China (Yunnan); Tibet; NE-India (Meghalaya); Myanmar (Kachin, etc.)
36 CHI TIB 40 IND 41 MYA
Remarks: subgen. 5 Niphopsis
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cyclophorus laevis (J.Sm. ex Bedd.) C.Chr.; Index Filic. 199 (1905)
= Niphobolus laevis J.Sm. ex Bedd.; Ferns Brit. For.: t. 161 (1866)
= Polypodium breve J.Sm.; Ann. Bot. (Oxford) 5: 472 (1891), orth. var.
= Polypodium jaintense C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 552, t. 82, f. 4 (1880)
= Polypodium laeve (J.Sm. ex Bedd.) Mett.; Linnaea 36: 139 (1869)
= Pyrrosia jaintensis (C.B.Clarke) Ching; Fl. Xizangica 1: 335 (1983) (fide Fraser-Jenkins 1997)
species :
Pyrrosia lanceolata (L.) Farw.; Amer. Midl. Naturalist 12: 245 (1931)
Distribution: China (Yunnan); Tibet; Ryukyu Isl.; Taiwan; Nepal; Bhutan; India (Uttarakhand, Sikkim, Darjeeling, Madhya Pradesh, Orissa, Tamil Nadu, Kerala, Arunachal Pradesh); Sri Lanka; Bangladesh; Andaman Isl.; Nicobar Isl.; Myanmar (Kachin, etc.); Thailand (widespread); Laos; Cambodia; Vietnam; peninsular Malaysia (widespread); Singapore; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak, Kalimantan); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok, Sumba, Sumbawa, Flores, Wetar, Timor, Alor); Moluccas (Seram, Ambon, etc.); Christmas Isl. [Austr.]; New Guinea (Irian Jaya, Papua New Guinea, Mapia Atoll); Bismarck Arch. (New Hanover, Admiralty Isl.); Solomon Isl. (Bougainville, Choiseul, Guadalcanal, Santa Ysabel, Kolombangara, Makira, Mono, Rendova, Tulagi, Ulawa, Malaita, Ranongga, Rennell, Tetepare, Vella Lavella); Australia (NE-Queensland); Vanuatu; Tonga (Tongatapu, ´Eua, Vava´u, Tafahi); Fiji (Viti Levu, Vanua Levu, Taveuni, Waya); Wallis & Futuna (Alofi, Futuna, ´Uvea, Wallis); American Samoa (Tutuila, Aunu´u, Ofu, Olosega, Ta´u); [Western] Samoa (Aleipata Isl., Savai´i, ´Upolu); Palau (Babeldaob, Ngaturel, Koror, Ngarakabesang, Ulebsechel, Urukthapel, Ngeanges, Tobi); Northern Marianas (Rota Isl., Saipan, Tinian); Guam; Martinique [I]
36 CHI TIB 38 NNS TAI 40 BAN BHU IND NEP SRL 41 AND CBD LAO MYA NCB THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SNG SUL SUM XMS 43 BIS SOL 50 AUS 60 ASA FIJ SAM TON VAN WAL 62 GUM MRN PLU 81 mar
Remarks: subgen. 4 Niphobolus. P. adnascens is kept separate by Flora of China but included here by Knapp (Ferns of Taiwan) and Fraser-Jenkins
Synonyms: hidden
= Acrostichum dubium Poir.; Lam., Encycl. suppl. 1: 120 (1810)
= Acrostichum lanceolatum L.; Sp. Pl. 2: 1067 (1753)
= Acrostichum linnaeanum (Fée) Hook.; 2. Cent. Ferns, t. 26 (1861)
= Bolbitis linnaeana (Fée) C.Chr.; Index Filic., Suppl. Tert.: 198 (1934)
= Campium linnaeanum (Fée) Copel.; Philipp. J. Sci. 37: 343 (1928)
= Candollea heterophylla Mirb.; Lam. & Mirb., Hist. Nat. Vég. Cl. Fam. 5: 87 (1802) (non Acrostichum heterophyllum L. (1753))
= Candollea lanceolata (L.) Mirb.; Lam. & Mirb., Hist. Nat. Vég. Cl. Fam. 5: 89 (1802)
= Chrysodium linnaeanum (Fée) Kuhn; Ann. Mus. Bot. Lugduno-Batavi 4: 293 (1869)
= Chrysodium linnaeanum var. sinuatum Kuhn; (1869)
= Cyclophorus cornutus Copel.; Brittonia 1: 77 (1931)
= Cyclophorus dimorphus Copel.; J. Arnold Arbor. 10: 180 (1929)
= Cyclophorus giesenhagenii (Christ) C.Chr.; Index Filic. 199 (1905)
= Cyclophorus heterophyllus (Mirb.) Desv.; Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 301 (1811)
= Cyclophorus lanceolatus (L.) Alston; J. Bot. 69: 102 (1931)
= Cyclophorus pachydermus (Baker) C.Chr.; Index Filic. 200 (1905)
= Cyclophorus stellatus Copel.; J. Arnold Arbor. 10: 179 (1929)
= Cyclophorus tener (Giesenh.) C.Chr.; Index Filic. 201 (1905)
= Cyclophorus varius var. flabelliformis Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 57 (1914)
= Cyclophorus vittarioides C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 129 (1851)
= Dendroglossa lanceolata (L.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 149 (1851)
= Dendroglossa linnaeana (Hook.) Fée; Mém. Foug. 5. (Gen. Filic.) 81 (1852)
= Drymoglossum martinicense Christ; Bot. Jahrb. Syst. 24: 137 (1897)
= Gymnopteris lanceolata (L.) T.Moore; Index Filic. (T. Moore) 10 (1857)
= Gymnopteris linnaeana (Hook.) Christ; J. Bot. (Morot) 19: 125 (1905)
= Leptochilus heteroclitus var. linnaeanus (Hook.) Christ; Philipp. J. Sci., Bot. 2: 160 (1907)
= Leptochilus lanceolatus (C.Presl) Zoll.; Syst. Verz. 33 (1854) (non Fée (1845))
= Leptochilus linnaeanus Fée; Mém. Foug. 2 [Hist. Acrostich.]: 87 (1845), nom. superfl.
= Niphobolus carnosus Blume; Enum. Pl. Javae 2: 105 (1828)
= Niphobolus caudatus Kaulf.; Epim. bot. 127 (1824)
= Niphobolus chamissoanus C.Presl; Tent. Pterid.: 127 (1836)
= Niphobolus elongatus Blume; Enum. Pl. Javae 2: 105 (1828)
= Niphobolus giesenhagenii Christ; Annuaire Conserv. Jard. Bot. Genève 7-8: 330 (1904)
= Niphobolus heterophyllus (Mirb.) Spreng.; Syst. Veg. ed. 16, 4(1): 44 (1827)
= Niphobolus koenigii Blume; Fl. Javae, Fil. 50 (1828)
= Niphobolus lanceolatus (L.) Trimen; J. Linn. Soc., Bot. 24: 152 (1886), non Keyserl. (1873)
= Niphobolus spathulifer Bory; Bél., Voy. Indes Or., Bot. 2: 24 (1833)
= Niphobolus tener Giesenh.; Die Farngattung Niphobolus 211 (1901)
= Niphobolus vittarioides (C.Presl) T.Moore; Index Filic. (T. Moore) 276 (1861), non C. Presl 1836
= Polypodium carnosum (Blume) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 124 (1856)
= Polypodium carnosum var. elongatum (Blume) Mett.; Abh. Senckenb. Naturf. Ges. (Polypodium) 124 (1856)
= Polypodium caudatum (Kaulf.) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 125, n. 255 (1857) (non Raddi (1819))
= Polypodium dubium (Poir.) Kuhn; Filic. Afr. 147 (1868), not Roxburgh (1844), nor (H. H. Karst.) Hook. (1864)
= Polypodium koenigii Blume; Fl. Javae, Fil. 50 (1828)
= Polypodium pachydermum Baker; J. Linn. Soc., Bot. 15: 108 (1876)
= Polypodium vittarioides (C.Presl) Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2: 256 (1858)
= Polypodium vittarioides Wall.; Numer. List no. 270 (1829), nomen
= Pteropsis martinicensis (Christ) Maxon; Contr. U.S. Natl. Herb. 16: 51 (1912)
= Pyrrosia caudata (Kaulf.) Ching; Bull. Chin. Bot. Soc. 1: 46 (1935)
= Pyrrosia cornuta (Copel.) Tagawa; J. Jap. Bot. 22: 163 (1949)
= Pyrrosia dimorpha (Copel.) Parris; Fern Gaz. 12(2): 119 (1980)
= Pyrrosia pachyderma (Baker) Ching; Bull. Chin. Bot. Soc. 1: 46 (1935)
= Pyrrosia stellata (Copel.) Parris; Fern Gaz. 12(2): 119 (1980)
species :
Pyrrosia linearifolia (Hook.) Ching; Bull. Chin. Bot. Soc. 1: 48 (1935)
Distribution: China (Jilin); North Korea; South Korea (incl. Cheju Isl.); Japan; Ryukyu Isl.; Taiwan
36 CHI 38 JAP KOR NKO NNS TAI
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Neoniphopsis linearifolia (Hook.) Nakai; Bot. Mag. (Tokyo) 42: 217 (1928)
= Niphobolus linearifolius Hook.; 2. Cent. Ferns, t. 58 (1861)
= Polypodium linearifolium (Hook.) Hook.; Sp. Fil. 5: 53 (1863)
= Pyrrosia linearifolia monstr. cristata (Akasawa) Nakaike
= Pyrrosia linearifolia f. cristata Akasawa
= Pyrrosia linearifolia var. heterolepis Tagawa; J. Jap. Bot. 24: 116 (1949)
species :
Pyrrosia lingua (Thunb.) Farw.; Amer. Midl. Naturalist 12: 245 (1931)
Name: Japanese felt fern, Tongue fern
Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangsu, Jiangxi, Liaoning, Sichuan, Yunnan, Zhejiang); South Korea (incl. Cheju Isl.); Taiwan; NE-India; Myanmar (Kachin, etc.); N-Thailand; Laos; Vietnam
36 CHI 38 KOR TAI 40 IND 41 LAO MYA THA VIE
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Acrostichum lingua Thunb.; Fl. Jap. 330, t. 33 (1784) et Linn. Syst. Veg. ed. 14: 928 (1784)
= Craspedaria chinensis Link; Fil. Spec. 118 (1841)
= Cyclophorus bodinieri H.Lév.; Fl. Kouy-Tchéou: 478 (1915)
= Cyclophorus lingua (Thunb.) Desv.; Mem. Soc. Linn. Paris 6, 2: 224 (1827)
= Cyclophorus lingua var. angustifrons Hayata; Icon. Pl. Formosan. 5: 164 (1915)
= Cyclophorus lingua var. attenuata Rosenst.; Hedwigia 56(5): 347 (1915)
= Cyclophorus martini (Christ) C.Chr.; Index Filic. 199 (1905)
= Cyclophorus taiwanensis (Christ) C.Chr.; Index Filic. 201 (1905)
= Niphobolus lingua (Thunb.) Spreng.; Syst. Veg. ed. 16, 4(1): 45 (1827)
= Niphobolus martini Christ; Mém. Soc. Bot. France 1: 23 (1905)
= Niphobolus sinensis Loudon; Loudon, Hort. Brit., Suppl. 2: 651 (1839)
= Polycampium lingua (Thunb.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 3: 496 (1845)
= Polypodium lingua (Thunb.) Sw.; Syn. Fil. 29 (1806)
= Polypodium taiwanense Christ; Monsunia 1: 60 (1900)
= Pyrrosia caudifrons Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64(1): 37 (1983)
= Pyrrosia martini (Christ) Ching; Acta Phytotax. Sin. 10: 304 (1965)
= Pyrrosia medogensis Ching & S.K.Wu; Fl. Xizangica 1: 335 (1983)
species :
Pyrrosia longifolia (Burm.f.) C.V.Morton; J. Washington Acad. Sci. 36: 168 (1946)
Distribution: China (Hainan, Yunnan); Myanmar; Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (widespread); Singapore; Sumatra; Borneo (Sabah, Sarawak); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Lombok, Bali, Flores); Moluccas (Aru Isl., Seram, Ambon, etc.); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Hanover); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, Makira, Malaita, Ndeni, Rennell, Tulagi); Australia (NE-Queensland); New Caledonia (Loyalty Isl.); Santa Cruz Isl.; Vanuatu; Society Isl. (Tahiti)
36 CHI 41 CBD LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SNG SUL SUM 43 BIS SOL 50 AUS 60 NWC SCZ VAN 61 SCI
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Acrostichum bicolor Cav.; Anal. Hist. Nat. 1: 103 (1799)
= Acrostichum longifolium Burm.f.; Fl. Ind. 228 (1768)
= Candollea longifolia (Burm.f.) Mirb.; Hist. Nat. Veg. 6. 88 (1803)
= Cyclophorus acrostichoides (G.Forst.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 130 (1851)
= Cyclophorus acrostichoides f. carnosus Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 23: 8 (1916)
= Cyclophorus acrostichoides var. backeri Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 10 (1912)
= Cyclophorus acrostichoides var. fissus (Blume) Bonap.; Notes Ptéridol. 7: 125 (1918)
= Cyclophorus acrostichoides var. gracilis Copel.; Leafl. Philipp. Bot. 1(13): 234 (1907)
= Cyclophorus cinnamomeus Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 6: 192 (1922)
= Cyclophorus induratus Christ; J. Bot. (Morot) 21: 238, 271 (1908)
= Cyclophorus longifolius (Burm.f.) Desv.; Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 301 (1811)
= Cyclophorus macropodus (Baker) C.Chr.; Index Filic. 199 (1905)
= Cyclophorus scolopendrium Desv.; Mem. Soc. Linn. Paris 6, 2: 225 (1827)
= Cyclophorus valleculosus Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 10 (1912)
= Gyrosorium fissum (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 141 (1851)
= Niphobolus acrostichoides (G.Forst.) Bedd.; Handb. Ferns Brit. India, t. 81 (1865), not Kaulfuss (1824)
= Niphobolus fissus Blume; Enum. Pl. Javae 2: 106 (1828)
= Niphobolus longifolius (Burm.f.) Spreng.; Syst. Veg. ed. 16, 4(1): 46 (1827)
= Niphobolus puberulus Blume; Enum. Pl. Javae 2: 108 (1828)
= Niphobolus scolopendrium (Desv.) T.Moore; Index Filic. (T. Moore) 276 (1861)
= Polypodium acrostichoides G.Forst.; Fl. Ins. Austr. 81 (1786)
= Polypodium fissum (Blume) Baker; Hook. & Baker, Syn. Fil. 351 (1867)
= Polypodium furfuraceum Wall.; Numer. List no. 278 (1829), nomen
= Polypodium macropodum Baker; J. Linn. Soc., Bot. 15: 108 (1876)
= Pyrrosia acrostichoides (G.Forst.) Ching; Bull. Chin. Bot. Soc. 1: 69 (1935)
= Pyrrosia coccideisquamata Gilli; Ann. Naturhist. Mus. Wien 81: 26 (1977 publ. 1978)
= Pyrrosia fissa (Blume) Mehra; Ferns of Mussoorie, Punjab. Univ. Publ. 26 (1939)
= Pyrrosia macropoda (Baker) Ching; Bull. Chin. Bot. Soc. 1: 70 (1935)
species :
Pyrrosia mannii (Giesenh.) Ching; Bull. Chin. Bot. Soc. 1: 55 (1933)
Distribution: China (Yunnan); Tibet; Nepal; Bhutan; N-India (Uttarakhand, Sikkim, Darjeeling, Meghalaya, Khasia Hills, Jaintla Hills, S-India, Arunachal Pradesh); Myanmar (Kachin, etc.); Thailand; Sumatra; Borneo; Java
36 CHI TIB 40 BHU IND NEP 41 MYA THA 42 BOR JAW SUM
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Niphobolus floccigerus Bedd.; Handb. Ferns Brit. India, Suppl. 22, t. 386 (1876)
= Niphobolus mannii Giesenh.; Farngat. Niphobolus: 107 (1901)
= Pyrrosia birii P.C.Pande & H.C.Pande; Indian Fern J. 11(1-2): 98 (1994 publ. 1995) (without latin descr. or type, fide Fraser-Jenkins (1997))
= Pyrrosia mannii var. birii P.C.Pande & H.C.Pande; Ill. Fern Fl. Kumaon Himalaya 1: 97 (2003) (fide Fraser-Jenkins 2008)
species :
Pyrrosia matsudae (Hayata) Tagawa; J. Jap. Bot. 24: 119 (1949)
Distribution: Taiwan
38 TAI
Remarks: included in P. davidii by Flora of China but kept separate by Knapp (Ferns of Taiwan, 2011 ff.)
Synonyms: hidden
= Cyclophorus matsudae Hayata; Icon. Pl. Formosan. 10: 73 t 48 (1921)
species :
Pyrrosia niphoboloides (Luerss.) M.G.Price; Kalikasan 3: 177 (1974)
Distribution: Madagascar
29 MDG
Remarks: subgen. 6 Solis
Synonyms: hidden
= Drymoglossum niphoboloides (Luerss.) Baker; Hooker´s Icon. Pl. t. 1686 (1887)
= Taenitis niphoboloides Luerss.; Abh. Nat. Ver. Bremen 7: 49, t. 1, f. 3-5 (1882)
species :
Pyrrosia novoguineae (Christ) M.G.Price; Kalikasan 3: 177 (1974)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Cyclophorus bamleri Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 339 (1912)
= Cyclophorus ledermannii Brause; Bot. Jahrb. Syst. 66: 206 (1920)
= Cyclophorus novoguineae (Christ) Nakai; Bot. Mag. (Tokyo) 40: 386 (1926), as ´novo-guinense´
= Drymoglossum crassifolium Brause; Bot. Jahrb. Syst. 49: 35 (1912)
= Drymoglossum novoguineae Christ; Fl. Schutzgeb. Südsee: 137 (1901)
species :
Pyrrosia nummulariifolia (Sw.) Ching; Bull. Chin. Bot. Soc. 1: 47 (1935)
Name: Creeping Button fern
Distribution: China (Yunnan); Bhutan; N-India; Bangladesh; Myanmar (Kachin, etc.); Thailand; ?Laos; ?Cambodia; Vietnam; peninsular Malaysia (Perlis, Kedah, Penang, Perak, Kelantan, Pahang, Johor); Borneo (Sabah, Sarawak); Sulawesi; Palawan; Philippines; Lesser Sunda Isl. (Bali, Lombok)
36 CHI 40 BAN BHU IND 41 MYA THA VIE 42 BOR LSI MLY PHI PLW SUL
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Acrostichum nummulariifolium Sw.; Syn. Fil. 191, 418, t. 2, f. 1 (1806)
= Acrostichum nummulariifolium var. subpeltatum Blume; Fl. Javae, Fil. 33 (1829)
= Acrostichum obovatum Blume; Enum. Pl. Javae 2: 102 (1828)
= Cyclophorus nummulariifolius (Sw.) C.Chr.; Index Filic. 200 (1903)
= Cyclophorus nummulariifolius var. obovatus (Blume) Bonap.; Notes Ptéridol. 7: 126 (1918)
= Cyclophorus nummulariifolius var. rufus Alderw.; Bull. Dép. Agric. Indes Néerl. 21: 8 (1908)
= Cyclophorus obovatus (Blume) Alderw.; Malayan Ferns: 685 (1909)
= Galeoglossa nummulariifolia (Sw.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 133 (1851)
= Galeoglossa obovata (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 133 (1851)
= Galeoglossa rotundifolia C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 133 (1851)
= Gymnopteris nummulariifolia (Sw.) C.Presl; Tent. Pterid.: 244 (1836)
= Niphobolus nummulariifolius (Sw.) Bedd.; Ferns South. India, t. 186 (1863), isonym
= Niphobolus nummulariifolius (Sw.) J.Sm.; J. Bot. (Hooker) 3: 396 (1841)
= Niphobolus obovatus (Blume) Kunze; Bot. Zeitung (Berlin) 120 (1848)
= Niphobolus rotundifolius (C.Presl) Fée; Mém. Foug. 5. (Gen. Filic.) 262 (1852)
= Polypodium nummulariifolium (Sw.) Mett.; Farngatt. Poly. 123, t. 3, f. 9, 10 (1857)
= Polypodium nummulariifolium var. obovatum (Blume) C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 554 (1880)
= Polypodium obovatum (Blume) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 124 (1856)
= Pteropsis nummulariifolius (Sw.) Desv.; (1827)
= Pyrrosia nummulariifolia var. rufa (Alderw.) Ching; Bull. Chin. Bot. Soc. 1(1): 47 (1935)
= Pyrrosia obovata (Blume) Ching; Bull. Chin. Bot. Soc. 147 (1935)
species :
Pyrrosia pannosa (Mett.) Ching; Bull. Chin. Bot. Soc. 1: 58 (1935)
Distribution: Sri Lanka
40 SRL
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Cyclophorus pannosus (Mett.) C.Chr.; Index Filic. 200 (1905)
= Niphobolus lingua Bedd.; Ferns South. India, t. 240 (1865) (non Spreng. (1827))
= Niphobolus pannosus (Mett.) Bedd.; Handb. Ferns Brit. India, Suppl. 22 (1876)
= Polypodium pannosum Mett.; Kuhn, Linnaea 36: 141 (1869)
species :
Pyrrosia penangiana (Hook.) Holttum; Rev. Fl. Mal. 2: 146, f. 62 (1955)
Distribution: Myanmar; peninsular Thailand; SW-Thailand; Vietnam; peninsular Malaysia; Sumatra; Java; Lesser Sunda Isl. (Lombok, Timor, Sumbawa, Bali, Flores, Alor)
41 MYA THA VIE 42 JAW LSI MLY SUM
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Cyclophorus brevipes Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 139 (1920)
= Cyclophorus mollis (Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 131 (1851)
= Cyclophorus penangianus (Hook.) C.Chr.; Index Filic. 200 (1905)
= Niphobolus mollis Kunze; Bot. Zeitung (Berlin) 6: 121 (1848)
= Niphobolus penangianus Hook.; Hooker´s Icon. Pl. 4: t. 203 (1840)
= Polycampium penangianum (Hook.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 136 (1851)
= Polypodium molle (Kunze) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 128, n. 261 (1857)
= Pyrrosia mollis (Kunze) Ching; Bull. Chin. Bot. Soc. 1: 53 (1935)
species :
Pyrrosia petiolosa (Christ) Ching; Bull. Chin. Bot. Soc. 1: 59 (1935)
Distribution: Russian Far East (Khabarovsk, Primorye); China (Guizhou, Hebei, Henan, Hubei, Jiangsu, Jilin, Shaanxi, Shandong, Shanxi, Sichuan, Yunnan, Zhejiang); North Korea; South Korea
31 RFE 36 CHI 38 KOR NKO
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Cyclophorus petiolosus (Christ) C.Chr.; Index Filic. 200 (1905)
= Niphobolus petiolosus (Christ) Diels; Bot. Jahrb. Syst. 29: 207 (1900)
= Polypodium petiolosum Christ; Nuovo Giorn. bot. It. n. s. 4: 96, t. 1, f. 2 (1897)
species :
Pyrrosia piloselloides (L.) M.G.Price; Kalikasan 3: 176 (1974)
Distribution: China (Hainan, Yunnan, Guangxi); Japan; India (throughout); Bangladesh; Andaman Isl.; Thailand; Laos; ?Cambodia; Vietnam; peninsular Malaysia (widespread); Singapore; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.
36 CHI 38 JAP 40 BAN IND 41 AND LAO THA VIE 42 BOR JAW LSI MLY MOL NWG PHI SNG SUL SUM 43 BIS
Remarks: subgen. 4 Niphobolus. Possibly a synonym of P. heterophylla (L.) M. G. Price, in which case P. heterophylla would have priority
Synonyms: hidden
= Drymoglossum assamense Gand.; Bull. Soc. Bot. France 66: 306 (1920)
= Drymoglossum piloselloides (L.) C.Presl; Tent. Pterid. 227, t. 10, f. 5,6 (1836)
= Drymoglossum piloselloides var. platycerioides Teruya; Acta Phytotax. Geobot. 1: 197, 199 (1932)
= Drymoglossum rotundifolium C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 157 (1851)
= Elaphoglossum piloselloides (L.) Keyserl.; Polyp. Herb. Bunge 36 (1873)
= Lemmaphyllum piloselloides (L.) Luerss.; Bot. Centralbl. 11: 78 (1882)
= Notholaena piloselloides (L.) Kaulf.; Enum. Filic. 133 (1824) (Nothochlaena)
= Oetosis piloselloides (L.) Kuntze; Revis. Gen. Pl. 2: 817 (1891)
= Pteris piloselloides L.; Sp. Pl. ed. II, 2: 1530 (1763)
= Pteropsis piloselloides (L.) Desv.; Mem. Soc. Linn. Paris 6, 2: 218 (1827)
= Taenitis piloselloides (L.) Mett.; Fil. Hort. Bot. Lips. 28, t. 10, f. 6-8 (1856)
species :
Pyrrosia platyphylla Hovenkamp; Blumea 30(1): 207 (1984)
Distribution: Borneo (Mt. Kinabalu, Sabah, Kalimantan)
42 BOR
Remarks: subgen. 1 Pyrrosia
species :
Pyrrosia polydactylos (Hance) Ching; Bull. Chin. Bot. Soc. 1: 48 (1935)
Distribution: ?South Korea; Taiwan
38 TAI
Remarks: subgen. 3 Neoniphopsis. Identity of South Korean populations need to be verified
Synonyms: hidden
= Cyclophorus polydactylos (Hance) C.Chr.; Index Filic. 200 (1905)
= Niphobolus polydactylos (Hance) Giesenh. ex Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 325 (1899)
= Polypodium polydactylon Hance; J. Bot. 21: 269 (1883)
= Pyrrosia ×pseudopolydactylis Seriz.; J. Jap. Bot. 45: 120 (1970)
species :
Pyrrosia porosa (C.Presl) Hovenkamp; Blumea 30(1): 208 (1984)
Distribution: China (Guangxi, Guizhou, Hainan, Sichuan, Yunnan); Tibet; Japan; Taiwan; Nepal; Bhutan; India (Jammu & Kashmir, Manipur, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Tamil Nadu, Kodaikanal, Nilgiri Hills, Kerala, S-India, Arunachal Pradesh); Sri Lanka; Myanmar; Thailand; Laos; Vietnam; Philippines (Luzon)
36 CHI TIB 38 JAP TAI 40 BHU IND NEP SRL 41 LAO MYA THA VIE 42 PHI
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Cyclophorus acrocarpus (Christ ex Giesenh.) C.Chr.; Index Filic. 197 (1905)
= Cyclophorus alcicornu Christ; J. Bot. (Morot) 21: 238, 270 (1908)
= Cyclophorus macrophyllus (Christ) C.Chr.; Index Filic. 199 (1905)
= Cyclophorus malacophyllus (Christ) C.Chr.; Index Filic. 4: 199 (1905)
= Cyclophorus porosus (C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 130 (1851)
= Cyclophorus schmidianus C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 132 (1851)
= Cyclophorus sticticus (Kunze) C.Chr.; Index Filic. 201 (1905)
= Cyclophorus xiphioides (Christ) C.Chr.; Index Filic. 202 (1905)
= Niphobolus acrocarpus Christ ex Giesenh.; Bull. Acad. Int. Géogr. Bot., sér. 3, 11: 221 (1902)
= Niphobolus fissus Bedd.; Ferns Brit. India, Correct.: II (1870), nom. illeg.
= Niphobolus porosus C.Presl; Tent. Pterid. 202 (1836)
= Niphobolus schmidianus Kunze; Bot. Zeitung (Berlin) 6: 121 (1848)
= Niphobolus sticticus Kunze; Linnaea 24: 257 (1851)
= Niphobolus sticticus var. major Christ; Bull. Acad. Int. Géogr. Bot., sér. 3, 16: 15: 109 (1906)
= Niphobolus xiphioides Christ; Bull. Soc. Bot. France 5, Mem. 2: 1. 25 (1905)
= Polypodium fissum Baker; Hook. & Baker, Syn. Fil. 351 (1867)
= Polypodium mollissimum Christ; Bull. Herb. Boissier 7: 5 (1899) (non Fée (1866))
= Polypodium porosum (C.Presl) Mett.; Abh. Senckenb. Naturf. Ges. (Polypodium) 128 (1856)
= Polypodium porosum Wall.; Numer. List no. 266 (1829), nom. nud.
= Polypodium porosum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2: 128, n. 259 (1857)
= Polypodium sticticum (Kunze) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 128, n. 260 (1857)
= Pyrrosia mollis f. alcicornu (Christ) Ching; Bull. Chin. Bot. Soc. 1: 54 (1935)
= Pyrrosia mollis var. mollissima Ching; Bull. Chin. Bot. Soc. 1: 54 (1935)
= Pyrrosia porosa var. mollissima (Ching) K.H.Shing; Vasc. Pl. Hengduan Mount. 1: 171 (1993)
= Pyrrosia scolopendrina Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 74 (1941)
= Pyrrosia stictica (Kunze) Holttum; Novit. Bot. Inst. Prag. 31 (1968) (fide Fraser-Jenkins 1997)
species :
Pyrrosia princeps (Mett.) C.V.Morton; Amer. Fern J. 60: 118 (1970)
Distribution: Sulawesi; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea)
42 MOL NWG SUL
Remarks: subgen. 1 Pyrrosia
Synonyms: hidden
= Cyclophorus aglaophyllus Copel.; J. Arnold Arbor. 10: 179 (1929)
= Cyclophorus princeps (Mett.) C.Chr.; Index Filic. 200 (1905)
= Niphobolus princeps (Mett.) Giesenh. ex Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 325 (1899)
= Polypodium princeps Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 232 (1866)
= Pyrrosia aglaophylla (Copel.) Copel.; Fern Fl. Philip. 3: 474 (1960)
species :
Pyrrosia rasamalae (Racib.) K.H.Shing; Amer. Fern J. 73(3): 78 (1983)
Distribution: NE-India (Arunachal Pradesh); Myanmar; Thailand; Laos; peninsular Malaysia (Perak, Selangor, Negeri Sembilan, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak); Philippines (N-Luzon); Java; Lesser Sunda Isl. (Bali, Lombok, Sumbawa, Flores, Timor)
40 IND 41 LAO MYA THA 42 BOR JAW LSI MLY PHI SUM
Remarks: subgen. 5 Niphopsis
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Antrophyum niphoboloides Kunze; Bot. Zeitung (Berlin) 209 (1848)
= Cyclophorus borneensis Copel.; Philipp. J. Sci., C 12: 64 (1917)
= Cyclophorus lanuginosus (Giesenh.) C.Chr.; Index Filic. 199 (1905)
= Cyclophorus rasamalae (Racib.) C.Chr.; Index Filic. 200 (1905)
= Cyclophorus rasamalae f. minor Alderw.; Bull. Dép. Agric. Indes Néerl. 21: 8 (1908)
= Niphobolus lanuginosus Giesenh.; Farngatt. Niphobolus 190 (1901)
= Niphobolus rasamalae (Racib.) Giesenh.; Farngatt. Niphobolus 187 (1901)
= Polypodium rasamalae Racib.; Pteridoph. Buitenzorg 99 (1898)
= Polypodium samarense Christ; Bull. Herb. Boissier 6: 199 (1898)
= Pyrrosia arunachalensis Sarn.Singh & Panigrahi; Ferns Fern-Allies Arunachal Pradesh 2: 526 (2005) (fide Fraser-Jenkins 2008)
= Pyrrosia borneensis (Copel.) K.H.Shing; Amer. Fern J. 73(3): 78 (1983)
= Pyrrosia lanuginosa (Giesenh.) Copel.; Fern Fl. Philip. 3: 472 (1960)
= Taenitis niphoboloides (Kunze) T.Moore; Index Filic. (T. Moore) 30 (1857)
species :
Pyrrosia rhodesiana (C.Chr.) Schelpe; J. S. Afr. Bot. 18: 126 (1952)
Distribution: Uganda; Tanzania; Malawi; Zimbabwe; Mozambique; Madagascar
25 TAN UGA 26 MLW MOZ ZIM 29 MDG
Remarks: subgen. 6 Solis
Synonyms: hidden
= Cyclophorus madagascariensis C.Chr.; Dansk Bot. Ark. 7: 161, t. 61, f. 7-10 (1932)
= Cyclophorus oblanceolatus C.Chr.; Dansk Bot. Ark. 7: 160, t. 61, f. 4-6 (1932)
= Cyclophorus rhodesianus C.Chr.; Dansk Bot. Ark. 7: 161 (1932)
= Pyrrosia madagascariensis (C.Chr.) Schelpe; J. S. Afr. Bot. 18: 133 (1952)
= Pyrrosia oblanceolata (C.Chr.) Tardieu; Humbert, Fl. Madag. Fam. 5, 2: 103 (1960)
species :
Pyrrosia rupestris (R.Br.) Ching; Bull. Chin. Bot. Soc. 1: 49 (1935)
Name: Felt fern
Distribution: Australia (E-Queensland, E-New South Wales, SE-Victoria)
50 AUS
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Craspedaria rupestris (R.Br.) Link; Fil. Spec. 118 (1841)
= Cyclophorus rupestris (R.Br.) C.Chr.; Index Filic. 200 (1905)
= Cyclophorus serpens var. rupestris (R.Br.) Domin; Biblioth. Bot. 20: 187 (1915)
= Niphobolus confluens (Hook.) F.M.Bailey; Handb. Queensl. Ferns 43 (1874) (non Spreng. (1827))
= Niphobolus rupestris (R.Br.) Spreng.; Syst. Veg. ed. 16, 4(1): 44 (1827)
= Polypodium confluens Hook.; Sp. Fil. 5: 46 (1863) (non R. Br (1810))
= Polypodium rupestre R.Br.; Prodr. Fl. Nov. Holland. 146 (1810)
species :
Pyrrosia samarensis (Mett.) Ching; Bull. Chin. Bot. Soc. 1: 49 (1935)
Distribution: Palawan; Philippines
42 PHI PLW
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Cyclophorus argyrolepis Christ; Bull. Herb. Boissier, sér, 2, 6: 991 (1906)
= Cyclophorus samarensis (C.Presl) C.Chr.; Index Filic. 200 (1905)
= Gyrosorium samarense C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 140 (1851)
= Niphobolus samarensis (C.Presl) Fée; Mém. Foug. 5 (Gen. Filic.): 263 (1852)
= Polypodium samarense (C.Presl) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 123, n. 246, t. 3, f. 1-4 (1857)
= Pyrrosia philippinensis Copel.; Philipp. J. Sci. 81: 43 (1952), nom. nov. superfl.
species :
Pyrrosia sarthalensis S.Singh, Sneha & B.Singh; Phytotaxa 522(3): 250 (2021)
Distribution: NW-India (Jammu & Kashmir)
40 IND
species :
Pyrrosia serpens (G.Forst.) Ching; Bull. Chin. Bot. Soc. 1: 49 (1935)
Distribution: Norfolk Isl.; Kermadec Isl.; Fiji (Viti Levu); Cook Isl. (Rarotonga, Mangaia, Aitutaki, Ngaputoru Group); Society Isl. (Tahiti, Maupiti, Bora Bora, Moorea); Austral Isl. (Rapa Iti, Raivavae, Rimatara, Tubuai, Rurutu); Pitcairn Isl.; Henderson Isl.
50 NFK 51 KER 60 FIJ 61 AUI COO HEN PIT SCI
Remarks: subgen. 4 Niphobolus
Synonyms: hidden
= Craspedaria ovalis C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 263 (1851)
= Craspedaria serpens (G.Forst.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 263 (1851)
= Cyclophorus blepharolepis C.Chr.; Index Filic. 198 (1905)
= Cyclophorus macrocarpus (Willd.) Copel.; Univ. Calif. Publ. Bot. 12: 381 (1931)
= Cyclophorus serpens (G.Forst.) C.Chr.; Index Filic. 201 (1905)
= Niphobolus bicolor Kaulf.; Enum. Filic. 128 (1824)
= Niphobolus macrocarpus (Willd.) Hook. & Arn.; Bot. Beech. Voy. 74, t. 18 (1832)
= Niphobolus serpens (G.Forst.) Endl.; Prodr. Fl. Norf. 8 (1883)
= Polypodium bicolor (Kaulf.) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 125, n. 251 (1857)
= Polypodium macrocarpum Willd.; Sp. Pl. 5 (1810)
= Polypodium serpens G.Forst.; Fl. Ins. Austr. 81 (1786) (non Sw. (1788))
= Polypodium serpens var. cuneatum F.Muell.; Fragm. 5: 141 (1865)
= Polypodium stellatum Vahl; Symb. Bot. 3: 104 (1794)
= Polypodium stoloniferum J.F.Gmel.; Syst. Nat. ed. 13[bis]. 2(2): 1305 (1792)
= Pyrrosia bicolor (Kaulf.) Ching; Bull. Chin. Bot. Soc. 1: 49 (1935)
= Pyrrosia blepharolepis (C.Chr.) Ching; Bull. Chin. Bot. Soc. 1: 49 (1935)
= Pyrrosia macrocarpa (Willd.) K.H.Shing; Amer. Fern J. 73: 78 (1983), nom. illeg.
= Pyrrosia macrocarpa (Willd.) Kaulf.; Enum. Filic. 245 (1824)
species :
Pyrrosia sheareri (Baker) Ching; Bull. Chin. Bot. Soc. 1: 64 (1935)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Jiangxi, Sichuan, Yunnan, Zhejiang); Taiwan; Vietnam
36 CHI 38 TAI 41 VIE
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Cyclophorus drakeanus f. maximus Y.C.Wu; Y. C. Wu, K. K. Wong & Pong, Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ. 3: 338, pl. 140 (1932)
= Cyclophorus grandissimus Hayata; Icon. Pl. Formosan. 4: 255, f. 179 (1914)
= Cyclophorus inaequalis (Christ) C.Chr.; Index Filic. 199 (1905)
= Cyclophorus sheareri (Baker) C.Chr.; Index Filic. 201 (1905)
= Cyclophorus sheareri f. maximus (Y.C.Wu) C.Chr.; Bull. Dept. Biol. Coll. Sci. Sun Yatsen 6: 18 (1933)
= Niphobolus inaequalis Christ; Bull. Soc. Bot. France 52, Mem. 1: 25 (1905)
= Niphobolus sheareri (Baker) Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 325 (1899)
= Polypodium sheareri Baker; J. Bot. 13: 201 (1875)
= Pyrrosia grandissima (Hayata) Ching; Bull. Chin. Bot. Soc. 1: 63 (1935)
= Pyrrosia nanchuanensis Ching; Bull. Chin. Bot. Soc. 1: 65 (1935)
species :
Pyrrosia similis Ching; Bull. Chin. Bot. Soc. 1: 56 (1935)
Distribution: China (Guangxi, Guizhou, Sichuan)
36 CHI
species :
Pyrrosia sphaerosticha (Mett.) Ching; Bull. Chin. Bot. Soc. 1: 62 (1935)
Distribution: Sulawesi; Philippines; Lesser Sunda Isl. (Timor); Moluccas (Bacan)
42 LSI MOL PHI SUL
Remarks: subgen. 5 Niphopsis
Synonyms: hidden
= Cyclophorus pseudolingua Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 11: 6 (1913)
= Cyclophorus sarasinorum (Giesenh.) C.Chr.; Index Filic. 200 (1905)
= Cyclophorus sphaerostichus (Mett.) C.Chr.; Index Filic. 201 (1905)
= Cyclophorus warburgii (Giesenh.) C.Chr.; Index Filic. 202 (1905)
= Niphobolus sarasinorum Giesenh.; Farngatt. Niphobolus 162 (1901)
= Niphobolus smithianus T.Moore; Index Filic. (T. Moore) 76 (1857)
= Niphobolus sphaerostichus (Mett.) Copel.; Polyp. Philipp. 114 (1905)
= Niphobolus warburgii Giesenh.; Farngatt. Niphobolus 163 (1901)
= Polypodium sphaerostichum Mett.; Abh. Senckenberg. Naturf. Ges. 2: 130, n. 265, t. 3, f. 13 (1857)
= Sphaerostichum acrostichoides C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 135 (1851) (non Cyclophorus)
species :
Pyrrosia spissa (Bory ex Willd.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 3: 262 (2020)
Distribution: Cameroon; Principe Isl.; São Tomé; Annobon Isl.; Gabon; Congo [Brazzaville]; Central African Republic; D.R.Congo [Zaire]; Uganda; Tanzania; Zimbabwe; Mozambique; Madagascar; Mauritius; La Réunion; Rodrigues; Seychelles
23 ANN CAF CMN CON GAB PRI SAO ZAI 25 TAN UGA 26 MOZ ZIM 29 MAU MDG REU ROD SEY
Remarks: subgen. 4 Niphobolus. Often combined with the Asian P. lanceolata
Synonyms: hidden
= Cyclophorus spissus (Bory ex Willd.) Desv.; Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 301 (1811)
= Cyclophorus spissus var. continentalis (Hieron. ex Engl.) Hieron.; Bot. Jahrb. Syst. 46: 399 (1911)
= Niphobolus adnascens var. spissus (Bory ex Willd.) Keyserl.; Polyp. Herb. Bunge 37 (1873)
= Niphobolus spissus (Bory ex Willd.) Kaulf.; Enum. Filic. 126 (1824)
= Niphobolus spissus var. continentalis Hieron. ex Engl.; Veg. d. Erde 9(2): 55, f. 53 (1908)
= Polypodium spissum Bory ex Willd.; Sp. Pl. ed. 4, 5: 146 (1810)
species :
Pyrrosia splendens (C.Presl) Ching; Bull. Chin. Bot. Soc. 1: 68 (1935)
Distribution: Philippines (Luzon, Samar, Leyte, Bohol, Mindanao)
42 PHI
Remarks: subgen. 1 Pyrrosia
Synonyms: hidden
= Apalophlebia splendens (J.Sm.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 138 (1851)
= Cyclophorus splendens (J.Sm.) C.Chr.; Index Filic. 201 (1905)
= Niphobolus splendens (C.Presl) J.Sm. ex T.Moore; Index Fil. (T. Moore) 2: lxxvi (1857)
= Polypodium nitens Baker; Hook. & Baker, Syn. Fil. ed. 2: 353 (1874) (non Desv. (1827))
= Polypodium splendens (J.Sm.) Hook.; Sp. 5: 52 (1862)
species :
Pyrrosia stenophylla (Bedd.) Ching; Bull. Chin. Bot. Soc. 1: 55 (1935)
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Yunnan); Tibet; Nepal; Bhutan; NE-India (Sikkim, Arunachal Pradesh, Manipur); Myanmar (Kachin); Thailand; Laos; Vietnam
36 CHI TIB 40 BHU IND NEP 41 LAO MYA THA VIE
Synonyms: hidden
= Cyclophorus stenophyllus (Bedd.) C.Chr.; Index Filic. 201 (1905)
= Cyclophorus tonkinensis (Giesenh.) C.Chr.; Index Filic. 201 (1905)
= Cyclophorus vittarioides Christ; Bull. Acad. Géogr. Bot. Mans (1909), Mém. 20: 175 (1909), nom. illeg.
= Niphobolus fissus var. stenophyllus Bedd.; Handb. Ferns Brit. India, Suppl. 92 (1892)
= Niphobolus stenophyllus (Bedd.) Giesenh.; Farngatt. Niphobolus 131 (1901)
= Niphobolus tonkinensis Giesenh.; Farngatt. Niphobolus 144 (1901)
= Polypodium fissum var. floccigerum C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 550 (1880)
= Polypodium mysurense B.Heyne ex Wall.; Numer. List no. 269 (1829), nomen
= Pyrrosia linearis Ching & S.K.Wu; Fl. Xizangica 1: 340 (1983)
= Pyrrosia nayariana Ching & P.Chandra ex P.Chandra; Amer. Fern J. 54: 62, f. 1-10 (1964) (fide Fraser-Jenkins 1997)
= Pyrrosia porosa var. stenophylla (Bedd.) Hovenkamp; Blumea 30(1): 208 (1984)
= Pyrrosia porosa var. tonkinensis (Giesenh.) Hovenkamp; Blumea 30(1): 208 (1984)
= Pyrrosia tibetica Ching; Lingnan Sci. J. 21: 34 (1945)
= Pyrrosia tibetica var. angustata Ching; Lingnan Sci. J. 21: 34 (1945)
= Pyrrosia tonkinensis (Giesenh.) Ching; Bull. Chin. Bot. Soc. 1: 55 (1935)
species :
Pyrrosia stigmosa (Sw.) Ching; Bull. Chin. Bot. Soc. 1: 67 (1935)
Distribution: China (Yunnan); Tibet; India (Arunachal Pradesh); Myanmar; Thailand (widespread); Laos; Cambodia; Vietnam; peninsular Malaysia (Perlis, Kedah, Langkawi Isl., Perak, Selangor, Kelantan, Terengganu, Pahang); Sumatra; S-Sulawesi; Lesser Sunda Isl. (Bali)
36 CHI TIB 40 IND 41 CBD LAO MYA THA VIE 42 LSI MLY SUL SUM
Remarks: subgen. 1 Pyrrosia
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Apalophlebia venosa (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 139 (1851)
= Cyclophorus stigmosus (Sw.) Desv.; Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 301 (1811)
= Niphobolus stigmosus (Sw.) T.Moore; Index Filic. (T. Moore) 276 (1861)
= Niphobolus venosus Blume; Fl. Javae, Fil. 63, t. 28, f. 1 (1828)
= Polypodium stigmosum Sw.; J. Bot. (Schrader) 1800 (2), 21 (1801) (p. p.)
= Pyrrosia chinensis Mirb.; Lam. & Mirb., Hist. Nat. Vég. Cl. Fam. 5: 92 (1802)
species :
Pyrrosia stolzii (Hieron.) Schelpe; J. S. Afr. Bot. 18: 133 (1952)
Distribution: Tanzania; Zambia; Malawi
25 TAN 26 MLW ZAM
Remarks: subgen. 6 Solis
Synonyms: hidden
= Cyclophorus stolzii (Hieron.) Hieron.; Bot. Jahrb. Syst. 46: 396 (1911)
= Niphobolus stoltzii Hieron.; Engl., Veg. d. Erde 9(2): 55 (1908), nomen
species :
Pyrrosia subfurfuracea (Hook.) Ching; Bull. Chin. Bot. Soc. 1: 68 (1935)
Distribution: China (Yunnan); Tibet; Bhutan; NE-India (Arunachal Pradesh, Nagaland, Manipur); Myanmar (Kachin, etc.); Laos; Vietnam
36 CHI TIB 40 BHU IND 41 LAO MYA VIE
Remarks: subgen. 3 Neoniphopsis
Synonyms: hidden
= Cyclophorus subfurfuraceus (Hook.) C.Chr.; Index Filic. 201 (1905)
= Niphobolus subfurfuraceus (Hook.) Bedd.; Handb. Ferns Brit. India, t. 259 (1869)
= Polypodium subfurfuraceum Hook.; Sp. Fil. 5: 52 (1863)
species :
Pyrrosia tricholepis (Carruth.) Ching; Acta Phytotax. Sin. 10: 301 (1965)
Distribution: Society Isl. (Tahiti)
61 SCI
Synonyms: hidden
= Niphobolus tricholepis Carruth.; Seem., Fl. Vit. 367 (1873)
= Polypodium tricholepis Mett.; Kuhn, Linnaea 36: 139 (1869) (non Schrad. (1824))
species :
Pyrrosia ×nipponica M.Beppu & Seriz.; J. Phytogeogr. Taxon. 30(1): 46 (1982)
Distribution: South Korea; Japan
38 JAP KOR
Remarks: Pyrrosia linearifolia x hastata
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A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

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Assembled through 40 years independently from other databases

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