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Inflorescence structure in species of Scleria subgenus Hypoporum and subgenus Scleria (Sclerieae-Cyperaceae)

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Abstract

The objective of this study was to realize a typology approach toward the inflorescences in Scleria in order to provide characters that would have potential use in further taxonomic and phylogenetic research and that would allow identification of the tendencies and processes that could have yielded the variations within Scleria inflorescences. The majority of species studied present a main florescence; Scleria reticularis and S. melanomphala have truncated inflorescences. The variations are related to the grade of development reached by the main florescence and the different parts of the paracladial zone. Major inflorescence variability was found in section Scleria. A change in spikelet sexuality can occur toward the distal parts. There were variations in the distinct species with respect to the hierarchy of the paracladia where the change occurs. Future investigations should center on the ontogenetic aspects of inflorescence development of Scleria, integrating these results, together with those of adult inflorescences, into taxonomic and phylogenetic investigations.

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Authors and Affiliations

  1. Botánica General, Facultad de Ciencias Agrarias, Universidad Nacional de Jujuy, Jujuy, Argentina

    Osvaldo Ahumada

  2. Morfología Vegetal, Facultad de Ciencias Agrarias, Universidad Nacional del Litoral, Esperanza, Santa Fe, Argentina

    Abelardo C. Vegetti

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  1. Osvaldo Ahumada
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  2. Abelardo C. Vegetti
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Correspondence to Abelardo C. Vegetti.

Appendix

Appendix

Section Hypoporum

Scleria composita (Nees) Boeck.: Ekman 1297 (B, SI); Steinbach 6981 (B, BM, G, SI); Glaziou 22354 (BR, G); Pedersen 816 (BR, CTES); Pedersen 4510 (BR, CTES, G, K, LP); Herrera 420 (CTES); Krapovickas et al. 24746 (CTES); Nicora 5257 (CTES, SI); Quarín 469 (CTES); Schinini et al. 21775 (CTES, JUA); Beck 13175 (GENT, JUA, LPB); Irwin et al. 24393 (K); Ahumada et al. 6664 (JUA); Ahumada et al. 6666 (JUA); Ahumada et al. 7633 (JUA); ídem. 7643 (JUA); Ahumada et al. 7886 (JUA); Guillén et al. 416 (JUA, MNES); Killeen 1597 (LPB); Guaglianone et al. 846 (SI); Pedersen 13466 (SI); Rojas 14051 (SI).

Scleria distans Poir.: Buchtein 7163 (B); Edwal 9206 (B); Ekman 1798 (B); Ekman 6205 (B); Herter 344 (B, CTES, G, LIL, M, SI); Lombardo Marchesi Mus. Botan. Montev. 1839 (B); Mutis 6109 (B); Osten 7414 (B); Osten 8835 (B, MVM); Osten 16088 (B); Steinbach 6861 (B, BM, G, K); Steinbach 6982 (B, BM, G, K); Partridge 60808 (BA); Rabello et Barbosa 197a (BA, MVM, SI); Hatschbach 2073 (BAA, SI); sin coleccionista (BAA 1054); Jörgensen-Hansen 702 (BAB); Martínez Crovetto et Milano 5001 (BAB); Balansa 448a (BAF, BM, G, K); Klein 3834 (BAF); Molfino s.n. (BAF); Reitz et Klein 16449 (BAF); Dusén 13490 (BM); Fiebrig 4840 (BM, G, K, P); Hassler 3575 (BM, G, K, P); Hassler 5633 (BM, G, K, P); Hassler 8254 (BM, G); Hassler 8794 (BM, G, K); Marín et Giménez GM518 (BM, SI); Morong 424 (BM, K); Spruce s.n. (BM); Zardini et Park guard 15500 (BM, PY); Balansa 448 (BR, G, K, P); P. Claussen 427 (BR); Kummrow 2129 (BR); Pedersen 499 (BR, G, K, LP, P); Pedersen 3652 (BR, G); Stephan s.n. (BR); Wullschlägel 655 (BR); Ahumada 1931 (CTES); Ahumada 2240 (CTES); Arbo et al. 1227 (CTES); Arbo et al. 6228 (CTES); Arbo et al. 6391 (CTES, JUA); Arbo et al. 6773 (CTES); Arbo et al. 7110 (CTES, JUA); Barros 2430 (CTES, LIL, SI); Bordas et Schmeda 4112 (CTES, FCQ); Carnevali 2404 (CTES); Carnevali 3901 (CTES); Carnevali 4380 (CTES); Carnevali 5482 (CTES); Carnevali 6231 (CTES); Carnevali 6332 (CTES); Fernández 977 (CTES); Ferrucci et al. 259 (CTES); Hatschbach 26214 (CTES); Jorgensen 3508 (CTES); Krapovickas et Cristóbal 13515 (CTES); Krapovickas et al. 25337 (CTES); Krapovickas et Schinini 36325 (CTES); Neiff 812 (CTES); Pedersen 7895 (CTES); Pedersen 12761 (CTES); Quarín et al. 2443 (CTES); Quarín et al. 2730 (CTES); Rambo 38818 (CTES); Rambo 44558 (CTES); Rosengurtt B-4319 (CTES, MVM, SI); Schinini 22943 (CTES); Schinini et al. 29334 (CTES, G, JUA); Schwindt 931 (CTES, LIL); Sparre et Vervoorst 986 (CTES); Swartz 9007 (CTES, LIL); Tressens et al. 920 (CTES); Cavalcante 926 (Herbario Emilio Göeldi; Basualdo 5442 (FCQ); Basualdo 5852 (FCQ); Degen 1731 (FCQ); Mereles 3479 (FCQ); Soria 4451 (FCQ); Soria 5834 (FCQ); Nicora et al. 9914 (FCQ, SI); Zardini et Velásquez 26597 (FCQ); Balansa 448b (G, P); Eggers 2491 (G, M); Fiebrig 437 (G, K, M, P); Hassler 32 (Herb. Hassler 2420) (G, SI); Hassler 1315 (G, K, P); Osten 4409 (G); Osten 5352 (G, MVM); Poeppig s.n. (G); Reitz et Klein 8424 (G); Riedel s.n. (G, SI 1805); Sellow s.n. (G); Steinbach 5327 (G, MVM); Steinbach 5495 (G); Beck 13137 (GENT); Ahumada 6340 (JUA); Ahumada et al. 6655 (JUA); Ahumada et al. 6660 (JUA); Ahumada et al. 6662 (JUA); Ahumada et al. 6663 (JUA); Ahumada et al. 7428 (JUA); Zardini et al. 641 (JUA, LP, SI); Troncoso et Bacigalupo 2985 (JUA, SI); Hasse 880 (K); Jorgensen 3588 (K); Lindeman 1899 (K); Montes 3291 (K, SI); Pedersen 7696 (K, P, SI); Wood 9373 (K); Wood et Goyder 15406 (K); Woolston 577 (K, SI); Woolston 1427 (K); Rambo 38514 (LIL); Duarte 6339 (LP, SI); Jorgensen 3588a (LP, SI); sin coleccionista 50729 (LP); Abott et Isaacs 16507 (LPB); Beck et al. 9867 (LPB); Beck 14860 (LPB); Beck 17196 (LPB); Beck 17458 (LPB); Beck 17561 (LPB); Beck 17866 (LPB); Beck 18381 (LPB); Bruderreck 16 (LPB); Buchtein 7165 (LPB); García et al. 2121 (LPB); Killeen 1763 (LPB); Müller 9456 (LPB); Legrand 22 (MVM); Osten 21865 (MVM, SI); Rosengurtt 971 (MVM); Rosengurtt 975 (MVM); Bonpland s.n. (P); Croat 24024 (P); Ellenberger 1457 (P); Poeppig 34 (P); Reitz et Klein 17825 (P, SI); Saint-Hilaire 2358 (P); Ule 1344 (P); Brian Abrell 68 (PY); Zardini et forest ranger 10277 (PY); Archer et Rojas 4852 (SI); Barrros 2405 (SI); Barros 2451 (SI); Bartlett 21235 (SI); Burkart 17322 (SI);Burkart 19419 (SI); Burkart 21534 (SI); Burkart 25074 (SI); Burkart et al. 29212 (SI); Burkart et al. 29213 (SI); Burkart et al. 29765 (SI); Burkart et al. 29779 (SI); Burkart et al. 30735 (SI); Chebataroff 727 (SI); Glaziou 13303 (SI); Hatschbach 2073 (SI); Hatschbach 3692 (SI); Hatschbach 6699 (SI); Joly 367 AB5 (SI); Klein 744 (SI); Klein 2545 (SI); Klein et al. 5924 (SI); MG N° 24670 (SI); Ocemago 12399 (SI); Rambo 32454 (SI); Reitz et Klein 399 (SI); Reitz 5338 (SI); Reitz et Klein 5668 (SI); Reitz et Klein 16593 (SI); Reitz et Klein 10304 (SI); Rojas 1032 (SI); Rojas 6661 (SI); Smith et Klein 7518 (SI); Smith et al. 9391 (SI); Türckheim 197 (SI); Zardini et Velázquez 24691 (SI); Zardini et Tilleria 31510 (SI).

Scleria leptostachya Kunth: Steinbach 5300 (B, G, MVM, SI); Steinbach 6864 (B, G, K); Steinbach 6927 (B, BM, G, K, SI); Reitz et Klein 16422 (BAF); Gardner s.n. (BM); Hassler 8210 (BM, G); Riedel 2990 (BM); Caballero Marmori 1353 (CTES); Hatschbach 26041 (CTES, LP); Schwindt 2348 (CTES, LIL); Balansa 456 (G, P).Sellow s.n. (G); Irwin et al. 11690 (GENT); Irwin et al. 25930 (GENT); Schessl 3142 (GENT); Ahumada 7578 (JUA); Burkart 14759 (JUA, K, SI); Glaziou 20052 (K); Ratter et Bertolda 1034 (K); Riedel 158 (K).Wood 14322 (K); Rambo 30816a (LIL); Regnell Ser. III, 1321 (LIL, P); Bruderreck 125 (LPB); Killeen 2041 (LPB); Duarte 7755 (M, SI);); Regnell s.n. (P); Riedel 2990 (P); Weddell s.n. (P); Araujo 58 (SI); Fontana F177-34 (SI); Hatschbach 3687 (SI); Irwin et al. 10234 (SI); Irwin et al. 16399 (SI); Irwin et al.18660 (SI); Klein 4604 (SI); Rambo 30816 (SI); Reitz et Klein 11983 (SI); Riedel 1833 (SI); Rojas 6846 (SI); Smith et Klein 10864 (SI); Smith et Klein 10991 (SI).

Scleria variegata (Nees) Steud: Brade 11191 (B); Hoehne 19492 (B); Hatschbach 24052 (CTES); Vanni et al. 3156 (CTES, JUA); Ahumada et al. 6658 (JUA); Martius s.n. (M); Herter 4319 (MVM); Atala 269 (SI); Castellanos 22697 (SI); Guaglianone et Gatusso 2504 (SI); Martinelli et al. 10860 (SI).

Section Ophryoscleria

Scleria macrophylla J. Presl & C. Presl: Pipoly et al.11702 (B); Sagot 1172 (BM, K, P); Steinbach 8021 (BM, G, K); Billiet et Jadin 6001 (BR); Morrone et M. Belgrano 5058 (CTES); Gardner 1858. (G); Gardner 1895 (G, K); Huker 7753 (G); Leblond s.n. (G); Pittier 4882 (G).Salzmann s.n. (G); Beck et al. 19107 (GENT); Beck 20717 (GENT); Croat et Rosas 62688A (GENT); Smith et al. 13645 (GENT, LPB); Beck et al. 19630 (K); Burchell 5489 (K); Gardner 1875 (K); Feurer 6480 (LPB); Gonzáles 85 (LPB); Wood 15012 (LPB); Janssen 581 (M); Martius s.n. (M); Prance et Silva 59012 (M); sin coleccionista s.n. (M); Raynal-Roques 20176 (P); Richard s.n. (P); sin coleccionista s.n. (P); Soubirou s.n. (P); Soubirou s.n. (P); Soubirou s.n. (P); Rojas 3689 (SI).

Scleria microcarpa Nees ex Kunth: Ekman 2115 (B); Ekman 4378 (B); Ekman 11232 (B); Ekman 13260 (B); Ekman 15893 (B); Ekman 17369 (B); Ekman 17115 (B); Killeen et Guillén 6837 (B); Pickel 335 (B); Pickel 3453 (B); Pickel 3160 (B); Rojas 7420 (B); Sampaio 5617 (B); Sampaio 5897 (B); Steinbach 7564 (B, BM, G, K); Hassler 8081 (BM, G, K, LIL, P); Ridley el al. s.n. (BM); Sagot 897 (BR, K, P); Schomburgk 660 (BR, K); Wright 723 (BR); Encarnación 26480 (G); Gehriger 279 (G); Kallunki et al. 483 (G); Poeppig 3062 (G); Anderson 10671 (GENT); Chonono et Saavedra 257 (GENT, LPB); Focke 910 (GENT); Focke 1069 (GENT); Jardim et al. 241 (GENT); Killeen et Guillén 6877 (GENT); Luceño 122 (GENT); Moraes 250 (GENT, LPB); Moraes et al. 1770 (GENT); Hasse 895 (K); Beck et al. 19348 (K); Lowe 4221 (K); Sperling 6559 (K, LPB); Traill 1206 (K, P); Ule 5298 (K); Martius s.n. (M); Sintenis 781 (M) Croat 24024 A (P); Benoist 1407 (P); Cremers 7836 (P); Leprieur 77 (P); Mélinon s.n. (P); Mélinon 18 (P); Poeppig s.n. (P); Schomburgk s.n. (P); Sintenis 78 (P); sin coleccionista s.n. (P); Anderson 10600 (SI); Rojas 1017 (SI); Rojas 2288 (SI).

Scleria mitis P.J. Bergius, Kongl.: Gregory et al.10313 (BAA); Hassler 10041 (BAF, G, K, LIL P); Sintenis 6659 (BAF); Jiménez et Marín, G 1955BJ (BM, SI); Weigelt s.n. (BM, G); Martius s.n. (BR); Ferrucci et al. 1426 (CTES, JUA); Ferrucci et al. 1634 (CTES, JUA); Hatschbach et al. 58618 (CTES); Krapovickas et Schinini 31779 (CTES); Krapovickas et Schinini 31994 (CTES); Mereles et Geisslu 3972 (CTES); Pedersen 12154 (CTES); Schinini et al. 29268 (CTES, G); Schinini et Dematteis 33297 (CTES); Mereles 3472 (FCQ); Broadway 7731 (G); Salzmann s.n. (G); Steinbach 5512 (G); Irwin et al. 14364 (GENT); Luceño 68 (GENT);W. Thomas et al. 11399 (GENT); Asplund 9328 (K); Asplund 19321 (K); Hage et al. 2292 (K); Kallunki et al. 483 (K); Spruce s.n. (K); Cabrera et Zardini 29985 (LP); Luetzelburg s.n. (M); Martius s.n. (M); Claussen 990 (P); Cremers et al.12486 (P); L. C. Richard s.n. (P); Sagot s.n. (P); Salzmann s.n. (P); Solomon et al. 6903 (PY); Filgueiras et Zuloaga 2192 SI).

Scleria obtusa Core: Fiebrig 4728. (G, K, M, P); Beck 3308 (GENT); Beck et Hasse 10131 (GENT, LPB); Beck 15032 (GENT, JUA, K); Beck et Hasse 9923 (LPB); Bruderreck 110 (LPB); Gutierrez et al. 1881 (LPB); Hasse 388 (LPB); Orellana et Beck 46 (LPB); Orellana et Beck 208 (LPB); Orellana 361 (LPB); Orellana et al. 566 (LPB); Orellana et al. 686 (LPB); Saavedra et Chonono 357 (LPB).

Section Schizolepis

Scleria latifolia Sw.: Barros 2343 (B, CTES, SI); Bartlett 11718 (B); Buchtein 388 (B); Buchtein 7161 (B); Camargo 2712 (B); Edwalli 9237 (B); Herter 4328 (B, MVM); Irwin et Soderstrom 6277 (B); Jensen-Jacobs et al. 257 (B); Jensen-Jacobs et al. 343 (B); Jürgens s.n. (acc. 1946 Herb. Rudolf Gross) (B); Loefgren et Edwalli 9232 (B); Rambo 57151 (B); Rambo 57157 (B); Swallen 3180 (B); Usteri 9247 (B); Walter 210 (B); Kermes 2463 bis (BAB, BAF); Múlgura et al. 595 (BAB, SI); Rodríguez 289 (BAB); Hassler 11302 (BAF, BM, G, K, P, SI); Hassler 8685 (BM, G, K, P, SI); Zardini et Velázquez 15780 (BM, PY); Ahumada 3689 (CTES); Araujo 808 et Peixoto 614, Herb. RB 172106 (CTES); Arbo 464 (CTES); Caballero Marmori s.n., Herb. Itaipú Binacional 1727 (CTES); Correa et al. 5417 (CTES); Guaglianone et al. 2911 (CTES, JUA, SI); Hatschbach et al. 35945 (CTES); Irwin et al. 18295 (CTES); João et Rohr s.n. (CTES 63168, LIL 201324); Krapovickas et al. 17214 (CTES); Krapovickas et al. 26431 (CTES); Krapovickas et Cristóbal 38532 (CTES); Lindeman et al., serie 26.13, Herb. ICN 28216 A y B (CTES); Pallarés 208 (CTES); Pedersen 5968 (CTES, K, P SI); Pedersen 9303 (CTES, K); Pérez 357 (CTES, PY); Pipoly et al.11216 (B, CTES).Quarín 480 (CTES, LP, P); Quarín 487 (CTES, LIL); Quarín et al. 2765 (CTES); Quarín 3376 (CTES); Schinini et Carnevali 10602 (CTES); Schinini et al. 11230 (CTES); Schinini et al. 23755 (CTES, JUA); Schinini et Caballero Marmori 27105 (CTES); Sparre et Vervoorst 518 (CTES); Tressens et al. 384 (CTES, K); Tressens et al. 883 (CTES); Tressens 2925 (CTES); Valls et Irgang 2979 A y B (CTES); Basualdo 2275 (FCQ); Mereles 3638 (FCQ); Zardini 6757 (FCQ, GENT, SI); Zardini et Aguayo 8494a (FCQ); Belanger 407 (G); Broadway 565 (G); Broadway 4755 (G, P); Broadway 5508 (G); Fiebrig 418 (G); Gaudichaud 85 (G, P); Hahn 1281 (G); Hart a.1896 (G); Hassler 10439 (G); Herminier 16 (G); Huber 1901 (G); Jardim et al. 737 (G); Kuniyoshi and Kuniyoshi 1620 (G); Molina et al. 17331 (G), Riedel s.n. (G); Schinini 4416 (G); Shotsky s.n. (G); Stutz 2079 (G); Tonduz 12776 (G); Tonduz 12853 (G); Weddell 1132 (G); Beck 27629 (GENT); Beck 13654 (GENT, JUA, LPB); Gentry et al. 59140 (GENT); Luceño 17 (GENT); Solomon 10818 (GENT); Zardini et Tilleria 28495 (GENT); Zardini et Aquino 32861 (GENT); Zardini et Vera 42643 (GENT); Ahumada 6338 (JUA); Burchell 980 (K); Lira et al. 71 (K); Rodríguez et al. 42 (K); Rodrigues et al. 49 (K); Sales de Melo et al. 228 (K); Silva et al. 125 (K); Tschá et al. 311 (K); Husnot 1868 (M); Martius s.n. (M); Meier et al. 2007 (M); Othmer s.n. (M); Prance et Silva 59590 (M); sin coleccionista s.n. (M); Swartz s.n. (M); Vogl A 21 (M); Rojas 2285 (MVM, SI); Dwyer et Lieser 12081 (P, SI); Glaziou s.n. (P); Lourteig 1935 (P); Nadeaud s.n. (P); Levy 1407 (P); Questel 4101 (P); Le Gallo 2847 (P); Raynal 15565 (P); Raynal 19571 (P); Raynal s.n. (P); Zardini et Benítez 3412 (PY); Zardini 7567 (PY); Zardini et park guard 11819 (PY); Zardini 15998 (PY); Joly et al. 681ABJ (SI); Koyama et al. 14141 (SI); Martinelli et al. 10950 a (SI); Montes 104 B (SI); Mori et al. 13773 (SI); Morrone et al. 1432 (SI); Múlgura et al. 1884 (JUA, SI); Múlgura et al. 1928 (SI); Múlgura et al. 2650 (SI); Múlgura et al. 3283 (JUA, SI); Muniz 354 (SI); Rambo 37920 (SI); Reitz et Klein 10960 (SI); Rojas 8095 (SI).

Scleria panicoides Kunth: Brade 10755 (B); Brade 10897 (B); Brade 12650 (B); Buchtein 5101 (B); Gehtr 7864 (B); Gross 20136 (B); Hoehne 3914 (B); Hoehne 9242 (B); Rambo 41854 (B); Hauman 22516 (BA); Jörgensen 4490 (BA); Pérez Moreau 21058 (BA); Bang 2350 (BM, G, K); Lindeman et Hass 1942 (BR); Orth 704 (BR, K); Amarilla et al. 75 (CTES); Castellanos 21662 (CTES); Eskuche et al. 5-126 (CTES, SI); Hatschbach 16491 (CTES); Honfi 53 (CTES, K); Keller et F. Moskovich (CTES); Keller 2797 (CTES); Herb. Orchioni 4913 (CTES); Pedersen 7791 (CTES); Quarín 1147 (CTES, LIL); Reitz 1796 (CTES); Schinini et al. 28746 (CTES, JUA); Schinini et al. 31390 (CTES); Tressens et al. 5389 (CTES); Tressens 5838 (CTES); Keel 1495 (FCQ); Reitz et Klein 8912 (G, SI); Solomon 18411 (GENT); Guaglianone et al. 2892 (JUA, SI); Rodríguez et al. 566 (JUA, MNES); Rodríguez et al. 847 (JUA, MNES); Ball s.n. (K); Custódio Filho et Franco 2743 (K); Glaziou 11658 (K, P); Hunt 6351 (K); sin coleccionista (K); Bertoni 2786 (LIL); Rodríguez 371 (LP, SI); Martius s.n. (M); Martius s.n. (M); Gaudichaud 84 (P); Lindeman et Hass 5111 (P); Lourteig 2349 (P); Mexia 4686 (P); Weddel 366 (P); Castellanos 24428 (SI); Deginani et al. 1258 (SI); Reitz et Klein 4293 (SI); Reitz et Klein 6789 (SI).

Scleria plusiophylla Steud.: Brade 12171 (B); Henkel et al. 5599 (B, GENT); Hoehne 9240 (B); Jörgensen 4490 (B, MVM, SI); Rojas 5030 (B, SI); Hauman 1923 (BA); Perrone 54115 (BA); Perrone 54117 (BA); Benítez 56 (BAA, CTES, LIL, LP); Correa et al. 5417 (BAB, CTES, SI); Fernández et al. 184 (BAB, CTES, LIL); Molina et al. 4103 (BAB, PY); Hassler 6365 (BM, G); Arenas 685 (CTES); Guaglianone et al. 2818 (CTES, JUA, SI); Krapovickas et Cristóbal 28753 (CTES); Mereles 3607 (CTES, FCQ); Ortiz 233 (CTES); Múlgura et al. 3514 (CTES, JUA, SI); Schinini 5606 (CTES); Schinini et Fernández 6076 (CTES); Schinini 8045 (CTES, G); Schinini et Bordas 20230 A y B (CTES); Schinini et al. 21927 (CTES); Schinini et Caballero Marmori 27023 (CTES); Schinini et M. Quintana 36342 (CTES); Vanni et al. 2719 (CTES, K, LIL); Ahumada 6343 (JUA); Ahumada et al. 6657 (JUA); Ahumada O. et al. 6668 (JUA); Beck 24722 (JUA, LPB); Cabral et al. 262 (JUA, CTES); Guaglianone et al. 3004 (JUA); Seidel et Vargas 1143 (JUA, LPB); Tur et Guaglianone 1974 (JUA, SI); Mereles 1125 (FCQ); Mereles 3606 (FCQ); Schinini 4416a (FCQ, G); Soria 1789 (FCQ, G); Soria 3086 (FCQ); Zardini et Degen 3667 (FCQ); Zardini et Salina 26153 (FCQ, GENT); Balansa 460 (G); Degen 223 (G, SI); Bertoni 976 y 976a (G); Chodat 269 (G); Fernández Casas et Molero 3755 (G); Hassler 901 (G); Hatschbach 45855 (G); Pedersen 14697 (G, SI); Beck 17294 (GENT, JUA, LPB); Zardini et Park guard 15511 (GENT, PY); Zardini et Gerrero 37110 (GENT); Glaziou 18593 (K, P); Hassler 8685b (K); Silva et al. 51 (K); Solomon 18411 (LPB); Brunner 1501 (PY); Zardini et forest ranger 10302 (PY); Zardini et Velásquez 15344 (PY); Zardini et Velásquez 15630 (PY); Boelcke 6743 (SI); Cabrera et al. 28894 (SI); Morrone et al. 1455 (SI); Morrone et al. 1633 (SI); Morrone et al. 2044 (SI); Pavetti et Rojas 9723 b (SI); Zardini et Aguayo 8494 (SI); Zardini et Benítez 47341 (SI); Zuloaga et al. 5162 (SI).

Section Scleria

Scleria bracteata Cav.: Bang 762 (BM, G); Buchtein 3652 (B, M); Buchtein 5102 (B); Buchtein 5103 (B); Buchtein 5104 (B); Buchtein 7163 (B); Buchtein 7164 (B); Buchtein 8067 (B); Buchtein 8068 (B); Faz, Loefgren 1359 (B); Hoffmannsegg s.n. (B–W 17341); Irwin et Soderstrom 6549 (B); Mass et al. 7204 (B); Mexia 6593 (B); Mexia 9145 (B); Pickel 644 (B); Pickel 3142 (B); Pilger 723 (B); Pipoly et al. 11458 (B); Sampaio 5567 (B); Steinbach 7038 (B, G, K, LIL); Ule 8540 (B, K); Hassler 11294 (BM, G, P); Ridley el al. s.n. (BM); Williams 1652 (BM); Martius s.n. (BR, K); Pittier et Durand 3693 (BR); Pohl s.n. (BR); Schomburgk 108 (BR, G, K); Wright 725 (BR).Blanchet 692 (G); Holton 109 (G); Johnston 203 (G); Jurgensen 659 (G); Krukoff 7135 (G); Pavon s.n. (G); Pittier 8093 (G); Pittier 9450 (G); Poeppig s.n. (G); Sagot 1350 (G).Salzmann s.n. (G, K); Salzmann 937 (G); Tnduz 4640 (G).Ule 8540 (G); Beck 12101 (GENT, LPB, SI); Beck et Ruthsatz 21813 (GENT); García 835 (GENT); Grifo et Solomon 809 (GENT); Luceño et Alves 87 (GENT); Luceño 133 (GENT); Mendes 195 (GENT); Mendes 2895 (GENT); Vargas et Seidel 441 (GENT); Beck 27883 (JUA, LPB); Bang 267 (K, M); da Silva et al. 3078 (K); Noblick et al. 2357 (K); Pulle 510 (K);Beck 1786 (LPB); Beck 2243 (LPB); Beck 17765 (LPB); Beck 22767 (LPB); Beck 24723 (LPB); Foster et Ribera 12323 (LPB); Lüetzelburg 134 (M); Lüetzelburg 136 (M); Lüetzelburg 192 (M); Martius 862 (M); Martius 2950 (M); Martius 2951 (M); Martius 2952 (M); Martius 2953 (M); Martius 2954 (M); Martius 2955 (M); Martius 2956; Spruce s.n. (M); Martius 2957. (M), Martius 2958 (M); Othmer s.n. (M); Pereira et al. 9221 (M); Prance et Silva 59495 (M); Reiche 424 (M); Wagner s.n. (M); Bonpland s.n. (P); de Grosourdy 18 (P); Pentland s.n. (P); Richard s.n. (P).

Scleria ciliata Michx.: Pedersen 4277 (BR, G, K, SI); Hatschbach 25353 (CTES); Pedersen 8518 (CTES, K); Tressens et al. 4202 (CTES); Davidse et al. 35117 (GENT); Ahumada 6341 (JUA); Ahumada et al. 6654 (JUA); Ahumada et al. 6661 (JUA); Ahumada et al. 6665 (JUA); Ahumada et al. 7406 (JUA); Ahumada et al. 7437 (JUA); Ahumada et al. 7454 (JUA); Ahumada et al. 7577 (JUA); Honfi 616 (JUA, MNES); Troncoso et al. 2012 (JUA, SI); Godfrey 48108 (LIL); Bacigalupo et al. 369 (SI); Bacigalupo et al. 554 (SI); Bacigalupo et al. 846 (SI); Bacigalupo et al. 1036 (SI); Burkart et al. 29214 (SI); Burkart et al. 29215 (SI); Fredholm 5041 (SI); Fredholm 5769 (SI); Guaglianone et al. 3049 (SI); Smith et Klein 7478 (SI); Smith et Klein 8558 (SI); Umbach 252 (SI).

Scleria flagellum-nigrorum P.J. Bergius: Schulz 6333 (BAA); Bertoni 5083 (CTES);Bertoni 5321 (CTES, LIL); Carnevali 3524 (CTES); Krapovickas et al. 24273 (CTES); Keel 1210 (FCQ); Keel et Vera 1277 (FCQ); Hassler 2904 (G); Mexia 6423 (G); Prance et al. 3489 (G); Tessmann 3277 (G); Simpson et al. 668 (G); Guillén et Coria 1635 (GENT); Schessl 2407 (GENT); Burchell 2723 (K); Sagot 991 (K); Sellow s.n. (K); Pott et al. 5833 (LP); Moraes et al. 1785 (LPB); Janssen et Gemtchujnicov 400 (M).

Scleria lacustris C. Wright: Ekman 11583 (B); Ekman 17741 (B); Ekman 17916 (B); Hassler 8081 a (BM, G); Beck 15035 (GENT, JUA, K, LPB); Franken 187 (GENT, LPB); Guareco 409 (GENT, LPB); Orellana 247 (GENT, LPB); Pott et al. 5739 (LP); Moraes et Sarmiento 1276 (LPB); Orellana 803 (LPB); Robinson 4700 (M); Weddell 3300 (P); Rojas 6817 (SI).

Scleria melaleuca Rchb. ex Schltdl. & Cham.: Buchtein 388 (B); Buchtein 4283 (B); Ekman 1801 (B); Ekman 2283 (B); Ekman 4327 (B); Dupré s.n. (B); Ekman 15816 (B); Günther 331 (B, P); Günther 334 (B); Günther 335 (B); Howard 9528 (B); Irwin et al. 21443 (B); Irwin al. 26255 (B); Pickel 3321 (B); sin coleccionista s. n. (B); Steinbach 6860 (B, BA, BM, G, K); Steinbach 14994 (B); Hauman 33278 (BA); Jörgensen 3312 (BA, SI); Jörgensen 3578 (BA, BAB, LIL, SI); Steinbach 2367 (BA); Nicora et Cámara Hernández 627 (BAA); Parodi 5456 (BAA, SI); Parodi 7088 (BAA); Bacigalupo et al. 1102 (BAB, FCQ); Cabral et Molina 1014 (BAB, CTES); Fortunato et al. 2491 (BAB); Kermes (404) 5.800 bis (BAB); Llamas BAB 27007 (BAB); Petetín et Molina 1465 (BAB); Ramella et al. 3033 (BAB); Spegazzini 17843 (BAB); Spegazzini, C. et P. Spegazzini 19332 (BAB); Spegazzini 19333 (BAB); Spegazzini 19355 (BAB); Spegazzini 19356 (BAB); Spegazzini 19357 (BAB); Balansa 3028 (BAF, P); Kermes s.n. (BAF); Hassler 491 (BM, G, K, P); Hassler 8667 (BM, G; K, P); Jiménez et Marín 65 (BM, CTES, JUA, SI); Morong 246 (BM, K); Schomburgk 876 (BM, K); Triana s.n. (BM); Zardini et Velázquez 16867 (BM); de Gryse 36 (BR); Husnot 41 (BR); Eggers 190 (BR); Johnson s.n. (BR); Martius s.n. (BR); Pedersen 533 (BR, K, LP, P); Pedersen 3203 (BR, G); Pedersen 3204 (BR); Pittier et Durand 2715 (BR); Sieber 269 (BR); Wright 726 (BR);Arenas 1147 (CTES); Bordas 3035 (CTES); Bordas 4169 (CTES); Bordón s.n. (CTES); Croat 18077 (CTES); Carnevali 6005 (CTES); Duré 482 (CTES); Heinonen et al. 74 (CTES); Honfi 65 (CTES, MNES); Krapovickas et Cristóbal 11342 (CTES); Krapovickas et al. 23904 (CTES); Krapovickas et al. 24274 (CTES, LIL); Krapovickas et al. 25467 (CTES); Krapovickas et Schinini 36319 (CTES); Liddell 274 (CTES); Mereles 794 (CTES, FCQ, G); Morel 3503 (CTES); Mroginski et al. 327 (CTES); Pedersen 15001 (CTES); Pedersen 16044 (CTES, G); Quarín 212 (CTES, LIL); Quarìn et Schinini 1068 (CTES); Quarín et al. 2603 (CTES); Ramalho Campelo et Costa Silva 1846 (CTES); Revilla 41 A (CTES); Schinini 5710 (CTES, FCQ); Schinini 5862 (CTES, G); Schinini 8161 (CTES); Schinini 21053 (CTES, JUA); Schinini 23144 (CTES, G, JUA); Schinini et Carnevali 23323 (CTES, JUA); Schinini et Bordas 25141 (CTES); Schinini et Vanni 25977 (CTES, G); Schinini 27542 (CTES); Schinini et al. 27708 (CTES); Schinini et al. 27871 (CTES); Schinini et Caballero Marmori 29947 (CTES); Schinini et Caballero Marmori 30225 (CTES, G); Schulz 2914 (CTES, SI); Schulz 6333 (CTES, JUA); Schulz 11741 (CTES); Smith 2195 (CTES);Sparre et Vervoorst 1018 (CTES); Sparre et Vervoorst 1652 (CTES); Schwarz 4046 (CTES); Schwarz 4336 (CTES); Schwarz 7088 (CTES); Schwarz 11739 (CTES); Tressens et al. 5189 (CTES, JUA, SI); Vanni et al. 1587 (CTES, LIL); Soria 1526 (FCQ); Mereles et al. 391 (FCQ, G); Mereles et al. 395 (FCQ); Mereles 1122 (FCQ); Mereles 3686 (FCQ); Mereles 3877 (FCQ); Mereles 4399 (FCQ); Zardini 7968 (b) (FCQ); Zardini et Velásquez 8944 (FCQ); Zardini et Velásquez 10755 (FCQ); Zardini et Velázquez 26702 (FCQ, GENT); Zardini et Velázquez 27120 (FCQ); Zardini et Velázquez 27205 (FCQ); Zardini et Velázquez 27332 (FCQ); Balansa 458 (G, K, P); Ekman 1271 (G); Fiebrig 418 b (G); Fiebrig 730 (G); Fiebrig 4728a (G); Garvizu et Fuentes. 387 (G); Hahn 2073 (G, PY); Hassler 2407 (G); Lossau 260 (G); Lossau 268 (G); Mamaní et al. 1400 (G); Mereles 2360 (G); Pittier et Tonduz 8393 (G); Piitier et Tonduz 8524 (G); Ritter 3121 (G); Rojas 169 (G); Schunke 2697 (G); Schunke 4540 (G); Solomon et al. 6868 (G, PY); Steinbach 5230 (G); Stern et al. 321 (G); Tonduz 8529 (G); Tonduz 14617 (G); Alves et al. 2295 (GENT); Beck et al. 372 (GENT); Beck 13192 (GENT, JUA, LPB); Chonono et Saavedra 257 (GENT, LPB); Focke 1220 (GENT); Gottsberger et Posey 43-22183 (GENT); Guareco 277 (GENT); Guillén et Choré 1525 (GENT); Guillén et Choré 3121 (GENT); Jardim et al. 173 (GENT); Luceño 16 (GENT); Schessl 315/1-3 (GENT); Schessl 268/1-6 (GENT); Schessl et Gottsberger 3035 (GENT); Solomon 17158 (GENT); Zardini et Velázquez 20505 (GENT); Zardini et Aquino 30195 (GENT, SI); Zardini et Aquino 31652 (GENT); Zardini et Guerrero 33763 (GENT); Zardini et Guerrero 41737 (GENT); Ahumada 6336 (JUA); Ahumada 6339 (JUA); Ahumada 6344 (JUA); Ahumada et al. 6659 (JUA); Ahumada et al. 6670 (JUA); Ahumada et al. 7384 (JUA); Ahumada et al. 7673 (JUA); Ahumada et al. 7772 (JUA); Honfi 258 (JUA, MNES); Múlgura et al. 3048 (JUA, SI); Seijo et al. 580 (JUA, MNES); André 494 (K); Baur 320 (K); Burchell 9938 (K); da Silva et al. 2917 (K); Fiebrig 418 (K); Glaziou 1424 (K); Glaziou 14373 (K); Guaglianone et al. 417 (K, SI); Martius s.n. (K); Pedersen 7548 (K, P); Sagot 627 (K); Salzmann s.n. (K, P); Sellow s.n. (K); Spruce s.n. (K); Stannard 122 (K); Woolston 463 (K); Villalba s.n. (LIL); Hatschbach 24325 (LP); Hatschbach 24170 (LP); La Plata 12857 (LP); Schwuabe 704 (LP); Beck 3266 (LPB); Chonono et Saavedra 124 (LPB); Luetzelburg 15604 (M); Martius s.n. (M);Ex herbario Schwargrichen 318 (M); Swartz s.n. (M); Debeaux 72 (P); Eggers 6596 (P); Ledru s.n. (P); Lepreieur s.n. (P); Mandon 96 (P); Raynal 15846 (P); Raynal 18275 (P); Riedel s.n. (P); Rodríguez 1560 (P); Rodríguez 2850 bis (P); sin coleccionista s.n. (P); Hahn et al. 1062 (PY); Hahn 2336 (PY); Soria 5497 (PY); Zardini et Velázquez 10387 (PY); Zardini et park guard 11798 (PY); Buchtein 1236 (SI);Filgueiras et Zuloaga 2244 (SI); Guaglianone et Múlgura 2117 (SI); Jiménez 52 (SI); Mereles 963 (SI); Meyer 255 (SI); Meyer 448 (SI); Morrone et Pensiero 321 (SI); Riedel 909 (SI); Smith et al. 6328 (SI);Tur et Guaglianone 2017 (SI); Zardini et Guerrero 35340 (SI); Zardini et Guerrero 37771 (SI); Zuloaga et al. 2244 (SI).

Scleria melanomphala Kunth: Ekman 1303 (B, G); Hassler 8889 (BM, G, K, P); Hassler 10144 (BM, G, K, P); Fiebrig 6298 a (G, SI); Callejas 7048 (GENT); Irwin et al. 16470 (GENT); Sneidern 1342 (G); Woolston 1104 (K, SI); Anderson 9554 (LIL); Hatschbach 34223 (LP); Guaglianone et al. 843 (SI).

Scleria reticularis Michx.: Ekman 11952 (B); Howard 9933 (B); Mass et al. 7362 (CTES); Pedersen 2611 (CTES); Quarín 471 (CTES); Steinbach 5425 (G); Ule 8065 (G); Ahumada et al. 9457 (JUA); Rambo 46182 (LIL); Sloughs, Umbach 4604 (LIL); Tracy 7127 (LIL); Sigle 19 (LPB); Bruderreck 241 (LPB); Assene 2686 (P); Assene 8434 (P); Feuillet 990 (P);Biltomore Herb. 3287a (SI); Black et al. 57-19758 (SI); Black 50-9989 (SI); Brevard 6038 (SI); Umbach 250 (SI).

Scleria scabra Willd.: Brade 9057 (B); Buchtein 330 (B); Ekman 17858 (B); Ekman 18063 (B); Spruce s.n.(BM, G, K); Schinini et Dematteis 33855 (CTES, JUA); Araujo et al. 1232 (G); Hatschbach 31900 (G); Hatschbach et al. 64884 (G); Hatschbach et al. 69016 (G); Ramella 2757 (G); Beck 18598 (GENT, JUA, LPB); Guillén et Centurión 874 (GENT); Luceño et al. 282 (GENT); Arbo et al. 5642 (K); Burchell 7013 (K); Hunt 5655 (K); Wood et Goyder 15383 (K); Loefgren 1223 (P); Huber 2135 (SI).

Scleria secans (L.) Urb.: Buchtein 329 (B, P); Ekman 2801 (B); Ekman 3878 (B);Ekman 11495 (B); Jörgensen 3585 (B, BA, MVM, SI); Jürgens 162 (B); Piruibe, Loefgren 9235 (B); Swallen 3135 (B); Schwacke 123 (B); Perez Moreau 48166 (BA, CTES); Balansa 3027 (BAF); Marín et Jiménez GM293 (BM); Spruce s.n. (BM, K); Williams 771 (BM); Husnot s.n. (BR); Alvim 270 (CTES); Eskuche 425 (CTES, SI); Gentry et Revilla 16197 (CTES); Hatschbach 25791 (CTES); Nee 8987 (CTES); Ratter et al. 6363 (CTES); Revilla 63 (CTES); Schinini 4322 (CTES, FCQ, LIL); Schinini et Bordas 20357 (CTES); Schinini et al. 21926 (CTES, JUA);Schinini 23126 (CTES, G); Schinini 23987 (CTES, G); Schinini et Caballero Marmori 27022 (CTES); Mereles 1124 (FCQ); Zardini 6724 (FCQ); Zardini 7433 (FCQ); Chodat 267 (G); Chodat 291 (G); Hahn 1740 (G, PY); L` Herminier s.n. (G); Salzmann s.n. (G); Schunke 3716 (G); Sellow s.n. (G); Steinbach 5308 (G, MVM);Webster et Wilson 5028 (G);Weigelt s.n. (G); Killeen et al. 4802 (GENT); Beck et Foster 13948 (GENT, JUA, LPB); Zardini et Velásquez 15759 (GENT, PY); Zardini et Telleria 28387 (GENT); Ahumada 6337 (JUA); Ahumada 6342 (JUA); Ahumada et al. 6669 (JUA); Ahumada et al. 7651 (JUA); Ahumada et al. 7778 (JUA); Guaglianone et al. 3006 (JUA, SI); Burchell 1844 (K); Burchell 9273 (K, P); Hostman 837 (K); Lanjouw et Lindeman 930 (K); Mueller s.n. (K); Noblick 1808 (K); Sagot 626 (K); Salzmann s.n. (K); Helme 432 (LPB); Gröger et al. 1334 (M); Martius s.n. (M); Pohl s.n. (M); Swartz s.n. (M); Leblond s.n. (P); Ledru 788 (P); Salzmann s.n. (P); Smith et Klein 7977 (P, SI); Zardini 7269 (PY, SI); Zardini 12609 (PY); Beck 7339 (SI); Burkart 15346 (SI); Guaglianone et al. 1076 (SI); Guaglianone et Múlgura 2168 (SI); Hatschbach 2511 (SI); Klein 2643 (SI); Klein et al. 5912 (SI); López et Conradt 203 (SI); Martinelli et al. 11046 (SI); Múlgura et al. 2806 (SI); Reitz 2269 (SI); Reitz et Klein 2292 (SI); Reitz et Klein 3794 (SI); Reitz et Klein 9191 (SI); Rojas 5256 (SI); Scolnik 1180 (SI); Smith et Klein 7572 (SI); Traill s.n. Santos et al. 1469 (SI).

Scleria sellowiana Kunth: Ekman 1282 (B, G); Herter 611a (B); Jürgens 38 (B); Lombardo 3563 (B); Osten 6572 (B, LIL); Osten 6932 (B, BA, LIL); Rambo 48857 (B); Hauman 33279 (BA); Parodi 12377 (BAA, SI); Pedersen 1881 (BR, CTES, LP, P); Pedersen 10013 (BR, CTES, G); Ahumada et al. 6656 (CTES); Fernández 473 (CTES); Goldfarb s.n. CTES. 353009 (CTES); Lindeman et al. 21161 (CTES); Montes 9819 (CTES, LIL); Montes 11106 (CTES, LP); Montes 12913 (CTES, LP); Montes 12949 (CTES, LP); Rambo 44237 (CTES, JUA); Rambo 49057 (CTES, SI); Schinini 16065 (CTES); Schinini et. Caballero Marmori 30163 (CTES, JUA); Schinini et al. 35.687 (CTES); Schulz 2971 (CTES, SI); Herter 4353 (G, M, SI); Beck 17863 (GENT); Ahumada et al. 6667 (JUA); Bacigalupo et al. 1060 (JUA, SI); Bacigalupo et Guaglianone 1523 (JUA, SI); Bacigalupo et Guaglianone 1526 (JUA, SI); Guaglianone et al. 787 (JUA, SI); Montes 15255 (JUA, SI); Balansa 459a (K); Castellanos 11754 (LIL, P); Jörgensen 3312 (LIL); Legrand 3612 (MVM); Osorio 225 (MVM); Rosengurt B-2768 (MVM, SI); Pérez et Aguayo 2371 (PY); Arechavaleta 4392 (SI); Barros 2429 (SI); Barros 2450 (SI); Burkart 7932 (SI); Chebataroff 4512 (SI); Gallinal et al. 4548 (SI); Klein 3716 (SI); Osten 22295 (SI); Rambo 54528 (SI); Reitz et Klein 5139 (SI); Rosengurtt B-422 (LP, SI); Rosengurtt B-3183 (SI); Rosengurtt B-4217 (SI); Sehnem 5039 (SI); Pire 1940 (UNR).

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Ahumada, O., Vegetti, A.C. Inflorescence structure in species of Scleria subgenus Hypoporum and subgenus Scleria (Sclerieae-Cyperaceae). Plant Syst Evol 281, 115–135 (2009). https://doi.org/10.1007/s00606-009-0194-4

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