Amblygonocarpus

Taxonomy

Amblygonocarpus H.A.T. Harms In Engler et Prantl, Nat. Pflanzenfam. Nachtr. II-IV 1: 191. Aug 1897.

Subfamily: Mimosoideae.
Phylogenetic Number: 2.3.11.
Tribe: Mimoseae.
Group: Adenanthera.
Species Studied - Species in Genus: 1 studied; 1 in genus.

Description

Fruit: A legume; unilocular; 8–20 cm long; 1.8–3.5 cm wide; 2–2.3 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight (or nearly so); not plicate; not twisted; symmetrical; oblong; not inflated; terete to quadrangular; without beak; blunt at apex to short tapered at apex; aligned with longitudinal axis of fruit; short tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; ligneous; seed chambers externally invisible (with regularly spaced depressions); margin not constricted; margin without sulcus; margin plain; wing(s) absent; stipitate; with the stipe 7–40 mm long; indehiscent (remaining attached to sutures). Replum invisible. Epicarp glossy; monochrome; brown to black; with surface texture uniform; glabrous; eglandular; without spines; smooth (but dimpled at regular intervals over seed chambers); not veined; not tuberculate; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; surface not veined; 1-layered; without balsamic vesicles; without fibers; without reniform canals; spongy (partially filling cavity). Endocarp present; visible; dull; opaque; monochrome; tan; smooth; without adhering pieces of testa; septate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 10–15; length transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus 10–15 mm long; of 1 length only; filiform; straight. Aril absent.

Seed: 10–13 mm long; 7–9 mm wide; 4–6 mm thick; not overgrown; not angular; symmetrical; oblong; compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; glossy; not modified by a bloom; colored; monochrome, or streaked; dark brown; with pale brown overlay; glabrous; smooth; osseous. Pleurogram present; 98 %. Pseudopleurogram absent. Fracture lines present. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; punctiform; apical at apex of radicle tip; flush; within halo (colored or textured). Lens discernible; 0.7 mm long; with margins straight; linear; not in groove of raphe; adjacent to hilum; flush (within hilum halo); similar color as testa; lighter than testa; tan; not within corona, halo, or rim. Endosperm present; thick; not pluglike and not resembling tip of radicle; covering entire embryo; adnate to embryo. Cotyledons smooth; both outer faces flat; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; partially concealing radicle; split over radicle; with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; parallel to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; straight with embryonic axis; centered between cotyledons. Plumule moderately developed; glabrous.

Distribution

Africa (Savannas).

Old World; Africa (savannas).

 Fruit and seed:  A. andongensis  (F.M.J. Welwitsch ex D. Oliver) Exell & Torre - bottom fruit, top left transverse section through seed chamber, top center and top right longitudinal section of fruit through seed chambers.
Fruit and seed: A. andongensis (F.M.J. Welwitsch ex D. Oliver) Exell & Torre - bottom fruit, top left transverse section through seed chamber, top center and top right longitudinal section of fruit through seed chambers.
 Seed, cotyledon, embryo, and testa:  A. andongensis  (F.M.J. Welwitsch ex D. Oliver) Exell & Torre - top left cotyledon concealing all but tip of radicle (L) and embryonic axis (R), bottom left seed topography, bottom right seeds, testa SEM.
Seed, cotyledon, embryo, and testa: A. andongensis (F.M.J. Welwitsch ex D. Oliver) Exell & Torre - top left cotyledon concealing all but tip of radicle (L) and embryonic axis (R), bottom left seed topography, bottom right seeds, testa SEM.