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Catalogue of the Diptera (Insecta) of Morocco— an annotated checklist, with distributions and a bibliography
expand article infoKawtar Kettani‡, Martin J. Ebejer§, David M. Ackland|, Gerhard Bächli, David Barraclough#, Miroslav Barták¤, Miguel Carles-Tolrá«, Milos Černý», Pierfilippo Cerretti˄, Peter Chandler˅, Mohamed Dakki¦, Christophe Daugeronˀ, Herman De Jongˁ, Josef Dils₵, Henry Disneyℓ, Boris Droz₰, Neal Evenhuis₱, Paul Gatt₳, Gustavo Graciolli₴, Igor Y. Grichanov₣, Jean-Paul Haenni₮, Martin Hauser₦, Oumnia Himmi¦, Iain MacGowan₭, Bruno Mathieu₲, Mohamed Mouna¦, Lorenzo Munari‽, Emilia P. Nartshuk₩, Oleg P. Negrobov₸, Pjotr Oosterbroekˁ, Thomas Pape‡‡, Adrian C. Pont|, Grigory V. Popov§§, Knut Rognes||, Marcela Skuhravᶶ, Vaclav Skuhravý¶¶, Martin Speight##, Guy Tomasovic¤¤, Bouchra Trari‡, Hans-Peter Tschorsnig««, Jean-Claude Vala»», Michael von Tschirnhaus˄˄, Rüdiger Wagner˅˅, Daniel Whitmore««, Andrzej J. Woźnica¦¦, Tadeusz Zatwarnickiˀˀ, Peter Zwickˁˁ
‡ Abdelmalek Essaadi University, Tetouan, Morocco
§ National Museum and Galleries of Wales, Cardiff, United Kingdom
| Oxford University Museum of Natural History, Oxford, United Kingdom
¶ Zoological Museum Winterthurerstr., Zürich, Switzerland
# University of Kwa Zulu-Natal, Durban, South Africa
¤ Czech University of Life Sciences Prague, Praha-Suchdol, Czech Republic
« Unaffiliated, Barcelona, Spain
» Unaffiliated, Halenkovice, Czech Republic
˄ Università di Roma “La Sapienza”, Rome, Italy
˅ Unaffiliated, Melksham, United Kingdom
¦ Université Mohammed V de Rabat, Rabat, Morocco
ˀ Muséum national d’Histoire naturelle, Paris, France
ˁ Naturalis Biodiversity Center, Leiden, Netherlands
₵ Unaffiliated, Hoevenen, Belgium
ℓ University of Cambridge, Cambridge, United Kingdom
₰ Unaffiliated, La Chaux-de-Fonds, Switzerland
₱ Bishop Museum, Honolulu, United States of America
₳ Unaffiliated, Wickford, United Kingdom
₴ Universidade Federal do Mato Grosso do Sul, Campo Grande, Brazil
₣ Institute of Plant Protection, St. Petersburg-Pushkin, Russia
₮ Muséum d’histoire naturelle, Neuchâtel, Switzerland
₦ California Department of Food & Agriculture, Sacramento, United States of America
₭ National Museums of Scotland, Edinburgh, United Kingdom
₲ Université de Strasbourg, Strasbourg, France
‽ Natural History Museum, Venezia, Italy
₩ Zoological Institute of Russian Academy of Sciences, St. Petersburg, Russia
₸ Voronezh State University, Voronezh, Russia
‡‡ Natural History Museum of Denmark, Copenhagen, Denmark
§§ I.I.Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kyiv, Ukraine
|| University of Stavanger, Stavanger, Norway
¶¶ Unaffiliated, Praha, Czech Republic
## Trinity College, Dublin, Ireland
¤¤ Université de Liège, Gembloux, Belgium
«« Staatliches Museum für Naturkunde Stuttgart, Stuttgart, Germany
»» Unaffiliated, Avignon, France
˄˄ Universität Bielefeld, Bielefeld, Germany
˅˅ Limnologische Fluss-Station des Max-Planck Instituts für Limnologie, Schlitz, Germany
¦¦ Wrocław University of Environmental & Life Sciences, Wrocław, Poland
ˀˀ Opole University, Opole, Poland
ˁˁ Unaffiliated, Schlitz, Germany

Corresponding author: Kawtar Kettani ( kettani.ka@gmail.com )

Academic editor: Torsten Dikow
© 2022 Kawtar Kettani, Martin J. Ebejer, David M. Ackland, Gerhard Bächli, David Barraclough, Miroslav Barták, Miguel Carles-Tolrá, Milos Černý, Pierfilippo Cerretti, Peter Chandler, Mohamed Dakki, Christophe Daugeron, Herman De Jong, Josef Dils, Henry Disney, Boris Droz, Neal Evenhuis, Paul Gatt, Gustavo Graciolli, Igor Y. Grichanov, Jean-Paul Haenni, Martin Hauser, Oumnia Himmi, Iain MacGowan, Bruno Mathieu, Mohamed Mouna, Lorenzo Munari, Emilia P. Nartshuk, Oleg P. Negrobov, Pjotr Oosterbroek, Thomas Pape, Adrian C. Pont, Grigory V. Popov, Knut Rognes, Marcela Skuhravá, Vaclav Skuhravý, Martin Speight, Guy Tomasovic, Bouchra Trari, Hans-Peter Tschorsnig, Jean-Claude Vala, Michael von Tschirnhaus, Rüdiger Wagner, Daniel Whitmore, Andrzej J. Woźnica, Tadeusz Zatwarnicki, Peter Zwick.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation: Kettani K, Ebejer MJ, Ackland DM, Bächli G, Barraclough D, Barták M, Carles-Tolrá M, Černý M, Cerretti P, Chandler P, Dakki M, Daugeron C, De Jong H, Dils J, Disney H, Droz B, Evenhuis N, Gatt P, Graciolli G, Grichanov IY, Haenni J-P, Hauser M, Himmi O, MacGowan I, Mathieu B, Mouna M, Munari L, Nartshuk EP, Negrobov OP, Oosterbroek P, Pape T, Pont AC, Popov GV, Rognes K, Skuhravá M, Skuhravý V, Speight M, Tomasovic G, Trari B, Tschorsnig H-P, Vala J-C, von Tschirnhaus M, Wagner R, Whitmore D, Woźnica AJ, Zatwarnicki T, Zwick P (2022) Catalogue of the Diptera (Insecta) of Morocco— an annotated checklist, with distributions and a bibliography. ZooKeys 1094: 1-466. https://doi.org/10.3897/zookeys.1094.62644
ZooBank: urn:lsid:zoobank.org:pub:1FC0B36F-B7BD-4305-9D8B-642A571A5F69
Open Access

Abstract

The faunistic knowledge of the Diptera of Morocco recorded from 1787 to 2021 is summarized and updated in this first catalogue of Moroccan Diptera species. A total of 3057 species, classified into 948 genera and 93 families (21 Nematocera and 72 Brachycera), are listed. Taxa (superfamily, family, genus and species) have been updated according to current interpretations, based on reviews in the literature, the expertise of authors and contributors, and recently conducted fieldwork. Data to compile this catalogue were primarily gathered from the literature. In total, 1225 references were consulted and some information was also obtained from online databases. Each family was reviewed and the checklist updated by the respective taxon expert(s), including the number of species that can be expected for that family in Morocco. For each valid species, synonyms known to have been used for published records from Morocco are listed under the currently accepted name. Where available, distribution within Morocco is also included. One new combination is proposed: Assuania melanoleuca (Séguy, 1941), comb. nov. (Chloropidae).

Keywords

Bibliography, classification, gnats, midges, new combination, new records, taxonomy, true flies

Foreword

Almost everyone is now aware that there is a biodiversity crisis, and even the popular press regularly reports on general declines in the diversity of species as well as more specific threats to the distribution and abundance of species. Despite this growing awareness of the importance of biodiversity to the health of our planet and thus to the welfare of humankind, relatively little attention has been paid to the glaringly obvious point that it is impossible to objectively assess changes such as declines in diversity in the absence of baseline data and a good taxonomic infrastructure. In other words, if you don’t know what you have to start with it is impossible to know what you are losing (or gaining). Recognition of invasions, extirpations, extinctions, range changes and changes in phenology all pivot on the documentation of what species occur in an area to begin with. Regional catalogues like this one provide both a baseline and a startling wake-up call; in this case with the announcements of 156 previously undocumented species, records of many hitherto unrecorded genera, and the first recognition of half a dozen families new to the country.

Documentation of the dipteran diversity of Morocco is of special significance both in terms of taxon and area. The taxon, Diptera or true flies, is one of the two or three largest orders of living things, with hundreds of thousands of undescribed species worldwide in addition to the approximately 160,000 currently named species. The Moroccan Diptera fauna is relatively well known compared to other parts of Africa, but nonetheless the current catalogue more than doubles the number of species known from the country as of the last inventories a couple of decades ago. This impressive advance in the documentation of Morocco’s significant dipteran fauna is of course in part due to the great dedication and Herculean effort on the part of the lead author, Prof. Kettani. But, it is also a great credit to the community of dipterists who rallied to the cause by meticulously cataloguing their chosen families.

Approximately 33 of the 60 contributors are retired entomologists or amateur dipterists who volunteered their time to contribute to this work. Sadly, a few have passed away since they made their contribution. It is both impressive and worrisome that more than half of those 60 top experts on the taxonomy of the region’s Diptera are retired or amateur, since there is a pressing and ongoing need for specialized expertise on all of the families of Diptera. Hopefully the framework, and the challenge, provided by this thorough description of our current state of knowledge of Moroccan flies will help and encourage new students of dipterology while at the same time providing a foundation for ongoing assessments of the dipteran diversity of North Africa.

I congratulate all involved for the production of such an important contribution to our understanding of the biodiversity of the region, and for the successful completion of the first full Diptera catalogue for any African country.

Stephen A. Marshall

Professor, University of Guelph, Canada

October 2020

Contributions

This catalogue was conceived by K. Kettani (KK) in 2011 and then further developed and planned together with Martin J. Ebejer (MJE). These plans included more fieldwork, predominantly by KK and MJE, with some contribution by students, carried out in 2012–2019, which resulted in about 1000 new species records for Morocco and 23 species (in 12 families) new to science. These records were published prior to the end of 2020 and the papers are included in the references and cited under the appropriate families. KK compiled the data and the preliminary list of species for each family. The taxonomic specialists (co-authors and contributors) revised any taxonomic issues, added data available to them and updated nomenclature. Contributors are thanked in the Acknowledgements, as well as mentioned in their sections of the checklist. The distributions of species in Morocco, with the appropriate citations, were added by KK. The introduction was written by KK and supplemented with additional information by MJE, who also checked the English language.

Introduction

Aims of the catalogue

Although the national study on biodiversity, Etude nationale sur la biodiversité (Dakki 1997; Mouna 1998), was the only important synthesis of entomological studies carried out in Morocco until now, important data on the Diptera are lacking. For example, no taxonomic rank such as subfamily and tribe, and no synonyms or revised generic combinations, names of museums or private collections that house species described from Morocco were included. No information was provided on the habitats and distribution of species in Morocco. Furthermore, the information resulting from the national study on biodiversity is now outdated. A large number of important recent publications, cited appropriately under each family, provide much additional data to the lists published in 1997 and 1998, almost doubling the number of known species. The main aims of this catalogue are to bring together all this information by providing an up-to-date checklist according to the current nomenclature and systematics, to provide synonyms used in older literature that dealt with the Moroccan fauna and to give regional distribution data. Finally, an essential comprehensive bibliography is provided.

This catalogue is the first for a North African country and it may prove useful also for neighbouring countries. It is hoped that it will provide a sound foundation for anyone choosing to study a group of Moroccan Diptera. It could indicate where the greatest need for investigation lies, both with regard to areas of the country that have been poorly studied and taxonomic groups (or families) that may benefit from more attention. It is undoubtedly not a complete inventory and many more discoveries are expected.

The faunistic knowledge of the Diptera of Morocco from 1787 to 2021 is summarized and updated, providing the first catalogue of the Moroccan species of Diptera. In total, 3057 species (975 Nematocera and 2082 Brachycera) in 93 families, classified into 949 genera (250 in Nematocera and 699 in Brachycera) are listed (Tables 1, 2).

Table 1.
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Families of Nematocera with contributing specialists and numbers of taxa.

Family Specialist(s) Subfamilies Genera Species
Anisopodidae Haenni 1 1 3
Bibionidae Haenni 1 2 10
Blephariceridae Zwick 1 1 4
Cecidomyiidae Skuhravá & Skuhravý 3 31 57
Ceratopogonidae Mathieu 3 6 62
Chaoboridae Wagner 1 2 2
Chironomidae - 7 100 412
Culicidae Trari, Himmi & Dakki 2 7 43
Dixidae Wagner - 2 12
Keroplatidae Chandler 1 2 2
Limoniidae Oosterbroek 4 22 67
Mycetophilidae Chandler 5 25 64
Pediciidae Oosterbroek 1 2 6
Psychodidae Wagner 2 12 51
Ptychopteridae Wagner 1 1 5
Scatopsidae Haenni 3 11 13
Sciaridae Heller - 12 70
Simuliidae - 1 6 43
Thaumaleidae Wagner - 1 2
Tipulidae Oosterbroek & de Jong 2 3 39
Trichoceridae Krzemińska 1 1 8
Total: 250 Total: 975
Table 2.
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Families of Brachycera with contributing specialists and numbers of taxa.

Family Specialist(s) Subfamilies Genera Species
Acroceridae Nartshuk 2 6 13
Agromyzidae Černý 2 12 62
Anthomyiidae Ackland 2 14 36
Anthomyzidae Ebejer - 2 2
Asilidae Tomasovic 9 42 131
Asteiidae Ebejer 1 2 5
Atelestidae Gatt - 1 1
Athericidae Mouna 1 1 2
Aulacigastridae Ebejer - 1 1
Bombyliidae Ebejer & Dils 11 48 248
Braulidae - 1 1 1
Calliphoridae Rognes 5 5 8
Camillidae Ebejer - 1 4
Canacidae Munari 2 3 15
Carnidae Ebejer - 1 3
Chamaemyiidae Ebejer 2 5 18
Chloropidae von Tschirnhaus & Ebejer 3 34 74
Chyromyidae Ebejer 2 3 22
Clusiidae Ebejer - 1 1
Coelopidae - 1 1 1
Conopidae - 5 10 34
Cryptochetidae Nartshuk - 1 1
Diastatidae Ebejer 2 2 2
Dolichopodidae Grichanov & Negrobov 12 36 112
Drosophilidae Bächli 2 9 26
Dryomyzidae - 1 1 1
Empididae Daugeron 3 8 40
Ephydridae Zatwarnicki 5 43 117
Fanniidae Pont - 1 10
Helcomyzidae - 1 1 1
Heleomyzidae Woźnica 3 5 19
Hippoboscidae Droz 3 10 17
Hybotidae Gatt 2 10 44
Lauxaniidae Ebejer 2 8 27
Lonchaeidae MacGowan 1 4 5
Lonchopteridae Barták 1 1 4
Micropezidae Ebejer - 1 1
Milichiidae - 3 6 8
Muscidae Pont 6 25 115
Mydidae Dikow 3 4 9
Mythicomyiidae Evenhuis 4 6 8
Nemestrinidae Barraclough 2 4 13
Nycteribiidae Graciolli 1 4 8
Odiniidae Ebejer 1 1 2
Oestridae Pape 2 5 10
Opomyzidae - - 2 5
Pallopteridae Ebejer - 1 1
Phoridae Disney - 2 3
Piophilidae Ebejer 1 3 3
Pipunculidae Ebejer 2 7 16
Platypezidae Ebejer 1 3 3
Platystomatidae Popov 1 2 4
Polleniidae Rognes - 1 12
Psilidae Ebejer - 1 1
Rhagionidae Ebejer 1 1 4
Rhiniidae Rognes 1 5 17
Rhinophoridae Pape 1 6 8
Sarcophagidae Whitmore & Pape 3 16 66
Scathophagidae - 1 2 3
Scenopinidae Carles-Tolrá 1 2 11
Sciomyzidae Vala 1 12 25
Sepsidae Haenni 1 4 12
Sphaeroceridae Gatt 3 29 67
Stratiomyidae Woodley 5 11 40
Streblidae Graciolli 1 2 2
Syrphidae Speight 2 50 166
Tabanidae - 3 11 69
Tachinidae Cerretti & Tschorsnig 4 88 147
Tephritidae Norrbom 3 29 69
Therevidae Hauser 2 9 27
Ulidiidae Ebejer 2 8 13
Vermileonidae Ebejer 1 2 6
Total: 698 Total: 2082

Brief history of Moroccan dipterology

The study of Diptera in Morocco is still far from being extensive despite the numerous investigations that have taken place so far. Until now, a synthesis of Moroccan Diptera had never been produced. The studies that have been devoted to this group of insects have been limited and sporadic. They were mainly concerned with either taxonomy and descriptions of new species, or faunal studies from specific areas of Morocco with little reference to the habitats of species. This was particularly so for early works such as Becker and Stein (1913), Séguy (1930a, 1934b, 1935a, 1941a, 1941d, 1949a, 1953a), Timon-David (1951), Vaillant (1956b) and, recently, Pârvu et al. (2006), Popescu-Mirceni (2011), and Ebejer et al. (2019). Very few recent works have provided relevant ecological data (Cassar et al. 2005, 2008). Becker and Stein (1913) were the first to publish a list of Moroccan Diptera, at that time comprising only 204 species. Séguy followed with a long series of valuable publications. In 1930 he produced his first list of Diptera, which included 471 species in 230 genera grouped into 37 families (Séguy 1930). Over the subsequent years, he contributed greatly to the knowledge of the Diptera of Morocco (Séguy 1926, 1934b, 1935a, 1941a, 1941d, 1949a, 1953a, 1957, 1961), where he also referenced an extensive bibliography. Later, Bailly-Choumara (1977) compiled a bibliography of Moroccan Diptera. It took until 1997–1998, with a national study on the biodiversity of Morocco co-ordinated by what was then the Ministry of the Environment, to update the list of the Moroccan insect fauna. This national study was dedicated to the compilation of the results of entomological research conducted in Morocco in order to highlight Moroccan biodiversity. It included a comprehensive inventory of published entomological research conducted in Morocco and was the last work to include a broad overview of Moroccan Diptera. It listed 623 species of aquatic Diptera in 210 genera and 25 families (Dakki 1997) and 928 species of terrestrial Diptera in 350 genera and 57 families (Mouna 1998).

Terrain and bioclimatic regions

Morocco, located in the westernmost corner of North Africa, constitutes a biogeographical crossroads between the Afrotropical and the western Palaearctic faunas, which may have allowed some exchange of genetic material. The country has a remarkable variety of bioclimates, ranging from humid in the Rif and the Middle and High Atlas to very arid in the Sahara in the south and to the sub-humid and semi-arid plains and foothills (Fig. 1). As a result of this diversity of landscapes and climates, there exists a great biological and ecological diversity in the country.

Figure 1. 

The major bioclimatic regions of the northern part of Morocco.

The geomorphological, orographic and bioclimatic diversity of the country is reflected in its seven distinct biogeographical areas: the Rif, the Atlantic Plain, Eastern Morocco, the Middle Atlas, the High Atlas, the Anti-Atlas and the Sahara. In the checklist, an abbreviated distribution (in bold) is given for each species according to these seven biogeographical areas (Fig. 2).

Figure 2. 

The seven biogeographical regions covering the studied area in the northern part of Morocco.

Rif: The Rif consists of a mountainous domain overlooking the Mediterranean Sea in the north of Morocco. It belongs to the Gibraltar Arc or Alborán Sea geological region, part of the Alpine orogenic belt, where the maximum altitude is 2,452 m in the Jbel Tidirrhine mountain range. The climate is mainly of the Mediterranean type and is characterised by high levels of precipitation. This domain is considered very rich and diverse in vegetation cover (Valdés et al. 2002), including nearly all of the Moroccan forest plant species (Médail and Quézel 1999) and integrating five bioclimatic stages.

Atlantic Plain (AP): Also known as the Moroccan Central Plateau, the Atlantic Plain is an ancient massif located in the northwest of Morocco, between the Atlantic coast and the Middle Atlas Mountains. It consists of a plateau surrounded by medium-altitude mountain massifs and covers an area of 8500 km². With an asymmetrical shape, the plateau culminates in the south-east at Jebel Mtourzgene (1627 m), located near the town of Oulmès. Its complex geomorphology presents tectonic troughs that have favoured the formation of lakes and rivers. The annual rainfall of this region, concentrated in winter and spring, totals an average of 500 mm, allowing the establishment of Holm Oak and Cork Oak forests in a semi-arid bioclimate (Piqué 1994). Towards the coast, the vegetation is mainly a steppe with Jujube shrubs and Dwarf Palms (Chamaerops humilis). The maritime fringe is underlined by cliffs and ancient consolidated dunes.

Eastern Morocco (EM): This is an arid region covering the Hauts Plateaux to the northeast of the country. It is the only region of Morocco subject to a true Mediterranean bioclimate, with some influence from the Sahara. The annual rainfall is less than 300 mm, encouraging the dominance of esparto steppe (Stipa tenacissima) (Piqué 1994). Its geomorphology is represented by a plateau at an altitude of 1000 m, which gradually decreases from 1500 to 500 m from the city of Midelt to the province of Guercif and is criss-crossed by the Moulouya wadi. In the northwestern part of Eastern Morocco is the Béni Snassen massif, beyond the Guercif depression.

Middle Atlas (MA): A mountainous massif located in the centre of Morocco, oriented from south-west to north-east and extending over 350 km (Piqué 1994). This mass of highlands is essentially composed of limestone layers, tabular and of medium altitude in the southwest (tabular Middle Atlas) and undulating in the northeast (folded Middle Atlas) to culminate at 3340 m at Jebel Bou Naceur, which overlooks the high plateaux of Eastern Morocco. The climate, generally subhumid to humid, is cold in winter, giving rise to cedar and Holm Oak forests covering the western slope, which is wetter than the eastern slope and where lakes are abundant and many rivers and streams have their source.

High Atlas (HA): The main mountainous massif of Morocco, oriented from west-south-west to east-north-east, is the High Atlas, stretching over more than 700 km from the Atlantic to the east of Morocco, with a north-south width of 50 to 100 km and reaching at Jebel Toubkal an altitude of 4165 m. The whole massif is high, incised by deep valleys that lead to passes through which the mountainous barrier is crossed: the Tizi-n’Test (2100 m) towards the Souss plain and the Tizi-n’Tichka (2200 m) towards the depression of Ouarzazate. The chain is formed by a massif of Palaeozoic rocks surrounded by sedimentary deposits. The relief of the High Atlas is divided into three different entities from west to east: the Western High Atlas, which is the oldest massif and is made up mainly of Jurassic or Cretaceous formations with deep valleys and culminates at Jebel Toubkal; the central High Atlas, which is an essentially limestone massif morphologically dominated by tabular zones culminating at an altitude of 2500 m, stretches between the towns of Azilal and Ouarzazate and is sheltered by the Jebel M’Goum (4071 m); and the Eastern High Atlas, which is formed by the vast high plateaux of the Upper Moulouya, stretches between the cities of Midelt and Imilchil and contains the Jebel Ayachi (3747 m) (Piqué 1994). The bioclimate of this area includes subhumid, humid and semi-arid zones, but with more than 800 mm of annual precipitation on the highest peaks and a significant snow cover from September to May, with temperatures dropping below -20 °C (AEFCS 1996) and bright sunshine even in winter. These climatic conditions have favoured large pine, Holm Oak, Thuja and thuriferous juniper forests (HCEFLCD 2017). The aridity of the southern slope of the High Atlas allows only steppe vegetation of sagebrush and esparto grass to grow.

Anti-Atlas (AA): This is the pre-Saharan area, which extends north of the Wadi Draa and rises to an average altitude of 2000 m. It is a semi-arid region with Jebel Bani and Jebel Ouarkziz as its main elevations, and is home to numerous oases. The western part of the Anti-Atlas is characterised by the Saharan plateau, with a temperate winter except in the high mountains. On the other hand, in the eastern part of the Anti-Atlas, the greatest arid area is marked by the northward extension of the Saharan landscape and a bioclimate of cool and temperate winters. These bioclimatic stages are characterised by plant communities such as wooded steppes with Acacia raddiana, Artemisia herba alba and Stipa tenuissima, interspersed with thuriferous juniper (Juniperus thurifera) at the top of the slopes (Piqué 1994). The vegetation is also represented by two formations that partially interpenetrate each other: groves of the Argan tree (Argania spinosa) and the Euphorbia steppe (Euphorbia echinus).

Sahara (SA): This area in the south of the country consists of a wide arid zone with a typical Saharan bioclimate of high daytime temperatures and very low precipitation. From a geological point of view it is part of the West African Craton and its margins. It is mainly ancient terrain with Precambrian basement rock and Palaeozoic cover (Piqué 1994). This area is represented by a peneplain of very modest altitudes from 200 to 500 m, established on ancient crystalline terrain. Precipitation from July to September is very low, of the order of 100 mm.

The dipteran species richness of each biogeographical region of Morocco is summarised in Table 3, which highlights the great disparity between the different regions and reflects the lack of knowledge about this insect order in some areas of Morocco. The Rif region in the north of the country has the greatest recorded diversity, but this undoubtedly reflects a greater sampling effort in this area, with large parts of the country still very poorly explored for Diptera. The catalogue does not include any data from the southern part of the Moroccan Sahara.

Table 3.
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Species richness of Diptera families each biogeographical region in Morocco. Rif; AP – Atlantic Plain; EM – Eastern Morocco; MA – Middle Atlas; HA – High Atlas; AA – Anti-Atlas; SA – Sahara. * No precise localities known.

Rif AP EM MA HA AA SA
Acroceridae 7 1 0 1 3 0 0
Agromyzidae 7 15 0 10 22 10 0
Anisopodidae *
Anthomyiidae 4 12 0 13 2 4 1
Anthomyzidae 1 0 0 0 1 0 0
Asilidae 34 27 4 35 25 24 5
Asteiidae 0 3 0 0 1 1 0
Atelestidae 1 0 0 0 0 0 0
Athericidae 0 0 0 1 2 0 0
Aulacigastridae 1 0 0 0 0 0 0
Bibionidae 4 2 0 3 1 2 2
Blephariceridae 1 0 0 0 3 0 0
Bombyliidae 35 80 12 59 48 56 19
Braulidae *
Calliphoridae 3 8 2 4 4 0 0
Camillidae 1 2 0 0 1 0 0
Canacidae 6 10 0 0 0 6 0
Carnidae 1 0 0 0 0 0 0
Cecidomyiidae 8 27 21 25 20 7 4
Ceratopogonidae 15 43 14 35 33 24 36
Chamaemyiidae 5 5 0 4 8 6 0
Chaoboridae *
Chironomidae 230 30 7 105 120 14 1
Chloropidae 35 22 0 7 6 25 1
Chyromyidae 6 6 0 0 1 12 1
Clusiidae 1 0 0 0 0 0 0
Coelopidae 1 0 0 0 0 0 0
Conopidae 7 13 0 16 7 9 0
Cryptochetidae 0 0 0 1 0 0 0
Culicidae 32 41 24 30 29 22 11
Diastatidae 0 1 0 0 1 0 0
Dixidae 2 2 0 2 5 0 0
Dolichopodidae 72 28 3 6 18 26 0
Drosophilidae 11 14 0 8 12 5 1
Dryomyzidae 1 0 0 0 0 0 0
Empididae 4 3 0 13 20 1 0
Ephydridae 47 44 1 60 9 20 1
Fanniidae 4 2 0 2 5 0 0
Helcomyzidae 1 0 0 0 0 0 0
Heleomyzidae 11 3 0 2 2 3 0
Hippoboscidae 0 2 3 4 4 4 1
Hybotidae 20 7 3 8 7 8 0
Keroplatidae 1 0 0 0 0 0 0
Lauxaniidae 23 3 2 1 8 2 0
Limoniidae 43 9 4 5 34 5 0
Lonchaeidae 3 3 0 1 1 0 0
Lonchopteridae 0 0 0 1 1 0 0
Micropezidae 1 0 0 0 0 0 0
Milichiidae 2 1 0 0 4 3 1
Muscidae 25 47 13 38 68 20 6
Mycetophilidae 60 3 7 3 0 0 0
Mydidae 0 3 2 1 0 3 1
Mythicomyiidae 2 0 0 0 1 2 1
Nemestrinidae 1 1 1 5 3 0 1
Nycteribiidae 6 5 0 6 0 2 1
Odiniidae 1 0 0 0 0 0 0
Oestridae 0 2 1 1 1 0 1
Opomyzidae 1 1 0 1 2 0 0
Pallopteridae 0 0 0 1 0 0 0
Pediciidae 3 1 0 0 0 0 0
Phoridae 0 2 0 0 0 0 0
Piophilidae 2 1 0 1 1 0 0
Pipunculidae 13 1 0 0 0 0 0
Platypezidae 2 1 0 0 0 0 0
Platystomatidae 0 2 1 1 0 0 0
Polleniidae 1 4 2 5 5 1 1
Psilidae 1 0 0 1 0 0 0
Psychodidae 25 12 12 12 34 9 2
Ptychopteridae *
Rhagionidae 2 0 0 0 2 0 0
Rhiniidae 2 5 3 5 0 5 4
Rhinophoridae 6 2 0 1 0 0 0
Sarcophagidae 11 19 1 13 8 9 2
Scathophagidae 3 1 0 1 0 0 0
Scatopsidae 10 2 0 0 1 3 0
Scenopinidae 3 4 1 1 0 0 1
Sciaridae 29 12 2 8 45 9 3
Sciomyzidae 8 4 2 11 18 3 0
Sepsidae 3 6 0 4 7 3 0
Simuliidae 32 8 0 19 32 6 0
Sphaeroceridae 42 3 2 10 1 3 3
Stratiomyidae 20 13 5 13 6 1 0
Streblidae 1 1 0 0 0 1 0
Syrphidae 70 49 11 82 64 25 4
Tabanidae 26 27 11 35 21 7 1
Tachinidae 18 27 4 46 43 29 3
Tephritidae 34 13 4 16 13 23 3
Thaumaleidae 0 0 0 0 2 0 0
Therevidae 9 7 2 9 4 5 2
Tipulidae 28 7 0 16 18 2 0
Trichoceridae 7 1 3 1 0 0 0
Ulidiidae 6 2 0 1 4 4 1
Vermileonidae 1 2 1 1 4 0 0
Total species 1126 770 191 831 876 474 126

Sources of data

Data for the present study were gathered from the literature, supplemented by information taken from a number of websites. The cut-off date for inclusion of species published in the literature is 30 June 2021. Taxonomic, nomenclatural, and distributional data for the present catalogue were obtained from:

  • studies by various foreign and Moroccan researchers published between 1787 and 2021, including theses, books, catalogues, checklists and notes from the 1997–1998 synthesis studies on Morocco’s biodiversity.
  • data from websites and organisations such as Global Species, the Global Biodiversity Information Facility, the BioSystematic Database of World Diptera, Systema Dipterorum, Catalogue of Life, Systema Naturae 2000 and others, or from relevant homepages specific to particular families.
  • the collection of Diptera in the National Museum of Morocco (Muséum National d’Histoire Naturelle à l’Institut Scientifique, Université Mohammed V, Rabat).
  • collaboration with specialists who communicated reliable identifications from their personal collections or provided data for species deposited in foreign museums.
  • published records of specimens in the personal collection of the lead author (PCKK), deposited at Abdelmalek Essaadi University, Tetouan, identified with the assistance of specialists.

Classification

The arrangement of families in this catalogue (Table 4) is mainly based on Pape et al. (2011) and Wiegmann et al. (2011). All families are listed alphabetically within each superfamily. Nomenclature follows the most recent version of Systema Dipterorum (Evenhuis and Pape 2021) and the Polleniidae are listed as a separate family (Gisondi et al. 2020).

Table 4.
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Diptera classification used in the catalogue.

Suborder NEMATOCERA
Tipuloidea Limoniidae
Pediciidae
Tipulidae
Trichoceridoidea Trichoceridae
Psychodoidea Psychodidae
Scatopsoidea Scatopsidae
Ptychopteridae
Culicoidea Chaoboridae
Culicidae
Dixidae
Chironomoidea Ceratopogonidae
Chironomidae
Simuliidae
Thaumaleidae
Blephariceridae
Bibionoidea Anisopodidae
Bibionidae
Sciaroidea Cecidomyiidae
Keroplatidae
Mycetophilidae
Sciaridae
Suborder BRACHYCERA
Stratiomyoidea Stratiomyidae
Tabanoidea Athericidae
Rhagionidae
Tabanidae
Vermileonidae
Nemestrinoidea Acroceridae
Nemestrinidae
Asiloidea Asilidae
Bombyliidae
Mydidae
Mythicomyiidae
Scenopinidae
Therevidae
Empidoidea Atelestidae
Empididae
Dolichopodidae
Hybotidae
Platypezoidea Phoridae
Platypezidae
Lonchopteridae
Syrphoidea Pipunculidae
Syrphidae
Conopoidea Conopidae
Nerioidea Micropezidae
Tanypezoidea Psilidae
Tephritoidea Lonchaeidae
Pallopteridae
Piophilidae
Platystomatidae
Tephritidae
Ulidiidae
Lauxanioidea Chamaemyiidae
Lauxaniidae
Sciomyzoidea Coelopidae
Dryomyzidae
Helcomyzidae
Sciomyzidae
Sepsidae
Opomyzoidea Agromyzidae
Anthomyzidae
Asteiidae
Aulacigastridae
Clusiidae
Odiniidae
Opomyzidae
Carnoidea Canacidae
Carnidae
Chloropidae
Milichiidae
Sphaeroceroidea Chyromyidae
Heleomyzidae
Sphaeroceridae
Ephydroidea Braulidae
Camillidae
Cryptochetidae
Diastatidae
Drosophilidae
Ephydridae
Hippoboscoidea Hippoboscidae
Nycteribiidae
Streblidae
Muscoidea Anthomyiidae
Fanniidae
Muscoidea Muscidae
Scathophagidae
Oestroidea Calliphoridae
Oestridae
Polleniidae
Rhiniidae
Rhinophoridae
Sarcophagidae
Tachinidae

Arrangement of the catalogue

The catalogue includes all names of taxonomically valid species of the Order Diptera so far known to occur in Morocco. For each family, synonyms and alternative combinations that have been published as records of Moroccan provenance are listed under the currently accepted name.

The transliteration of Moroccan place names in the literature, maps, and sometimes on data labels presented a small problem. Although it would have been preferable to use standard names, this would have meant changing some place names from their original spellings in the literature. Most transliteration used the French language for spelling and Roman letters, but not all. English was sometimes used, and occasionally also Spanish. Readers are therefore advised to take this into account when searching for place names given in the catalogue and in the cited references. Some common examples are “Jebel”, “Jbel”, “Djebel” or “Dj.”, used for “mountain”, or “Kbir” and “Kebir”, for “large” or “great”. The article “the” in Arabic is El or L; it is sometimes separated by a hyphen from its associated noun or joined to it, sometimes capitalised while the noun is not, both are capitalised, or only the noun is. When the article “el” is followed by a noun beginning with a certain consonant, like n, r, s and t, the l is changed to match the noun’s initial, for example “Er-Rifiyine”. In transliteration, Arabic speakers would make this change, whereas non-Arabic speakers may not. In cases where the article is joined to the noun, the resulting versions may appear as being different place names; for example, El Rachidia, Er Rachidia and Errachidia are alternative transliterations of the same place name. In some entries, the French terms “affluent” and “aval affluent” are left as in the original citation, because “tributary” may not be an accurate translation in all instances. For some of the sites, the French term refers to a downstream overflow of water into a side tributary or waterfall, whereas for others it refers to an actual tributary feeding into a main river or stream.

Older authors were imprecise with localities, and many references did not give the coordinates of localities from which species were collected. We could not reliably provide accurate coordinates for each species. Where known, species depositories are included. Individual family checklists may include, at the end, taxonomic notes and comments on species doubtfully present in the country. New records are indicated with an asterisk (*) and the relevant data listed at the end of the family section.

Limitations

One of the major difficulties encountered while compiling the catalogue was the limited ability to check old identifications. In some cases, identifications were accepted as given in the original sources, even when doubtful, as it was usually not possible to re-examine specimens. The inaccessibility of some references was also an obstacle to updating the checklists for some families.

Collections cited

The following museums and collections house some of the specimens mentioned in this catalogue. The National Museum of Natural History of Morocco is located in the Institut Scientifique, Université Mohammed V, Rabat. However, the Diptera collection housed in that museum is not yet included in any world database such as the GBIF Registry of Scientific Collections, https://www.gbif.org/grscicoll, or the Insect and Spider collections of the World Website, http://hbs.bishopmuseum.org/codens/.

HNHM Hungarian Natural History Museum, Budapest, Hungary;

INHS Illinois Natural History Survey, Urbana, USA;

IRSNB Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium;

MfN Museum für Naturkunde, Berlin, Germany;

MHNN Muséum d’Histoire Naturelle de Neuchâtel, Neuchâtel, Switzerland;

MISR Museum National d’Histoire Naturelle à l’Institut Scientifique, Université Mohammed V, Rabat, Morocco;

MNCN Museo Nacional de Ciencias Naturales, Madrid, Spain;

MNHN Muséum National d’Histoire Naturelle, Paris, France;

NHMD Natural History Museum of Denmark, Copenhagen, Denmark;

NHMUK Natural History Museum, London, United Kingdom;

NMWC National Museum of Wales, Cardiff, United Kingdom;

OUMNH Oxford University Museum of Natural History, Oxford, United Kingdom;

RMNH Rijksmuseum voor Natuurlijke Historie, Leiden, The Netherlands;

SMNS Staatliches Museum für Naturkunde Stuttgart, Stuttgart, Germany;

ZMUM Zoological Museum of Moscow University, Moscow, Russia;

ZSM Zoologische Staatssammlung, Munich, Germany;

ZSM Zoologische Staatssammlung, Munich, Germany.

The personal collection of Dr Michael von Tschirnhaus has been donated to the ZSM, but is currently on loan to Dr von Tschirnhaus.

Other abbreviations

HT Holotype;

NPH National Park of Al Hoceima;

NPHAO National Park of Haut Atlas Oriental;

NPI National Park of Ifrane;

NPTNational Park of Talassemtane;

PCJD Private Collection of Jos Dils, Belgium;

PCKK Private Collection of Kawtar Kettani, Morocco;

PNPB Project of Natural Park of Bouhachem.

Acknowledgements

The authors are deeply grateful to the reviewers for their constructive comments, which helped to improve the manuscript. Our sincere thanks are expressed to the subject editor, Torsten Dikow, for his significant contribution and support as editor.

We are most grateful to the following specialists who generously contributed in various ways to this catalogue by sharing information and knowledge of their group or by reviewing a family section: Patrick Ashe (Dublin, Ireland)—Chironomidae; Torsten Dikow (Washington, D.C., USA)—Mydidae; the late Amnon Freidberg (Tel Aviv, Israel)—Tephritidae; Kai Heller (Heikendorf, Germany)—Sciaridae; Gail Kampmeier (Champaign, USA)—Therevidae; Valery Korneyev (Kiev, Ukraine)—Tephritidae; Ewa Krzemińska (Kraków, Poland)—Trichoceridae; Allen Norrbom (Washington, D.C., USA)—Tephritidae; Adrian R. Plant (Mahasarakham, Thailand)—Empididae, Hemerodrominae; Bradley J. Sinclair (Ottawa, Canada)—Empididae, Clinocerinae; Jens-Hermann Stuke (Leer, Germany)—Conopidae; Norman Woodley (Hereford, AZ, USA)—Stratiomyidae. Mohamed Mouna and Mohamed Dakki are acknowledged and thanked for their encouragement and inspiration that led the first author to commence this project, as well as for generously donating to her a large bibliography on the subject. Mohamed Mouna was instrumental in ensuring that she had full access to the collections and library and full cooperation from the staff of these departments. He is immensely thanked by the first author for donating to her his library, consisting of many books and publications.

Mohammed Fekhaoui, Abdellatif Bayed, Mohamed-Aziz El Agbani, Mohamed Arahou from the administration of Museum National d’Histoire Naturelle à l’Institut Scientifique, Université Mohammed V, Rabat are warmly thanked for their valuable assistance in providing access to the library and Diptera collections of the museum. Thanks also to Amina Zouida, the librarian at the said institute, for her tireless and precious help in searching for obscure publications, and to Abderrahmane Mataame (responsible for the zoological collections of the museum) for his kind and patient assistance during the first author’s frequent visits to the museum. Yassine Fekrani is acknowledged for his kind help producing the maps.

Suborder NEMATOCERA

Tipuloidea

LIMONIIDAE

K. Kettani, P. Oosterbroek

Number of species: 67. Expected: 85

Faunistic knowledge of the family in Morocco: moderate

Chioneinae

Baeoura Alexander, 1924

Baeoura ebenina Starý, 1981

Driauach and Belqat 2015, Rif, Oued Tazarine (Mezine village); Dayat near Aïn Afersiw; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: Rif)

Baeoura staryi Driauach & Belqat, 2015

Driauach and Belqat 2015, Rif, Jnane Niche; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: Rif)

Cheilotrichia Rossi, 1848

Cheilotrichia (Empeda) cinerascens (Meigen, 1804)

Driauach et al. 2013, Rif, Ikadjiouen; Driauach and Belqat 2016, Rif, Oued Ouara, Oued Amsa, Oued Maggou, Aïn Quanquben (Jebel Bou Bessoui), EM, Grotte du Chameau (Zegzel, Béni Snassen); Oosterbroek 2020 (CCW: Rif; EM)

Cheilotrichia (Empeda) fuscohalterata (Strobl, 1906)

Driauach and Belqat 2016, Rif, Dayat Fifi, tributary of Oued El Fondak, Barrage Ajras, Dayat Mghara, Oued Zarka, Oued Tizekhte; Oosterbroek 2020 (CCW: Rif)

Cheilotrichia (Empeda) minima (Strobl, 1898)

Driauach and Belqat 2016, Rif, Oued Zendoula, Oued Jnane Niche; Oosterbroek 2020 (CCW: Rif)

Ellipteroides Becker, 1907

Ellipteroides (Ellipteroides) lateralis (Macquart, 1835)

= Gonomyia lateralis Macquart, in Pierre 1922a: 22, Dakki 1997: 62

= Gonomyia cincta Egger, in Séguy 1941a: 26

Pierre 1922a, AP, Dradek (near Rabat); Lackschewitz 1940a, HA, Tachdirt (2200–2900 m); Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Savchenko et al. 1992; Dakki 1997; Starý and Freidberg 2007; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Dayat Afrate; Oosterbroek 2020 (CCW: Rif)

Ellipteroides (Protogonomyia) alboscutellatus (von Roser, 1840)

Lachschewitz 1940a, HA; Savchenko et al. 1992; Starý and Oosterbroek 2008; Starý 2009a; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Ellipteroides (Protogonomyia) hutsoni (Starý, 1971)

Starý 1971, HA, Jebel Ayachi; Savchenko et al. 1992, HA, Jebel Ayachi; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Erioconopa Starý, 1976

Erioconopa diuturna (Walker, 1848)

Driauach et al. 2013; Driauach and Belqat 2016, Rif, Dayat Jebel Zemzem, Oued Mezine, Aïn Bab Tariouant; Oosterbroek 2020 (CCW: Rif)

Erioconopa symplectoides (Kuntze, 1914)

= Dactylolabis symplectoides Egger, in Pierre 1922a: 23

Pierre 1922a, HA, Marrakech; Savchenko et al. 1992; Starý and Oosterbroek 2008, HA; Gavryushin pers. comm. 2012, HA; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Erioptera Meigen, 1803

Erioptera (Erioptera) fuscipennis Meigen, 1818

Pierre 1922a, AP, Casablanca (Oued Guerera); Pierre 1922b, HA, Savchenko et al. 1992; Dakki 1997; Driauach et al. 2013, Rif, Sidi Brahim Ben Arrif (Bab Hachef Aissa); Driauach and Belqat 2016, Rif, Aïn El Ma Bared, Oued Amsemlil, Oued Maggou (Bridge), Oued Tkarae, Dayat near Aïn Afersiw, Aïn Afersiw, Oued Abou Bnar, Oued Maggou (Zaouiet El Habtiyne), Dayat Amsemlil, Dayat Aïn Jdi­oui; Dayat Afrate, Oued Mezine; Oosterbroek 2020 (CCW: Rif, HA)

Erioptera (Erioptera) lutea lutea Meigen, 1804

Driauach and Belqat 2016, Rif, Aïn Boughaba; Oosterbroek 2020 (CCW: Rif)

Erioptera (Erioptera) transmarina Bergroth, 1889

= Mesocyphona transmarina Bergroth, in Pierre 1922a: 22, Pierre 1922b: 148, Dakki 1997: 62

Pierre 1922a, HA, Marrakech; Pierre 1922b, HA, Tannaout (1000 m), Marrakech; Dakki 1997; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Gonomyia Meigen, 1818

Gonomyia (Gonomyia) abscondita Lackschewitz, 1935

Driauach and Belqat 2016, Rif, maison forestière; Oosterbroek 2020 (CCW: Rif)

Gonomyia (Gonomyia) sicula Lackschewitz, 1940

Driauach and Belqat 2016, Rif, Oued Kbir, Dayat Jebel Zemzem, Aïn Sidi Brahim Ben Arrif, Dayat Tazia, Aïn El Maounzil; Oosterbroek 2020 (CCW: Rif)

Gonomyia (Gonomyia) subtenella Savchenko, 1972

Starý and Oosterbroek 2008, HA, Massif Toubkal, Aghbalou, 43 km S Marrakech (1000 m); Starý 2009a; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Jnane Niche, Oued Sidi Yahia Aârab, EM, Oued Béni Ouachekradi; Oosterbroek 2020 (CCW: Rif, EM)

Gonomyia (Gonomyia) tenella (Meigen, 1818)

Pierre 1924a, HA, Asni (1200 m); Séguy 1930a, HA, Asni; Savchenko et al. 1992; Dakki 1997; Starý 2009a; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Hoplolabis Osten-Sacken, 1869

Hoplolabis (Parilisia) obtusiapex (Savchenko, 1982)

Starý 2006, HA, Oasis Meski, AA; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Hoplolabis (Parilisia) punctigera (Lackschewitz, 1940)

Starý 2006, HA, Oasis Meski, AA; Koçak and Kemal 2010; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Hoplolabis (Parilisia) sororcula (Lackschewitz, 1940)

Driauach et al. 2013; Driauach and Belqat 2016, Rif, Barrage Ajras, Oued Ouringa, Oued El Kanar, Oued Maggou; Oosterbroek 2020 (CCW: Rif)

Idiocera Dale, 1842

Idiocera (Euptilostena) jucunda (Loew, 1873)

= Gonomyia jucunda Loew, in Pierre 1924a: 201, Séguy 1930a: 22

Pierre 1924a, HA, Asni (1200 m); Séguy 1930a, HA, Asni; Savchenko et al. 1992; Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Idiocera (Idiocera) ampullifera (Starý, 1979)

Driauach and Belqat 2016, AA, Oued Zag; Oosterbroek 2020 (CCW: AA)

Idiocera (Idiocera) pulchripennis (Loew, 1856)

= Gonomyia sexpunctata Dale, in Pierre 1922b: 148

= Gonomyia pulchripennis (Loew), in Dakki 1997: 62

Pierre 1922b, AP, Atlantic coast; Ramdani 1981, AP, Merja Sidi Boughaba; Savchenko et al. 1992; Dakki 1997; Koçak and Kemal 2010; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Jnane Niche, Oued Aârkoub, Aïn Jdioui; Oosterbroek 2020 (CCW)

Idiocera (Idiocera) sziladyi (Lackschewitz, 1940)

Driauach and Belqat 2016, Rif, Oued Zarka; Oosterbroek 2020 (CCW: Rif)

Ilisia Rondani, 1856

Ilisia maculata (Meigen, 1804)

Driauach and Belqat 2016, Rif, Oued Maggou, EM, Oued Tafoughalt; Oosterbroek 2020 (CCW: Rif; EM)

Molophilus Curtis, 1833

Molophilus (Molophilus) ibericus Starý, 2011

Starý 2011, HA; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Molophilus (Molophilus) obscurus (Meigen, 1818)

Starý and Oosterbroek 2008, HA, Massif Toubkal, Oukaimeden, 2500–2800 m; Driauach et al. 2013; Driauach and Belqat 2016, Rif, tributary of Oued Ouara, tributary of Oued Maggou, Dayat El Ânassar, Dayat Jebel Zemzem, Aïn El Ma Bared, Dayat Rmali, Oued Amsemlil, Aïn Mâaze, Dayat Afrate, Dayat Lemtahane; Oosterbroek 2020 (CCW: Rif)

Molophilus (Molophilus) propinquus propinquus (Egger, 1863)

Savchenko et al. 1992; Starý and Oosterbroek 2008, HA, Oukaimeden (2500–2800 m); Starý 2011; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Ouara, Oued Farda, tributary of Oued Taida, Oued Maggou; Oosterbroek 2020 (CCW: Rif)

Molophilus (Molophilus) testaceus Lackschewitz, 1940

Driauach and Belqat 2016, Rif, Dayat Amsemlil, Dayat Lemtahane, Marj El kheyl, Oued Tkarae, Dayat near Aïn Afersiw, Dayat Mezine, Dayat Tazia, Dayat Rmali, Dayat Amsemlil, Dayat near Aïn Afersiw; Oosterbroek 2020 (CCW: Rif)

Symplecta Meigen, 1830

Symplecta (Symplecta) grata Loew, 1873

Ebejer et al. 2020, Rif, Aïn Jdioui (Tahaddart, 8 m)

Symplecta (Symplecta) hybrida (Meigen, 1804)

Lackshewitz 1940a, HA, Goundafa (1200 m); Séguy 1941a, HA, Taroudant; Savchenko et al. 1992; Oosterbroek et al. 2007; Koçak and Kemal 2010; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Barrage Ajras, Oued El Kanar, Oued Aârkoub, Oued Jnane Niche, MA, Barrage Allal El Fassi; Oosterbroek 2020 (CCW: Rif; MA)

Symplecta (Trimicra) pilipes (Fabricius, 1787)

= Trimicra pilipes Fabricius, in Pierre 1922a: 23

= Trimicra andalusiaca Strobl, in Pierre 1922b: 149

= Trimicra hirsutipes Macquart, in Séguy 1930a: 22

Pierre 1922a, AP, Dradek (near Rabat), HA, Marrakech; Pierre 1922b, MA, Volubilis, HA, Oued Tensift; Séguy 1930a; Séguy 1941d, AA, Taroudant; Savchenko et al. 1992; Dakki 1997; Pârvu and Zaharia 2007; Driauach et al. 2013; Driauach and Belqat 2016, Rif, tributary of Oued Hachef, Oued Aârk­ob, Oued El Kanar, Oued Mezine, AP, Aïn Chouk (Larache); Oosterbroek 2020 (CCW: Rif)

Tasiocera (Dasymolophilus) murina (Meigen, 1818)

Driauach and Belqat 2016, Rif, Oued Farda, Oued Amsemlil; Oosterbroek 2020 (CCW: Rif)

Dactylolabinae

Dactylolabis Osten-Sacken, 1860

Dactylolabis (Dactylolabis) symplectoidea Egger, 1863

Pierre 1922a, AP, Around Casablanca (coastal meseta); Savchenko et al. 1992; Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Limnophilinae

Austrolimnophila Alexander, 1920

Austrolimnophila (Austrolimnophila) latistyla Starý, 1977

Driauach and Belqat 2016, Rif, Oued Maggou; Oosterbroek 2020 (CCW: Rif)

Dicranophragma Osten-Sacken, 1860

Dicranophragma (Brachylimnophila) adjunctum (Walker, 1848)

= Neolimnomyia adjuncta (Walker, 1848), in Pârvu et al. 2006: 273

Pârvu et al. 2006, AP, Merja Zerga; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranophragma (Brachylimnophila) nemorale (Meigen, 1818)

Lackschewitz 1940b, HA, Tachdirt (2200–2700 m); Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Aïn Sidi Brahim Ben Arrif, tributary Oued Ouara, Oued Maggou; Oosterbroek 2020 (CCW: Rif)

Eloeophila Rondani, 1856

Eloeophila maroccana Starý, 2009

Starý 2009b, HA, Okaïmeden (2500–2800 m); Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Euphylidorea Alexander, 1972

Euphylidorea (Euphylidorea) crocotula (Séguy, 1941)

= Phylidorea crocotula (Séguy), in Séguy 1941a: 28

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Savchenko et al. 1992, HA, Tachdirt (Toubkal); Starý and Oosterbroek 2008, HA, Oukaimeden (2500 m–2800 m), Imlil (1400 m); Driauach et al. 2013; Driauach and Belqat 2016, Rif, Aïn Bab Tariouant; Oosterbroek 2020 (CCW: Rif; HA)

Euphylidorea (Euphylidorea) dispar (Meigen, 1818)

Driauach and Belqat 2016, Rif, Oued at 15 km from Fifi; Oosterbroek 2020 (CCW: Rif)

Euphylidorea (Euphylidorea) lineola (Meigen, 1804)

Lackschewitz 1940b, HA; Savchenko et al. 1992; Starý and Freidberg 2007; Driauch et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Hexatoma Latreille, 1809

Hexatoma (Hexatoma) bicolor (Meigen, 1818)

Driauach and Belqat 2016, Rif, Oued at 15 km from Fifi, tributary of Oued Ouara, Oued Madissouka, Oued Maggou, Aïn Quanquben (Jebel Bou Bessoui), Oued Tkarae, Oued Tamerte; Oosterbroek 2020 (CCW: Rif)

Hexatoma (Hexatoma) gaedii (Meigen, 1830)

Lackschewitz 1940b, HA, Tachdirt (2200 m–2700 m); Savchenko et al. 1992; Starý and Freidberg 2007; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Pseudolimnophila Alexander, 1919

Pseudolimnophila (Pseudolimnophila) sepium (Verrall, 1886)

Driauach et al. 2013, Rif, Guelta Tazia; Driauach and Belqat 2016, Rif, Dayat Tazia, Aïn Sidi Brahim Ben Ar­rif, Aïn Afersiw, Oued Maggou, Dayat Tazia, Oued Taida, Aïn Sidi Brahim Ben Arrif; Oosterbroek 2020 (CCW: Rif)

Limoniinae

Dicranomyia Stephens, 1829

Dicranomyia (Dicranomyia) affinis (Schummel, 1829)

Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Amsemlil, Dayat Lemtahane, Oued Tkarae, Marj El Kheyl, Aïn Sidi Brahim Ben Arrif, Dayat Tazia, Aïn El Maounzil, Aïn El Malâab, Dayat near Aïn Afersiw, Aïn Bab Tariouant; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Dicranomyia) chorea (Meigen, 1818)

Pierre 1922b, HA, Haute Réghaya, Tannaout, Asni (1000 m–1250 m); Savchenko et al. 1992; Dakki 1997; Starý and Freidberg 2007; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Farda, Oued Kelâa, Aïn Ras el Ma, Oued Jnane Niche, Cascade Chrafate, maison forestière, Oued El Kanar, tributary of Oued Zarka, HA, Oued Sidi Fares (National Park of Toubkal); Oosterbroek 2020 (CCW: Rif; HA)

Dicranomyia (Dicranomyia) didyma (Meigen, 1804)

Pierre 1922b, HA, Haute Réghaya (2000 m); Dakki 1997; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Dicranomyia (Dicranomyia) goritiensis (Mik, 1864)

Lackschewitz 1940b; Vaillant 1956b; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Maggou, Cascade Chrafate, Oued El Koub, Cascade Zarka, Âounsar Aheramen, maison forestière; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Dicranomyia) longicollis (Macquart, 1846)

= Telecephala longicollis (Macquart), in Pierre 1922a: 21, Pierre 1922b: 148, Séguy 1930a: 22

Séguy 1930a, AP, Dradek (near Rabat); Dakki 1997; Pierre 1922a, AP, Dradek, HA, Marrakech; Pierre 1922b, AP, Rabat; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Aârate, Barrage Moulay Bouchta, Oued Kbir, MA, Barrage Allal El Fassi; Oosterbroek 2020 (CCW)

Dicranomyia (Dicranomyia) mitis (Meigen, 1830)

= Dicranomyia hygropetrica Vaillant, in Vaillant 1956b: 42

Vaillant 1956b, HA, Asif Tessaout (M’Goum), Izourar, Tahanaout, Tamesrit, Imi-N’Ifri, Aguelmous, Sidi Chamarouch, Lac Tamhda (Anremer), Oukaimeden; Savchenko et al. 1992; Pârvu et al. 2006, AA, near Agadir, Souss plain to High Atlas occidental; Pârvu and Zaharia 2007; Gavryushin pers. comm. 2012, HA; Driauach et al. 2013, Rif, Tazia, Tisgris; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranomyia (Dicranomyia) modesta (Meigen, 1818)

Driauach and Belqat 2016, Rif, Oued Farda, Oued El Kanar, Oued Jnane Niche, Oued Sidi Yahia Aârab; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Dicranomyia) novemmaculata (Strobl, 1906)

Driauach and Belqat 2016, Rif, tributary of Oued el Fondak, Oued Aârate; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Dicranomyia) ventralis (Schummel, 1829)

Driauach and Belqat 2016, Rif, Lake Badriouen; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Glochina) sericata (Meigen, 1830)

Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranomyia (Melanolimonia) hamata Becker, 1908

Driauach and Belqat 2016, Rif, tributary of Oued Kbir, Oued Aârate, Aïn Sidi Brahim Ben Arrif, Dayat Tazia; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Melanolimonia) morio (Fabricius, 1787)

= Limonia pauliani Séguy, in Séguy 1941a: 26

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Savchenko et al. 1992, HA, Tachdirt (Toubkal); Driauach et al. 2013; Driauach and Belqat 2016, Rif, Seguia Lemtahane, Dayat near Aïn Afersiw, Dayat Jebel Zemzem; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranomyia majuscula Pierre, 19241

Pierre 1924a, HA, Haut Imminen (2400 m); Séguy 1930a, HA, Haut Imminen; Dakki 1997; Savchenko et al. 1992, HA, Haut Imminen; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranoptycha Osten-Sacken, 1860

Dicranoptycha fuscescens (Schummel, 1829)

Eiroa 2000, Rif; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Mlilah, Oued Zendoula, AP, Oued Loukous; Oosterbroek 2020 (CCW)

Geranomyia Haliday, 1833

Geranomyia caloptera (Mik, 1867)

Driauach et al. 2013, HA, Setti Fatma; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Geranomyia obscura Strobl, 1900

Vaillant 1956b, HA, Lac Tamhda (Anremer), Oukaimeden, Izourar, Sidi Chamarouch, Tamesrit; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Helius Lepeletier & Serville, 1828

Helius (Helius) hispanicus Lackschewitz, 1928

Starý and Oosterbroek 2008, HA, Massif Toubkal, Imlil (17 km S Asni, 1700–1900 m); Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Amsemlil; Oosterbroek 2020 (CCW: Rif)

Helius (Helius) pallirostris Edwards, 1921

Driauach and Belqat 2016, AP, Aïn Chouk (Larache); Oosterbroek 2020 (CCW: Rif)

Limonia Meigen, 1803

Limonia flavipes (Fabricius, 1787)

Pierre 1922b, HA, Haute Réghaya, Asni (1250 m); Savchenko et al. 1992; Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Limonia hercegovinae (Strobl, 1898)

Starý and Oosterbroek 2008, MA, Ifrane (1700 m), HA; Gavryushin pers. comm. 2012, HA; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Limonia macrostigma (Schummel, 1829)

Starý and Oosterbroek 2008, HA, 5 km from Oukaimeden (2350 m); Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Limonia nubeculosa Meigen, 1804

Pierre 1922b, HA, Haute Réghaya, Asni (1250 m); Savchenko et al. 1992; Dakki 1997; Gavryushin pers. comm. 2012, HA; Driauach and Belqat 2016, Rif, tributary of Oued El Fondak, Aïn Ras el Ma, Aïn Boughaba, Âounsar Aheramen, Aïn Takhninjoute, maison forestière, Oued Madissouka, Aïn Quan­quben (Jebel Bou Bessoui), EM, Oued Azila; Oosterbroek 2020 (CCW: Rif; HA)

Limonia phragmitidis (Schrank, 1781)

Starý and Oosterbroek 2008, MA, Ifrane (1700 m); Driauach and Belqat 2016, Rif, Aïn Quanquben (Jebel Bou Bessoui); Oosterbroek 2020 (CCW)

PEDICIIDAE

K. Kettani, P. Oosterbroek

Number of species: 6. Expected: 10

Faunistic knowledge of the family in Morocco: moderate

Pediciinae

Dicranota Zetterstedt, 1838

Dicranota (Dicranota) bimaculata (Schummel, 1829)

Pierre 1922b, HA, Haute Réghaya (1800 m); Savchenko et al. 1992 (? Morocco); Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranota (Dicranota) irregularis Pierre, 1921

Pierre 1922b, HA, Haute Réghaya (1800 m); Savchenko et al. 1992, HA, Cirque d’Arround (Haute Réghaya); Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranota (Ludicia) claripennis (Verrall, 1888)

Driauach and Belqat 2016, Rif, Oued Amsemlil, maison forestière; Oosterbroek 2020 (CCW)

Dicranota (Paradicranota) candelisequa Starý, 1981

Pârvu et al. 2006, AP, Merja Zerga; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranota (Paradicranota) landrocki Czižek, 1931

Driauach et al. 2013, Rif, Fifi (1252 m); Driauach and Belqat 2016, Rif, Oued Ouara, Oued Taida, Aïn Sidi Brahim Ben Arrif, Âounsar Aheramen, Oued Tizekhte, Oued Mezine, maison forestière, Aïn Bab Tariouant, HA, Imlil (Assif Haouz); Oosterbroek 2020 (CCW)

Tricyphona Zetterstedt, 1838

Tricyphona (Tricyphona) immaculata (Meigen, 1804)

Driauach and Belqat 2016, Rif, maison forestière, Dayat Lemtahane; Oosterbroek 2020 (CCW)

TIPULIDAE

K. Kettani, P. Oosterbroek, H. de Jong

Number of species: 39. Expected: 42

Faunistic knowledge of the family in Morocco: moderate

Dolichopezinae

Dolichopeza Curtis, 1825

Dolichopeza (Dolichopeza) hispanica Mannheims, 1951

Theowald and Oosterbroek 1980, HA, Aghbalou, Oukaimeden, Imlil, Tadmant; Oosterbroek and Theowald 1992; Oosterbroek and Lantsov 2011, HA, Oukaimeden (2300 m), Aghbalou (Massif Toubkal), 43 km S Marrakech (1000 m), Imlil (17 km S Asni, 1700–1900 m), Tadmant (17 km E Asni); Adghir et al. 2018, Rif, Kitane, Aîn Ras el Ma (Chefchaouen); Oosterbroek 2020 (CCW)

Tipulinae

Nephrotoma Meigen, 1803

Nephrotoma alluaudi (Pierre, 1922)

= Pachyrhina lunulicornis Schummel, in Pierre 1922a: 24

= Pachyrhina alluaudi Pierre, in Pierre 1922b: 150, Dakki 1997: 62

= Pales alluaudi (Pierre), in Mannheims 1951: 47

Pierre 1922b, HA, Tannaout (1000 m); Mannheims 1951, Rif, Beni Seddat, AP, Lagune Guedira; HA, Taddert north of Marrakech, Goundafa (1200 m); AA, Llano Amarillo, Tlata Reisana; Oosterbroek 1979b, Rif, HA, Imlil (1400 m), Tizi-n’Tichka (2200 m); Theowald and Oosterbroek 1980, AP, Rabat, Guedira lagoon, MA, Immouzer, Ifrane, Timahdit, Aghbalou, Tizi-n’Zou, HA, Marrakech, Goundafa, Taddert, Dayat, Tizi-n’Test, Tizi-n’Tichka, Asni, Imlil, Oukaimeden, Setti Fatma, Tinmel, Acif Tifni, AA, Taroudant, Mikdana, Sidi Said bou Merdoul, Tlata Reisana, Llano Amarillo; Eiroa 1990, MA, Azrou, Ajabo; Oosterbroek and Theowald 1992, HA, Tannaout; Dakki 1997; Mouna 1998; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen), Oued Laou, Dardara, Douar Mouarâa, Oued Zandoula, Laghmari-Rmal, Douar Louamera, Douar Laheyayda, Oued Jnane Niche, Cabo Negro, Oued Beni Said, Oued El Kanar, Oued Amsemlil; Oosterbroek 2020 (CCW) – MISR (HA), MHNV, MNCNM, MAKB

Nephrotoma appendiculata pertenua Oosterbroek, 1978

Oosterbroek 1978, Rif, 9 km SW Chefchaouen; Theowald and Oosterbroek 1980, Rif, Dardara, MA, Fès, Immouzer, Khemisset; Oosterbroek and Theowald 1992; de Jong 1993, 1998, Rif; Adghir et al. 2018, Rif, Oued Ametrasse, Dayat Aïn Jdioui, Barrage Moulay Bouchta, Oued Sahil, Dayat Jebel Zemzem, Aïn Sidi Brahim Ben Arrif, Oued Nakhla, Oued Tizekhte, Lot Hemmadi, Douar Ayacha, Douar Louamera, Dayat Mezine, Aïn El Malâab, Aïn Takhninjoute, Dayat Tazia, Oued Taida, maison forestière Tazia, Tourbière Amesmlil, Oued El Hamma, Oued Kbir, Jebel Lakraâ; Oosterbroek 2020 (CCW)

Nephrotoma astigma Pierre, 1925

Theowald and Oosterbroek 1980, Rif, Dardara, MA, Taza; Adghir et al. 2018, Rif, Oued Tabandout, Etang Maggou, Aïn El Malâab, Douar Remla; Oosterbroek 2020 (CCW)

Nephrotoma fontana Oosterbroek, 1978

de Jong 1998, Rif; Adghir et al. 2018, Rif, Ketama, maison forestière Tazia, Dayat Tazia; Oosterbroek 2020 (CCW)

Nephrotoma guestfalica vaillanti de Jong, Adghir & Bosch, 2021

de Jong et al. 2021, Rif, Ras el Ma (Chefchaouen, 500 m), Fomento, Oued Laou (6 km north-west of Chefchaouen, 200 m), Jebel Tissouka (3 km south of Chefchaouen, 500 m; 3 km south of Chefchaouen, 700 m; 4 km south-east of Chefchaouen, 700–800 m; 5 km southeast of Chefchaouen, 700–900 m; 5 km south of Chefchaouen, 900–1000 m), Bab Taza (25 km south-east of Chefchaouen, 750–800 m), Dardara (10 km south of Chefchaouen, 300 m), Oued Martil, Nakhla, Âounsar Aheramen, Boumerouil, Oued Maggou, Oued Maggou (Aïn Ras el Ma), Douar Kitane, Oued Sidi Yahia Aârab, Oued Tamerte, Oued Zandoula, Oued El Hamma, Lot Hemmadi, Oued Tamerte, Oued Sidi Mohamed Saâda, Oued El Koub, Douar Iholebatine, Dayat Tazia, Belouazen, Oued Lemtahane, Oued Siflaou, Oued Amsemlil, AP, Oued Loukous, Aïn el-Aouda, MA, Ifrane (road to Mischliffen, 1680 m), N.S. and W of Ifrane (1400–1800 m), Khemisset, Azrou, Oum-er-Rbia, HA, M’semrir (bord de l’Oued), Rich, Haute Rhégaya (identified by Mannheims in 1951 as surcoufi)

Nephrotoma luteata (Meigen, 1818)

Oosterbroek 1979a, Rif, Targuist, Chefchaouen, HA, Kasba Taguendaft, near Oukaimeden; Theowald and Oosterbroek 1980, Rif, Targuist, Chefchaouen, HA, Kasba Taguendaft, near Oukaimeden; Oosterbroek and Theowald 1992; Pârvu et al. 2006, AP, Merja Zerga; Adghir et al. 2018, Rif, Oued Laou, Dardera, Oued Nakhla; Oosterbroek 2020 (CCW)

Nephrotoma subanalis (Mannheims, 1951)

= ? Pachyrhina analis Schummel, in Pierre 1922a: 24, Pierre 1922b: 150

= Pales subanalis Mannheims, in Mannheims 1951: 56

Mannheims 1951, HA, Tachdirt (2200–2900 m); Vaillant 1956b, HA, Oukaimeden (2250 m); Oosterbroek 1979c, HA, Tachdirt (2200–2900 m); Theowald and Oosterbroek 1980, HA, Tachdirt, Oukaimeden, M’Goum, Tadmant, Imlil, Tizi-N’Tichka; Oosterbroek and Theowald 1992, HA, Tachdirt; Oosterbroek 2020 (CCW)

Nephrotoma submaculosa Edwards, 1928

Oosterbroek 1982, Rif, Ketama, Dardara, MA, Azrou; Oosterbroek and Theowald 1992; de Jong 1998, Rif, Atlas; Oosterbroek 2011; Adghir et al. 2018, Rif, Jebel Dahedouh, Ketama, Aïn Sidi Brahim Ben Arrif, Oued Taida, tributary of Oued Ouara, Sidi Chouiref, Dayat Tazia; Oosterbroek 2020 (CCW)

Nephrotoma sullingtonensis Edwards, 1938

Oosterbroek 1978; Theowald and Oosterbroek 1980, Rif, Bab Berred; Oosterbroek 1982 Rif, Bab Berred; de Jong 1993, 1998, Rif; Oosterbroek and Theowald 1992; Adghir et al. 2018, Rif, 4 km SE Ketama, Barrage Moulay Bouchta, Oued Aârate, Aïn Takhninjoute, Oued Jbara, Aïn El Malâab, Aïn El Ma Bared, Oued Lemtahane, maison forestière Tazia, Oued Taida, Dayat Tazia; Oosterbroek 2020 (CCW)

Tipula Linnaeus, 1758

Tipula (Acutipula) anormalipennis Pierre, 1924

Pierre 1924b, HA, Haut Imminen; Séguy 1930a, HA, Haut Imminen; Séguy 1941a, HA, Tachdirt (2500 m), Haut Imminen; Mannheims 1952, HA, Tachdirt (2500 m), Imminen (2400–2500 m), Arround (1950 m); Vaillant 1956b, HA, Lake of Tamhda (Anremer, 2900 m); Theowald and Oosterbroek 1980, HA, Anremer, Haut-Immenen, Tachdirt, Arround, Oukaimeden; Vermoolen 1983, HA, Haut Imminen, Oukaimeden (2500–2800 m), Arround (1950 m), Tachdirt (2500 m); Oosterbroek and Theowald 1992; de Jong 1994a, HA; de Jong 1998, HA; Dakki 1997; Oosterbroek 2020 (CCW)

Tipula (Acutipula) repentina Mannheims, 1952

= Tipula maxima Poda, in Séguy 1941a: 26

Séguy 1941a, HA, Tachdirt (2500 m); Mannheims 1952, HA, Tachdirt (2200–2700 m); Vaillant 1956b, HA, Lac Tamhda (Anremer, 2900), M’Goum (2500 m); Theowald and Oosterbroek 1980, HA, Anremer, M’Goum, Tachdirt, Tizi-N’Tichka, Asni, Oukaimeden, Setti Fatma, Imlil, Tadmant; Vermoolen 1983, MA, Ifrane, HA, Androment, M’Goum, Tadmant, Tizi-N’Test, Setti Fatma, Oukaimeden, Tizi-N’Tichka; Oosterbroek and Theowald 1992, HA, Tachdirt; de Jong 1994a, MA, HA; de Jong 1998, HA; Dakki 1997; Oosterbroek 2020 (CCW)

Tipula (Acutipula) rifensis Theowald & Oosterbroek, 1980

Theowald and Oosterbroek 1980, Rif, Targuist; Vermoolen 1983, Rif, Targuist, Tidiguin (90 km E. of Ouezzane, 2350 m); Oosterbroek and Theowald 1992, Rif; de Jong 1994a, 1998, Rif; Adghir et al. 2018; Oosterbroek 2020 (CCW)

Tipula (Emodotipula) leo Dufour, 1991

= Tipula (Emodotipula) obscuriventris Strobl2, in Oosterbroek and Theowald 1992: 99

Oosterbroek and Theowald 1992 (?); Dufour 2003, Rif; Adghir et al. 2018, Rif, Jebel Tissouka; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) bivittata Pierre, 1922

Pierre 1922a, AP, Maâmora between Kénitra and Oued Beth, Dradek (near Rabat); Pierre 1922b, AP, Rabat; Mannheims 1968; Theowald and Oosterbroek 1980, AP, forest of Maâmora, Dradek; Oosterbroek and Theowald 1992, AP, forest of Maâmora, Dradek; Dakki 1997; Oosterbroek 2020 (CCW) – MISR (AP, Kénitra, Dradek)

Tipula (Lunatipula) cinereicolor Pierre, 1924

Pierre 1924b, HA, Haut Imminen; Séguy 1930a, HA, Tachdirt (3100–3200 m); Theowald 1973, HA, Haut Imminen (2400 m); Theowald and Oosterbroek 1980, MA, Ifrane, HA, Oukaimeden, Tachdirt, Haut-Imminen; Eiroa 1990, MA, Ajabo; Oosterbroek and Theowald 1992, HA, Imminen; Dakki 1997; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) cornicula Pierre, 1922

Pierre 1922b, HA, Arround (2000 m); Séguy 1930a, HA, Tachdirt (3100–3200 m); Theowald 1973, HA, Cirque d’Arround (2000 m), Tachdirt (2200–2700 m), Goundafa (1200 m); Theowald and Oosterbroek 1980, HA, Oukaimeden, Tachdirt, Goundafa; Oosterbroek and Theowald 1992, HA, Tachdirt; Dakki 1997; Oosterbroek 2020 (CCW) – MISR (HA)

Tipula (Lunatipula) fabiola Mannheims, 1968

Theowald and Oosterbroek 1980, Rif, Bab Berred, Ras El Ma (Chefchaouen), MA, Jebel Abad, Ifrane; Oosterbroek and Theowald 1992; Adghir et al. 2018; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) hermes Theischinger, 1977

Theischinger 1977, Rif, north of Ouezzane; Theowald 1980; Theowald and Oosterbroek 1980, Rif, north of Ouezzane; Oosterbroek and Theowald 1992, Rif, Ouezzane; Adghir et al. 2018, Rif, 4 km SE Ketama, Oued Ametrasse, Aïn El Ma Bared; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) iberica iberica Mannheims, 1963

= Tipula lunata Linnaeus, in Pierre 1922b: 149, Séguy 1930a: 23

Pierre 1922b, HA, Haute Réghaya, Tannaout (1000 m); Séguy 1930a, HA, Haut Imminen; Mannheims 1963, MA; Theowald and Oosterbroek 1980, Rif, Bab Berred, MA, Ifrane, Taounate; Eiroa 1990, MA, Ifrane; Oosterbroek and Theowald 1992; Oosterbroek 2009; Adghir et al. 2018, Rif, Ketama; Oosterbroek 2020 (CCW) – MISR (HA)

Tipula (Lunatipula) iberica spinula Theischinger, 1980

Theischinger 1980, HA, Oukaimeden (1300–2800 m); Theowald and Oosterbroek 1980, HA, Oukaimeden; Oosterbroek and Theowald 1992, HA, Oukaimeden; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) peliostigma peliostigma Schummel, 1833

Eiroa 1990, MA, Azrou; Mouna 1997; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) pjotri de Jong & Adghir, 2018

Adghir et al. 2018, Rif, Jebel El Kelâa (Talassemtane, 1340 m)

Tipula (Lunatipula) pseudocinerascens Strobl, 1906

Adghir et al. 2018, Rif, Stehat, Oued Taida, Perdicaris Park, Dayat Tazia

Tipula (Lunatipula) rocina Theischinger, 1979

Theowald and Oosterbroek 1980, Rif, Tétouan; Oosterbroek and Theowald 1992; Oosterbroek 2009; Adghir et al. 2018; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) selenaria Mannheims, 1967

Mannheims 1967, MA, Jebel Tazzeka (1500–1989 m), HA, Goundafa (1200 m); Theowald and Oosterbroek 1980, MA, Tazzeka, HA, Foum Keneg, Goundafa, Oukaimeden; Eiroa 1990, MA, Ajabo; Oosterbroek and Theowald 1992, HA, Goundafa; de Jong 1995, MA, HA, Oukaimeden; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) stimulosa Mannheims, 1973

Adghir et al. 2018, Rif, Jebel El Kelâa (Talassemtane, 1340 m)

Tipula (Lunatipula) subfalcata Mannheims, 1967

de Jong 1995, 1998, Rif; Adghir et al. 2018, Rif, Jebel Tissouka, Oued Tamerte, Oued El Koub; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) subpustulata Mannheims, 1963

= Tipula pustulata Pierre, in Pierre 1922b: 150

Pierre 1922b, AP, Mogador; Vaillant 1956b, HA, M’Goun (2500 m); Mannheims 1963, AP, Aïn el Aouda, MA, Jebel Tazzeka (1600–1989 m), HA, Goundafa (1200 m), Tachdirt (2200–2900 m), AA, Lac Goulmima; Theowald 1972; Theowald and Oosterbroek 1980, AP, Aïn el Aouda, MA, Iebel Tazzeka, HA, M’Goum, Goundafa, Tachdirt, Oukaimeden, Imlil; Eiroa 1990, MA, Ajabo; Oosterbroek and Theowald 1992, HA, Goundafa; Dakki 1997; Adghir et al. 2018, Rif, 4 km SE Ketama, Aïn El Ma Bared; Oosterbroek 2020 (CCW) – MISR (AP, Mogador)

Tipula (Lunatipula) tazzekai Theowald, 1973

Theowald 1973, MA, Jebel Tazzeka (1600–1989 m); Theowald and Oosterbroek 1980, MA, Jebel Tazzeka; Eiroa 1990, MA, Jebel Hebri; Oosterbroek and Theowald 1992, MA, Jebel Tazzeka; Oosterbroek 2020 (CCW)

Tipula (Savtshenkia) atlas Pierre, 1924

Pierre 1924b, HA, Tachdirt (3100–3250 m); Séguy 1930a, HA, Tachdirt (3100–3200 m); Vaillant 1956b, HA, Cascade Siroua (3000 m), M’Goun (2500 m), Toubkal (3350 m), Lake of Tamhda (Anremer, 2900 m); Mannheims 1964, HA; Theowald 1973, HA; Theowald 1980, HA; Theowald and Oosterbroek 1980, HA, Toubkal, Sources de Tessaouts, M’Goum, Siroua, Anremer, Tachdirt, Oukaimeden, Tizi-N’Tichka, Tadmant; Eiroa 1990, MA, Oum-Er-Rbia; Oosterbroek and Theowald 1992, HA, Tachdirt; de Jong 1994b; de Jong 1998, Atlas; Dakki 1997; Oosterbroek 2020 (CCW)

Tipula (Savtshenkia) breviantennata Lackschewitz, 1933

de Jong 1998, Rif; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen), Oued Maggou, Douar Aouzighen; Oosterbroek 2020 (CCW)

Tipula (Savtshenkia) confusa van der Wulp, 1883

Adghir et al. 2018, Rif, maison forestière (Talassemtane)

Tipula (Savtshenkia) rufina rufina Meigen, 1818

Theowald and Oosterbroek 1980, HA, Oukaimeden; Theowald and Oosterbroek 1983, Rif, Atlas; Oosterbroek and Theowald 1992; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen); Oosterbroek 2020 (CCW)

Tipula (Tipula) mediterranea Lackschewitz, 1930

Vaillant 1956b, HA, Oukaimeden (2250 m), M’Goun (2500 m); Mannheims 1952; Theowald and Oosterbroek 1980, MA, Ifrane, HA, Oukaimeden, M’Goum, Asni, M’Semrir, Ifni, Bab-Rou-Idie, Tizi-N’Tichka, Setti Fatma; Theowald 1984, Rif; Eiroa 1990, MA, Azrou, Oum-Er-Rbia; Oosterbroek and Theowald 1992; Pârvu et al. 2006 AP, Merja Zerga; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen), Jebel Tissouka, Bab Taza, Dardara, 4 km SE Ketama, Aïn Afersiw, Douar Kitane, Dayat Jebel Zemzem, Wilaya (Tétouan), Aïn El Ma Bared, Oued Tamerte, Douar Louamera, Tourbière Amesmlil, Dayat Mezine, Hejar Nehal, tributary of Oued Ouara, Oued Tkarae, Oued Jnane Niche, Dayat Afrate, Oued Ametrasse; Oosterbroek 2020 (CCW)

Tipula (Tipula) oleracea Linnaeus, 1758

Mannheims 1952, Rif, Tlata Ketama; Theowald and Oosterbroek 1980, Rif, Tlata Ketama; Theowald 1984, Rif; Oosterbroek and Theowald 1992; Dakki 1997; Pârvu and Zaharia 2007; Oosterbroek 2011; Adghir et al. 2018, Rif, Oued Laou, Dardara, Jebel Tissouka, Ksar Rimal, 15 km from Fifi, Oued Aârate, Oued El Hamma, Douar Louamera, Oued Zaouya, Près de Beni Said, Wilaya (Tétouan), Tourbière Amesmlil, Oued Smir, Aïn Jdida; Oosterbroek 2020 (CCW) – MISR

Tipula (Vestiplex) vaillanti vaillanti Theowald, 1977

Adghir et al. 2018, Rif, Jebel El Kelâa (Talassemtane, 1340 m), Douar Kitane

Tipula (Yamatotipula) afriberia afriberia Theowald & Oosterbroek, 1980

Theowald and Oosterbroek 1980, HA, Oukaimeden; Oosterbroek and Theowald 1992, HA, Oukaimeden; Oosterbroek 1994a; Adghir et al. 2018, Rif, Jebel Tissouka, Dardara, Douar Mokedassen, Oued Zarka; Oosterbroek 2020 (CCW)

Tipula (Yamatotipula) barbarensis Theowald & Oosterbroek, 1980

= Tipula lateralis Meigen, in Pierre 1922a: 23, Pierre 1922b: 150, Séguy 1930a: 23, Séguy 1941a: 26, Mannheims 1952: 98 (in part), Vaillant 1956b: 238

Pierre 1922a, AP, Dradek (Rabat), MA, Azrou (riverside of Oued Tigrigra); Pierre 1922b, AP, Mogador, MA, Beni Méllal, HA, Asni; Séguy 1930a; Séguy 1941a, HA, Imi n’Ouaka (1500 m); Mannheims 1952; Vaillant 1956b, HA, Oukaimeden; Theowald and Oosterbroek 1980, Rif, Dardara, MA, Ifrane, Aghbalou, HA, Asni, Oukaimeden, Setti Fatma, Imlil, Tizi-N’Tichka, Tadmant, Tachdirt; Eiroa 1990, MA, Azrou, Oum-er-Rbia, Ifrane; Oosterbroek and Theowald 1992, HA, Setti Fatma; Oosterbroek 1994a, Rif, Dardara, MA, Ifrane, Aghbalou, HA, Oukaimeden, Setti Fatma, Imlil, Tizi-N’Tichka, Tadmant, Asni, Tachdirt; Dakki 1997; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen), Jebel Tissouka, Bab Taza, Dardara, 4 km SE Ketama, Aïn El Manzela, Aïn Bab Tariouante, Dayat Aïn Afersiw, Oued El Kanar, Dayat Afrate, Douar Kitane, Wilaya (Tétouan), 15 km from Fifi, Aïn Sidi Brahim Ben Arrif, Oued Nakhla, Âounsar Aheramen, Oued Boumerouil, Douar Zaouya, Oued Tizekhte, Oued Samsa, Dayat Aïn Jdioui, Douar Ouled Laghmari-Rmal, maison forestière Tazia, Hejar Nehal, Lot Hemmadi, tributary of Oued Ouara, Oued Sidi Mohamed Saâda, Oued Amsemlil, Tourbière Amesmlil, Beni Salah, Oued Jnane Niche, Oued Maggou, Souk Lhed Beni Darkoul, Oued Imassouden, Aïn Helouma, Source Zarka, Aïn Kchour; Oosterbroek 2020 (CCW) – MISR

Trichoceridoidea

TRICHOCERIDAE

K. Kettani, E. Krzemińska

Number of species: 8. Expected: 10

Faunistic knowledge of the family in Morocco: good

Trichocerinae

Trichocera Meigen, 1803

Trichocera (Trichocera) hiemalis (DeGeer, 1776)

Pierre 1922b (AP, Rabat); Dakki 1997; Mouna 1998; AP (Rabat) – MISR

Trichocera (Trichocera) marocana Driauach, Krzemińska & Belqat, 2015

Driauach et al. 2015, Rif, Oued Akrir

Trichocera (Saltrichocera) annulata Meigen, 1818

Driauach et al. 2015, Rif, affluent Oued Akrir, Dayat Fifi, Oued Taria, Oued Kelâa, Oued Ouara, Oued à 15 km de Fifi, Aïn Mâaze, Oued Maggou, Aïn Quanquben, Oued Tizekhte, Forêt Jebel Bouhachem, EM, Grotte du Chameau, Oued Zegzel, Aïn Sidi Yahia, Cascade Grotte des pigeons, Oued Tafoughalt

Trichocera (Saltrichocera) pappi Krzemińska, 2003

Driauach et al. 2015, Rif, Dayat Fifi, affluent Oued Akrir, Oued Amsemlil, Ruisseau Agoummir, maison forestière, EM, Cascade Grotte des Pigeons, MA, Seguia El Hajeb

Trichocera (Saltrichocera) saltator (Harris, 1776)

Driauach et al. 2015, Rif, Dayat Fifi, Oued à 15 km de Fifi, Forêt Jebel Bouhachem, maison forestière, EM, Cascade Grotte des pigeons

Trichocera (Saltrichocera) sardiniensis Petrašiūnas, 2009

Driauach et al. 2015, Rif, Oued à 15 km de Fifi, affluent Oued Akrir, Oued Amsemlil, maison forestière

Trichocera (Saltrichocera) regelationis (Linnaeus, 1758)

Driauach et al. 2015, Rif, Oued Ouara, Aïn el Ma Bared, Oued à 15 km de Fifi, Cascade Chrafate, Aïn Quanquben

Trichocera (Saltrichocera) rufescens Edwards, 1921

Driauach et al. 2015, Rif, affluent Oued Akrir, Dayat Fifi, Oued Tassikeste, Oued Farda, Aïn Mâaze

Psychodoidea

PSYCHODIDAE

K. Kettani, R. Wagner

Number of species: 51. Expected: 70

Faunistic knowledge of the family in Morocco: moderate

Phlebotominae

Phlebotomus Loew, 1845

Phlebotomus (Larroussius) ariasi Tonnoir, 1921

Gaud 1947a; Gaud and Laurent 1952, AP, Rabat; Bailly-Choumara et al. 1971, AP, MA, HA; Rioux et al. 1974; Mouna 1998; Guernaoui et al. 2005, MA, HA; Bounamous 2010; Boussaa et al. 2005; Boussaa et al. 2010

Phlebotomus (Larroussius) chadlii Rioux, Juminer & Gibily, 1966

Rioux et al. 1974, HA; Rioux et al. 1975; Croset et al. 1978; Mouna 1998; Bounamous 2010

Phlebotomus (Larroussius) langeroni Nitzulescu, 1930

Rislorcelli 1941, HA; Bailly-Choumara et al. 1971, AP, EM; Croset et al. 1978; Mouna 1998

Phlebotomus (Larroussius) longicuspis Nitzulescu, 1930

Rislorcelli 1941, HA; Gaud and Laurent 1952, AP, Rabat; Bailly-Choumara et al. 1971, Rif, EM, AP, MA, HA, AA; Rioux et al. 1974, HA; Croset et al. 1978; Mouna 1998; Guernaoui et al. 2005, Rif, Chefchaouen, HA, AA, Agadir; Boussaa et al. 2005; Boussaa 2008; Boussaa et al. 2008; Boussaa et al. 2010; Bounamous 2010; Zarrouk et al. 2016

Phlebotomus (Larroussius) mariae Rioux, Croset, Léger and Bailly-Choumara, 1974

Hervy et al. 1994; Rioux et al. 1974, HA; Mouna 1998; Guernaoui et al. 2005, MA, HA

Phlebotomus (Larroussius) perfiliewi Parrot, 1930

Rioux et al. 1977; Croset et al. 1978; Mouna 1998

Phlebotomus (Larroussius) perniciosus Newstead, 1911

Séguy 1930a; Gaud and Laurent 1952, AP, Rabat; Bailly-Choumara et al. 1971, AP, EM, MA, HA, AA; Mouna 1998; Guernaoui et al. 2005, Rif, Chefchaouen, HA; Boussaa et al. 2008; Bounamous 2010; Boussaa et al. 2010; Zarrouk et al. 2016

Phlebotomus (Paraphlebotomus) alexandri Sinton, 1928

Bailly-Choumara et al. 1971, AA; Abonnenc 1972; Rioux et al. 1974, HA; Croset et al. 1978; Mouna 1998; Colacicco-Mayhugh et al. 2010, EM; Boussaa et al. 2010; Bounamous 2010

Phlebotomus (Paraphlebotomus) chabaudii Croset, Abonnenc & Rioux, 1970

Rioux et al. 1974, HA; Rioux et al. 1975; Mouna 1998

Phlebotmus (Paraphlebotomus) kazeruni Theodor & Mesghali, 1964

Mouna 1998

Phlebotomus (Paraphlebotomus) riouxi Depaquit, Killick-Kendrick & Léger, 1998

Bounamous 2010

Phlebotomus (Paraphlebotomus) sergenti Parrot, 1917

Séguy 1930a; Rislorcelli 1941; Rislorcelli 1947; Gaud and Laurent 1952, AP, Rabat; Bailly-Choumara et al. 1971, Rif, AP, EM, MA, HA, AA; Abonnenc 1972; Rioux et al. 1974, HA; Mouna 1998; Boussaa et al. 2009; Bounamous 2010

Phlebotomus (Phlebotomus) bergeroti Parrot, 1934

Rioux et al. 1975, HA; Mouna 1998; Bounamous 2010

Phlebotomus (Phlebotomus) papatasi (Scopoli, 1786)

Séguy 1930a; Rislorcelli 1941, Rislorcelli 1947, HA; Bailly-Choumara et al. 1971, AP, EM, MA, HA, AA; Abonnenc 1972; Rioux et al. 1974, HA; Croset et al. 1978; Mouna 1998; Boussaa et al. 2005; Boussaa 2008; Colacicco-Mayhugh et al. 2010, Mediterranean region; Bounamous 2010; Boussaa et al. 2010; Prudhomme et al. 2012

Phlebotomus clydei Sinton, 1928

Bailly-Choumara et al. 1971, EM; Mouna 1998

Phlebotomus lewisi Parrot, 1948

Bailly-Choumara et al. 1971, South AP; Mouna 1998

Sergentomyia França & Parrot, 1920

Sergentomyia (Grassomyia) dreyfussi (Parrot, 1933)

Rislorcelli 1941, Rislorcelli 1947, HA; Bailly-Choumara et al. 1971, AP, EM, MA, HA; Abonnenc 1972; Rioux et al. 1974, HA; Croset et al. 1978; Mouna 1998; Boumezzough et al. 2009, HA, Marrakech; Bounamous 2010

Sergentomyia (Parrotomyia) africana (Newstead, 1912)

= Phlebotomus (Parrotomyia) africana (Newstead), in Rislorcelli 1941: 522, Bailly-Choumara et al. 1971: 454; Rioux et al. 1974: 99

Séguy 1930a (as subspecies of minutus Rondani: 43), HA, Marrakech; Rislorcelli 1941 (as subspecies of minutus Rondani: 528), AA, Ksar es Souk; Gaud and Laurent 1952 (as subspecies of minutus Rondani: 75), AP, Rabat; Bailly-Choumara et al. 1971 (as subspecies of minutus Rondani: 438), AP, AA; Rioux et al. 1974, HA; Croset et al. 1978 (as subspecies of minutus Rondani: 722); Mouna 1998; Boussaa et al. 2005; Boumezzough et al. 2009, HA, Marrakech; Bounamous 2010; Boussaa et al. 2010

Sergentomyia (Sergentomyia) antennata (Newstead, 1912)

= Phlebotomus cinctus Parrot & Martin, 1944, in Mouna 1998: 86

= Phlebotomus signatipennis Newstead, 1920, in Mouna 1998: 86

Bailly-Choumara et al. 1971, EM; Rioux et al. 1974, HA; Léger et al. 1974, AA (south of Morocco); Mouna 1998

Sergentomyia (Sergentomyia) fallax (Parrot, 1921)

Rislorcelli 1947; Gaud 1954; Bailly-Choumara et al. 1971, AP, EM, MA, AA; Rioux et al. 1974, HA; Mouna 1998; Guernaoui et al. 2005; Boussaa et al. 2005; Boussaa et al. 2007; Boumezzough et al. 2009, HA, Marrakech; Boussaa et al. 2010; Bounamous 2010

Sergentomyia (Sergentomyia) minuta (Rondani, 1843)

= Phlebotomus minutus Rondani, in Gaud and Laurent 1952: 75, Mouna 1998: 86

= Phlebotomus (Sergentomyia) parroti (Adler and Theodor), in Rislorcelli 1941: 526, Rislorcelli 1947: 487, Bailly-Choumara et al. 1971: 450, Rioux et al. 1974, 99

Séguy 1930a, HA, Marrakech; Gaud and Laurent 1952, AP, Rabat; Rislorcelli 1941; Rislorcelli 1947, HA; Bailly-Choumara et al. 1971, Rif, AP, EM, MA, HA; Rioux et al. 1974, HA; Croset et al. 1978 (from the mediterranean region to the Sahara); Mouna 1998; Boussaa et al. 2005; Boumezzough et al. 2009, HA, Marrakech; Bounamous 2010; Boussaa et al. 2010; Depaquit et al. 2015, Rif, Chefchaouen

Sergentomyia (Sergentomyia) schwetzi Adier, Theodor & Parrot, 1929

Bailly-Choumara and Léger 1976, SA, Aouinet-Torkoz; Mouna 1998

Sergentomyia (Sintonius) christophersi (Sinton, 1927)

Rioux et al. 1974, HA; Rioux et al. 1975; Croset et al. 1978; Mouna 1998

Psychodinae

Maruinini

Tonnoiriella Vaillant, 1982

Tonnoiriella paveli Ježek, 1999

Ježek 1999, HA, AA; Afzan and Belqat 2016

Tonnoiriella pulchra (Eaton, 1893) (?) [probably mis-identification]

Wagner 1990; Ježek and Hájek 2007; Afzan and Belqat 2016

Mormiini

Mormia Enderlein, 1935

Mormia tenebricosa Vaillant, 1954

= Telmatoscopus (Mormia) tenebricosus Vaillant, in Vaillant 1956b: 244

Vaillant 1956b, HA, Imi-N’Ifri; Afzan and Belqat 2016, Rif, Oued Achekrade

Paramormiini

Clogmia Enderlein, 1937

Clogmia albipunctata (Williston, 1893)

Afzan and Belqat 2016, Rif, Douar Kitane, Douar Moukhlata, Oued Mhannech, AP, Douar Aoulad Ali (Central Plateau (Coastal region))

Panimerus Eaton, 1913

Panimerus thienemanni (Vaillant, 1954)

= Panimerus maynei (Tonnoir, 1919), in Dakki 1997: 62

Boumezzough and Vaillant 1986b, HA, Assif Réghaya; Afzan and Belqat 2016

Paramormia Enderlein, 1935

Paramormia ustulata (Walker, 1856)

Vaillant 1956b, HA; Mouna 1998; Ježek and Yağcı 2005; Omelková and Ježek 2012a; Afzan and Belqat 2016, Rif, Seguia Barrage Dar Chaoui, Douar Kitane, Oued Jnane Niche

Pericomini

Bazarella Vaillant, 1964

Bazarella atra (Vaillant, 1955)

= Pericoma atra Vaillant, in Vaillant 1956b: 234, 238, 242

Vaillant 1956b, HA, Assif Tasouat (M’Goum), Siroua, Imi-N’Ifri, Aguelmous, Sidi Chamarouch, Lac Tamhda (Anremer), Oukaimeden; Boumezzough and Thomas 1987, HA, Oued Réghaya (Imlil, 1750 m); Dakki and Himmi 2008; Afzan and Belqat 2016, Rif, Oued Inesmane, Oued Madissouka, Aïn Quanquben

Pericoma Walker, 1856

Pericoma (Pachypericoma) blandula Eaton, 1893

Boumezzough and Vaillant 1986b, HA; Dakki 1997; Ježek 2004, Rif; Ježek and Hájek 2007; Dakki and Himmi 2008, MA, Oued Sebou; Omelková and Ježek 2012a; Afzan and Belqat 2016, Rif, Oued Taida, Âounsar Aherman, Oued Beni Ouachekradi, Oued Aâyaden, Cascade Ras el Ma

Pericoma (Pericoma) barbarica Vaillant, 1955

Vaillant 1956b, HA, M’Goum; Afzan and Belqat 2016, Rif, Oued Taida, Douar Taria, Cascade Grotte des pigeons

Pericoma (Pericoma) granadica Vaillant, 1978

Boumezzough and Vaillant 1986b, HA; Vaillant and Moubayed 1987; Dakki 1997; Dakki and Himmi 2008, MA, Oued Sebou; Afzan and Belqat 2016, Rif, Oued Taida, Ametrasse, Oued Farda, Oued Aâyaden, Oued Ras el Ma, MA, Aïn Vittel, HA, Cascade sur sol cuivreux, Oued Réghaya

Pericoma (Pericoma) diversa Tonnoir, 1920

Vaillant 1978, HA; Afzan and Belqat 2016, Rif, Cascade Chrafate

Pericoma (Pericoma) exquisita Eaton, 1893

Ježek 2004, Rif, HA; Afzan and Belqat 2016

Pericoma (Pericoma) latina Sarà, 1954

Vaillant 1955, HA; Afzan and Belqat 2016, Rif, Cascade Chrafate, Oued Maggou, Nord Village Maggou, Oued Kelâa, Oued Talembote, Oued associé à Dayat Fifi, Oued Tiffert, Oued à 20 km de Fifi, Oued El Kanar, Beni Fenzar

Pericoma (Pericoma) maroccana Vaillant, 1955

= Pericoma numidica var. marocana Vaillant, 1955

Boumezzough and Vaillant 1986b, HA, Tissaout; Dakki 1997; Afzan and Belqat 2016, Rif, Cascade Chrafate, ruisseau maison forestière; Dakki and Himmi 2008, MA, Oued Sebou

Pericoma (Pericoma) modesta Tonnoir, 1922

= Pericoma numidica Vaillant, in Vaillant 1956b: 236, 237, 240

Vaillant 1956b, HA, Assif Tassouat (M’Goum), Lac Tamhda (L’Anremer); Boumezzough and Vaillant 1986b, HA; Dakki 1997; Dakki and Himmi 2008, MA, Oued Sebou; Afzan and Belqat 2016

Pericoma pseudexquisita Tonnoir, 1940

Afzan and Belqat 2016, Rif, Oued Azila

Pneumia Enderlein, 1935

Pneumia nubila (Meigen, 1818)

Afzan and Belqat 2016, Rif, Aïn Mâaze

Pneumia pilularia (Tonnoir, 1940)

Ježek 2004; Ježek and Hájek 2007; Omelková and Ježek 2012a

Pneumia propinqua (Satchell, 1955)

Afzan and Belqat 2016, Rif, Chrafate, Oued Zarka

Pneumia reghayana (Boumezzough & Vaillant, 1986)

= Satchelliella reghayana Boumezzough & Vaillant, 1986, in Boumezzough and Vaillant 1986b: 238

Boumezzough and Vaillant 1986b, HA; Dakki 1997; Dakki and Himmi 2008; Afzan and Belqat 2016

Pneumia toubkalensis Omelková & Ježek, 2012

Omelková and Ježek 2012b, HA, Toubkal; Afzan and Belqat 2016, Rif, Oued Aâyaden, Aïn Ras el Ma

Psychodini

Philosepedon Eaton, 1904

Philosepedon (Philosepedon) humeralis (Meigen, 1818)

Afzan and Belqat 2016, Rif, Oued Hachef, Cascade Ras el Ma, Oued El Kanar, 2 km de Douar Assoul, Oued Aâyaden

Psychoda Latreille, 1796

Psychoda (Logima) albipennis (Zettersdedt, 1850)

= Logima albipennis (Zettersdedt), in Mouna 1998: 86

Mouna 1998

Psychoda (Psycha) grisescens Tonnoir, 1922

Ježek 2004, Rif; Afzan and Belqat 2016, Rif, Douar Kitane, MA, Gîte Aït Ayoub

Psychoda (Psychoda) uniformata Haseman, 1907

Ježek 2004, Rif; Afzan and Belqat 2016

Psychoda (Psychodocha) cinerea Banks, 1894

Boumezzough and Thomas 1987, HA, Azib Oukaimeden (2730 m); Dakki 1997; Dakki and Himmi 2008; Afzan and Belqat 2016, Rif, Oued Tazzarine, Douar Taria, Douar Kitane, Oued Chrafate, Oued Aâyaden, EM, Cascade Grotte des Pigeons (Béni Snassen)

Psychoda (Psychodocha) gemina (Eaton, 1904)

Afzan and Belqat 2016, Rif, Dayat Fifi, Oued Zarka, Douar Kitane, Oued Aâyaden

Psychoda (Tinearia) alternata Say, 1824

Tonnoir 1920, HA, La Maire; Boumezzough and Thomas 1987, HA, Oued Réghaya (1740 m), Imlil; Dakki 1997; Dakki and Himmi 2008; Afzan and Belqat 2016, Rif, Oued Nakhla, Oued Farda, Oued Ouara, Oued Ametrasse, Oued Chrafate, Douar Derâa, Douar Ihermochene, Douar Ikhlafene, Douar Taria, Douar Idrene, Douar Kitane, Oued 2 km de Douar Assoul, Oued Aâyaden, ruisseau maison forestière, Oued Mhannech, Aïn Sidi Yahia, MA, Gîte Aït Ayoub

Scatopsoidea

SCATOPSIDAE3

K. Kettani, J.P. Haenni

Number of species: 13. Expected: 30–40

Faunistic knowledge of the family in Morocco: poor

Ectaetiinae

Ectaetia Enderlein, 1912

Ectaetia clavipes (Loew, 1846)

= Scatopse clavipes Loew, in Mouna 1998: 84

Mouna 1998; Haenni and Kettani 2016, Rif, Amsa – MISR

Psectrosciarinae

Anapausis Enderlein, 1912

Anapausis sp.*

Rif, Source Aïn Tissemlal

Psectrosciara Kieffer, 1911

Psectrosciara sp. 1*

Rif, Adrou

Psectrosciara sp. 2*

Rif, Adrou

Scatopsinae

Coboldia Melander, 1916

Coboldia fuscipes (Meigen, 1830)

Haenni and Kettani 2011, Rif, Kitane, AP, El Jadida, Rabat, AA, Souss; Haenni and Kettani 2016, Rif, Maggou, Aârkob, Jnane Niche; Rif (M’Diq farm) – MISR

Parascatopse Cook, 1955

Parascatopse sp.

Haenni and Kettani 2011, AA, Ouarzazate ­– MHNN

Quateiella Cook, 1975

Quateiella inexpectata Haenni, 1988

Haenni and Kettani 2016, Rif, Afertane

Reichertella Enderlein, 1912

Reichertella geniculata (Zetterstedt, 1850)

Haenni and Kettani 2016, Rif, Jnane Niche

Reichertella maroccana Haenni, 2011

Haenni and Kettani 2011, HA, Oukaimeden – RMNH

Rhegmoclemina Enderlein, 1936

Rhegmoclemina lunensis Haenni & Godfrey, 2009

Haenni and Kettani 2011, Rif, Boujdad

Rhexoza Enderlein, 1936

Rhexoza freyi (Duda, 1936)

Haenni and Kettani 2011, AA, Agadir – MHNN

Scatopse Geoffroy, 1762

Scatopse notata (Linnaeus, 1758)

Mouna 1998; Haenni and Kettani 2016, Rif, Onsar Lile – MISR

Swammerdamella Enderlein, 1912

Swammerdamella brevicornis (Meigen, 1830)

Haenni and Kettani 2011, Rif, Kitane, Oued Laou, Ketama, AP, El Jadida, Essaouira, HA, Ouarzazate; Haenni and Kettani 2016, Rif, Amsa, Jnane Niche, Ametrasse – MISR

New records for Morocco

An undescribed species of Anapausis has been collected in the Rif mountains in 2014 by K. Kettani and will be described elsewhere.

Anapausis sp.

Rif: Forêt Azilane (NPT), Source Aïn Tissemlal, 1255 m, 35°11.67N, 5°15.20W, 7.vi.2014, Sapinière à Abies maroccana et Pinus nigra, 1♂.

Two undescribed species of Psectrosciara have been collected in the Rif mountains in 2013 by K. Kettani and will be described elsewhere.

Psectrosciara sp. 1

Rif: Taghzout (PNPB), Adrou, 556 m, 35°22.39N, 05°32.28W, chênes-liège, 14.vii–15.viii.2013, 6♂♂.

Psectrosciara sp. 2

Rif: Taghzout (PNPB), Adrou, 556m, 35°22.39N, 05°32.28W, chênes-liège, 14.vii–15.viii.2013, 3♂♂.

PTYCHOPTERIDAE

K. Kettani, R. Wagner

Number of species: 5. Expected: 8

Faunistic knowledge of the family in Morocco: moderate

Ptychopterinae

Ptychoptera Meigen, 1803

Ptychoptera albimana (Fabricius, 1787)

MISR (No locality given)

Ptychoptera contaminata (Linnaeus, 1758)

MISR (No locality given)

Ptychoptera lacustris Meigen, 1830

MISR (No locality given)

Ptychoptera paludosa Meigen, 1804

MISR (No locality given)

Ptychoptera scutellaris Meigen, 1818

MISR (No locality given)

Culicoidea

CHAOBORIDAE

K. Kettani, R. Wagner

Number of species: 2. Expected: 4

Faunistic knowledge of the family in Morocco: poor

Chaoborinae

Chaoborus Lichtenstein, 1800

Chaoborus crystallinus (De Geer, 1776)

Dakki 1997: 60

Mochlonyx Loew, 1844

Mochlonyx culiciformis (De Geer, 1776)

Dakki 1997: 60

CULICIDAE

K. Kettani, B. Trari, O. Himmi, M. Dakki

Number of species: 43. Expected: 60

Faunistic knowledge of the family in Morocco: good

Anophelinae

Anopheles Meigen, 1818

Anopheles (Anopheles) algeriensis Theobald, 1903

Viallate 1922, AP, Kénitra; Séguy 1930a; Bonjean 1947, EM, MA; Gaud 1957a, HA, north of High Atlas; Guy 1959a, MA, Béni Mellal, HA; Guy 1959b, HA; Guy 1959a, MA, Béni Mellal; Bailly-Choumara 1967a, MA, Ghorm El Alem; Benmansour et al. 1972, MA, Barrage Bin El Ouidane; Bailly-Choumara 1973b, AP, Sidi Yahia du Gharb; Metge 1986, AP, Casablanca; Trari and Himmi 1987, AP; Himmi et al. 1995; Louah 1995, Rif, Tahaddart, Schroda; Dakki 1997; Ramdani 1997, AP, Skhirat, Casablanca; Trari et al. 2002; Trari et al. 2004b; Himmi 2007, Rif, Chefchaouen; El Ouali Lalami et al. 2010 a,b, MA, Fès, Boulmane; El Ouali Lalami 2012, MA, Fès; Trari 2017, Rif, Chefchaouen, AA, Tiznit; Trari and Dakki 2017a, Rif, Chefchaouen, AA, Tiznit; Trari and Dakki 2017b, AA, Tiznit; Trari et al. 2017

Anopheles (Anopheles) claviger (Meigen, 1804)

Vialatte 1923, AP, Kénitra; Séguy 1930a, AP, Rabat; Langeron 1938, HA, Tounfit, Massou, Anefgou, Tirghist, Tighermine, Louggouargh; Callot 1940, HA, Anefgou, Tirghist; Bonjean 1947, EM, MA; Gaud 1947c, MA, Sefrou, Meknès; Gaud 1948, AP, Rabat, MA, El Hajeb, AA, Errachidia, Tadla; Gaud et al. 1948, AP, Skhirat; Guy 1963, Rif, Taounate, MA, Meknès, Ifrane; Bailly-Choumara 1967a, MA, piste Tafechna-Taoujgelt, maison forestière Ouiouane, piste Tafechna-Senoual-Itzer, piste Ksiba-Naour, piste Naour-Arbala, Zaouia Cheikh, Oued Sarif (environs El Ksiba), Dayet Aoua (environs Ifrane), Boulemane; Bailly-Choumara 1967b, EM, 8 km N Itzer; Bailly-Choumara 1967c, Rif, piste Ketama-Mt Tiguidin; Guy 1967, HA, Marrakech, AA, Tafilalt; Guy and Holstein 1968, SA; Metge 1986, AP, Casablanca; Himmi et al. 1995; Louah 1995, Rif, Haidra, Marina Smir; Dakki 1997; Ramdani 1997, AP, Skhirat, Tamaris drains, Tamaris merja; Trari et al. 2002; Trari et al. 2004b; Himmi 2007, Rif, Bab Berred, Tanakoub; Faraj et al. 2008b, Rif, Assoul, Mizgane; El Ouali Lalami et al. 2010a, MA, Boulmane; Larhbali et al. 2010, MA, Zhiliga, Boukachmir, Aït Ichou; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Anopheles) labranchiae Falleroni, 1926

d’Anfreville 1916, AP, Salé; Delanoe 1917, AP, El Jadida; Viallate 1922, AP, Rabat, Boulhaut, Bouznika, Gharb, MA, Sidi Kacem, Tiflet, Fès, Taza; Charrier 1924, Rif, Tanger; Séguy 1930a, MA; Roubaud 1935, AP, Rabat; Sicault et al. 1935, AP, Merja Ras Eddaoura, Merja Zerga, dayas entre Sebou et Maâmora, Dar bel Hamri (entre barrage El Kansera et Sidi Slimane), à proximité de Merja Zerga, Douar Anabsa; Langeron 1938, AA; Callot 1940, AA; Ristorcelli 1946a, b, HA, Oued Tensift, Oued Issil; Bonjean 1947, AP, Gharb; Gaud 1947c, MA, Oulmès, HA, Marrakech; Gaud et al. 1948, AP, Merja Ras Eddaoura; Gaud et al. 1949, AP, Salé, SA, Foum Zguid, Tagounit; Gaud 1953a, AP, from Tanger to El Jadida, MA, Tissa, Timhadit, Bekrit, Meknès, Ifrane (1700 m), HA, Sidi Aissa, Tizi-n’Tichka; Guy 1958, HA, Oued N’fis; Sacca and Guy 1960b, Rif, Tétouan, AP, Skhirat, Sidi Yahia, Sidi Bettache, Mazagan, Braila (près Sidi Allal Tazi), Aït Lahsen, MA, Meknès, HA, Marrakech; Guy 1962, Rif, Taounate, AP, Gharb, HA, Marrakech (ville et banlieu); Guy 1963, Rif, Tanger, Tétouan, EM, Berkane, Debdou, Oujda, AP, Kénitra, Souk Larba, Settat, Chamaîa, Safi, Casablanca, Rabat, Essaouira, MA, Meknès, Fès, Azrou, Oued Zem, Béni Mellal, HA, Kelaâ of Sraghna; Bailly-Choumara 1967a, MA, piste Tafechna-Znan Imes, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, Aguelmane Azigza, piste Aguelmane Azigza-Aïn Leuh, maison forestière Ouiouane, route Khénifra-Tafechna, piste Tafechna-Senoual-Itzer, Ajdir, Itzer, Bords Moulouya, piste Itzer-Boumia, RN P.33 Boumia-El Kbab, Ouaoumana, piste Ksiba-Naour, piste Naour-Arbala, piste Arbala-El Kbab, Kafensour, Oued Sarif (environs El Ksiba), Dayet Aoua, Ifrane, Imouzer Marmoucha; Bailly-Choumara 1967b, EM, route Itzer-Midelt, Douar Sherba, Douar Aïn Shebbak, Douar Aïn Zabia, Douar Madarh, Douar Sidi Hashas, Mechraa safsaf; Bailly-Choumara 1967c, Rif, route Chaouen-Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, piste Bab Taza-Bab Berred, Anasar, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, Al Hoceima Club Med, Béni Bouayache, Targuist et environs; Guy 1967, HA, Marrakech, AA, Tafilalt; Bailly-Choumara 1968b, AP, Larache; Guy and Holstein 1968, AA, Ouarzazate; Bailly-Choumara 1970, Rif, Tétouan, EM, Berkane, AP, Larache, Sidi Yahia du Gharb, HA, Marrakech; Benmansour et al. 1972, AA, Agadir; Bailly-Choumara 1972a, AP, Merja Sheishat; Bailly-Choumara 1972b, AP, Merja Sheishat; Bailly-Choumara 1973a, AP, Merja de l’Oued Smir; Bailly-Choumara 1973b, Rif, Tétouan, EM, Berkane, AP, Merja Sheishat, HA, Souk des Oudaias; de Zulueta et al. 1983, Rif; Ibn Jilali 1984, AP, Maâmora; Metge 1986, AP, Entre Oulad Dlim et Al Ara’ra; Trari and Himmi 1987, AP, Gharb; Himmi 1991, AP, Kénitra, Maâmora; Metge 1991, AP, Sidi Bettache; Trari 1991, Rif, Chefchaouen, Tanger, Taounate, EM, Oujda, AP, Sidi Amira, Sidi Boughaba, Merja Zerga, Sidi Allal Tazi, Aïn Chouk, Oued Loukous, Merja Oulad Skhar, MA, Khémissat, Béni Mellal, Khouribga, HA, El Kelaâ des Sraghrna, AA, Ouarzazate; El Bermaki 1993, AP, entre l’aeroport Anfa et l’aménagement d’El Oulfa, Sidi Maârouf, El Oulfa; Chlaida and Bouzidi 1995, AP, Aïn Blal, Oued Sidi Messoud; Louah 1995, Rif, Tres piedras, Marina Smir, Bouzerlal, Oued Maleh, Zekri, Lajour, Azla, Tahaddart, Skroda, Stehat, Moulay Bouchta, Kebbache, Talembote, Loubart, Chefchaouen, Oued Maggou, Bab Berred, Sidi Kankoch, Oued kbir, Oued Jebel Lehbib; Louah et al. 1995; Chlaida 1997, AP, Aïn Blal, Oued Sidi Messoud, Douar Chlihat, Barrage Al Massira; Moussalim 1997, AP, Sidi Allal Tazi, Maâmora; Ramdani 1997, AP, Tamaris, Skhirat; Faraj et al. 1997, AP, Ouled Moussa; Handaq 1998, HA, Oukaimeden, Amizmiz, Tiguenziouine; Himmi et al. 1998, AP, Dayat d’El Menzeh (north-east of Kénitra), Sidi Boughaba; Alaoui Slimani et al. 1999, AP, Bou-Regreg Salé, Sidi Bouguettaya, Quartier industriel Takaddoum, Marjane; Alaoui Slimani 2002, AP, Rabat-Salé; Trari et al. 2002; Faraj et al. 2003, Rif, Azib Bouflou, Azib Jrou, Imzouren, Amezzaourou, Tizi-Tamalout, MA, Aït Abdelsalam, Aït Lamfadel; Faraj et al. 2004, Rif, Azib Jrou, Tanghaya Akarkar, Amezzaourou, Ouled Nsar, AP, Fedalate, MA, Talaa Chougaga, Aïn Smen (Fès), Aït Lamfadel; Trari et al. 2004a, Rif, Larache; Trari et al. 2004b, Rif, Ketama, Gzenaya, Taounate, Bouzaghlal, Oued Laou, Smir (Merja), Béni Hassane, Azib Jrou, Azib Bouflou, AP, Laaouamra, Aarabat Sidi Abdelaziz, Moukaouama, Dar Belamri, Laksibia, Mgadid, Beggara, Rabat (Chellah), MA, Adouz, Rommani (Khémisset), Aït Abderrahmane, Aït Ishaq, Oulad Messoud, Ouled Fennane, El karma, Oulad Zguida, Oulad Abbou; Aouinty et al. 2006, AP, Mohammedia; Himmi 2007, Rif, Bab Berred, Bab Taza, Tanakoub, AP, Skhirate, Maâmora, Oulja, Bouknadel, Sidi Azzouz, Tamaris, MA, Khémisset; Faraj et al. 2008a, AP, Laouamra, Boucharen, MA, Béni Khlef, Talaa Chougaga; Faraj et al. 2008b, Rif, Assoul, Mizgane; El Ouali Lalami et al. 2010b, MA, Fès; Faraj et al. 2010, AP, Begara, Boucharen, Ben Slimane, Skhirat, Rabat, Sehoul, MA, Sidi Allal Msader, Aïn Aghbal, Aïn Elouali, Sidi Kacem; Larhbali et al. 2010, MA, Aït Haddou Said; Adlaoui et al. 2011, AP, Larache; Larhbali et al. 2011, MA, Oulmès, Aït Yadin, Sfassif, Mâaziz, Rommani, Laghoualem, Ezzhiliga, Sidi Allal Bahraoui, Boukachmine, Aït Malek, Sidi Boukhalkhal, Bni Ounzar, Ganzra, Aït Siberne, Sidi Allal Msader, El Ghandour; El Ouali Lalami 2012, MA, Pont Diamant vert, Sidi Harazem, Oued El Himmer, Moulay Yakoub, Oued Sebou, Aïn Kansara, Oued Aïn Chkef, Sefrou, Boulemane; Laboudi et al. 2012, AP, Larache; Hadji et al. 2013, AP, Sidi Slimane; El Joubari et al. 2014, Rif, Smir lagoon; Laboudi et al. 2014, Rif, Tétouan, Tanger, Chefchaouen, Al Hoceima, AP, Larache, Salé, MA, Taza, Khémissat; El Joubari et al. 2015a, Rif, Smir lagoon; El Joubari et al. 2015b, Rif, Smir lagoon; Marc et al. 2016, AP, Kénitra; Trari 2017, Rif, Chefchaouen, Tétouan, AP, Larache, Rabat, Settat, EM, Oujda, MA, Khémisset, Meknès, Khouribga, HA, Marrakech, AA, Tiznit, Ouarzazate; Trari and Dakki 2017a, Rif, Chefchaouen, Tétouan, AP, Larache, Rabat, Settat, EM, Oujda, MA, Khémisset, Meknès, Khouribga, HA, Marrakech, AA, Tiznit, Ouarzazate; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Anopheles) marteri Senevet & Prunnelle, 1927

Gaud 1945b, MA, El Hajeb, Khénifra, HA, Tizi-n’test, Tillougite; Gaud et al. 1949, HA, Tizi-n’test; Gaud 1953b, HA, Tizi-n’test; Bailly-Choumara 1967b, EM, Grotte du Zegzel; Bailly-Choumara 1967c, Rif, route Chaouen-Bab Taza, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Taza-Asifane, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, Boured; Benmansour et al. 1972, MA, Taza; Trari 1991, Rif, Taounate; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Himmi 2007; Trari 2017, Rif, Chefchaouen, AP, Settat; Trari and Dakki 2017a, Rif, Chefchaouen, AP, Settat; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Anopheles) ziemanni Grünberg, 1902

Senevet 1935, MA; Gaud et al. 1949, HA, plain of south and north of the Occidental Atlas; Gaud et al. 1950, HA, plain of south and north of the Occidental Atlas; Guy 1958, HA, Marrakech; Guy et al. 1958, HA, Oued N’fis; Guy 1967, EM, Oujda, AP, Rabat, HA, Marrakech, AA, Tafilelt; Bailly-Choumara 1970, HA, Marrakech; Benmansour et al. 1972, MA, Taza, HA, Haouz, Tadla Azilal; Bailly-Choumara 1973b, HA, Souk des Oudaias (Souk Tnine des Oudaias, 470 m); Trari 1991, MA, Tissa; Moussalim 1997, AP, 7.5 km de Sidi Allal Tazi; Trari et al. 2002; Trari et al. 2004b; Himmi 2007; Trari 2017, Rif, Tétouan, AA, Tiznit; Trari and Dakki 2017a, Rif, Tétouan, AA, Tiznit; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Cellia) cinereus Theobald, 1901

Viallate 1922, MA, Sefrou, Sidi Lamine, HA, Mtougui; Sicault et al. 1935, AP, Souk Larba of Gharb; Senevet 1935, AP, Souk Larba of Zemmour; Langeron 1938, HA, Anefgou, Tirghist, Valley of Sidi Yahia Ouyoussef, Tighermine, Louggouargh, Massou; Callot 1940, HA, Anefgou, Tirghist; Gaud 1945a, Rif, Meridional Rif, EM, Moulouya, HA, Marrakech, AA, Tansikht, valley de Sous; Gaud and Duthu 1945, HA, Marrakech; Viamonte and Ramirez 1945, Rif; Viamonte and Ramirez 1946, Rif; Ristorcelli 1946a, HA, Oued Tensift, Oued Issil; Ristorcelli 1946b, HA, Oued Tensift, Oued Issil; Gaud 1945b, AA, Tansikht; Gaud et al. 1949, Rif, Meridional Rif, EM, Moulouya, HA, Marrakech, AA, Tansikht, valley de Sous; Gaud et al. 1950, Rif, Meridional Rif, EM, Moulouya, HA, Marrakech, AA, Tansikht, valley de Sous; Gaud 1953a; Gaud 1958, HA, Marrakech; Guy 1962, Rif, Taounate, HA, Marrakech; Guy 1963, MA, Midelt, AA, Hamada of Draa; Bailly-Choumara 1965, EM, Aman d’Aït Oussa, Tiglit, El Megrinat, Taskala, Aïn Aït delouine, Oued mesdourt, Talmesdourt, Assa, AA, Aït Melloul, Oued Teima, Issen, Taroudant, Talaint, Tiznit, Oued Assaka, Anezi, Pont de la route Agadir-Tiznit, valley of low Draa, Tafraoute, Tacharicht, Bou Izakarn, Jemâa N’tirhirte, Aït Erkha,Tazert, Barrage Taourirt, AA, Goulmima, SA, Aouinet Torkoz, Tirh Mzoun; Bailly-Choumara 1966, AA, piste Foum Zguid-Lac Iriqui, Agadir-Tissint, Akka-Iguiren, Tirhem, Taoujgelt, Souk El Khémis Dades, Akka, Aït Ouabelli, Foum-el-Hassan, Tarhjicht, Aït melloul, Taliouine, Tazenakht (Rocade of Draa); Bailly-Choumara 1967a, MA, route Azrou-Khénifra, Jnane Imasse, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, Sources Oum-er-Rbia, Itzer, piste Itzer-Boumia, Ouaoumana, piste Ghorm El Alem-El Ksiba, piste Naour-Arbala, Zaouia Cheich, kafensour, Ifrane, Imouzzer Marmoucha; Bailly-Choumara 1967b, EM, 7 km N Itzer, 8 km N Itzer, route Itzer-Midelt, Aïn Srouna, Gouttitir, Cascade Oued Za, Grotte du Zegzel, Douar Aïn Soultane, Mechraa Safsaf, Oujda, Berguent (valley of Moulouya), Figuig; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Berred-Assifane, Anasar, piste Bab berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, route Al Hoceima-Arba Taourirt, Arba Taourirt, Targuist et environs, route Targuist Al Hoceima, Jebha, Oued Ouergha, route Aknoul-Al Hoceima; Guy 1967, AP, Rabat, EM, Oujda, HA, Marrakech, AA, Tafilalet; Bailly-Choumara 1970, HA, Marrakech; Bailly-Choumara 1973b, HA, Souk des Oudaias; Trari 1991, Rif, Al Hoceima, Chefchaouen, Taounate, EM, Nador, Oujda, Figuig, AP, Larache, Settat, Ben Slimane, MA, Khénifra, Taza, Khouribga, HA, Kelaâ of Sraghna, AA, Ouarzazate, Goulmima; Bouallam 1992, HA, Oued N’fis; Louah 1995, Rif, Marina Smir, Bouzerlal, Tahaddart, Stehat, Moulay Bouchta, Kebbache, Loubart, 0ued Maggou; Louah et al. 1995; Bouallam et al. 1997b, HA, Marrakech; Handaq 1998, HA, Amizmiz, Tiguenziouine; Bouallam 2001, HA, Oued N’fis; Trari et al. 2002; Trari et al. 2004b, Rif, Ketama, Azib Jrou, Sidi Mokhfi, MA, Taghzirt, Aghbala, Aït Shak, Smaala, Mlalih, Ouled Fennane, Béni Khlef, Tachrafte; Faraj et al. 2007, Rif, Assoul, Mizgane; Himmi 2007, Rif, Bab Berred, Bab Taza, Stehat, Tanakoub; Faraj et al. 2008, Rif, Assoul, Mizgane; El Ouali Lalami et al. 2010a, MA, Fès; El Ouali Lalami et al. 2010b, MA, Oued El Himmer; Larhbali et al. 2010, MA, Roumani Aïn Sbite, Jamâa M. B., Ghoualem, Zhiliga, Oulmès, Tarmilate, Boukachmir, Mrirte, Aït Ichou, Mâaziz, Tiddas, Bni Ounzar, Ganzra; El Ouali Lalami 2012, MA, Oued El Himmer; Trari 2017, Rif, Chefchaouen, Tétouan, MA, Khouribga, AA, Tiznit; Trari and Dakki 2017a, Rif, Chefchaouen, Tétouan, MA, Khouribga, AA, Tiznit; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Anopheles (Cellia) dthali Patton, 1905

Saccà 1960, AA, Aoufous, Meski, Erfoud, Agdz, Zogora, Tagounit, Tamsrruth; Guy 1961, HA, Sud de Zagora (en bordure Hamada du Draa); Guy 1963, AA, Zagora and south of the High Atlas (at edge of Hamada of Draa), Oued Ziz; Bailly-Choumara 1965c, AA, Tiznit, EM, Aïn Aït Delouine, Aouinet Torkoz, Rich Tamlougout (eastern borders of Jebel Bani); Bailly-Choumara 1966, AA, piste Foum Zguid au Lac Iriqui, Agadir-Tissint, Akka-Iguiren, Souk El Khémis Dades, Akka et Environs, Aït Ouabelli, Tarhjicht, piste Tazenakhte à Foum Zguid (Rocade du Draa); Bailly-Choumara 1967b, EM, valley of Moulouya; Guy 1967, HA, Marrakech, AA, Tafilalet; Guy and Holstein 1968, EM, N Outat El Haj (valley of Moulouya), Gouttitir (environs de Taourirt); Bailly-Choumara 1970, AA, Foum Zquid; Bailly-Choumara 1973b, AA, Foum Zquid; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Trari et al. 2004b; Faraj et al. 2007, Rif, Assoul, Mizgane (SE of Bab Berred); Faraj et al. 2008, Rif, Assoul, Mizgane (SE Bab Berred); Himmi 2007; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Cellia) multicolor Cambouliu, 1902

Messerlin and Treillard 1938, HA, Marrakech; Viamonte and Ramirez 1945, Rif; Guy 1963, SA; Bailly-Choumara 1965c, EM, Aït Oussa, Aman d’Aït Oussa, Aïn Aït Delouine, Aouinet Torkoz, Rich tamlougout (Confins orientaux du Jebel Bani), AA, Tafnidilt, Guelta Zerga, Aïn Temda (valley of low Draa), Tirhmert (Goulmima), SA, Vallée et embouchure de l’Oued Assaka, Tantan ville, Tirh Mzoun; Bailly-Choumara 1966, AA, Rocade of Draa; Bailly-Choumara 1967b, EM, 40 km N de Outat El Haj, Gouttitir, Cascade Oued Za, Douar Aïn Shebbak (valley of Moulouya); Guy 1967, HA, Marrakech, AA, Tafilelt; Guy and Holstein 1968, HA, south of Atlas, AP, plain located between Marrakech and the Atlantic from Tanger along the length of the Mediterranean; Bailly-Choumara 1970, AA, Foum Zquid; Bailly-Choumara 1973b, AA, Foum Zquid; Metge 1986, AP, Casablanca; Trari 1991, Rif, Al Hoceima, Taounate, Oued Maleh, Bab Berred, AP, Tamaris Merja, AA, Ouarzazate; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Trari et al. 2004b; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Cellia) sergentii (Theobald, 1907)

Séguy 1930a; Messerlin and Treillard 1938, HA, Tamelelt; Langeron 1938, Rif, Targhist; Callot 1940, AA, Taghjicht; Gaud 1947c, AA, Wadi Draa; Gaud et al. 1949, Rif, Zoumi, HA, Zaouia Sidi Hamza, Tizi-n’test (1700 m), Tillougit (1800 m); Gaud et al. 1950, Rif, Zoumi, MA; Guy et al. 1958, HA, Oued N’fis; Guy 1961, Rif, AP, south of Casablanca, AA, Sud de Zagora; Guy 1962, Rif, AP, south of Casablanca, HA, Marrakech; Guy 1963, Rif, Tanger, EM, Berkane, AP, south of Casablanca, MA, Béni Mellal, HA, Oued Tensift, Oued Ziz, Marrakech, Chichaoua, AA, Oued Draâ, Oued Dades, Goulmima, sud de Zagora, SA, Foum Zguid; Bailly-Choumara 1965c, EM, iglit, Aman d’Aït Oussa, Oued Isker, El Megrinat, Aïn Aït Delouine, Oued Mesdourt, almesdourt, Aouinet Torkoz, Bouanama, Rich Tamlougout, Assa, AA, Tiznit, Tafraoute, Oued Izi, Bou Izakarn, Abeino (region of Goulmima), SA, Aouzeroual, Tirhmert, Tacharicht, Jebel Bani, Tantan; Bailly-Choumara 1966, AA, Rocade of Draa, Taliouine; Bailly-Choumara 1967a, Rif, AP, south of Casablanca, MA, Itzer, Ghorm El Alem, Zaouia Cheikh, Kafensour, HA, north of High Atlas; Bailly-Choumara 1967b, EM, Aïn Srouna, Grotte du Zegzel, Douar Mardarh, Douar Aïn Soultane, Merja Boubker, Selouane, Driouch (valley of Moulouya); Bailly-Choumara 1967c, Rif, route nationale Jebha, Targuist-Béni Boufrah, Al Hoceima, Béni Bouayache, Marchica, Had El Rouadi, Pont du Srah; Guy 1967, HA, Marrakech, AA, Tafilelt; Guy and Holstein 1968, AP, Casablanca; Bailly-Choumara 1970, EM, Berkane, HA, Marrakech; Bailly-Choumara 1973b, EM, Berkane, HA, Marrakech; Metge 1986, AP, Casablanca; Trari 1991, Rif, Al Hoceima, Taounate, AP, Larache, AA, Ouarzazate; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Faraj et al. 2003, Rif, Al Hoceima, Chefchaouen, Taounate, HA, Khouribga; Trari et al. 2004b, Rif, Ketama, Sidi Mokhfi; Faraj et al. 2007, Rif, Assoul, Mizgane; Faraj et al. 2008, Rif, Assoul, Mizgane; El Ouali Lalami et al. 2010a, MA, Fès; Larhbaliet al. 2010, MA, Oulmès, Ganzra; Trari 2017, Rif, Chefchaouen, Tétouan, EM, Oujda, MA, Khouribga; Trari and Dakki 2017a, Rif, Chefchaouen, Tétouan, EM, Oujda, MA, Khouribga; Trari and Dakki 2017b; Trari et al. 2017; Benabdelkrim Filali et al. 2018; Mouatassem et al. 2019, MA, Fès

Culicinae

Aedini

Aedes Meigen, 1818

Aedes (Acartomyia) mariae (Sergent & Sergent, 1903)

Séguy 1930b; Messerlin 1938, AP, Rabat; Séguy 1930a, Rif, littoral méditerranéen; Bailly-Choumara 1967b, Rif, Al Hoceima; Bailly-Choumara 1967c, Rif, Al Hoceima; Bailly-Choumara 1968a, Rif, Al Hoceima, AP, Larache, Sidi Yahia, Sidi Allal Tazi, Rabat, HA, Marrakech, AA, Tiznit; Himmi et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Aedimorphus) vexans (Meigen, 1830)

Gaud 1947c, AP, Sidi Allal Tazi, MA, Khémisset; Metge 1986, AP, Littoral Casablanca; Himmi et al. 1995; Handaq 1998, AP, Gharbia; Dakki 1997; Trari et al. 2002; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Dahliana) echinus (Edwards, 1920)

Séguy 1924; Séguy 1930a; Gaud 1953a, AP, Rabat, Sidi Yahia, MA, Moulay Bouazza, Taza, Fès, Meknès, Ifrane; Bailly-Choumara 1965b, AP, Maâmora; Bailly-Choumara 1967c, Rif, piste Bab Taza-Talassemtane; Himmi et al. 1995; Trari et al. 2002; Nikookar et al. 2010; El Joubari et al. 2014; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Dahliana) geniculatus (Olivier, 1791)

Séguy 1924, MA; Séguy 1930a; Metge and El Alaoui 1987, AP, Subéraies de Béni Abid-Benslimane (Casablanca); Metge and Belakoul 1989, AP, Sidi Bettache; Himmi et al. 1995; El Ouali Lalami et al. 2010a, MA, Fès Boulmane; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Ochlerotatus) berlandi (Séguy, 1921)

Séguy 1930a, AP, Rabat; Gaud 1953a, MA, Fès; Bailly-Choumara 1967a, MA, Jnane Imasse, piste Tafechna-Taoujgelt; Bailly-Choumara 1967c, Rif, piste Bab Taza-Béni Ahmed; Belakoul 1985, AP, Benslimane, Sidi Bettache; Metge and El Alaoui 1987, AP, Casablanca; Metge and Belakoul 1989, AP, Benslimane, Sidi Bettache; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Ochlerotatus) caspius (Pallas, 1771)

Séguy 1930b; Viamonte et Ramirez 1946, AP, Larache; Gaud 1952, AP, Rabat, Casablanca; Gaud 1953a, AP, Rabat, Casablanca; Senevet and Andarelli 1954, EM, Embouchure de la Moulouya, Figuig, AP, Jorf Lasfar, Mohammedia, Rabat, MA, Meknès, Fès, Taza, HA, Marrakech, Midelt, AA, Tiznit; Bailly-Choumara 1966, AA, environs de Tiznit; Bailly-Choumara 1967a, EM, Bords Moulouya (près Itzer), Cherarba; Bailly-Choumara 1967b, EM, Cherarba, Aïn Shebbak, Saidia, Berguent; Bailly-Choumara 1967c, Rif, piste Al Hoceima-Arba Taourirt; Bailly-Choumara 1972a, AP, Merja Sheishat; Bailly-Choumara 1973b, Rif, Merja de l’Oued Smir; El Kaim 1972, AP, Bou-Regreg; Rioux et al. 1975, AP, Rabat-Salé; Metge 1986, AP, Littoral casablancais; Himmi 1991, AP, Sidi Boughaba; Trari 1991, AP, Sidi Boughaba, Merja Zerga, Oued Loukous; Himmi et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi, Kénitra; Ramdani 1997, AP, Tamaris Merja; Handaq 1998, AP, Zemamra, B. Iffou (Entre El Oualidia et Youssoufia), MA, Béni Mellal, HA, Marrakech, Zaouiet Ben Sassi, Bengrir; Himmi et al. 1998, AP, Sidi Boughaba; Trari et al. 2002; Himmi 2007, AP, Sidi Boughaba; Alaoui Slimani 2002, AP, Rabat; Aouinty et al. 2006, AP, Mohammedia; El Joubari et al. 2014, Rif, Smir lagoon; El Joubari et al. 2015a, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Ochlerotatus) coluzzii Rioux, Guilvard & Pasteur, 1998 et Aedes (Ochlerotatus) detritus (Haliday, 1833) [Complexe detritus]

Charrier 1924, Rif, Tanger; Séguy 1930a; Gaud 1953a, EM, Saidia, AP, Kénitra, Rabat, Bouznika, El jadida, Oualidia, HA, Marrakech, AA, Agadir, Tafnidilt; Bailly-Choumara 1965c, EM, Aïn Aït delouine, SA, Tirhmert; Bailly-Choumara 1970, Rif, Tétouan; Knight 1971, AP, Kénitra; El Kaim 1972, AP, Bou-Regreg; Bailly-Choumara 1973b, Rif, Merja de l’Oued Smir; Rioux et al. 1975, AP, Rabat; Pasteur et al. 1978, AP, Bou-Regreg; Metge 1986, AP, Littoral casablancais; Himmi 1991, AP, Sidi Boughaba; Trari 1991, AP, Sidi Boughaba, Merja Zerga, Oued Loukous; Louah 1995, Rif, Tres piedras, Cabo Negro, Lajour, Azla, Tahaddart; Himmi et al. 1995; Louah et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi, Rabat; Ramdani 1997, AP, Tamaris Merja; Handaq 1998, AP, Essaouira, Zima-Chemaîa; Himmi et al. 1998, AP, Sidi Boughaba; Himmi 2007, AP, Sidi Boughaba; Alaoui Slimani 2002, AP, Rabat; Trari et al. 2002; El Joubari et al. 2014, Rif, Smir lagoon; El Joubari et al. 2015a, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Ochlerotatus) pulchritarsis (Rondani, 1872)

Gaud 1953a, AP, Benslimane, Sidi Yahia, Rabat, MA, Oued Zem, Khénifra, Fès; Metge and El Alaoui 1987, AP, Benslimane; Himmi et al. 1995; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Rusticoidus) rusticus (Rossi, 1790)

Viamonte and Ramirez 1946, Rif, Tétouan, AP, Larache; Gaud 1953a, MA, Taza; Himmi et al. 1995; Handaq 1998, HA, Bengrir; Trari et al. 2002; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Stegomyia) aegypti (Linnaeus in Hasselquist, 1762)

d’Anfreville 1916, AP, Salé; Vialatte 1923, AP, Rabat, Casablanca, HA, Marrakech; Charrier 1924, Rif, Tanger; Gaud 1953a, AP, Salé, HA, Marrakech; Himmi et al. 1995; Dakki 1997; Handaq 1998, HA, Bengrir; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Stegomyia) albopictus (Skuse, 1895)

Bennouna et al. 2016, AP, Agdal (Rabat); Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Faraj et al. 2018; Amraoui et al. 2019

Culicini

Culex Linnaeus, 1758

Culex (Barraudius) modestus Ficalbi, 1889

Séguy 1930a; Bailly-Choumara 1968a, AP, Larache; Trari 1991, AP, Gharb; Himmi et al. 1995; Dakki 1997; Handaq 1998, HA, Bengrir; Trari et al. 2002; Himmi 2007, Rif, Bab Berred, AP, Maâmora; Hadji et al. 2013, AP, Sidi yahia du Gharb, Kcebia, Sidi Hagouch (Sidi Slimane); Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) brumpti Galliard, 1931

Bailly-Choumara 1968a, AP, Merja Bokka, Larache, HA, Marrakech; Bailly-Choumara 1972a, AP, Merja Sheishat; Himmi et al. 1995; Dakki 1997; Himmi 2007; Trari et al. 2002; El Ouali Lalami et al. 2010a, MA, Fès; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) laticintus Edwards, 1913

Charrier 1924, Rif, Tanger; Callot 1940, AA, Goulmima (mares); Gaud 1953a, HA, Marrakech, AA, Agadir; Gaud 1957a, EM, Nador; Bailly-Choumara 1965c, EM, Oued Isker, Aïn Aït delouine, Talmesdourt, AA, Ouled Teima, ounaamane, Bou Izakarn, Agunil Khnufa, Akka-guiren; Bailly-Choumara 1966, AA, Akka-Iguiren, Tirherm, Taoujgelt, Aït Ouabelli, Anamere-Smougue, Aït melloul, Tiznit (Rocade de Draa); Bailly-Choumara 1967b, Rif, Béni Bouayache, Targuist et environs, EM, Grotte du Zegzel; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Himmi 2007, AP, Skhirat; Faraj et al. 2008b, AP, Louamra; Hadji et al. 2013, AP, Sidi Hagouch (Sidi Slimane); Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) mimeticus Noè, 1899

Séguy 1930a; Viamonte and Ramirez 1946, Rif, Béni Ider, Fnideq, Khemis Anjra, Ketama, Oued Amsa, Oued Krikra, Oued Martil, Oued Laou; Gaud 1953a, Rif, Ouezzane, Ghafsai, EM, Berkane, Martinpray (près Berkane), El Aïoun Sidi Mellouk, AP, Tamri, MA, Meknès, Fès, Ifrane, Taza, Béni Mellal, HA, Midelt, Marrakech, Azilal, AA, Tinghir, Tichka; Guy et al. 1958, HA, Oued N’fis; Bailly-Choumara 1965c, AA, Oued Noun, Anezi, Tafraoute; Bailly-Choumara 1966, AA, Agadir; Bailly-Choumara 1967a, MA, route Azrou-Khénifra, piste Tafechna-Taoujgelt, Source Oumerrbia, Ghorm El Alem, piste Ksiba-Naour, piste Naour-Arbala, Zaouia Cheikh, Oued Sarif; Bailly-Choumara 1967b, EM, 7 km N d’Itzer, 8 km d’Itzer; Bailly-Choumara 1967c, Rif, route Chaouen-Bab Taza, Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Taza-Asifane, piste Bab Berred-Tamorote, Ketama, piste Ketama-Jebel Tidighine, route Ketama-Targuist, route nationale Jebha, Al Hoceima, Béni Bouayache, Marchica, Targuist et environs, Jebha, Boured; Trari 1991, AP, Sidi Yahia du Gharb; Himmi et al. 1995; Louah 1995, Rif, Riffien, Tres piedras, Marina Smir, Bouzaghlal, M’diq, Oued Maleh, Azla, Tahaddart, Moulay Bouchta, Schroda, Kebbache, Talambote, Oued Maggou; Louah et al. 1995; Chlaida 1997, AP, Oued Sidi Messoud, Aïn Blal, Douar Chlihat, Barrage Al Massira; Chlaida and Bouzidi 1995, AP, Barrage El Massira; Dakki 1997; Ramdani 1997, AP, Skhirat, Tamaris Merja; Handaq 1998, HA, Oukaimeden, Amizmiz, Tiguenziouine (près Oued N’fis); Trari et al. 2002; Himmi 2007, Rif, Bab Berred, AP, Bouknadel, Douar jdid (Skhirat); El Ouali Lalami et al. 2010a, MA, Fès; Larhbali et al. 2010, MA, Oulmès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) perexiguus Theobald, 1903

Séguy 1930a; Callot 1940, AA, Assa; Senevet and Andarelli 1959a, EM, Oujda, Taourirt, AP, Aïn el Aouda, Arbaoua, Had Kourt, Oued Beht, Rabat, Allal Tazi, Oued Sahli, Zaouia Ech cheikh, Taghzirt, MA, Béni Mellal, Foum Zabel, Ifrane, Meknès, Aït Atta du Rteb, Fès, Sidi Mokhfi, Tahala, HA, Tazert, Midelt, AA, akka, Tamri; Bailly-Choumara 1965c, EM, Assa; Bailly-Choumara 1966, AA, Souk El Khémis Dades, Aït Ouabelli, Tarhjicht, Tirherm, Taoujgelt, Aït melloul; Bailly-Choumara 1967a, MA, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, Ajdir; Bailly-Choumara 1967b, EM, Madagh, Merja Boubker, Aïn Béni Mathar; Bailly-Choumara 1967c, Rif, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, Anasar, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, Béni Bouayache, Targusit et environs; Bailly-Choumara 1972a, AP, Merja Sheishat; Louah 1995, Rif, Riffien, Tres piedras, Marina Smir, Bouzeghlal, Oued Maleh, Tahaddart, Talembote, Schroda, Tanger; Louah et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi, El Oulja, Fouarate; Ramdani 1997, AP, Skhirat, Tamaris; Handaq 1998, HA, Marrakech, entre Oued N’fis et Chichaoua, Kelaâ Sraghna; Alaoui Slimani 2002, AP, Rabat, Salé; Trari et al. 2002; Himmi 2007, Rif, Bab Berred, Bab Taza; Faraj et al. 2008c, AP, Larache Louamra; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Culex (Culex) pipiens Linnaeus, 1758

d’Anfreville 1916, AP, Salé; Charrier 1924, Rif, Tanger; Séguy 1930a; Callot 1940, SA, Goulimine; Viamonte and Ramirez 1946, Rif, Boudinar, Dar Benkarrich, Tétouan, Tanger, Asilah, Ksar El Kébir, Chefchaouen, Ketama, Nador; Gaud 1952, AP, Gharb; Séguy 1953a, SA, Tindouf; Guy 1958, HA, Oued N’fis; Guy et al. 1958, HA, Oued N’fis; Bailly-Choumara 1965c, EM, Aouinet Aït Oussa, Aïn Isker, Aïn Aït Delouine, Oued Mesdourt, Talmesdourt, Toudi, AA, Aït Onmar, Oulad Teima, Taroudant, Tiznit ville, Talaint, Hassi Tafnidilt, Zaouiat Cheikh, Aïn Guerzim, Tafraout ville, SA, Goulimine ville, Vallée de l’Oued Assaka, Ouaroun, Zriouila, Labyar, Tighmert, Abeino, Tantan ville, Zag; Bailly-Choumara 1966, AA, piste d’Akka au Draa, Akka et environs, Aït Ouabelli, Anamere-Smougue, Tarhjicht, Aït Melloul, Tiznit et environs; Bailly-Choumara 1967a, MA, route Azrou-Khénifra, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, maison forestière Ouiouane, piste Tafechna-Senoual-Itzer, Ajdir, Itzer, piste Itzer-Boumia, Ghorm El Alem, Ghorm El Alem-El Ksiba, El Ksiba, piste Ksiba-Naour, Zaouia Cheikh, Oued Sarif, Dayet Aoua, Pont Tarmilate, Ifrane, Imilchil; Bailly-Choumara 1967b, EM, route Itzer-Midelt, Gaada de Debdou, Guercif ville, Cascade Oued Za, Grotte du Zegzel, Environs Saidia, Douar Aïn Shebbak, Douar Aïn Zabia, Douar Mardarh, Douar Sidi Hashas, Saidia, Merja Boubker, Berguent, Tendrara, Figuig; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, Anasar, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, Al Hoceima, Al Hoceima Club Med, Béni Bouayache, Marchica, Targuist et environs, Jebha, Ghafsai; Bailly-Choumara 1972a, AP, Merja Sheishat; Bailly-Choumara 1973b, AP, Merja Bokka, Merja Qodiya; Metge and Belakoul 1989, AP, Benslimane, Sidi Bettache; Himmi 1991, AP, Sidi Boughaba, Sidi Amira; Trari 1991, AP, Maâmora, El Menzeh, Sidi Boughaba, Chkaïfien, Sidi Yahia du Gharb, Bokka, Merja Zerga, Sidi Allal Tazi, Oued Loukous, Aïn Chouk, Merja Bargha, Merja Oulad Skhar; Bouallam and Ramdani 1992, HA, Marrakech; El Bermaki 1993, AP, Sidi Maârouf; Himmi et al. 1995; Louah 1995, Rif, Fnideq, Riffien, Tres piedras, Marina Smir, M’diq, Cabo Negro, Zekri, Azla, Tahaddart, Schroda, Kebbache, Ouadras, Punta cirres, Sidi Kankoch, Oued Kbir; Louah et al. 1995; Bouallam et al. 1997b, HA, Marrakech; Dakki 1997; Mestari 1997, AP, Mohammedia; Himmi et al. 1998, AP, Sidi Boughaba, Puits de Sidi Amira (forest of Maâmora); Moussalim 1997, AP, El Oulja, Maâmora, Fouarate, Sidi Boughaba; Ramdani 1997, AP, meseta côtière (Témara-Casablanca); Faraj et al. 1997, AP, Kénitra; Handaq 1998, AP, Zemamra, MA, Oued Zem, HA, Marrakech, Zaouiet Ben Sassi, Sidi Bou Othmane, Kelaâ Sraghna, Bengrir; Bouallam 2001, HA, Marrakech; Alaoui Slimani 2002, AP, Rabat; Trari et al. 2002; Aouinty et al. 2006, AP, Mohammedia; Faraj et al. 2006, AP, Salé; Himmi 2007, Rif, Bab Berred, Bab Taza, Stehat, AP, Skhirate, forest of Maâmora, forest of Hilton, Sidi Boughaba, El Oulja, Ouled Salem, Ouled dlim, Douar Ould Yahia Ben Ali, Larouaza, Douar Elarja, Douar Jdid, Douar Jnaja; Faraj et al. 2008b, AP, Louamra; El Ouali Lalami et al. 2010a, MA, Fès; Larhbali et al. 2010, MA, Oulmès, Tarmilate, Bni Ounzar, Ganzra;Louali Lalami et al. 2010b, MA, Oued El Himmer; Amraoui 2012, HA, Marrakech; Amraoui et al. 2012, Rif, Tanger, AP, Mohammedia, Casablanca, HA, Marrakech; Amraoui et al. 2012, AP, Mohammedia, Casablanca; El Ouali lalami 2012, MA, route de Sidi Harazem; Hadji et al. 2013, AP, Sidi Slimane; El Joubari et al. 2014, Rif, Smir lagoon; El Joubari et al. 2015a, Rif, Smir lagoon; Marc et al. 2016, AP, Kénitra; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Bkhache et al. 2018, Rif, Tanger, AP, Rabat, Mohammedia, HA, Marrakech; Tmimi et al. 2018, AP, Mohammedia; Mouatassem et al. 2019, MA, Fès

Culex (Culex) simpsoni Theobald, 1905

Callot 1940, AA, Taghicht; Senevet et al. 1949, AA, Oued Noun, Tafnidelt, Akka; Gaud 1953a, AA, Imsouane, Aït Melloul, Akka, O’Noun, Taghjicht, Assa, Tafnidilt; Bailly-Choumara 1965c, EM, Aman d’Aït Oussa, El Megrinat, AA, Oued Izi, Oued Massa-Pont de la route Agadir-Tiznit, Guelta Zerga, Tafnidilt, SA, Poste militaire de Boujrif, Tirhmert, Taourirt-Barrage, Aouinet Torkoz; Bailly-Choumara 1966, AA, Tirherm, Taoujgelt, Taghjicht, Aït melloul, Tiznit; Chlaida and Bouzidi 1995, AP, Sidi M’barek, Oued Sidi Messoud, Mechrâa, Sidi Boulâarais, Douar Chlihat (south of Settat); Chlaida 1997, AP, Sidi M’barek, Mechrâa Settir, Sidi Boulâarais, Douar Chlihat (south of Settat); Dakki 1997; Handaq 1998, HA, Sidi Bou Othmane (Marrakech); Trari et al. 2002; Himmi 2007, Rif, Bab Berred, MA, Khémisset, AA, sud Anti Atlas; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) theileri Theobald, 1903

Callot 1940, AA, Taghjicht; Viamonte and Ramirez 1946, Rif, Dar Benkarrich, Boudinar, Ketama, Tanger, Tétouan, Chefchaouen, AP, Larache; Guy 1958, HA, Oued N’fis; Guy et al. 1958, HA, Oued N’fis; Bailly-Choumara 1965c, EM, Aouinet Aït Oussa, Aïn Oumesdour, Aouinet Torkoz, SA, Tirh Mzoun; Bailly-Choumara 1966, AA, Souk El Khémis Dades, Akka et environs, Aït Melloul, Tazenakhte, piste Tiznit-Tafraout, SA, Goulimine; Bailly-Choumara 1967a, MA, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, Aguelmane Azigza, piste Aguelmane Azigza-Aïn Leuh, maison forestière Ouiouane, Khénifra, piste Tafechna-Senoual-Itzer, Ajdir, Itzer, Col du Zad, piste Itzer-Boumia, Ghorm El Alem, piste Ghorm El Alem-El Ksiba, El Ksiba, piste Ksiba-Naour, Aguelmane Moulay Yakoub, Zaouia Cheich, Kafensour, Dayet Aoua, Pont Tarmilate; Bailly-Choumara 1967b, EM, route Itzer-Midelt, 40 km N Outat El Haj, Taourirt, Driouch, Figuig; Bailly-Choumara 1967c, Rif, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, route Bab Taza-Bab Berred, Anasar, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, ArbaaTaourirt, Al Hoceima, Targuist et environs, Ghafsai; Bailly-Choumara 1972a, AP, Merja Sheishat; Bailly-Choumara 1973b, AP, Merja Sheishat, Merja Bokka; Himmi 1991, AP, Sidi Boughaba, El menzeh, Sidi Amira; Trari 1991, AP, Maâmora, Oued Sebou, Bordure Oued Loukous, Sidi Yahia du Gharb, Bokka, Entre Moulay Bousselham et Larache, Merja Zerga, Larache; El Bermaki 1993, AP, Sidi Maârouf, El Oulfa; Louah 1995, Rif, Bouzeghlal, M’Diq, Oued Maleh, Zekri, Lajour, Tahaddart, Loubart, Chefchaouen, Tanger; Himmi et al. 1995; Louah et al. 1995; Dakki 1997; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi, Maâmora, Rabat, Kénitra; Ramdani 1997, AP, Skhirat, Tamaris; Handaq 1998, AP, Zemamra, MA, Béni Mellal, HA, Oukaimeden, Amizmiz, Tiguenziouine, Marrakech, Sidi Bou Othmane, Seguarta-Kelaâ, Had Mhara, Aîn Äounate, Bengrir; Himmi et al. 1998, AP, Sidi Boughaba, Dayat d’El Menzeh (north east of Kénitra), Gharb; Alaoui Slimani 2002, AP, Rabat; Trari et al. 2002; Himmi 2007, Rif, Bab Berred, Tanakoub, AP, Skhirate, forest of Maâmora, Chiahna, Sidi Amira, Sidi Boughaba, Bouknadel, Sidi Azzouz, Ehssaïne, Bettana, Sidi Yahia, Sebbah, Larouaza, Douar Elarja, Douar Jdid, Douar Jnaja; Faraj et al. 2008b, AP, Larache Louamra; El Ouali Lalami et al. 2010a, AP, Boucharen;Larhbali et al. 2010, MA, Roumani, Aïn Sbite, Ezzhiliga, Oulmès, Tarmilate, M’rirt, Bni Ounzar, Ganzra (Khémisset); Hadji et al. 2013, AP, Sidi yahia du Gharb, Kcebia, Sidi Hagouch, Dar Belamri (Sidi Slimane); El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Culex (Maillotia) deserticola Kirkpatrick, 1925

Gaud 1947c, AA, Tansikht; Gaud 1953a, EM, Figuig, Berguent, Anoual, Boudnib, Aïn Chair, Aoufous, Tarhit, HA, Tazenakht, Tichka, Tizi-n’Telghemt, AA, Ouarzazate, Zagora; Bailly-Choumara 1965c, EM, Oued Isker, El Megrinat, Taskala, Aïn Aït Delouine, Talmesdourt, Aouinet Torkoz, Bouanama, Rich Tamlougout, Assa, AA, Tafraout ville, Oued Jemâa Idaousmaal, Aït abdallah, Issedrim Igmur Igues, SA, Vallée de l’Oued Assaka, Tacharicht, Bou Izakarn, Aïn Erkha; Chlaida and Bouzidi 1995, AP, Sud de Settat; Himmi et al. 1995; Chlaida 1997, AP, Barrage Al Massira; Dakki 1997; Trari et al. 2002; Himmi 2007; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Maillotia) hortensis Ficalbi, 1889

Charrier 1924, Rif, Tanger; Séguy 1930a; Langeron 1938, HA, Tounfite; Callot 1940, HA, Anefgou (2500 m), Tirghist (2500 m), Tighermine (2500 m); Gaud 1953a, HA, Tizi-n’Telghemt, Tizi-n’Tichka; Bailly-Choumara 1967a, MA, piste tafechna-Znan Imes, Khénifra, piste Tafechna-Senoual-Itzer, Ajdir, Itzer, Ghorm El Alem, El Ksiba, piste Ksiba-Naour, Piste Naour-Arbala, piste Arbala-El Kbab, Zouia Cheikh, Oued Sarif, Ifrane, Boulmane, Imilchil; Bailly-Choumara 1967b, EM, Tafraout, Grotte du Zegzel; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Berred-Tamorote, route Bab Berred-Ketama, Ketama, piste Ketama-Mt Tiguidin, route nationale Jebha, route Targuist-Béni Boufrah,Trari 1991, AP, Gharb; El Bermaki 1993, AP, Casablanca; Himmi et al. 1995; Louah 1995, Rif, Riffien, Marina Smir, M’diq, Oued Maleh, Lajour, Tahaddart, Schroda, Talembote, Bab Berred, Punta Cirres, Sidi Kankoch, Oued kbir, Oued Jebel Lehbib; Louah et al. 1995; Chlaida and Bouzidi 1995, AP, Barrage Al Massira; Chlaida 1997, AP, Barrage Al Massira; Dakki 1997; Ramdani 1997, AP, Skhirat, Tamaris; Handaq 1998, HA, Amizmiz, Sidi Bou Othmane, Had Mhara, Bengrir; Trari et al. 2002; Himmi 2007, Rif, Bab Berred, Bab Taza, AP, Skhirate; El Ouali Lalami et al. 2010b, MA, Oued Fès, Oued El Himmer, Camping Sidi Harazem; Larhbali et al. 2010, MA, Merchouch, Ghoualem, Ezzhiliga, Oulmès, Tarmilate, M’rirt, Bni Ounzar, Ganzra (Khémisset); El Ouali Lalami 2012, MA, Oued Fès; Oued El Himmer, route de Sidi Harazem, Camping Sidi Harazem; Hadji et al. 2013, AP, Sidi Yahia du Gharb, Kcebia, Sidi Hagouch, Dar Belamri, Lalla Itto, Soualem (Sidi Slimane); Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Culex (Neoculex) impudicus Ficalbi, 1890

Charrier 1924, Rif, Tanger; Séguy 1930a; Bailly-Choumara 1966, AA, Taliouine; Bailly-Choumara 1967a, MA, route Azrou-Khénifra, piste Tafechna-Senoual-Itzer, Itzer, piste Naour-Arbala, Zaouia Cheikh, Pont Tarmilate; Bailly-Choumara 1967b, EM, route Itzer-Midelt, Cascade Oued Za, Grotte du Zegzel; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Berred-Asifane, piste Bab Berred-Tamorote, Ketama, route nationale Jebha, Al Hoceima Club Med, Targuist et environs; Trari 1991, AP, Sidi Amira, El Menzeh, Oued Sebou, Sidi Yahia du Gharb, Moulay Bousselham, Larache, Bordure Oued Loukous; Dakki 1997; Himmi et al. 1998, AP, Sidi Boughaba, Dayat d’El Menzeh (north east of Kénitra), Puits de Sidi Amira (forest of Maâmora); Trari et al. 2002; Himmi 1991, AP, Sidi Boughaba, El Menzeh, Sidi Amira; Himmi et al. 1995; Louah 1995, Rif, Haidra, Marina Smir, Tahaddart, Tanger; Louah et al. 1995; Ramdani 1997, AP, Tamaris; Alaoui Slimani 2002, AP, Rabat; Himmi 2007, Rif, Bab Berred, Bab Taza, AP, forest Maâmora; El Ouali Lalami et al. 2010a, MA, Fès; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Neoculex) martinii Medschid, 1930

Bailly-Choumara 1968a, Rif, Al Hoceima, AP, Sidi Yahia du Gharb, Sidi Allal Tazi, Larache, Rabat, HA, Marrakech, AA, Tiznit; Himmi et al. 1995; Louah 1995, Rif, Haidra, Marina Smir, Cabo Negro, Oued Maleh; Louah et al. 1995; Dakki 1997; Trari et al. 2002; Himmi 2007; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culisetini

Culiseta Felt, 1904

Culiseta (Allotheobaldia) longiareolata (Macquart, 1838)

d’Anfreville 1916, AP, Salé; Séguy 1930a; Gaud 1947c, AA, Tansikht; Gaud 1953a, HA, Marrakech; Bailly-Choumara 1965c, EM, El Aïoun du Draa, Aïn Aït Delouine, Talmesdourt, Aouinet Torkoz, Rich Tamlougout, AA, Aït Onmar, Ouled Teima, Tiznit ville, Bounaamane, Id Baha, Tafnidilt, Guelta Zerga, Aïn Kerma, Tafraout ville, Oued Jemâa Idaousmaal, Toudi, Aït Abdallah, Igherm, Issedrim Igmur Igues, SA, Goulimine ville, Poste militaire de Boujrif, Embouchure de l’Oued Assaka, Ouaroun, Labyar, Asrir, Tacharicht, Bou Izakarn ville, Jemâa N’Tirhirte, Aït Erkha, Tantan ville; Bailly-Choumara 1966, AA, Aït melloul, Tiznit; Bailly-Choumara 1967a, MA, Jnane Imasse, Khénifra, piste Tafechna-Senoual-Itzer, Itzer, Ghorm El Alem, El ksiba, piste Naour-Arbala, Ifrane, Pont Tarmilate, Imouzzer Marmoucha; Bailly-Choumara 1967b, EM, Tafraout, Guercif ville, Grotte du Zegzel, Berguent, Tendrara; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, Bab Taza, route Bab Taza-Bab Berred, route Ketama-Targuist, route nationale Jebha, Targuist et environs, Aïn Hamra; Bailly-Choumara 1968a, AP, Meja Bokka, AA, Tiznit; Himmi 1991, AP, Sidi Boughaba, El Menzeh; Trari 1991, AP, El Menzeh, Gharb; El Bermaki 1993, AP, Sidi Maârouf; Himmi et al. 1995; Louah 1995, Rif, Fnideq, Riffien, Marina Smir, M’diq, Cabo Negro, Lajour, Ouadras, Punta Cirres, Ksar Sghir, Tanger; Louah et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Maâmora, Fouarate; Ramdani 1997, AP, Témara, Skhirat, Tamaris; Dakki 1997; Handaq 1998, HA, Marrakech, Kelaâ Sraghna, Bengrir Himmi et al. 1998, AP, Sidi Boughaba, Dayat d’El Menzeh (north east of Kénitra); Bouallam 2001, HA, Bordure Oued N’fis; Alaoui Slimani 2002, AP, Rabat; Trari et al. 2002; Aouinty et al. 2006, AP, Mohammedia; Himmi 2007, Rif, Bab Berred, AP, Skhirate, Maâmora, forest of Hilton; Koçak and Kemal 2013; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Culiseta (Culicella) fumipennis (Stephens, 1825)

Gaud 1947c, AP, Rabat, Sidi Allal Tazi, MA, Khémisset; Senevet and Andarelli 1959, AP, Rabat, Casablanca, Bouznika, MA, Fès; Bailly-Choumara 1967a, MA, Jnane Imasse; Bailly-Choumara 1967c, Rif, route Bab Taza-Bab Berred; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culiseta (Culicella) litorea (Shute, 1928)

Metge 1986, AP, Casablanca; Dakki 1997; Carles-Tolrá 2002; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culiseta (Culiseta) annulata (Schrank, 1776)

d’Anfreville 1916, AP, Salé; Charrier 1924, Rif, Tanger; Séguy 1930a, Rif, Tanger; Viamonte and Ramirez 1946, Rif, Ben Karrich, Asilah, Ketama, Malaliene, Tétouan, AP, Larache; Gaud 1952, AP, Souk El Hadd (Gharb); Gaud 1953a, Rif, Tanger, Tétouan, EM, Saidia, Talsint, AP, Casablanca, Rabat, Kénitra, MA, Meknès, Fès, Ifrane, Taza, HA, Marrakech, Aït Bouguemez, Midelt; Gaud 1957b, Rif, Tétouan, Tanger, AP, Sidi Allal Tazi, Rabat, Casablanca, MA, Meknès, Ifrane; Bailly-Choumara 1967a, MA, Jnane Imasse, maison forestière Ouiouane, piste Tafechna-Senoual-Itzer, Ghorm El Alem, Zaouia Cheikh, Pont Tarmilate, Ifrane; Bailly-Choumara 1967b, EM, Tafraout, route Itzer-Midelt; Bailly-Choumara 1967c, Rif, piste Bab Taza-Talassemtane; Bailly-Choumara 1972a, AP, Merja Sheishat; Himmi et al. 1995; Louah 1995, Rif, Riffien, M’diq, Tanger; Louah et al. 1995; Chlaida 1997, AP, Barrage Al Massira; Dakki 1997; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Rabat; Ramdani 1997, AP, meseta côtière (Casablanca-Rabat); Himmi et al. 1998, AP, Sidi Boughaba; Himmi 1991, AP, Sidi Boughaba, El Menzeh; Trari 1991, AP, Sidi Boughaba, El Menzeh; Trari et al. 2002; Himmi 2007, AP, Skhirate, Maâmora, forest of Hilton, Oulja; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culiseta (Culiseta) subochrea (Edwards, 1921)

Gaud 1952, AP, Gharb; Gaud 1957b, Rif, Tanger, Chefchaouen, EM, Oujda, AP, Sidi Allal Tazi, Casablanca; Bailly-Choumara 1967a, MA, Imilchil; Bailly-Choumara 1967c, Rif, Ketama; Bailly-Choumara 1972a, AP, Merja Sheishat; Himmi 1991, AP, Sidi Boughaba; Trari 1991, AP, Sidi Boughaba; Louah 1995, Rif, Riffien, Tres piedras, M’diq, Cabo Negro, Oued Maleh, Tanger; Louah et al. 1995; Dakki 1997; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Kénitra; Ramdani 1997, AP, Témara, Skhirat, Casablanca; Handaq 1998, AP, El Jadida, HA, Bengrir; Himmi et al. 1998, AP, Sidi Boughaba; Trari et al. 2002; Himmi 2007, AP, Skhirat, Dayet Aïn Chems, Maâmora, forest of Hilton; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Mansoniini

Coquillettidia Dyar, 1905

Coquillettidia (Coquillettidia) buxtoni (Edwards, 1923)

Bailly-Choumara 1965a, AP, Merja Bokka; Bailly-Choumara 1970, AP, Merja Bokka; Bailly-Choumara 1972a, AP, Merja Sheishat (Larache); Bailly-Choumara 1973b, AP, Merja Bokka, Larache; Himmi et al. 1995; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Coquillettidia (Coquillettidia) richiardii (Ficalbi, 1889)

Bailly-Choumara 1965a, AP, Merja Bokka; Bailly-Choumara 1967a, MA, Zaouia Cheikh; Bailly-Choumara 1970, AP, Merja Bokka; Bailly-Choumara 1972a, AP, Merja Sheishat (Larache); Bailly-Choumara 1973b, AP, Merja Bokka; Trari 1991, AP, Sidi Yahia du Gharb, Aïn Chouk, Merja Bargha; Himmi et al. 1995; Dakki 1997; Moussalim 1997, AP, Fouarate sur les bordures de la Merja; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Orthopodomyiini

Orthopodomyia Theobald, 1904

Orthopodomyia pulcripalpis (Rondani, 1872)

Bailly-Choumara 1965b, AP, Maâmora; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Uranotaeniini

Uranotaenia Lynch Arribálzaga, 1891

Uranotaenia (Pseudoficalbia) unguiculata Edwards, 1913

Séguy 1930a; Gaud 1953a, HA, Marrakech; Senevet and Andarelli 1959a, EM, Oujda, Berguent, Guercif, AP, Sidi Allal Tazi, Bouznika, Casablanca, Béni Moussa, MA, Oulad Massine, Meknès, Moulay Yacoub, Béni Mellal, AA, Ksar Mzizel, Aït Melloul; Bailly-Choumara 1965c, EM, Aïn Aït Delouine, Talmesdourt, Rich Tamlougout; Bailly-Choumara 1966, AA, Aït Ouabelli; Bailly-Choumara 1967a, MA, Pont Tarmilate; Bailly-Choumara 1967b, EM, environs de Taourirt, Madagh, Merja Boubker; Bailly-Choumara 1967c, Rif, route Bab Taza-Bab Berred, piste Ketama-Jebel Tidighine; Bailly-Choumara 1968a, AP, Oued Loukous, Merja Bokka, Kénitra, Oued Bou-Regreg; Himmi 1991, AP, Sidi Boughaba; Trari 1991, AP, Oued Sebou; El Bermaki 1993, AP, El Oulfa (Casablanca); Himmi et al. 1995; Dakki 1997; Ramdani 1997, AP, Tamaris; Handaq 1998, AP, Sidi Bennour, HA, Zima Chemaîa (Bengrir); Trari et al. 2002; Himmi 2007, AP, Sidi Boughaba; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Uranotaenia (Uranotaenia) balfouri Theobald, 1904

Bailly-Choumara 1968a, AP, Merja Bokka, Oued Loukous, Sidi Yahia, Oued Sebou (Kénitra), Oued Bou-Regreg; Himmi et al. 1995; Dakki 1997; Moussalim 1997, AP, Sidi Allal Tazi, Fouarate, Sidi Boughaba; Trari et al. 2002; Himmi 2007, AP, Sidi Boughaba; El Ouali Lalami et al. 2010a, MA, Fès Boulmane; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

DIXIDAE

K. Kettani, R. Wagner

Number of species: 12. Expected: 15

Faunistic knowledge of the family in Morocco: moderate

Dixa Meigen, 1818

Dixa caudatula Séguy, 1928

Séguy 1930a, HA, Arround, Skoutana (2400 m); Vaillant 1959; Dakki 1997; Mouna 1998

Dixa dilatata Strobl, 1900

= Dixa riparia Vaillant, in Vaillant 1959: 180

Vaillant 1959, HA, Source de M’Goum (2500 m), Gorges d’Imi-N’Ifri (1050 m); Vaillant 1965; Dakki 1997

Dixa maculata Meigen, 1818

Séguy 1930a; Dakki 1997; Mouna 1998

Dixa mera Séguy, 1930

Séguy 1930a, MA, forest of Timelilt (1900 m); Vaillant 1959; Dakki 1997; Mouna 1998

Dixa nebulosa Meigen, 1830

Séguy 1930a, HA; Dakki 1997; Mouna 1998

Dixa perexilis Séguy, 1928

Séguy 1930a, HA, riverside of Oued Imminen (Tachdirt, 2400 m); Vaillant 1959; Dakki 1997; Mouna 1998

Dixa puberula Loew, 1849

Vaillant 1959, HA, headwaters of Asif M’Goum (2500 m); Vaillant 1965; Dakki 1997; Mouna 1998

Dixa submaculata Edwards, 1920

Séguy 1930a, MA, Sidi Yahia, Talzent (1800 m); Dakki 1997; Mouna 1998

Dixella Dyar & Shannon, 1924

Dixella aestivalis (Meigen, 1818)

Séguy 1930a, AP, Merja Boughaba; Vaillant 1965; Ramdani 1981; Ramdani and Tourenq 1982; Mouna 1998

Dixella attica (Pandazis, 1933)

= Dixella numidica (Sicart, 1955)

Ebejer et al. 2019, Rif, Cabo Negro (indoors: 10 m)

Dixella martinii (Peus, 1934)

Ebejer et al. 2019, Rif, Moulay Abdelsalam (965 m)

Dixella serotina (Meigen, 1818)

= Dixa serotina Wied, in Mouna 1998: 85

Séguy 1930a, AP, Casablanca (between Kénitra and Oued Beth); Dakki 1997; Mouna 1998

Chironomoidea

CERATOPOGONIDAE

K. Kettani, B. Mathieu

Number of species: 62. Expected: 80

Faunistic knowledge of the family in Morocco: moderate

Ceratopogoninae

Culicoidini

Culicoides Latreille, 1809

Culicoides (Avaritia) imicola Kieffer, 1913

Kremer et al. 1971, MA, Fès-Meknès, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, SA, Guelmim-Oued Noun; Chaker et al. 1980, MA, Fès, Rhafsai, AA, Torkoz, Tarhjisht; Remm 1988a; Dakki 1997; Bouayoune et al. 1998, Rif, Tanger-Tétouan-Al Hoceima, EM, Oriental, AP, Rabat-Salé-Kénitra, Safi, MA, Fès-Meknès, Béni Mellal-Khénifra, HA, Marrakech, AA, Draa-Tafilalet, Souss-Massa, SA, Guelmim-Oued Noun; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Lhor et al. 2015, Rif, Sahel Chamali, MA, Sidi Hammadi, Benioukil, Aïn Leuh, Ait Siberne, Meknès, AA, Errachidia, Sidi Dahmane, SA, Foum El Oued; Bourquia et al. 2019, AP, Rabat

Culicoides (Avaritia) montanus Shakirzjanova, 1962

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Safi, HA, Marrakech; Chaker et al. 1980, Rif, Al Hoceima, AP, Oued Cherrat, HA, Souk Tnine de Oudaias (Haouz), Marrakech, AA, Torkoz; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Avaritia) obsoletus (Meigen, 1818)

Callot et al. 1968, Rif, Al Hoceima, AP, Merja Bokka, Sidi Yahia du Gharb, Sidi-Bettache (Zaeir), Rabat-Salé-Kénitra, HA, El Harcha (plateau central); Bailly-Choumara and Kremer 1970, AP, estuaire de Bou-Regreg; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Safi, MA, Béni Mellal-Khénifra, HA, Marrakech; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, HA, Marrakech; Chaker et al. 1980, Rif, Al Hoceima, AP, Zaers, Sidi Bettache, HA, El Harcha, Talet Inaouane (Haouz); Remm 1988a; Dakki 1997; Bouayoune et al. 1998, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, MA, Fès-Meknès, Béni Mellal-Khénifra, HA, Marrakech, AA, Souss-Massa, SA, Guelmim-Oued Noun; Cêtre-Sossah and Baldet 2004; Lhor et al. 2015, Rif, Sahel Chamali, MA, Ait Siberne; Bourquia et al. 2019, AP, Rabat

Culicoides (Avaritia) scoticus Downes & Kettle, 1952

Kremer et al. 1971, AP, Safi, MA, Béni Mellal-Khénifra, HA, El Harcha, Talet Inaouane (Haouz), Marrakech; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Casablanca-Settat, Safi, HA, Marrakech; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Beltranmyia) circumscriptus Kieffer, 1918

Callot et al. 1968; Bailly-Choumara and Kremer 1970, Rif, Smir lagoon, Oued Negro, EM, Merja Boubker (Berkane), Gouttitir (NE Guercif), AP, Merja Sheishat (Larache), Aïn Muelha (near Oued Sidi Allal Tazi, estuaire Oued Bou-Regreg, Dayat Qoudiya (Sidi Yahia Gharb); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, MA, Fès-Meknès, HA, Marrakech, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, MA, Fès-Meknès, AA, Souss Massa; Chaker et al. 1980, Rif, Al Hoceima, AP, Merja Qoudiya, Merja Bokka, Sidi Yahia du Gharb, MA, Aïn Karma (Saiss), Oulmès, HA, Setti Fatma, AA, Aït Melloul (Souss); Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Lhor et al. 2015, Rif, Sahel Chamali, MA, Sidi Hammadi, Benioukil, Meknès, AA, Errachidia, SA, Foum El Oued; Bourquia et al. 2019, AP, Rabat

Culicoides (Culicoides) fagineus Edwards, in Edwards et al. 1939

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Safi, MA, Béni Mellal-Khénifra, HA, Marrakech; Kremer et al. 1975; Chaker et al. 1979, AP, Rabat-Salé-Kénitra; Kremer et al. 1979; Chaker et al. 1980, AP, Rabat, Sidi Bettache, MA, Khemisset, HA, Marrakech; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Culicoides) newsteadi Austen, 1921

= Culicoides (Culicoides) halophilus Kieffer, in Callot et al. 1968: 886, Bailly-Choumara and Kremer 1970: 386, Dakki 1997: 60

Callot et al. 1968, Rif, Cabo Negro (Ferma), Tétouan, Talerhza, Tanger-Tétouan-Al Hoceima, AP, Larache, Merja Bokka (Gharb), Aïn Chok, HA, Talet-Inaouan (Haouz); Bailly-Choumara and Kremer 1970, Rif, Smir lagoon, Oued Negro, AP, Merja Sheishat (Larache), Aïn Muelha (near Oued Sidi Allal Tazi, estuaire Oued Bou-Regreg, Dayat Qoudiya (Sidi Yahia du Gharb), EM, Merja Boubker (Berkane), Ksabi (NE Midelt), HA, Souk Tnine des Oudaias (bordure Oued N’fis), AA, Aïn Sefra (south Foum Zquid); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Casablanca, Settat, MA, Fès-Meknès, Béni Mellal-Khénifra, SA, Guelmim-Oued Noun; Kremer et al. 1975; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Safi, Casablanca, Settat, MA, Béni Mellal- Khénifra, HA, Marrakech; Kremer et al. 1979; Baylis et al. 1997, Rif, Tanger, HA, Marrakech; Remm 1988a; Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Lhor et al. 2015, MA, Aïn Leuh, Ait Siberne, Meknès, SA, Foum El Oued; Bourquia et al. 2019, AP, Rabat

Culicoides (Culicoides) pulicaris (Linnaeus, 1758)

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, Béni Mellal-Khénifra, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech, SA, Guelmim-Oued Noun; Chaker et al. 1980, MA, Lalla Outka, Khénifra, Oulmès, HA, Talet Inaouan (Haouz), AA, Aouinet-Torkoz, Tarhjicht; Remm 1988a; Dakki 1997; Bouayoune et al. 1998, Rif, Tanger-Tétouan-Al Hoceima, EM, Oriental, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, MA, Fès-Meknès, Béni Mellal-Khénifra, HA, Marrakech, AA, Drâa-Tafilalet, Souss-Massa; Cêtre-Sossah and Baldet 2004; Lhor et al. 2015, Rif, Sahel Chamali, MA, Aïn Leuh, Aït Siberne, Meknès; Bourquia et al. 2019

Culicoides (Culicoides) punctatus (Meigen, 1804)

Callot et al. 1968, Rif, Tanger-Tétouan-Al Hoceima; Bailly-Choumara and Kremer 1970, Rif, Merja Smir, Oued Negro, AP, Merja Sheishat (Larache), estuaire Bou-Regreg, EM, Merja Boubker (Berkane); Kremer et al. 1971, Rif, Cabo Negro (Ferma), Tétouan, EM, Berkane, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, AA, Foum Zguid; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Casablanca-Settat; Dakki 1997; Lhor et al. 2015, Rif, Sahel Chamali, MA, Sidi Hammadi, Benioukil, Aïn Leuh, Meknès; Bourquia et al. 2019

Culicoides (Culicoides) subfagineus Delécolle & Ortega, 1998

Bourquia et al. 2019, AP, Rabat

Culicoides (Monoculicoides) parroti Kieffer, 1922

Bailly-Choumara and Kremer 1970, HA, Dar Saâda (Haouz); Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979, AP, Safi, HA, Marrakech; Chakeret al. 1980, AP, Rabat, HA, Marrakech, Souk Tnine des Oudaias (Haouz); Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Monoculicoides) puncticollis (Becker, 1903)

Callot et al. 1968, AP, Merja Qoudiya, Sidi Yahia du Gharb, Romani (Zaers), Rabat-Salé-Kénitra, MA, Aïn Karma (Saiss), HA, Souk Tnine des Oudaias (Haouz); Bailly-Choumara and Kremer 1970 (reported as C. riethi and corrected by Kremer et al. 1971), Rif, Merja Smir, AP, Merja Sheishat (Larache), estuaire Bou-Regreg, Dayat Qoudiya (Sidi Yahia du Gharb), HA, Souk Tnine des Oudaias (bordure de l’Oued N’fis), Dar Saâda (Haouz), Talet Inouane (bordure marécageuse du lac du Barrage Lalla Taguergoust); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès; Kremer et al. 1975; Kremer et al. 1979, AP, Rabat-Salé-Kénitra, Safi, MA, Fès-Meknès, HA, Marrakech; Chaker et al. 1980, AP, Aïn Karma, Zaers, Roumani, HA, Souk-Tnine des Oudaias; Remm 1988a; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019, AP, Rabat

Culicoides (Oecacta) azerbajdzhanicus Dhzafarov, 1962

Bailly-Choumara and Kremer 1970, HA, Souk Tnine des Oudaias (bordure Oued N’fis); Kremer et al. 1971, AP, Safi, MA, Fès-Meknès, Beni Mellal-Khénifra, HA, Marrakech, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979, AA, Souss-Massa SA, Guelmim-Oued Noun; Chaker et al. 1980, MA, Kkénifra, Rhafsai, HA, Marrakech, AA, Torkoz, Tarhjisht; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) longipennis Khalaf, 1957

Kremer et al. 1971, EM, Berkane AP, Safi, MA, Fès-Meknès, HA, Marrakech; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Oecacta) marcleti Callot, Kremer & Basset, 1968

Kremer et al. 1971, MA, Rhafsai, Fès-Meknès, SA, Guelmim-Oued Noun; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) pallidus Khalaf, 1957

= Culicoides stackelbergi Dhzafarov, in Kremer et al. 1971: 662, Dakki 1997: 61

Kremer et al. 1971, AA, Torkoz, SA, Guelmim-Oued Noun; Kremer et al. 1971; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) ravus De Meillon, 1936

= Culicoides (Synhelea) subravus Cornet and Château, in Kremer et al. 1971: 664, Chaker et al. 1980: 85, Dakki 1997: 61

Kremer et al. 1971, AP, Safi, HA, Marrakech, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AA, Souss-Massa, SA, Guelmim-Oued Noun; Chaker et al. 1980, HA, Marrakech, AA, Torkoz, Tarhjicht, Aït Ouaballi (Draa); Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) sahariensis Kieffer, 1923

= Culicoides colluzzii Callot, Kremer and Bailly-Choumara, in Bailly-Choumara and Kremer 1970: 386, Chaker et al. 1980: 83, Dakki 1997: 61

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache), EM, Merja Boubker (Berkane), HA, Souk Tnine des Oudaias (bordure Oued N’fis); Callot et al. 1970, AP, Larache, HA, Marrakech; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Rabat-Salé-Kénitra, SA, Guelmim-Oued Noun; Chaker et al. 1980, AP, Merja Bokka (Gharb), Rabat (Zaers), EM, Berkane, MA, Fès, Khémisset, AA, Tarhjisht; Baylis et al. 1997; Dakki 1997; Bouayoune et al. 1998; Cêtre-Sossah and Baldet 2004; Bourquia et al. 2019, AP, Rabat

Culicoides (Oecacta) santonicus Callot, Kremer, Rault & Bach, 1966

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache); Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Rabat-Salé-Kénitra; Bailly-Choumara et al. 1980, AP, Larache, Sidi Bettache, MA, Oulmès, EM, El-Harcha; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) semimaculatus Clastrier, 1958

Kremer et al. 1975, Rif, Tanger-Tétouan-Al Hoceima, AP, Larache, Casablanca-Settat, MA, Plateau Central (Khatouate); Chaker et al. 1979; Remm 1988a; Dakki 1997; Szadziewski and Dominiak 2006; Bourquia et al. 2019

Culicoides (Oecacta) sergenti Kieffer, 1921

= Culicoides (Oecacta) mosulensis Khalaf, in Chaker et al. 1980: 84, Dakki 1997: 60

Kremer et al. 1979, EM, Oriental, SA, Guelmim-Oued Noun; Chaker et al. 1980, AA, Tarhjicht, SA, Bou-Arfa; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) similis Carter, Ingram & Macfie, 1920

Kremer et al. 1971, AA, Torkoz, SA, Guelmim-Oued Noun; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) truncorum Edwards, 1939

= Culicoides (Oecacta) sylvarum Callot and Kremer, in Kremer et al. 1971: 662, Remm 1988a: 65, Dakki 1997: 61

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Béni Mellal- Khénifra; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Pontoculicoides) saevus Kieffer, 1922

Callot et al. 1968, AP, Sidi Yahia du Gharb, Rabat-Salé-Kénitra, Safi, MA, Aïn Karma (Saiss), HA, Talet Inouane, Marrakech, Souk Tnine des Oudaias (Haouz), AA, Aït Melloul (Souss), Ksar er Souk (Tafilalt), Tarhjicht, SA, Bou-Arfa; Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache), HA, Souk Tnine des Oudaias (bordure de l’Oued N’fis); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, EM, Oriental, AP, Safi, MA, Fès-Meknès, HA, Marrakech, AA, Drâa-Tafilalet, Souss-Massa, SA, Guelmim-Oued Noun; Bailly-Choumara and Kremer 1980; Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019

Culicoides (Pontoculicoides) sejfadinei Dzhafarov, 1958

Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, EM, Oriental, AA, Drâa-Tafilalet; Bourquia et al. 2019

Culicoides (Remmia) kingi Austen, 1912

Bailly-Choumara and Kremer 1970, AA, Mrimima (Oued de Foum Zquid), Souss-Massa; Chaker et al. 1980, MA, Meknès, AA, Tarhjisht; Cornet and Brunhes 1994; Bourquia et al. 2019, AP, Rabat

Culicoides (Remmia) schultzei (Enderlein, 1908)

Callot et al. 1968, AP, Safi, HA, Talet Inouane, Marrakech; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) badooshensis Khalaf, 1961

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache), HA, Marrakech, Souk Tnine des Oudaias; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Safi, MA, Fès-Meknès; Kremer et al. 1975; Kremer et al. 1979, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech; Dakki 1997; Chaker et al. 1980, AP, Oued Cherrat, Bousselham, Sidi Yahia, MA, Fès, HA, Marrakech; Bourquia et al. 2019

Culicoides (Sensiculicoides) cataneii Clastrier, 1957

Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra; Chaker et al. 1980, AP, Oued Cherrat, Rabat; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) clastrieri Callot, Kremer & Deduit, 1962

Bourquia et al. 2019, MA, Fès-Meknès

Culicoides (Sensiculicoides) derisor Callot & Kremer, 1965

Chaker et al. 1980, AP, Rabat; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès; Kremer et al. 1975; Chaker et al. 1979, AP, Rabat-Salé-Kénitra; Kremer et al. 1979; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) duddingstoni Kettle & Lawson, 1955

Bourquia et al. 2019, MA, Fès-Meknès

Culicoides (Sensiculicoides) dzhafarovi Remm, 1967

Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, SA, Guelmim-Oued Noun; Chaker et al. 1980, AP, Oued Cherrat, AA, Tarhjisht; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) festivipennis Kieffer, 1914

= Culicoides (Oecacta) odibilis Austen, in Bailly-Choumara and Kremer 1970: 387, Chaker et al. 1980: 84, Dakki 1997: 61

Callot et al. 1968, Rif, Tanger-Tétouan-Al Hoceima; Bailly-Choumara and Kremer 1970, Rif, Merja Smir, HA, Souk Tnine des Oudaias (bordure Oued N‘fis); Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Safi, Casablanca-Settat, HA, Marrakech; Chaker et al. 1980, Rif, Tétouan, EM, El-Harcha, AP, Aïn Chok, Rabat, Larache, MA, Oulmès, HA, Marrakech, Talet-Inaouan (Haouz); Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) heteroclitus Kremer and Callot, in Callot & Kremer, 1965

Kremer et al. 1975, AP, Safi, HA, Marrakech, AA, Tafraout, Tiznit, Souss-Massa; Chaker et al. 1979; HA, Haouz; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) jumineri Callot & Kremer, 1969

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, EM, Oriental, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech, AA, Souss-Massa, SA, Guelmim-Oued Noun; Chaket et al. 1980, AP, Oued Cherrat, Merja Bokka, Rabat, MA, Fès, HA, Talet-Inaouan (Haouz), AA, Torkoz, Tarhjisht, Aït Oubelli, SA, Bou-Arfa; Remm 1988a; Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) kibunensis Tokunaga, 1937

= Culicoides (Oecata) cubitalis Edwards, in Kremer et al. 1975: 205, Dakki 1997: 60

Kremer et al. 1975, AP, Safi, MA, Ifrane, Imouzzer-du-Kander, Fès-Meknès, HA, Haouz, Marrakech; Chaker et al. 1979; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) kurensis Dzhafarov in Gutsevich, 1960

Remm 1988a; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019

Culicoides (Sensiculicoides) landauae Kremer, Rebholtz-Hirtzel & Bailly-Choumara, 1975

Kremer et al. 1975, MA, Imouzzer-du-Kander, Sefrou, Fès-Meknès; Chaker et al. 1979; Hervy et al. 1994; Borkent and Wirth 1997; Remm 1988a; Dakki 1997; Chilasse and Dakki 2004, MA; Borkent 2012; Bourquia et al. 2019

Culicoides (Sensiculicoides) langeroni Kieffer, 1921

Bailly-Choumara and Kremer 1970, HA, Tnine des Oudaias (bordure de Oued N‘fis); Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Bailly-Choumara and Kremer 1980, MA, Khénifra, AA, Tarhjicht, Torkoz (Draa); Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) maritimus Kieffer, 1924

Bailly-Choumara and Kremer 1970, AP, estuaire de Bou-Regreg; Kremer et al. 1971, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Bailly-Choumara and Kremer 1980, Rif, Tétouan, AP, Larache, Rabat, Sidi Bettache, HA, Talet-Inaouan (Haouz), Souk Tnine des Oudaias (Haouz); Remm 1988; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) odiatus Austen, 1921

= Culicoides lailae Khalaf, in Bailly-Choumara and Kremer 1970: 387, Dakki 1997: 60

= Culicoides indistinctus Khalaf, in Kremer et al. 1975: 206, Dakki 1997: 60

Bailly-Choumara and Kremer 1970, HA, Tnine des Oudaias (bordure de Oued N‘fis); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, AA, Torkoz, SA, Guelmim-Oued Noun; Kremer et al. 1975, AA, Tafraout, Souss-Massa; Kremer et al. 1979; Chaker et al. 1979, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech, SA, Guelmim-Oued Noun; Chaker et al. 1980, AP, Kénitra, Merja Bokka (Gharb), HA, Souk Tnine des Oudaias (Haouz), AA, Tarhjicht; Remm 1988a; Dakki 1997; Baylis et al. 1997; Bouayoune et al. 1998; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Sarvašová et al. 2014; Bourquia et al. 2019

Culicoides (Sensiculicoides) paolae Boorman, 1996

Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) pictipennis (Staeger, 1839)

Bailly-Choumara and Kremer 1970, AP, estuaire de Bou-Regreg; Kremer et al. 1971; Remm 1988a; Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) pseudopallidus Khalaf, 1961

Bailly-Choumara and Kremer 1970, HA, Souk Tnine des Oudaias (bordure Oued N’fis); Kremer et al. 1971, AP, Safi, MA, Rhafsai, Fès-Meknès, HA, Marrakech; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) shaklawensis Khalaf, 1957

Kremer et al. 1975, AP, Safi, MA, Sefrou, Fès-Meknès, HA, Setti Fatma, Marrakech; Chaker et al. 1979; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) simulator Edwards, 1939

Kremer et al. 1975, MA, Ifrane, Fès-Meknès, HA, Setti Fatma; Chaker et al. 1979; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) univittatus Vimmer, 1932

= Culicoides agathensis Callot, Kremer and Rioux, in Bailly-Choumara and Kremer 1970: 386, Kremer et al. 1971: 663, Chaker et al. 1980: 82, Dakki 1997: 60

Bailly-Choumara and Kremer 1970, AP, estuaire Bou-Regreg; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Rif, Tanger-Tétouan-Al Hoceima; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Safi, MA, Fès-Meknès, HA, Marrakech; Chaker et al. 1980, Rif, Tétouan, AP, Larache, Sidi Bettache MA, Oulmès; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) vidourlensis Callot, Kremer, Molet & Bach, 1968

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache), estuaire de Oued Bou-Regreg, HA, Souk Tnine des Oudaias (bordure de Oued N’fis); Kremer et al. 1971; Remm 1988a; Bourquia et al. 2019

Culicoides (Silvaticulicoides) pallidicornis Kieffer, 1919

Bailly-Choumara and Kremer 1970, HA, Souk Tnine des Oudaias (bordure Oued N’fis); Kremer et al. 1971, AA, Torkoz, SA, Guelmim-Oued Noun; Dakki 1997; Balenghien et al. 2014, SA, Guelmim-Oued Noun; Bourquia et al. 2019

Culicoides (Silvaticulicoides) picturatus Kremer & Deduit, 1961

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache); Kremer et al. 1971, Rif, Talerhza, EM, El-Harcha, AP, Bousselham, Rabat-Salé-Kénitra, MA, Oulmès; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Casablanca-Settat, MA, Béni Mellal-Khénifra; Remm 1988a; Dakki 1997; Sarvašová et al. 2014; Bourquia et al. 2019

Culicoides (Silvaticulicoides) subfasciipennis Kieffer, 1919

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, Béni Mellal-Khénifra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima; Bailly-Choumara and Kremer 1980, AP, Larache, Zaers, Rabat, Aïn Chok, MA, Sefrou; Remm 1988a; Hervy et al. 1994; Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019

Culicoides (Wirthomyia) faghihi Navai, 1971

Kremer et al. 1975, AA, Tafraout, Souss-Massa; Chaker et al. 1979; Remm 1988a; Hervy et al. 1994; Dakki 1997; Bourquia et al. 2019

Culicoides (Wirthomyia) minutissimus (Zetterstedt, 1855)

Referred as C. pumilus

Culicoides (Wirthomyia) pumilus (Winnertz, 1852)

Kremer et al. 1975, AP, Safi, MA, Ifrane, Imouzzer-du-Kander, Fès-Meknès, HA, Setti Fatma, Marrakech; Chaker et al. 1979; Dakki 1997; Bourquia et al. 2019

Culicoides calloti Kremer, Delécolle, Bailly-Choumara & Chaker, 1979

Chaker et al. 1979, AA, Souss Massa, SA, Guelmim-Oued Noun; Kremer et al. 1979, AA, Tarhjigt, Aït Ouaballi, Souss Massa, SA, Guelmim-Oued Noun; Chaker et al. 1980; Remm 1988a; Hervy et al. 1994; Borkent and Wirth 1997; Baylis et al. 1997; Dakki 1997; Koçak and Kemal 2010; Borkent 2012; Bourquia et al. 2019

Ceratopogonini

Alluaudomyia Kieffer, 1913

Alluaudomyia hygropetrica Vaillant, 1954

Vaillant 1956b, HA, Sidi Chamarouch

Palpomyiini

Bezzia Kieffer, 1899

Bezzia (Bezzia) nigritula (Zetterstedt, 1838)

= Palpomyia tenebricosa Goetghebuer, 1912, in Vaillant 1956b: 241

Vaillant 1956b, HA, Tamesrit

Palpomyia Meigen, 1818

Palpomyia helviscutellata Borkent, in Borkent & Wirth 1997

= Dasyhelea flavoscutellata (Zetterstedt, 1850), in Vaillant 1956b: 244

Vaillant 1956b, HA, Tahanaout

Forcipomyiinae

Dasyheleini

Dasyhelea Kieffer, 1911

Dasyhelea (Prokempia) flaviventris (Goetghebuer, 1910)

Dominiak 2012

Dasyhelea (Pseudoculicoides) turficola Kieffer, 1925

Dominiak 2012

Leptoconopinae

Leptoconops Skuse, 1889

Leptoconops (Holoconops) laurae (Weiss, 1912)

Remm 1988b

CHIRONOMIDAE

K. Kettani

Number of species: 412. Expected: 600

Faunistic knowledge of the family in Morocco: good

Buchonomyiinae

Buchonomyia Fittkau, 1955

Buchonomyia thienemanni Fittkau, 1955

Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012; Ashe et al. 2015, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Moubayed-Breil 2018, Rif

Podonominae

Paraboreochlus Thienemann, 1939

Paraboreochlus minutissimus (Strobl, 1895)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Tanypodinae

Macropelopiini

Apsectrotanypus Fittkau, 1962

Apsectrotanypus trifascipennis (Zetterstedt, 1838)

Kettani et al. 2010, Rif, Aïn Abou Hayane (Tiouertiouane, 880 m), Oued Maggou (Maggou village, 777 m), Oued Kanar (Gorges Kanar, 280 m); Kettani and Langton 2012

Macropelopia Thienemann, 1916

Macropelopia adaucta Kieffer, 1916

Kettani and Langton 2011, Rif, Fifi, Issaguen; Kettani and Langton 2012

Macropelopia nebulosa (Meigen, 1804)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Psectrotanypus Kieffer, 1909

Psectrotanypus varius (Fabricius, 1787)

Kettani et al. 1996; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012

Pentaneurini

Ablabesmyia Johannsen, 1905

Ablabesmyia (Ablabesmyia) ebbae Lehmann, 1981

Lehmann 1981; Azzouzi and Laville 1987; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Ablabesmyia (Ablabesmyia) longistyla Fittkau, 1962

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Barrage Talembote, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, source Maggou (Maggou, 1300 m), Oued Talembote; Kettani and Langton 2012

Ablabesmyia (Ablabesmyia) monilis (Linnaeus, 1758)

Reiss 1977, Rif, Tétouan, HA, kranichsee (Dra-Tal); Azzouzi and Laville 1987, Rif, retenue El Makhazine; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Naya 1988, MA, Haut Sebou; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Conchapelopia Fittkau, 1957

Conchapelopia (Conchapelopia) melanops (Meigen, 1818)

Kettani et al. 1996; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Conchapelopia (Conchapelopia) pallidula (Meigen, 1818)

Kettani and Moubayed-Breil 2018, Rif

Conchapelopia (Conchapelopia) viator (Kieffer, 1911)

= Conchapelopia Pe 1 Langton 1991 in Kettani et al. 1994: 28, Kettani et al. 1995: 256

Azzouzi et al. 1992, HA, Gorges de Dadès (Imdiazen, 1900 m); Kettani et al. 1994; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Ashe and O’Connor 2009; Kettani and Langton 2012

Larsia Fittkau, 1962

Larsia atrocincta (Goetghebuer, 1942)

Azzouzi et al. 1992, HA, Gorges de Dadès (Imdiazen, 1900 m); Kettani et al. 1994, Rif, Oued Moulay Bouchta; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Larsia curticalcar (Kieffer, 1918)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Nord Maggou village (Maggou, 905 m); Kettani and Langton 2012

Nilotanypus Kieffer, 1923

Nilotanypus dubius (Meigen, 1804)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Oued Siflaou, Oued Moulay Bouchta, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Kanar (Gorges Kanar, 280 m), Oued Tassikeste (Afechtal, 240 m), Oued Talembote (Usine électrique, 120 m), Oued Laou (Afertane, 55 m); Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paramerina Fittkau, 1962

Paramerina cingulata (Walker, 1856)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paramerina divisa (Walker, 1856)

Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and Langton 2012

Paramerina mauretanica Fittkau, 1962

Fittkau 1962, Atlas (850 m), SA; Azzouzi and Laville 1987; Ashe and Cranston 1990, EM, Figuig; Kettani et al. 2001; Ashe and O’Connor 2009, EM, Figuig; Kettani et al. 2010, Rif, Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012

Paramerina spec. Greichenland (Fittkau, 1962)

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Laville and Langton 2002; Kettani et al. 2010, Rif, Oued Chrafat (Armotah, 900 m), Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Moubayed-Breil 2018, Rif

Pentaneurella Fittkau & Murray, 1983

Pentaneurella sp. Ourika

Azzouzi et al. 1992, HA; Kettani et al. 2001

Rheopelopia Fittkau, 1962

Rheopelopia maculipennis (Zetterstedt, 1838)

Naya 1988, MA, Haut et Moyen Sebou; Azzouzi and Laville 1987, MA, Oum-er-Rbia, HA, Tensift; Kettani et al. 1994, Rif, Oued Siflaou, Oued Moulay Bouchta; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Ruisselet maison forestière (Talassemtane, 1683 m), Source Maggou (Maggou, 1300 m), Oued Talembote (avant village Talembote, 320 m), Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheopelopia murrayi Dowling, 1983

Dowling 1983, AA, Tata (Moyen Draa); Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; 2012; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheopelopia ornata (Meigen, 1838)

Azzouzi and Laville 1987, MA, Oued Fès; Kettani et al. 1995, Rif, Oued El Kbir, amont Oued Nakhla, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Telopelopia Roback, 1971

Telopelopia fascigera (Verneaux, 1970)

= Telopelopia maroccana Murray, 1980, in Reiss 1977: 91, Murray 1980: 151, Azzouzi and Laville 1987: 218, Ashe and Cranston 1990: 133

Reiss 1977, AP, Larache, HA, Dra-Tal; Murray 1980, AP, Larache, HA, Dra-Tal; Azzouzi and Laville 1987; Ashe and Cranston 1990; Azzouzi and Laville 1987, MA, Oum-er-Rbia; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Telmatopelopia Fittkau, 1962

Telmatopelopia nemorum (Goetghebuer, 1921)

Kettani et al. 1996, Rif, Oued Khizana (Oued Laou); Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Thienemannimyia Fittkau, 1957

Thienemannimyia (Thienemannimyia) berkanea Dowling, 1987

Dowling 1987, EM, Berkane; Azzouzi et al. 1992, EM, Environs de Berkane, HA, Ouarzazate (1160 m), Oasis Meski (1160 m), Aït Saoun; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Thienemannimyia (Thienemannimyia) carnea (Fabricius, 1805)

Kettani and Langton 2012, Rif

Thienemannimyia (Thienemannimyia) choumara Dowling, 1983

Dowling 1983, EM, Environ de Berkane (Monts de Bni Snassen), HA, Souk des Judais (Marrakech); Azzouzi et al. 1987, HA, Dra-Tal; Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Thienemannimyia (Thienemannimyia) geijskesi (Goetghebuer, 1934)

Kettani and Langton 2012, Rif, Oued Zarka

Thienemannimyia (Thienemannimyia) laeta (Meigen, 1818)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Thienemannimyia (Thienemannimyia) lentiginosa (Fries, 1823)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Thienemannimyia (Thienemannimyia) northumbrica (Edwards, 1929)

Fittkau 1962; Azzouzi and Laville 1987, MA, Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2009; Kettani and Langton 2012

Trissopelopia Kieffer, 1923

Trissopelopia longimana (Staeger, 1839)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Xenopelopia Fittkau, 1962

Xenopelopia falcigera (Kieffer, 1911)

Kettani and Langton 2011, Rif, Anasser, Fifi, AP, marais de Loukous; Kettani and Langton 2012

Xenopelopia nigricans (Goetghebuer, 1927)

Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique); Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Zavrelimyia Fittkau, 1962

Zavrelimyia (Zavrelimyia) barbatipes (Kieffer, 1911)

Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Ashe and O’Connor 2009 (?); Kettani et al. 2010, Rif, Oued Tiffert (Tiffert Talassemtane, 1230 m), Aïn Abou Hayane (Tiouertiouane, 880 m), Oued Abiyati (Ifansa, 140 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Zavrelimyia (Zavrelimyia) berberi Fittkau, 1962

Azzouzi and Laville 1987; Ashe and Cranston 1990, HA, Tamhda; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Zavrelimyia (Zavrelimyia) hirtimana (Kieffer, 1918)

Kettani and Langton 2012

Zavrelimyia (Zavrelimyia) melanura (Meigen, 1804)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Zavrelimyia (Zavrelimyia) nubila (Meigen, 1830)

Kettani and Langton 2011, Rif, marais de Lemtahane (PNPB), Dayat Aïn Rami, Dayat Amlay; Kettani and Langton 2012

Procladiini

Procladius Skuse, 1889

Procladius (Holotanypus) brevipetiolatus (Goetghebuer, 1935)

Azzouzi et al. 1992, HA, Oued Meski (1160 m), Khettaras de Marrakech; Kettani et al. 2001; Kettani and Langton 2012

Procladius (Holotanypus) choreus (Meigen, 1804)

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, AP, Merja Sidi Boughaba; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote; Kettani et al. 1995, Rif, amont Oued Nakhla, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2010, Rif, Aïn Talassemtane (Talassemtane, 1700 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Procladius (Holotanypus) culiciformis (Linnaeus, 1767)

Kettani and Moubayed-Breil 2018, Rif

Procladius (Holotanypus) noctivagus (Kieffer, 1910)

Azzouzi et al. 1992, HA, Ouarzazate (1160 m); Kettani et al. 2001; Kettani and Langton 2012

Procladius (Holotanypus) sagittalis (Kieffer, 1909)

Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Procladius (Psilotanypus) anomalus Kieffer, 1906

Nomen dubium in Ashe and O’Connor 2009: 213

Naya 1988, MA; Kettani et al. 2001; Kettani and Langton 2012

Procladius Pe 3 Langton 1991

Kettani et al. 1994, Kettani et al. 1995, Rif, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 2001; Dakki 1997

Tanypodini

Tanypus Meigen, 1803

Tanypus (Tanypus) brevipalpis (Kieffer, 1923)

Reiss 1977, EM, Berkane; Ashe and O’Connor 2009 (?); Kettani and Langton 2012

Tanypus (Tanypus) kraatzi (Kieffer, 1912)

Azzouzi et al. 1992, HA, Oasis Meski; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Tanypus (Tanypus) punctipennis Meigen, 1818

Reiss 1977, EM, Berkane; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesinae

Boreoheptagyiini

Boreoheptagyia Brundin, 1966

Boreoheptagyia legeri (Goetghebuer, 1933)

= Boreoheptagyia punctulata (Goetghebuer, 1934), in Kettani et al. 2001: 327

Ashe and Cranston 1990; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesini

Diamesa Meigen, 1835

Diamesa aberrata Lundbeck, 1898

Saether 1968; Serra-Tosio 1973; Fittkau and Reiss 1987; Serra-Tosio 1983; Azzouzi and Laville 1987, HA (2500–3350 m); Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Diamesa bertrami Edwards, 1935

Serra-Tosio 1983, HA, Gorges de Todra (2500 m); Azzouzi and Laville 1987, HA, Gorges Todra; Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Diamesa hamaticornis Kieffer, 1924

Reiss 1977; Serra-Tosio 1983, HA, M’Goum; Azzouzi and Laville 1987, HA, M’Goum; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesa insignipes Kieffer, 1908

Serra-Tosio 1983, HA (2500 m); Azzouzi and Laville 1987; Naya 1988, MA, Haut and Moyen Sebou; Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Diamesa latitarsis (Goetghebuer, 1921)

Vaillant 1955b; Vaillant 1956b, HA, Asif Tessaout (M’Goum), Lac Tamhda (Anremer); Serra-Tosio 1967; Serra-Tosio 1967; Saether 1968; Serra-Tosio 1973; Azzouzi and Laville 1987, HA; Ashe and Cranston 1990; Kettani et al. 2001, Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesa steinboecki Goetghebuer, 1933

Vaillant 1956b, HA, Cascade Siroua, Oukaimeden, Sidi Chamarouch

Diamesa tonsa (Haliday in Walker, 1856)

= Diamesa thienemanni Kieffer, 1909

Naya 1988, MA, Haut Sebou (Arhbalou Yahya, Oued Arbi, Pont Aït hamza); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2011, Rif, Oued Ketama, Oued Sgara; Kettani and Langton 2012

Diamesa vaillanti Serra-Tosio, 1972

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Diamesa veletensis Serra-Tosio, 1971

Serra-Tosio 1983, HA (2500 m); Azzouzi and Laville 1987, HA; Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesa zernyi Edwards, 1933

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Potthastia Kieffer, 1922

Potthastia gaedii (Meigen, 1838)

Azzouzi and Laville 1987, MA, oued Boufekrane, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Laou, Oued Afertane; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Potthastia pastoris (Edwards, 1933)

Kettani and Moubayed-Breil 2018, Rif

Pseudodiamesa Goetghebuer, 1939

Pseudodiamesa (Pseudodiamesa) branickii (Nowicki, 1873)

Naya 1988, MA, Haut Sebou; Ashe and Cranston 1990; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2001; Kettani and Langton 2012

Pseudodiamesa (Pseudodiamesa) nivosa (Goetghebuer, 1928)

Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2009; Kettani and Langton 2012

Sympothastia Pagast, 1947

Sympothastia zavreli Pagast, 1947

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1995, Rif, aval Oued Krikra; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Syndiamesa Kieffer, 1918

Syndiamesa hygropterica (Kieffer, 1909)

Naya 1988, MA, Moyen Sebou (Sidi Abdellah, Dar El Arsa, Pont Oulad Slimane, Pont Portugais); Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Protanypini

Protanypus Kieffer, 1906

Protanypus morio (Zetterstedt, 1838)

Naya 1988, MA, Moyen Sebou; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Prodiamesinae

Odontomesa Pagast, 1947

Odontomesa fulva (Kieffer, 1919)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Prodiamesa Kieffer, 1906

Prodiamesa olivacea (Meigen, 1818)

Naya 1988, MA, Haut Sebou (Haut Guigou); Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Maggou village, Ifansa; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladiinae

Orthocladiini

Acricotopus Kieffer, 1921

Acricotopus lucens (Zetterstedt, 1850)

Kettani and Moubayed-Breil 2018, Rif

Brilla Kieffer, 1913

Brillia bifida (Kieffer, 1909)

= Brilla modesta (Meigen, 1830)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1996; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Tamaridine (Zaouiet El Habtiyine, 819 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Brillia flavifrons (Johannsen, 1905)

Kettani and Langton 2012

Brilla longifurca Kieffer, 1921

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Sebou, Oued Oum-er-Rbia; Kettani et al. 1995, Rif, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m), Oued Talembote (Usine électrique, 120 m); Kettani and Langton 2012; Ashe and O’Connor 2012

Bryophaenocladius Thienemann, 1934

Bryophaenocladius aestivus (Brundin, 1947)

Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Bryophaenocladius flexidens (Brundin, 1947)

Kettani and Moubayed-Breil 2018, Rif

Bryophaenocladius cf. furcatus Thienemann & Strenzke, 1940

Kettani and Moubayed-Breil 2018, Rif

Bryophaenocladius illimbatus (Edwards, 1929)

Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Bryophaenocladius muscicola (Kieffer, 1906)

Kettani and Moubayed-Breil 2018, Rif

Bryophaenocladius nidorum (Edwards, 1929)

Kettani and Moubayed-Breil 2018, Rif

Bryophaenocladius subvernalis (Edwards, 1929)

Azzouzi and Laville 1987, MA, Oued Boufekrane; Kettani et al. 2001; Kettani and Langton 2011, Rif, Oued Taida (Moulay Abdelsalam); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Bryophaenocladius tuberculatus (Edwards 1929)

Kettani and Moubayed-Breil 2018, Rif

Camptocladius Wulp, 1874

Camptocladius stercorarius (De Geer, 1976)

Kettani and Moubayed-Breil 2018, Rif

Cardiocladius Kieffer, 1912

Cardiocladius capucinus (Zetterstedt, 1850)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, Oued Afertane, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Tassikeste (Afechtal, 240 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cardiocladius fuscus Kieffer, 1924

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Oum-er-Rbia; Naya 1988, MA, Haut Sebou (Amont de Aïn Tadout, Skhounate, amont confluence avec Oued Atchane, Pont Aït Hamza); Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Chaetocladius Kieffer, 1911

Chaetocladius (Chaetocladius) acuticornis (Kieffer in Potthast, 1914)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chaetocladius dentiforceps (Edwards, 1929)

Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Chaetocladius dissipatus (Edwards, 1929)

Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Chaetocladius (Chaetocladius) melaleucus (Meigen, 1818)

Kettani and Langton 2011, Rif, Oued Sgara, Bab Tariouant, Bouztata; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Chaetocladius piger (Goetghebuer, 1913)

Kettani and Moubayed-Breil 2018, Rif

Chaetocladius (Chaetocladius) perennis (Meigen, 1830)

Kettani and Langton 2011, Rif, Oued Hamma; Kettani and Langton 2012

Chaetocladius (Chaetocladius) vitellinus (Kieffer in Kieffer & Thienemann, 1908)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Corynoneura Winnertz, 1846

Corynoneura carriana Edwards, 1924

Naya 1988, MA, Haut Sebou (Haut Guigou, Aïn Nokra); Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Corynoneura celtica Edwards, 1924

Kettani and Langton 2011, Rif, Oued Hamma; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Corynoneura coronata Edwards, 1924

Kettani and Langton 2011, Rif, Oued Hamma; Kettani and Langton 2012; Ashe and O’Connor 2012

Corynoneura edwardsi Brundin, 1949

Kettani and Langton 2012

Corynoneura lacustris Edwards, 1924

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Corynoneura lobata Edwards, 1924

Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Azzouzi et al. 1992, HA; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Corynoneura scutellata Winnertz, 1846

Kettani and Moubayed-Breil 2018, Rif

Corynoneura Pe 2 Langton 1991

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Corynoneurella Brundin, 1949

Corynoneurella paludosa Brundin, 1949

Kettani et al. 2010, Rif, Nord Maggou village (Maggou, 905 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (avant village Talembote, 320 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus van der Wulp, 1874

Cricotopus (Cricotopus) albiforceps (Kieffer in Thienemann and Kieffer 1916)

Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) annulator Goetghebuer, 1927

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m), Oued Tassikeste (Afechtal, 240 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2011, Rif, Oued Sgara; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) beckeri Hirvenoja, 1973

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1995, Rif, aval Oued El Kbir; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Langton and Laville 2002; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) bicinctus (Meigen, 1818)

Azzouzi and Laville 1987, MA, Oued Boufekrane; Naya 1988, MA, Haut et Moyen Sebou; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, Oued Afertane, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) caducus Hirvenoja, 1973

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) ephippium (Zetterstedt, 1838)

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) levantinus Moubayed & Hirvenoja, 1986

Kettani et al. 1996, Rif, Haut Maggou; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Laville and Langton 2002; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Inesmane (Adeldal, 1173 m); Kettani and Langton 2012; Ashe and O’Connor 2012

Cricotopus (Cricotopus) pallidipes Edwards, 1929

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1995, Rif, Oued Martil (Tamuda); Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) pulchripes Verrall, 1912

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) similis Goetgnebuer, 1921

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Afertane, aval Oued Talembote (usine éléctrique), aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) tremulus (Linnaeus, 1758)

Kettani et al. 2010, Rif, Oued Maggou (Maggou village, 777 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) triannulatus (Macquart, 1826)

Kettani et al. 1994, Rif, Haut Laou, Oued Moulay Bouchta, Oued Afertane; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) trifascia Edwards, 1929

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Maggou (Maggou village, 777 m), Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) vierriensis Goetghebuer, 1935

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, Oued Sebou, Oued Oum-er-Rbia, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Isocladius) brevipalpis Kieffer, 1909

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Cricotopus (Isocladius) laetus Hirvenoja, 1973

Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Cricotopus (Isocladius) ornatus (Meigen, 1818)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Cricotopus (Isocladius) sylvestris (Fabricius, 1794)

Fittkau and Reiss 1978; Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Isocladius) tricinctus (Meigen, 1818)

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Paratrichocladius) micans (Kieffer, 1918)

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Haut Maggou (1300 m), Oued Talembote, Oued Laou (Afertane, 56 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Paratrichocladius) osellai Rossaro, 1990

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Paratrichocladius) rufiventris (Meigen, 1830)

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Fès, Oued Boufekrane, Oued Sebou, Oued Oum-er-Rbia, HA, Oued Tensift; (Dar El Arsa, Pont oulad Slimane, Pont portugais); Naya 1988, MA, Haut et Moyen Sebou; Kettani et al. 1995, Rif, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m), Oued Talembote; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Paratrichocladius) skirwithensis (Edwards, 1929)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukieferiella Thienemann, 1926

Eukieferiella ancyla Svensson, 1986

Kettani and Langton 2011, Rif, Oued Tkarae; Kettani and Langton 2012; Ashe and O’Connor 2012

Eukiefferiella bedmari Vilchez-Quero & Laville, 1988

Azzouzi and Laville 1987, MA, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Laville and Langton 2002; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella brehmi Gowin, 1943

Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (avant village Talembote, 320 m), Oued Talembote (Usine électrique, 120 m)

Eukiefferiella brevicalcar (Kieffer, 1911)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani et al. 2010, Rif, Nord Maggou village (Maggou, 905 m), Oued Ametrasse (Ametrasse, 820 m); Kettani and Langton 2011, Rif, Oued Issaguen, Oued Ketama, Oued Sgara; Bab Tariouant, Bouztate (Fifi); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella claripennis (Lundbeck, 1898)

Fittkau and Reiss 1978; Naya 1988, MA, Moyen Sebou (Dar Cheik Harazem); Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukieffeiella clypeata (Thienemann, 1919)

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Oued Talembote (usine éléctrique); Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella coerulescens (Kieffer in Zavřel, 1926)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Naya 1988, MA, Haut Sebou (Skhounate, Arhbalou Aberchane); Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Nord Maggou village (Maggou, 905 m), Oued Tamaridine (Zaouiet El Habtiyine, 819 m), Oued Maggou (Maggou village, 777 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella cyanea Thienemann, 1936

Vaillant 1955b, HA; Fittkau and Reiss 1978; Azzouzi and Laville 1987, HA; Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella devonica (Edwards, 1929)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Eukiefferiella dittmari Lehmann, 1972

Kettani and Langton 2011, Rif, Oued Boujdad, Fifi; Kettani and Langton 2012, Rif, Oued Zarka; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella fittkaui Lehmann, 1972

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Eukiefferiella fuldensis Lehmann, 1972

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Eukiefferiella gracei (Edwards, 1929)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, aval Oued Tassikeste; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Eukiefferiella ilkleyensis (Edwards, 1929)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella lobifera Goetghebuer, 1934

Azzouzi and Laville 1987, MA, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2011, Rif, Oued Ketama, Oued Sgara; Kettani and Langton 2012; Ashe and O’Connor 2012

Eukiefferiella minor (Edwards, 1929)

Vaillant 1955b, HA (1050 m); Vaillant 1956b, HA, Imi-N’Ifri; Azzouzi and Laville 1987, HA; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Eukiefferiella pseudomontana Goetghebuer, 1935

Kettani et al. 2010, Rif, Oued Madissouka (Talassemtane, 1530 m), Oued Dchar d’Amran (Béni M’Hamed, 1180 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella similis Goetghebuer, 1939

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, Oued Sebou, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella tirolensis Goetghebuer, 1938

Kettani et al. 1994, Rif, Oued Afertane; Kettani et al. 1996; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m); Azzouzi et al. 1992, HA, Oued Tensift; Dakki 1997; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella Pe 2 Langton 1991

Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012

Halocladius Hirvenoja, 1973

Halocladius (Halocladius) varians (Staeger, 1839)

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m); Ashe and Cranston 1990; Kettani and Langton 2012; Ashe and O’Connor 2012

Heleniella Gowin, 1943

Heleniella dorieri Serra-Tosio, 1967

Kettani and Langton 2012

Heleniella ornaticollis (Edwards, 1929)

Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012; Ashe and O’Connor 2012

Heleniella serratosioi Ringe, 1976

Kettani and Langton 2011, Rif, Oued Hamma, Bouztate (Fifi); Kettani and Langton 2012; Ashe and O’Connor 2012

Heterotrissocladius Spärck, 1923

Heterotrissocladius marcidus (Walker, 1856)

Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Kettani and Langton 2012; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2012

Hydrobaenus Fries, 1830

Hydrobaenus conformis (Holmgren, 1869)

Kettani and Moubayed-Breil 2018, Rif

Hydrosmittia Ferrington & Sæther, 2011

Hydrosmittia oxoniana (Edwards, 1929)

= Pseudosmittia recta (Edwards, 1929), in Azzouzi and Laville 1987: 218, Kettani et al. 2001: 330, Kettani and Langton 2012: 422

Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Hydrosmittia ruttneri (Strenzke & Thienemann, 1942)

Kettani and Moubayed-Breil 2018, Rif

Krenosmittia Thienemann & Krüger, 1939

Krenosmittia boreoalpina (Goetghebuer, 1944)

Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani and Langton 2012; Ashe and O’Connor 2012

Krenosmittia camptophleps (Edwards, 1929)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2011, Rif, Oued Ketama, Oued Sgara; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Krenosmittia halvorseni (Cranston & Sæther, 1986)

Kettani et al. 1995, Rif, aval Oued El Kbir; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Krenosmittia hispanica Wülker, 1957

Kettani et al. 2001; Laville and Langton 2002; Ashe and O’Connor 2012

Limnophyes Eaton, 1875

Limnophyes difficilis Brunidin, 1947

Kettani and Moubayed-Breil 2018, Rif

Limnophyes gelasinus Saether, 1990

Kettani and Moubayed-Breil 2018, Rif

Limnophyes habilis (Walker, 1856)

Kettani and Moubayed-Breil 2018, Rif

Limnophyes madeirae Sæther, 1985

Kettani and Moubayed-Breil 2018, Rif

Limnophyes minimus (Meigen, 1818)

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987, MA, Lac Aguelmane Azigza; Kettani et al. 2001; Kettani and Langton 2011; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Limnophyes natalensis (Kieffer, 1914)

Kettani and Moubayed-Breil 2018, Rif

Limnophyes ninae Sæther, 1975

Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Limnophyes pentaplastus (Kieffer, 1921)

Kettani and Moubayed-Breil 2018, Rif

Limnophyes pumilio (Holmgren, 1869)

Kettani and Moubayed-Breil 2018, Rif

Limnophyes punctipennis (Goetghebuer, 1919)

Kettani and Langton 2012

Limnophyes Pe 1a Langton 1991

Kettani et al. 2010, Rif, Oued Talembote (Usine électrique, 120 m); Kettani and Langton 2012

Metriocnemus van der Wulp, 1874

Metriocnemus (Metriocnemus) albolineatus Meigen, 1818

Kettani and Moubayed-Breil 2018, Rif

Metriocnemus (Metriocnemus) eurynotus (Holmgren, 1883)

= Metriocnemus hygropetricus Kieffer, 1912, in Ashe and O’Connor 2012: 372

= Metriocnemus (Metriocnemus) obscuripes (Holmgren, 1869), in Azzouzi et al. 1992: 229, Kettani et al. 2001: 329, Kettani and Langton 2012: 421

Boumezzough and Thomas 1987, HA, Oued Réghaya (1740 m), Imlil; Azzouzi and Laville 1987, HA, Oued Tensift; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Metriocnemus (Metriocnemus) fuscipes (Meigen, 1818)

Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Metriocnemus (Metriocnemus) hirticollis (Staeger, 1839)

Kettani and Moubayed-Breil 2018, Rif

Metriocnemus (Metriocnemus) ursinus Holmgren, 1869

Kettani and Moubayed-Breil 2018, Rif

Nanocladius Kieffer, 1913

Nanocladius (Nanocladius) balticus (Palmén, 1959)

Kettani et al. 1995, Rif, Oued El Kbir, amont Oued Nakhla; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Nanocladius (Nanocladius) dichromus (Kieffer 1906)

Kettani and Moubayed-Breil 2018, Rif

Nanocladius (Nanocladius) parvulus (Kieffer 1909)

Kettani and Moubayed-Breil 2018, Rif

Nanocladius (Nanocladius) rectinervis (Kieffer, 1911)

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Inesmane (Adeldal, 1173 m), Oued Talembote (aval Barrage Talembote, 245 m); Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius van der Wulp, 1874

Orthocladius (Eudactylocladius) fuscimanus (Kieffer, 1908)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Oued Talembote (usine éléctrique); Kettani et al. 1995, Rif, aval Oued Krikra, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani et al. 2010, Rif, source Maggou (Maggou, 1300 m), Oued Chrafat (Armotah, 900 m), Oued Tamaridine (Zaouiet El Habtiyine, 819 m), Oued Talembote (Usine électrique, 120 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Euorthocladius) ashei Soponis, 1990

= Orthocladius luteipes Goetghebuer, in Azzouzi and Laville 1987: 218

= Orthocladius rivicola Kieffer, in Azzouzi and Laville 1987: 218

Azzouzi and Laville 1987, MA, Oued Fès, Oued boufekrane, Oued Sebou, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique), aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m), Oued Kanar (Gorges Kanar, 280 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (Usine électrique, 120 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2011, Rif, Bouztate (Fifi); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Euorthocladius) rivulorum Kieffer, 1909

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1995, Rif, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor; 2012 Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Euorthocladius) thienemanni Kieffer, 1906

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Tamaridine (Zaouiet El Habtiyine, 819 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (avant village Talembote, 320 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Tassikeste (Afechtal, 240 m), Oued Talembote (Usine électrique, 120 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Mesorthocladius) frigidus (Zetterstedt, 1838)

Vaillant 1955, HA (2900 m); Vaillant 1956b, HA, Lac Tamhda (Anremer); Azzouzi and Laville 1987; Naya 1988, MA, Haut Sebou (Haut Guigou); Fekhaoui et al. 1993; Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Dchar d’Amran (Béni M’Hamed, 1180 m), Nord Maggou village (Maggou, 905 m), Oued Tamaridine (Zaouiet El Habtiyine, 819 m), Oued Maggou (Maggou village, 777 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Orthocladius (Orthocladius) oblidens (Walker, 1856)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2011, Rif, Bouztate (Fifi); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Orthocladius) obumbratus Johannsen, 1905

= Orthocladius excavatus Brundin, in Azzouzi and Laville 1987: 218

Azzouzi and Laville 1987, MA, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012

Orthocladius (Orthocladius) pedestris Kieffer, 1909

Kettani et al. 2010, Rif, Oued Tassikeste (Afechtal, 240 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Orthocladius) rubicundus (Meigen, 1818)

= Orthocladius saxicola Kieffer, in Azzouzi and Laville 1987: 218

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Sebou, Oued Oum-er-Rbia, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m), Oued Kanar (Gorges Kanar, 280 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (Usine électrique, 120 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Orthocladius) vaillanti Langton & Cranston, 1991

Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Symposiocladius) lignicola Kieffer in Potthast, 1914

= Symposiocladius lignicola Kieffer, in Kettani et al. 2010: 70

Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012; Ashe and O’Connor 2012

Orthocladius (Symposiocladius) ruffoi Rossaro & Prato, 1991

= Orthocladius Pe 1 Langton 1991, in Azzouzi and Laville 1987: 218

= Rheortocladius sp A Langton 1991, in Kettani et al. 1995: 257

= Rheorthocladius ruffoi Rossaro, in Kettani et al. 1997: 184

Azzouzi and Laville 1987, MA, Oum-er-Rbia; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Tamaridine (Zaouiet El Habtiyine, 819 m), Oued Maggou (Maggou village, 777 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (avant village Talembote, 320 m), Oued Kanar (Gorges Kanar, 280 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Tassikeste (Afechtal, 240 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Ifansa, 105 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012

Paracricotopus Brundin, 1956

Paracricotopus niger (Kieffer, 1913)

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès; Kettani et al. 1994, Rif, Oued Afertane; Kettani et al. 1995, Rif, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou); Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2001; Kettani et al. 2010, Rif, Haut Maggou, Oued Tamaridine (Zaouit et Habtyiène, 819 m), Oued Kelaâ (Akoumi, 400 m), Oued Kanar (Gorges Kanar, 280 m), Oued Talembote (155 m), Oued Tassikeste (240 m), Oued Laou (Ifansa, 105 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Parakiefferiella Thienemann, 1936

Parakiefferiella coronata (Edwards, 1929)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Parakiefferiella wuelkeri Moubayed, 1994

= Parakiefferiella sp. d Wülker, in Azzouzi et al. 1992: 230

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1995, Rif, aval Oued El Kbir; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Parametriocnemus Goetghebuer, 1932

Parametriocnemus boreoalpinus Gowin & Thienemann, 1942

Kettani and Langton 2011, Rif, Oued Taida (Moulay Abdelsalam); Kettani and Langton 2012; Ashe and O’Connor 2012

Parametriocnemus stylatus (Spärck, 1923)

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Béni M’Hamed (1330 m), Haut Maggou (1300 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (320 m), Oued Tassikeste (Afechtal, 240 m), Oued Laou (Afertane, 56 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Parametriocnemus valescurensis Moubayed & Langton, 1999

Kettani and Langton 2011, Rif, Oued Issaguen; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Parametriocnemus Pe 1 Langton 1991

Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Haut Maggou (1300 m), Oued Talembote (320 m); Kettani and Langton 2012

Paraphaenocladius Thienemann, 1924

Paraphaenocladius exagitans ssp. 1

Kettani and Moubayed-Breil 2018, Rif

Paraphaenocladius impensus impensus (Walker, 1856)

Kettani and Moubayed-Breil 2018, Rif

Paraphaenocladius irritus Walker, 1856

Kettani and Moubayed-Breil 2018, Rif

Paraphaenocladius pseudirritus Strenzke, 1950

Kettani and Moubayed-Breil 2018, Rif

Paratrissocladius Zavřel, 1937

Paratrissocladius excerptus (Walker, 1856)

Kettani et al. 1996, Rif, Ras el Ma (Chefchaouen); Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Parorthocladius Thienemann, 1935

Parorthocladius nudipennis (Kieffer in Kieffer & Thienemann 1908)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Psecrocladius Kieffer, 1906

Psectrocladius (Allopsectrocladius) obvius (Walker, 1856)

= Psectrocladius dilatatus (van der Wulp, 1859), in Naya 1988: 48

Naya 1988, MA, Moyen Sebou; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2011, Rif, sources de Issaguen; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Psectrocladius (Allopsectrocladius) platypus (Edwards, 1929)

Kettani and Moubayed-Breil 2018, Rif

Psectrocladius (Mesopsectrocladius) barbatipes Kieffer, 1923

Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique); Kettani et al. 1995, Rif, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Haut Maggou (1300 m), Oued Laou (Afertane, 56 m); Dakki 1997; Kettani and Langton 2012; Ashe and O’Connor 2012

Psecrocladius (Psectrocladius) brehmi Kieffer, 1923

Kettani et al. 1995, Rif, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Psectrocladius (Psectrocladius) fennicus Storå, 1939

Kettani and Langton 2012

Psectrocladius (Psectrocladius) limbatellus (Holmgren, 1869)

Wülker 1959; Azzouzi and Laville 1987, HA, Lac Tamhda (2800 m); Kettani et al. 2001; Kettani and Langton 2011, AP, marais de Loukous; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Psectrocladius (Psectrocladius) octomoculatus Wülker, 1956

Kettani et al. 1995, Rif, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Psectrocladius (Psectrocladius) sordidellus (Zetterstedt, 1838)

Azzouzi and Laville 1987, MA, Lac Aguelmane Azigza; Kettani et al. 2001; Kettani and Langton 2011, AP, marais de Loukous (NE Boucharene); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Psectrocladius (Psectrocladius) ventricosus Kieffer, 1925

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Pseudosmittia Edwards, 1932

Pseudosmittia albipennis (Goetghebuer, 1921)

Kettani and Moubayed-Breil 2018, Rif

Pseudosmittia baueri Strenzke, 1960

Kettani and Moubayed-Breil 2018, Rif

Pseudosmittia danconai (Marcuzzi, 1947)

Kettani and Moubayed-Breil 2018, Rif

Pseudosmittia holsata Thienemann & Stenzke, 1940

Kettani and Moubayed-Breil 2018, Rif

Pseudosmittia obtusa Strenzke, 1960

Kettani and Moubayed-Breil 2018, Rif

Pseudosmittia trilobata Edwards, 1929

Kettani and Moubayed-Breil 2018, Rif

Pseudorthocladius Goetghebuer, 1943

Pseudorthocladius (Pseudorthocladius) berthelemyi Moubayed, 1990

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Laville and Langton 2002; Kettani and Langton 2012; Ashe and O’Connor 2012; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Pseudorthocladius (Pseudorthocladius) curtistylus (Goetghebuer, 1921)

Azzouzi et al. 1992, HA, Oasis Meski (1160 m); Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Pseudorthocladius near Pe 3 Langton 1991

Kettani and Langton 2011, Rif, Bouztate (Fifi); Kettani and Langton 2012

Rheocricotopus Brundin, 1956

Rheocricotopus (Psilocricotopus) atripes (Kieffer, 1913)

= Rheocricotopus (Psilocricotopus) foveatusfoveatus (Edwards, 1929), in Naya 1988: 40

Naya 1988, MA, Haut Sebou (Haut Guigou); Azzouzi et al. 1992, HA, Oued Tensift, Gorges de Dadès (Imdiazen, 1900 m); Kettani et al. 1994, Rif, Haut Laou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote, Oued Tassikeste (Afechtal, 240 m), Oued Laou (Afertane, 56 m); Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Rheocricotopus (Psilocricotopus) chalybeatus subsp. chalybeatus (Edwards, 1929)

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Sebou, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, Oued Afertane, aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Tamaridine (Zaouiet et Habtiyiène, 819 m); Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Rheocricotopus (Psilocricotopus) gallicus Lehamnn 1969

Kettani and Moubayed-Breil 2018, Rif

Rheocricotopus (Psilocricotopus) glabricollis (Meigen, 1830)

Azzouzi and Laville 1987, MA, Oued Boufekrane; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Ametrasse (Ametrasse, 820 m), Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Rheocricotopus (Psilocricotopus) meridionalis Moubayed-Breil, 2016

Kettani and Moubayed-Breil 2018, Rif

Rheocricotopus (Psilocricotopus) tirolus Lehmann, 1969

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Azzouzi et al. 1992, HA, Oued Tensift; Dakki 1997; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Rheocricotopus (Rheocricotopus) effusus (Walker, 1856)

Reiss 1977; Naya 1988, MA, Haut et Moyen Sebou; Fekhaoui et al. 1993; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Tamaridine (Zaouiet et Habtiyiène, 819 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Rheocricotopus (Rheocricotopus) fuscipes (Kieffer, 1909)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani et al. 2010, Rif, Maggou (905 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Rheocricotopus (Rheocricotopus) rifensis Moubayed & Kettani, 2019

Moubayed-Breil and Kettani, Rif, Chrafate, Challal Sghir (Akchour)

Synorthocladius Thienemann, 1935

Synorthocladius semivirens (Kieffer, 1909)

Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Azzouzi et al. 1992, HA, Oued Tensift; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Smittia Holmgren, 1869

Smittia alpicola Goetghebuer, 1941

Kettani and Moubayed-Breil 2018, Rif

Smittia aterrima Meigen, 1818

Kettani and Moubayed-Breil 2018, Rif

Smittia contingens Walker, 1856

Kettani and Moubayed-Breil 2018, Rif

Smittia foliacea (Kieffer, 1921)

Kettani and Moubayed-Breil 2018, Rif

Smittia pratorum Goetghebuer, 1927

Kettani and Moubayed-Breil 2018, Rif

Thienemannia Kieffer, 1909

Thienemannia cf. fulvofasciata (Kieffer, 1921)

Kettani and Moubayed-Breil 2018, Rif

Thienemannia gracilis Kieffer, 1909

Kettani and Moubayed-Breil 2018, Rif

Thienemanniella Kieffer, 1911

Thienemanniella acuticornis (Kieffer, 1912)

Fittkau and Reiss 1978; Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m), Oued Talembote (320 m); Kettani and Langton 2011, Rif, Oued Hamma, Oued Ketama, Oued Sgara; Kettani and Langton 2012; Ashe and O’Connor 2012

Thienemanniella clavicornis (Kieffer, 1911)

Kettani and Moubayed-Breil 2018, Rif

Thienemanniella majuscula (Edwards, 1924)

Kettani et al. 1995, Rif, aval Oued El Kbir; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Thienemanniella vittata (Edwards, 1924)

Kettani et al. 1996, Rif, Haut Maggou; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Haut Maggou (1300 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Thienemanniella Pe 2a Langton 1991

Kettani et al. 2010, Rif, Oued Maggou (905 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote; Kettani and Langton 2012

Thienemanniella Pe 2b Langton 1991

Kettani et al. 2010, Rif, Oued Maggou (905 m), Oued Talembote; Kettani and Langton 2012

Trissocladius Kieffer, 1908

Trissocladius brevipalpis Kieffer in Kieffer & Thienemann 1908

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Tvetenia Kieffer, 1922

Tvetenia bavarica (Goetghebuer, 1934)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Tvetenia calvescens (Edwards, 1929)

Naya 1988, MA, Moyen Sebou; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Tamaridine (Zaouiet et Habtiyiène, 819 m), Oued Talembote (245 m), Oued Laou (Afertane, 56 m); Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Tvetenia discoloripes (Goetghebuer & Thienemann in Thienemann, 1936)

Kettani and Langton 2011, Rif, Oued Nakhla, Bouztate (Fifi); Kettani and Langton 2012; Ashe and O’Connor 2012

Tvetenia verralli (Edwards, 1929)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta; Kettani et al. 1995, Rif, aval Oued El Kbir, amont Oued Nakhla; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, ruisselet maison forestière Talassemtane (1683 m), Oued Tamaridine (Zaouiet et Habtiyiène, 819 m), Oued Talembote (245 m), Oued Laou (Afertane, 56 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Zalutschia Lipina, 1939

Zalutschia humphriesiae Dowling & Murray, 1980

Kettani and Langton 2011, Rif, marais de Lemtahane (PNPB), Dayat Fifi; Kettani and Langton 2012; Ashe and O’Connor 2012

Chironominae

Chironomini

Chironomus Meigen, 1803

Chironomus (Baeotendipes) noctivagus (Kieffer, 1911)

Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) annularius Meigen, 1818

Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) aprilinus sensu Meigen, 1818

= Chironomus halophilus Kieffer, in Ramdani and Tourenq 1982: 180, in Naya 1988: 50

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Naya 1988, MA, Haut Sebou; Fekhaoui et al. 1993; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) bernensis Klötzli, 1973

= Chironomus sp 1 Kettani 1994

Kettani et al. 1994; Kettani et al. 1996; Kettani et al. 2001; Dakki 1997; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) calipterus Kieffer, 1908

Reiss 1977, AP, Larache; Fittkau and Reiss 1978; Ramdani and Tourenq 1982 AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) longistylus Goetghebuer, 1921

Kettani et al. 2011, Rif, Oued Ketama; Kettani and Langton 2012

Chironomus (Chironomus) luridus Strenzke, 1959

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Kettani et al. 2001; Kettani and Langton 2011, Rif, merja Mtalssi (Tamuda, 31 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) nuditarsis Keyl, 1961

Kettani et al. 2011, Rif, Oued Boujdad (Kitane, 42 m), Oued El Hatba (SIBE Jebel Moussa, 165 m); Kettani and Langton 2012, Rif, SIBE Jebel Moussa

Chironomus (Chironomus) piger (Strenzke, 1956)

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) plumosus (Linnaeus, 1758)

Reiss 1977, AP, Larache, AA, Dra-Tal; Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987; Naya 1988, MA, Sidi Abdellah, Dar Cheih Harazem, Dar El Arsa; Fekhaoui et al. 1993; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani et al. 2010, Rif, Aïn Tissmelal (Tissmelal, 1046 m); Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) prasinus sensu Pinder, 1978

Kettani et al. 2011, Rif, merja Mtalssi (Tamuda, 31 m); Kettani and Langton 2012

Chironomus (Chironomus) riparius Meigen, 1804

= Chironomus thummi Kieffer, in Naya 1988: 51, Fekhaoui et al. 1993: 26

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, Oued Sebou; Naya 1988, MA, Moyen Sebou; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Barrage Talembote; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) salinarius Kieffer, 1915

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Kettani et al. 2001; Kettani and Langton 2011, Rif, merja Mtalssi (Tamuda, 31 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chironomus (Chironomus) tentans Fabricius, 1805

= Camptochironomus tentans Fabricius, 1805, in Naya 1988: 50

Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Kettani and Langton 2012

Cladopelma Kieffer, 1921

Cladopelma virescens (Meigen, 1818)

Kettani and Moubayed-Breil 2018, Rif

Cryptochironomus Kieffer, 1918

Cryptochironomus (Cryptochironomus) albofasciatus (Staeger, 1839)

= Cryptochironomus obreptans Walker, 1856, in Kettani 1994: 28

Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Kettani et al. 2001; Dakki 1997; Kettani and Langton 2012

Cryptochironomus (Cryptochironomus) psittacinus (Meigen, 1830)

Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Kettani and Langton 2012

Cryptochironomus (Cryptochironomus) rostratus Kieffer, 1921

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Fès, Oued Sebou, oued Oum-er-Rbia, Oued Boufekrane, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Cryptochironomus (Cryptochironomus) supplicans (Meigen, 1830)

Kettani and Moubayed-Breil 2018, Rif

Cryptochironomus Pe 5 Langton 1991

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and Langton 2012

Demicryptochironomus Lenz, 1941

Demicryptochironomus (Demicryptochironomus) vulneratus (Zetterstedt, 1838)

Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani et al. 2010, Rif, Nord Maggou village (Maggou, 905 m); Kettani and Langton 2012

Demicryptochironomus (Irmakia) neglectus Reiss, 1988

Kettani and Moubayed-Breil 2018, Rif

Demicryptochironomus (Irmakia) Pe 1 Langton 1991

Kettani et al. 1995, Rif, aval Oued El Kbir, Oued El Kbir, aval Oued Khemis; Kettani et al. 2001; Kettani and Langton 2012

Dicrotendipes Kieffer, 1913

Dicrotendipes collarti (Goetghebuer, 1936)

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Kettani et al. 2001; Kettani and Langton 2012

Dicrotendipes cordatus Kieffer, 1922

Kettani et al. 1996, Rif, Oued Khizana (Oued Laou); Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and Langton 2012

Dicrotendipes fusconotatus (Kieffer, 1922)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Barrage Talembote; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Dicrotendipes modestus (Say, 1823)

Kettani et al. 2011, Rif, merja Mtalssi (Tamuda, 31 m); Kettani and Langton 2012

Dicrotendipes nervosus (Staeger, 1839)

= Limnochirononomus nervosus Staeger, in Naya 1988: 53

Naya 1988, MA, Moyen Sebou (Sidi Abdellah); Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Dicrotendipes notatus (Meigen, 1818)

Kettani and Moubayed-Breil 2018, Rif

Dicrotendipes pallidicornis (Goetghebuer, 1934)

Azzouzi and Laville 1987, Rif, Retenue El Makhazine, MA, Oued Boufekrane; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Dicrotendipes peringueyanus Kieffer, 1924

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Barrage Talembote; Kettani et al. 1995, Rif, aval Oued Krikra, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012

Dicrotendipes septemmaculatus (Becker, 1908)

= Dicrotendipes pilosimanus Kieffer, in Reiss 1977: 91, Azzouzi and Laville 1987: 219

Reiss 1977, AP, Larache; Fittkau and Reiss 1978; Azzouzi and Laville 1987, AP, Larache; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Barrage Talembote, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued Krikra, amont Oued Nakhla, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012

Endochironomus Kieffer, 1918

Endochironomus albipennis (Meigen, 1830)

Naya 1988, MA, Haut Sebou (Skhounata); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Endochironomus tendens (Fabricius, 1775)

Naya 1988, MA, Moyen Sebou (Gantra Mdez, Azzaba); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Glyptotendipes Kieffer, 1913

Glyptotendipes (Caulochironomus) viridis (Macquart, 1834)

Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Glyptotendipes (Glyptotendipes) cauliginellus (Kieffer, 1913)

= Glyptotendipes gripekoveni (Kieffer)

Naya 1988, MA, Haut Sebout (Guigou); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Glyptotendipes (Glyptotendipes) pallens (Meigen, 1804)

Azzouzi and Laville 1987, Rif, Retenue El Makhazine; Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Glyptotendipes sp A Langton 1991

Naya 1988, MA, Oued Sebou; Kettani et al. 2001; Kettani and Langton 2012

Glyptotendipes sp B Langton 1991

Naya 1988, MA, Oued Sebou; Kettani et al. 2001; Kettani and Langton 2012

Harnischia Kieffer, 1921

Harnischia curtilamellata (Malloch, 1915)

Azzouzi and Laville 1987, MA, Oued Fès, Oued Sebou; Kettani et al. 1994, Rif, Oued Siflaou, Oued Afertane, aval Oued Laou; Kettani et al. 1995, Rif, amont Oued Nakhla, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Harnischia fuscimanus Kieffer, 1921

Azzouzi and Laville 1987, Rif, Retenue El Makhazine, MA, Oued Boufekrane; Kettani et al. 1995, Rif, Oued El Kbir, amont Oued Nakhla, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Kiefferulus Goetghebuer, 1922

Kiefferulus (Kiefferulus) tendipediformis (Goetghebuer, 1921)

Reiss 1977, Rif, Tétouan; Fittkau and Reiss 1978; Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 2010, Rif, Guelta 1 km après Amariguen (Jebel Setsou, 1280 m); Kettani et al. 2001; Kettani and Langton 2011, Rif, Dayat Dalia (SIBE Jebel Moussa, 169 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Kloosia Kruseman, 1933

Kloosia pusilla (Linnaeus, 1767)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Lauterborniella Thienemann & Bause, 1913

Lauterborniella agrayloides (Kieffer, 1911)

Naya 1988, MA, Haut Sebou; Kettani et al. 2001; Kettani and Langton 2012

Microchironomus Kieffer, 1918

Microchironomus deribae (Freeman, 1957)

= Leptochirononomus deribae Freeman, in Reiss 1977: 91, Ramdani and Tourenq 1982: 180

Reiss 1977, AP, Rabat; Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 2001; Kettani and Langton 2012

Microchironomus lendli (Kieffer, 1918)

Reiss 1986, AA, Oasis Meski; Azzouzi and Laville 1987; Kettani et al. 2001; Kettani and Langton 2012

Microchironomus tener (Kieffer, 1918)

Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Azzouzi et al. 1992, HA, Oued Tensift, Barrage Lalla Takerkoust; Kettani and Langton 2012

Microtendipes Kieffer, 1915

Microtendipes britteni (Edwards, 1929)

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012

Microtendipes chloris (Meigen, 1818)

Kettani et al. 2011, Rif, Dayat En-Nâsser (Khandek En-Nâsser, 1177 m), source Bab Karn (Fifi, 1216 m), Dayat Fifi (1179 m); Kettani and Langton 2012

Microtendipes confinis (Meigen, 1830)

Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Azzouzi et al. 1992, HA, Oued Tensift; Kettani and Langton 2012

Microtendipes diffinis (Edwards, 1929)

Reiss 1977, AA, Dra-Tal; Fittkau and Reiss 1978; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 2001; Kettani et al. 2011, Rif, Dayat En-Nâsser (Khandek En-Nâsser, 1177 m), Dayat Aïn Rami, source Bab Karn (Fifi, 1216 m); Kettani and Langton 2012

Microtendipes pedellus (De Geer, 1776)

Reiss 1977, Rif, Environ de Tétouan; Fittkau and Reiss 1978; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, Rif, Tétouan, HA; Naya 1988, MA, Haut Sebou (amont Aîn Tadout, Skhounate, Arhbalou Aberchane); Ashe and Cranston 1990; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Nubensia Spies, 2015

Nubensia nubens (Edwards, 1929)

= Polypedilum nubens (Edwards, 1929), in Azzouzi and Laville 1987: 219; Kettani et al. 1994: 28, 1995: 257, 1996: 137, 1997: 184, 2001: 331, 2010: 70; Dakki 1997: 65; Dakki et al. 2008: 32, Kettani and Langton 2012: 423

Azzouzi and Laville 1987, MA, Oued Sebou; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, Oued Afertane, aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Laou (Ifansa, 105 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Parachironomus Lenz, 1921

Parachironomus frequens (Johannsen, 1905)

Kettani et al. 1995, Rif, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012

Parachironomus parilis (Walker, 1856)

Reiss 1977, AP, Environ de Larache; Azzouzi and Laville 1987, MA, Lac Aguelmane Azigza; Ashe and Cranston 1990; Kettani et al. 1995, Rif, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Paracladopelma Harnisch, 1923

Paracladopelma camptolabis (Kieffer, 1913)

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Barrage Talembote; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paracladopelma galaptera (Townes, 1945)

Azzouzi et al. 1992, HA, Ouarzazate (1140 m), Gorges de Todra (1400 m); Kettani et al. 2001; Kettani and Langton 2012

Paracladopelma graminicolor (Kieffer, 1925)

= Cryptotendipes graminicolor (Kieffer), in Azzouzi et al. 1992: 230

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Paracladopelma laminatum (Kieffer, 1921)

Reiss 1977, AA, Dra-Tal; Fittkau and Reiss 1978; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paracladopelma mikianum (Goetghebuer, 1937)

Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Azzouzi et al. 1992, HA, Oued Tensift; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paralauterborniella Lenz, 1941

Paralauterborniella nigrohalteralis (Malloch, 1915)

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, Oued Sebou; Kettani et al. 2001; Kettani and Langton 2012

Paratendipes Kieffer, 1911

Paratendipes albimanus (Meigen, 1818)

Naya 1988, MA, Moyen Sebou (Mdez); Kettani et al. 1994, Rif, aval Barrage Talembote; Kettani et al. 1995, Rif, aval Oued Krikra, Oued Mhajrat, aval Oued Khemis, Oued Laou (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paratendipes nudisquama (Edwards, 1929)

Kettani and Moubayed-Breil 2018, Rif

Paratendipes striatus (Kieffer, 1925)

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Kettani et al. 2001; Kettani and Langton 2012

Phaenopsectra Kieffer, 1921

Phaenopsectra flavipes (Meigen, 1818)

Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Polypedilum Kieffer, 1912

Polypedilum (Pentapedilum) ruandae Freeman, 1955

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Kettani et al. 2001; Kettani and Langton 2012

Polypedilum (Pentapedilum) sordens (van der Wulp, 1875)

= Polypedilum sp 1, in Kettani et al. 1994: 28

Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Polypedilum (Pentapedilum) uncinatum (Goetghebuer, 1921)

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès; Kettani et al. 2001; Kettani and Langton 2012

Polypedilum (Polypedilum) albicorne (Meigen, 1838)

Naya 1988, MA, Haut Sebou; Kettani et al. 1995, Rif, aval Oued Krikra, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Polypedilum (Polypedilum) arundineti (Goetghebuer, 1921)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012

Polypedilum (Polypedilum) laetum (Meigen, 1818)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Polypedilum (Polypedilum) nubeculosum (Meigen, 1804)

Reiss 1977, Rif, Environ de Tétouan; Fittkau and Reiss 1978; Azzouzi and Laville 1987, Rif, Environ Tétouan, MA, Oued Sebou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Ashe and Cranston 1990; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Polypedilum (Polypedilum) nubifer (Skuse, 1889)

= Polypedilum pharao Kieffer, in Reiss 1977: 91, Naya 1998: 55, Ramdani and Tourenq 1982: 180

Kügler and Wool 1968; Reiss 1977, AP, Larache, Rabat; Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987, AP, Environ de Larache, Rabat, Merja Sidi Boughaba; Naya 1988, MA, Haut Sebou; Ashe and Cranston 1990; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Polypedilum (Polypedilum) pedestre (Meigen, 1830)

Reiss 1977; Azzouzi and Laville 1987, MA, Oued Boufekrane; Kettani et al. 1994, Rif, aval Barrage Talembote; Kettani et al. 1995, Rif, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani et al. 2005, Rif, Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Polypedilum (Tripodura) acifer Townes, 1945

Reiss 1977, AA, Dra-Tal; Azzouzi and Laville 1987, MA, Oued Boufekroune, Oued Fès, Oued Sebou; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012

Polypedilum (Tripodura) aegyptium Kieffer, 1925

= Polypedilum pruina Freeman, in Reiss 1977: 91

Reiss 1977, AP, Larache, HA, Marrakech, AA, Dra-Tal; Reiss 1985; Azzouzi and Laville 1987, AP, Larache, HA, Marrakech, AA, Gorges de Todra; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Oued Talembote (usine éléctrique), Oued Afertane; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Kettani et al. 2010, Rif, Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012

Polypedilum (Tripodura) bicrenatum Kieffer, 1921

Azzouzi and Laville 1987, MA, Oued Sebou; Kettani et al. 2001; Kettani and Langton 2012

Polypedilum (Tripodura) pullum (Zetterstedt, 1838)

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Oum-er-Rbia, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Polypedilum (Tripodura) quadriguttatum Kieffer, 1921

Naya 1988, MA, Moyen Sebou; Kettani et al. 1995, Rif, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Polypedilum (Tripodura) scalaenum (Schrank, 1803)

Reiss 1977, AA, Dra-Tal; Fittkau and Reiss 1978; Azzouzi and Laville 1987; Kettani et al. 1996, Rif, Ras el Ma (Chefchaouen); Ashe and Cranston 1990; Kettani et al. 2001; Kettani and Langton 2012

Polypedilum (Tripodura) tetracrenatum Hirvenoja, 1962

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Polypedilum (Tripodura) tridens Freeman, 1955

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Kettani et al. 2001; Kettani and Langton 2012

Polypedilum (Uresipedilum) convictum (Walker, 1856)

Reiss 1977, AP, Environ de Larache; Fittkau and Reiss 1978; Azzouzi and Laville 1987, MA, Oued Boufekrane (Gantra Mdez), Naya 1988, MA, Haut Sebou; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued pont Béni M’Hamed (Béni M’Hamed, 1330 m), Oued Talembote (avant village Talembote, 320 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Polypedilum (Uresipedilum) cultellatum Goetghebuer, 1931

Fittkau and Reiss 1978; Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Kettani and Langton 2012

Polypedilum ontario-group sp. 1

Kettani et al. 1995, Rif, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012

Rheomus Laville & Reiss, 1988

Rheomus alatus Laville & Reiss, 1988

Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Rheomus yahiae Laville & Reiss, 1988

Azzouzi and Laville 1987, MA, Oued Fès; Kettani et al. 2001; Kettani and Langton 2012

Stenochironomus Kieffer, 1919

Stenochironomus gibbus Fabricius, 1794

Kettani and Moubayed-Breil 2018, Rif

Stictochironomus Kieffer, 1919

Stictochironomus caffrarius (Kieffer, 1921)

Reiss 1977; Azzouzi and Laville 1987; Kettani et al. 2001; Kettani and Langton 2012

Stictochironomus maculipennis (Meigen, 1818)

Azzouzi and Laville 1987, MA, Oued Sebou; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Afertane; Kettani et al. 1995, Rif, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Laou (Afertane, 55 m); Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Stictochironomus pictulus (Meigen, 1830)

Reiss 1977, AP, Environ de Larache; Fittkau and Reiss 1978; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou; Kettani et al. 1995, Rif, aval Oued Kbir, aval Oued Krikra, Oued El Kbir; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, Oued Nakhla; Kettani and Langton 2012

Stictochironomus rosenschoeldi Zetterstedt, 1838

Kettani and Moubayed-Breil 2018, Rif

Stictochironomus reissi Cranston, 1989

= Stictochironomus sp. nov. Reiss, in Reiss 1977: 91

Reiss 1977; Azzouzi and Laville 1987, AA, M’Hamid, Dra-Tal; Kettani et al. 2001; Kettani and Langton 2012

Stictochironomus sticticus (Fabricius, 1781)

= Stictochironomus histrio (Fabricius, 1794), in Kettani et al. 1996: 138

Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2011, Rif, Oued Berranda (Bouztate, 1259 m), Dayat Dalia (SIBE Jebel Moussa); Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012

Stictochironomus Pe 2 Langton 1991

Kettani et al. 2001

Xenochironomus Kieffer, 1921

Xenochironomus xenolabis (Kieffer, 1916)

Azzouzi and Laville 1987, MA, Oued Fès; Kettani et al. 2001; Kettani and Langton 2012

Tanytarsini

Cladotanytarsus Kieffer, 1921

Cladotanytarsus (Cladotanytarsus) atridorsum Kieffer, 1924

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Azzouzi et al. 1992, HA, Aït Saoun, Gorges de Dadès (1900 m), vallée de Drâa, Marrakech; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Barrage Talembote, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, amont Oued Nakhla, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Cladotanytarsus (Cladotanytarsus) capensis (Freeman, 1954)

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Kettani et al. 2001; Kettani and Langton 2012

Cladotanytarsus (Cladotanytarsus) ecristatus Reiss, 1991

= Tanytarsus sp. nov. (Morokko) Reiss, in Azzouzi and Laville 1987: 219

Reiss 1977, AA, Dra-Tal; Azzouzi and Laville 1987, EM, Berkane; Reiss 1991; Azzouzi et al. 1992, HA; Kettani et al. 2001; Kettani and Langton 2012

Cladotanytarsus (Cladotanytarsus) mancus (Walker, 1856)

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Cladotanytarsus (Cladotanytarsus) pallidus Kieffer, 1922

= Cladotanytarsus Pe 5 Langton 1984

Azzouzi and Laville 1987, MA, Oued Sebou, Oum Rbia; Kettani et al. 2001; Kettani and Langton 2012

Cladotanytarsus (Cladotanytarsus) vanderwulpi (Edwards, 1929)

Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued Mhajrat, Oued Martil (Tamuda); Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Lithotanytarsus Thienemann, 1933

Lithotanytarsus dadesi Reiss, 1991

Reiss 1991; Azzouzi et al. 1992, HA, Gorges de Dadès (Imdiazen, 1900 m); Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique), Oued Afertane; Kettani et al. 1995, Rif, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Tassikeste (Afechtal, 240 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Lithotanytarsus emarginatus (Goetghebuer, 1933)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani and Langton 2012

Micropsectra Kieffer, 1909

Micropsectra andalusiaca Marcuzzi, 1950

Kettani and Moubayed-Breil 2018, Rif

Micropsectra apposita (Walker, 1856)

= Micropsectra contracta Reiss, 1965

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Chrafat (Armotah, 900 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Micropsectra aristata Pinder, 1976

Kettani and Langton 2012, Rif, Oued Zarka

Micropsectra atrofasciata (Kieffer, 1911)

= Micropsectra bidentata (Goetghebuer, 1921), in Azzouzi et al. 1992: 230; Kettani et al. 2001: 332; Kettani and Langton 2011: 590, 2012: 424

Fittkau and Reiss 1978; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Sebou (Arhbalou Aberchane), Oued Oum-er-Rbia; Naya 1988, MA, Haut Sebou; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Oued Talembote (usine éléctrique); Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Madissouka (Talassemtane, 1530 m), Oued Chrafat (Armotah, 900 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (aval affluent Talembote, 155 m); Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2011, Rif, Oued Taida (Moulay Abdelsalam, 650 m), cascade Zarka, Dayat En-Nâsser (Khandek En-Nâsser, 1177 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Micropsectra junci (Meigen, 1818)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Micropsectra lacustris Säwedal, 1975

Kettani and Langton 2012, Rif, Oued Zarka

Micropsectra lindrothi Goetghebuer, 1931

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Micropsectra notescens (Walker, 1856)

Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2011, Rif, Oued Ketama, Oued Sgara, ruisselet Bab Tariouant, Oued Berranda (Bouztate, 1259 m), source Bab Karn (Fifi, 1220 m), Dayat Fifi (Fifi, 1179); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Micropsectra pallidula (Meigen, 1830)

Kettani and Moubayed-Breil 2018, Rif

Micropsectra schrankelae Stur & Ekrem, 2006

Kettani and Moubayed-Breil 2018, Rif

Micropsectra zernyi Marcuzzi, 1950

Kettani and Moubayed-Breil 2018, Rif

Paratanytarsus Thienemann & Bause, 1913

Paratanytarsus bituberculatus (Edwards, 1929)

Azzouzi et al. 1992, MA, Lac Aguelmane Azigza (1510 m); Kettani et al. 1995, Rif, Oued Martil (Tamuda); Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and Langton 2012

Paratanytarsus dissimilis (Johannsen, 1905)

= Paratanytarsus confusus Palmén, 1960, in Naya 1988: 40; Dakki et al. 2008: 32; Kettani et al. 2001: 332; Kettani and Langton 2012: 423

Naya 1988, MA, Haut Sebou; Dakki et al. 2008, MA, Oued Sebou; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paratanytarsus grimmii (Schneider, 1885)

Kettani et al. 2010, Rif, Oued Laou (Afertane, 55 m); Kettani and Langton 2012

Paratanytarsus inopertus (Walker, 1856)

Reiss 1977, Rif, Environ Tétouan; Fittkau and Reiss 1978; Reiss and Säwedal 1981; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 2001; Kettani and Langton 2011, Rif, merja Mtalssi (Tamuda, 31 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paratanytarsus mediterraneus Reiss & Säwedal, 1981

Reiss and Säwedal 1981, Rif, Estuaire Oued Mharka (Tanger), AP, Oued Loukous; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 2001; Kettani and Langton 2011, AP, marais de Loukous; Kettani and Langton 2012

Paratanytarsus tenellulus (Goetghebuer, 1921)

= Microspsectra tenellula Reiss 1977: 91; Azzouzi and Laville 1987: 219

Reiss 1977, MA, Lac Kranichsee; Fittkau and Reiss 1978; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 2001; Kettani and Langton 2012

Paratanytarsus tenuis (Meigen, 1830)

= Tanytarsus tenuis Meigen, in Naya 1988: 57

Naya 1988, MA, Moyen Sebou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Rheotanytarsus Thienemann & Bause, 1913

Rheotanytarsus ceratophylli Dejoux, 1973

Naya 1988, MA, Moyen et Bas Sebou; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Rheotanytarsus curtistylus (Goetghebuer, 1921)

Azzouzi et al. 1992, HA, Oasis Meski (1160 m); Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheotanytarsus langtoni Moubayed & Kettani, 2018

Moubayed-Breil and Kettani 2018, Rif, Oued Farda; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Rheotanytarsus muscicola Thienemann, 1929

Reiss 1977, AP, Environ de Larache, AA, Dra-Tal (Tissint Moyen Dra); Fittkau and Reiss 1978; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheotanytarsus nigricauda Fittkau, 1960

Kettani and Moubayed-Breil 2018, Rif

Rheotanytarsus pellucidus (Walker, 1818)

= Rheotanytarsus distinctissimus (Brundin, 1947), in Kettani et al. 1995: 258; Kettani et al. 1996: 138, 1997: 185; Kettani and Langton 2012: 424

Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheotanytarsus pentapoda (Kieffer, 1909)

= Rheotanytarsus sp 1, in Kettani et al. 1994: 28

Kettani et al. 1994, Rif, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote; Kettani et al. 1995, Rif, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Talembote (avant village Talembote, 320 m), Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Rheotanytarsus photophilus (Goetghebuer, 1921)

Naya 1988, MA, Haut Sebou; Kettani et al. 2001; Kettani and Langton 2012

Rheotanytarsus procerus Reiss, 1991

Reiss 1991, HA; Azzouzi et al. 1992, HA, Gorges de Dadès (Imdiazen, 1900 m); Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheotanytarsus reissi Lehmann, 1970

Lehmann, 1970; Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Tassikeste (Afechtal, 240 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheotanytarsus rhenanus Klink, 1983

Kettani and Moubayed-Breil 2018, Rif

Rheotanytarsus ringei Lehmann, 1970

Lehmann, 1970; Reiss 1977, Rif, Environ Tétouan; Fittkau and Reiss 1978; Azzouzi and Laville 1987, Rif, Tétouan, MA, Oued Boufekrane, Oued Fès, Oued Sebou, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheotanytarsus Pe 3 Langton 1991

Kettani et al. 2010; Kettani and Langton 2011, Rif, Oued Sgara (Ketama, 1300 m); Kettani and Langton 2012

Stempellina Thienemann & Bause, 1913

Stempellina almi Brundin, 1947

Fittkau and Reiss 1978; Azzouzi and Laville 1987, MA, Oued Boufekrane; Kettani et al. 2001; Kettani and Langton 2012

Stempellina bausei (Kieffer, 1911)

Kettani and Langton 2012, Rif, Ketama; Kettani and Moubayed-Breil 2018, Rif

Stempellinella Brundin, 1947

Stempellinella brevis (Edwards, 1929)

Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012

Tanytarsus van der Wulp, 1874

Tanytarsus brundini Lindeberg, 1963

Kettani et al. 1994, Rif, Oued Moulay Bouchta, aval Oued Laou; Kettani et al. 1995, Rif, amont Oued Nakhla, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Dakki 1997; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Tanytarsus chinyensis Goetghebuer, 1934

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2011, Rif, Dayat Fifi (Fifi, 1179 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Tanytarsus cretensis Reiss, 1987

= Tanytarsus sp. nov. (creticus), in Reiss 1977: 91; Azzouzi and Laville 1987: 219

= Cladotanytarsus sp 1, in Kettani et al. 1995: 258

Reiss and Fittkau 1971; Reiss 1977, EM, Environ de Berkane; Reiss 1987; Azzouzi and Laville 1987, Rif, Tétouan, AP, Larache, Kénitra; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012

Tanytarsus dibranchius Kieffer, 1926

= Tanytarsus separabilis Brundin, 1947, in Kettani et al. 1994: 29; Kettani et al. 1995: 258, 1996: 138, 2001: 332; Dakki 1997: 63; Kettani and Langton 2012: 424

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Barrage Talembote, aval Oued Talembote (usine éléctrique); Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and Langton 2012

Tanytarsus ejuncidus (Walker, 1856)

Kettani and Moubayed-Breil 2018, Rif

Tanytarsus eminulus (Walker, 1856)

Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Tanytarsus formosanus Kieffer, 1912

= Tanytarsus horni Goetghebuer, 1934, in Reiss and Fittkau 1971: 122; Reiss 1977: 91; Fittkau and Reiss 1978: 439; Ramdani and Tourenq 1982: 180; El Mezdi and Giudicelli 1985: 292; Azzouzi and Laville 1987: 219; Ashe and Cranston 1990: 341; Kettani et al. 2001: 332; Kettani and Langton 2012: 424

Reiss and Fittkau 1971, Rif, M’Diq; Reiss 1977, Rif, Environ Tétouan, AP, Larache, Rabat, Kénitra; Fittkau and Reiss 1978; Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, HA, Oued Tensift; Ashe and Cranston 1990; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Tanytarsus gregarius Kieffer, 1909

Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012

Tanytarsus heusdensis Goetghebuer, 1923

Reiss 1977, AA, Dra-Tal; Fittkau and Reiss 1978; Azzouzi and Laville 1987; Ashe and Cranston 1990; Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m), Oued Talembote (avant village Talembote, 320 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Ifansa, 105 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Tanytarsus mendax Kieffer, 1925

Kettani and Moubayed-Breil 2018, Rif

Tanytarsus medius Reiss & Fittkau, 1971

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Tanytarsus palettaris Verneaux, 1969

Kettani et al. 1994, Rif, aval Oued Laou; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Tanytarsus pallidicornis (Walker, 1856)

Kettani and Langton 2011, Rif, Dayat Fifi (Fifi, 1179 m), Oued El Hatba (SIBE Jebel Moussa, 165 m); Kettani and Langton 2012

Tanytarsus recurvatus Brundin, 1947

Kettani and Langton 2011, Rif, Oued El Hamma (El Hamma, 240 m); Kettani and Langton 2012

Tanytarsus signatus (van der Wulp, 1859)

= Tanytarsus Pe 5 Langton 1991, in Azzouzi and Laville 1987: 219

Kügler and Reiss 1973; Reiss 1977, AA, Dra-Tal; Azzouzi and Laville 1987; Kettani et al. 2001; Kettani and Langton 2011, Rif, Dayat Aïn Rami (373 m), Dayat Amlay (258 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Tanytarsus verralli Goetghebuer, 1928

Kettani and Langton 2011, Rif, Oued Taida (650 m); Kettani and Langton 2012

Tanytarsus volgensis Miseiko, 1967

= Tanytarsus fimbriatus Reiss & Fittkau, 1971, in Fittkau and Reiss 1978: 439; Azzouzi and Laville 1987: 219; Kettani et al. 2001: 332; Kettani and Langton 2012: 424

Fittkau and Reiss 1978; Azzouzi and Laville 1987, MA, Oued Fès, Oued Sebou, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Tanytarsus Pe 14 Langton 1991

Kettani and Langton 2011, Rif, source Issaguen (Ketama, 1600 m); Kettani and Langton 2012

Tanytarsus Pe 23 Langton 1991

Kettani and Langton 2011, Rif, Oued El Hamma (El Hamma, 240 m); Kettani and Langton 2012

Virgatanytarsus Pinder, 1982

Virgatanytarsus albisutus (Santos-Abreu, 1918)

= Virgatanytarsus maroccanus Kügler and Reiss, in Azzouzi and Laville 1987: 219

Fittkau and Reiss 1978; Reiss and Schurch 1984, AA, Dra-Tal; Reiss 1986; Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia, AA, Dra-Tal; Ashe and Cranston 1990; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Virgatanytarsus ansatus Reiss & Schürch, 1984

Reiss and Schurch 1984, HA; Azzouzi and Laville 1987, MA, Lac Aguelmane Azigza; Ashe and Cranston 1990; Kettani et al. 2001; Kettani and Langton 2012

Virgatanytarsus arduennensis (Goetghebuer, 1922)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Virgatanytarsus triangularis (Goetghebuer, 1928)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Virgatanytarsus Pe 1 Langton 1991

Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani and Langton 2012

Zavrelia Kieffer, Thienemann & Bause, 1913

Zavrelia pentatoma Kieffer & Bause, 1913

Kettani and Langton 2012

Zavrelia Pe 1 Langton, 1991

Kettani and Langton 2011, Rif, Oued Berranda (Bouztate, 1259 m); Kettani and Langton 2012

Acknowledgment

We gratefully acknowledge the invaluable assistance and cooperation of Patrick Ashe (Dublin, Ireland) who contributed greatly to the revision of this family.

SIMULIIDAE

K. Kettani

Number of species: 43.

Faunistic knowledge of the family in Morocco: good

Simulinae

Prosimuliini

Helodon Enderlein, 1921

Helodon laamii (Beaucournu-Saguez and Bailly-Choumara, 1981)

Beaucournu-Saguez and Bailly-Choumara 1981, Rif; Clergue-Gazeau et al. 1991; Hervy et al. 1994; Belqat et al. 2001a; Belqat 2002; Belqat and Dakki 2004; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Prosimulium Roubaud, 1906

Prosimulium hirtipes species group

Bailly-Choumara and Beaucournu-Saguez 1981: 53–54: groupe latimucro (species nova ?); Beaucournu-Saguez and Bailly-Choumara 1981: 119: groupe latimucro, groupe tomosvaryi and groupe rufipes-hirtipes; Clergue-Gazeau et al. 1991: 54 as «sp. gr. Hirtipes»

Prosimulium latimucro (Enderlein, 1925)4

Bailly-Choumara and Beaucournu-Saguez 1981; Beaucournu-Saguez and Bailly-Choumara 1981; Giudicelli and Thiery 1985, HA; Giudicelli and Bouzidi 1989, HA; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’Goum en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Adler and Belqat 2001, Rif, Oued Iouchirene, Oued Ketama (Al Hoceima); Belqat et al. 2001a, Rif, HA; Belqat and Adler 2001, Rif, Aïn Khandek En Nâsser, Oued Iouchirene, Oued Ketama; Belqat 2002, Rif; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Koçak and Kemal 2010; Belqat et al. 2011; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Prosimulium rachiliense Djafarov, 1954 (complex)5

Beaucournu-Saguez and Bailly-Choumara 1981; Adler and Belqat 2001; Belqat and Adler 2001; Belqat 2002; Belqat and Dakki 2004; Belqat et al. 2005; Belqat et al. 2008; Belqat et al. 2011; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Prosimulium tomosvaryi (Enderlein, 1921)

Beaucournu-Saguez and Bailly-Choumara 1981; Giudicelli and Thiery 1985, HA; Giudicelli and Bouzidi 1989, Giudicelli et al. 2000; Adler and Belqat 2001, Rif, Oued Iouchirene (Al Hoceima); Belqat and Adler 2001, Rif, Oued Ouringa Tamdâ, oued Iouchirene, Oued Mrinet, Oued Ketama, Aîn Ksour, Oued Tisgris, Aîn Sidi Brahim Ben Arrif, Oued Hannacha; Belqat et al. 2001a, Rif; Belqat et al. 2001b; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Koçak and Kemal 2010; Belqat et al. 2011; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Urosimulium Contini, 1963

Urosimulium faurei (Bernard, Grenier & Bailly-Choumara, 1972)

Grenier et al. 1957, MA; Bernard et al. 1972: 63–68 (original description), MA, Plateau de Talerhza (environ de Meknès); Clergue-Gazeau et al. 1991, MA; Hervy et al. 1994; Belqat and Adler 2001, Rif, Oued Iouchirene, Oued Mrinet, Oued Biyada, Oued Hannacha, Oued Ankouda; Belqat et al. 2001a, Rif, MA; Belqat 2002, Rif, MA; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif, MA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simuliini

Greniera Doby & David, 1959

Greniera fabri Doby & David, 1959

Clergue-Gazeau et al. 1991, MA; Dakki 1997; Belqat et al. 2001a, Rif; Belqat 2002, Rif; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif, MA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Metacnephia Crosskey, 1969

Metacnephia blanci (Grenier & Théodoridès, 1953)

= Cnephia sp. in Grenier 1953: 157

= Cnephia blanci Grenier and Théodoridès, in Grenier and Théodoridès 1953: 430–435

= Eusimulium latinum Rubzov, in Benhoussa et al. 1988: 160–164

Grenier 1953, HA; Grenier and Théodoridès 1953, HA; Grenier et al. 1957, MA; Bailly-Choumara and Beaucournu-Saguez 1978, Rif; Bailly-Choumara and Beaucournu-Saguez 1981, HA; Benhoussa et al. 1988, AP, Oued Bou-Regreg; Clergue-Gazeau et al. 1991, AA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane: 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, MA, HA, AA; Belqat 2002, Rif, MA, HA, AA; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Belqat et al. 2008; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif, AP, MA, HA, AA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Metacnephia nuragica Rivosecchi, Raastad & Contini, 19756

= Cnephia tredecimatum (Edwards), in Grenier et al. 1957: 226

Grenier et al. 1957, AP, Coastal meseta (region of Rabat); Belqat et al. 2001a, AP, Rabat; Belqat 2002, AP, Rabat; Belqat and Dakki 2004, AP, Rabat; Belqat et al. 2011, AP; Belqat et al. 2018

Simulium Latreille, 1802

Simulium (Crosskeyellum) gracilipes Edwards, 1921

Edwards 1921: 143 (original description), MA; Séguy 1925: 233, MA; Séguy 1930a, MA; Grenier 1953, MA; Crosskey 1964, MA, Fès; Grenier and Bailly-Choumara 1970: 96–102 (original description of subgenus Crosskeyellum, description of gracilipes), MA; Clergue-Gazeau et al. 1991, MA; Hervy et al. 1994; Dakki 1997; Belqat et al. 2001a, MA; Belqat 2002, MA; Belqat and Dakki 2004, MA; Belqat et al. 2011, MA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Eusimulium) angustipes Edwards, 1915

Clergue-Gazeau et al. 1991, MA, HA; Dakki 1997; Belqat et al. 2001a, MA, HA; Belqat 2002, MA, HA; Belqat and Dakki 2004, MA, HA; Dakki et al. 2008, MA, O. Sebou; Koçak and Kemal 2010; Belqat et al. 2011, MA, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Eusimulium) mellah Giudicelli & Bouzidi, 2000 [in Giudicelli, Bouzidi and Abdelaali 2000]

Giudicelli et al. 2000: 63 (original description), HA, Oued Mellah (Bassin Draa); Belqat et al. 2001a, HA; Belqat 2002, MA, HA; Belqat and Dakki 2004, MA, HA; Dakki et al. 2008, MA, Oued Sebou; Koçak and Kemal 2010; Belqat et al. 2011, MA, HA; Adler et al. 2015, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Eusimulium) petricolum (Rivosecchi, 1963)

= Simulium latizonum Bailly-Choumara and Beaucournu-Saguez, in Bailly-Choumara and Beaucournu-Saguez 1978: 143–144 (misidentified); Bailly-Choumara and Beaucournu-Saguez 1981: 53–54

Bailly-Choumara and Beaucournu-Saguez 1978, Rif, HA; Bailly-Choumara and Beaucournu-Saguez 1981, HA; Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, HA; Belqat 2002, Rif, HA; Belqat and Dakki 2004, Rif, HA; Belqat et al. 2005, Rif; Dakki et al. 2008, Rif, MA, Oued Sebou; Koçak and Kemal 2010; Belqat et al. 2011, HA; Adler et al. 2015, Rif, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Eusimulium) rubzovianum (Sherban, 1961)

Adler et al. 2015, Rif, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Eusimulium) velutinum sensu stricto (Santos Abreu, 1922)

= Eusimilium latinum Rubzov, in El Mezdi and Giudicelli 1985: 292–295; Benhoussa et al. 1988: 160–164

Bailly-Choumara and Beaucournu-Saguez 1978, Rif; Bailly-Choumara and Beaucournu-Saguez 1981, HA; El Mezdi and Giudicelli 1985, HA, Khettaras of Marrakech; Benhoussa et al. 1988, AP, Oued Bou-Regreg; Clergue-Gazeau et al. 1991, AA; Benhoussa et al. 1993, AP, Oued Bou-Regreg; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, MA, HA, AA; Belqat 2002, Rif, MA, HA, AA; Belqat and Dakki 2004, Rif, MA, HA, AA; Belqat et al. 2005, Rif; Dakki et al. 2008, MA, Oued Sebou; Koçak and Kemal 2010; Belqat et al. 2011, Rif, AP, MA, HA, AA; Adler et al. 2015, Rif, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Eusimulium) velutinum cytospecies ‘5’

Adler et al. 2015, Rif, Tanger-Anjra, HA, Marrakech; Belqat et al. 2018

Simulium (Nevermannia) ruficorne species group

Simulium (Nevermannia) angustitarse (Lundström, 1911)

Belqat et al. 2001a, Rif; Belqat et al. 2001b, Rif; Belqat 2002, Rif; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Belqat et al. 2011, Rif; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Nevermannia) ibleum (Rivosecchi, 1966)

Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, HA; Belqat 2002, Rif, HA; Belqat and Dakki 2004, Rif, HA; Belqat et al. 2005, Rif; Belqat et al. 2011, Rif, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Nevermannia) lundstromi (Enderlein, 1921)

Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA; Belqat et al. 2001a, HA; Belqat 2002, HA; Belqat and Dakki 2004, HA; Belqat et al. 2011, Rif, Kanar (280 m), Majjo (905 m), 10 km before the Issaguen source (1200 m), HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Nevermannia) ruficorne Macquart, 1838

= Eusimulium ruficorne Macquart, in El Mezdi and Giudicelli 1985: 292, 294–295

Grenier et al. 1957, AA; Bailly-Choumara and Beaucournu-Saguez 1978, Rif; Bailly-Choumara and Beaucournu-Saguez 1981, HA; El Mezdi and Giudicelli 1985, HA, Khettaras of Marrakech; Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, HA, AA; Belqat 2002, Rif, HA, AA; Crosskey et al. 2002; Belqat and Dakki 2004, Rif, HA, AA; Belqat et al. 2005, Rif; Belqat et al. 2011, Rif, AP, HA, AA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Nevermannia) vernum species group

Simulium (Nevermannia) brevidens (Rubtsov, 1956)

Clergue-Gazeau et al. 1991, HA; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, HA; Belqat 2002, HA; Belqat and Dakki 2004, HA; Belqat et al. 2011, HA; Belqat et al. 2018; Adler 2019

Simulium (Nevermannia) carthusiense (Grenier & Dorier, 1959)

Giudicelli and Dakki 1984, Rif; Dakki 1997; Belqat et al. 2001a, Rif; Belqat 2002, Rif; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Belqat et al. 2008, Rif; Belqat et al. 2011, Rif; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Nevermannia) costatum Friederichs, 1920

Grenier et al. 1957, Rif, Pré-Rif, MA; Bailly-Choumara and Beaucournu-Saguez 1981, HA; Giudicelli and Bouzidi 1989, HA; Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, MA, HA; Belqat 2002, Rif, MA, HA; Belqat and Dakki 2004, Rif, MA, HA; Belqat et al. 2005, Rif; Belqat et al. 2008, Rif; Belqat et al. 2011, Rif, MA, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Nevermannia) cryophilum (Rubtsov, 1959) (complex)

= Simulium pusillum Fries, in Séguy 1930a: 52 (misidentification); Grenier 1953: 159 (after Séguy)

Séguy 1930a, HA; Grenier 1953, Rif, HA, Lac Ifni; Bouzidi and Giudicelli 1986, HA; Bouzidi and Giudicelli 1989, HA; Clergue-Gazeau et al. 1991, HA; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Giudicelli Belqat et al. 2001a, Rif, HA; Belqat 2002, Rif, HA; Belqat and Dakki 2004, Rif, HA; Belqat et al. 2005, Rif; Belqat et al. 2008, Rif; Belqat et al. 2011; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Nevermannia) toubkal (Bouzidi & Giudicelli, 1986)

Bouzidi and Giudicelli 1986: 41–52 (original description), HA, assif n’Ouarzane (Oued Nfis); Giudicelli and Bouzidi 1989, HA; Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, HA; Belqat 2002, HA; Belqat and Dakki 2004, HA; Belqat et al. 2011, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Nevermannia) vernum Macquart, 1826 (complex) [latipes authors pre-1972, not Meigen]

Clergue-Gazeau et al. 1991, HA; Dakki 1997; Belqat et al. 2001a, Rif, HA; Belqat 2002, Rif; Belqat and Dakki 2004, Rif, HA; Belqat et al. 2005, Rif; Belqat et al. 2011, Rif, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Rubzovia) knidirii (Giudicelli & Thiery, 1985)

Giudicelli and Thiery 1985: 109–123 (original description in new subgenus Simulium (Crenosimulium), HA; Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, HA; Belqat 2002, HA; Belqat and Dakki 2004, HA; Belqat et al. 2011, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Rubzovia) lamachi (Doby & David, 1960)

Giudicelli and Dakki 1984, Rif; Giudicelli and Thiery 1985, Rif; Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane: 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, HA; Belqat 2002, Rif, HA; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Belqat et al. 2011, Rif, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Simulium) bezzii species group

Simulium (Simulium) bezzii (Corti, 1914) (complex)

= Simulium atlas Séguy, 1930, in Séguy 1930a: 50 (original description); Grenier 1953: 158 (synonymy of atlas Séguy with bezzii suggested)

Séguy 1930a, MA; Grenier 1953; Grenier 1953, MA, HA; Grenier and Théodoridès 1953; Grenier et al. 1957, AA; Bailly-Choumara and Beaucournu-Saguez 1978, Rif; Bailly-Choumara and Beaucournu-Saguez 1981, HA; Bouzidi and Giudicelli 1986, HA; Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, MA, HA, AA; Belqat 2002, Rif, MA, HA, AA; Belqat and Dakki 2004, Rif, MA, HA, AA; Belqat et al. 2005, Rif; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif, MA, HA, AA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Simulium) ornatum species group

Simulium (Simulium) egregium Séguy, 1930

Grenier 1930, HA; Séguy 1930a: 51 (original description), HA; Belqat et al. 2001a, HA; Belqat 2002, HA; Belqat and Dakki 2004, HA; Belqat et al. 2011, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Simulium) intermedium Roubaud, 1906

= Simulium reptans var. fasciatum Séguy, in Séguy 1930a: 52 (misidentification)

= Simulium ornatum var. nitidifrons Edwards, in Grenier 1953: 159, Grenier and Théodoridès 1953: 441, Grenier and Faure 1957 [1956]: 840, Grenier and Bailly-Choumara 1970: 102, Bailly-Choumara and Beaucournu-Saguez 1978: 143–144

= Odagmia nitidifrons Edwards, in Giudicelli and Dakki 1984: 95, Benhoussa et al. 1988: 160–164

= Simulium nitidifrons Edwards, in El Mezdi and Giudicelli 1985: 292, 294–295

Séguy 1930a, HA; Grenier 1953, MA, HA; Grenier and Théodoridès 1953, MA; Grenier and Faure 1957 [1956], Rif, Pré-Rif, AP, S Rabat; MA, Plain of Meknès; Grenier et al. 1957, Rif, Pré-Rif, HA; Grenier and Bailly-Choumara 1970, MA; Bernard et al. 1972, MA; Bailly-Choumara and Beaucournu-Saguez 1978, Rif; Giudicelli and Dakki 1984, Rif, MA; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Benhoussa et al. 1988, AP, Oued Bou-Regreg; Giudicelli and Bouzidi 1989, HA; Clergue-Gazeau et al. 1991, MA, HA; Benhoussa et al. 1993, AP, Oued Bou-Regreg; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’oun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, MA, HA; Belqat 2002, Rif, MA, HA; Belqat and Dakki 2004, Rif, MA, HA; Belqat et al. 2005, Rif; Belqat et al. 2008, Rif; Belqat et al. 2011, Rif, AP, MA, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Simulium) ornatum Meigen, 1818 (complex)

= reptans var fasciatum, in Séguy 1930: 52

[subornatum: Séguy 1925/1930, not Edwards]

Séguy 1930a: 52 (ornatum and subornatum records), HA; Grenier 1953, HA; Bailly-Choumara and Beaucournu-Saguez 1978, Rif; Clergue-Gazeau et al. 1991, MA, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, MA, HA, AA; Belqat 2002, Rif, MA, HA, AA; Belqat and Dakki 2004, Rif, MA, HA, AA; Belqat et al. 2005, Rif; Belqat et al. 2008, Rif; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif, MA, HA, AA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Simulium) trifasciatum Curtis, 1839

Belqat et al. 2001a, Rif; 2001b, Rif; Belqat 2002, Rif; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Belqat et al. 2008, Rif; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Simulium) variegatum species group

Bailly-Choumara and Beaucournu-Saguez (1981: 52–54): groupe monticola (“sp. nova A” and “sp. nova B”)

Simulium (Simulium) atlasicum Giudicelli & Bouzidi, 1989

Giudicelli and Bouzid 1989: 146–151 (original description), HA, near village Aguelmous; Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, HA; Belqat 2002, HA; Belqat and Dakki 2004, HA; Belqat et al. 2011, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Simulium) berberum Giudicelli & Bouzidi, 1989

Giudicelli and Bouzidi 1989: 151–156 (original description), HA, assif n’Ouarzane; Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, HA; Belqat 2002, HA; Belqat and Dakki 2004, HA; Belqat et al. 2011, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Simulium) variegatum Meigen, 1818

Bailly-Choumara and Beaucournu-Saguez 1978, Rif, HA; Bailly-Choumara and Beaucournu-Saguez 1981, HA; Giudicelli and Bouzidi 1989; Clergue-Gazeau et al. 1991; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, HA; Belqat 2002, Rif, HA; Belqat and Dakki 2004, Rif, HA; Belqat et al. 2005, Rif, HA; Belqat et al. 2011, Rif, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Simulium) xanthinum Edwards, 1933

= Simulium gaudi Grenier and Faure, in Grenier and Faure 1957 [1956]: 838–840

Grenier and Faure 1957 [1956], Rif, Pré-Rif, HA; Grenier et al. 1957; Bailly-Choumara and Beaucournu-Saguez 1978, Rif; Clergue-Gazeau et al. 1991, MA; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, MA, HA; Belqat 2002, Rif, MA, HA; Carles-Tolrá 2002; Belqat and Dakki 2004, Rif, MA, HA; Belqat et al. 2005, Rif; Belqat et al. 2008, Rif; Belqat et al. 2011, Rif, MA, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Trichodagmia) albellum species group

Simulium (Trichodagmia) auricoma Meigen, 1818

= Simulium (Obuchovia) auricoma Meigen, 1818, in Belqat et al. 2011: 52

Belqat 2000, Rif; Belqat et al. 2001a, Rif; Belqat 2002, Rif; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Belqat et al. 2011, Rif; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Trichodagmia) galloprovinciale Giudicelli, 1963 [1962]

= Simulium (Obuchovia) galloprovinciale Giudicelli, 1963, in Belqat et al. 2011: 52

Belqat 2000, Rif; Belqat et al. 2001a, Rif; Belqat 2002, Rif; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Belqat et al. 2011, Rif; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Trichodagmia) marocanum Bouzidi & Giudicelli, 1988 [1987]

= Simulium (Obuchovia) marocanum Bouzidi & Giudicelli, 1987, in Belqat et al. 2011: 52

Bouzidi and Giudicelli 1987: 185–195 (original description), Rif, near village Bou Adel, HA, Oued Rdat (affluent de l’Oued Tensift); Clergue-Gazeau et al. 1991, HA; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, HA; Belqat 2002, HA; Belqat and Dakki 2004, HA; Belqat et al. 2011, Rif, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Wilhelmia) equinum species group

Simulium (Wilhelmia) equinum (Linnaeus, 1758)

= Simulium equinum Linnaeus, in Grenier et al. 1957: 231–232

Grenier et al. 1957, MA; Bailly-Choumara and Beaucournu-Saguez 1981, HA; Dakki 1997; Belqat et al. 2001a, HA; Belqat 2002, HA; Belqat and Dakki 2004, HA; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, MA, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Wilhelmia) pseudequinum Séguy, 1921

= Simulium barbaricum Séguy, in Séguy 1930a: 51

= Simulium equinum var. mediterraneum Puri, in Grenier 1953: 145–148; Grenier and Théodoridès 1953: 436

= Simulium equinum mediterraneum Puri, in Grenier and Faure 1957 [1956]: 840; Grenier et al. 1957: 232–234

= Wilhelmia pseudequinum Séguy, in Benhoussa et al. 1988: 160–164

Séguy 1930a, HA; Grenier 1953, HA; Grenier and Théodoridès 1953, HA; Grenier and Faure 1957 [1956], Rif, Pré-Rif, AP, HA, AA; Meknès; Grenier et al. 1957, HA; Bailly-Choumara and Beaucournu-Saguez 1978, Rif; Bailly-Choumara and Beaucournu-Saguez 1981, HA; Benhoussa et al. 1988, AP, Oued Bou-Regreg; Benhoussa et al. 1993, AP, Oued Bou-Regreg; Dakki 1997; Clergue-Gazeau et al. 1991, HA; Benhoussa et al. 1993, AP, Oued Bou-Regreg; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, MA, HA, AA; Belqat 2002, Rif, MA, HA, AA; Belqat and Dakki 2004, Rif, MA, HA, AA; Belqat et al. 2005, Rif; Belqat et al. 2008, Rif; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif, AP, MA, HA, AA; Adler and Crosskey 2017; Belqat et al. 2018

Simulium (Wilhelmia) quadrifila Grenier, Faure & Laurent, 1957 [1956]

Grenier et al. 1957: 238–239 (original description as form of sergenti), Rif, Pré-Rif, AP, S Casablanca, MA, Meknès, HA; Bailly-Choumara and Beaucournu-Saguez 1978, Rif, MA, HA; Clergue-Gazeau et al. 1991, Rif, Pré-Rif; Dakki 1997; Belqat et al. 2001a, Rif; Belqat 2002, Rif, AP, S Casablanca, MA, HA; Belqat and Dakki 2004, Rif; Belqat et al. 2005, Rif; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif, AP, MA, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

Simulium (Wilhelmia) sergenti (Edwards, 1923)

= Simulium ariasi Séguy, in Séguy 1925: 231–238; Séguy 1930a: 50; Grenier 1953: 144

= Simulium equinum mediterraneum Puri, in Grenier and Faure 1957 [1956]: 840; Grenier et al. 1957: 238–240

= Wilhelmia sergenti Edwards, in Benhoussa et al. 1993: 249

Séguy 1930a, MA; Grenier 1953, MA; Grenier and Théodoridès 1953, HA; Grenier and Faure 1957 [1956], Rif, Pré-Rif; Grenier et al. 1957, Rif, Pré-Rif, AP, S Casablanca, MA, HA; Bailly-Choumara and Beaucournu-Saguez 1978, Rif, AP, MA, HA; Bailly-Choumara and Beaucournu-Saguez 1981, HA; Clergue-Gazeau et al. 1991, Rif, Pré-Rif, HA; Benhoussa et al. 1993, AP, Oued Bou-Regreg; Dakki 1997; Giudicelli et al. 2000, HA, Oued Réghaya (Neltner, 3800 m), Oued Réghaya (Sidi Chamharouch, 2300 m), Oued Réghaya (lmlil, 1740 m), Oued Réghaya (Aguersioual, 1550 m), Oued Réghaya (Moulay Brahim, 1200 m), Oued Réghaya (Tahanaout, 890 m), ruisselet émissaire de source débouchant dans Oued Réghaya en amont d’lmlil (1750 m), ruisselet émissaire de source débouchant dans l’assif M’zik (1850 m), ruisselet émissaire de source débouchant dans l’assif N’Ouarzane (3000 m), ruisseau émissaire de source (assif N’Ouarzane, 3000 m), assif N’Ouarzane (Irhoulidene, 2800 m), ruisseau affluent en rive droite de l’assif N’Ouarzane (2400 m), Oued N’fis (amont Ijoukak, 1600 m), Oued N’fis (amont Wirgan, 1200 m), Oued N’fis (980 m), Oued N’fis (amont retenue Lalla Takerkoust, 660 m), ruisseau de Tinzart (émissaire de source: 2850 m), ruisseau de Tifni (émissaire de source: 2780 m), ruisseau de Likemt (émissaire de source: 2670 m), ruisseau de Tougroudadene (émissaire de source: 2660 m), assif Oukaimeden (2600 m), source hélocrène au niveau du cirque d’Oukaimeden (2660 m), assif Tiferguine (2500 m), assif Oukaimeden (2450 m), ruisseau émissaire de source débouchant dans l’assif Oukaimeden (1740 m), complexe rhéocrène formé par des émissaires de source débouchant dans l’assif Oukaimeden (1730 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1630 m), affluent temporaire en rive gauche de l’assif Oukaimeden (1360 m), affluent temporaire en rive droite de l’assif Oukaimeden (1260 m), affluent en rive droite de l’assif Oukaimeden (1300 m), assif Tarzaza (1200 m), assif Tarzaza (1000 m), cours inférieur de l’oued Ourika (850 m), Oued Rdat en amont de Taddert (1850 m), affluent temporaire en rive gauche de Oued Rdat (1400 m), Oued Rdat (1600 m), Oued Rdat (1230 m), Oued Tensift (600–700 m), khetarras (450–600 m), Oued Tessaout au niveau d’Aït Tamli (1620 m), Oued Lakdar en aval de la retenue de Sidi Driss (1030 m), ruisseau émissaire de source formant le début de l’assif Imini (2090 m), assif Imini (1560 m), Oued Ounila (1820 m), ruisseau affluent en rive gauche de l’oued Ounila (1820 m), Oued Ounila (Timhlt, 1600 m), Oued Mellah (Anghessa, 1400 m), Oued Dadès en amont des gorges (1630 m), Oued Dadès (Boumalne, 1530 m), Oued Dadès (Sidi Flah, 1100 m), Oued M’Goun (1530 m), Oued M’Goun en aval de Kelaâ (1370 m), ruisseau émissaire de source débouchant dans un affluent de l’Oued Souss (2350 m); Belqat et al. 2001a, Rif, MA, HA; Belqat 2002, Rif, MA, HA; Belqat and Dakki 2004, Rif, MA, HA; Belqat et al. 2005, Rif; Belqat et al. 2008, Rif; Dakki et al. 2008, MA, Oued Sebou; Belqat et al. 2011, Rif, AP, MA, HA; Adler and Crosskey 2017; Belqat et al. 2018; Adler 2019

THAUMALEIDAE

K. Kettani, R. Wagner

Number of species: 2. Expected: 10

Faunistic knowledge of the family in Morocco: poor

Thaumalea Ruthe, 1831

Thaumalea bernardi Vaillant, 1956

Vaillant 1956b, HA, Toubkal, Siroua, Lac Tamhda (Anremer), Sidi Chamarouch, Izourar, M’Goum, Oukaimeden; Dakki 1997

Thaumalea spinata Vaillant, 19547

Vaillant 1954a, HA, M’Goum, springs powering d’Ameskeur el Fougani, springs powering Izourar lagoon (Azourki), springs powering the lake Tamhda (Anremer), torrent at the bottom of Jebel Siroua, Oukaimeden (Toubkal), Jebel Toubkal, Atend (Sidi Chamarouch)

BLEPHARICERIDAE

K. Kettani, P. Zwick

Number of species: 4

Blepharicerinae

Liponeura Loew, 1844

Liponeura alticola Giudicelli & Bouzidi, 1987

Giudicelli and Bouzidi 1987, HA, Oued Réghaya; Dakki 1997

Liponeura megalatlantica (Vaillant, 1956)

Vaillant 1956c, HA, Izourar, Imi-N’Ifri; Giudicelli and Lavandier 1974; Dakki 1997

Liponeura rifincola Zwick, 2013

Zwick 2013, Rif, Issaguen (Ketama, 1800 m)

Liponeura sirouana (Vaillant, 1956)

= Cardiocrepsis sirouana Vaillant, in Vaillant 1956b: 234

Vaillant 1956b, HA, Siroua (3000 m); Giudicelli and Lavandier 1974; Dakki 1997

Bibionoidea

ANISOPODIDAE8

K. Kettani, J.-P. Haenni

Number of species: 3. Expected: 7–8

Faunistic knowledge of the family in Morocco: poor

Anisopodinae

Sylvicola Harris, 1780

Sylvicola fenestralis (Scopoli, 1763)9

= Rhyphus fenestralis (Scopoli, 1763), in Mouna 1998: 86

Mouna 1998 (no locality given) – MISR

Sylvicola fuscatus (Fabricius, 1775)1

= Rhyphus fuscatus (Fabricius, 1775), in Mouna 1998: 86

Mouna 1998 (no locality given) – MISR

Sylvicola punctatus (Fabricius, 1787)1

= Rhyphus punctatus (Fabricius, 1787), in Mouna 1998: 86

Mouna 1998 (no locality given) – MISR

BIBIONIDAE10

K. Kettani, J.-P. Haenni

Number of species: 10. Expected: 20

Faunistic knowledge of the family in Morocco: poor

Bibioninae

Bibio Geoffroy, 1762

Bibio hortulanus (Linnaeus, 1758)

= Bibio hortularum Linnaeus, 1758, in Becker and Stein 1913: 85

Becker and Stein 1913, Rif, Tanger; Séguy 1949a, SA, Foum Zguid

Bibio lanigerus Meigen, 1818

Ebejer et al. 2019, Rif, Amsemlil (PNPB, 1067 m)

Bibio laufferi Strobl, 1906

Ebejer et al. 2019, MA, 3.5 km S of Azrou (Ifrane, 1450 m), 20 km S of Azrou (Ifrane, 1720 m)

Bibio leucopterus (Meigen, 1804)

Ebejer et al. 2019, MA, 6 km S of Azrou (Ifrane, 1610 m)

Bibio marci (Linnaeus, 1758)

Becker and Stein 1913, Rif, Tanger; Haenni 2009, Rif, Tanger, MA, Ifrane

Dilophus Meigen, 1803

Dilophus antipedalis Wiedemann in Meigen, 1818

Ebejer et al. 2019, Rif, Oued Azla (Nwawel, 57 m), Oued Azla (Hallila, 95 m)

Dilophus (cf. bispinosus Lundström, 1914)11

Séguy 1953a, AA, Aïn Chaib (Souss)

Dilophus febrilis (Linnaeus, 1758)

= Dilophus vulgaris Meigen, 1818, in Séguy 1949a: 153

Séguy 1949a, AA, Goulimine, Foum-el-Hassan

Dilophus femoratus Meigen, 1804

= Philia femorata Meigen, 1804, in Séguy 1941: 2

Séguy 1941d, HA, Tizi-n’Test (2000 m); Pârvu et al. 2006, AP, Merja Zerga

Dilophus tridentatus Walker, 1848

Ebejer et al. 2019, AP, Sidi Mokhtar (Essaouira) – MHNN (J.-P. Haenni leg.), MNHN (SA, Foum-el-Hassan)

Sciaroidea

CECIDOMYIIDAE

K. Kettani, M. Skuhravá, V. Skuhravý

Number of species: 57. Expected: 100

Faunistic knowledge of the family in Morocco: moderate

Lestremiinae

Lestremia Macquart, 1826

Lestremia parvostylia Jaschhof, 1994

Jaschhof 1994, SA, Abeino, 15 km N Goulimine; Papp 2007; Gagné 2010; Gagné and Jaschhof 2014; Skuhravá et al. 2017

Micromyinae

Campylomyza Meigen, 1818

Campylomyza flavipes Meigen, 1818

Jaschhof 1998, HA, Telouet; Skuhravá et al. 2017

Campylomyza fusca Winnertz, 1853

Jaschhof 1998, HA, Telouet; Skuhravá et al. 2017

Campylomyza mohrigi Jaschhof, 2009

Jaschhof 2009 (south Morocco); Gagné 2010; Skuhravá et al. 2017

Monardia Kieffer, 1895

Monardia (Xylopriona) toxicodendri (Felt, 1907)

Jaschhof 1998 (South Morocco); Jaschhof 2009; Gagné 2010; Skuhravá et al. 2017

Cecidomyiinae

Asphondylia Loew, 1850

Asphondylia capparis Rübsaamen, 1893

Houard 1921, MA, Fès; Skuhravá et al. 1984; Skuhravá 1986; Skuhravá et al. 2017

Asphondylia cytisi Frauenfeld, 1873

Mimeur 1949, HA, Tanzat (1800 m), AA, Jebel Sargho, Amalou Bou Mansour (2000 m); Skuhravá et al. 2017

Asphondylia punica Marchal, 1897

= Asphondylia conglomerata De Stefani, 1900

Houard 1922, EM, Zousfana (Jebel Tagla); Mimeur 1949, AA, Agdz; Mouna 1998; Skuhravá et al. 2017

Asphondylia scrophulariae Schiner, 1856

Mimeur 1949, AP, Rabat, Arcilla; Skuhravá et al. 2017

Asphondylia verbasci (Vallot, 1827)

Mimeur 1949, AP, Maâmora, Rabat, Zaërs; Skuhravá et al. 2017

Baldratia Kieffer, 1897

Baldratia salicorniae Kieffer, 1897

Mimeur 1949, AP, Rabat, Salé, Bou-Regreg; Möhn 1966, EM, Melilla, Bocona, AP, Rabat; Skuhravá et al. 1984; Skuhravá 1986; Skuhravá et al. 2014a; Skuhravá et al. 2017

Bayeriola Gagné, 1991

Bayeriola thymicola (Kieffer, 1888)

Houard 1923, HA, Réghaya; Mimeur 1949, EM, Sidi Ali Oujda, Jebel Hamra, Itzer, MA, Ifrane, Azrou, Bordj-Doumergue, Timhadite, Aguelmane; Skuhravá et al. 1984; Skuhravá 1986; Skuhravá et al. 1993; Gagné 2010 (south of Morocco); Bruun et al. 2012; Gagné and Jaschhof 2014; Skuhravá et al. 2017

Blastomyia Kieffer, 1913

Blastomyia origani (Tavares, 1901)

Houard 1922, MA, Col de Bouchtata, Zalagh (Mouret); Skuhravá et al. 1984; Skuhravá 1986; Gagné 2010 (south of Morocco); Gagné and Jaschhof 2014; Skuhravá et al. 2017

Braueriella Kieffer, 1896

Braueriella phillyreae (Löw, 1877)

Houard 1922, Rif, Jebel Kébir; Houard 1923, AP, Cap Ghir (south of Morocco); Mimeur 1949, Rif, Zoumi, Ouezzane, EM, Béni Snassen, AP, Larache, Zaërs, Mehdia, Sehoul, MA, Jebel Said, Taza, Tahala, Tadla; Mouna 1998; Skuhravá et al. 2017 – MNHN (AP, Mehdia)

Contarinia Rondani, 1860

Contarinia ilicis Kieffer, 1898

Houard 1919, MA, Immouzer; Mimeur 1949, EM, Béni Snassen, Ras Foughal, El-Harcha, MA, Ifrane, Aït Bou-Mzil, Monts Zaian, Agoumi-n´Aït Mguild; Skuhravá et al. 2017

Contarinia luteola Tavares, 1902

Mimeur 1949, EM, El Harcha, MA, Ifrane, Djaba, Imouzzer-du-Kandar, Tafechna; Mouna 1998; Skuhravá et al. 2017

Contarinia nasturtii (Kieffer, 1888)

Mouna 1998: 85 (no accurate locality); Skuhravá et al. 2017

Contarinia pyrivora (Riley, 1886)

= Diplosis pirivora Riley, in Mouna 1998: 85

Mouna 1998: 85 (no accurate locality); Skuhravá et al. 2017

Dasineura Rondani, 1840

Dasineura affinis (Kieffer, 1886)

= Perrisia affinis (Kieffer), in Mouna 1998: 85

Mimeur 1949, Rif, Tanger, EM, Oujda, AP, Gharb, Port-Lyautey, Rabat, Fedala, Casablanca, Mazagan, Settat, Oued-Zem, Mogador, MA, Fès, Tahala, Marchand; Mouna 1998; Skuhravá et al. 2017; AP (Rabat) – MISR

Dasineura asparagi (Tavares, 1902)

Mimeur 1949, AP, Rabat, Zaërs, HA, Chaouia; Skuhravá et al. 2017

Dasineura crataegi (Winnertz, 1853)

Mimeur 1949, EM, Chaouia des Béni Snassen, AP, Rabat, Zaërs, MA, Ifrane, Tahala Aguelmane de Sidi Ali, Fès, Agoumi-n´Aït Mguild, Tarhzirt, AA, Argana, Imi-n-Tanoute; Skuhravá et al. 2017

Dasineura ericaescopariae (Dufour, 1837)

Houard 1912, Rif, Cap Spartel; Rübsaamen 1899; Skuhravá et al. 2017

Dasineura helianthemi (Hardy, 1850)

= Contarinia helianthemi (Hardy, 1850)

Mimeur 1949, AP, Gharb, Maâmora, Zaërs; Skuhravá et al. 2017

Dasineura napi (Loew, 1850)

= Dasineura brassicae (Winnertz, 1853) in Mouna 1998: 85

Mouna 1998 (no accurate locality); Skuhravá et al. 2017

Dasineura periclymeni (Rübsaamen, 1889)

Mimeur 1949, AP, Rabat, Chellah, Yquem, Grou, Korifla, EM, Berkane; Skuhravá et al. 2017

Dasineura plicatrix (Loew, 1850)

Mimeur 1949, EM, Massif des Béni Snassen, El-Harcha, AP, Larache, Korifla, Grou, Yquem, Malah, Rabat, MA, Azrou, Ifrane, Monts Zaian, Oulmés, Khénifra, HA, Tizi-Machou, AA, Bigoudine; Skuhravá et al. 2017

Dasineura rosae (Loew, 1850)

= Wachtliella rosarum (Hardy, 1850)

Mimeur 1949, MA, Ifrane, Aguelmane de Sidi Ali, Aguelmane Azigza, Zaad, Bordj-Doumergue, Bekrite, HA, Tizi-Machou; Skuhravá et al. 2017

Dicrodiplosis Kieffer, 1895

Dicrodiplosis pseudococci (Felt, 1914)

Harris 1968, AP, Rabat, Salé, HA, Asni; Skuhravá et al. 1984; Skuhravá 1986; Skuhravá et al. 2017

Dryomyia Kieffer, 1898

Dryomyia lichtensteinii (Löw, 1878)

Mimeur 1949, EM, Debdou, Ras Foughal, El-Harcha, MA, Taza, Oulmès, Azrou, Aguelmane Azigza, Ifrane, Imouzzer-du-Kandar, Tizi-n´Tretten, Ida-ou Tanane, Azilal, HA, Ayachi; Skuhravá et al. 1984; Skuhravá 1986; Oosterbroek 2007; Gagné and Jaschhof 2014; Skuhravá et al. 2017; MA (Oulmès) – MISR

Etsuhoa Inouye, 1959

Etsuhoa thuriferae Skuhravá, 1996

Mimeur 1949, HA, Sidi-Chamharouch, Aït Bou-Jafar; Skuhravá 1995; Skuhravá et al. 2017

Feltiella Rübsaamen, 1910

Feltiella acarisuga (Vallot, 1827)

Gagné 1995, AP, Rabat; Osborne et al. 2002 (productive areas of cereals); Gagné 2004; Gagné and Jaschhof 2014; Skuhravá et al. 2017

Gephyraulus Rübsaamen, 1916

Gephyraulus diplotaxis (Solinas, 1982)

Houard 1922, EM, Oasis de Figuig, Jebel Ouazzani; Mimeur 1949, AP, Vallée de l´Oued Korifla, Sidi Bouknadel; Skuhravá et al. 2017

Gephyraulus raphanistri (Kieffer, 1886)

Houard 1923, AP, Mogador; Mimeur 1949, AP, Gharb, Skhirat, Bouznika; Skuhravá et al. 2017

Houardiella Kieffer, 1912

Houardiella salicorniae Kieffer, 1912

Trotter 1904, Rif, Tingis; Houard 1912, Rif, Tanger; Mimeur 1949, AP, Loukous, Larache, Ksob, Ameur, Mogador, Rabat, Salé, Bou-Regreg, EM, Moulouya, AA, Agadir; Skuhravá et al. 2017

Iteomyia Kieffer, 1913

Iteomyia major (Kieffer, 1889)

Mimeur 1949, AP, Korifla, Grou, MA, Leghzel; Skuhravá et al. 2017

Jaapiella Rübsaamen, 1916

Jaapiella bryoniae (Bouché, 1847)

Mimeur 1949, Rif, Ouezzane, AP, Sidi Bouknadel, Mehdia, Temara, Skhirat, Pont-Blondin, Larache, Monod, Boulhaut, Safi, MA, Tahala, Fès, Khemisset, Marchand, AA, Agadir; Skuhravá et al. 2017

Lasioptera Meigen, 1818

Lasioptera berlesiana Paoli, 1907

Mimeur 1949; Skuhravá et al. 1984; Skuhravá 1986; Skuhravá et al. 2017

Lasioptera rubi (Schrank, 1803)

Mimeur 1949, AP, Tinkert, MA, Ifrane, Taza, Tarhzirt, HA, N´Fis, Oumer-Rbia, Jebel Tardema; Skuhravá et al. 2017

Lasioptera thapsiae Kieffer, 1898

= Lasioptera carophila F. Löw, in Houard 1923: 699

Houard 1921; Houard 1923, Rif, Tanger; Mimeur 1949, EM, Béni Snassen, Guercif, AP, Settat, MA, Tiddas; Mouna 1998; Skuhravá et al. 2017

Lestodiplosis Kieffer, 1894

Lestodiplosis aonidiellae Harris, 1968

Harris 1968, EM, Oujda, AP, Rabat, MA, Fès; Skuhravá et al. 2017

Mayetiola Kieffer, 1896

Mayetiola avenae (Marchal, 1895)

Mouna 1998; Lahloui et al. 2005, AP, Settat, Safi, El Jadida, MA, Béni Mellal, Khouribga, HA, Marrakech, El Kelaâ; Skuhravá et al. 2017

Mayetiola destructor (Say, 1817)

Vayssière 1920, EM, Oujda; Jourdan 1937; Hudault and Zelensky 1939; Mimeur 1949; Skuhravá et al. 1984; Skuhravá 1986; Gagné et al. 1991; Amri et al. 1992 (productive areas of cereals); El Bouhssini et al. 1992a, 1992b; Lhaloui et al. 1992; El Bouhssini et al. 1996a, 1996b; Khalifi et al. 1996; Azzam et al. 1997; El Bouhssini et al. 1997, 1999; El Bouhssini et al. 1998; Mouna 1998; Naber et al. 2000, 2003; Lhaloui et al. 2001; Lhaloui et al. 2005; Gagné 2010; Gagné and Jaschhof 2014; Skuhravá et al. 2017; AP (Rabat), MA – MISR

Mayetiola hordei Kieffer, 1909

Mouna 1998; Gagné et al. 1991; Lhaloui et al. 2005, AP, Settat, Safi, El Jadida, MA, Béni Mellal, Khouribga, HA, Marrakech, El Kelaâ; Skuhravá et al. 2005; Gagné 2010; Gagné and Jaschhof 2014; Skuhravá et al. 2017

Oligotrophus Latreille, 1805

Oligotrophus panteli Kieffer, 1898

Mimeur 1949, EM, Béni Snassen, MA, Aguelmane Azigza, Azrou, HA, Ayachi, Aït Bou-Jafar; Skuhravá et al. 2017

Oligotrophus valerii (Tavares, 1904)

= Arceuthomyia valerii (Tavares, 1904)

Mimeur 1949, HA, Ayachi, Aït Bou-Jafar; Skuhravá et al. 2017

Orseolia Kieffer & Massalongo, 1902

Orseolia cynodontis Kieffer & Massalongo, 1902

Houard 1922, Rif, Tanger, Aïn Dalia; Skuhravá et al. 1984; Skuhravá 1986; Gagné 2010; Gagné and Jaschhof 2014; Skuhravá et al. 2017

Phyllodiplosis Kieffer, 1912

Phyllodiplosis cocciferae (Tavares, 1902)

= Blastodiplosis cocciferae (Tavares, 1902)

Trotter 1904, Rif, Cap Spartel; Houard 1912; Mimeur 1949, Rif, Cap Spartel, Ouezzane, EM, Béni Snassen, El-Harcha, AP, Larache, Maâmora, Boulhaut, Zaërs, MA, Taza, Djaba, Ifrane, Imouzzer-du-Kandar, Dayet Achlaf, Dayet Ifrah, Michlifen, Bordj-Doumergue, Zaad, Bekrite, Aguelmane Azigza, Agoumi-n´Aït Mguild, Bou-Mzil, HA, Bou-Jafar, Jebel Tardema; Skuhravá et al. 1984; Skuhravá 1986; Gagné 2010; Gagné and Jaschhof 2014

Psectrosema Kieffer, 1904

Psectrosema tamaricum (Kieffer, 1912)

= Amblardiella tamaricum Kieffer, in Mouna 1998: 85

Houard 1922, EM, Zousfana, near Sidi Youssef; Mimeur 1949, Morocco Mediterranean, continental and sub-saharan, EM, Berkane, Moulouya, AP, MA, Tadla, HA, Haouz, AA, Tafilalet, SA, Agad; Skuhravá et al. 1984; Skuhravá 1986; Mouna 1998; Skuhravá et al. 2014a; Skuhravá et al. 2017

Resseliella Seitner, 1906

Resseliella oleisuga (Targioni-Tozzetti, 1887)

= Clinodiplosis oleisuga Targioni-Tozzetti, in Mouna 1998: 85

Hafraoui 1966; Skuhravá et al. 1984; Skuhravá 1986; Mouna 1998; Gagné 2010; Gagné and Jaschhof 2014; Skuhravá et al. 2017

Rhopalomyia Rübsaamen, 1892

Rhopalomyia navasi Tavares, 1904

Houard 1922, EM, Jebel Mais, Djahifa, HA, Aït Ameli; Houard 1923; Mimeur 1949, EM, Figuig, Jebel Nokra, Ifkern, Haute Moulouya, MA, Tadla; Skuhravá et al. 1993; Skuhravá et al. 2014b; Skuhravá et al. 2017

Schizomyia Kieffer, 1889

Schizomyia buboniae (Frauenfeld, 1859)

Houard 1917, EM, Djorf de Taouriet; Houard 1922, EM, Jebel Tagla, Aïn Yalon; Skuhravá et al. 2014a; Skuhravá et al. 2017

Stefaniella Kieffer, 1898

Stefaniella trinacriae De Stefani, 1900

Mimeur 1949, AP, Oualidia, Zima, Casablanca; Skuhravá et al. 2017

Stefaniola Kieffer, 1913

Stefaniola africana Möhn, 1971

Mimeur 1949, AP, Rabat, Salé, Bou-Regreg; Skuhravá et al. 2017

Stefaniola bilobata (Kieffer, 1913)

Houard 1922–1923; Mimeur 1949, MA, Tadla, HA, Ksar-es-Souk, AA, Haouz; Skuhravá et al. 1984; Skuhravá 1986; Skuhravá et al. 1993; Gagné 2010; Gagné and Jaschhof 2014; Skuhravá et al. 2017

Stefaniola opulenta Möhn, 1971

Möhn 1971; Skuhravá et al. 1984; Skuhravá 1986; Gagné 2010; Gagné and Jaschhof 2014; Skuhravá et al. 2017; Pape and Thompson 2019

Stefaniola ventriosa Möhn, 1971

Möhn 1971, MA, Oued Gheris; Skuhravá et al. 1984; Skuhravá 1986; Gagné 2010; Gagné and Jaschhof 2014; Skuhravá et al. 2014a; Skuhravá et al. 2017

Thecodiplosis Kieffer, 1895

Thecodiplosis brachyntera (Schwägrichen, 1835)

Mouna 1998: 85 (no accurate locality); Skuhravá et al. 2017

KEROPLATIDAE

K. Kettani, P.J. Chandler

Number of species: 2. Expected: 50

Faunistic knowledge of the family in Morocco: poor

Keroplatinae

Keroplatus Bosc, 1792

Keroplatus reaumurii (Dufour, 1839)

Matile 1986; Chandler et al. 2005; Evenhuis 2006

Macrocera Meigen, 1803

Macrocera fasciata Meigen, 1804

Becker and Stein 1913, Rif, Tanger; Chandler and Ribeiro 1995; Evenhuis 2006

MYCETOPHILIDAE

K. Kettani, P.J. Chandler

Number of species: 64. Expected: 250

Faunistic knowledge of the family in Morocco: poor

Mycetophilinae

Exechiini

Allodiopsis Tuomikoski, 1966

Allodiopsis rustica Edwards, 1941

Banamar et al. 2020, Rif, Dayat Tazia

Anatella Winnertz, 1864

Anatella concava Plassmann, 1990

Banamar et al. 2020, Rif, Oued Aârate

Brevicornu Marshall, 1896

Brevicornu intermedium (Santos Abréu, 1920)

Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, Oued Tisgris, Oued Aârate, Dayat Tazia, Oued Maggou (Maggou Village), Douar Tizga, maison forestière de Talassemtane, Dayat Jebel Zemzem, Aïn Takhninjoute, Forêt Adrou, Dayat Amsemlil

Brevicornu griseicolle Staeger, 1840

Banamar et al. 2020, Rif, Dayat Fifi, Oued Aârate, Dayat Tazia, Oued Maggou (Maggou Village), Aïn El Malaâb, maison forestière de Talassemtane, Douar Tizga, Aïn Takhninjoute, Dayat Amsemlil, Dayat Lemtahane, Dayat avant Taida

Brevicornu sericoma (Meigen, 1830)

Banamar et al. 2020, Rif, Chefchaouen, Dayat Fifi, Forêt Jebel Lakraâ, Dayat Fifi, Oued Maggou (Maggou Village), maison forestière de Talassemtane, Douar Tizga, Oued Amsemlil, Grotte d’Hercule, Forêt Adrou, Cascade Chrafate, Oued Aârate, Bab el Karn, Dayat Amsemlil – MNHN (coll. J. Beaucournu)

Brevicornu verralli (Edwards, 1925)

Banamar et al. 2020, Rif, Dayat Afersiw, Forêt Jebel Lakraâ

Cordyla Meigen, 1803

Cordyla crassicornis Meigen, 1818

Chandler and Ribeiro 1995; Banamar et al. 2020, Rif, Douar Abou Boubnar, maison forestière de Talassemtane, Forêt Adrou, Oued Sidi Yahia Aârab, EM, Oued Tafoughalt; Rif (Chefchaouen) – MNHN

Cordyla insons Laštovka & Matile, 1974

Banamar et al. 2020, Rif, Oued Maggou

Cordyla murina Winnertz, 1864

Banamar et al. 2020, Rif, Forêt Aïn Boughaba

Cordyla styliforceps (Bukowski, 1934)

Banamar et al. 2020, Rif, maison forestière de Talassemtane, Oued Tkaraâ, Oued Sidi Yahia Aârab

Exechia Winnertz, 1863

Exechia bicincta (Staeger, 1840)

Banamar et al. 2020, Rif, Oued Kelaâ, Oued Aârate, EM, Grotte des Pigeons, Oued Tafoughalt

Exechia dorsalis (Staeger, 1840)

Banamar et al. 2020, Rif, Bab el Karn

Exechia fulva Santos Abreu, 1920

= Rymosia exornata Séguy, in Séguy 1941a: 26

Séguy 1941a, HA, Toubkal; Chandler and Ribeiro 1995; Banamar et al. 2020, Rif, Oued Kelaâ, Forêt Aïn Boughaba, Forêt Jebel Lakraâ, Dayat Fifi, Aïn Sidi Brahim Ben Arrif, Oued Maggou (Maggou Village), Aïn Takhninjoute, maison forestière de Talassemtane, Oued Amsemlil, Dayat Jebel Zemzem, Aïn Takhninjoute, Bab el Karn, Dayat Fifi, Dayat Amsemlil; Rif (Chefchaouen, coll. J. Beaucournu) – MNHN; Rif (20 km west of Targuist, coll. A.M. Hutson) – NHMUK

Exechia fusca (Meigen, 1804)

Banamar et al. 2020, Rif, Oued Kelaâ, Forêt Jebel Lakraâ, Douar Kitane, Dayat Amsemlil

Exechiopsis Tuomikoski, 1966

Exechiopsis coremura (Edwards, 1928)

Banamar et al. 2020, Rif, Cascade Chrafate

Pseudexechia Tuomikoski, 1966

Pseudexechia tuomikoskii (Kjærandsen, 2009)

Banamar et al. 2020, Rif, Source Aheramen

Rymosia Winnertz, 1864

Rymosia affinis Winnertz, 1864

Banamar et al. 2020, Rif, maison forestière de Talassemtane, Dayat Amsemlil, Aïn Tiouila, Dayat Fifi

Rymosia beaucournui Matile, 1963

Chandler 1994; Chandler and Ribeiro 1995; Chandler et al. 2005; Banamar et al. 2020, EM, Grotte des Pigeons; AP (Oued y Kern, coll. H. Choumara) – MNHN

Rymosia pseudocretensis Burghele-Balacesco, 1966

Chandler 1994; Chandler et al. 2005; Banamar et al. 2020; AP (Oued y Kern, coll. H. Choumara) – MNHN

Stigmatomeria Tuomikoski, 1966

Stigmatomeria crassicornis (Stannius, 1831)

Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, Dayat Fifi, Aïn Takhninjoute, maison forestière de Talassemtane, Dayat Amsemlil; Rif (Chefchaouen, coll. J. Beaucournu) – MNHN

Tarnania Tuomikoski, 1966

Tarnania dziedzickii (Edwards, 1941)

Banamar et al. 2020, Rif, maison forestière de Talassemtane, Dayat Amsemlil, Cascade Chrafate, Dayat Amsemlil, Grotte Aïn El-Aouda – MNHN

Mycetophilini

Mycetophila Meigen, 1804

Mycetophila alea Laffoon, 1965

Banamar et al. 2020, Rif, Aïn el Ma Bared, maison forestière de Talassemtane, Oued Aârate, Bab el Karn; Rif (Chefchaouen, coll. J. Beaucournu) – MNHN

Mycetophila britannica Laštovka & Kidd, 1975

Chandler and Ribeiro 1995; Banamar et al. 2020, Rif, Forêt Aïn Boughaba, Forêt Jebel Lakraâ, Douar Kitane, Aïn El Malaâb, oued à 15 km de Fifi, Aïn El Ma Bared, maison forestière de Talassemtane, Oued Maggou (Maggou Village), Aïn Takhninjoute, Grotte d’Hercule, Oued Aârate, Bab el Karn, Dayat Amsemlil, Dayat Lemtahane, Dayat avant Taida, Forêt Taghzout – MNHN

Mycetophila deflexa Chandler, 2001

Banamar et al. 2020, Rif, Forêt Taghzout, route Ksar el Kebir–Chefchaouen

Mycetophila edwardsi Lundström, 1913

Banamar et al. 2020, Rif, Dayat Fifi, Dayat Tazia, Oued Tkarae, Forêt Jebel Lakraâ, Forêt Taghzout, Grotte d’Hercule, EM, Béni Snassen

Mycetophila formosa Lundström, 1911

Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, Oued Amsemlil, Dayat Amsemlil, Dayat avant Taida

Mycetophila marginata Winnertz, 1864

Banamar et al. 2020, Rif, Aïn Ras el Ma, Oued Maggou (Maggou Village), maison forestière de Talassemtane, Dayat Amsemlil; Rif (Chefchaouen, coll. J. Beaucournu) – MNHN

Mycetophila perpallida Chandler, 1993

Banamar et al. 2020, Rif, Forêt Aïn Boughaba, Aïn El Malaâb, maison forestière de Talassemtane, Bab el Karn, Dayat Amsemlil, Douar Kitane

Mycetophila pictula Meigen, 1830

Chandler and Ribeiro 1995; Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, Oued Maggou (Maggou Village); Rif (Chefchaouen, coll. J. Beaucournu) – MNHN

Mycetophila sordida van der Wulp, 1874

Chandler 1994; Banamar et al. 2020, Rif, Forêt Jebel Lakraâ; Oued Maggou (Maggou Village); MA (Khenolap-el-Ouaer, 1580 m, coll. J. Beaucournu) – MNHN

Mycetophila spectabilis Winnertz, 1864

Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, Forêt Aïn Boughaba, oued à 15 km de Fifi, Aïn El Malaâb

Mycetophila strigatoides (Landrock, 1927)

Banamar et al. 2020, Rif, Forêt Taghzout, Oued Aârate

Mycetophila unicolor Stannius, 1831

Banamar et al. 2020, Rif, Oued Kelaâ

Mycetophila vittipes Zetterstedt, 1852

Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, maison forestière de Talassemtane, Dayat Amsemlil

Phronia Winnertz, 1863

Phronia biarcuata (Becker, 1908)

Chandler 1994; Chandler and Ribeiro 1995; Chandler et al. 2005; Banamar et al. 2020, Rif, Dayat Tazia, Aïn el Ma Bared, Aïn El Malaâb, Aïn Takhninjoute, maison forestière de Talassemtane, Grotte d’Hercule, Dayat Amsemlil; Rif (Chefchaouen, coll. J. Beaucournu) – MNHN

Phronia cinerascens Winnertz, 1864

Banamar et al. 2020, Rif, Dayat Amsemlil

Phronia nitidiventris (van der Wulp, 1858)

Banamar et al. 2020, Rif, Dayat Afersiw, Oued Aârate

Phronia tenuis Winnertz, 1864

Banamar et al. 2020, Rif, Oued Kelaâ, Oued Aârate, Oued Maggou (Maggou Village), Aïn Takhninjoute, Grotte d’Hercule

Phronia tyrrhenica Edwards, 1928

Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, maison forestière de Talassemtane, Dayat Amsemlil, Aïn Takhninjoute

Phronia willistoni Dziedzicki, 1889

Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, Oued Aârate, maison forestière de Talassemtane, Dayat Amsemlil, Cascade Chrafate, Bab el Karn

Sceptonia Winnertz, 1864

Sceptonia intestata Plassmann & Schacht, 1990

Banamar et al. 2020, Rif, maison forestière de Talassemtane, Aïn El Malaâb

Sceptonia membranacea Edwards, 1925

Banamar et al. 2020, Rif, oued à 15 km de Fifi, Oued Aârate

Trichonta Winnertz, 1864

Trichonta foeda Loew, 1869

Banamar et al. 2020, Rif, Aïn Takhninjoute, Bab el Karn, Dayat Amsemlil, Oued Tkarae

Trichonta icenica Edwards, 1925

Banamar et al. 2020, EM, Grotte du Chameau

Trichonta vitta (Meigen, 1830)

Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, Bab el Karn

Trichonta vulcani Dziedzicki, 1889

Banamar et al. 2020, EM, Grotte du Chameau

Zygomyia Winnertz, 1864

Zygomyia humeralis (Wiedemann, 1817)

Banamar et al. 2020, Rif, maison forestière de Talassemtane

Zygomyia valida Winnertz, 1864

Banamar et al. 2020, Rif, Aïn Ras el Ma, Oued Aârate

Leiinae

Docosia Winnertz, 1864

Docosia gilvipes (Walker, 1856)

Banamar et al. 2020, Rif, Forêt Jebel Lakraâ, maison forestière de Talassemtane, Aïn Takhninjoute, Dayat Amsemlil

Leia Meigen, 1818

Leia arsona Hutson, 1978

Banamar et al. 2020, Rif, Oued Maggou (Maggou village), Oued Sidi Yahia Aârab, MA, Aïn Walili

Leia beckeri Landrock, 1940

Banamar et al. 2020, Rif, Aïn Ras el Ma

Leia bimaculata (Meigen, 1804)

Chandler 1994; Chandler et al. 2005; Banamar et al. 2020, Rif, Dayat Tazia, maison forestière de Talassemtane, Aïn el Ma Bared, MA, Forêt 3.5 km S Azrou; MA (Forêt Ifrane, coll. P.N. Lawrence) – NHMUK

Novakia Strobl, 1893

Novakia scatopsiformis Strobl, 1893

Banamar et al. 2020, Rif, maison forestière de Talassemtane

Novakia simillima Strobl, 1910

Banamar et al. 2020, Rif, Oued Aârate, maison forestière de Talassemtane

Gnoristinae

Boletina Staeger, 1840

Boletina gripha Dziedzicki, 1885

Banamar et al. 2020, Rif, Dayat Fifi, Oued Aârate, Aïn Sidi Brahim Ben Arrif, Dayat Amsemlil, Grotte d’Hercule, Cascade Chrafate, Oued Sidi Yahia Aârab, Aïn el Ma Bared, Bab el Karn, MA, forêt 3.5 km S Azrou

Coelosia Winnertz, 1864

Coelosia fusca Bezzi, 1892

Banamar et al. 2020, Rif, Dayat Fifi, Oued Kelaâ, Oued Amsemlil, Dayat Amsemlil, Aïn Takhninjoute, Cascade Chrafate, Bab el Karn, Dayat avant Taida

Synapha Meigen, 1818

Synapha fasciata Meigen, 1818

Banamar et al. 2020, Rif, Aïn Sidi Brahim Ben Arrif, Dayat Tazia, Forêt Adrou, Dayat Amsemlil, Dayat avant Taida

Synapha vitripennis (Meigen, 1818)

Banamar et al. 2020, Rif, Dayat Amsemlil

Tetragoneura Winnertz, 1846

Tetragoneura ambigua Grzegorzek, 1885

Banamar et al. 2020, Rif, Forêt Aïn Boughaba, EM, Oued Tafoughalt

Mycomyinae

Mycomya Rondani, 1856

Mycomya flavicollis (Zetterstedt, 1852)

Banamar et al. 2020, Rif, Aïn El Malaâb, maison forestière de Talassemtane

Mycomya pygmalion Väisänen, 1984

Banamar et al. 2020, Rif, Oued Amsemlil, Aïn Sidi Brahim Ben Arrif

Mycomya tumida (Winnertz, 1864)

Banamar et al. 2020, Rif, Dayat Fifi

Sciophilinae

Azana Walker, 1856

Azana anomala Staeger, 1840

Banamar et al. 2020, Rif, Oued Maggou (Maggou village), maison forestière de Talassemtane

Sciophila Meigen, 1818

Sciophila iberolutea Chandler & Blasco-Zumeta, 2001

Chandler and Gatt 2000, AP, Oued y Kern; Chandler and Blasco-Zumeta 2001, AP, Oued y Kern; Bechev and Koç 2006; Banamar et al. 2020, Rif, maison forestière de Talassemtane, Dayat Jebel Zemzem, Oued Sidi Yahia Aârab, Bab el Karn, Marabout el Khaloua; AP (Oued y Kern, coll. H. Choumara) – MNHN

SCIARIDAE

K. Kettani, K. Heller

Number of species: 70. Expected: 200–500

Faunistic knowledge of the family in Morocco: poor

Austrosciara Schmitz & Mjöberg, 1924

Austrosciara hyalipennis (Meigen, 1804)

El Ouazzani et al. 2019, Rif, Douar El Hamma, MA, Lac Ouiouane

Bradysia Winnertz, 1867

Bradysia alpicola (Winnertz, 1867)

El Ouazzani et al. 2019, MA, Lac Ouiouane

Bradysia bulbigera Mohrig & Kauschke, 1994

El Ouazzani et al. 2019, Rif, Oued Ouara, Oued Ametrasse, Merzouk Bni Salah, Dayat Bayn widane, HA, Ouirgane

Bradysia cavernicola Mohrig & Eckert, 1999

Menzel and Heller 2004, HA, Ouirgane

Bradysia cinerascens (Grzegorzek, 1884)

El Ouazzani et al. 2019, Rif, Issaguen, Anissar (PNPB)

Bradysia crinita Mohrig, 1992

El Ouazzani et al. 2019, Rif, Issaguen, Douar El Hamma

Bradysia fenestralis (Zetterstedt, 1838)

El Ouazzani et al. 2019, Rif, Forêt R’milat

Bradysia fenestrata (Meigen, 1818)

El Ouazzani et al. 2019, Rif, Jebel Zemzem, Oued Tkarâa, Ben Karrich, Dayat Tazia, Perdicaris Park, Tourbière Amsemlil, Dayat Tazia

Bradysia flavipila Tuomikoski, 1960

El Ouazzani et al. 2019, HA, Ouirgane

Bradysia iberiana Rudzinski & Baumjohann, 2009

El Ouazzani et al. 2019, HA, Ouirgane

Bradysia lembkei Mohrig & Menzel, 1990

El Ouazzani et al. 2019, Rif, Oued Maggou, Dayat Tazia, AP, Forêt Maâmora, HA, Ouirgane, Gerifodene

Bradysia lucichaeta Mohrig & Krivosheina, 1989

Mohrig et al. 1997, AA, Sidi Rbat (40 km S Agadir)

Bradysia mediterranea Mohrig & Menzel, 1992

Mohrig and Menzel 1992, HA

Bradysia nigrispina Menzel, 2006

El Ouazzani et al. 2019, HA, Gerifodene

Bradysia pectoralis (Staeger, 1840)

El Ouazzani et al. 2019, MA, Lac Ouiouane, HA, Ouirgane

Bradysia placida (Winnertz, 1867)

El Ouazzani et al. 2019, HA, Ouirgane

Bradysia promissa Mohrig & Röschmann, 1999

El Ouazzani et al. 2019, Rif, Beni Barou, Anissar (PNPB), Oued Tkarâa (PNPB), Taida, Marabout Moulay Abdelsalam

Bradysia reflexa Tuomikoski, 1960

El Ouazzani et al. 2019, HA, Ouirgane

Bradysia regularis (Lengersdorf, 1934)

El Ouazzani et al. 2019, Rif, Talassemtane (maison forestière), HA, Ouirgane

Bradysia ruginosa Mohrig, 1994

Mohrig et al. 1997, SA, Ablino (15 km N Goulimine); El Ouazzani et al. 2019, Rif, Jebel Lakraâ, Aïn El Fakir, HA, Amizmiz

Bradysia santorina Mohrig & Menzel, 1992

Mohrig et al. 1997, AA, Sidi Rbat (40 km S Agadir)

Bradysia scabricornis Tuomikoski, 1960

El Ouazzani et al. 2019, Rif, Oued Ouara, Maggou, Oued Azla, Douar El Hamma, HA, Ouirgane, Setti Fatma, Oued Imlil

Bradysia subrufescens Mohrig & Krivosheina, 1989

El Ouazzani et al. 2019, HA, Ouirgane

Bradysia subsantorina Mohrig & Kauschke, 1997

El Ouazzani et al. 2019, HA, Ouirgane

Bradysia tilicola (Loew, 1850)

El Ouazzani et al. 2019, Rif, Douar Tissouka, MA, Lac Ouiouane, HA, Ouirgane

Bradysia transitata Rudzinski & Baumjohann, 2013

El Ouazzani et al. 2019, Rif, Oued Ez-Zarka, Oued Tkarâa (PNPB), Oued Laou, AP, Larache (Strawberry farm), Forêt Maâmora, AA, Barrage Aoulouz, Assif Tifnout

Bradysia trivittata (Staeger, 1840)

Mohrig and Röschmann 1993, HA, AA, Sidi Rbat (40 km S Agadir); El Ouazzani et al. 2019, Rif, Aïn Tayattine, Oued Ez-Zarka, Oued Bayine, Beni Barou, Douar El Hamma, Douar Mouarâa, Tétouan, MA, Lac Ouiouane, HA, Télouet, Ouirgane

Bradysia vagans (Winnertz, 1868)

misidentified as Bradysia rufescens (Zetterstedt, 1852) in Röschmann and Mohrig 1993: 111

Röschmann and Mohrig 1993, HA, Talouete; El Ouazzani et al. 2019, Rif, Aïn Fouara, HA, Ouirgane

Bradysia xenoreflexa Mohrig & Menzel, 1993

El Ouazzani et al. 2019, AP, Forêt Maâmora

Bradysiopsis Tuomikoski, 1960

Bradysiopsis vittata (Meigen, 1830)

El Ouazzani et al. 2019, HA, Setti Fatma

Camptochaeta Hippa & Vilkamaa, 1994

Camptochaeta jeskei (Mohrig & Röschmann, 1993)

= Corynoptera jeskei Mohrig and Röschmann, in Röschmann and Mohrig 1993: 109

Röschmann and Mohrig 1993, HA, Talouete (1800 m)

Corynoptera Winnertz, 1867

Corynoptera andalusica Hippa, Vilkamaa & Heller, 2010

El Ouazzani et al. 2019, MA, Lac Ouiouane, HA, Ouirgane

Corynoptera bicuspidata (Lengersdorf, 1926)

Hippa et al. 2010, HA, Ouirgane, Lac Ouiouane

Corynoptera bispinulosa Mohrig & Dimitrova, 1992

El Ouazzani et al. 2019, EM, Tafoughalt

Corynoptera caesula Hippa & Menzel, 2004

El Ouazzani et al. 2019, Rif, Aïn Kchour, AP, Forêt Maâmora

Corynoptera cincinnata Mohrig & Blasco-Zumeta, 1996

Hippa et al. 2010, HA, Ouirgane

Corynoptera dentiforceps (Bukowski & Lengersdorf, 1936)

El Ouazzani et al. 2019, HA, Ouirgane

Corynoptera deserta Heller & Menzel, 2006

El Ouazzani et al. 2019, HA, Ouirgane

Corynoptera fatigans (Johannsen, 1912)

= Corynoptera perpusilla Winnertz, 1867, in Mohrig et al. 1997: 384

Mohrig et al. 1997, HA, Anezal; Hippa et al. 2010 [for nomenclature see Mohrig et al. 2013]

Corynoptera gemina (Hippa & Vilkamaa, 1994)

El Ouazzani et al. 2019, HA, Ouirgane

Corynoptera globiformis (Frey, 1945)

El Ouazzani et al. 2019, Rif, Talassemtane (maison forestière)

Corynoptera hemiacantha Mohrig & Mamaev, 1992

El Ouazzani et al. 2019, HA, Ouirgane

Corynoptera iberica Hippa, Vilkamaa & Heller, 2010

El Ouazzani et al. 2019, AP, Forêt Maâmora, Sidi Boughaba

Corynoptera inclinata Hippa, Vilkamaa & Heller, 2010

Hippa et al. 2010, HA, Ouirgane

Corynoptera irmgardis (Lengersdorf, 1930)

Hippa et al. 2010, HA, Ouirgane

Corynoptera postglobiformis Mohrig, 1993

El Ouazzani et al. 2019, Rif, Talassemtane (maison forestière), HA, Ouirgane

Corynoptera praeparvula Mohrig & Krivosheina, 1983

El Ouazzani et al. 2019, HA, Ouirgane, Amizmiz, Gerifodene

Corynoptera saccata Tuomikoski, 1960

Hippa et al. 2010, SA, Goulimine; Mohrig et al. 2012

Corynoptera semipedestris Mohrig & Blasco-Zumeta, 1996

Mohrig et al. 1997, SA, Ablino (15 km N Goulimine)

Corynoptera spiciceps Hippa, Vilkamaa & Heller, 2010

Hippa et al. 2010, HA, Ouirgane

Corynoptera stipidaria Mohrig, 1994

Hippa et al. 2010, HA, Ouirgane

Corynoptera subcavipes Menzel & Smith, 2007

El Ouazzani et al. 2019, Rif, Douar El Hamma, Talassemtane (maison forestière)

Corynoptera subparvula Tuomikoski, 1960

El Ouazzani et al. 2019, HA, Ouirgane

Epidapus Haliday, 1851

Epidapus atomarius (De Geer, 1778)

El Ouazzani et al. 2019, Rif, Aïn el Ma Bared (Fifi)

Leptosciarella Tuomikoski, 1960

Leptosciarella dives (Johannsen, 1912)

Mohrig et al. 2012, HA, Ouirgane

Leptosciarella parcepilosa (Strobl, 1900)

El Ouazzani et al. 2019, AP, Sidi Boughaba

Leptosciarella subviatica Mohrig & Menzel, 1997

El Ouazzani et al. 2019, HA, Ouirgane

Leptosciarella tomentosa (Mohrig & Kauschke, 1994)

El Ouazzani et al. 2019, AP, Forêt Maâmora

Lycoriella Frey, 1942

Lycoriella agraria (Felt, 1898)

El Ouazzani et al. 2019, Rif, Douar Tissouka

Lycoriella sativae (Johannsen, 1912)

El Ouazzani et al. 2019, Rif, M’Diq, Oued Zaouya, AP, Larache (strawberry farm), HA, Ouirgane

Pseudolycoriella Menzel & Mohrig, 1998

Pseudolycoriella morenae (Strobl, 1900)

El Ouazzani et al. 2019, Rif, Perdicaris Park, Bab Tariouant, Oued Maggou, EM, Zegzel, HA, Ouirgane

Scatopsciara Edwards, 1927

Scatopsciara (Scatopsciara) atomaria (Zetterstedt, 1851)

= Scatopsciara vivida (Winnertz, 1867), in Röschmann and Mohrig 1993: 111

Röschmann and Mohrig 1993, HA, Talouete; El Ouazzani et al. 2019, Rif, Oued Ez-Zarka, Oued Tkarâa (PNPB), Oued Guallet, Marécage Lemtahane (PNPB), Oued Ametrasse, Issaguen, Talassemtane (maison forestière), Douar El Hamma, Aïn Fouara, Oued Souk El Had, Merzouk Bni Salah, Oued Maggou, AP, Sidi Boughaba, HA, Assif Tifnout, Gerifodene, Armed, Amzmiz, Lac Tislit, AA, Barrage Mokhtar Soussi

Scatopsciara (Scatopsciara) maroccoensis Mohrig & Jaschhof, 1997

Mohrig et al. 1997, AA, Sidi Rbat (40 km S Agadir)

Scatopsiara (Scatopsciara) nana (Winnertz, 1871)

El Ouazzani et al. 2019, Rif, Oued Ez-Zarka, Oued Maâmala, Oued Aârkob, Ben Karrich, Merja Sidi Lhaj Merzouk, Tétouan, HA, Ouirgane, Anafgou

Scatopsciara (Scatopsciara) vitripennis (Meigen, 1818)

El Ouazzani et al. 2019, Rif, Oued Ouarra, Oued Tkarâa (PNPB), Oued Aoudour, Oued Ametrasse, Oued Aârkob, Oued Boumarouil, Ben Karrich, Dayat Tazia, Aïn El Fakir, Azib de Khmis Mdik, Merzouk Bni Salah, Oued Souk El Had, Oued Maggou, El Malâab (Talassemtane), AP, Sidi Boughaba, MA, Lac Ouiouane, HA, Ouirgane, AA, Barrage Aoulouz, Assif Tifnout, Barrage Mokhtar Soussi

Scatopsciara (Xenopygina) curvilinea (Lengersdorf, 1934)

El Ouazzani et al. 2019, AP, Sidi Boughaba, HA, Aïn Taferaout, Amzmiz, AA, Assif Tifnout, Barrage Mokhtar Soussi

Scatopsciara (Xenopygina) subarmata Mohrig & Mamaev, 1983

El Ouazzani et al. 2019, Rif, Oued Amsa, AP, Larache, MA, Mont Habri, HA, Ouirgane, AA, Id Aissa, Tissint

Schwenckfeldina Frey, 1942

Schwenckfeldina carbonaria (Meigen, 1830)

= Sciara carbonaria Meigen, in Séguy 1941d: 2

Séguy 1941d, HA, Tizi-n’Test (2000 m)

Sciara Meigen, 1803

Sciara flavimana Zetterstedt, 1851

El Ouazzani et al. 2019, Rif, Douar El Hamma

Sciara hemerobioides (Scopoli, 1763)

= Lycoria (Sciara) thomae Linnaeus, in Becker and Stein 1913: 85

Becker and Stein 1913, Rif, Tanger

Suborder BRACHYCERA

Stratiomyoidea

STRATIOMYIDAE

K. Kettani, N. Woodley

Number of species: 40. Expected: 50–60

Faunistic knowledge of the family in Morocco: moderate

Beridinae

Beris Latreille, 1802

Beris rozkosnyi Kassebeer, 1996

Kassebeer 1996; Woodley 2001, MA, Meknès, Ifrane; Kehlmaier 2004; Yimlahi et al. 2017

Chorisops Rondani, 1856

Chorisops tunisiae (Becker, 1915)

Haenni 1990, Rif, Tanger; Woodley 2001; Kehlmaier 2004; Mason et al. 2006; Koçak and Kemal 2010; Yimlahi et al. 2017; Lebard et al. 2020

Clitellariinae

Pycnomalla Gerstaecker, 1857

Pycnomalla aterrima Sack, 1912

Séguy 1930a, MA, Tizi-s’Tkrine (1700 m); Séguy 1953a, MA, Dayat Aoua; Woodley 2001; Yimlahi et al. 2017

Pycnomalla auriflua (Erichson, 1841)

Séguy 1930a, MA, Soufouloud (1900–2100 m), Boulhaut; Duisit 1960, AP, Cap Cantin; Woodley 2001

Pycnomalla splendens (Fabricius, 1787)

Séguy 1930a; Séguy 1953a, MA, Dayat Aoua; Duisit 1960, AP, forest of Maâmora, Casablanca, Cap Cantin; Rozkošný 1983, AP, Cap Cantin; Woodley 2001; Kehlmaier 2004; Koçak and Kemal 2010; Yimlahi et al. 2017

Nemotelinae

Lasiopa Brullé, 1832

Lasiopa benoisti Séguy, 1930

Séguy 1930a, MA, Meknès; Duisit 1960, EM, Aïn Guettara (Middle Moulouya); Woodley 2001; Koçak and Kemal 2010; Yimlahi et al. 2017

Lasiopa pantherina Séguy, 1930

Séguy 1930a, EM, Maharidja; Woodley 2001; Yimlahi et al. 2017

Nemotelus Geoffroy, 1762

Nemotelus (Camptopelta) nigrinus Fallén, 1817

Duisit 1960, AP, Khatouat (S Rabat); Dakki 1997; Woodley 2001; Yimlahi et al. 2017

Nemotelus (Nemotelus) atriceps Loew, 1856

Yimlahi et al. 2017, AA, village Massa

Nemotelus (Nemotelus) cingulatus Dufour, 1852

Rozkošný 1977, AP, Skhirat; Rozkošný 1983, Rif, Tanger; Woodley 2001; Yimlahi et al. 2017, Rif, Dayat Afrate, Oued Koub

Nemotelus (Nemotelus) cylindricornis Rozkošný, 1977

Faucheux 2009, AP, Oualidia

Nemotelus (Nemotelus) danielssoni Mason, 1989

Yimlahi et al. 2017, Rif, Oued Izelfane (Beni Boufrah)

Nemotelus (Nemotelus) latiusculus Loew, 1871

Lindner 1949; Rozkošný 1977; Rozkošný 1983, Rif, Tanger; Woodley 2001; Kehlmaier 2004; Yimlahi et al. 2017, Rif, Barrage Moulay Bouchta

Nemotelus (Nemotelus) longirostris (Wiedemann, 1824)

Becker and Stein 1912, 1913, Rif, Tanger; Séguy 1930a, Rif, Tanger; Duisit 1960, AP, Mechra-bel-Ksiri (Gharb), EM, Saïdia, Rozkošný 1977; Dakki 1997; Woodley 2001; Yimlahi et al. 2017

Nemotelus (Nemotelus) maculiventris Bigot, 1861

Yimlahi et al. 2017, Rif, Oued Zandoula

Nemotelus (Nemotelus) nigrifrons Loew, 1846

Linder 1936; Becker and Stein 1912, Rif, Tanger; Rozkošný 1977, Rif, Tanger; Woodley 2001; Yimlahi et al. 2017, Rif, affluent Tarmast (NPH)

Nemotelus (Nemotelus) pantherinus (Linnaeus, 1758)

Séguy 1930a, Rif, Tanger; Duisit 1960, AP, Zëar; Rozkošný 1977; Dakki 1997; Woodley 2001; Koçak and Kemal 2010; Üstüner and Hasbenli 2013; Yimlahi et al. 2017

Nemotelus (Nemotelus) proboscideus Loew, 1846

Linder 1936; Yimlahi et al. 2017

Nemotelus (Nemotelus) subuliginosus Rozkošný, 1974

Rozkošný 1977; Woodley 2001, Rif, Tanger; Yimlahi et al. 2017

Nemotelus (Nemotelus) ventralis Meigen, 1830

Woodley 2001, AP, Essaouira; Yimlahi et al. 2017

Nemotelus (Nemotelus) uliginosus (Linnaeus, 1767)

Duisit 1960, AP, Dradek

Pachygastrinae

Pachygaster Meigen, 1803

Pachygaster atra Panzer, 1798

Yimlahi et al. 2017, Rif, Dayat Mezine

Pachygaster maura Lindner, 1939

Lindner 1939; Woodley 2001, MA, Tagzirt; Yimlahi et al. 2017

Sarginae

Chloromyia Duncan, 1837

Chloromyia formosa (Scopoli, 1763)

Becker and Stein 1912, 1913, Rif, Tanger; Séguy 1930a, HA, M’Rassine; Duisit 1960, AP, Rabat, Korifla, Khatouat; Woodley 2001; Pârvu et al. 2006, AP, Merja Zerga; Pârvu and Zaharia 2007; Popescu-Mirceni 2011; Yimlahi et al. 2017, Rif, Taghbalout, Lac Ametrasse, Douar Kitane

Stratiomyinae

Oxycerini

Oxycera Meigen, 1803

Oxycera germanica (Szilády, 1932)

= Hermione dorieri var. barbarica, in Vaillant 1956b: 232, 237, 242

Vaillant 1956b, HA, Lac Tamhda (Anremer), Tahanaout, Sidi Chamarouch, Aguelmous

Oxycera ochracea (Vaillant, 1950)

= Hermione ochracea Vaillant, in Vaillant 1956b: 237, 244

Vaillant 1956b, HA, Lac Tamhda (Anremer), Imi-N’Ifri

Oxycera pardalina (Meigen, 1822)

Yimlahi et al. 2017, Rif, Oued Abou Bnar (NPT), Oued Maggou, Oued Achekrade, Ruisseau maison forestière (NPT), MA, Cascade Aïn Vittel, Mchacha Aïn Vittel

Oxycera rara (Scopoli, 1763)

= Hermione pulchella var. similis Vaillant, in Vaillant 1956b: 242

Vaillant 1956b, HA, Sidi Chamarouch, Imi-N’Ifri

Oxycera terminata Meigen, 1822

Yimlahi et al. 2017, Rif, Cascade Chrafate

Oxycera torrentium (Vaillant, 1950)

= Hermione torrentium Vaillant, in Vaillant 1956b: 240, 241

Vaillant 1956b, HA, Izourar

Oxycera trilineata (Linnaeus, 1767)

= Hermione bucheti Séguy, in Séguy 1939: 62

= Hermione trilineata var. algira Vaillant, in Vaillant 1956b: 244

Becker and Stein 1912, 1913, Rif, Tanger; Séguy 1930a; Villeneuve 1933; Vaillant 1956b, HA, Imi-N’Ifri; Woodley 2001; Üstüner and Hasbenli 2013; Yimlahi et al. 2017, Rif, Dayat Aïn Jdioui

Vanoyia Villeneuve, 1908

Vanoyia tenuicornis (Macquart, 1834)

Lindner 1936; Woodley 2001, Rif, Tanger; Yimlahi et al. 2017

Stratiomyini

Odontomyia Meigen, 1803

Odontomyia alolena (Séguy, 1930)

= Eulalia alolena Séguy, in Séguy 1930a: 65

Séguy 1930a, Rif, Tanger, EM, Maharidja, AP, Casablanca, MA, Aïn Leuh (1400–1500 m); Dakki 1997; Woodley 2001, Rif, Tanger, EM, Maharidja, AP, Casablanca, MA, Aïn Leuh; Yimlahi et al. 2017

Odontomyia angulata (Panzer, 1798)

Becker and Stein 1912, 1913, Rif, Tanger; Woodley 2001; Koçak and Kemal 2010; Mohammadi and Khaghaninia 2015; Yimlahi et al. 2017

Odontomyia discolor (Loew, 1846)

Becker and Stein 1912, 1913, Rif, Tanger; Rozkošný 1982, Rif, Tanger; Woodley 2001; Koçak and Kemal 2010; Üstüner and Hasbenli 2013; Yimlahi et al. 2017

Odontomyia flavissima (Rossi, 1790)

= Hadracantha flavissina nigripes Pleske, in Pleske 1925: 27, 32; Séguy 1930a: 66

= Eulalia flavissima Rossi, in Duisit 1960: 121

Séguy 1926a; Séguy 1930a; Duisit 1960, MA, Boulhaut; Dakki 1997; Woodley 2001; Yimlahi et al. 2017

Odontomyia limbata (Wiedemann, 1822)

= Eulalia limbata Wiedemann, in Séguy 1930a: 65

Becker and Stein 1912, 1913, Rif, Tanger; Séguy 1930a, Rif, Tanger, MA, Meskedell (1800–1900 m); Dakki 1997; Woodley 2001; Yimlahi et al. 2017, Rif, Lac Ametrasse, Aïn Sidi Brahim Ben Arrif, Dayat Afrate, ruisseau mai­son forestière (NPT), Aïn El Malaâb (NPT), Dayat Rmali El Malaâb (NPT), Dayat Tazia; Rif (Tahaddart) – MISR

Odontomyia microcera (Séguy, 1930)

= Eulalia microcera Séguy, in Séguy 1930a: 65

Séguy 1930a; Dakki 1997; Woodley 2001, MA, Meknès (550 m); Yimlahi et al. 2017

Stratiomys Geoffroy, 1762

Stratiomys cenisia Meigen, 1822

Séguy 1930a, Rif, Tanger, AP, Rabat, Sidi Bettache, MA, Tizi-s’Tkrine (Jebel Ahmar, 1700 m), Meknès, Aïn Sferguila, HA, Timhadit; Duisit 1960, AP, Maâmora, MA, Arhbala (1700 m); Dakki 1997; Woodley 2001; Koçak and Kemal 2010; Mohammadi and Khaghaninia 2015; Yimlahi et al. 2017

Stratiomys longicornis (Scopoli, 1763)

= Stratiomys (Hirtea) anubis Wiedemann, in Séguy 1930a: 63

Séguy 1930a, EM, Itzer (Haute Moulouya), AP, Chellah (Rabat), Casablanca, MA, Ras el Ksar (1900 m), HA, Marrakech; Duisit 1960, AP, Rabat, MA, Aguelmane Azigza (1800 m); Dakki 1997; Woodley 2001; Barták and Kubik 2005; Koçak and Kemal 2010; Üstüner and Hasbenli 2013; Yimlahi et al. 2017

Tabanoidea

ATHERICIDAE

K. Kettani, M. Mouna

Number of species: 2. Expected: 2

Faunistic knowledge of the family in Morocco: good

Athericinae

Atherix Meigen, 1803

Atherix amicorum (Thomas, 1985)

= Ibisia amicorum Thomas, in Thomas 1985: 89

Thomas 1985, HA, Oued Réghaya near Marabout Sidi Chamarouch (Toubkal, 2310 m); Boumezzough and Thomas 1987

Atherix maroccana (Séguy, 1930)

= Ibisia maroccana Séguy, in Thomas et al. 1995: 64

Séguy 1930a, MA, Oued Tigrigra; Thomas et al. 1995, MA, Oued Tigrigra (900 m), Timahdit (1830 m), HA, Asif Aït Bou Guemmaz (1900 m); Dakki 1997

RHAGIONIDAE

K. Kettani, M.J. Ebejer

Number of species: 4. Expected: 7

Faunistic knowledge of the family in Morocco: poor

Rhagioninae

Chrysopilus Macquart, 1826

Chrysopilus asiliformis (Preyssler, 1791)

= Chrysopilus aureus (Meigen, 1804), in Séguy 1941a: 29; Dakki 1997: 62

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Dakki 1997

Chrysopilus pullus Loew, 1869

Ebejer et al. 2019, Rif, Jebel Lakraâ (NPT, 1377–1541 m), Adrou (PNPB, 556 m)

Chrysopilus splendidus (Meigen, 1820)

Ebejer et al. 2019, Rif, Oued Kbir (Béni Ratene, 157 m)

Chrysopilus tsacasi Thomas, 1979

Thomas 1979, HA, Jebel Toubkal (Tachdirt, 2500 m); Boumezzough and Thomas 1987, HA, Oued Réghaya (Imlil, 1750 m), l’azib Oukaimeden (2730 m); Dakki 1997; Kerr 2004

TABANIDAE

K. Kettani

Number of species: 69. Expected: 75

Faunistic knowledge of the family in Morocco: good

Chrysopsinae

Chrysopsini

Chrysops Meigen, 1803

Chrysops caecutiens Linnaeus, 1758

El Haouari and Kettani 2014, Rif, Oued Rha, Oued Samsa, Oued Laou (Afertane), Oued Jnane Niche, Oued Berranda, Oued Biyada; El Haouari et al. 2014, HA, Imi-n’Tadart

Chrysops connexus Loew, 1858

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA, Timhadit, Oued Yquem, Volubilis, Kenitra, Meknès, Rif, Tanger; Leclercq 1967; Leclercq and Maldès 1987; Chvála et al. 1972; SA (Guelmim) – MISR

Chrysops flavipes Meigen, 1804

= Heterochrysops perspicillaris Loew, in Séguy 1930a: 79

= Chrysops punctifer Loew, in Séguy 1930a, 79

Séguy 1930a, AP, Mogador, EM, Haute Moulouya, MA, Fès, Volubilis, AA, Taroudant; Leclercq 1967, AA, Agadir-Tissint (Rocade du Draa); Chvála et al. 1972; Leclercq and Maldès 1987; Kiliç 1999; Müller et al. 2012

Chrysops italicus Meigen, 1804

Chvála et al. 1972; Leclercq and Maldès 1987; Müller et al. 2012

Chrysops mauritanicus Costa, 1893

Séguy 1930a, AP, Rabat, Fedhala, Larache, MA, Itzer, HA, Haute Réghaya; Leclercq 1967, AP, Rabat (salt marshes on Salicornia); Chvála et al. 1972; Leclercq and Maldès 1987; AP (Kénitra) – MISR

Chrysops pallidiventris Kröber, 1922

Séguy 1930a, AP, Mogador, MA, Fès; Leclercq 1967; Chvála et al. 1972; Leclercq and Maldès 1987; Pape and Thompson 2019

Chrysops relictus Meigen, 1820

Chvála et al. 1972; El Haouari and Kettani 2014, Rif, Oued Kbir (Tamuda), Oued Kelaâ (Talembote), Oued Bou Ahmed, Oued Jnane Niche, Oued Berranda, Oued Biyada; El Haouari et al. 2014, HA, Imi-n’Tadart, Oulmès, Setti Fatma, Tafza

Chrysops viduatus (Fabricius, 1794)

El Haouari et al. 2014, HA, Setti Fatma

Silvius Meigen, 1920

Silvius algirus Meigen, 1830

Séguy 1930a; Leclercq and Maldès 1987; Kiliç 1999

Silvius alpinus (Scopoli, 1763)

= Silvius vituli Fabricius, 1805, in Séguy 1930a: 78

Séguy 1930a, MA, Meknès, Forêt Zaers; Leclercq and Maldès 1987, AP, Rabat, EM, Béni Snassen, Haute Moulouya, MA, Aïn Leuh

Silvius variegatus (Fabricius, 1805)

= Diachlorus maroccanus Bigot, in Surcouf 1921: 143; Séguy 1930a: 78

Surcouf 1921, Rif, Tanger; Becker and Stein 1913, Rif, Tanger; Séguy 1930a, Rif, Tanger, AP, Rabat, EM, Haute Moulouya; Leclercq 1960, Rif, Tanger, AP, Larache, Rabat, Salé EM, Haute Moulouya; Leclercq 1967; Leclercq and Maldès 1987; Koçak and Kemal 2013a; AP (Rabat, Larache) – MISR

Pangoniinae

Pangoniini

Pangonius Latreille, 1802

Pangonius alluaudi Séguy, 1930

Séguy 1930a, MA, Azrou, Aïn Leuh, Timhadit, Tasrah des Ighrezrane, Talzent, Aharmoumou; Leclercq 1967, MA, Ifrane; Leclercq and Maldès 1987; MA – MISR

Pangonius brevicornis (Kröber, 1921)

Leclercq 1967; Leclercq and Maldès 1987

Pangonius hassani (Leclercq, 1968)

Leclercq 1968, MA, Ifrane, Dayat Aoua; Leclercq and Olsufjev 1975; Leclercq and Maldès 1987, MA, Sidi Allal El Bahraoui

Pangonius haustellatus (Fabricius, 1781)

= Pangonius marginata (Fabricius, 1805), in Séguy 1930a: 74

= Pangonius aterrima Dufour 1853, in Séguy 1930a: 74

= Pangonius funebris Macquart, 1846, in Séguy 1930a: 74

Séguy 1930a, MA, Volubilis, Tizi-s’Tkrine, Aïn Leuh, Azrou, HA, Asni; Leclercq 1961b, MA, Ifrane; Leclercq 1967, 1968; Chvála and Lyneborg 1970; Leclercq and Maldès 1987, MA, Sidi Allal El Bahraoui (forest of Quercus suber of Maâmora); Müller et al. 2012; AP (Dradek near Rabat, Kénitra), MA (wide distribution between Azrou and Ras el Ma), HA – MISR

Pangonius mauritanus (Linnaeus, 1767)

= Pangonius funebris Fabricius, 1794, in Séguy 1930a: 76

= Pangonius maculatus (Fabricius), in Séguy 1953a: 78

Séguy 1930a; Séguy 1953a, AP, Cap Ghir; Séguy 1949a, AA, Guelmim; Leclercq 1967; Leclercq and Maldès 1987, MA, Maamar (800 m); AP (Dradek, El Maazi, Mazagan) – MISR

Pangonius micans Meigen, 1820

Leclercq and Maldès 1987

Pangonius powelli Séguy, 193012

= Pangonius sobradieli Séguy, 1934e: 21

Séguy 1930a, MA, Bekrit, Tizi-s’Tkrine, Soufouloud; Séguy 1934e; Séguy 1949a, AA, Guelmim; Leclercq and Maldès 1987

Pangonius raclinae Leclercq, 1960

Leclercq 1960, HA, Tifni by Demnate; Leclercq 1967; Leclercq and Maldès 1987; Bisby et al. 2011

Philolichini

Ectinocerella Séguy, 1929

Ectinocerella surcoufi Séguy, 1929

= Pangonius ectinocerella surcoufi Séguy, in Séguy 1929a: 100

Séguy 1929a, MA, Azrou, Ank El Djemel AA, Agadir; Séguy 1930a, MA, Azrou, AA, Agadir; Leclercq and Maldès 1987, MA, From Meknès to Khemisset, near Beth river; Leclercq 1967; HA (Tifni) – MISR

Tabaninae

Diachlorini

Dasybasis Macquart, 1847

Dasybasis barbata Coscaron & Philip, 1967

= Surcoufia barbata Bigot, 1892, in Séguy 1930a: 78

= Surcoufia paradoxa Kröber, 1925, in Séguy 1930a: 78

Séguy 1930a, Rif, Tanger

Dasyrhamphis Enderlein, 1922

Dasyrhamphis algirus (Macquart, 1838)

= Atylotus algirus Auct, in Séguy 1930a: 82

Séguy 1930a, AP, Dradek, EM, Oujda, HA, Talouet Glaoua; Chvála et al. 1972; Leclercq and Maldès 1987; AP (Sibara) – MISR

Dasyrhamphis anthracinus (Meigen, 1820)

= Atylotus anthracinus Surcouf, 1924, in Séguy 1930a: 82

Séguy 1930a, AP, Rabat, Sidi Bettache, MA, M’Rirt, Aïn Sferguila, Volubilis

Dasyrhamphis ater (Rossi, 1790)

= Tabanus ater (Rossi, 1790), in Becker and Stein 1913: 77

= Atylotus ater Barotte, 1926, in Séguy 1930a: 82

= Dasyrhamphis ater (Rossi, 1790), in Leclercq and Maldès 1987: 80

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA; Leclercq 1967, MA, Ifrane; Leclercq and Maldès 1987, HA, Jebel Tazzeka, Bab Ahzar (1200 m), Idni (1700 m); MA

– MISR

Dasyrhamphis tomentosus (Macquart, 1846)

= Atylotus tomentosus Macquart, in Séguy 1930a: 84

Séguy 1930a, AP, Rabat, Oued Cherrat, MA, Azrou, El Hajeb, Meknès, Aïn Leuh, Tizi-S’Tkrine, Forêt Tiffert, Talzent, Tazarine, Meskedall; Séguy 1949a, AA, Guelmim; Leclercq and Maldès 1987

Dasyrhamphis villosus (Macquart, 1838)

= Atylotus villosus Macquart, 1838, in Séguy 1930a: 84

Séguy 1930a, MA, Tameghilt; Leclercq 1967; Leclercq and Maldès 1987; MA – MISR

Dasyrhamphis nigritus (Fabricius, 1794)

= Therioplectes alexandrinus Wiedemann, 1830, in Séguy 1930a: 83

Séguy 1930a, MA, Aïn Leuh, El Hajeb, M’Rirt, Dar M’Tougui, Dar Kaid M’Tougui, EM, Oujda; Leclercq and Maldès 1987

Haematopotini

Haematopota Meigen, 1803

Haematopota algira Kröber, 1922

Séguy 1930a; Leclercq 1961b, MA, Dayat Aoua; Leclercq 1967; Leclercq 1968, MA, Bab Ferrich, Dayat Aoua; Leclercq and Maldès 1987

Haematopota benoisti Séguy, 1930

Séguy 1930a, AP, Rabat, MA, M’Rirt; Leclercq 1967; Leclercq and Maldès 1987; Pape and Thompson 2019

Haematopota bigoti Gobert, 1880

Séguy 1930a, MA, Volubilis; Séguy 1926a; Leclercq 1961b, MA, Dayat Aoua; Leclercq 1967; MA (Ifrane) – MISR

Haematopota crassicornis Wahlberg, 1848

Becker and Stein 1913, Rif, Tanger; Séguy 1930a

Haematopota fuscicornis Becker, 1914

= Chrysozona fuscicornis Povolny, in Becker and Stein 1913: 78

Becker and Stein 1913, Rif, Tanger; Séguy 1930a (sic! fusicornis), MA, Fès; Chvála et al. 1972; Leclercq and Maldès 1987

Haematopota grandis Meigen, 1820

Leclercq 1967, AP, Kénitra; Chvála et al. 1972; Leclercq and Maldès 1987

Haematopota italica Meigen, 1804

= Haematopota tenuicornis Macquart, 1834, in Séguy 1930a: 81

= Haematopota longicornis Macquart, 1834, in Séguy 1930a: 81

Becker and Stein 1913, Rif, Tanger; Séguy 1930a; Chvála et al. 1972

Haematopota lambi Villeneuve, 1921

Leclercq 1961b, MA, Dayat Aoua; Leclercq 1967, 1968; Leclercq and Maldès 1987

Haematopota ocelligera (Kröber, 1922)

Leclercq 1961b, MA, Dayat Aoua (on a horse); Leclercq 1967, AP, Sidi Yahia du Gharb (on Juncus acutus); Leclercq 1968, MA, Azrou; Leclercq and Maldès 1987

Haematopota pluvialis (Linnaeus, 1758)

Becker and Stein 1913, Rif, Tanger; Leclercq and Maldès 1987; El Haouari and Kettani 2014, Rif, Oued Rha, marshes of Smir, Oued Kbir (Tamuda), Oued Moukhlata (Boujdad), Oued Azla (Mokdassen Oulya), Oued Moulay Bouchta, Oued Jnane Niche, Oued Koudiat Shiba; El Haouari et al. 2014, HA, Oulmès, Tafza

Haematopota pseudolusitanica Szilády, 1923

= Chrysozona lusitanica Guérin, 1835, in Séguy 1930a: 81

Séguy 1930a, MA, M’Rirt, Sebou; Leclercq 1967, MA, Sebou

Haematopota subcylindrica Pandellé, 1888

Leclercq 1967, AP, Sidi Yahia du Gharb; El Haouari and Kettani 2014, Rif, Oued Rha, Oued Boumarouil, Oued Jnane Niche; El Haouari et al. 2014, HA, Tafza

Heptatoma Meigen, 1803

Heptatoma pellucens (Fabricuis, 1779)

El Haouari and Kettani 2014, Rif, Oued Rha, Oued Achiar (Bounezzal), Oued Azla (Mokdassen Oulya), Oued Azla (Mokdassen soufla), Oued Imsa (Centre Imsa), bog of Amsemlil, Oued Ouara (Khizana), Oued Boumarouil, Oued Laou (Siflaou), Oued Talembote, Oued Laou (Afertane), Oued Tizharine, Oued Bouhya (Kanar), Bab Tariouant, Oued Taysra (Ketama), Oued Srâ (Ketama); El Haouari et al. 2014, HA, Imi-n’Tadart, Oulmès

Tabanini

Atylotus Osten-Sacken, 1876

Atylotus agrestis (Wiedemann, 1828)

Ovazza et al. 1968; Pape and Thompson 2019

Atylotus agricola Wiedemann, 1828

= Tabanus agricola var. Kröberi Surcouf, in Séguy 1953a: 78

Séguy 1953a, SA, entre Tagounit et Zegdou

Atylotus fulvus (Meigen, 1804)

Leclercq 1961b, MA, Aïn Leuh, Bordj Doumergue; Leclercq 1967, Rif, Ketama; Chvála et al. 1972; Leclercq and Maldès 1987; Barták and Kubik 2005

Atylotus latistriatus (Brauer, 1880)

= Dasystipia nigrifacies Gobert, 1881, in Séguy 1930a: 84

Séguy 1930a, MA, Aïn Leuh; Chvála et al. 1972; Kiliç 1999

Atylotus loewianus (Villeneuve, 1920)

Leclercq 1967, MA, Aguelmane Azigza (marshy meadow), Aguelmane de Sidi Ali Leclercq 1968; Chvála et al. 1972

Atylotus pulchellus Loew, 1858

Becker and Stein 1913 Rif, Tanger; Chvála et al. 1972

Atylotus quadrifarius (Loew, 1874)

Chvála et al. 1972; Müller et al. 2011

Atylotus sublunaticornis (Zetterstedt, 1842)

El Haouari and Kettani 2014, Rif, Oued Rha, Oued Kbir (Koudiat Krikra), Oued Martil, Oued Khizana, Oued Laou (Ifansa), Oued Bou Ahmed, Oued Biyada; El Haouari et al. 2014, HA, Imi-n’Tadart, Oulmès

Hybomitra Enderlein, 1922

Hybomitra arpadi Szilády 1923

El Haouari et al. 2014, HA, Oulmès

Hybomitra bimaculata Macquart, 1826

Ježek 1995; El Haouari and Kettani 2014, Rif, Oued Rha, marshes of Lemtahane (PNPB), Oued Raouz, Oued Zarka, Oued Mokhlata (Boujdad), Oued Amsa (Er-Rifiyine), bog of Amsemlil, Oued Talembote (Talembote), Oued Jnane Niche, Oued Biyada, Oued Aârkob, Oued Sidi Yahia Aârab; El Haouari et al. 2014, HA, Oulmès

Hybomitra distinguenda (Verrall, 1909)

Ježek et al. 2012; El Haouari et al. 2014, HA, Imi-n’Tadart

Hybomitra vittata (Fabricius, 1794)

= Straba vittata Fabricius, 1794, in Séguy 1930a: 83

= Tabanus spectabilis Loew, 1858, in Séguy 1930a: 83

Séguy 1930a, Rif, Tanger AP, Maâmora, Rabat, Casablanca, MA, Oued Yquem, M’Rirt; Chvála and Lyneborg 1970, Rif, Tanger, EM, Haute Moulouya; Leclercq and Maldès 1987

Tabanus Linnaeus, 1758

Tabanus autumnalis Linnaeus, 1761

= Straba autumnalis Linnaeus, 1761, in Séguy 1930a: 82

= Straba automnalis var. brunnescens Szilády, 1941, in Séguy 1930a: 82

= Straba automnalis var. molestans Becker, 1914, in Séguy 1930a: 83

Loew 1860; Becker and Stein 1913, Rif, Tanger; Séguy 1930a, AP, Rabat, EM, Béni Snassen, Itzer (Haute Moulouya), MA, Aïn Leuh; Séguy 1953a, HA, Ksar-es-Souk; Leclercq 1961b, MA, Dayat Aoua, Aïn Leuh, Immouzer Kander; Leclercq 1967, AP, Kénitra (on Mimosa grove), Gharb (Sidi Yahia, Sidi Allal Tazi), MA, Adjir by Khenifra (on livestock), Leclercq 1968, MA, Dayat Aoua; Leclercq and Maldès 1987, HA, edges near river Tessaout, Kelaâ des Sraghna; Pârvu et al. 2006, AP, Merja Zerga, MA, Kasba Tadla; El Haouari et al. 2014, HA, Setti Fatma; MA (Allal Tazi) – MISR

Tabanus barbarus Coquebert, 1804

Becker and Stein 1913, Rif, Tanger; Leclercq 1961b Rif, Azib de Ketama; Chvála and Lyneborg 1970, Rif, Tanger; Portillo 1982; Leclercq and Maldès 1987, MA, marshes around Kasba Tadla; Popescu-Mirceni 2011, AP, Merja Zerga

Tabanus bifarius Loew, 1858

= Atylotus bifarius Loew, 1858, in Séguy 1930a: 83

Séguy 1930a, EM, Haute Moulouya

Tabanus bovinus Linnaeus, 1758

Loew 1860; Séguy 1930a; Leclercq 1967; Portillo 1982; Leclercq and Maldès 1987; Barták and Kubik 2005, MA, M’Rirt; El Haouari and Kettani 2014, Rif, Oued Khemis (Khemis Anjra), Oued Kelaâ (Akchour)

Tabanus bromius Linnaeus, 1758

= Straba bromius Linnaeus, 1758, in Séguy 1930a: 83

Loew 1860; Séguy 1930a, EM, Haute Moulouya, MA, Aïn Leuh, Ras El Ksar; Leclercq 1961b, Rif, Azib de Ketama, MA, Ifrane, Immouzer, Azrou, Bordj Doumergue, Dayat Aoua, Aïn Leuh; Leclercq 1967, MA, Immouzer des Marmoucha, Adjir by Khenifra; Leclercq 1968, Rif, Ketama, MA, Bab-Bou-Idir, Bab Ferrich, Ifrane, Col du Zad; Pernot-Visentin and Beaucournu-Saguez 1974, Rif; Leclercq and Maldès 1987; El Haouari and Kettani 2014, Rif, Oued Samsa, Oued Raouz, Oued Zarka, Oued Khemis (Khemis Anjra), Oued Azla (Mokdassen Oulya), Oued Kelaâ (Akchour), Oued Laou (Sifalaou), Oued Jnane Niche, Oued Berranda; El Haouari et al. 2014, HA, Oulmès; AA (Errachidia) – MISR

Tabanus choumarae Leclercq, 1967

Leclercq 1967, AA, Aouinet Torkoz (down Draa); Leclercq and Olsufjev 1975; Leclercq and Maldès 1987; Pârvu et al. 2006, AA, Ouarzazate

Tabanus cordiger Meigen, 1820

Leclercq 1961b, MA, Bordj Doumergue, Dayat Aoua; Leclercq 1967, Rif, Had el Rouadi, MA, Arbala par Ksiba, Boulemane, AA, Agadir-Tissint (Rocade du Draa), Akka; Leclercq 1968, Rif, Ketama, MA, Bab Termas, Taza, Doniet; Leclercq 1986; Leclercq and Maldès 1987, AA, Tinmal, edges of Draa river, Agdz, Ourika near Ouarzazate; Barták and Kubik 2005; Müller et al. 2011; Popescu-Mirceni 2011, HA, Ouarzazate; El Haouari and Kettani 2014, Rif, Oued Berranda; El Haouari et al. 2014, Rif, Oued Oueghra

Tabanus darimonti Leclercq, 1964

Leclercq 1967, MA, Aïn Leuh; Leclercq 1968, MA, Bab Boudir, forêt Bab-Azhar, Immouzer Kander; Portillo 1982; Leclercq and Maldès 1987; Mikuška et al. 2008

Tabanus eggeri Schiner, 1868

= Tabanus intermedius Egger, 1859, in Séguy 1930a: 83

Séguy 1930a, MA, M’Rirt; Leclercq 1968, Rif, M’Diq, MA, Bab Boudir, El Hajeb, Azrou, Miscliffen; Portillo 1982; Leclercq and Maldès 1987; Mikuška et al. 2008

Tabanus leleani Austen, 1920

= Atylotus leleani Austen, 1920, in Séguy 1930a: 84

Séguy 1930a, HA, upstream of Réghaya; Leclercq 1967; Leclercq and Maldès 1987; MA – MISR

Tabanus lunatus Fabricius, 1794

= Atylotus lunatus Fabricius, 1974, in Séguy 1930a: 84

Séguy 1930a, EM, Haute Moulouya, MA, Meknès; Leclercq 1961b, MA, Ifrane, Immouzer, Bordj Doumergue, Dayat Aoua, Aïn Leuh; Leclercq 1967, EM, Haute Moulouya, MA, Ajdir by Khenifra (on livestock); Leclercq 1968, Rif, Ketama, MA, Bab Bouder, Immouzer Kander, El Hajeb, Mishliffen, Jebel Hebri; Portillo 1982; Leclercq and Maldès 1987, MA, Afourer (800 m) – MISR

Tabanus maculicornis Zetterstedt, 1842

El Haouari and Kettani 2014, Rif, Oued Rha, marshes of Lemtahane, Oued Boumarouil, Oued Laou (Dardara), Oued Kanar, Oued Jnane Niche; El Haouari et al. 2014, HA, Imi-n’Tadart, Oulmès, Tafza

Tabanus miki Brauer, 1880

El Haouari and Kettani 2014, Rif, Oued Khemis, Oued Boumarouil, Oued Berranda, Oued Biyada; El Haouari et al. 2014, HA, Setti Fatma

Tabanus nemoralis Meigen, 1820

Leclercq 1968, Rif, Ketama; Portillo 1982; Leclercq and Maldès 1987

Tabanus quatuornotatus Meigen, 1820

Portillo 1982; Leclercq and Maldès 1987; Koçak and Kemal 2010; Müller et al. 2011; El Haouari and Kettani 2014, Rif, Oued Maggou, Oued Ouara (Khizana); El Haouari et al. 2014, HA, Setti Fatma; Rif (Talassemtane) – MISR

Tabanus sudeticus Zeller, 1842

Leclercq 1967; Portillo 1982; Leclercq and Maldès 1987; Gioia Martins-Neto 2003

Tabanus tinctus Walker, 1850

Leclercq 1961b, MA, forêt Bab Azhar, Aïn Leuh; Pernot-Visentin and Beaucournu-Saguez 1974, MA, HA (2300 m); Portillo 1982; Leclercq and Maldès 1987; MA (Azrou) – MISR

VERMILEONIDAE

K. Kettani, M.J. Ebejer

Number of species: 6. Expected: 6

Faunistic knowledge of the family in Morocco: good

Vermileoninae

Lampromyia Macquart, 1835

Lampromyia cylindrica (Fabricius, 1794)

Stuckenberg 1998, HA

Lampromyia lecerfi Séguy, 1928

= Lampromyia Le Cerfi Séguy, in Séguy 1928a: 45

Séguy 1928a, HA, Tinmel (Goundafa), Asni; Stuckenberg 1998, HA; Kirk-Spriggs and McGregor 2009; Kehlmaier 2014, HA; AP (Tamri) – MHNN

Lampromyia nigripennis Séguy, 1930

Séguy 1930a, EM, Berkane, grove in Tlet n’Rhohr; Stuckenberg 1998, MA; Kirk-Spriggs and McGregor 2009; Kehlmaier 2014, MA, south of Azrou (1500 m), HA

Lampromyia pallida Macquart, 1835

Stuckenberg 1998, HA

Vermileo Macquart, 1834

Vermileo vermileo (Linnaeus, 1758)13

= Vermileo degeeri Macquart 1834, in Séguy 1953a: 78

Séguy 1953a, AP, Rabat

Vermileo nigriventris Strobl, 1906

Ebejer et al. 2019, Rif, Cap Spartel (Tanger, 15 m), Anissar (PNPB, 987 m)

Nemestrinoidea

ACROCERIDAE

K. Kettani, E.P. Nartshuk

Number of species: 13. Expected: 25

Faunistic knowledge of the family in Morocco: poor

Acrocerinae

Acrocera Meigen, 1803

Acrocera orbicula (Fabricius, 1787)

Weinberg and Bächli 2002, HA, Marrakech, Ouirgane (1000 m)

Cyrtus Latreille, 1796

Cyrtus gibbus (Fabricius, 1794)

Becker and Stein 1913, Rif, Tanger; Séguy 1926, Rif; Pleske 1930, Rif, Tanger; Schlinger 1972; Mouna 1998

Cyrtus maroccanus Séguy, 1930

Schlinger 1972, Rif; Mouna 1998

Cyrtus pallidus Gil Collado, 1929

Gil Collado 1929b, Rif, Tanger; Schlinger 1972; Mouna 1998

Cyrtus pusillus Macquart, 1834

Pleske 1930, Rif, Tanger; Schlinger 1972, Rif, Tanger; Mouna 1998

Ogcodes Latreille, 1797

Ogcodes zonatus (Erichson, 1840)

Becker and Stein 1913, Rif, Tanger; Séguy 1926; Pleske 1930

Opsebius Costa, 1856

Opsebius cyrtus Séguy, 1930

Mouna 1998; HA (Lac Ifni) – MISR

Opsebius formosus Loew, 1871

Becker and Stein 1913, Rif, Tanger

Opsebius inclinatus Séguy, 1930

Mouna 1998

Opsebius inflatus (Loew, 1857)

Weinberg and Bächli 2002, MA, Azrou, Timahdit Ighboula (1850 m), HA, Marrakech, Ouirgane (1000 m)

Opsebius pepo Loew, 1870

Pleske 1930, Rif, Tanger

Panopinae

Astomella Latreille, 1809

Astomella hispaniae Lamarck, 1816

Gil Collado 1929b, AP, Mogador; Mouna 1998

Physegastrella Brunetti, 1926

Physegastrella maroccana Brunetti, 1926

Brunetti 1926; Pleske 1930; Mouna 1998

NEMESTRINIDAE

K. Kettani, D. Barraclough

Number of species: 13

Faunistic knowledge of the family in Morocco: poor

Nemestrininae

Nemestrinus Latreille, 1802

Nemestrinus aegyptiacus (Wiedemann, 1828)

Timon-David 1951; Séguy 1953a, HA, Amsed; Bernardi 1973; Mouna 1998; SA (Oued el Ma) – MISR

Nemestrinus ater (Olivier, 1810)

Mouna 1998; EM (Zaouillet El Atenf) – MISR

Nemestrinus escalerai Arias, 1913

Paramonow 1945; Bernardi 1973, HA, Marrakech; Mouna 1998

Nemestrinus exalbidus (Lichtwardt, 1907)

Mouna 1998

Nemestrinus fasciatus (Olivier, 1810)

= Rhynchocephalus fasciatus Olivier, in Mouna 1998: 86

Bernardi 1973; Mouna 1998; MA (Immouzer) – MISR

Nemestrinus nigrovillosus Lichtwardt, 1909

Arias 1913; Séguy 1930a, MA, Ras el Ma, Azrou, Forêt Tiffert (2000–2200 m), HA, Tizi-n’Test, Jebel Imdress (2000–2450 m), Goundafa; Séguy 1941d, HA, Tizi-n’Test; Timon-David 1951, MA, Tizi-n’Tretten; Bernardi 1973; Mouna 1998

Nemestrinus pieltaini (Gil Collado, 1934)

= Nemestrellus pieltaini Gil Collado, in Gil Collado 1934: 325

Gil Collado 1934, Rif, Imasinen, Bab Chiquer, Bab Bagla; Bernardi 1973; Mouna 1998

Nemestrinus ruficornis (Macquart, 1840)

Mouna 1998

Nemestrinus rufipes (Olivier, 1810)

Mouna 1998; MA (Timahdit) – MISR

Nemestrinus striatus (Lichtwardt, 1907)

Mouna 1998

Trichopsideinae

Fallenia Meigen, 1820

Fallenia fasciata (Fabricius, 1805)

Arias 1913; Séguy 1930a, AP, Casablanca, Rabat, Bou Knadel, MA, M’Rirt (1200 m); Timon-David 1951, AP, Forêt Maâmora; Mouna 1998; AP (Rabat, Bou Knadel), MA (Aïn Leuh) – MISR

Neorhynchocephalus Lichtwardt, 1909

Neorhynchocephalus tauscheri (Fisher, 1812)

= Rhynchocephalus tauscheri Fischer, in Mouna 1998: 86

Mouna 1998

Trichopsidea Westwood, 1839

Trichopsidea costata (Loew, 1857)

= Symmictus costatus Loew, in Arias 1913: 26, Séguy 1930a: 89

Arias 1913; Séguy 1930a, MA, Tameghilt (1900 m); Mouna 1998

Asiloidea

ASILIDAE

K. Kettani, G. Tomasovic

Number of species: 131. Expected: 230

Faunistic knowledge of the family in Morocco: moderate

Apocleinae

Apoclea Macquart, 1838

Apoclea algira (Linnaeus, 1767)

Séguy 1953a, AA, Tata; Tomasovic 1997; Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014

Apoclea micracantha Loew, 1856

Tomasovic 1997, HA, Sidi Mhejmed Ou Said; Geller-Grimm 2007; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014

Eremonotus Theodor, 1980

Eremonotus hauseri Geller-Grimm & Hradský, 1998

Geller-Grimm and Hradský 1998, HA; Geller-Grimm 2007, AA, Agadir

Asilinae

Afroepitriptus Lehr, 1992

Afroepitriptus beckeri Lehr, 1992

Geller-Grimm 2007; Koçak and Kemal 2010

Antiphrisson Loew, 1849

Antiphrisson trifarius Loew, 1849

Tomasovic 1997, HA, Errachidia, Ziz, Oasis Zouala; Geller-Grimm 2007; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014; HA (Asni) – MISR

Asilus Linnaeus, 1758

Asilus barbarus Linnaeus, 1758

Becker and Stein 1913, Rif, Tanger; Séguy 1930a; Séguy 1941a, HA, Tizi-Tamatert (Toubkal, 2250 m); Mouna 1998; Weinberg and Blasco-Zumeta 2004; Geller-Grimm 2007; Koçak and Kemal 2010; Ghahari et al. 2014

Asilus crabroniformis Linnaeus, 1758

Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; Ghahari et al. 2014

Asilus tingitanus Boisduval, 1835

Geller-Grimm 2007, Rif, Tanger

Dysmachus Loew, 1860

Dysmachus albisetosus (Macquart, 1850)

Geller-Grimm 2007

Dysmachus cochleatus (Loew, 1854)

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007

Dysmachus cristatus (Wiedemann, 1820)

= Dysmachus dasynotus Loew, in Becker and Stein 1913: 72, Timon-David 1951: 138

Becker and Stein 1913, Rif, Tanger; Timon-David 1951, AP, Rabat, Harcha, Salé, Oued Ksab, MA, Ifrane; Mouna 1998; Geller-Grimm 2007; AP (Rabat, Cap Cantia) – MISR

Dysmachus digitulus Becker, 1923

Geller-Grimm 2007

Dysmachus elapsus Villeneuve, 1933

Villeneuve 1933, AP, Mazagan, Mogador; Mouna 1998; Tomasovic 2001b; Geller-Grimm 2007; AP (Cap Cantia) – MISR

Dysmachus evanescens Villeneuve, 1912

Timon-David 1951, AP, Sehoul; Mouna 1998; Geller-Grimm 2007

Dysmachus trigonus (Meigen, 1804)

Timon-David 1951, AP, Rabat, Chellah, Forêt Maâmora, Ras el Arba, Sehoul, Zaër

Eccoptopus Loew, 1860

Eccoptopus longitarsis (Macquart, 1838)

Timon-David 1951, AA, Zagora; Mouna 1998; Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014

Engelepogon Lehr, 1992

Engelepogon brunnipes (Fabricius, 1794)

= Heligmoneura brunnipes Fabricius, in Séguy 1930a: 125

= Acanthopleura brunnipes Fabricius, in Timon-David 1951: 137

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA, Meknès; Timon-David 1951, MA, Meknès; Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010; Ghahari et al. 2014

Epitriptus Loew, 1849

Epitriptus cingulatus (Fabricius, 1871)

Séguy 1941a, AA, Agadir; Mouna 1998

Eremisca Hull, 1962

Eremisca heleni heleni (Efflatoun, 1934)

Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014

Eremisca osiris (Wiedemann, 1828)

Geller-Grimm 2007; El Hawagry 2011

Eutolmus Loew, 1848

Eutolmus wahisi Tomasovic, 2001

Tomasovic 2001a, Rif, Tétouan (Jebel Tazout, 1650 m); Geller-Grimm 2007; Koçak and Kemal 2010

Filiolus Lehr, 1967

Filiolus apicalis (Becker in Becker & Stein, 1913)

= Eutolmus apicalis Becker, in Becker and Stein 1913: 75

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007

Machimus Loew, 1849

Machimus cribratus (Loew, 1849)

Geller-Grimm 2007; AP (Cap Cantia) – MISR

Machimus fimbriatus (Meigen, 1804)

Geller-Grimm 2007

Machimus fortis (Loew, 1849)

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, AP, Rabat; Mouna 1998; Geller-Grimm 2007

Machimus gonatistes (Zeller, 1840)

Geller-Grimm 2007

Machimus mauritanicus Bequaert, 1964

Tomasovic 2003; Geller-Grimm 2007; AP (Forêt Boulhaut, Salé) – MISR

Machimus nigrosetosus Séguy, 1941

Séguy 1941d AA, Agadir; Mouna 1998; Geller-Grimm 2007

Machimus perplexus Becker, 1915

Geller-Grimm 2007

Machimus pilipes (Meigen, 1820)

= Eutolmus hispanus Loew, in Becker and Stein 1913: 74

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA, Forêt Zaers, Tizi-n’Bouftene (2300 m), HA, bords de l’Imminen (Tachdirt: 2400–2600 m); Geller-Grimm 2007

Machimus pseudogonatistes Villeneuve, 1930

= Machimus ermineus Becker, in Mouna 1998: 84

Villeneuve 1933; Mouna 1998; Geller-Grimm 2007

Neoepitriptus Lehr, 1992

Neoepitriptus inconstans (Wiedemann in Meigen, 1820)

= Machimus micropyga Becker, in Becker and Stein 1913: 74

Becker and Stein 1913, Rif, Tanger; Mouna 1998; Geller-Grimm 2007; El Hawagry 2011, Rif, Tanger

Neoepitriptus minusculus (Bezzi, 1898)

= Machimus minusculus Bezzi, in Timon-David 1951: 138, Mouna 1998: 84

Timon-David 1951, MA, Ifrane; Mouna 1998; Geller-Grimm 2007

Neomochtherus Osten-Sacken, 1878

Neomochterus brevipennis Séguy, 1932

Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010

Neomochtherus grandicollis (Becker, 1914)

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007

Neomochterus ochriventris (Loew, 1854)

Timon-David 1951, AP, Sidi Moussa el Harati; Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010

Pashtshenkoa Lehr, 1995

Pashtshenkoa clypeatus maroccanus (Tsacas, 1968)

Geller-Grimm 2007

Phileris Tsacas & Weinberg, 1976

Phileris haplopygus Tsacas & Weinberg, 1976

Geller-Grimm 2007

Phileris pilosus Tsacas & Weinberg, 1976

Geller-Grimm 2007

Satanas Jacobson, 1908

Satanas gigas (Eversmann, 1855)

Maldès 2000, ME, Oujda, HA, Errachidia, Meski

Turka Őzdikmen, 2008

Turka cervinus (Loew, 1856)

= Stenopogon cervinus Loew, in Séguy 1930a: 122

Séguy 1930a, MA, pont de l’Oued Korifla (Zaers), HA, Sidi Bou Rziguine; Geller-Grimm 2007; Hayat et al. 2008; Özdikmen 2008; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014

Dasypogoninae

Dasypogon Meigen, 1803

Dasypogon atratus (Fabricius, 1794)

= Selidopogon atratus Meigen, in Séguy 1930a: 118

= Selidopogon atratus Fabricius, in Timon-David 1951: 136

Séguy 1930a, MA; Timon-David 1951, Rif, Ouezzane, AP, Rabat MA, Oued Beth; Mouna 1998; Geller-Grimm 2007

Dasypogon auripilus (Séguy, 1934)

Mouna 1998; Geller-Grimm 2007; AP (Casablanca) – MISR

Dasypogon crassus Macquart in Lucas, 1849

= Selidopogon crassus Macquart, in Séguy 1930a: 119, Timon-David 1951: 136

Séguy 1930a, Rif, Tanger, MA, Meknès; Timon-David 1951, AP, M’Soun, Guerrouaou; Mouna 1998; Geller-Grimm 2007

Dasypogon diadema (Fabricius, 1781)

= Selidopogon cylindricus Fabricius, in Séguy 1930a: 119

= Selidopogon diadema Fabricius, in Séguy 1930a: 118

= Selidopogon sicanus Costa, 1853, in Hayat et al. 2008: 183

Séguy 1930a, AP, Dar Salem, Tarfaya, Oued Korifla (Zaers), HA, Bou Tazzert; Timon-David 1951, AP, Port Lyautey; Mouna 1998; Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; Ghahari et al. 2014

Dasypogon gougeleti (Bigot, 1878)

= Selidopogon gougeleti Bigot, in Timon-David 1951: 136

Becker and Stein 1913, Rif, Tanger; Timon-David 1951, AP, Oued Korifla; Mouna 1998; Geller-Grimm 2007

Dasypogon olcesci (Bigot, 1878)

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007

Dasypogon rubinipes (Becker in Becker & Stein, 1913)

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007

Dasypogon ruficauda (Fabricius, 1805)

Geller-Grimm 2007

Saropogon Loew, 1847

Saropogon aretalogus Séguy, 1953

Séguy 1953a, MA, Ifrane; Geller-Grimm 2007

Saropogon aurifrons (Macquart in Lucas, 1850)

Timon-David 1951, AP, Zaers; Mouna 1998; Geller-Grimm 2007; El Hawagry 2011

Saropogon clausus Becker, 1906

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, EM, Itzer, Moulay Aïn Djemine (Haute Moulouya); Timon-David 1951, AP, Zaers; Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010; Ghahari et al. 2014

Saropogon jugulum (Loew, 1847)

Timon-David 1951, AP, Zaers; Mouna 1998; Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014

Saropogon leucocephalus (Meigen, 1820)

Séguy 1930a, MA, Forêt Tiffert (2000–2200 m); Mouna 1998; Geller-Grimm 2007; El Hawagry 2011; Ghahari et al. 2014

Saropogon maroccanus Séguy, 1930

Séguy 1930a, MA, Ras El Ksar (1900 m); Séguy 1949a, SA, Goulimine; Mouna 1998; Carles-Tolrá 2002; Geller-Grimm 2007

Saropogon obscuripennis (Macquart in Lucas, 1849)

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, AP, Rabat, MA, Aïn Leuh, Tizi-s’Tkrine (1700 m), HA, Imi-M’Tanout, Dar M’Tougui; Séguy 1941d, AA, Agadir; Timon-David 1951, EM, Guenfouda; Mouna 1998; Geller-Grimm 2007

Saropogon philocalus Séguy, 1941

Séguy 1941d, AA, Agadir; Mouna 1998; Geller-Grimm 2007

Saropogon rufipes (Gimmerthal, 1847)

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007

Saropogon tassilaensis Séguy, 1953

Séguy 1953a, AA, Tassila (Souss); Geller-Grimm 2007

Dioctriinae

Dioctria Meigen, 1803

Dioctria atrorubens Séguy, 1930

Séguy 1930a, MA, Tizi-s’Tkine (1700 m); Villeneuve 1933; Mouna 1998; Geller-Grimm 2007

Dioctria cothurnata Meigen, 1820

Ebejer et al. 2019, Rif, Dardara (484 m)

Dioctria fuscipes Macquart, 1834

Timon-David 1951, MA, Aguelmane Sidi Ali (2070 m)

Dioctria gagates Wiedemann in Meigen, 1820

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007

Dioctria notha Séguy, 1941

Séguy 1941d, AA, Agadir; Mouna 1998; Geller-Grimm 2007

Dioctria rufa Strobl, 1906

Ebejer et al. 2019, Rif, Dardara (484 m)

Dioctria rungsi Timon-David, 1951

Timon-David 1951, MA, Ifrane (1650 m); Mouna 1998; Geller-Grimm 2007

Laphriinae

Glyphotriclis Hermann, 1920

Glyphotriclis ornatus (Schiner, 1868)

= Triclis ornatus Schiner, in Becker and Stein 1913: 67

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, HA, Marrakech; Mouna 1998; Geller-Grimm 2007; El Hawagry 2011

Laphria Meigen, 1803

Laphria bomboides Macquart, 1849

= Laphria praelusia Séguy, in Séguy 1930a: 124

Séguy 1930a, MA, Soufouloud (1900–2100 m); Mouna 1998; MA (Meghraona, Tamtraekt) – MISR

Pogonosoma Rondani, 1856

Pogonosoma maroccanum (Fabricius, 1794)

Loew 1860; Becker and Stein 1913, Rif, Tanger; Mouna 1998; Carles-Tolrá 2002; Geller-Grimm 2004; Geller-Grimm 2007; Ghahari et al. 2007; Koçak and Kemal 2010; Koçak and Kemal 2013a; Ghahari et al. 2014

Stiphrolamyra Engel, 1928

Stiphrolamyra rubicunda Oldroyd, 1947

Timon-David 1951, AP, Sidi Moussa el Harati; Mouna 1998; Geller-Grimm 2007; El Hawagry 2011; Ghahari et al. 2014

Stiphrolamyra vitai Hradský & Geller-Grimm, 1997

Hradský and Geller-Grimm 1997, HA, Taroudant; Geller-Grimm 2007

Laphystiinae

Perasis Hermann, 1905

Perasis sareptana Hermann, 1906

Séguy 1930a, HA, Asni; Mouna 1998

Scytomedes Röder, 1882

Scytomedes haemorrhoidalis (Fabricius, 1794)

= Triclis haemorrhoidalis Fabricius, in Mouna 1998: 84

Séguy 1930a, MA, Meknès; Mouna 1998; Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; Ghahari et al. 2014

Trichardis Hermann, 1906

Trichardis leucocomus (Van der Wulp, 1899)

= Trichardis leucicoma Van der Wulp, in Timon-David 1951: 132

Timon-David 1951, AA, Tata, piste de Fask Tahrjicht; Mouna 1998; Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014

Leptogastrinae

Leptogaster Meigen, 1803

Leptogaster cylindrica (De Geer, 1776)

= Leptogaster hispanica Meigen, in Séguy 1930a: 117

Séguy 1930a, MA, Meknès; Mouna 1998; Tomasovic 2006, Rif; Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; Ghahari et al. 2014

Leptogaster pedunculata Loew, 1847

= Gonypes pedunculatus Loew, in Becker and Stein 1913: 72

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, HA, Haute Réghaya; Mouna 1998

Leptogaster straminea Becker, 1907

Timon-David 1951, MA, Aguelmane Sidi Ali (2070 m); Mouna 1998; Geller-Grimm 2007

Stenopogoninae

Afroholopogon Londt, 1994

Afroholopogon waltlii (Meigen, 1838)

= Heteropogon waltlii Meigen, in Séguy 1930a: 123

Séguy 1930a, MA, Meknès; Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010

Amphisbetetus Hermann, 1906

Amphisbetetus sexspinus Tomasovic, 2008

Tomasovic and Weyer 2008, AA, Imsouane (Agadir); Geller-Grimm 2007

Ancylorhynchus Berthold in Latreille, 1827

Ancylorrhyncus gummigutta (Becker, 1906)

Séguy 1930a, Rif, Tanger; Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010

Ancylorrhyncus limbatus (Fabricius, 1794)

Séguy 1930a, MA, Meknès, Timhadit (2000 m); Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010

Ancylorrhyncus vultur Séguy, 1930

Séguy 1930a, MA, Timhadit (2000 m); Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010

Eriopogon Loew, 1847

Eriopogon jubatus Becker, 1906

Timon-David 1951, Rif, Tanger, AP, Rabat; Hradský and Hüttinger 1995, AP, Rabat; Mouna 1998; Geller-Grimm 2007; AP (Forêt Temara) – MISR

Eriopogon laniger Meigen, 1804

= Holopogon flavescens Jaennicke, in Séguy 1930a: 123

Séguy 1930a, HA, Aguergour; Mouna 1998; Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; Ghahari et al. 2014

Eriopogon spatenkai Hradský & Hüttinger, 1995

Geller-Grimm 2007; Hradský and Hüttinger 1995, MA, Mishliffen

Galactopogon Engel, 1929

Galactopogon hispidus Engel, 1929

Ebejer et al. 2019, AA, 23 km S of Rich (Errachidia, 2012 m)

Habropogon Loew, 1847

Habropogon aerivagus (Séguy, 1953)

Séguy 1953a, SA, Aouletis

Habropogon appendiculatus Schiner, 1867

Timon-David 1951, AA, Aïn Chaïb; Mouna 1998; Weinberg and Blasco-Zumeta 2004; Hradský and Geller-Grimm 2005; Geller-Grimm 2007; Koçak and Kemal 2010; El Hawagry 2011

Habropogon bacescui Weinberg & Tsacas, 1973

Geller-Grimm 2007; Koçak and Kemal 2010

Habropogon distipilosus Weinberg & Tsacas, 1973

Geller-Grimm 2007

Habropogon hauseri Hradský & Geller-Grimm, 2005

Hradský and Geller-Grimm 2005, HA, Tizi-n’Test; Geller-Grimm 2007; Koçak and Kemal 2010

Habropogon odontophallus Weinberg & Tsacas, 1973

Geller-Grimm 2007

Habropogon parappendiculatus Weinberg & Tsacas, 1973

Hradský and Geller-Grimm 2005, HA, Aït Saoun; Geller-Grimm 2007; Kirk-Spriggs and McGregor 2009, HA; Koçak and Kemal 2010

Habropogon prionophallus Weinberg & Tsacas, 1973

Geller-Grimm 2007

Habropogon pyrrhophaeus Weinberg & Tsacas, 1973

Geller-Grimm 2007

Habropogon rubriventris Macquart, 1849

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, AP, Aïn el Hadjar (near Mogador), MA, Meknès, Tlet n’Rhohr, EM, Berkane (1350–1400 m); Mouna 1998

Habropogon senilis Wulp, 1899

Geller-Grimm 2007

Habropogon spissipes Hermann, 1909

Geller-Grimm 2007; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014

Habropogon striatus (Fabricius, 1794)

= Habropogon heteroneurus Timon-David, in Timon-David: 135

Séguy 1941d, AA, Agadir; Mouna 1998; Geller-Grimm 2007; El Hawagry 2011, AP, Rabat

Heteropogon Loew, 1847

Heteropogon biplex Becker, in Becker & Stein 1913: 65

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007

Heteropogon manicatus (Meigen, 1820)

Séguy 1930a, MA, Azrou, Meknès, Aïn Leuh, HA, Asni; Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010; MA (Ifrane) – MISR

Heteropogon nubilus (Meigen, 1820)

= Isopogon brevis Schiner, in Becker and Stein 1913: 64

= Sisyrnodytes brevis Macquart, in Timon-David 1951: 134, Séguy 1953a: 79

Becker and Stein 1913, Rif, Tanger; Séguy 1953a, AA, Imiter; Mouna 1998; Geller-Grimm 2007; Koçak and Kemal 2010; Shoeibi and Karimpour 2010; Ghahari et al. 2014

Holopogon Loew, 1847

Holopogon dimidiatus (Meigen, 1820)

Séguy 1941d, AA, Agadir

Holopogon dusmeti Strobl in Czerny & Strobl 1909

= Eriopogon dusmeti Strobl, in Timon-David 1951: 132

Timon-David 1951, EM, Guenfouda, HA, Tifni; Mouna 1998; Geller-Grimm 2007; HA (Tifni Demnat) – MISR

Holopogon melaleucus (Meigen, 1820)

Séguy 1930a, AP, Forêt Maâmora, Dar Salem (Rabat); Séguy 1941d, AA, Agadir; Mouna 1998; Geller-Grimm 2007

Holopogon pusillus (Macquart, 1838)

= Habropogon pusillus (Macquart), in Séguy 1949a: 154

Séguy 1949a, SA, Goulimine; Mouna 1998

Holopogon quadrinotatus Séguy, 1953

Séguy 1953a, SA, Amguilli Sguelma

Acnephalum Macquart, 1838

= Pycnopogon Loew, 1847 in Londt 2010

Acnephalum apiformis (Macquart in Lucas, 1849)

Séguy 1930a, MA, Timhadit (2000 m), Meskedall (1800–1900 m); Timon-David 1951, MA, Ifrane (1650 m); Mouna 1998; Geller-Grimm 2007

Acnephalum denudatus (Séguy, 1949)

= Stenopogon denudatus Loew, in Séguy 1930a: 123, Séguy 1934b: 162

Séguy 1930a, MA, Tizi-n’Tkrine; Séguy 1934b, HA, Haute Réghaya; Séguy 1949b, AP, Bou Tazzert near Mogador; Séguy 1953a, AA, Oasis du Ferkla; Mouna 1998; Geller-Grimm 2007

Acnephalum fasciculatus (Loew, 1847)

Séguy 1930a, MA, Azrou, Timelilt, Sidi Bettache, HA, Asni, bords Imminen (Tachdirt), Likount (2500–2800 m), Lac Ifni (Skoutana), SA, Béni Mgild; Timon-David 1951, AP, Oued Korifla, MA, Lac Aguelmane Sidi Ali (2070 m), Oued N’Zala; Mouna 1998; Geller-Grimm 2007; Lehr et al. 2007; Koçak and Kemal 2010; Ghahari et al. 2014; MA (Ras el Ma) – MISR

Stenopogon Loew, 1847

Stenopogon costatus Loew, 1871

= Stenopogon costarus Loew, in Mouna 1998: 84

Séguy 1930a, MA, Tizi-n’Tkrine (Jebel Ahmar, 1700 m); Mouna 1998

Stenopogon gracilis (Macquart, 1838)

= Stenopogon fumipenis Becker, in Becker and Stein 1913: 68

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007

Stenopogon heteroneurus (Macquart, 1838)

Timon-David 1951, AP, Forêt Maâmora, Oued Akreuch, HA, Mouldikht; Mouna 1998; Hayat et al. 2008; Geller-Grimm 2007; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014

Stenopogon iphippus Séguy, 1932

Séguy 1932b, MA, Volubilis; Mouna 1998; Geller-Grimm 2007

Stenopogon iphis Séguy, 1932

Séguy 1932b, MA, Azrou; Timon-David 1951, Rif, Plateau de Tisserouine (2000 m), MA, Ifrane (1650 m), Ito (Rabat); Geller-Grimm 2007

Stenopogon ischyrus Séguy, 1932

Séguy 1932b, MA, Tizi-s’Tkrine (Jebel Ahmar, 1700 m); Mouna 1998; Geller-Grimm 2007

Stenopogon junceus (Wiedemann in Meigen, 1820)

Timon-David 1951, AP, Oued Akreuch, Zaër, MA, Sefrou; Mouna 1998; Geller-Grimm 2007; Hayat et al. 2008; Koçak and Kemal 2010; Ghahari et al. 2014

Stenopogon kocheri Timon-David, 1951

Timon-David 1951, HA, Tifni; Mouna 1998; Geller-Grimm 2007

Stenopogon porcus Loew, 1871

Séguy 1949a, AA, Akka; Mouna 1998; Geller-Grimm 2007

Stenopogon werneri Engel, 1933

Geller-Grimm 2007; Koçak and Kemal 2010, MA, Fès, Zalagh

Sisyrnodytes Loew, 1856

Sisyrnodytes leucophaetus Séguy, 1930

Séguy 1930a, MA, Béni Berberi; Mouna 1998; Geller-Grimm 2007; Londt 2009

Sisyrnodytes nilicola (Rondani, 1850)

Oldroyd 1980; Londt 1987; Geller-Grimm 2007; Londt 2009, AA, Ifni, Tiznit, Tata, Tazegzout; El Hawagry 2011; Samin et al. 2011; Ghahari et al. 2014

Stichopogoninae

Stichopogon Loew, 1847

Stichopogon albellus Loew, 1856

Séguy 1949a, AA, Foum-el-Hassan; Mouna 1998

Stichopogon albofasciatus (Meigen, 1820)

Séguy 1930a, HA, Kasba Taguendaft (Goundafa); Mouna 1998; Geller-Grimm 2007

Stichopogon elegantulus (Wiedemann, 1820)

Séguy 1930a, HA, Kasba Taguendaft (Goundafa), Aguerd el Had, AA, Talekjount (Souss); Mouna 1998; Geller-Grimm 2007; Ghahari et al. 2007; Koçak and Kemal 2010; El Hawagry 2011; Ghahari et al. 2014; AP (Kénitra) – MISR

Stichopogon inaequalis Loew, 1847

Séguy 1930a, HA, Aguerd el Had, AA, Talekjount (Souss); Mouna 1998; Geller-Grimm 2007

Stichopogon maroccanus (Becker, 1914)

Becker and Stein 1913, Rif, Tanger; Geller-Grimm 2007; El Hawagry 2011, Rif, Tanger

Stichopogon punctiferus Bigot, 1878

Geller-Grimm 2007

Stichopogon pusio (Macquart in Lucas, 1849)

Séguy 1930a, HA, Kasba Taguendaft (Goundafa); Mouna 1998; Geller-Grimm 2007; El Hawagry 2011

Stichopogon schineri Koch, 1872

Timon-David 1951, AA, Backkoum (Jebel Siroua); Mouna 1998; Hayat et al. 2008; Geller-Grimm 2007; Koçak and Kemal 2010; Ghahari et al. 2014

BOMBYLIIDAE

K. Kettani, M.J. Ebejer, J. Dils

Number of species: 248. Expected: 270

Faunistic knowledge of the family in Morocco: good

Usiinae

Apolysis Loew, 1860

Apolysis eremophila Loew, 1873

= Usia tomentosa Engel, in Paramonow 1947: 209

= Parageron ornata Engel, in Zaitzev 2007: 160

Paramonow 1947; Mouna 1998; Zaitzev 2007, AP (south), Tamri; Koçak and Kemal 2010; El Hawagri 2011; Evenhuis and Greathead 2015

Parageron Paramonov, 1929

Parageron gratus (Loew, 1856)

= Usia grata Loew, in Timon-David 1951: 143

Timon-David 1951, AP, Oued Grou; Mouna 1998; Bader and Arabyat 2004; Zaitzev 2007, SA; Koçak and Kemal 2010; El Hawagri 2011; Evenhuis and Greathead 2015 – MISR

Parageron griseus Paramonov, 1947

Zaitzev 2007, SA

Parageron hyalipennis (Séguy, 1941)

= Oligodranes hyalipennis Séguy, in Séguy 1941d: 9

Séguy 1941d, AA, Agadir (Forêt Admine); Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015, HA

Parageron incisus (Wiedemann, 1830)

= Usia incisa Wiedemann, in Timon-David 1951: 143

Séguy 1930a, AP, Mogador, Casablanca, Sidi Bettache, Aïn Sferguila, MA, Forêt Zaers, Ras el Ma, Aïn Leuh, HA, Tenfecht; Timon-David 1951, AP, Rabat, Sehoul, MA, Ifrane; Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Parageron major Macquart, 1840

Mouna 1998; Pârvu and Zaharia 2007; Zaitzev 2007, AP, Rabat, Nkheila; Evenhuis and Greathead 2015

Usia Latreille, 1802

Usia (Micrusia) aurata (Fabricius, 1794)

= Usia (Micrusia) taeniolata Costa, 1883, in, Koçak and Kemal 2010

Séguy 1930a, Rif, Tanger, AP, Rabat, Sidi Bettache; Paramonov 1950, Rif, Tanger; Mouna 1998; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015

Usia (Micrusia) crispa Gibbs, 2011

Gibbs 2011, HA, Marrakech, AA, Agadir, Taroudant, Tafraoute, Ouarzazate

Usia (Micrusia) cryptocrispa Gibbs, 2011

Gibbs 2011, AP, Ben Slimane, Rabat

Usia (Micrusia) dilsi Gibbs, 2011

Gibbs 2011, Rif, Tétouan, Al Hoceima, HA, Taourirt

Usia (Micrusia) echinus Gibbs, 2011

Gibbs 2011, AP, Agadir, Guelmim, Sidi Ifni, Cap Ghir, MA, Tafraoute, HA, Marrakech, Tizi-n-Test, AA, Tafingoult

Usia (Micrusia) falcata Gibbs, 2011

Gibbs 2011, MA, Azrou, Ifrane

Usia (Micrusia) forcipata Brullé, 183314

Mouna 1998: 84

Usia (Micrusia) globicauda Gibbs, 2011

Gibbs 2011, AP, Essaouira

Usia (Micrusia) loewi Becker, 1906

Zaitzev 2007, Rif, Tanger, AP, Skhirate, Nkheila, MA, Taferiate, HA, Taourirt

Usia (Micrusia) novakii Strobl, 1902

Séguy 1941d, HA, Tizi-n’Test; Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Usia (Micrusia) parascripa Gibbs, 2011

Gibbs 2011, MA, Ifrane, Mischliffen (2200 m)

Usia (Micrusia) pusilla Meigen, 1820

Séguy 1930a, AP, Rabat, MA, Azrou, HA, Tafingoult (Goundafa, 1500–1600 m); Séguy 1934b, AP, Rabat (on Calendula); Séguy 1953a, AP, Cap Ghir; Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Usia (Micrusia) versicolor (Fabricius, 1787)

Séguy 1930a, AP, Berrchid, Casablanca, M’Rassine; Timon-David 1951, AP, Rabat, MA, Oulmès; Mouna 1998; Koçak and Kemal 2010; Gibbs 2011, HA; Evenhuis and Greathead 2015; EM (Oujda) – MNHNR

Usia (Usia) aenea (Rossi, 1794)

Séguy 1949a, AA, Foum-el-Hassan; Mouna 1998; Bader and Arabyat 2004; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Usia (Usia) angustifrons Becker, 1906

Mouna 1998

Usia (Usia) atrata (Fabricius, 1798)

= Voluccella atrata Fabricius, in Fabricius 1798: 570

= Usia claripennis Macquart, in Macquart 1840: 105

Meigen 1820; Séguy 1930a, Rif, Tanger, MA, Aïn Leuh; Timon-David 1951, AP, Rabat, Guerrouaou; Mouna 1998; Zaitzev 2007, HA, Tizi-n’Tichka; Koçak and Kemal 2010; Gibbs 2014, AP, Mogador, Arbaa-Sahel (320 m), Tamri (215 m), MA, Khemisset, Oulmès (700 m), Ifrane, El Merabtine, HA, Marrakech, Aït Ourirr (530 m), Oukaimeden (2200 m), Tizi-n’Test (1450 m), Timzit (1700 m), AA, Agadir, Tiznit, Igherm (1660 m), Taroudant, Tata, Iguiour (1260 m), SA, Bou Jarif, Goulimine; Evenhuis and Greathead 2015

Usia (Usia) bicolor Macquart, 185515

Mouna 1998: 84

Usia (Usia) cornigera Gibbs, 2014

Gibbs 2014, Rif, Tanger, AP, Sidi Bettache, Rabat, MA, Meknès (550 m), Aïn Leuh (1350 m), HA, Dar Kaid M’tougui

Usia (Usia) florea (Fabricius, 1794)

= Volucella florea Fabricius, in Becker 1906: 203

= Usia cuprea Macquart, 1834, in Becker 1906: 203

Becker 1906a; Séguy 1930a, Rif, Tanger, AP, Mogador, Sidi Bettache, HA, Tinmel (Goundafa), around (Skoutana); Timon-David 1951, EM, Oued Moulouya; Mouna 1998; Pârvu and Zaharia 2007; Koçak and Kemal 2010; Gibbs 2014; Evenhuis and Greathead 2015

Usia (Usia) ignorata Becker, 1906

Becker 1906a; Mouna 1998; Bader and Arabyat 2004; Pârvu and Zaharia 2007; Koçak and Kemal 2010; El Hawagry 2011

Usia (Usia) maghrebensis Gibbs, 2014

Gibbs 2014, Rif, Tanger, Tétouan, El Biutz (150 m), AP, Mogador, MA, Aïn Leuh (1350 m)

Usia (Usia) vestita Macquart, 1846

Mouna 1998: 84; Gibbs 2011

Phthiriinae

Phthiria Meigen, 1802

Phthiria albogilva Séguy, 1941

Séguy 1941d, AA, Agadir; Mouna 1998; Evenhuis and Greathead 2015

Phthiria gaedii Wiedemann in Meigen, 1820

Séguy 1930a, MA, Foum Keneg; Timon-David 1951, AP, Zaers, MA, Ifrane; Mouna 1998; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015 – MISR (MA, Ifrane)

Phthiria maroccana Zaitzev, 2005

= Phthiria maroccana Zaitzev, in Zaitzev 2005: 667

Zaitzev 2005, MA, Taferiate, HA, Taourirt; Zaitzev 2007, MA, Taferiate, HA, Taourirt

Phthiria merlei Zaitzev, 2005

= Phthiria merlei Zaitzev, in Zaitzev 2005: 665

Zaitzev 2005, AP (south), Tamri, Inchaden (south of Aït Melloul); Zaitzev 2007, AP (south), Tamri, Inchaden (south of Aït Melloul)

Phthiria minuta (Fabricius, 1805)

Séguy 1930a, HA, Tenfecht, AA, Souss; Mouna 1998; Koçak and Kemal 2010; El Hawagry 2011

Phthiria pulicaria var. flavofasciata Strobl in Morge, 1976

Mouna 1998; Zaitzev 2007, AA, Tizi-n’Tiniggigt (1600 m); Koçak and Kemal 2010; Evenhuis and Greathead 2015

Phthiria scutellaris Wiedemann in Meigen, 1820

Séguy 1930a, MA, Meknès; Séguy 1941a, MA, Meknès, HA, Imi-n’Ouaka (1500 m); Mouna 1998; Evenhuis and Greathead 2015; Rif (Sapinière Talassemtane) – MISR

Phthiria simonyi Becker, 1908

Séguy 1949a, SA, Guelmim; Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015, MA, Meknès

Phthiria umbripennis Loew, 1846

Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015, MA, Meknès

Phthiria vagans Loew, 1846

Zaitzev 2007, HA, Taourirt, AA, Tizi-n’Taratine

Toxophorinae

Geron Meigen, 1820

Geron intonsus Bezzi, 1925*

MA, HA

Geron macquarti Greathead in Evenhuis & Greathead 1999

Dils and Özbek 2006; Koçak and Kemal 2010; El Hawagry 2011

Geron subflavofemoratus Andréu Rubio, 1959

Andréu Rubio 1959; Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Toxophora Meigen, 1803

Toxophora fasciculata (Villers, 1789)

Séguy 1930a, AP, Rabat; Mouna 1998; Dils and Özbek 2006; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015; AP (Rabat) – MISR

Toxophora fuscipennis (Macquart, 1840)

Mouna 1998: 84

Toxophora pauli Zaitzev, 2005

Zaitzev 2005, AA, Ouarzazate, Jebel Tighermine (SE of Ouarzazate); Zaitzev 2007, AA, Ouarzazate, Jebel Tighermine (SE of Ouarzazate)

Toxophora shelkovnikovi Paramonov, 1933

Zaitzev 2007, AA, Ouarzazate

Heterotropinae

Heterotropus Loew, 1873

Heterotropus atlanticus Séguy, 1930

Séguy 1930a, AP, Mogador; Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Heterotropus longitarsus Séguy, 1930

Koçak and Kemal 2010; Evenhuis and Greathead 2015

Heterotropus maroccanus Zaitzev, 2003

Zaitzev 2003, AA, Jebel Tighermine (SE of Ouarzazate); Zaitzev 2007, AA, Jebel Tighermine (SE of Ouarzazate)

Bombyliinae

Anastoechus Osten-Sacken, 1877

Anastoechus bahirae Becker, 1915

Mouna 1998; Zaitzev 2007, AA, Jebel Tighermine (SE of Ouarzazate)

Anastoechus hyrcanus Pallas & Wiedemann in Wiedemann, 1818

Mouna 1998: 84

Anastoechus latifrons (Macquart, 1839)

Timon-David 1951, AP, Dradek; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Anastoechus nitidulus ssp. nitidulus Fabricius, 1794

Mouna 1998: 84

Anastoechus stramineus Wiedemann in Meigen, 1820

Mouna 1998: 84

Anastoechus trisignatus (Portschinsky, 1881)

Bader and Arabyat 2004; Ziatzev 2007, AP, Rabat, AA, Jebel Tighermine (SE of Ouarzazate), AA, 20 km SW Goulmima, SA, Tan-Tan, Between Guelmim and Tan-Tan (90 km from Guelmim), Taganint (south of Bou-Izakarn); Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015

Bombomyia Greathead, 1995

Bombomyia discoidea (Fabricius, 1794)

Séguy 1930a, AP, Oued Korifla (Zaers), Sidi Bettache; Bader and Arabyat 2004; Dils and Özbek 2006; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Bombomyia stictica Boisduval, 1835

Zaitzev 2007, MA, col de Zeggota (N Meknès), Oulmès

Bombomyia vertebralis (Dufour, 1833)

= Bombylius punctatus Fabricius, in Timon-David 1951: 144

Becker and Stein 1913, Rif, Tanger; Séguy 1930a AP, Dradek near Rabat; Timon-David 1951, MA, Volubilis; Mouna 1998; Bader and Arabyat 2004; Koçak and Kemal 2010 Evenhuis and Greathead 2015; AP (Dradek, Casablanca), EM (Oujda), MA (Volubilis, Aïn Leuh) – MISR

Bombylisoma Rondani, 1856

Bombylisoma algirum (Macquart, 1840)

= Bombylius nigrifrons Becker, in Becker and Stein 1913: 83

Becker and Stein 1913, Rif, Tanger; Mouna 1998; Zaitzev 2007, Rif, Tanger; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Bombylisoma breviusculum (Loew, 1855)

Dils and Özbek 2006; Zaitzev 2007; Evenhuis and Greathead 2015

Bombylisoma flavibarbum Loew, 1855

Mouna 1998: 84

Bombylisoma melanocephalum Fabricius, 1794

Zaitzev 2007, HA, Taourirt, south of Tizi-n’Test

Bombylius Linnaeus, 1758

Bombylius (Bombylius) albaminis Séguy, 1949

Séguy 1949a, HA, Alnif; Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Bombylius (Bombylius) ambustus Pallas & Wiedemann, 1818

Mouna 1998: 84

Bombylius (Bombylius) analis (Olivier, 1789)

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, AP, Oued Korifla, Rabat, Sidi Bettache, Aïn Sferguila; Timon-David 1951, AP, Rabat; Mouna 1998; Zaitzev 2007, MA, route Fès-Sidi Kacem (30 km from Fès); Koçak and Kemal 2010; Evenhuis and Greathead 2015, AP (Rabat, Casablanca) – MISR

Bombylius (Bombylius) audcenti Bowden, 1984

Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Bombylius (Bombylius) canescens Mikan, 1796

Becker and Stein 1913, Rif, Tanger; Mouna 1998; Zaitzev 2007, Rif, Cap Malabata (Tanger), MA, Tachguelt, route Fès-Sidi Kacem (30 km from Fès)

Bombylius (Bombylius) cinerascens Mikan, 1796

Mouna 1998; Bader and Arabyat 2004

Bombylius (Bombylius) discolor Mikan, 1796

Mouna 1998; Zaitzev 2007, MA, route El Hajeb-Ifrane (1 km from Ifrane)

Bombylius (Bombylius) eploceus Séguy, 1949

Séguy 1949a, SA, Guelmim; Mouna 1998; Evenhuis and Greathead 2015

Bombylius (Bombylius) fimbriatus Meigen, 1820

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA, Tizi-s’Tkrine, Jebel Ahmar (1700 m); Mouna 1998; Pârvu and Zaharia 2007; Zaitzev 2007; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015

Bombylius (Bombylius) flavipes Wiedemann, 1828

Becker and Stein 1913, Rif, Tanger; Mouna 1998; Zaitzev 2007; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015

Bombylius (Bombylius) fulvescens Wiedemann in Meigen, 1820

Becker and Stein 1913, Rif, Tanger; Séguy 1953a, AP, Cap Ghir; Séguy 1941d, AA, Agadir; Mouna 1998

Bombylius (Bombylius) fuscus Fabricius, 1781

Mouna 1998: 84

Bombylius (Bombylius) major (Linnaeus, 1758)

Becker and Stein 1913, Rif, Tanger; Séguy 1953a, MA, Oulmès; Bader and Arabyat 2004; Zaitzev 2007; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015; AP (Kénitra) – MISR

Bombylius (Bombylius) mauritanus Olivier, 1789

Koçak and Kemal 2010; Evenhuis and Greathead 2015, HA

Bombylius (Bombylius) medius (Linnaeus, 1758)

Becker and Stein 1913, Rif, Tanger; Timon-David 1951, AP, Sehoul; Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; Pârvu and Zaharia 2007; Zaitzev 2007; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015; AP (Oued Yquem, Dradek) – MISR

Bombylius (Unplaced) megacephalus Portschinsky, 1887*

EM, AA

Bombylius (Bombylius) minor Linnaeus, 1758

Mouna 1998; Zaitzev 2007, Rif, Tanger; MA (Aguelmane Azigza), SA

– MISR

Bombylius (Bombylius) mus Bigot, 1862

Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015

Bombylius (Bombylius) niveus Meigen, 1804

Mouna 1998: 84; AP (Mogador) – MISR

Bombylius (Bombylius) numidus Macquart, 1846

Séguy 1953a, MA, Ifrane; El Hawagry 2011; Evenhuis and Greathead 2015

Bombylius (Bombylius) pauli Zaitzev, 2003

Zaitzev 2003, MA, route Fès-Sidi Kacem (30 km from Fès); Zaitzev 2007, MA, route Fès-Sidi Kacem (30 km from Fès)

Bombylius (Bombylius) posticus (Fabricius, 1805)

Dils and Özbek 2006; Zaitzev 2007; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015

Bombylius (Bombylius) postversicolor Evenhuis & Greathead, 1999

= Bombylius versicolor Fabricius, 1805

Meigen 1820; Bezzi 1906; Séguy 1930a, AP, Mogador; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Bombylius (Bombylius) pumilus Meigen, 1820

Mouna 1998: 84

Bombylius (Bombylius) semifuscus (Meigen, 1820)

Séguy 1953a, AP, Cap Ghir; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Bombylius (Bombylius) torquatus Loew, 1855

Séguy 1930a, HA, Ouaounzert (Glaoua), Arround (Skoutana), Tachdirt (bord de l’Imminen, 2400–2600 m); Timon-David 1951, AP, Rabat; Mouna 1998; Evenhuis and Greathead 2015; AP (Mogador) – MISR

Bombylius (Bombylius) undatus Mikan, 1796

Pârvu and Zaharia 2007

Bombylius (Bombylius) vagans Meigen, 1830

Koçak and Kemal 2010; Evenhuis and Greathead 2015

Bombylius (Bombylius) venosus Mikan, 1796

Mouna 1998; Zaitzev 2007, AP, El Koudia (30 km SW from Rabat); AP (Dradek) – MISR

Bombylius (Zephyrectes) cruciatus Fabricius, 1798

Séguy 1930a, MA, Aharmoumou (1100 m), Azrou, Ras el Ma, HA, Tizi-n’Test, Jebel Imdress (Goundafa, 2000–2450 m); Mouna 1998; Zaitzev 2007; Koçak and Kemal 2010; Evenhuis and Greathead 2015; Rif (Talassemtane), AP (Mogador), MA (Sefrou) – MISR

Bombylius (Zephyrectes) leucopygus (Macquart, 1846)

Becker and Stein 1913, Rif, Tanger; Mouna 1998; Zaitzev 2007, AP, Larache, MA, Moulay Idris (900 m); Evenhuis and Greathead 2015, SA, Erfoud; MA (Ifrane) – MISR

Conophorus Meigen, 1803

Conophorus bellus Becker, 1906*

HA

Conophorus fuliginosus (Wiedemann in Meigen, 1820)

= Ploas fuliginisa (Meigen), in Séguy 1953a: 83

Séguy 1930a, MA, Aharmoumou (1100 m); Séguy 1953a, MA, Ahermoumou (1100 m); Timon-David 1951, AP, Dradek, MA, Sefrou, HA, Marrakech; Mouna 1998; Zaitzev 2007, Rif, Tanger; Evenhuis and Greathead 2015; AP (Salé, Mogador) – MISR

Conophorus fuscipennis (Macquart, 1840)

Séguy 1930a, HA, Tizi-n’Test, Jebel Imdress (2000–2450 m), Goundafa; Mouna 1998; Evenhuis and Greathead 2015

Conophorus griseus (Fabricius, 1787)

Mouna 1998; Zaitzev 2007; Evenhuis and Greathead 2015

Conophorus hamilkar Paramonov, 1929

Timon-David 1951, AP, Mogador; Mouna 1998; Evenhuis and Greathead 2015; AP (Mogador) – MISR

Conophorus macroglossus (Dufour, 1852)

Mouna 1998; Zaitzev 2007; Evenhuis and Greathead 201; AP (Mogador) – MISR

Conophorus mauritanicus Bigot, 1892

= Conophorus heteropilosus Timon-David, in Timon-David 1951: 141; Mouna 1998: 84; Evenhuis and Greathead 2015: 192

Timon-David 1951, MA, Oulmès; Mouna 1998; Zaitzev 2007, AP, El Koudia (30 km SW from Rabat), Forêt Zaer (35 km SW from Rabat), N Tretten; Koçak and Kemal 2010; Dils 2013, MA, Mrirt; Evenhuis and Greathead 2015

Conophorus rossicus Paramonow, 1929

Dils and Özbek 2006

Dischistus Loew, 1855

Dischistus albatus (Séguy, 1934)

= Acanthogeron albatus Séguy, 1934, in Séguy 1934d: 73; Zaitzev 2007: 162

Séguy 1934d; Zaitzev 2007, SA, 30 km S Tata

Dischistus auripilus (Séguy, 1930)

= Acanthogeron auripilus Séguy, 1930, in Séguy 1930a: 104

Séguy 1930a, AP, Mogador; Séguy 1934b, AP, Zaers; Timon-David 1951, AP, Oued Korifla; Mouna 1998; Evenhuis and Greathead 2015

Dischistus maroccanus (Séguy, 1930)

= Acanthogeron maroccanus Séguy, 1930, in Séguy 1930a: 106

Séguy 1930a, AP, Mogador; Mouna 1998; Zaitzev 2007, HA, Tazzarine; Evenhuis and Greathead 2015

Dischistus mittrei (Séguy, 1930)

= Acanthogeron mittrei Séguy, in Séguy 1930a: 105

Séguy 1930a, AP, Mogador; Mouna 1998; Evenhuis and Greathead 2015

Dischistus perniveus (Bezzi, 1925)

= Acanthogeron perniveus Bezzi, in Timon-David 1951: 143

Timon-David 1951, AP, Djamda de M’Tal; Mouna 1998; El Hawagry 2011; Evenhuis and Greathead 2015

Dischistus senex (Wiedemann in Meigen, 1820)

= Acanthogeron senex Meigen, 1820, in Séguy 1953a: 83, Mouna 1998: 84, Zaitzev 2007: 162

Séguy 1930a, HA, Tafingoult (Goundafa, 1500–1600 m); Villeneuve 1933; Séguy 1953a, HA, Aït Ourir; Mouna 1998; Zaitzev 2003, HA, Taourirt; Zaitzev 2007, HA, Taourirt; El Hawagry 2011; Evenhuis and Greathead 2015; AP (Dradek), MA (Sefrou) – MISR

Dischistus separatus (Becker, 1906)

= Acanthogeron talboti Séguy, 1930, in Séguy 1930a: 106

Evenhuis and Greathead 2015

Efflatounia Bezzi, 1925

Efflatounia berbera Bowden, 1973

Ebejer et al. 2019, AA, Agadir – NMWC

Legnotomyia Bezzi, 1902

Legnotomyia fascipennis Bezzi, 1925*

SA

Merleus Zaitzev, 2003

Merleus punctipennis Zaitzev, 2003

= Merleus punctipennis Zaitzev 2003: 599

Zaitzev 2003, AP, Skhirate; Zaitzev 2007, AP, Skhirate

Prorachthes Loew, 1869

Prorachthes crassipalpis Villeneuve, 1930

Evenhuis and Greathead 2015

Systoechus Loew, 1855

Systoechus ctenopterus (Mikan, 1796)

Timon-David 1951, MA, Ifrane; Mouna 1998; Dils and Özbek 2006; Zaitzev 2007; El Hawagry 2011; Evenhuis and Greathead 2015

Systoechus gomezmenori Andréu Rubio, 1959

Carles-Tolrá 2002; Evenhuis and Greathead 2015

Systoechus gradatus (Wiedemann in Meigen, 1820)

Timon-David 1951, AP, Mouldikht; Mouna 1998; Zaitzev 2007, MA, Taferiat, HA, Taourirt; El Hawagry 2011; Evenhuis and Greathead 2015

Systoechus mixtus Wiedemann, 1821

= Bombylius stylicornis Macquart in Séguy 1941: 10

Séguy 1941d, AA, Agadir; Mouna 1998

Systoechus pumilio Becker, 1915

Mouna 1998: 84

Triplasius Loew, 1855

Triplasius boghariensis (Lucas, 1852)

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, EM, Oujda; Mouna 1998; Pârvu and Zaharia 2007; Evenhuis and Greathead 2015

Triplasius maculipennis (Macquart, 1846)

= Bombylius maculipennis var. melanopus Timon-David, in Timon-David 1951: 144

= Bombylius (Triplasius) maculipennis Macquart, 1849, in Zaitzev 2007: 166

Timon-David 1951, MA, Azrou; Zaitzev 2007, MA, route El Hachef, Criosement route Raubei Idris-Merhassine; Evenhuis and Greathead 2015

Ecliminae

Eclimus Loew, 1844

Eclimus gracilis Loew, 1844

Séguy 1930a, MA, Ras el Ma; Timon-David 1951, AP, Oued Korifla; Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; Zaitzev 2007, MA, Maaziz; Evenhuis and Greathead 2015

Thevenetimyia Bigot, 1875

Thevenetimyia quedenfeldti (Engel, 1885)*

Rif, AP, MA

Crocidiinae

Crocidium Loew, 1860

Crocidium aegyptiacum Bezzi, 1925*

SA

Crocidium nudum Efflatoun, 1945*

EM, AA

Semiramis Becker in Becker and Stein 1914

Semiramis punctipennis Becker, 1914

Zaitzev 2007, AA, Aoulouz

Cythereinae

Amictus Wiedemann, 1817

Amictus castaneus (Macquart, 1840)

Séguy 1930a, AP, Rabat, HA, Ank el Djemal; Mouna 1998; Evenhuis and Greathead 2015

Amictus compressus (Fabricius, 1805)

Evenhuis and Greathead 2015

Amictus heteropterus Macquart, 1838

Zaitzev 2007, Rif, Tanger, AP, Rabat, HA, S Tizi-n’Test, AA, Tizi-n’Taratine

Amictus oblongus (Fabricius, 1805)

= Bombylius oblongus Fabricius, in Macquart 1834: 390

Macquart 1834

Amictus pulchellus Macquart, 1846

Séguy 1930a, AP, Rabat, Maâmora; Mouna 1998; Zaitzev 2007, HA, Taourirt; El Hawagry 2011; Evenhuis and Greathead 2015; AP (Rabat) – MISR

Amictus setosus Loew, 1869*

AP

Amictus tener Becker, 1906

Zaitzev 2007, AP, Rabat

Amictus validus Loew, 1869

Bader and Arabyat 2004; Dils and Özbek 2006; Karimpour 2012; Evenhuis and Greathead 2015

Amictus variegatus Meigen in Waltl, 1835

Mouna 1998: 84

Chalcochiton Loew, 1844

Chalcochiton argentifrons (Macquart in Lucas, 1849)

Séguy 1953a, AP, Cap Ghir, Salé, Sidi Battache, MA, Tizi-n’Bou Zabal (2300 m), AA, Aïn Chaïb (Souss); Evenhuis and Greathead 2015

Chalcochiton argyrocephalus (Macquart, 1840)

= Chalcochiton (Anthrax) argyrocephala (Macquart), in Engel 1938: 328

Engel 1938; Séguy 1953a, AA, Agadir; El Hawagry 2011; Evenhuis and Greathead 2015

Chalcochiton atlantica Dils, 2008

Dils 2008, SA, Guelmim

Chalcochiton holosericeus (Fabricius, 1794)

= Chalcochiton semiargentaea Macquart, in Zaitsev 2007: 172

Séguy 1930a, AP, Maâmora, Sidi Bettache, HA, Tizi-n’Test, Jebal Imdress (2000–2450 m), Tafingoult (Goundafa, 1500–1600 m); Séguy 1941d; Mouna 1998; Zaitzev 2007, Rif, Tanger, AP, Skhirate, EM, Taourirt, MA, Taferiat, Meknès-Moulay Idriss, Merhassine, AA, Agadir, Ouarzazate; Evenhuis and Greathead 2015; AP (Salé, Forêt Temara, Oued Yquem, Meshra) – MISR

Chalcochiton maghrebi Dils, 2017

Dils 2017, Rif, Souk El Kolla, Bab Taza, 10 km S of Mjara, AP, Sidi Bettache, Temsia, Imsouane, Mansouria, Rommani, Béni Slimane, Tioulit, EM, El Aioun, MA, Béni Mellal, el Ksiba, 10 km SE Bir Tamtam, Merchouch, Mrirt, Fès, HA, Azilal, Asni, Tizi-Mlil, AA, Taroudant, Tizi-n’Test, Tiznit, Agouim, Sidi Ifni, Mesti, Tafinegoult, Tizi-n’Tinififft, El Mrabtine, SA, Semara

Chalcochiton maroccanus Zaitzev, 2006

Séguy 1953a, HA, Tafingoult (Goundafa, 1500–1600 m); Zaitzev 2006, AP (south), Aït Melloul; Zaitzev 2007, AP (south), Aït Melloul

Chalcochiton merlei Zaitzev, 2006

Zaitzev 2006, AP, Skhirate; Zaitzev 2007, AP, Skhirate

Chalcochiton pallasii Loew, 1856

Bader and Arabyat 2004; Dils and Özbek 2006; Zaitzev 2007; Karimpour 2012; Evenhuis and Greathead 2015

Callostoma Macquart, 1840

Callostoma fascipenne Macquart, 1840

Bader and Arabyat 2004

Cyllenia Latreille, 1802

Cyllenia rustica Rossi, 1790

Mouna 1998; Zaitzev 2007; AP (Mogador) – MISR

Cytherea Fabricius, 1794

Cytherea albolineata Bezzi, 1925*

SA

Cytherea alexandrina Becker, 1902

Becker 1902: 30; Zaitzev 2007, AA, Jebel Tighermine (SE of Ouarzazate)

Cytherea aurea (Fabricius, 1794)

Séguy 1930a, AP, Rabat, HA; Mouna 1998; Bader and Arabyat 2004; Zaitzev 2007, AA, Tizi-n’Taratine; El Hawagry 2011; Evenhuis and Greathead 2015, HA, Tafingoult (Goundafa, 1500–1600 m); AP (Rabat, Oued Cherrat) – MISR

Cytherea cinerea Fabricius, 1805

= Mulio delicatus Becker, 1906

Becker 1906b: 153; Timon-David 1951, AP, Meshra; Mouna 1998; Bader and Arabyat 2004; El Hawagry 2011; Evenhuis and Greathead 2015

Cytherea delicata Becker, 1906

Zaitzev 2007, HA, S Tizi-n’Test, AA, Zagora, Taroudant, Tizi-n’Taratine

Cytherea dispar (Loew, 1873)

Bader and Arabyat 2004; Dils and Özbek 2006; Evenhuis and Greathead 2015

Cytherea fenestrata (Loew, 1873)

Bader and Arabyat 2004; Evenhuis and Greathead 2015

Cytherea infuscata (Meigen, 1820)

Séguy 1930a, EM, Itzr (Haute Moulouya), MA, Forêt Timelilt (1900 m), HA, Aït el Hadj, Marrakech; Mouna 1998; Evenhuis and Greathead 2015; AP (Meskara) – MISR

Cytherea maroccana (Becker, 1903)

= Mulio maroccanus Becker, in Becker 1903: 89

Becker 1903, Rif, Tanger; Bezzi 1906: 249; Timon-David 1951, AP, Azemmour; Mouna 1998; El Hawagry 2011; Evenhuis and Greathead 2015

Cytherea obscura Fabricius, 1794

Séguy 1930a, EM, Haute Moulouya, AP, Sidi Bettache, HA, Ouaouenzert; Séguy 1941d; Mouna 1998; Dils and Özbek 2006; Zaitzev 2007, MA, Taferiat, AA, Agadir, Amredi, Jebel Tighermine (SE of Ouarzazate), Tizi-n’Tiniggigt, Tizi-n’Taratine, Tizi-n’Bachkoun; Karimpour 2012; Evenhuis and Greathead 2015

Cytherea rungsi Timon-David, 1951

Timon-David 1951, EM, Guenfouda; Mouna 1998; Evenhuis and Greathead 2015

Cytherea thyridophora (Bezzi, 1925)

Ebejer et al. 2019, Rif, Moulay Abdelsalam (Jebel Bouhachem, 965 m)

Cytherea trifaria (Becker, 1906)

Evenhuis and Greathead 2015

Lomatiinae

Lomatia Meigen, 1820

Lomatia abbreviata Villeneuve, 1911

Séguy 1930a, MA, Forêt Zaers; Timon-David 1951, EM, Guercif; Mouna 1998; Bader and Arabyat 2004; Evenhuis and Greathead 2015; AP (Maâmora, Oued Cherrat, Dradek), HA – MISR

Lomatia belzebul paramonovi Fabricius, 1794

Séguy 1930a, AP, Dar Salem, MA, Timhadit, Meknès, Aïn Leuh; Mouna 1998; Dils and Özbek 2006; Zaitzev 2008; Karimpour 2012; Evenhuis and Greathead 2015

Lomatia erynnis (Loew, 1869)

Mouna 1998; Dils and Özbek 2006; Zaitzev 2008, AP, Rabat; Evenhuis and Greathead 2015

Lomatia hamifera Becker, 1915

Mouna 1998: 84

Lomatia lachesis Egger, 1859

Dils and Özbek 2006; Evenhuis and Greathead 2015

Lomatia lateralis (Meigen, 1820)

Séguy 1930a, MA, Ras el Ma, HA, Forêt Timelilt; Mouna 1998; Evenhuis and Greathead 2015; AP (Rabat), MA (Volubilis, Ras el Ma) – MISR

Lomatia obscuripennis Loew, 1869

Zaitzev 2008, AP, Nkheila; Evenhuis and Greathead 2015

Lomatia sabaea (Fabricius, 1781)

Mouna 1998: 84

Lomatia tysiphone Loew, 1860

Zaitzev 2008, MA, Azrou, AA, Tizi-n’Taratine

Antoniinae

Antonia Loew, 1856

Antonia bouillonae Séguy, 1932

Evenhuis and Greathead 2015

Anthracinae

Aphoebantini

Aphoebantus Loew, 1872

Aphoebantus wadensis Becker, 1925*

SA

Anthracini

Anthrax Scopoli, 1763

Anthrax aethiops (Fabricius, 1781)

Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; Evenhuis and Greathead 2015; AP (Forêt Maâmora) – MISR

Anthrax anthrax (Schrank, 1781)

= Argyramoeba anthrax Schrank, in Séguy 1930a: 93

Séguy 1930a, MA, Aïn Leuh, Soufouloud (1900–2100 m), HA, Tizi-n’Test, Jebel Imdress (2000–2450 m), Goundafa; Timon-David 1951, MA, El Ksiba, Ifrane; Mouna 1998; Dils and Özbek 2006; Evenhuis and Greathead 2015

Anthrax binotatus (Wiedemann in Meigen, 1820)

= Argyramoeba binotata Meigen, in Séguy 1926: 209, Séguy 1930a: 94

Séguy 1926; Séguy 1930a, AP, Rabat, HA, Tizi-n’Test, Jebel Imdress (2000–2450 m); Séguy 1949a, HA, Alnif; Mouna 1998; Dils and Özbek 2006; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015; AP (Rabat) – MISR

Anthrax dentatus (Becker, 1906)

Bader and Arabyat 2004; Zaitzev 2008, AA, Tizi-n’Tiniggigt; El Hawagry 2011; Evenhuis and Greathead 2015

Anthrax hemimelas Speiser, 1910

Zaitzev 2008, AA, Ouarzazate, Jebel Tighermine (SE of Ouarzazate)

Anthrax kiritshenkoi Paramonov, 1935

Evenhuis and Greathead 2015

Anthrax lucidus (Becker, 1902)

Ebejer et al. 2019, AA, Ziz river (13 km N of Erfoud, 800 m)

Anthrax morio Fabricius, 1775

Mouna 1998; MA (Ifrane, Azrou) – MISR

Anthrax trifasciatus (Meigen, 1804)

= Argyramoeba trifasciata Meigen, in Timon-David 1951: 139

Séguy 1930a, MA, Meknès; Timon-David 1951, AP, south of Rabat; Mouna 1998; Dils and Özbek 2006; Koçak and Kemal 2010; El Hawagry 2011; Evenhuis and Greathead 2015

Anthrax varius Fabricius, 1794

Séguy 1930a, AP, Rabat; Mouna 1998; Evenhuis and Greathead 2015 – MISR

Anthrax virgo Egger, 1859

= Argyramoeba virgo Egger, in Séguy 1930a: 94

Séguy 1930a, AP, Rabat; Zaitzev 2008, MA, Taferiat, AA, Jebel Tighermine (SE of Ouarzazate)

Cononedys Hermann, 1907

Cononedys efflatouni (Bezzi, 1925)*

SA

Cononedys escheri Bezzi, 1908

Zaitzev 2008, AP, Skhirate, Rabat

Cononedys lyneborgi (François, 1969)

Evenhuis and Greathead 2015

Cononedys scutellatus Meigen, 1835

Zaitzev 2008, Rif, Jebala, Haouta el Kazdir, AA, Aouzlida near Aoulouz

Satyramoeba Sack, 1909

Satyramoeba hetrusca (Fabricius, 1794)

Mouna 1998: 84

Spogostylum Macquart, 1840

Spogostylum isis (Meigen, 1820)

Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; El Hawagry 2011; Karimpour 2012; Evenhuis and Greathead 2015

Spogostylum trinotatum Dufour, 1852

Mouna 1998: 84

Spogostylum tripunctatum (Pallas in Wiedemann, 1818)

Timon-David 1951, HA, Aït Mhamed Sgatt; Mouna 1998; Dils and Özbek 2006; Zaitzev 2008, AA, Jebel Tighermine (SE of Ouarzazate); El Hawagry 2011; Karimpour 2012; Evenhuis and Greathead 2015; HA (Aïn Mhamed Sgatt) – MISR

Turkmeniella Paramonov, 1940

Turkmeniella crosi (Villeneuve, 1910)

Evenhuis and Greathead 2015

Exoprosopa Macquart, 1840

Exoprosopa aeacus Meigen, 1804

Mouna 1998: 84

Exoprosopa baccha Loew, 1869

Mouna 1998; Zaitzev 1999; Dils and Özbek 2006; Zaitzev 2008; Evenhuis and Greathead 2015

Exoprosopa capucina (Fabricius, 1871)

Mouna 1998: 84

Exoprosopa circeoides Paramonov, 1928

Zaitzev 2008, AA, Jebel Tighermine (SE of Ouarzazate)

Exoprosopa cleomene Egger, 1859

Mouna 1998: 84

Exoprosopa decrepita (Wiedemann, 1828)

Zaitzev 2008, AA, Zagora

Exoprosopa efflatouni Bezzi, 1925

Zaitzev 2008, AA, Jebel Tighermine (SE of Ouarzazate), Ouarzazate, SA, Taganint (south of Bou-Izakarn)

Exoprosopa grandis Wiedemann in Meigen, 1820

Mouna 1998; Zaitzev 2008, HA, Tishka (2200 m)

Exoprosopa italica (Rossi, 1794)

Zaitzev 2008, HA, Taourirt, AA, Tizi-n’Taratine, Jebel Tighermine (SE of Ouarzazate), SA, Taganint (south of Bou-Izakarn)

Exoprosopa jacchus (Fabricius, 1805)

Séguy 1930a, AP, Mogador, Sidi Taibi, MA, Tizi-s’Tkrine (1700 m), Dar Salem, Aïn Leuh, HA, Bou Tazzert; Mouna 1998; Mirceni and Pârvu 2009; Evenhuis and Greathead 2015; Rif (Talassemtane, Forêt Izarine, road of Jebha, Zoumi) – MISR

Exoprosopa minos (Meigen, 1804)

Séguy 1949a, AA, Tata; Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; Zaitzev 2008, MA, Taferiat; El Hawagry 2011; El Hawagry and Dhafer 2015; Evenhuis and Greathead 2015; MA (Jebel Lachhab) – MISR

Exoprosopa pandora (Fabricius, 1805)

Greathead 2001; Bader and Arabyat 2004; Evenhuis and Greathead 2015

Exoprosopa rutila (Pallas & Wiedemann, 1818)

Evenhuis and Greathead 2015

Micomitra Bowden, 1964

Micomitra stupida Rossi, 1790

= Exoprosopa stupida Rossi, in Mouna 1998: 84

Mouna 1998

Plesiocera Macquart, 1840

Plesiocera algira (Macquart, 1840)

Zaitzev 2008, MA, Taferiat; Evenhuis and Greathead 2015

Heteralonia Rondani, 1863

Heteralonia (Homolonia) megerlei (Hoffmansegg in Wiedemann, 1818)

Zaitzev 2008, SA, Goulimine

Heteralonia (Mesoclis) pygmalion (Fabricius, 1805)

= Exoprosopa pygmalion Fabricius, in Timon-David 1951: 139

= Mesoclis pygmalion Fabricius, 1805, in Zaitzev 2008: 191

Séguy 1930a, Rif, Tanger, AP, Maâmora, Rabat, MA, Aïn Leuh; Timon-David 1951, AP, Temara; Mouna 1998; Zaitzev 2008, AP, Cherrat El Hawagry 2011; Evenhuis and Greathead 2015

Heteralonia (Zygodipla) algira (Fabricius, 1794)

Séguy 1930a, Rif, Tanger, AP, Mogador, HA, Bou Tazzert; Mouna 1998; Zaitzev 2008, HA, Tifnite (south of Aït Melloul); El Hawagry 2011; Evenhuis and Greathead 2015

Heteralonia (Zygodipla) bagdadensis (Macquart, 1840)

Zaitzev 2008, AA, Zagora

Heteralonia (Zygodipla) singularis (Macquart, 1840)

Bader and Arabyat 2004; Evenhuis and Greathead 2015

Heteralonia arenacea Becker, 1906

Evenhuis and Greathead 2015

Heteralonia dispar (Loew, 1869)

= Exoprosopa dispar Loew, in Timon-David 1951: 139

Timon-David 1951, HA, Marrakech; Mouna 1998; Dils and Özbek 2006; Evenhuis and Greathead 2015

Heteralonia rivularis (Meigen, 1820)

= Exoprosopa rivularis Meigen, in Timon-David 1951: 139

Séguy 1930a, AP, Rabat, Maâmora; Timon-David 1951, AP, Oued Akreuch; Mouna 1998; Zaitzev 1999; Bader and Arabyat 2004; Zaitzev 2008, AP, Rabat

Oestranthrax Bezzi, 1921

Oestranthrax brunnescens (Loew, 1857)

Bader and Arabyat 2004

Oestranthrax pallifrons Bezzi, 1926

Evenhuis and Greathead 2015

Pachyanthrax François, 1964

Pachyanthrax albosegmentatus (Engel, 1936)

Zaitzev 2008, AA, Jebel Tighermine (south of Ouarzazate)

Pachyanthrax nomadorum (Greathead, 1970)

Koçak and Kemal 2010; Evenhuis and Greathead 2015

Exhyalanthrax Becker, 1916

Exhyalanthrax afer (Fabricius, 1794)

= Anthrax tangerinus Bigot, 1892

Bezzi 1906; Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; Zaitzev 2008; El Hawagry 2011; Evenhuis and Greathead 2015, Rif, Tanger; MA (Ifrane) – MISR

Hemipenthes Loew, 1869

Hemipenthes morio (Linnaeus, 1758)

Séguy 1930a, MA, Azrou, HA, Arround (Skoutana, 2000–2400 m); Dils and Özbek 2006; Koçak and Kemal 2010; Karimpour 2012; Evenhuis and Greathead 2015; MA (Azrou, Ifrane) – MISR

Hemipenthes velutinus (Meigen, 1820)

Séguy 1930a, MA, Azrou; Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Thyridanthrax Osten-Sacken, 1886

Thyridanthrax alphonsi Sánchez Terrón and Roldan Bravo, 2000

Sánchez Terrón and Roldan Bravo 2000

Thyridanthrax elegans ssp. elegans (Wiedemann in Meigen, 1820)

Séguy 1930a, AP, Rabat; Mouna 1998; Dils and Özbek 2006; El Hawagry 2011; Evenhuis and Greathead 2015; AP (Oued Cherrat, Rabat), MA (Volubilis) – MISR

Thyridanthrax fenestratus (Fallén, 1814)

Séguy 1926; Séguy 1930a, EM, Berkane (1350–1400 m); Mouna 1998; El Hawagry 2011; Evenhuis and Greathead 2015; Rif (Tomorot) – MISR

Thyridanthrax griseolus Klug, 1832

Zaitzev 2008, SA, Taganint (south of Bou-Izakarn)

Thyridanthrax hispanus (Loew, 1869)

Becker and Stein 1913, Rif, Tanger; Sánchez Terrón and Roldan Bravo 2000

Thyridanthrax incanus (Klug, 1832)

= Anthrax incana Klug, 1832, in Séguy 1953a: 83

Séguy 1930a, AP, Oued Korifla (Zaers); Timon-David 1951, AP, Zaer; Séguy 1953a, MA, Tarda; Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; El Hawagry 2011; Karimpour 2012; Evenhuis and Greathead 2015

Thyridanthrax loustaui Andréu Rubio, 1961

Sánchez Terrón and Roldan Bravo 2000

Thyridanthrax maroccanus Dils, 2012

Dils 2012, AA, Ouarzazate, Skoura (1250 m), Amerzgane (1350 m)

Thyridanthrax mutilus (Loew, 1869)*

AA

Thyridanthrax nebulosus (Dufour, 1852)

Becker and Stein 1913, Rif, Tanger; Andréu Rubio 1959; Mouna 1998; Sánchez Terrón and Roldan Bravo 2000, Rif, Benibuifrur, Melilla, Restinga; Evenhuis and Greathead 2015

Thyridanthrax perspicillaris ssp. perspicillaris (Loew, 1869)

Séguy 1930a, MA, Aïn Leuh, Forêt Azrou, HA, Tizi-n’Test, Jebel Imdress (2000–2450 m), Goundafa; Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; El Hawagry 2011; Evenhuis and Greathead 2015

Thyridanthrax polyphemus (Hoffmansegg, 1819)

Séguy 1930a, MA, Volubilis (400 m); Mouna 1998; Bader and Arabyat 2004; Dils and Özbek 2006; Karimpour 2012; Evenhuis and Greathead 2015

Veribubo Evenhuis, 1978

Veribubo angusteoculatus (Becker, 1902)

Zaitzev 2008, AA, Zagora

Veribubo saudensis (François, 1970)*

AA

Veribubo tabaninus (François, 1970)*

AA, SA

Villa Lioy, 1864

Villa brunnea Becker, 1916

Mouna 1998: 84

Villa ceballosi Andréu Rubio, 1959

Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Villa cingulata Meigen, 1804

Mouna 1998; AP (Rabat, Casablanca), MA (Volubilis, Fès) – MISR

Villa distincta (Meigen in Waltl, 1835)

Dils and Özbek 2006; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Villa fasciata (Meigen, 1804)

= Villa circumdata (Meigen), in Séguy 1941a: 29

Séguy 1930a, AP, Rabat; Séguy 1941a, AP, Rabat, HA, Tachdirt (Toubkal, 2500 m); Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Villa hottentotta (Linnaeus, 1758)

= Anthrax hottentotus Linnaeus, in Séguy 1926: 198, Séguy 1930a: 92, Bléton and Fleuzet 1939: 64

Séguy 1930a, AP, Rabat, MA, Aïn Leuh; Bléton and Fleuzet 1939, MA, Fès; Séguy 1941d, HA, Tizi-n’Test; Mouna 1998; Dils and Özbek 2006; Evenhuis and Greathead 2015 – MISR

Villa ixion (Fabricius, 1794)

Dils and Özbek 2006; Koçak and Kemal 2010; Karimpour 2012; Evenhuis and Greathead 2015

Villa laevis Becker, 1915

Bader and Arabyat 2004; Dils and Özbek 2006; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Villa leucostoma (Meigen, 1820)

Mouna 1998: 84; AP (Bou-Regreg) – MISR

Villa luculenta Séguy, 1941

Séguy 1941d, AA, Taroudant; Mouna 1998; Koçak and Kemal 2010; Evenhuis and Greathead 2015

Villa niphobleta (Loew, 1869)

Bader and Arabyat 2004; Koçak and Kemal 2010; Karimpour 2012; Evenhuis and Greathead 2015

Villa venusta (Meigen, 1820)

Mouna 1998: 84

Desmatoneura Williston, 1895

Desmatoneura albifacies (Macquart, 1840)

Ebejer et al. 2019, AA, Merzouga (714 m)

Desmatoneura flavifrons Becker, 1915

Zaitzev 2008, AA, Ouarzazate, Taroudant, Jebel Tighermine (SE of Ouarzazate)

Petrorossia Bezzi, 1908

Petrorossia albula Zaitzev, 1962

Zaitzev 1999; Bader and Arabyat 2004; Zaitzev 2008, AA, Jebel Tighermine (SE of Ouarzazate); El Hawagry 2011; Evenhuis and Greathead 2015

Petrorossia freidbergi Zaitzev, 1999

Zaitzev 2008, AA, Jebel Tighermine (SE of Ouarzazate)

Petrorossia hespera (Rossi, 1790)

Séguy 1949a, AA, Tata; Mouna 1998; Zaitzev 1999; Dils and Özbek 2006; El Hawagry 2011; Evenhuis and Greathead 2015; AP (Bou-Regreg), MA (Timahdit) – MISR

Petrorossia margaritae Zaitzev, 1999

Zaitzev 2008, AA, Ouarzazate, Jebel Tighermine (SE of Ouarzazate)

New records for Morocco

Amictus setosus Loew, 1869

Atlantic Plain: Rommani, Marmouch, 33.568°N, 06.533°W, 400 m, 1♂1♀, Dils J.- Faes J., coll: PCJD.

Aphoebantus wadensis Becker, 1925

Sahara: Tata, 9 km. W Tissint, 29.851°N, 07.265°W, 535 m, 1♂1♀, 03.iii.2007, Dils J.- Faes J., coll: PCJD.

Bombylius (Unplaced) megacephalus Portschinsky, 1887

Eastern Morocco: Figuig, Abbou Lakhal, 32.1587°N, 01.507°W, 1050 m, 1♀, 07.iii.2009, Dils J.- Faes J., coll: PCJD.

Anti Atlas: Tiznit, 84 km. SSE Guelmim, 28.631°N, 10.75522°W, 235 m, 1♂, 27.ii.2007, Dils J.- Faes J., coll: PCJD; Tiznit, Abaynou, 29.057°N, 10.026°W, 360 m, 1♀, 13.iii.2009, Dils J.- Faes J., coll: PCJD.

Cononedys efflatouni Bezzi, 1925

Sahara: Guelmim, Souk Tnine Nouaday, 29.166°N, 09.279°W, 680 m, 2♂3♀, 07.iv.2015, Dils J.- Faes J., coll: PCJD.

Conophorus bellus Becker, 1906

High Atlas: Marrakech, Oukaimeden, 31.233°N, 07.817°W, 2200 m, 3♂, 06.iv.2006, Dils J.- Faes J., coll: PCJD.

Crocidium aegyptiacum Bezzi, 1925

Anti Atlas: Tiznit, Mesti, 29.274°N, 10.139°W, 280 m, 1♂, 23.iii.2006, Dils J.- Faes J., coll: PCJD.

Sahara: Tata, 28 km E of Tachjicht, 29.106°N, 09.149°W, 700 m, 1♀, 02.iii.2007, Dils J.- Faes J., coll: PCJD.

Crocidium nudum Efflatoun, 1945

Eastern Morocco: Oujda, Plateau du Rekkam, 33.839°N, 02.55781°W, 1150 m, 1♀, 25.iv.2010, Dils J.- Faes J., coll: PCJD.

Anti Atlas: Agadir, Imsouane, 30.885°N, 09.780°W, 270 m, 3♂13♀, 09.iv.2009, Dils J.- Faes J., coll: PCJD; Ouarzazate, Amerzgane, 31.024°N, 07.223°W, 1370 m, 17♂12♀, 31.iii.2006, Dils J.- Faes J., coll: PCJD; Taliouine, 18 km. W of Taliouine, 30.6003°N, 08.082°W, 900 m, 2♀, 24.iii.2009, Dils J.- Faes J., coll: PCJD; Taroudant, Tafinegoult, 30.734°N, 08.430°W, 680 m, 3♀, 24.iii.2009, Dils J.- Faes J., coll: PCJD; Tiznit, Arbaa Sahel, 29.657°N, 09.869°W, 320 m, 11♂26♀, 21.iii.2006, Dils J.- Faes J., coll: PCJD.

Cytherea albolineata Bezzi, 1925

Sahara: Guelmim, Tainzirt, 29.121°N, 09.333°W, 670 m, 1♀, 31.iii.2010, Dils J.- Faes J., coll: PCJD.

Geron intonsus Bezzi, 1925

Middle Atlas: Khenifra, Boulôjoul, 32.873°N, 04.945°W, 1500 m, 7♂10♀, 26.iv.2008, Dils J.- Faes J., coll: PCJD.

High Atlas: Midelt, 32.680°N, 04.677°W, 1400 m, 2♂2♀, 20.iv.2015, Dils J.- Faes J., coll: PCJD; Midelt, Zeïda, 32.781°N, 04.964°W, 1500 m, 9♂11♀, 24.iv.2015, Dils J.- Faes J., coll: PCJD.

Legnotomyia fascipennis Bezzi, 1925

Anti Atlas: Zagora, Tazarinne, 30.798°N, 05.584°W, 900 m, 1♂, 07.iii.2007, Dils J.- Faes J., coll: PCJD.

Sahara: Tata, 9 km W of Tissint, 29.851°N, 07.265°W, 535 m, 2♂1♀, 03.iii.2007, Dils J.- Faes J., coll: PCJD.

Thevenetimyia quedenfeldti (Engel, 1885)

Rif: Tanger-Tétouan, Souk El Kolla (Quolla), 35.083°N, 05.538°W, 150 m, 5♂4♀, 30.iv.2017, Dils J.- Faes J., coll: PCJD.

Atlantic Plain: Rommani, Merchouch, 33.568°N, 06.753°W, 400 m, 5♂22♀, 04.v.2010, Dils J.- Faes J., coll: PCJD.

Middle Atlas: Tadla-Azilal, Afourer, 32.180°N, 06.520°W, 1150 m, 5♂10♀, 07.v.2008, Dils J.- Faes J., coll: PCJD; Béni Mellal, El Ksiba, 32.576°N, 06.050°W, 870 m, 7♂23♀, 23.iv.2008, Dils J.- Faes J., coll: PCJD.

Thyridanthrax mutilus Loew, 1869

Anti Atlas: Tiznit, Sidi Ifni, 29.384°N, 10.172°W, 0 m, 7♂1♀, 10.iv.2008, Dils J.- Faes J., coll: PCJD.

Veribubo saudensis François, 1970

Anti Atlas: Erfoud, Tikkert-N-Ouchane, 31.223°N, 04.784°W, 830 m, 1♂3♀, 03.iv.2009, Dils J.- Faes J., coll: PCJD.

Veribubo tabaninus François, 1970

Anti Atlas: Ouarzazate, Amerzgane, 31.024°N, 07.223°W, 1370 m, 2♂9♀, 31.iii.2006, Dils J.- Faes J., coll: PCJD; Erfoud, Tikkert-N-Ouchane, 31.250°N, 04.617°W, 860 m, 1♀, 07.iii.2007, Dils J.- Faes J., coll: PCJD; Ouarzazate, Amerzgane, 31.017°N, 07.229°W, 1350 m, 6♂27♀, 25.iii.2009, Dils J.- Faes J., coll: PCJD; Ouarzazate, Amerzgane, 31.017°N, 07.229°W, 1350 m, 12♂8♀, 25.iii.2009, Dils J.- Faes J., coll: PCJD; Ouarzazate, 30.847°N, 06.817°W, 1200 m, 1♀, 30.iii.2009, Dils J.- Faes J., coll: PCJD.

Sahara: Guelmim, Tainzirt, 29.121°N, 09.333°W, 670 m, 22♀, 31.iii.2010, Dils J.- Faes J., coll: PCJD.

MYDIDAE

K. Kettani, T. Dikow

Number of species: 9. Expected: 10

Faunistic knowledge of the family in Morocco: moderate

Leptomydinae

Leptomydas Gerstaecker, 1868

Leptomydas lusitanicus (Wiedemann, 1820)

Mouna 1998

Rhopaliinae

Rhopalia Macquart, 1838

Rhopalia berlandi Séguy, 1949a: 153

Séguy 1949a, AA, Tagounit, Asni; Mouna 1998; Dikow 2017

Perissocerus Gerstaecker, 1868

Perissocerus rungsi Séguy, 1953

Séguy 1953a, SA

Syllegomydinae

Syllegomydini

Syllegomydas Becker, 1906

Syllegomydas algiricus (Gerstaecker, 1868)

= Rhopalia algirica Gerstaecker, in Séguy 1928c: 149

Gerstaecker 1868, AP, Casablanca; Séguy 1928c, AP, Rabat; Séguy 1930a, AP, Rabat; Mouna 1998; El Hawagry 2011; Dikow 2017

Syllegomydas berlandi (Séguy, 1941)

Séguy 1941, AA, Agadir; Dikow 2017

Syllegomydas bueni Arias, 1914

Arias 1914, AA, Tafilalt; Séguy 1928c; Séguy 1930a; Carles-Tolrá 2015; Carles-Tolrá 2017, EM, Mariouri, Trifa; Dikow 2017

Syllegomydas cinctus Macquart, 1835

Macquart 1835, MA, Immouzer road, AA, Agadir, Taroudant; Séguy 1930a; Mouna 1998; Carles-Tolrá 2017, EM, Quebdana, douar Shila, AA, Agadir coast; Dikow 2017

Syllegomydas maroccanus Séguy, 1928

Séguy 1928c, AP, Kénitra, Rabat, Oued Korifla, Forêt Zaers; Séguy 1930a, AP, Rabat, Temara, Oued Korifla, Forêt Zaers; Séguy 1932c; Mouna 1998; Carles-Tolrá 2017, AP, Larache, Ras Remel; Dikow 2017; AP (Kénitra) – MISR

Syllegomydas merceti Arias, 1914

Arias 1914, AP, Mogador; Séguy 1930a; Mouna 1998; El Hawagry 2011, AP, Mogador; Dikow 2017

MYTHICOMYIIDAE

K. Kettani, N. Evenhuis

Number of species: 8. Expected: 15

Faunistic knowledge of the family in Morocco: poor

Empidideicinae

Empidideicus Becker, 1907

Empidideicus crocea Séguy, 1949

= Cyrtosia crocea Séguy, in Séguy 1949a: 85

Séguy 1949a, SA, Guelmim; Séguy 1949c; Mouna 1998; Evenhuis 2002

Glabellulinae

Glabellula Bezzi, 1902

Glabellula maroccana Evenhuis & Kettani, 2018

Evenhuis and Kettani 2018, Rif, Adrou (PNPB) – BPBM, MISR

Leylaiya Efflatoun, 1945

Leylaiya pellea Evenhuis & Kettani, 2018

Evenhuis and Kettani 2018, AA, Tiznit – BPBM

Mythicomyiinae

Mythenteles Hall & Evenhuis, 1991

Mythenteles signifera Evenhuis & Kettani, 2018

Evenhuis and Kettani 2018, Rif, Talassemtane (maison forestière, 1699 m) – BPBM, MISR

Platypyginae

Cyrtisiopsis Séguy, 1930

Cyrtisiopsis melleus (Loew, 1856)

Evenhuis 2002; Zaitzav 2008, AA, Jebel Tighermine (SE Ouarzazate); Koçak and Kemal 2010; El Hawagry 2011

Cyrtisiopsis singularis Séguy, 1930

Evenhuis 2002

Cyrtosia Perris, 1839

Cyrtosia aglota Séguy, 1930

Evenhuis 2002

Cyrtosia marginata Perris, 1839

Séguy 1930a, HA; Mouna 1998; Evenhuis 2002; Evenhuis and David 2004

SCENOPINIDAE

K. Kettani, M. Carles-Tolrá

Number of species: 8 (+3 unidentified). Expected: 12

Faunistic knowledge of the family in Morocco: good

Scenopininae

Scenopinus Latreille, 1802

Scenopinus albicinctus (Rossi, 1794)

= Omphrale albicincta Rossi, in Séguy 1930a: 110

Séguy 1930a; Mouna 1998

Scenopinus fenestralis (Linnaeus, 1758)

= Omphrale fenestralis Linnaeus, in Séguy 1930a: 110

Séguy 1930a; Mouna 1998

Scenopinus glabrifrons Meigen, 1824

= Omphrale glabrifrons Meigen, in Séguy 1930a: 110

Séguy 1930a, MA, Aïn Leuh; Mouna 1998

Scenopinus niger (De Geer, 1776)

Becker and Stein 1913, Rif, Tanger; Mouna 1998

Scenopinus parallelus Kelsey, 1969

Kelsey 1969, AP, Villa Cisneros (Dakhla), SA, Río de Oro (Oued Eddahab)

Scenopinus physadius (Séguy, 1930)

= Omphrale physadia Séguy, in Séguy 1930a: 111

Séguy, 1930, EM, Bou Denib; Kelsey 1969, EM, Bou Denib; Mouna 1998

Scenopinus pilosus (Séguy, 1930)

= Omphrale pilosa Séguy, in Séguy 1930a: 111

Séguy 1930a, AP, Bou Knadel; Kelsey 1969, AP, Bou Knadel; Mouna 1998; Carles-Tolrá 2001

Scenopinus undescribed sp. 1

Ebejer et al. 2019, Rif, Martil (9 m)

Scenopinus undescribed sp. 2

Ebejer et al. 2019, Rif, Adrou (556 m)

Stenomphrale Kröber, 1937

Stenomphrale teutankhameni (Kröber, 1923)

Ebejer et al. 2019, AP, forest of Maâmora (56 m)

Stenomphrale sp.

AP (Essaouira (J.-P. Haenni leg.)) – MHNN Neuchâtel

THEREVIDAE

K. Kettani, M. Hauser

Number of species: 27.

Faunistic knowledge of the family in Morocco: moderate

Phycusinae

Phycusini

Actorthia Kröber, 1912

Actorthia micans (Kröber, 1924)

Kröber 1924, AA, Errachidia (45 km S Erfoud), Merzouga

Phycus Walker, 1850

Phycus lacteipennis Lyneborg, 2002

Lyneborg 2002, AA, 25 km S Goulmima (1000 m), SA, Mekn s-Tafilalet; Winterton et al. 2012; Badrawy and Mohammad 2013

Salentia Costa, 1857

Salentia anancitis (Séguy, 1941)

= Apioeicoceras anancitis Séguy, in Séguy 1941d: 10

Séguy 1941d, AA, Agadir; Mouna 1998

Salentia costalis (Wiedemann, 1824)

= Apioeicoceras costalis Wiedemann, in Séguy 1930a: 108

Wiedemann 1824, AP, Mogador, HA, Marrakech-Tensift-Al Haouz; Séguy 1930a; Mouna 1998; Koçak and Kemal 2010

Salentia fuscipennis Costa, 1857

Costa 1857, Rif, Tanger, AA, Tagadirt (Agadir); Becker and Stein 1913, Rif, Tanger; Séguy 1930a, HA, Bou Knadel, Tinmel; Mouna 1998

Therevinae

Therevini

Acanthothereva Séguy, 1935

Acanthothereva rungsi Séguy, 1935

Séguy 1935b, AP, Mehdia (20 km S Rabat); Lyneborg 1968; Koçak and Kemal 2010; Rif (Cap Spartel) – MISR

Acrosathe Irwin & Lyneborg, 1981

Acrosathe annulata (Fabricius, 1805)

Ebejer et al. 2019, Rif, Oued Kbir (Béni Ratene, 157 m)

Chrysanthemyia Becker, 1912

Chrysanthemyia chrysanthemi (Fabricius, 1787)

Fabricius 1787, EM, Béni Snassen Mountains, Tafouralt (800 m); Séguy 1930a, MA, Meknès, Berrechid; Mouna 1998; MA (Oued Grou, Timahdit) – MISR

Chrysanthemyia velutinifrons (Becker, 1912)

= Chrysanthemyia lucidifrons Becker 1912: 81

= Oedicera velutinifrons Becker, in Becker and Stein 1913: 82

Becker 1912, Rif, Region de Tanger-Tétouan, Cercle d’Ouezzane (300 m), AP, 3 km S Settat, EM, Figuig, MA, Fès-Boulmane, Sidi Harazem (223 m); Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA, Meknès; Mouna 1998; MA (Oued Grou) – MISR

Hoplosathe Lyneborg & Zaitsev, 1980

Hoplosathe distincta Lyneborg & Zaitsev, 1980

Lyneborg and Zaitsev 1980, HA, Oued Tensift (Marrakech)

Neotherevella Lyneborg, 1978

Neotherevella macularis (Wiedemann, 1828)

Hauser et al. 2017, AA, Tifnite (10 Km S Agadir), Merzouga (45 Km S Erfoud)

Thereva Latreille, 1797

Thereva atra Kröber, 1913

El Hawagry 2011

Thereva aureoscutellata Kröber, 1914

Ebejer et al. 2019, Rif, Moulay Abdelsalam (965 m)

Thereva binotata Loew, 1847

Koçak and Kemal 2010

Thereva bipunctata Meigen, 182016

Ebejer et al. 2019, MA, Khénifra (17 km SW of Midelt, 1940 m; 17 km NW of Zaida, 1878 m; 28 km S of Timahdit, 2100 m), Lac Aguelmane Afennourir (30 km SW of Azrou, 2050 m)

Thereva brevicornis Loew, 184717

Pârvu et al. 2006, AP, Cap Sim; Popescu-Mirceni 2011

Thereva chrysargyrea Séguy, 1953

Séguy 1953a, SA, Amguilli Sguelma

Thereva cincta Meigen, 1820

Ebejer et al. 2019, Rif, Oued Aliane (Ksar Sghir, 1 m)

Thereva funebris Meigen, 182018

= Thereva lugubris Meigen

Mouna 1998; AP (Rabat), MA (Ifrane) – MISR

Thereva graeca Kröber, 19123

Séguy 1930a, Rif, Tanger; Mouna 1998

Thereva plebeja (Linnaeus, 1758)3

Séguy 1953a, AA, Tifnit (Souss); Mouna 1998; MA (Ras el Ma) – MISR

Thereva powelli Séguy, 1930

Séguy 1930a, MA, Forêt Azrou; MA (Ras el Ma) – MISR

Thereva spiloptera Wiedemann, 1824

Wiedemann 1824, HA, Ouirgane (Marrakech, 1000 m); Séguy 1930a, Rif, Tanger, AP, Mogador, MA, Meknès; Séguy 1953a, AP, Temara; Mouna 1998; El Hawagry 2011

Thereva spinulosa Loew, 1847

Loew 1847, AP, Maâmora, MA, Khemisset

Thereva stigmatica Kröber, 1912

EL-Hawagy 2011, Rif, Tanger

Thereva strigata (Fabricius, 1794)19

Koçak and Kemal 2010

Thereva tuberculata Loew, 1847

= Thereva algirica Kröber, 1913, in Séguy 1953a: 83

Loew 1847, AP, Salé; Séguy 1930a, MA, Meknès; Séguy 1953a, AP, Salé; Mouna 1998; Koçak and Kemal 2010

Acknowledgments

We gratefully acknowledge Martin Ebejer (UK) for material, comments and cooperation, as well as Gail Kampmeier (USA) for sharing data of Moroccan Therevidae out of the mandala database (http://wwx.inhs.illinois.edu/research/mandala/about/).

Empidoidea

ATELESTIDAE

K. Kettani, P. Gatt

Number of species: 1. Expected: 1

Faunistic knowledge of the family in Morocco: poor

Atelestus Walker, 1837

Atelestus sp. nov.

Ebejer et al. 2019, Rif, Dardara (484–730 m), Aïn Tissemlal (Azilane, 1255 m), Douar El Hamma (338 m), Chrabkha pond (Al Manzla, 58 m)

EMPIDIDAE

K. Kettani, C. Daugeron

Number of species: 40. Expected: 100

Faunistic knowledge of the family in Morocco: poor

Clinocerinae

Clinocera Meigen, 1803

Clinocera maroccana (Séguy, 1941): 29 (= Hydrodromia)

= Atalanta (Hydrodromia) algira (Vaillant, 1952): 65

Séguy 1941a, HA, Anrhemer (Toubkal, 2500 m); Vaillant 1956b, HA, Sidi Chamarouch; Vaillant 1964; Dakki 1997

Clinocera megalatlantica (Vaillant, 1957): 65 (= Atalanta (Atalanta))

Vaillant 1956b, HA; Dakki 1997

Clinocera nigra Meigen, 1804: 292

= Heleodromia unicolor (Curtis, 1834): plate 513, Paramesia roberti (Macquart, 1835): 657

Vaillant 1956b, HA, Izourar, Sidi Chamarouch, Aguelmous; Dakki 1997

Dolichocephala Meigen, 1803

Dolichocephala ocellata (Costa, 1858): 7 (= Ardoptera)

= oculata (Loew, 1858~7): 7 (= Ardoptera)

= novemguttata (Strobl, 1893): 98 (= Ardoptera)

= albohalterata (Strobl, 1898): 399 (= Ardoptera)

= barbarica Vaillant, 1952: 65

= algira Vaillant, 1957: 64

Vaillant 1956b, HA, Imi-N’Ifri; Dakki 1997

Dolichocephala pavonica Vaillant & Gagneur, 1998: 380

Vaillant and Gagneur 1998, HA, Demnat

Kowarzia Mik, 1881

Kowarzia barbatula (Mik, 1880): 347 (= Clinocera)

= Clinocera dorieri (Vaillant, 1968): 88

Vaillant 1956b, HA, Cascade Siroua, Lac Tamhda (Anremer), Aguelmous, Sidi Chamarouch, Izourar, Oukaimeden; Dakki 1997; Vaillant and Moubayed 1998

Kowarzia bipunctata (Haliday, 1833) (= Heleodromia)

Vaillant 1956b, HA, Oukaimeden; Vaillant 1964; Dakki 1997

Kowarzia dieuzedei Vaillant, 1953: 60

Vaillant 1956b, HA, Lac Tamhda (Anremer); Vaillant 1964, HA; Dakki 1997

Kowarzia madicola (Vaillant, 1965): 152 (= Atalanta)

Vaillant 1956b, HA, Tahanaout

Kowarzia tenella (Wahlberg, 1844): 107 (= Parmesia)

= Heleodromia zetterstedti (Walker, 1851): 105

= Wiedemannia securigera (Engel, 1918): 70

Vaillant 1956b, HA, Cascade Siroua, Lac Tamhda (Anremer), Aguelmous, Sidi Chamarouch, Izourar, Oukaimeden; Dakki 1997; Vaillant and Moubayed 1998

Wiedemannia Zetterstedt, 1838

Wiedemannia (Chamaedipsia) beckeri (Mik, 1889): 71 (= Chamaedipsia)

= jugorum (Strobl, 1893): 105 (= Chamaedipsia)

= crinita Engel, 1918: 217 (as var. of W. beckeri)

= alticola Vaillant, 195l: 54

= alpina Vaillant, 1967: 274 (as ssp. of W. beckeri)

= glaciola Wagner, 1985: 86 (as ssp. of W. beckeri; new name for W. beckeri alpina Vaillant)

Dakki 1997

Wiedemannia (Chamaedipsia) mgounica Vaillant, 1957: 69

Vaillant 1956a, HA, M’Goum; Dakki 1997

Wiedemannia (Philolutra) azurea (Vaillant, 1951): 3 (= Philolutra)

El Mezdi and Giudicelli 1985, HA, Khettaras Marrakech; Dakki 1997; Vaillant and Moubayed 1998

Wiedemannia (Philolutra) fallaciosa Loew, 1873: 44

Vaillant 1964, HA; Dakki 1997

Wiedemannia (Philolutra) fallaciosa ssp. litardierei Vaillant, 1957: 67

Vaillant 1956a, HA; Dakki 1997

Wiedemannia (Roederella) ouedorum Vaillant, 1952: 371

= ovedorum (error) Vaillant, 1978: 469

Vaillant 1964, HA; Dakki 1997

Hemerodromiinae

Hemerodromia Meigen, 1822

Hemerodromia bethiana Vaillant & Gagneur, 1998

Vaillant and Gagneur 1998, MA, Tigrigra (Azrou)

Hemerodromia subapicalis Yang, Zhang & Zhang, 2007

= Hemerodromia apicalis Vaillant and Gagneur 1998: 372 (preoccupied by Smith, 1969)

Vaillant and Gagneur 1998, HA, Oum-er-Rbia; Yang et al. 2007

Hemerodromia tigrigrana Vaillant & Gagneur, 1998

Vaillant and Gagneur 1998, MA, Tigrigra (Azrou)

Hemerodromia todrhana (Vaillant, 1956)

Vaillant 1956, HA, Todrha; El Mezdi and Guidicelli 1985; Dakki 1997; Vaillant and Gagneur 1998

Hemerodromia zarcana Vaillant & Moubayed, 1998

Vaillant and Gagneur 1998, MA, Tigrigra (Azrou)

Empidinae

Empidini

Empis Linnaeus, 1758

Empis (Coptophlebia) confluens Becker, 1907

MA (Meknès), 19.v.1997, K. Deneš leg. – OUMNH

Empis (Empis) decora Meigen, 1822

Bahid 2018, Rif, Oued Tkarae (PNPB); Ebejer et al. 2019, Rif, Oued Nakhla, Moulay Abdelsalam

Empis (Empis) nikita Shamshev, 2018

Shamshev 2018, AP, Essaouira

Empis (Kritempis) taffertensis Daugeron, 2009

Daugeron 2009, MA, forest of Taffert; Bahid 2018

Empis (Leptempis) tenuis Bahid & Daugeron, 2017

Bahid et al. 2017, MA, Tizi-s’Tkrine (Jebel Amar, 1760 m); Bahid 2018

Empis (Pachymeria) suberis Becker, 1907

Ebejer et al. 2019, Rif, Moulay Abdelsalam, Issaguen, Bab Berred, Jebel Lakraâ

Empis (Polyblepharis) eumera Loew, 1866

Ebejer et al. 2019, MA, Ifrane

Empis (Xanthempis) chopardi Daugeron, 1997

Daugeron 1997, MA, Ifrane; Daugeron 2000; Bahid 2018

Empis (Xanthempis) edithae Daugeron, 1997

Daugeron 1997, HA, High Imminen, Tachdirt; Daugeron 2000; Bahid 2018

Empis (Xanthempis) ifranensis Daugeron, 1997

Daugeron 1997, MA, Ifrane; Daugeron 2000; Bahid 2018

Empis (Xanthempis) styriaca (Strobl, 1893)

Chvála and Wagner 1989, HA [doubtful record]; Bahid 2018

Empis (Xanthempis) widanensis Bahid & Daugeron, 2018

Bahid et al. 2018, Rif, Dayat Bayan Widane, Aïn Sedraouia, Tazia, Anissar, Lalla Outka

Rhamphomyia Meigen, 1822

Rhamphomyia (Rhamphomyia) maroccana Collin, 2009

Chvála and Wagner 1989, MA, Ifrane; Collin 2009, MA, Ifrane; Bahid 2018, Rif, Oued Akrir (Fifi)

Rhamphomyia (Holoclera) tenuipes Becker, 1907*

AP, Haenni pers. comm.

Hilarini

Hilara Meigen, 1822

Hilara algecirasensis Strobl, 1899

Ebejer et al. 2019, MA, Lac Aguelmane Afennourir, HA, Ziz river

Hilara almeriensis Strobl, 1906

Chvála 2008, AA, Tifoultoute (1146 m)

Hilara fusitibia Strobl, 1899

Chvála 2008, MA, Ifrane (Forêt de Cédres, 1500 m)

Hilara longeciliata Strobl, 1906

Chvála 2008, AP, Rabat (near Oued Bou-Regreg, 0–10 m)

Hilara schachti Chvála, 2008

Chvála 2008, MA, Ifrane (Ghabat al Behar, 1650–1700 m); Bahid 2018

New record for Morocco

Rhamphomyia (Holoclera) tenuipes Becker, 1907

Atlantic Plain: Essaouira, 6 km W, 3.iv.2002, Forêt de genévriers pâturée, 1♂1♀, J.-P. Haenni leg., coll. MHNN.

Acknowledgements

We are very grateful to Bradley Sinclair (Canadian Food Inspection Agency, Canada) and Adrian Plant (Mahasarakham University, Thailand) for reviewing parts of this family.

DOLICHOPODIDAE

K. Kettani, I.Ya. Grichanov, O.P. Negrobov

Number of species: 112. Expected: 300

Faunistic knowledge of the family in Morocco: poor

Diaphorinae

Argyra Macquart, 1834

Argyra argentina Meigen, 1824

Parent 1924, Rif, Cap Spartel, Tétouan; Parent 1927, Rif, Tétouan

Argyra argyria (Meigen, 1824)

Parent 1924 (females only), Rif, Cap Spartel, Tétouan; Kechev and Ivanova 2015

Argyra biseta Parent, 1929

Parent 1929a, Rif, Tanger; Vaillant 1955a

Argyra grata Loew, 185720

Negrobov 1991 (no material provided)

Asyndetus Loew, 1869

Asyndetus separatus (Becker, 1902)

Ebejer et al. 2019, AA, 14 km E of Rich (Errachidia, 1278 m)

Chrysotus Meigen, 1824

Chrysotus albibarbus Loew, 1857

Ebejer et al. 2019, MA, Lac Aguelmane Afennourir (30 km SW of Azrou, 1760 m); Grichanov 2019, AA, Aït Melloul

Chrysotus gramineus (Fallén, 1823)

Parent 1924, Rif, Tanger; Parent 1927

Chrysotus larachensis Grichanov, Nourti & Kettani, 2020

Grichanov et al. 2020b, Rif, El Hamma (338 m)

Chrysotus pennatus Lichtwardt, 1902

Ebejer et al. 2019, Rif, Smir Barrage (145 m), AA, 1 km N of Tarda (Errachidia, 1023 m)

Chrysotus suavis Loew, 1857

Grichanov 2009, HA, Asni area (1100–1400 m); Nourti et al. 2019a, Rif, Amsemlil env. (1067 m), MA, Dayat Ifrane (1607 m), HA, Tahanout (956 m); Dawah et al. 2020

Diaphorus Meigen, 1824

Diaphorus africus Parent, 1924

Parent 1924, Rif, Tétouan, Tanger; Parent 1927, Rif, Tanger; Ebejer et al. 2019, Rif, Oued Siflaou (281 m); Grichanov 2019, AA, Ouarzazate (1100 m)

Diaphorus vitripennis Loew, 1859

Grichanov 2019, AA, Aït Melloul

Dolichopodinae

Dolichopus Latreille, 1796

Dolichopus andalusiacus (Strobl, 1899)

Ebejer et al. 2019, AP, Loukous marsh (2 m); Nourti et al. 2019a, Rif, plage Stihat (0 m)

Dolichopus griseipennis Stannius, 1831

Parent 1924, Rif, Tanger; Parent 1927; Séguy 1930a, Rif, Tanger; Nourti et al. 2019a, Rif, Adrou (PNPB, 556 m)

Dolichopus sabinus Haliday, 1838

Ebejer et al. 2019, Rif, Martil (9 m), Oued Laou (2 m); Nourti et al. 2019a, Rif, plage Stihat (4 m)

Dolichopus scutopilosus Parent, 1933

Parent 1933, HA, Arround

Dolichopus signifer Haliday, 1832

Parent 1929b, “Maroc”; Séguy 1930a, MA, Ras el Ma; Grichanov 2019, HA, Oukaimeden (2600 m)

Dolichopus strigipes Verrall, 1875

Grichanov 2009, AP, 40 km S Larache (0–20 m)

Gymnopternus Loew, 1857

Gymnopternus assimilis (Staeger, 1842)

Nourti et al. 2019a, Rif, Amsemlil (1067 m)

Hercostomus Loew, 1857

Hercostomus apollo (Loew, 1869)

Nourti et al. 2019a, Rif, Talassemtane (1696 m), Adrou (PNPB, 556 m), Amsemlil (PNPB, 1067 m)

Hercostomus canariensis Santos Abreu, 1929

Grichanov et al. 2020a, Rif, Pont de Dieu (Akchour, 536 m); Nourti et al. 2019a (as H. aff. exarticulatoides Stackelberg, 1949)

Hercostomus chetifer (Haliday, 1849)

Ebejer et al. 2019, Rif, Sidi Yahia Aarab (377 m)

Hercostomus discriminatus Parent, 1925

Parent 1925, Rif, “Favier, Environs de Tanger”; Parent 1927, Rif, Tanger; Vaillant 1950, Rif, Tanger

Hercostomus exarticulatus (Loew, 1857)

Vaillant 1956b, HA, Lac Tamhda (Anremer), Aguelmous; Grichanov et al. 2020a

Hercostomus excipiens Becker, 1907

Parent 1924, Rif, Tétouan; Parent 1927; Séguy 1930a, Rif, Oued Judios (Tanger); Nourti et al. 2019a, Rif, Talembote (440 m)

Hercostomus germanus (Wiedemann, 1817)

Parent 1924, Rif, Cap Spartel; Parent 1927; Kettani and Negrobov 2016, Rif, Chefchaouen, Ketama – MISR (Rif, Ketama)

Hercostomus longiventris (Loew, 1857)

Vaillant 1956b, HA, Lac Tamhda (Anremer), Izourar

Muscidideicus Becker, 1917

Muscidideicus praetextatus (Haliday, 1855)

Grichanov 2019, AP, Oualidia lagune

Ortochile Berthold, 1827

Ortochile morenae (Strobl, 1899)

= Hercostomus morenae (Strobl, 1899), in Becker 1917: 225; Nourti et al. 2019a: 124

Grichanov and Nourti 2021; Nourti et al. 2019a, Rif, Mnezla (74 m), Talassemtane (980 m), Oued Ametrasse (841 m), estuary Tahaddart (dune marshland, 0 m)

Ortochile nigrocaerulea Latreille, 1779

Parent 1924, Rif, Tanger, Cap Spartel, Tétouan, Béni Hozmar; Parent 1927, Rif, Tanger; Séguy 1930a, Rif, Oued Judios (Tanger); Grichanov 2009, Rif, Ouezzane (300 m); Nourti et al. 2019a, Rif, Douar El Hamma (338 m), Triwa Bni Hassane (654 m), Taida (501 m)

Platyopsis Parent, 1929

Platyopsis maroccanus (Parent, 1929)

Parent 1929b, Rif, Tanger; Vaillant 1950, Rif, Tanger

Poecilobothrus Mik, 1878

Poecilobothrus appendiculatus (Loew, 1859)

= Hercostomus appendiculatus (Loew): Ebejer et al. 2019: 146

Parent 1924, Rif, Tanger, Cap Spartel; Ebejer et al. 2019, Rif, Oued Nakhla (200 m), Moulay Abdelsalam (965 m), Dardara (730 m), Cap Spartel (155 m); Nourti et al. 2019a, Rif, Perdicaris Park (223 m)

Poecilobothrus infuscatus (Stannius, 1831)

Ebejer et al. 2019, Rif, Tahaddart (2 m); Rif (Tahaddart) – MISR

Sybistroma Meigen, 1824

Sybistroma dufouri Macquart, 1838

= Haltericerus spathulatus Loew, in Becker and Stein 1913: 86

Becker and Stein 1913, Rif, Tanger

Sybistroma obscurellum Fallén, 182320

= Hypophyllus obscurellus Fallén, in Dakki 1997: 61

Dakki 1997 (no material provided)

Sybistroma quadrifilatum (Strobl, 1899)

= Sybistroma parvulum (Parent, 1927), in Grichanov and Nourti 2021: 190

Grichanov and Nourti 2021, Rif, Fahs Anjra (372 m)

Sybistroma theodori Grichanov & Nourti, 2021

Grichanov and Nourti 2021, Rif, Moulay Abdelsalam (649 m)

Tachytrechus Haliday, 1851

Tachytrechus consobrinus (Haliday, 1851)20

Parent 1938 (no material provided)

Tachytrechus goudoti (Macquart, 1842)

= Dolichopus goudoti Macquart, 1842

Macquart 1842, Rif, Tanger; Parent 1926 (redescription), 1927

Tachytrechus insignis (Stannius, 1831)

Parent 1927, “Maroc”; Séguy 1930a, Rif, Tanger, HA, Aguerd el Had, Talekjount (1000–1100 m); Vaillant 1956b, HA, Lac Tamhda (Anremer); Popescu-Mirceni 2011, AP, Merja Zerga; Grichanov 2019, AP, Essaouira

Tachytrechus notatus (Stannius, 1831)

Vaillant 1950 (no material provided), 1956b, HA, Lac Tamhda (Anremer); Grichanov 2009, AA, 15 km SW Tazenakcht; Dawah et al. 2020

Tachytrechus planitarsis Becker, 1907

Vaillant 1950, HA, Touggourt; Grichanov 2009, AA, 15 km SW Tazenakcht; Grichanov 2019, AA, Ouarzazate (1100 m)

Hydrophorinae

Anahydrophorus Becker, 1917

Anahydrophorus cinereus (Fabricius, 1805)

= Scatophaga cinerea Fabricius, 1805: 205

Fabricius 1805, AP, Mogador (Essaouira); Séguy 1930a, Rif, Tanger; Vaillant 1955a, AP, Temara, Port-Lyautey; Boumezzough and Vaillant 1986a, AP, beach of Rabat; Kettani and Negrobov 2016, AP, Skhirat; AP (Skhirat) – MISR

Aphrosylus Haliday, 1851

Aphrosylus maroccanus Vaillant, 1955

Vaillant 1955a, AP, Port Lyautey

Aphrosylus mitis Verrall, 1912

Grichanov 2019, AP, Oualidia lagune

Aphrosylus raptorluteipes Parent, 1929

Parent 1929b, AP, Mogador (as a variation of Aphrosylus raptor Haliday, 1851); Vaillant 1955a; Negrobov, 1979 (as a subspecies of Aphrosylus raptor Haliday, 1851); Kettani and Negrobov 2016 (as Aphrosylus raptor Haliday, 1851); Grichanov 2019, AP, Oualidia lagune

Aphrosylus temaranus Vaillant, 1955

Vaillant 1955a, AP, Temara; Grichanov 2019, AP, Oualidia lagune

Aphrosylus venator Loew, 1857

Parent 1927, Rif, Tanger; Séguy 1930a, Rif, Tanger

Epithalassius Mik, 1891

Epithalassius corsicanus Becker, 1910

Pârvu 2008, AP, Merja Zergha, Cap Sim, Essaouira

Hydrophoprus Fallén, 1823

Hydrophorus balticus (Meigen, 1824)

Vaillant 1956b, HA, Jebel Toubkal, Lac Tamhda (Anremer), Oukaimeden, Izourar; Boumezzough and Vaillant 1986a, HA, Jebel Toubkal (3100 m); Grichanov 2019, HA, Aguelmouss (2050 m), Oukaimeden (2600 m); Nourti et al. 2019a, MA, Mont Habri (2071 m)

Hydrophorus nilicola Parent, 1927

= Hydrophorus viridis nilicola Parent, in Boumezzough and Vaillant 1986a: 297

Boumezzough and Vaillant 1986a, MA, Tizi-n’Imdrhas (1800 m), HA, Oued N’fis (650 m), AA, near Agadir N’oussbai (400 m); Grichanov 2019, AP, Essaouira

Hydrophorus oceanus (Macquart, 1838)

= Hydrophorus bisetus Loew, 1857, in Parent 1927, Séguy 1930a, Grichanov 2019

Parent 1927, AP, Rabat; Séguy 1930a, Rif, Tandja el Balia (Tanger) (Hydrophorus bisetus Loew); Vaillant 1955a, AP, Port-Lyautey; Boumezzough and Vaillant 1986a, AP, beach of Rabat

Hydrophorus praecox (Lehmann, 1822)

Parent 1924, Rif, Cap Spartel, de Tanger à Tétouan, Rincón de Medik, Dar Riffien (Ceuta); Parent 1927; Séguy 1941a, HA, Toubkal (2500 m); Boumezzough and Vaillant 1986a, HA, Lac Tamhda, Lac Tamdhanit (Massif Anremer, 2900 m), Lac Izourar (Massif Azourki, 2650 m)

Hydrophorus viridis (Meigen, 1824)21

Parent 1927, AP, Rabat; Boumezzough and Vaillant 1986a: 297

Liancalus Loew, 1857

Liancalus virens (Scopoli, 1763)

Vaillant 1956b, HA, Toubkal, Assif Tassouat (M’Goum), Aguelmous, Sidi Chamarouch, Imi-N’Ifri; Boumezzough and Vaillant 1986a, HA, Tahanaout (750 m), Adrar Anremer (2900 m), AA, Jebel Siroua (3000 m); Kettani and Negrobov 2016, AP, S-Tifni; Nourti et al. 2019a, Rif, Amsemlil env. (1059 m) – MISR (AP, S Tifni)

Machaerium Haliday, 1832

Machaerium maritimae Haliday, 1832

Parent 1927, Rif, Tanger; Séguy 1930a, Rif, Tanger; Kettani and Negrobov 2016, AP, Oued Bou-Regreg; Grichanov 2019, AP, Oualidia lagune; AP (Oued Bou-Regreg) – MISR

Orthoceratium Schrank, 1803

Orthoceratium sabulosum (Becker, 1907)

= Orthoceratium lacustre (Scopoli, 1763)22, in Ebejer et al. 2019: 146

Ebejer et al. 2019, Rif, Jebel Lakraâ (Talassemtane, 1541 m)

Thinophilus Wahlberg, 1844

Thinophilus (Thinophilus) flavipalpis (Zetterstedt, 1843)

Pârvu et al. 2006, AA, Lac Edehby, Ouarzazate; Grichanov 2009, AP, 40 km S Larache (0–20 m)

Thinophilus (Thinophilus) indigenus Becker, 1902

Ebejer et al. 2019, AA, 14 km E of Rich (Errachidia, 1278 m), Oued Laou (2 m); Grichanov 2019, AA, Ouarzazate (572 m)

Thinophilus (Thinophilus) mirandus Becker, 1907

Negrobov 1971, Rif, Tanger; Grichanov 2019, AA, Ouarzazate (572 m)

Thinophilus (Schoenophilus) versutus Haliday, 1851

= Schoenophilus versutus (Haliday, 1851), in Parent 1924, 1927

Parent 1924, Rif, Cap Spartel, Tétouan; Parent 1927; Pârvu et al. 2006, AP, Merja Zerga; Nourti et al. 2019a, Rif, Oued Souk Lhad (613 m), Dayat Tazia (733 m)

Medeterinae

Medetera Fisher, 1819

Medetera diadema (Linnaeus, 1767)

= Medeterus (cf. diadema), in Séguy 1953a: 84

Parent 1938 (no material provided); Séguy 1953a, AP, Rabat; Nourti et al. 2019a, Rif, Kitane (49 m), Rif, plage Stihat (beach)

Medetera media Parent, 1925

Nourti et al. 2019a, Rif, Faculty of Sciences of Tétouan (garden: on trunk of olive tree, 14 m)

Medetera micacea Loew, 1857

Ebejer et al. 2019, Rif, Dardara (730 m); Nourti et al. 2019a, Rif, Issaguen (1547 m),

HA, Lac Tislit (Imilchil, 2254 m)

Medetera pallipes (Zetterstedt, 1843)23

Nourti et al. 2019a, Rif, Douar El Hamma (338 m)

Medetera petrophila Kowarz, 187720

Parent 1938 (no material provided)

Medetera petrophiloides Parent, 1925

Nourti et al. 2019a, Rif, Issaguen (1547 m)

Medetera aff. roghii Rampini & Canzoneri, 1979

Nourti et al. 2019a, Rif, Douar El Hamma (338 m)

Medetera truncorum Meigen, 182424

Ebejer et al. 2019a, Rif, Dardara (484 m)

Medetera varvara Grichanov & Vikhrev, 2009

Grichanov and Vikhrev 2009, AP, Essaouira

Thrypticus Gerstäcker, 1864

Thrypticus bellus Loew, 1869

Séguy 1930a, EM, Dayat Sidi Kacem; Nourti et al. 2019a, Rif, Amsemlil env. (1067 m), estuary Oued Tahaddart (0 m), Barrage 9 Avril, plage Stihat (0 m)

Microphorinae

Schistostoma Becker, 1902

Schistostoma eremita Becker, 1902

Ebejer et al. 2019, AA, Ziz river (12 km S of Rissani, 737 m), Lac Tiffert (4 km W of Merzouga, 702 m)

Neurigoninae

Neurigona Rondani, 1856

Neurigona solodovnikovi Grichanov, 2010

= Neurigona punctifera Becker, 1907, in Kettani and Negrobov 2016 (misidentification)

Grichanov 2010, AP, 40 km S Larache; Kettani and Negrobov 2016, Rif, Aïn Tissemlal (Azilane, 1255 m); Nourti et al. 2019a, Rif, Perdicaris Park (223 m), Taria Bni Faghloum (894 m), Chrafate (832 m)

Parathalassiinae

Microphorella Becker, 1909

Microphorella ulrichi Gatt, 2003

Gatt 2003, Rif, Tanger, Oued Armal (Ksar Sghir)

Parathalassius Mik, 1891

Parathalassius blasigii Mik, 1891

Ebejer et al. 2019, AP, Larache (5 m)

Peloropeodinae

Chrysotimus Loew, 1857

Chrysotimus molliculoides Parent, 1937

Parent 1937a, MA, Ifrane (1600 m); Parent 1937b; Nourti et al. 2019a, Rif, Dayat Tazia (733 m)

Micromorphus Mik, 1878

Micromorphus albipes (Zetterstedt, 1843)

Parent 1924, Rif, Béni Hozmar (Tétouan); Parent 1927, Rif, Tétouan; Kechev and Ivanova 2015; Nourti et al. 2019a, Rif, Oued Souk Lhad (613 m), estuary Oued Tahaddart (0 m), Barrage 9 Avril, Kharouba, plage Stihat (0 m), Aïn Jdioui, EM, Bouanane (Figuig, 855 m), AA, Boudnib (951 m); Dawah et al. 2020

Micromorphus minusculus Negrobov, 2000

Nourti et al. 2019a, EM, Bouanane (Figuig, 855 m), AA, Taliouine (Taroudant, 1014 m); Grichanov 2019, AA, Ouarzazate

Rhaphiinae

Rhaphium Meigen, 1803

Rhaphium appendiculatum Zetterstedt, 184920

= Rhaphium macrocerum (Parent, 1925), in Parent 1938, Grichanov 2019

Parent 1938 (no material provided)

Rhaphium brevicorne Curtis, 1835

= Xiphandrium pectinatum Becker, in Vaillant 1956b: 112, Grichanov 2019

Vaillant 1956b, AP, Rabat, HA, Oukaimeden; Kettani and Negrobov 2016, AP, Rabat; Grichanov 2019, HA, Oukaimeden (1000 m); Nourti et al. 2019a, Rif, Pont de Dieu (Akchour, 536 m), Dayat Tazia (733 m), Moulay Abdelsalam (1177 m); AP (Rabat) – MISR

Rhaphium caliginosum Meigen, 1824

= Raphium lanceolatum Loew, 1850, in Kazerani et al. 2013

Parent 1924, Rif, Cap Spartel; Kazerani et al. 2013 (no material provided); Kechev 2017; Nourti et al. 2019a, Rif, Amsemlil env. (1067 m), Dbani (Bni Selmane, 1046 m)

Rhaphium fascipes (Meigen, 1824)

= Porphyrops fascipes Meigen, 1824, in Parent 1927

Parent 1927, Rif, Tétouan

Rhaphium fissum Loew, 1850

Grichanov 2009, AA, Tizi-n’Test pass (2100 m)

Sciapodinae

Sciapus Zeller, 1842

Sciapus adumbratus Becker, 1902

Grichanov and Negrobov 2014, AP, near Essaouira, AA, Oued Souss, near Ouarzazate (1100 m)

Sciapus costae (Mik, 1890)

Negrobov 1991 (no material provided); Ebejer et al. 2019, Rif, Oued Laou (30 m)

Sciapus euzonus (Loew, 1859)

= Psilopus euzonus Loew, in Dakki 1997: 62

Parent 1927, Rif, El Mahadi; Séguy 1930a, Rif, Tanger; Séguy 1941d, AA, Taroudant

Sciapus glaucescens (Loew, 1856)

Grichanov and Negrobov 2014, AP, Oualidia

Sciapus heteropygus Parent, 1926

Nourti et al. 2019c, Rif, Talassemtane National Park (1696 m)

Sciapus holoxanthos Parent, 1926

Nourti et al. 2019c, Rif, Jbel Bouhachem, Adrou (556 m), Talassemtane National Park (1696 m)

Sciapus laetus (Meigen, 1838)

Grichanov 2009, AA, 40 km S Larache (0–20 m); Ebejer et al. 2019, Rif, Martil (9 m)

Sciapus longulus (Fallén, 1823)25

Pârvu et al. 2006, AP, Merja Zerga

Sciapus aff. negrobovi Naglis & Barták, 2015

Nourti et al. 2019a, Rif, plage Stihat (0 m), Kitane (49 m) (as Sciapus aff. negrobovi); Nourti et al. 2019c

Sympycninae

Campsicnemus Haliday in Walker, 1851

Campsicnemus crinitarsis Strobl, 1906

Dakki 1997 (no material provided); Grichanov 2012, AP, Essaouira; Kettani and Negrobov 2016, Rif, Oued Amsa

Campsicnemus curvipes (Fallén, 1823)26

Frey 1936 (no material provided)

Campsicnemus filipes Loew, 1859

Grichanov 2009, AA, 40 km S Larache (0–20 m)

Campsicnemus loripes (Haliday, 1832)

Ebejer et al. 2019, Rif, Moulay Abdelsalam (965 m)

Campsicnemus magius (Loew, 1845)

Pârvu et al. 2006, AA, Lac Edehby, Ouarzazate; Grichanov 2019, AP, Essaouira

Campsicnemus simplicissimus Strobl, 1906

Nourti et al. 2019a, Rif, plage Stihat (0 m)

Sympycnus Loew, 1857

Sympycnus pulicarius (Fallén, 1823)

Nourti et al. 2019a, HA, Aïn Taferaout (Sidi Masali, 1237 m)

Syntormon Loew, 1857

Syntormon aulicus Meigen, 1824

= Eutarsus aulicus Meigen, in Parent 1924, 1927; Séguy 1930a: 125

Parent 1927, Rif, Tanger; Séguy 1930a, Rif, Tanger; Vaillant 1952, 1956b, HA, Imi-N’Ifri

Syntormon codinai Parent, 1924

Parent 1924, Rif, Cap Spartel, Tanger; Parent, 1927, Rif, Tanger

Syntormon denticulatus (Zetterstedt, 1843)

= Syntormon pumilus Parent, 1925 (nec Meigen, 1824; misidentification), in Pârvu et al. 2006, Grichanov 2019

Parent 1924, Rif, Tétouan; Parent 1927, Rif, Tétouan; Séguy 1930a, Rif, Tanger; Pârvu et al. 2006, AA, Lac Edehby, Ouarzazate; Nourti et al. 2019a, Rif, plage Stihat (0 m), Amlay (294 m), Koudiat Taifour (100 m), Amsemlil env. (1067 m), Dayat Tazia (733 m), Oued Souk Lhad (613 m), HA, Aïn Taferaout (Sidi Masali, 1237 m), AA, Taliouine (1014 m)

Syntormon mikii Strobl, 1899

Parent 1927, “Maroc”; Kechev and Ivanova 2015; Nourti et al. 2019a, Rif, Pont de Dieu (Akchour, 536 m), Moulay Abdelsalam (1177 m)

Syntormon monilis (Haliday, 1851)

Parent 1924, Rif, Cap Spartel; Parent 1927, Rif, Cap Spartel

Syntormon pallipes (Fabricius, 1794)

Parent 1924, Rif, Cap Spartel, Tétouan, Chefchaouen; Parent 1927, Rif, Cap Spartel; Nourti et al. 2019a, Rif, Oued Souk Lhad (613 m), Talassemtane (339 m), Perdicaris Park (223 m), Amsemlil env. (1067 m); Dawah et al. 2020

Syntormon pilitibia Grichanov, 2013

Nourti et al. 2019a, Rif, Amsemlil (PNPB, 1067 m)

Syntormon pumilus (Meigen, 1824)

= Syntormon rufipes auctt. (nec Meigen, 1824; misidentification), in Pârvu et al. 2006; Grichanov 2019

Pârvu et al. 2006, AA, Lac Edehby, Ouarzazate

Syntormon zelleri (Loew, 1850)

Vaillant 1956b, HA, Oukaimeden, Izourar; Pârvu et al. 2006, AP, Merja Zerga; Nourti et al. 2019a, Rif, Pont de Dieu (Akchour, 536 m), Amsemlil env. (1059–1067 m)

Teuchophorus Loew, 1857

Teuchophorus cristulatus Mueffels & Grootaert, 1990

Ebejer et al. 2019, AA, 14 km E of Rich (Errachidia, 1278 m)

Teuchophorus rifensis Nourti, Grichanov & Kettani, 2019

Nourti et al. 2019b, Rif, Oued Souk Lhad (613 m)

Teuchophorus spinigerellus (Zetterstedt, 1843)

Pârvu et al. 2006, AA, Lac Edehby, Ouarzazate

Xanthochlorinae

Xanthochlorus Loew, 1857

Xanthochlorus tenellus (Wiedemann, 1817)

Grichanov 2009, AA, 40 km S Larache (0–20 m); Ebejer et al. 2019, Rif, Moulay Abdelsalam (1180 m)

HYBOTIDAE

K. Kettani, P. Gatt

Number of species: 44. Expected: 120

Faunistic knowledge of the family in Morocco: poor

Ocydromiinae

Bicellariini

Bicellaria Macquart, 1823

Bicellaria spuria Fallén, 1816

Becker and Stein 1913, Rif, Tanger

Ocydromiini

Ocydromia Meigen, 1820

Ocydromia glabricula (Fallén, 1816)

Becker and Stein 1913, Rif, Tanger

Oropezella Collin, 1926

Oropezella sphenoptera (Loew, 1873)

Cassar et al. 2005, Rif, lagoon Smir

Tachydromiinae

Drapetini

Chersodromia Haliday in Walker, 1851

Chersodromia albopilosa Chvála, 1970

Cassar et al. 2008, Rif, Basin Laou

Chersodromia nigrosetosa Chvála, 1970

Chvála 1981 (Ceuta); Chvála and Kovalev 1989

Chersodromia pseudohirta Chvála, 1970

Ebejer et al. 2019, Rif, Kabila beach

Crossopalpus Bigot, 1857

Crossopalpus aeneus (Walker, 1871)27

Shamshev et al. 2005, HA, Marrakech, Ouirgane

Crossopalpus dilutipes (Strobl, 1906)

Ebejer et al. 2019, AP, 9 km SE of Aïn Chouk (Lower Loukous marsh, 6 m)

Crossopalpus nigritellus (Zetterstedt, 1842)

Ebejer et al. 2019, Rif, Talassemtane (maison forestière), Issaguen (1620 m)

Crossopalpus setiger (Loew, 1859)

Ebejer et al. 2019 Rif, Smir lagoon

Drapetis Meigen, 1822

Drapetis disparilis Frey, 1936

Chvála 1981 (Ceuta); Chvála and Kovalev 1989

Drapetis laevis Becker, 1914

Becker and Stein 1913, Rif, Tanger; Kovalev 1970; Chvála and Kovalev 1989

Elaphropeza Macquart, 1827

Elaphropeza boergei Chvála, 1971

Ebejer et al. 2019, Rif, Smir lagoon, Oued Laou (saltmarsh)

Elaphropeza hutsoni Smith, 1967

Ebejer et al. 2019, Rif, Jnane Niche (46 m)

Stilpon Loew, 1859

Stilpon demnatensis Vaillant, 195628

Vaillant 1956b: 244, HA, Imi-N’Ifri

Stilpon moroccensis Grootaert & Zouhair, 2021

Grootaert et al. 2021, Rif, beach of Stehat, Bab Tariouant, Amsemlil, EM, Bouanane

Stilpon subnubilus Chvála, 1988

Ebejer et al. 2019, Rif, Smir lagoon, M’Diq (Kabila beach and dunes), Martil (beach and dunes)

Tachydromiini

Platypalpus Macquart, 1827

Platypalpus alluaudi Grootaert & Chvála, 1992

Grootaert and Chvála 1992, HA, Chichaoua; Vaňhara and Rozkošný 1997

Platypalpus anomalicerus (Becker, 1902)

= Coryneta aerivaga Séguy, in Séguy 1941d: 11

Séguy 1941d, EM, Oued Guir, AA, Agadir; Grootaert and Chvála 1992, EM, Oued Guir, AA, Agadir

Platypalpus annulatus (Fallén, 1815)

Pârvu et al. 2006, AP, Merja Zerga; Popescu-Mirceni 2011

Platypalpus anomalitarsis Chvála & Kovalev, 1974

Ebejer et al. 2019, MA, 10 km S of Azrou (1775 m), 10 km S of Azrou (1720 m), AA, Ziz river (30 km N of Erfoud, 894 m)

Platypalpus approximatus (Becker, 1902)

Grootaert and Chvála 1992, AP, Casablanca

Platypalpus asniensis Grootaert & Chvála, 1992

Grootaert and Chvála 1992, HA, Asni; Vaňhara and Rozkošný 1997

Platypalpus calceatus (Meigen, 1822)

Pârvu et al. 2006, MA, Meknès, AA, Foum Zghouig

Platypalpus chillcotti Chvála, 1981

Grootaert and Chvála 1992, MA, Ifrane

Platypalpus chrysonotus (Strobl, 1899)

Ebejer et al. 2019, Rif, Oued Laou (saltmarsh)

Platypalpus desertorum (Becker, 1907)

Ebejer et al. 2019, AA, 14 km E of Rich (Errachidia, 1278 m)

Platypalpus distichus Grootaert & Chvála, 1992

Ebejer et al. 2019, AP, 9 km SE of Aïn Chouk (Lower Loukous marsh, 6 m), AA, Ziz river (10 km S of Errachidia, 1008 m)

Platypalpus flavicornis (Meigen, 1822)

Ebejer et al. 2019, AA, 2 km N of Erfoud (818 m), Ziz river (30 km N of Erfoud, 894 m)

Platypalpus longicauda Grootaert & Chvála, 1992

Ebejer et al. 2019, Rif, Smir lagoon

Platypalpus lyneborgi Chvála, 1981

Grootaert and Chvála 1992, AP, Dradek, MA, Azrou

Platypalpus nigritarsis (Fallén, 1816)

Pârvu et al. 2006, MA, Ifrane; Popescu-Mirceni 2011

Platypalpus obscuripes (Strobl, 1899)

Ebejer et al. 2019, Rif, Martil (9 m), AP, Larache (Loukous marsh, 2 m)

Platypalpus ostiorum (Becker, 1902)

Grootaert and Chvála 1992

Platypalpus pachycerus (Collin, 1949)

Ebejer et al. 2019, AA, Ziz river (10 km S of Errachidia, 1008 m)

Platypalpus pallidiventris (Meigen, 1822)

Grootaert and Chvála 1992, MA, Ifrane; Maarouf 2003, HA, Chaouia

Platypalpus pseudoexiguus (Strobl, 1909)

Ebejer et al. 2019, Rif, Oued Laou (saltmarsh)

Platypalpus pseudounguiculatus (Strobl, 1909)

= Tachydromia pseudounguiculata Strobl 1909

Grootaert and Chvála 1992

Platypalpus riojaensis Chvála, 1981

Grootaert and Chvála 1992, EM, Oujda, MA, Meknès, HA, Chichaoua

Platypalpus turgidus (Becker, 1907)

Grootaert and Chvála 1992, MA, Takkat-n- Sountat

Platypalpus vockerothi Chvála, 1981

Grootaert and Chvála 1992, HA, Asni

Tachydromia Meigen, 1803

Tachydromia arrogans (Linnaeus, 1761)

Ebejer et al. 2019, Rif, Oued Laou, El-Fahsa (maquis)

Tachydromia annulimana Meigen, 182229

= Tachista annulimana Meigen, in Becker and Stein 1913: 84

Becker and Stein 1913, Rif, Tanger

Tachydromia undulata (Strobl, 1906)

Chvála 1969

Platypezoidea

PHORIDAE

K. Kettani, H. Disney

Number of species: 3. Expected: >400

Faunistic knowledge of the family in Morocco: very poor

Diplonevra Lioy, 1864

Diplonerva crassicornis (Meigen, 1830)

= Phora crassicornis Meigen, in Meigen 1830: 220

= Dohrniphora dudai Schmitz, in Schmitz 1920: 100

Raclin 1957; Mouna 1998; AP (Rabat) – MISR

Diplonevra tangeriana (Becker, 1913)30

= Phora tangeriana Becker and Stein, in Becker and Stein 1913: 90; Schmitz 1949: 238 (Species incerta)

Becker and Stein 1913, Rif, Tanger; Schmitz 1949

Megaselia Rondani, 1856

Megaselia minor (Zetterstedt, 1848)

= Trineura minor Zetterstedt, in Zetterstedt 1848: 2864; Wood 1912: 167

= Aphiochaeta angustifrons Wood, in Wood 1912: 167; Disney 1984: 239

= Phora minor Shob, in Mouna 1998: 86

Zetterstedt 1848; Wood 1912; Disney 1984; Mouna 1998; AP (Rabat) – MISR

PLATYPEZIDAE

K. Kettani, M.J. Ebejer

Number of species: 3. Expected: 6

Faunistic knowledge of the family in Morocco: poor

Platypezinae

Lindneromyia Meigen, 1804

Lindneromyia dorsalis (Meigen, 1804)

Chandler 2001, Rif, Chefchaouen (600 m), AP, Rabat, Maâmora; Tkoč and Roháček 2014; AP (Maâmora) – MISR

Microsania Zetterstedt, 1837

Microsania raclinae Collart31

Mouna 1998: 86

Protoclythia Kessel, 1949

Protoclythia rufa (Meigen, 1830)

Ebejer et al. 2019, Rif, Tahaddart (1 m)

LONCHOPTERIDAE

K. Kettani, M. Barták

Number of species: 4. Expected: 5

Faunistic knowledge of the family in Morocco: moderate

Lonchopterinae

Lonchoptera Meigen, 1803

Lonchoptera bifurcata (Fallén, 1810)

= Dispa furcata Fallén, in Vaillant 1989: 217

= Muscidora furcata Fallén, in Mouna 1998: 86

Vaillant 1989; Mouna 1998

Lonchoptera fallax De Meijere, 1906

= Muscidora fallax Meigen, in Mouna 1998: 86

Mouna 1998

Lonchoptera lutea Panzer, 1809

= Muscidora lutea Panzer, in Mouna 1998: 86

Vaillant 1989, HA (>3000 m); Mouna 1998; Pârvu et al. 2006, MA, Ifrane; Popescu-Mirceni 2011; MA – MISR

Lonchoptera tristis Meigen, 1824

= Muscidora tristis Meigen, in Mouna 1998: 86

Mouna 1998

Syrphoidea

PIPUNCULIDAE

K. Kettani, M.J. Ebejer

Number of species: 16. Expected: 50

Faunistic knowledge of the family in Morocco: poor

Chalarinae

Chalarus Walker, 1834

Chalarus brevicaudis Jervis, 1992

Ebejer and Kettani 2019b, Rif, Dardara (730 m), Azilane (1255 m), El Hamma (338 m)

Chalarus sp. aff. brevicaudis Jervis, 1992

Ebejer and Kettani 2019b, Rif, Azilane (1255 m)

Pipunculinae

Eudorylini

Claraeola Aczél, 1940

Claraeola sp. aff. halterata (Meigen, 1838)

Ebejer and Kettani 2019b, Rif, Akchour (424 m)

Clistoabdominalis Skevington, 2001

Clistoabdominalis dilatatus (De Meyer, 1997)

Ebejer and Kettani 2019b, Rif, Jebel Talassemtane (1546 m), El Hamma (338 m)

Dasydorylas Skevington, 2001

Dasydorylas setosus (Becker, 1908)

Kehlmaier 2005; Motamedinia et al. 2017

Eudorylas Aczél, 1940

Eudorylas ibericus Kehlmaier, 2005

Ebejer and Kettani 2019b, Rif, Jebel Talassemtane (1546 m)

Pipunculini

Pipunculus Latreille, 1802

Pipunculus carlestolrai Kuznetzov, 1993

Ebejer and Kettani 2019b, Rif, Jebel Talassemtane (1546 m)

Tomosvaryellini

Tomosvaryella Aczél, 1939

Tomosvaryella cilifemorata (Becker, 1907)

Ebejer and Kettani 2019b, Rif, Adrou (PNPB, 556 m)

Tomosvaryella debruyni De Meyer, 1995

Ebejer and Kettani 2019b, Rif, Adrou (PNPB, 556 m)

Tomosvaryella frontata (Becker, 1897)

Ebejer and Kettani 2019b, Rif, Oued Mhajrate (Ben Karrich, 67 m)

Tomosvaryella geniculata (Meigen, 1824)

Ebejer and Kettani 2019b, HA, Lac Tislite (Imilchil, 2254 m)

Tomosvaryella kuthyi Aczél, 1944

Ebejer and Kettani 2019b, Rif, El Hamma (338 m), Akchour (424 m), Barrage Smir (27 m)

Tomosvaryella minima (Becker, 1897)

Ebejer and Kettani 2019b, Rif, Koudiat Taifour (100 m)

Tomosvaryella mutata (Becker, 1898)

Ebejer and Kettani 2019b, Rif, Jebel Lakraâ (Talassemtane, 1596 m)

Tomosvaryella trichotibialis De Meyer, 1995

Ebejer and Kettani 2019b, Rif, Koudiat Taifour (100 m)

Tomosvaryella  sp. subvirescens group

Ebejer and Kettani 2019b, AP, Loukous marsh (Larache)

SYRPHIDAE

K. Kettani, M.C.D. Speight

Number of species: 166. Expected: more than 200

Faunistic knowledge of the family in Morocco: moderate

Eristalinae

Brachyopini

Brachyopa Meigen, 1822

Brachyopa atlantea Kassebeer, 2000

Kassebeer 2000, HA, Ouirgane (1000 m); Speight 2018; Sahib et al. 2020

Chrysogaster Meigen, 1803

Chrysogaster basalis Loew, 1857

Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Kassebeer 1999d; Sahib et al. 2020

Ighboulomyia Kassebeer, 1999

Ighboulomyia atlasi Kassebeer, 1999

Kassebeer 1999c, MA, Azrou, Umgebung, Timahdit, Ighböula Ulaichuor, Quellteich; Sahib et al. 2020

Myolepta Newman, 1838

Myolepta difformis Strobl in Czerny & Strobl 1909

= Myolepta philonis Séguy, 1961, in Dirickx 1994: 93

Dirickx 1994, HA; Reemer et al. 2004, MA, HA; Speight 2013, 2018; Sahib et al. 2020

Neoascia Williston, 1886

Neoascia clausseni Hauser & Kassebeer, 1998

= Neoascia podagrica (Fabricius, 1775), in Gil Collado 1929: 40; Claussen 1989b: 373

Gil Collado 1929a; Claußen 1989b; Dirickx 1994; Hauser and Kassebeer 1998, MA, HA, Taroudant (1800 m); Pârvu et al. 2006, AP, Merja Zerga; Popescu-Mirceni 2011; Sahib et al. 2020, Rif, Oued Jnane Niche, Oued Maggou

Orthonevra Macquart, 1829

Orthonevra bouazzai Kassebeer, 1999

Kassebeer 1999d, MA; Sahib et al. 2020

Orthonevra brevicornis (Loew, 1843)32

Sahib et al. 2020, Rif, Aïn Afersiw

Orthonevra elegans (Meigen, 1822)

Becker and Stein 1913, Rif; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Orthonevra schachti Claußen, 1989b

Claußen 1989b, HA, Oukaimeden (2600 m); Dirickx 1994; Schmid 1995; Kassebeer 1999d, MA; Sahib et al. 2020

Riponnensia Maibach, Goeldlin & Speight, 1994

Riponnensia longicornis (Loew, 1843)

= Orthonevra longicornis Loew, in Kanervo 1939: 2; Séguy 1961: 23; Kassebeer 1999c: 162

Kanervo 1939; Séguy 1961, AP; Claußen 1989b; Kassebeer 1999c, MA, HA; Dirickx 1994; Dousti and Hayat 2006; Speight 2013, 2018; Sahib et al. 2020

Riponnensia splendens (Meigen, 1822)

= Chrysogaster splendens (Meigen), in Gil Collado 1929: 405

= Orthonevra splendens (Meigen), in Claußen 1989b: 363, 373; Dirickx 1994: 97

Gil Collado 1929a, Rif; Mouna 1998; Claußen 1989b, HA, Tizi-n’Test (1900 m); Dirickx 1994; Kassebeer 1999c, MA; Sahib et al. 2020, MA, Douar Zaouiat Cheikh; AP (Dradek) – MISR

Callicerini

Callicera Panzer, 1806

Callicera fagesi Guérin-Meneville, 1844

Kassebeer 1998a, HA, Ouirgane (1000 m); Sahib et al. 2020

Callicera rufa Schummel, 1842

Gil Collado 1929a, Rif, Tanger; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Cerioidini

Ceriana Rafinesque, 1815

Ceriana conopsoides (Linnaeus, 1758)

= Cerioides conopsoides Linnaeus, in Séguy 1930a: 131

Séguy 1930a, MA, Ras El Ksar; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020; AP (Maghrawa, Maâmora) – MISR

Ceriana vespiformis (Latreille, 1804)

= Cerioides vespiformis Latreille, in Becker and Stein 1913: 88; Gil Collado 1929: 414; Séguy 1930a: 131; Kanervo 1939: 5; Leclercq 1961: 242

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a, Rif, HA; Séguy 1930a, AP, Rabat, Casablanca, MA, Tizi-s’Tkrine, Aïn Leuh, Meknès; Kanervo 1939, MA; Leclercq 1961a, Rif, Melillia, MA, Dayat Aoua, Aïn Leuh; Claußen 1989b; Dirickx 1994; Steenis et al. 2016; Speight 2018; El Hawagry and Gilbert 2019; Sahib et al. 2020, Rif, 1 km after Dardara, Meadow Mizoghar, Oued Achekrade

Sphiximorpha Rondani, 1850

Sphiximorpha subsessilis (Illiger in Rossi, 1807)

Steenis et al. 2016

Eristalini

Anasimyia Schiner, 1864

Anasimyia contracta Caußen & Torp, 1980

Kassebeer 1998a, MA, Timahdit (1850 m); Sahib et al. 2020

Eristalinus Rondani, 1845

Eristalinus aeneus (Scopoli, 1763)

= Lumpetia aenea (Scopoli), in Becker and Stein 1913: 86

= Eristalis aeneus (Scopoli), in Gil Collado 1929: 406, 407; Leclercq 1961: 242

= Lathyrophtalmus aeneus (Scopoli), in Séguy 1930: 129; Kanervo 1939: 5

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a, Rif, Tanger, AP, Mogador; Séguy 1930a, AP, Casablanca; Kanervo 1939; Timon-David 1951, AP, Rabat; Leclercq 1961a, EM, Melilla; Claußen 1989b; Dirickx 1994; Pârvu and Zaharia 2007; Sahib et al. 2020, Rif, Jumb Kitane, HA, vicinity of Asni

Eristalinus megacephalus (Rossi, 1794)

= Eristalis quinquelineatus Fabricius, in Becker and Stein 1913: 85, Gil Collado 1929: 407,

= Lathyrophthalmus quinquelineatus Fabricius, in Séguy 1930a: 129

Gil Collado 1929a, Rif, Tanger; Séguy 1930a, AP, Rabat, Oued Korifla, Sidi Bettache; Séguy 1961; Dakki 1997; Claußen 1989b; Dirickx 1994; Dousti and Hayat 2006; Sahib et al. 2020; AP (Rabat) – MISR

Eristalinus sepulchralis (Linnaeus, 1758)

= Eristalis sepuleralis Linnaeus, in Becker and Stein 1913: 85, Gil Collado 1929: 406

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a, AP, Claußen 1989b, HA, Ansegmir-Tal W Midelt (1400 m); Dirickx 1994; El Hawagry and Gilbert 2019; Sahib et al. 2020

Eristalinus taeniops (Wiedemann, 1818)

= Eristalis taeniops Wiedemann, in Becker and Stein 1913: 85

= Eristalodes taeniops Wiedemann, in Séguy 1930a: 130

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, AP, Oued Korifla, Rabat, HA, Tenfecht (Takeljount); Leclercq 1961a, MA, Dayet Aoua; Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Dakki 1997; Dousti and Hayat 2006; Koçak and Kemal 2010; Speight 2013, 2018; El Hawagry and Gilbert 2019; Sahib et al. 2020, Rif, Oued Martil, Halouma Kitane, Oued Sidi Yahia Aârab, MA, bridge Oued Oum-er-Rbia (Douar Ahl Souss), HA, Lac Oukaimeden; AP (Rabat), MA (Volubilis) – MISR

Eristalis Latreille, 1804

Eristalis arbustorum (Linnaeus, 1758)

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a, Rif; Séguy 1930a, MA, Aïn Leuh, Ras El Ksar, forest of Taffert; Kanervo 1939, Rif; Timon-David 1951, AP, Rabat, MA, Ifrane; Claußen 1989b, MA, Azrou (1700 m); Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Dakki 1997; El Hawagry and Gilbert 2019; Sahib et al. 2020, Rif, Aïn Sidi Brahim Ben Arrif, MA, vicinity of Ifrane, HA, vicinity of Asni, Lac Oukaimeden, AA, Douar Issafen, Douar Issafen; MA, HA – MISR

Eristalis jugorum Egger, 185833

Dakki 1997

Eristalis pertinax (Scopoli, 1763)

Séguy 1930a, AP, Oued Korifla, Sidi Bettache, MA, Forêt de Timelilt; Claußen 1989b; Dirickx 1994; Dakki 1997; Sahib et al. 2020

Eristalis similis (Fallén, 1817)

= Eristalis pratorum Meigen, in Gil Collado 1929: 407

Gil Collado 1929a, Rif; Séguy 1961; Claußen 1989b, HA, Oukaimeden (2600 m); Dirickx 1994; Pârvu and Zaharia 2007; Sahib et al. 2020, Rif, maison forestière, HA, Douar Zaouiat Cheikh, Lac Oukaimeden

Eristalis tenax (Linnaeus, 1758)

= Eristalomyia tenax Linnaeus, in Séguy 1930a: 130; Timon-David 1951: 146

Gil Collado 1929a, Rif, AP; Kanervo 1939, MA; Séguy 1949a, AA; Timon-David 1951; Leclercq 1961a; Claußen 1989b; Dirickx 1994; Sahib et al. 2020, Rif, Village Sebt Zinnat, Belyounech, Aïn Takhninjoute, maison forestière, Jumb Kitane, meadow Fahs Lmhar, HA, Douar Zaouiat Cheikh, vicinity of Asni, Lac Oukaimeden

Helophilus Meigen, 1822

Helophilus trivittatus (Fabricius, 1805)

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Mallota Meigen, 1822

Mallota cimbiciformis (Fallén, 1817)

= Mallota eristaloides Loew, in Becker and Stein 1913: 85

Becker and Stein 1913, Rif, Tange; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Mallota dusmeti Andreu, 1926

Kassebeer 1998a, HA, Ouirgane; Sahib et al. 2020

Melanogaster Rondani, 1857

Melanogaster lindbergi Kassebeer, 1999

= Chrysogaster macquardti Loew, in Becker and Stein 1913: 87

= Chrysogaster viduata Meigen, in Kanervo 1939: 2, Séguy 1961: 27, 28

= Chrysogaster lucida (Scopoli), in Claußen 1989b: 372

Becker and Stein 1913, Rif; Kanervo 1939, MA; Séguy 1961; Claußen 1989b; Dirickx 1994; Kassebeer 1999d, MA; Popov et al. 2020; Sahib et al. 2020

Myathropa Rondani, 1845

Myathropa florea (Linnaeus, 1758)

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a; Séguy 1930a, MA, Aïn Leuh; Claußen 1989b, MA, Azrou (1700 m); Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Dakki 1997; Sahib et al. 2020, Rif, Oued à 15 km de Fifi

Parhelophilus Girschner, 1897

Parhelophilus versicolor (Fabricius, 1794)

= Helophilus versicolor (Fabricius), in Gil Collado 1929: 407

Gil Collado 1929a, AP, Oulad Mesbah; Claußen 1989b; Dirickx 1994; El Hawagry and Gilbert 2019; Sahib et al. 2020

Eumerini

Eumerus Meigen, 1822

Eumerus amoenus Loew, 1848

Séguy 1961; Claußen 1989b; Dirickx 1994; Dousti and Hayat 2006; Speight 2013, 2018; Sahib et al. 2020, AA, Douar Aourir, beach of Tamelallt

Eumerus barbarus (Coquebert, 1804)

= Eumerus australis Meigen, in Gil Collado 1929: 412

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a, Rif; Séguy 1961; Claußen 1989b, MA, Azrou (1900 m); Dirickx 1994; Mouna 1998; Speight 2013, 2018; Steenis et al. 2017, AA, 11 km NW Taliouine, S Aït-Baha, 10 km NE Tafraoute; El Hawagry and Gilbert 2019; Sahib et al. 2020; AP (Cap Cantin) – MISR

Eumerus basalis Loew, 1848

= Eumerus angusticornis Rondani, in Séguy 1930a: 130

= Eumerus basalis Loew, in Mouna 1998: 86

Séguy 1930a, MA, forest of Timelilt; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020

Eumerus caballeroi Gil Collado, 1929

Gil Collado 1929a, AP, Laguna Gedira; Claußen 1989b; Dirickx 1994; Speight 2013, 2018; Sahib et al. 2020

Eumerus hungaricus Szilády, 1940

Speight 2018

Eumerus lunatus (Fabricius, 1794)

= Eumerus lunulatus Fabricius, in Becker and Stein 1913: 86

Becker and Stein 1913, Rif, Tanger; Leclercq 1961a, EM, Melilla; Claußen 1989b; Dirickx 1994; Mouna 1998; Dousti and Hayat 2006; Speight 2013, 2018; Sahib et al. 2020

Eumerus melotus (Séguy, 1941)

= Lampetia melota Séguy, in Séguy 1941d: 13

Séguy 1941d, AA, Agadir; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020

Eumerus nudus Loew, 1848

Becker and Stein 1913, Rif, Tanger; Claußen 1989b; Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Speight 2013, 2018; Sahib et al. 2020

Eumerus obliquus (Fabricius, 1805)

Sahib et al. 2020, Rif, Oued Jnane Niche, EM, Oued Khemis

Eumerus ornatus Meigen, 1822

Séguy 1930a, MA, Aïn Leuh, forest of Timelilt; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020; AP (Oued Cherrat) – MISR

Eumerus pulchellus Loew, 1848

Séguy 1961; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Eumerus punctifrons Loew, 1857

Leclercq 1961a, EM, Melilla; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020

Eumerus pusillus Loew, 1848

Claußen 1989b, HA, Tizi-n’Test (1900 m); Dirickx 1994; Sahib et al. 2020

Eumerus sabulonum (Fallén, 1817)

Séguy 1961; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Eumerus schmideggeri Steenis, Hauser & Zuijen, 2017

Steenis et al. 2017, AA, Sidi R’bat (37 km S Agadir); Sahib et al. 2020

Eumerus strigatus (Fallén, 1817)34

Kanervo 1939, Rif, HA; Timon-David 1951, AP, Rabat, Sehoul; Claußen 1989b, MA, Azrou (1700 m), HA, Oukaimeden (2200 m), Ansegmir-Tal W Midelt; Claußen and Hauser 1990, MA, Ifrane, Hajeb; Dirickx 1994; Mouna 1998; Speight 2013; Sahib et al. 2020

Eumerus subornatus Claußen, 1989b

Claußen 1989b, HA, Tizi-n’Test (1900 m); Schmid 1995; Speight 2013, 2018; Dirickx 1994; Sahib et al. 2020

Eumerus truncatus Rondani, 1868

Steenis et al. 2017, AA, S. Aït-Baha, 11 km NW Taliouine, 25 km NE Tizinit, 20 km E Tizinit, Assaka; Speight 2018; Sahib et al. 2020

Merodon Meigen, 180335

Merodon aberrans Egger, 1860

= Lampetia aberrans Egger, in Séguy 1961: 174

Séguy 1961; Claußen 1989b; Dirickx 1994; Marcos-Garcia et al. 2007; Koçak and Kemal 2010; Vujic et al. 2011; Speight 2013, 2018; Sahib et al. 2020

Merodon aeneus Meigen, 182236

= Lampetia aenea Meigen, in Becker and Stein 1913: 86; Kanervo 1939: 5; Timon-David 1951: 146

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a; Séguy 1930a, AP, Vallée Oued Korifla, MA, Tizi-s’Tkrine; Kanervo 1939, Rif; Timon-David 1951; Séguy 1961; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020

Merodon arrasus Becker, 192137

Becker 1921, Rif, Tanger; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Merodon aurifer Loew, 1862

= Merodon distinctus Palma, 1864

= Lampetia distincta Palm, in Timon-David 1951: 146

Timon-David 1951, AP, Zaer, MA, Ifrane; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020; Vujić et al. 2021c

Merodon avidus Rossi, 178238

= Lampetia spinipes (Fabricius), in Becker and Stein 1913: 86; Timon-David 1951: 146

= Merodon spinipes (Fabricius), in Gil Collado 1929: 409

= Lampetia avida Rossi, in Séguy 1961: 176

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a, Rif, HA; Séguy 1930a, AP, Chellah (Rabat), Oued Korifla, MA, Aïn Leuh; Kanervo 1939; Timon-David 1951, AP; Séguy 1961; Claußen 1989b, HA, Oukaimeden (2600 m); Hurkmans 1993; Dirickx 1994; Mouna 1998; Koçak and Kemal 2010; Likov et al. 2020; Sahib et al. 2020, HA, Lac Oukaimeden

Merodon bequaerti Hurkmans, 1993

Vujić et al. 2020a, EM, Mountain of Beni-Snassen, MA, Azrou

Merodon cabanerensis Marcos-García, Vujić & Mengual, 2007

Vujić et al. 2018, HA, Ait Mhamed (Azilal, 1700 m); Speight 2018; Sahib et al. 2020; Vujić et al. 2021

Merodon calcaratus (Fabricius, 1794)

Vujić et al. 2021b, EM, Mountains of Béni Snassen, near Nador, AP, 38 km SW of El Jadida, Garbouz

Merodon chalybeus Wiedemann in Meigen, 1822

= Lampetia spicata Becker, in Timon-David 1951: 146

= Merodon spicatus Becker, in Claußen 1989b: 365, 373

Timon-David 1951, AP, forest of Maâmora; Claußen 1989b, HA (2500 m); Dirickx 1994; Mouna 1998; Marcos-Garcia et al. 2007; Speight 2013, 2018; Sahib et al. 2020; AP (Cap Cantin) – MISR

Merodon clavipes (Fabricius, 1781)

Hurkmans 1993, MA; Marcos-García et al. 2007; Koçak and Kemal 2010; Sahib et al. 2020

Merodon eques Fabricius, 1805

= Lampetia eques (Fabricius), in Séguy 1961: 178

Séguy 1961; Claußen 1989b; Dirickx 1994; Ebejer and Bensusan 2010, AA; El Hawagry and Gilbert 2019; Sahib et al. 2020

Merodon equestris (Fabricius, 1794)

= Eristalis ferrugineus (Fabricius), in Fabricius 1805: 240

= Lampetia equestris (Fabricius), in Séguy 1961: 178, 179

Fabricius 1805, AP; Séguy 1961; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Merodon escalerai Gil Collado, 1929

Gil Collado 1929a, AP, Essaouira; Claußen 1989b; Dirickx 1994; Speight 2018; Sahib et al. 2020

Merodon femoratus Sack, 1913

= Merodon biarcuatus Curran, 1939, in Curran 1939: 6, 7; Claußen 1989: 373; Dirickx 1994: 79; Koçak and Kemal 2010: 1199; Speight 2018: 137

= Merodon elegans Hurkmans, 1993, in Hurkmans 1993: 195; Schmid 1995; Marcos-Garcia et al. 2007: 553; Speight 2018: 141

Curran 1939, AP, forest of Maâmora (Rabat); Claußen 1989b; Hurkmans 1993, AP; Dirickx 1994; Schmid 1995; Marcos-Garcia et al. 2007; Koçak and Kemal 2010; Speight 2018; Likov et al. 2020

Merodon geniculatus Strobl, 1909

Gil Collado 1929a, Rif; Claußen 1989b, HA (2500 m); Dirickx 1994; Mouna 1998; Marcos-Garcia et al. 2007; Koçak and Kemal 2010; Speight 2013, 2018; Sahib et al. 2020, HA, Lac Oukaimeden – MISR

Merodon hurkmansi Marcos-García, Vujić & Mengual, 2007

Marcos-García et al. 2007

Merodon ibericus Vujić, 2015

= Merodon bicolor Gil Collado, 1930

Popović et al. 2015, MA, Azrou, Ifrane; Acanski et al. 2016; Speight 2018; Sahib et al. 2020

Merodon italicus Rondani, 1845

Claußen and Hauser 1990, MA, Ifrane (1750 m); Mouna 1998

Merodon longicornis Sack, 1913

Claußen and Hauser 1990, MA; Dirickx 1994; Sahib et al. 2020

Merodon maroccanus Gil Collado, 1929

Gil Collado 1929a, AP, Essaouira; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Merodon minutus Strobl, 1893

= Lampetia minutus Strobl, in Séguy 1961: 180

Séguy 1961; Leclercq 1961a; Claußen 1989b; Dirickx 1994; Speight 2013, 2018; Sahib et al. 2020

Merodon monticolus Villeneuve, 1924

Kassebeer 1998a, HA, Ouirgane, Taftraoute, Taliouine; Sahib et al. 2020

Merodon murorum (Fabricius, 1794)

= Syrphus murorum (Fabricius), in Fabricius 1794: 288

= Merodon auripilus (Meigen), in Meigen 1830: 354

= Lampetia auripila Meigen, in Séguy 1941d: 13; Mouna 1998: 86

Fabricius 1794; Meigen 1830; Séguy 1941d, AA, Agadir; Séguy 1961; Claußen 1989b; Dirickx 1994; Mouna 1998; Vujić et al. 2018, AP, Essaouira; Sahib et al. 2020

Merodon pruni (Rossi, 1790)

= Lampetia pruni (Rossi), in Becker and Stein 1913: 86

= Merodon pruni var. obscurus Gil Collado, in Gil Collado 1929: 407, 408

Becker and Stein 1913; Gil Collado 1929a, Rif, Tanger; Claußen 1989b; Hurkmans, 1993; Dirickx 1994; Koçak and Kemal 2010; Sahib et al. 2020

Merodon pumilus Macquart, 1849

Ebejer et al. 2019, Rif, Moulay Abdelsalam (965 m); Sahib et al. 2020

Merodon rufus Meigen, 1838

= Lampetia rufa Meigen, 1838, in Becker and Stein 1913: 86

Becker and Stein 1913, Rif; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Merodon segetum (Fabricius, 1794)

Peck 1988; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Merodon serrulatus Wiedemann in Meigen, 1822

Hurkmans 1993; Speight 2013, 2018; Sahib et al. 2020; Vujić et al. 2020a

Merodon sophron Hurkmans, 1993

Hurkmans 1993, MA, Azrou; Schmid 1995; Koçak and Kemal 2010; Vujić et al. 2020a, MA, Azrou

Merodon tangerensis Hurkmans, 1993

Hurkmans 1993, Rif, Tanger; Schmid 1995; Koçak and Kemal 2010; Sahib et al. 2020

Merodon tricinctus Sack, 1913

= Lampetia tricincta Sack, in Timon-David 1951: 146

Timon-David 1951, MA, Ifrane; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020

Merodon unguicornis Strobl, 1909

Ebejer et al. 2019, MA, 10 km S of Azrou (1775 m); Sahib et al. 2020, Rif, maison forestière

Platynochaetus Wiedemann, 1830

Platynochaetus rufus Macquart, 1835

Gil Collado 1929a, AP, Mogador; Dirickx 1994; Sahib et al. 2020

Platynochaetus setosus (Fabricius, 1794)

Gil Collado 1929a, HA, Marrakech; Séguy 1953a, AA, Souss: Aïn Chaib; Claußen 1989b; Dirickx 1994; Speight 2013, 2018; Sahib et al. 2020

Milesiini

Milesia Latreille, 1804

Milesia crabroniformis (Fabricius, 1775)

Claußen and Hauser 1990, MA, Ifrane, Hajeb; Dirickx 1994; Sahib et al. 2020

Spilomyia Meigen, 1803

Spilomyia maroccana Kuznetzov, 1997

= Spilomyia digitata (Rondani), in Becker and Stein 1913: 86

Becker and Stein 1913, Rif, Tanger; Claußen 1989b, HA, Tizi-n’Test (1900 m); Claußen and Hauser 1990, MA, Ifrane, Hajeb; Kuznetzov 1997; Dirickx 1994; Kassebeer 1999d; Steenis 2000; Sahib et al. 2020

Syritta Le Peletier & Serville, 1828

Syritta flaviventris Macquart, 1842

Claußen 1989b, AP, Kénitra; Dirickx 1994; Sahib et al. 2020, EM, farm Saf-Saf

Syritta pipiens (Linnaeus, 1758)

Becker and Stein 1913, Rif, Tanger; Gill Collado 1929a; Séguy 1930a, AP, Rabat, Casablanca, MA, Meknès; Kanervo 1939; Timon-David 1951, AP, Rabat, AA, Agdz; Leclercq 1961a, EM, Melilla; Claußen 1989b, HA, Tizi-n’Test (1900 m); Claußen and Hauser 1990, MA, Ifrane, Hajeb; Dirickx 1994; Mouna 1998; Pârvu et al. 2006, AP, Merja Zerga; Popescu-Mirceni 2011; Sahib et al. 2020, Rif, Bni Maaden, dam Moulay Bouchta, Oued Koub, Douar Kitane, Oued Maggou, EM, farm Saf-Saf, MA, vicinity of Ifrane, HA, vicinity of Asni, Tizi-n’Test, AA, Oued Assa; AP (Rabat), MA (Aïn Leuh), SA – MISR

Temnostoma Le Peletier & Serville, 1828

Temnostoma bombylans (Fabricius, 1805)

Séguy 1961, MA; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Xylota Meigen, 1822

Xylota segnis (Linnaeus, 1758)

= Zelima (Xylota) segnis Linnaeus, in Becker and Stein 1913: 86; Timon-David 1951: 147

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a; Timon-David 1951, HA, Zaouia Ahansal; Leclercq 1961a, Rif, Azib de Ketama; Claußen 1989b; Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Mouna 1998; Sahib et al. 2020, Rif, Aïn El Maounzil, Oued Koub, Oued Sidi Ben Saâda, HA, vicinity of Asni; HA (Zaouiet Ahansal) – MISR

Rhingiini

Cheilosia Meigen, 1822

Cheilosia brunnipennis Becker, 1894

= Chilosia flavipes (Panzer), in Kanervo 1939: 2

Kanervo 1939, HA; Kassebeer, 1998c, MA; Séguy 1961; Claußen 1989b; Speight 2013, 2018; Sahib et al. 2020

Cheilosia flavipes (Panzer, 1798)

Mouna 1998: 86

Cheilosia grossa (Fallén, 1817)

Kassebeer 1998c, HA, Asif Mellah, Tizi-n’Tichka; Speight 2013, 2018; Sahib et al. 2020

Cheilosia latifrons (Zetterstedt, 1843)

= Cheilosia intonsa Loew, in Timon-David 1951: 144

Timon-David 1951, AP, Sehoul; Claußen 1989b; Dirickx 1994; Mouna 1998; Kassebeer 1998c, Rif, Ouezzane, AP, Oued Loukous, Larache, MA, Ifrane, HA, Oukaimeden; Sahib et al. 2020

Cheilosia mutabilis (Fallén, 1817)

Speight 2013, 2018

Cheilosia griseiventris Loew, 1857

= Chilosia marokkana Becker, in Becker 1894: 395; Becker and Stein 1913: 87

= Cheilosia maroccana Becker, 1894, in Gil Collado 1929: 405

Becker 1894; Becker and Stein 1913; Gil Collado 1929a; Séguy 1961; Claußen 1989b; Dirickx 1994; Kassebeer 1998c, MA, HA; Sahib et al. 2020

Cheilosia paralobi Malski, 1962

= Cheilosia longula (Zetterstedt, 1838), in Gil Collado 1929: 405

Gil Collado 1929a; Claußen 1989b, MA, Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Kassebeer 1998c, HA; Speight 2013, 2018; Sahib et al. 2020

Cheilosia parva Kassebeer, 1998

Kassebeer 1998c, MA, Azrou, Ifrane (1650 m); Claußen and Speight 2007; Sahib et al. 2020

Cheilosia rodgersi Wainwright, 1911

Becker and Stein 1913, Rif, Tanger; Claußen 1989a; Dirickx 1994; Kassebeer 1998c, Rif, Tanger; Speight 2013, 2018; Sahib et al. 2020

Cheilosia scutellata (Fallén, 1817)

Gil Collado 1929a, Rif; Claußen 1989b; Dirickx 1994; Kassebeer 1998c, Rif, Chefchaouen, MA, Ouiouane, Ifrane; Sahib et al. 2020

Cheilosia soror (Zetterstedt, 1843)

= Cheilosia rufipes (Preyssler, 1793), in Claussen and Hauser 1990: 436, Kassebeer 1998c: 65

Claußen and Hauser 1990, MA, Ifrane (1750 m); Kassebeer 1998c, MA, Ifrane; Dirickx 1994; Sahib et al. 2020

Cheilosia variabilis (Panzer, 1798)

Kassebeer 1998c, MA, Ifrane; Speight 2013, 2018; Khaganinia and Kazerani 2014; Sahib et al. 2020

Ferdinandea Rondani, 1844

Ferdinandea fumipennis Kassebeer, 1999

Kassebeer 1999b, MA, Ifrane, Azrou, HA, Marrakech, Ouirgane; Speight 2013, 2018; Sahib et al. 2020

Volucellini

Volucella Geoffroy, 1762

Volucella inanis (Linnaeus, 1758)

Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Sahib et al. 2020

Volucella liquida Erichson, 1841

Gil Collado 1929a, Rif; Séguy 1930a, AP, Mogador, MA, Azrou, Bekrit; Kanervo 1939, MA; Séguy 1953a, MA, Ifrane; Timon-David 1951, MA, Ifrane, Azrou, HA, Aït Mohamed Sgatt; Leclercq 1961a, Rif, Azib de Ketama, MA, Dayat Aoua, Ifrane, Azrou; Claußen 1989b; Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Mouna 1998; Sahib et al. 2020, Rif, jumb Kitane – MISR

Volucella zonaria Poda, 1761

Séguy 1930a, AP, Casablanca; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020; AP (Rabat, Casablanca) – MISR

Brachypalpus Macquart, 1834

Brachypalpus valgus (Panzer, 1798)

Kassebeer 1998a, MA, Ifrane, HA, Ouirgane; Sahib et al. 2020

Psilotini

Psilota Fallén, 1823

Psilota atra (Fallén, 1817)

= Psilota toubkalana Kassebeer, 1995, in Kassebeer 1995: 395–400

Kassebeer 1995, HA; Smit and Vujic 2008, HA, Ouirgane, Marrakech; Speight 2018; Sahib et al. 2020, MA, Douar Zaouiat Cheikh

Pipizinae

Pipizini

Heringia Rondani, 1856

Heringia heringi (Zetterstedt, 1843)

Kassebeer 1998a, HA, Tahanaout, Ouirgane; Sahib et al. 2020

Pipizella Rondani, 1856

Pipizella thapsiana Kassebeer, 1995

Kassebeer 1995, HA (1000 m); Speight 2013, 2018; Sahib et al. 2020, MA, Douar Zaouiat Cheikh

Triglyphus Loew, 1840

Triglyphus escalerai Gil Collado, 1929

Gil Collado 1929a, Rif, Tanger; Dirickx 1994; Speight 2013; Sahib et al. 2020

Syrphinae

Bacchini

Melanostoma Schiner, 1860

Melanostoma mellinum (Linnaeus, 1758)

Becker and Stein 1913, Rif, Tanger; Séguy 1934b, AP, Korifla; Timon-David 1951, MA, Ifrane, Azrou, Aïn Leuh, HA, Aït Mizane; Leclercq 1961a, MA, Ifrane; Claußen 1989b, HA, Ansegmir-Tal W Midelt (1400); Claußen and Hauser 1990, MA, Ifrane (1750 m); Mouna 1998; Pârvu and Zaharia 2007; Sahib et al. 2020, Rif, Dayat Rahrah, Aïn el Ma Bared, Oued Dardara, Dayat El Ânassar, Dayat Lemtahane, Garden Ksar Al Rimal, tributary Oued Tazarine, Oued Farda, Dayat El Birdiyel, maison forestière, stream at 1 km from Sidi Yahia Aârab, Dayat Amsemlil, EM, farm Saf-Saf, MA, Oued d’Ifrane, HA, Lac Oukaimeden – MISR

Melanostoma mundum Czerny & Strobl, 1909

Dakki 1997; Mouna 1998

Melanostoma scalare (Fabricius, 1794)

Gil Collado 1929a, Rif; Claußen 1989b; Dirickx 1994; Sahib et al. 2020, Rif, Aïn el Ma Bared, Oued Mezine, Aïn Quanquben, Aïn Takhninjoute, Oued Koub

Platycheirus Le Peletier & Serville, 1828

Platycheirus albimanus (Fabricius, 1781)39

Séguy 1930a, HA, Tizi-n’Test, Jebel Imdress; Mouna 1998; Kassebeer 1998b

Platycheirus ambiguus (Fallén, 1817)

Kassebeer 1998b, HA; Sahib et al. 2020

Platycheirus atlasi Kassebeer, 1998

Kassebeer 1998b, MA, Azrou, Ifrane; Sahib et al. 2020

Platycheirus fulviventris (Macquart, 1829)

Ebejer et al. 2019, AA, 14 km E of Rich (Errachidia, 1278 m); Sahib et al. 2020

Platycheirus manicatus (Meigen, 1822)

Séguy 1961; Claußen 1989b; Dirickx 1994; Mouna 1998; Kassebeer 1998b, HA, Toubkal; Sahib et al. 2020

Platycheirus marokkanus Kassebeer, 1998

Kassebeer 1998b, MA, HA; Speight 2018; Sahib et al. 2020, Rif, Aïn Takhninjoute, HA, Douar Akhlij Tnine Ourika

Xanthandrus Verrall, 1901

Xanthandrus comtus (Harris, 1776)

Gil Collado 1929a, Rif, Tanger; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Paragini

Paragus Latreille, 1804

Paragus albifrons (Fallén, 1817)

Kanervo 1939; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020

Paragus atlasi Claußen, 1989

Claußen 1989b, HA, Tizi-n’Test (1900 m); Dirickx 1994; Schmid 1995; Speight 2013, 2018; Sahib et al. 2020

Paragus bicolor (Fabricius, 1794)

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA, Tizi-s’Tkrine, forest of Timelilt, Azrou; Kanervo 1939, AP, HA; Séguy 1949a, SA, Guelmim; Timon-David 1951, MA, Ifrane; Claußen 1989b; Claußen and Hauser 1990, HA, Tizi-n’Test; Dirickx 1994; Dakki 1997; Sahib et al. 2020, Rif, Douar Dacheryène, HA, vicinity of Asni; MA (Fès, Ifrane, Azrou) – MISR

Paragus cinctus Schiner & Egger, 1853

Claußen 1989b, HA, Tizi-n’Test (1900 m); Dirickx 1994; Speight 2013, 2018; Sahib et al. 2020, HA, vicinity of Asni

Paragus coadunatus Rondani, 1847

Claußen and Hauser 1990, MA, Ifrane, Hajeb; Dirickx 1994; Speight 2013, 2018; Sahib et al. 2020

Paragus flammeus Goeldlin, 1971

Claußen and Hauser 1990; Dirickx 1994; Speight 2013, 2018; Sahib et al. 2020

Paragus haemorrhous Meigen, 1844

Claußen 1989b, MA, Azrou (1700 m); Dirickx 1994; Sahib et al. 2020, Rif, Oued Mharhar

Paragus hermonensis Kaplan, 1981

Claußen 1989b, MA, Azrou (1700 m); Dirickx 1994; Sahib et al. 2020

Paragus quadrifasciatus Meigen, 1822

= Paragus pulcherrimus Strobl, in Timon-David 1951: 144

Timon-David 1951, MA, Ifrane; Claußen 1989b, AP, Kénitra; Claußen and Hauser 1990, MA, Ifrane, Hajeb; Dirickx 1994; Mouna 1998; Sahib et al. 2020, Rif, Douar Kitane, AA, Agadir airport; MA (Ifrane) – MISR

Paragus majoranae Rondani, 1857

Claußen and Hauser 1990, MA; Dirickx 1994; Sahib et al. 2020

Paragus pecchiolii Rondani, 1857

Claußen and Hauser 1990, MA, Ifrane (1750 m), Hajeb

Paragus strigatus Meigen, 1822

= Paragus bimaculatus Meigen, in Wiedemann 1824: 33

Wiedemann 1824, AP; Claußen 1989b; Claußen and Hauser 1990, MA, Ifrane (1750 m), Hajeb; Dirickx 1994; Speight 2018; Sahib et al. 2020

Paragus tibialis (Fallén, 1817)

= Paragus tibialis meridionalis Becker, in Becker and Stein 1913: 88; Gil Collado 1929: 403; Leclercq 1961: 241

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a; Kanervo 1939; Séguy 1949a, SA, Guelmim; Leclercq 1961a, Rif, Bab Taza, MA, Taza; Claußen 1989b; Claußen and Hauser 1990, HA, Tizi-n’Test (2000 m); Dirickx 1994; Dakki 1997; Grabener 2017; Sahib et al. 2020, HA, vicinity of Asni, Ijoukak vicinity; AP (Rabat) – MISR

Paragus vandergooti Marcos-Garcia, 1986

Claußen 1989b, HA, Tizi-n’Test à (1900 m); Claußen and Hauser 1990, MA, Ifrane, Hajeb; Dirickx 1994; Speight 2013, 2018; Sahib et al. 2020

Syrphini

Chrysotoxum Meigen, 1803

Chrysotoxum bicinctum (Linnaeus, 1758)

Timon-David 1951, MA, Ifrane, HA, Haute Réghaya; Leclercq 1961a, MA, Mischliffen (2019 m); Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020; MA, HA – MISR

Chrysotoxum intermedium Meigen, 1822

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a; Séguy 1930a, MA, Tizi-S’Tkrine, Aïn Leuh, forest of Taffert, HA, Tizi-n’Test, Goundafa; Kanervo 1939; Claußen 1989b; Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Mouna 1998; Pârvu et al. 2006, AP, Cap Bedouza; Dousti and Hayat 2006; Pârvu and Zaharia 2007; Kazerani et al. 2013b; Sahib et al. 2020, Rif, 1 km after Dardara, Oued Azila, Dayat Jebel Zemzem, Aïn El Maounzil, Oued Tafoughalt, MA, Douar Zaouiat Cheikh, HA, vicinity of Asni, AA Douar Issafen; AP (Rabat), MA (Aïn Leuh), HA (Réghaya) – MISR

Chrysotoxum volaticum Séguy, 1961

Séguy 1961, MA; Claußen 1989b, HA, Oukaimeden (2600 m); Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Sahib et al. 2020

Dasysyrphus Enderlein, 1938

Dasysyrphus albostriatus (Fallén, 1817)

Kassebeer 1998a, HA, Bin-el-Ouidane, Ouirgane, Imlil, Asni; Sahib et al. 2020

Epistrophe Walker, 1852

Epistrophe eligans (Harris, 1780)

= Syrphus ochrostoma (Zetterstedt), in Becker and Stein 1913: 88; Claußen 1989b: 372

Becker and Stein 1913, Rif, Tanger; Claußen 1989b; Kassebeer 1998a, MA, Ifrane, HA, Imlil, Asni, Ouirgane; Sahib et al. 2020, Rif, Dayat Tazia

Epistrophe eligans (Harris, 1870) var. trifasciata Strobl

Ebejer and Bensusan 2010, AA; Djellab et al. 2013

Episyrphus Matsumura & Adachi, 1917

Episyrphus balteatus (De Geer, 1776)

= Syrphus balteatus De Geer, in Becker and Stein 1913: 88; Séguy 1930a: 129

= Epistrophe balteata (De Geer), in Gil Collado 1929: 406; Kanervo 1939: 3; Timon-David 1951: 144

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a; Séguy 1930a, AP, Rabat, MA, Aïn Sferguila; Timon-David 1951, AP, forest of Maâmora, Rabat, HA, Marrakech; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020, Rif, Sebt Zinate, Aïn Sidi Brahim Ben Arrif, dam Nakhla, Aïn Boughaba, Garden Ksar Al Rimal, Oued Aârkoub, Dayat Rahrah, Oued Sahel, dam Moulay Bouchta, maison forestière, Ksar El Kébir, Dayat Jebel Zemzem, Oued Taida, stream at 1 km from Sidi Yahia Aârab, Aïn Quanquben, Oued Maggou, Forest Bab El Karn, Douar Kitane, forest El Mahfoura, HA, Aïn Zarka of Meski, AA, Douar Zaouia; AP (Rabat) – MISR

Eupeodes Osten-Sacken, 1877

Eupeodes corollae (Fabricius, 1794)

= Syrphus berber Bigot, in Bigot 1884: 88

= Syrphus corollae Meigen, in Becker and Stein 1913: 88; Séguy 1930a: 129, Kanervo 1939: 3; Gil Collado 1929: 406

= Syrphus corollae Fabricius, in Timon-David 1951: 144

= Metasyrphus corollae (Fabricius), in Dirickx 1994: 89

Bigot 1884; Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a; Séguy 1930a, HA, Tizi-n’Test, Jebel Imdress, Goundafa; Kanervo 1939; Timon-David 1951, AP, forest of Maâmora, Rabat, Sidi Taibi, HA, Réghaya, Tazzarine, AA, Agdz, Plaine de Souss (Taroudant); Leclercq 1961a, MA, Ifrane; Claußen and Hauser 1990, MA, Ifrane (1750 m), HA, Oukaimeden (3200 m), AA, Tan-Tan; Dirickx 1994; Mouna 1998; Pârvu and Zaharia 2007; Grabener 2017; El Hawagry and Gilbert 2019; Sahib et al. 2020, Rif, Village Sebt Zinate, Aïn el Ma Bared, Garden Ksar Al Rimal, Oued Bin EL Ouidane, Oued Sahel, Aïn Takhninjoute, stream at 1 km from Sidi Yahia Aârab, Oued Jnane Niche, dam Smir, Oued Boumarouil, Dayat Jebel Zemzem, Oued Maggou, Meadow Fahs Lmhar, Douar Kitane, forest El Mahfoura, MA, Douar Zaouiat Cheikh, HA, vicinity of Asni, Ijoukak vicinity, Lac Oukaimeden; AA Agdz – MISR

Eupeodes latifasciatus (Macquart, 1829)

= Syrphus latifasciatus Macquart, in Séguy 1949: 156; Séguy 1953a: 84

= Metasyrphus latifasciatus (Macquart), in Dirickx 1994: 89

Séguy 1949a, SA, Guelmim; Séguy 1953a, AA, Oued Khoref; Claußen 1989b; Dirickx 1994; Dakki 1997; Sahib et al. 2020, Rif, Douar Kitane, maison forestière, Oued Ametrasse

Eupeodes luniger (Meigen, 1822)

= Metasyrphus luniger (Meigen), in Dirickx 1994: 90, 237

= Syrphus luniger Meigen, in Gil Collado 1929: 406

Gil Collado 1929a, AP; Claussen 1989b; Dirickx 1994; Sahib et al. 2020, Rif, Oued Maggou, Oued Martil, Belyounech, Douar Kitane, MA, Douar Ben Smim, HA, vicinity of Asni

Eupeodes nuba (Wiedemann, 1830)

= Syrphus rufinasutus Bigot, in Bigot 1884: 88, Séguy 1961: 107

= Metasyrphus nuba (Wiedemann), in Dirickx 1994: 90, 238

Bigot 1884; Séguy 1961; Claußen 1989b, HA, Ansegmir-Tal W Midelt (1400 m); Dirickx 1994; Dousti and Hayat 2006; Ehteshamnia et al. 2010; Naderloo et al. 2011; Speight 2013, 2018; Kazerani et al. 2013b; El Hawagry and Gilbert 2019; Sahib et al. 2020

Eupeodes punctifer (Frey, 1934)40

Mouna 1998: 86

Ischiodon Sack, 1913

Ischiodon aegyptius (Wiedemann, 1830)

= Simosyrphus aegyptius (Wiedemann, 1830)

Gil Collado 1929a; Timon-David 1951, AP, Rabat, AA, Agdz; Mouna 1998; Grabener 2017; Mengual 2018; Sahib et al. 2020; AP (Rabat) – MISR

Lapposyrphus Dušek & Láska, 1967

Lapposyrphus lapponicus (Zetterstedt, 1838)

= Syrphus arcuatus Fallén, 1817, in Becker and Stein 1913: 88

= Metasyrphus lapponicus (Zetterstedt, 1838), in Dirickx 1994: 89

= Eupeodes lapponicus (Zetterstedt, 1838), in Claußen 1989b: 372

Becker and Stein 1913, Rif; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Meliscaeva Frey, 1946

Meliscaeva auricollis (Meigen, 1822)

= Epistrophe auricollis Meigen, in Becker and Stein 1913: 89; Gil Collado 1929: 406; Timon-David 1951: 145

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a; Timon-David 1951, AP, Oued Korifla, Zaers, Rabat; Claußen 1989b; Dirickx 1994; Mouna 1998; El Hawagry and Gilbert 2019; Sahib et al. 2020, Rif, Aïn el Ma Bared, Dayat El Ânassar, Belyounech, Oued Mezine, dam Moulay Bouchta, Aïn Afersiw, dam Entrasol, Oued à 15 km de Fifi, jumb Kitane, Oued Maggou, Douar Kitane, Oued Sahel, MA, Aïn Ouilili; AP (Rabat, Zaers) – MISR

Meliscaeva cinctella (Zetterstedt, 1843)

= Syrphus cinctellus Zeterstedt, in Séguy 1934b: 162

Séguy 1934b, MA, Oued Leben (Taounate); Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020 – MISR

Scaeva Fabricius, 1850

Scaeva albomaculata (Macquart, 1842)

= Lasiopticus albomaculata (Macquart), in Gil Collado 1929: 405; Timon-David 1951: 145

= Lasiophthicus albomaculatus Macquart, in Séguy 1953a: 84

Gil Collado 1929a; Séguy 1953a, MA, Immouzer; Timon-David 1951, AP, Rabat, MA, El Harcha; Leclercq 1961a, MA, Azrou; Claußen 1989b, MA, Azrou (1700 m); Claußen and Hauser 1990, MA, Ifrane; Dirickx 1994; Mouna 1998; Dousti and Hayat 2006; Ehteshamnia et al. 2010; Naderloo et al. 2011; Speight 2013, 2018; Kazerani et al. 2013b; Grabener 2017; El Hawagry and Gilbert 2019; Sahib et al. 2020, Rif, Belyounech, HA, Aïn Zarka of Meski, Lac Oukaimeden; AP (Rabat, Cap Cantin), EM (Debdou) – MISR

Scaeva dignota (Rondani, 1857)

Claußen and Hauser 1990, MA, Ifrane (1750 m); Dirickx 1994; Sahib et al. 2020, Rif, Oued Maggou, Dayat Lemtahane

Scaeva mecogramma (Bigot, 1860)

Dirickx 1994; Kassebeer 1998a, Rif, Chefchaouen, AP, Kénitra, HA, Ouirgane; Sahib et al. 2020

Scaeva pyrastri (Linnaeus, 1758)

= Catabomba pyrastri Linnaeus, in Becker and Stein 1913: 88

= Lasiophthicus pyrastri Linnaeus, in Séguy 1930a: 128

= Lasiopticus pyrastri Linnaeus, in Gil Collado 1929: 405, Timon-David 1951: 145

Becker and Stein 1913, Rif; Gil Collado 1929a, AP; Séguy 1930a, AP, forest of Zaers, forest of Maâmora, MA, Tizi-s’Tkrine; Timon-David 1951, AP, Rabat, MA, Ifrane; Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020, Rif, Dayat Jebel Zemzem, stream at 1 km from Oued Sidi Yahia Aârab, AA, 1 km before Douar Aïn Lahmar; AP (Rabat, Cap Cantin) – MISR

Scaeva selenitica (Meigen, 1822)

= Lasiophthicus seleniticus Meigen, in Séguy 1930a: 128

Séguy 1930a, HA, Aguerd el Had, AA, Talekjount (Souss); Claußen 1989b; Dirickx 1994; Mouna 1998; Sahib et al. 2020

Sphaerophoria Le Peletier & Serville, 1828

Sphaerophoria interrupta (Fabricius, 1805)

= Sphaerophoria menthastri (Linnaeus), in Becker and Stein 1913: 87; Kanervo 1939: 3; Timon-David 1951: 145; Séguy 1961: 109

Kanervo 1939, AP, MA, HA; Becker and Stein 1913, Rif; Timon-David 1951, AP, Kénitra, Rabat, MA, Harcha, Ifrane, Sefrou, HA, Agdz; Claußen 1989b; Dirickx 1994; Mouna 1998; El Hawagry and Gilbert 2019; Sahib et al. 2020; AP (Rabat), MA (Ifrane, Meknès) – MISR

Sphaerophoria rueppelli (Wiedemann, 1830)

Kanervo 1939, Rif, HA; Timon-David 1951, AP, Rabat, AA, Agadir, Agdz, Zagora; Séguy 1961; Claußen 1989b, HA, Ansegmir-Tal W Midelt (1400 m); Dirickx 1994; Mouna 1998; El Hawagry and Gilbert 2019; Sahib et al. 2020, Rif, Tarmast tributary, Oued Sidi Ben Saâda, Dayat Amsemlil, EM, farm Saf-Saf, MA, Oued d’Ifrane; AP (Rabat), AA (Agadir, Agdz, Zagora) – MISR

Sphaerophoria scripta (Linnaeus, 1758)

= Sphaerophoria dispar (Meigen), in Timon-David 1951: 145

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a, Rif; Séguy 1930a, AP, Tlet n’Rhohr, MA, forest of Timelilt, Aïn Leuh, EM, Berkane; Kanervo 1939, Rif, AP, MA, HA; Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Timon-David 1951, AP, Rabat, MA, Harcha, Sefrou, Ifrane; Leclercq 1961a, MA, Dayat Aoua, Ifrane, Azrou; Claußen 1989b, MA, Azrou (1700 m), HA, Oukaimeden, Tizi-n’Test (1900 m), Ansegmir-Tal W midelt (1400 m); Claußen and Hauser 1990, MA, Ifrane, Hajeb (1750 m), HA, Oukaimeden (3200 m); Dirickx 1994; Mouna 1998; Pârvu and Zaharia 2007; El Hawagry and Gilbert 2019; Sahib et al. 2020, Rif, Oued Aârkoub, dam Nakhla, meadow Mizoghar, Oued Dardara, 1 km after Dardara, Dayat El Birdiyel, palm grove Igrane, Aïn Quanquben, maison forestière, Oued Sidi Ben Saâda, Dayat Lemtahane, Dayat Amsemlil, forest El Mahfoura, EM, farm Saf-Saf, MA, Oued d’Ifrane, HA, vicinity of Asni, Lac Oukaimeden, AA, Msidira – MISR

Sphaerophoria taeniata (Meigen, 1822)

= Sphaerophoria menthastri var. taeniata Meigen, in Timon-David 1951: 10

Timon-David 1951, AP, Rabat, saline mud; Claußen 1989b; Dirickx 1994; Sahib et al. 2020; MA (Aïn Leuh, Azrou, Timahdit) – MISR

Syrphus Fabricius, 1775

Syrphus ribesii (Linnaeus, 1758)

Kassebeer 1998a, Rif, Chefchaouen, Tétouan; Sahib et al. 2020, Rif, Douar Kitane

Syrphus vitripennis Meigen, 1822

Gil Collado 1929a, Rif, Tanger; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Xanthogramma Schiner, 1860

Xanthogramma dives (Rondani, 1857)

Ebejer et al. 2019, AA, 29 km N of Rich (Errachidia, 1570 m); Sahib et al. 2020

Xanthogramma evanescens Becker, 1913

Becker and Stein 1913, Rif; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Xanthogramma marginale (Loew, 1854)

= Xanthogramma marginale var. morenae Loew, in Becker and Stein 1913: 86, Gil Collado 1929: 406

Becker and Stein 1913, Rif, Tanger; Gil Collado 1929a, Rif; Kanervo 1939, Rif, HA; Séguy 1961; Claußen and Hauser 1990, MA, Ifrane (1750 m); Mouna 1998; Claußen 1989b; Dirickx 1994; Ebejer and Bensusan 2010, AA; Speight 2013, 2018; Sahib et al. 2020, Rif, village Sebt Zinate, Oued Maggou, Oued Ametrasse, Douar Kitane

Xanthogramma pedissequum (Harris, 1776)

= Xanthogramma ornatum (Meigen, 1822), in Gil Collado 1929: 406

Gil Collado 1929a, Rif, Tanger; Claußen 1989b; Dirickx 1994; Sahib et al. 2020

Conopoidea

CONOPIDAE41

K. Kettani

Number of species: 34. Expected: 40

Faunistic knowledge of the family in Morocco: good

Conopinae

Conopini

Conops Linnaeus, 1758

Conops aegyptiacus (Rondani, 1850)

Kröber 1915, 1928

Conops ceriaeformis Meigen, 1804

= Conops acuticornis Loew, 1847, in Becker and Stein 1913: 89

Becker and Stein 1913, Rif, Tanger; Kröber 1924, 1928, Rif, Tanger

Conops djanetianus Séguy, 1938

Mouna 1998: 85

Conops elegans Meigen, 1824

= Conops semifumosus Adams, in Séguy 1934b: 162

= Conops ruficornis Becker, 1913, in Becker and Stein 1913: 89; Kröber 1924: 69

Becker and Stein 1913, Rif, Tanger; Kröber 1924, 1927, AP, Casablanca; Séguy 1934b, Rif, Tanger; Stuke 2016; El Hawagry et al. 2021

Conops nubeculipennis Bezzi, 1901

= Conops atrogonius Séguy, 1930, in Séguy 1930a: 134; Séguy 1953a: 85; Mouna 1998: 85

Séguy 1930a, AP, Rabat, Mogador; Séguy 1953a, MA, Dayat Ifrah; Mouna 1998; Stuke and Schmid-Egger 2015, AA, 1.5 km S of Tissint, 14 km NW of Icht; Stuke 2016; El Hawagry et al. 2021

Conops theryi Séguy, 1928

Séguy 1928d; Séguy 1930a, AP, Rabat, Casablanca; Mouna 1998

Conops tifedarius Séguy, 1928

Séguy 1928d; Séguy 1930a, AP, Rabat; Mouna 1998

Leopoldius Rondani, 1843

Leopoldius coronatus (Rondani, 1857)

= Brachyglossum coronatum Rondani, in Séguy 1930a: 132; Mouna 1998: 85

Séguy 1930a, MA, Aïn Leuh (1400–1500 m); Mouna 1998

Physocephalini

Physocephala Schiner, 1861

Physocephala chrysorrhoea (Meigen, 1824)

Séguy 1930a, AP, Sidi Bettache; Mouna 1998; El Hawagry et al. 2021

Physocephala laticincta (Brullé, 1832)

Séguy 1930a, MA, Aïn Leuh (1400–1500 m); Bei-Bienko and Steyskal 1989; Mouna 1998

Physocephala maculigera Kröber, 1915

Séguy 1941d, AA, Agadir; Mouna 1998

Physocephala nigra (De Geer, 1776)

Séguy 1930a, AP, Sidi Bettache

Physocephala pusilla (Meigen, 1804)

Séguy 1928d; Séguy 1930a, MA, Meknès, Ras el Ksar (1900 m), HA, Asni; Mouna 1998; Stuke and Schmid-Egger 2015, AA, Aoulouz, 2 km NW of Tissint; El Hawagry et al. 2021

Physocephala rufipes (Fabricius, 1781)

Mouna 1998; AP (Tagulet (Essaouira)) – MISR

Physocephala vittata (Fabricius, 1794)

Séguy 1928d; Séguy 1930a, AP, Rabat, Casablanca, MA, Meknès, Forêt Zaers; Mouna 1998; Stuke and Schmid-Egger 2015, AA, Aoersi (15 km NE of Agadir), AA, Oued near beach (19 km W of Tiznit); El Hawagry et al. 2021; HA – MISR

Pseudophysocephala Kröber, 1940

Pseudophysocephala bouvieri (Séguy, 1936)

= Conops bouvieri Séguy, in Séguy 1936a: 299

Séguy 1936a, MA, Meknès (550 m); Sidney 2001, MA, Meknès

Dalmanninae

Dalmannini

Dalmannia Robineau-Desvoidy, 1830

Dalmannia aculeata (Linnaeus, 1761)

Séguy 1928d; Séguy 1930a, MA, Aïn Leuh, Meknès; Mouna 1998 – MISR

Dalmannia dorsalis (Fabricius, 1794)

= Dalmannia flavescens (Meigen), in Becker and Stein 1913: 90

Becker and Stein 1913, Rif, Tanger; Stuke and Kehlmaier 2008, MA, Fès

Myopinae

Myopini

Melanosoma Robineau-Desvoidy, 1853

Melanosoma bicolor (Meigen, 1824)

Séguy 1941d, AA, Agadir (Admine forest); Mouna 1998

Melanosoma mundum Czerny & Strobl, 1909

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, HA, Tafingoult (Goundafa, 1500–1600 m), AA, Tenfeht (Souss, 1000–1500 m); Séguy 1953a, HA, Aït Ourir; Séguy 1949a, AA, Foum-el-Hassan; Mouna 1998; Stuke and Schmid-Egger 2015, AA, SE of Awir (10 km NNW of Agadir), Talmakant (80 km NE of Agadir), AA, 19 km W of Tiznit, Massa river (25 km NE of Tiznit), Imitek (30 km WSW of Tata), Issafen (55 km WNW of Tata); Grabener 2017

Myopa Camras, 1953

Myopa buccata (Linnaeus, 1758)

Mouna 1998; MA (Oulmès) – MISR

Myopa dorsalis Fabricius, 1794

Séguy 1930a, MA, Jebel Ahmar (1750 m); Mouna 1998; MA (Oulmès); El Hawagry et al. 2021 – MISR

Myopa hirsuta Stuke & Clements, 2008

Stuke and Clements 2008, MA, Azrou, Ifrane

Myopa nigrita Wiedemann, 1824

Wiedemann 1824, 1830; Kröber 1916, 1928

Myopa pellucida Robineau-Desvoidy, 1830

Stuke and Clements 2008, Rif, Chefchaouen, HA, Ourika

Myopa picta Panzer, 1798

Séguy 1930a, AP, Casablanca, MA, Meknès; Mouna 1998; El Hawagry et al. 2021

Myopa stigma Meigen, 1824

Becker and Stein 1913, Rif, Tanger; El Hawagry et al. 2021

Myopa testacea (Linnaeus, 1767)

Séguy 1930a, AP, Casablanca, Fouarat, HA, Arround (Skoutana: 2000–2400 m), Tachdirt, Jebel Likount; Mouna 1998; AP (Fouarat) – MISR

Myopa palliceps (Bigot, 1887)

= Myopa minor Strobl, in Clements 2000: 239

= Myopa vaulogeri Séguy, in Séguy 1930a: 136; Clements 2000: 236

Séguy 1930a, AP, Casablanca; Villeneuve 1933; Mouna 1998; Clements 2000, AP, Casablanca, HA, Marrakech, Ourigane (1000 m), AA, Ammelental (10 km NE of Tafraoute)

Thecophora Rondani, 1845

Thecophora atra (Fabricius, 1775)

= Occemyia atra Fabricius, in Séguy 1930a: 137; Becker and Stein 1913: 90

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, AP, Sidi Bettache, MA, Meknès, HA, Asni, AA, Taroudant (Souss); Mouna 1998; HA (Mouldikht (Marrakech)); El Hawagry et al. 2021 – MISR

Thecophora cinerascens (Meigen, 1804)

= Thecophora pusilla (Meigen, 1824), in Pârvu et al. 2006: 276; Popescu-Mirceni 2011: 35

Pârvu et al. 2006, MA, Ifrane; Popescu-Mirceni 2011, MA, Ifrane

Sicini

Sicus Scopoli, 1763

Sicus ferrugineus (Linnaeus, 1761)

Séguy 1930a, MA, Aïn Leuh; Zimina 1975; Mouna 1998; MA (Tizi-n’Ifrah, Guisser (1400 m)) – MISR

Zodioninae

Zodionini

Zodion Latreille, 1797

Zodion cinereum (Fabricius, 1794)

Séguy 1930a, MA, Meknès, HA, Around (Skoutana); Mouna 1998; Grabener 2017

Zodion erythrurum Rondani, 1865

Séguy 1930a, AP, Casablanca, HA, Dar Caïd M’Tougui; Mouna 1998; Stuke and Schmid-Egger 2015, AA, Tamzergoute (10 km N of Agadir); El Hawagry et al. 2021

Nerioidea

MICROPEZIDAE

K. Kettani, M.J. Ebejer

Number of species: 1. Expected: 2

Faunistic knowledge of the family in Morocco: poor

Micropeza Meigen, 1803

Micropeza kettaniae Ebejer, 2019

Ebejer 2019b, Rif, Oued Kbir (Béni Ratene, 157 m), Dayat Tazia (Tazia, 733 m) – MISR, NHMUK

Tanypezoidea

PSILIDAE

K. Kettani, M.J. Ebejer

Number of species: 1. Expected: 5

Faunistic knowledge of the family in Morocco: poor

Chamaepsila Hendel, 1917

Chamaepsila nigricornis (Meigen, 1826)

Ebejer et al. 2019, Rif, Tétouan, Onsar Lile (349 m), Aïn Tissemlal (Azilane, 1255 m), MA, 17 km SW of Midelt (1940 m), Lac Aguelmane Afennourir (30 km SW of Azrou, 2050 m)

Tephritoidea

LONCHAEIDAE

K. Kettani, I. MacGowan

Number of species: 5. Expected: 30

Faunistic knowledge of the family in Morocco: poor

Dasiopinae

Dasiops Rondani, 1856

Dasiops latifrons (Meigen, 1826)

Séguy 1934a; Mouna 1998; MacGowan and Freidberg 2008; AP (Rabat) – MISR; Rif (Tanger), AP (Rabat) – MHNP

Lonchaeinae

Lamprolonchaea Bezzi, 1920

Lamprolonchaea smaragdi (Walker, 1849)

= Lonchea aurea Macquart, 1851, in Séguy 1953a: 85

Becker and Stein 1913, Rif, Tanger; Séguy 1934a; Séguy 1953a, AP, Rabat, MA, Fès; Rungs 1952, HA, Arganier; Mouna 1998

Lonchaea Fallén, 1820

Lonchaea tarsata Fallén, 1820

MacGowan and Freidberg 2008

Lonchaea sp.

= Recorded as Lonchaea laticornis Meigen, 1826 but almost certainly not this species

Becker and Stein 1913, Rif, Tanger

Silba Macquart, 1851

Silba adipata McAlpine, 1956

= Lonchaea aristella Becker, 1903, in Séguy 1934b: 162

Séguy 1934b, AP, Rabat; MacGowan and Freidberg 2008; AP (Rabat) – MNHN

PALLOPTERIDAE

K. Kettani, M.J. Ebejer

Number of species: 1. Expected: 3

Faunistic knowledge of the family in Morocco: poor

Palloptera Fallén, 1820

Palloptera ustulata (Fallén, 1820)

Ebejer et al. 2019, MA, Zaouia d’Ifrane (Ifrane, 1603 m)

PIOPHILIDAE

K. Kettani, M.J. Ebejer

Number of species: 3. Expected: 6

Faunistic knowledge of the family in Morocco: poor

Piophilinae

Mycetaulus Loew, 1845

Mycetaulus hispanicus Duda, 1927

Mouna 1998; Carles-Tolrá 2002

Piophila Fallén, 1810

Piophila casei (Linnaeus, 1758)

Séguy 1930a, Rif, Tanger, AP, Rabat, MA, Meknès; Mouna 1998; Rif (Tanger), AP (Rabat), MA (Meknès) – MISR

Prochyliza Walker, 1849

Prochyliza nigrimana (Meigen 1826)

Ebejer et al. 2019, Rif, Aïn Tissemlal (Azilane, 1255 m)

PLATYSTOMATIDAE

K. Kettani, G.V. Popov

Number of species: 4. Expected: 4

Faunistic knowledge of the family in Morocco: moderate

Platystomatinae

Platystoma Meigen, 1803

Platystoma idia Séguy, 193442

Séguy 1934a, AP, Aïn Sferguila (Forêt Zaers); Hennig 1945, AP, Aïn Sferguila (Forêt Zaers); Soós 1984a

Platystoma meridionale Hendel, 1913

= Platystoma seminationis (Fabricius), in Becker 1907: 385

Becker 1907; Hendel 1913, AP, Mogador; Hennig 1945, AP, Mogador; Soós 1984a, AP, Mogador

Rivellia Robineau-Desvoidy, 1830

Rivellia hispanica Lyneborg, 1969

Ebejer et al. 2019, EM, Tafoughalt

Rivellia syngenesiae (Fabricius, 1781)

Becker and Stein 1913, Rif, Tanger; Mouna 1998; MA (Oulmès, Meknès) – MISR

TEPHRITIDAE

K. Kettani, A.L. Norrbom

Number of species: 69. Expected: 75

Faunistic knowledge of the family in Morocco: moderate

Dacinae

Ceratitidini

Capparimyia Bezzi, 1920

Capparimyia savastanii (Martelli, 1911)

= Capparimyia savastanii (Martelli), in Séguy 1953a: 85

Séguy 1953a, AA, Tiznit (on Capparis spinosa); El Harym and Belqat 2017

Ceratitis MacLeay, 1829

Ceratitis capitata (Wiedemann, 1824)

Becker and Stein 1913, Rif, Tanger; Vayssière 1920, AP, Rabat; Séguy 1930a (common in all of Morocco); Rungs 1952, HA, Arganier; Harris et al. 1980; Mouna 1998; De Meyer 2000, AP, Ouadj-Ouli-Mohamed, env. Settat, Insgane; Vidal et al. 2008; Aboussaid et al. 2009; Koçak and Kemal 2010; El Harym and Belqat 2017, Rif, Kitane, El Haouta, MA, Sensla, AA, Environs Massa, Oued Massa, Douar Sidi Abou, Douar Tighrimt, Douar Zaouia; Elaini and Mazih 2018; Elaini et al. 2019, HA, Arganeraie; AP (Safi) – MHNNR

Dacini

Bactrocera Macquart, 1835

Bactrocera oleae (Rossi, 1790)

= Dacus oleae Rossi, in Becker and Stein 1913: 94, Vayssière 1920: 256

= Dacus oleae Gmelin, in Séguy 1930a: 168

Becker and Stein 1913, Rif, Tanger; Vayssière 1920, EM, Oujda; Séguy 1930a; Mouna 1998; Savio 2011, EM, Oujda; El Harym and Belqat 2017, Rif, El Haouta, Oued Maâza, Cascade Chrafate, Koudiat El Aouinate, Lâazaba, Dhar Sbagh Mâasra, El Hajria, MA, Sensla, AA, route Bab El Khemis

Dacus Fabricius, 1805

Dacus frontalis (Becker, 1922)

El Harym and Belqat 2017, AA, Oued Foum Ziguid (Douar Ouaiftoute), Oued Draa (Ikhf Mezrou), Isdaoun

Dacus longistylus (Wiedemann, 1830)

El Harym and Belqat 2017, AA, Oued Tata, Douar Tighrimt, Oued Foum Ziguid (Douar Ouaiftoute)

Tephritinae

Dithrycini

Oedaspis Loew, 1862

Oedaspis daphnea Séguy, 1930

Séguy 1930a, AP, El Mers (Rabat); Foote 1984; Soós 1984b; Mouna 1998; Norrbom et al. 1999; El Harym and Belqat 2017

Oedaspis multifasciata (Loew, 1850)

= Oedaspis multifasciatus (Loew), in Séguy 1953a: 85

Séguy 1953a, EM, Itzer (Haute Moulouya); El Harym and Belqat 2017

Oedaspis trotteriana Bezzi, 1913

Soós 1984b; Ribera and Blasco-Zumeta 1998; Norrbom et al. 1999; El Harym and Belqat 2017

Myopitini

Myopites Blot, 1827

Myopites cypriacus Hering 1938

El Harym et al. 2020, Rif, Douar Halila, Dam Nakhla, Marabout Sidi Bou Hadjel

Myopites inulaedyssentericae Blot, 182743

= Myopites apicatus Freidberg, 1980, in El Harym and Belqat 2017: 140

El Harym and Belqat 2017, Rif, affluent Tarmast, Aïn Afersiw

Myopites longirostris (Loew, 1846)

El Harym et al. 2020, Rif, Oued Tahaddart, Douar Kouf, Mkhinak

Myopites stylatus Fabricius, 1794

El Harym and Belqat 2017, Rif, affluent Tarmast, Oued El Hamma, El Haouta

Myopites variofasciatus Becker, 1903

= Myopites variofasciata Becker, in Séguy 1941a: 32

Séguy 1941a, HA, Imi-n’Ouaka (1500 m), Mouna 1998

Urophora Robineau-Desvoidy, 1830

Urophora congrua Loew, 186244

= Euribia congrua Loew, in Séguy 1941d: 14

Séguy 1941d, AA, Taroudant; Mouna 1998; El Harym and Belqat 2017

Urophora jaculata Rondani, 1870

= Urophora mauritanica Macquart, in El Harym and Belqat 2017: 149 (misidentification)

Urophora mauritanica Macquart, 1851

= Euribia algira Macquart, in Séguy 1930a: 169

= Urophora algira Macquart, in Séguy 1934a: 98, Mouna 1998: 87

= Urophora macrura Loew, in Séguy 1934a: 100, Mouna 1998: 87

Séguy 1930a, HA, Imi-N’Takandout, Dar Kaid M’Tougui; Séguy 1934a; White and Korneyev 1989, HA, Ito; Mouna 1998; Norrbom et al. 1999; Koçak and Kemal 2010; Koçak and Kemal 2013b

Urophora quadrifasciata algerica (Hering, 1941)

= Euribia quadrifasciata Meigen, in Séguy 1930a: 169

Séguy 1930a (common in all North Africa); Mouna 1998; El Harym and Belqat 2017

El Harym et al. 2020, MA, Douar Oulad Abdoune, Mlakite, Tirra, Douar Oulad Amar, Tihli, Douar Oulad Amar

Urophora solstitialis (Linnaeus, 1758)

= Euribia solstitialis Linnaeus, in Séguy 1930a: 169

Séguy 1930a, HA, Haute Réghaya; Mouna 1998; El Harym and Belqat 2017

Noeetini

Ensina Robineau-Desvoidy, 1830

Ensina sonchi (Linnaeus, 1767)

El Harym and Belqat 2017, Rif, Ksar Rimal, Douar Tizga, Oued Kbir, MA, Sensla, AA, Oued Massa (Pont Aghbalou), Centre Sidi Ouassay, Aïn Boharroch, Atbane, Oued Tamanarne, Oued Draa (Tahtah), Jnane Makadir, Kasbah Asma, Ait Aissa O Brahim, Oued Ziz (Pont Errachidia), Oued Ouarzazate; AA (Tafraout (Al Ourir, 12 km E)) – MHNNR

Hypenidium Loew, 1862

Hypenidium graecum Loew, 1862

= Stephanaciura bipartita Séguy, in Séguy 1930a: 171

Séguy 1930a, MA, Tiffert (2000–2200 m); Villeneuve 1933; Soós 1984b; Mouna 1998: 87; Norrbom et al. 1999; El Harym and Belqat 2017

Tephrellini

Aciura Robineau-Desvoidy

Aciura coryli (Rossi, 1790)

= Aciura Powelli Séguy, in Séguy 1930a: 170, Séguy 1953a: 85, Mouna 1998: 87

Séguy 1930a, MA, Azrou (larvae from Phlomis crinita Cav.); Séguy 1953a, AP, Korifla; Mouna 1998; Norrbom et al. 1999, MA, Azrou; El Harym and Belqat 2017; El Harym et al. 2020, Rif, Arhil

Oxyaciura Hendel, 1927

Oxyaciura tibialis (Robineau-Desvoidy, 1830)

= Aciura tibialis Robineau-Desvoidy, in Becker and Stein 1913: 94

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA, Sker; Séguy 1934a; Mouna 1998; Koçak and Kemal 2010; Koçak and Kemal 2013b; El Harym and Belqat 2017, Rif, Dayat El Birdiyel, Oued Azila, maison forestière; Oued Maâza (Tarik El Ouasâa); AP (Tamri, 10 km S) – MHNNR

Sphaeniscus Becker, 1908

Sphaeniscus filiolus (Loew, 1869)

= Spheniscomyia filiola Loew, in Séguy 1930a: 170; Mouna 1998: 87

= Spheniscomyia aegyptiaca Efflatoun, in Séguy 1949a: 157; Mouna 1998: 87

Séguy 1930a; Séguy 1949a, SA, Guelmim; Mouna 1998; El Harym and Belqat 2017, Rif, affluent Tarmast, Oued Maâza (Tarik El Ouasâa)

Tephritini

Acanthiophilus Becker, 1908

Acanthiophilus helianthi (Rossi, 1790)

= Tephritis eluta Meigen, in Becker and Stein 1913: 94

= Orellia eluta Meigen, in Mouna 1998: 87

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, Rif, Tanger (Sarf, route Spartel), MA, Tizi s’Tkrine (Jebel Ahmar, 1700 m); Séguy 1949a, AA, Alnif, Foum-el-Hassan, SA, Guelmim; Mouna 1998; Koçak and Kemal 2013b; El Harym and Belqat 2017, Rif, Oued Zinat, Ksar Rimal, Oued Dardara, affluent Tarmast, Dayat El Birdiyel, Oued Azila, Dayat Jebel Zemzem, Oued Boumarouil, Douar Abou Boubnar (Mara­bout Sidi Gile), El Haouta, Oued Maâza (Tarik El Ouasâa), Douar Tizga, Dayat Afrate, Oued Mezine, Aïn El Malaâb, Oued Jnane Niche, MA, Oued Oum-er-Rbia, AA, Centre Sidi Ouassay, Avant Sidi Bin­zarne, route Bab El Khemis, airport Sidi Ifni, Oued Tisla, Oued Sayad, Oued Tamanarne, Oued Foum Ziguid (Douar Ouaiftoute), Jnane Makadir, Douar Rggaga, Oued Tinghir, Oued Ouarzazate; AP (Essaouira (Cap Hadid), Tamri, 10 km S) – MHNNR

Campiglossa Rondani, 1870

Campiglossa martii (Becker, 1908)

El Harym and Belqat 2017, Rif, Oued Kbir, AA, Centre Sidi Ouassay

Campiglossa producta (Loew, 1844)

= Oxyna tessellata Loew, in Becker and Stein 1913: 94 (misidentification)

= Paroxyna tessellata (Loew), in Séguy 1930a: 174; Mouna 1998: 87; Koçak and Kemal 2013b: 38 (misidentification)

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, Rif, Tanger, AP, Mogador, HA, Telouet Glaoua; Séguy 1934a; Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Mouna 1998; Koçak and Kemal 2013b, HA; El Harym and Belqat 2017, Rif, Oued Al Mizzine, Aïn El Malaâb, Dayat El Hajjami

Campiglossa sororcula (Wiedemann, 1830)

= Dioxyna sororcula (Wiedemann), in El Harym and Belqat 2017: 155

El Harym and Belqat 2017, Rif, Ksar Rimal, Oued Jnane Niche, Oued Halila, Oued Zarka, Oued Martil, Oued Amsa, Oued Sahel, Dayat Jebel Zemzem, Oued Maggou, Dhar Sbagh Mâasra, Douar Kitane

Capitites Foote & Freidberg, 1981

Capitites augur (Frauenfeld, 1857)

= Trypanea augur (Frauenfeld), in Séguy 1930a: 176; Mouna 1998: 87

Séguy 1930a, MA, Forêt Azrou, Tizi-s’Tkrine (Jebel Ahmar, 1700 m), HA, Tenfecht, AA, Souss; Mouna 1998; Pârvu et al. 2006, AA, Ouarzazate, Lac Edehby; Popescu-Mirceni 2011, MA, Tizi-s’Tkrine, Azrou, HA, Tenfecht, AA, Ouarzazate

Capitites ramulosa (Loew, 1844)

= Acanthiophilus ramulosus Loew, in Séguy 1930a: 177; Séguy 1941d: 15; Séguy 1949a: 157; Mouna 1998: 87

Séguy 1930a, MA, Timelilt, HA, Tizi-n’Test (Jebel Imdress, 2000–2450 m); Séguy 1941d, AA, Taroudant; Séguy 1949a, AA, Foum-el-Hassan, Akka, Agdz, Alnif; Mouna 1998; El Harym and Belqat 2017; El Harym et al. 2020, Rif, Amsa, Koudiat Taifour

Desmella Munro, 1957

Desmella rostellata (Séguy, 1941)

= Paroxyna rostellata Séguy, in Séguy 1941d: 14; Mouna 1998: 87

Séguy 1941d, AP, Agadir; Soós 1984b; Mouna 1998; Norrbom et al. 1999, AP, Agadir; El Harym and Belqat 2017

Euaresta Loew, 1873

Euaresta bullans (Wiedemann, 1830)

Herman and Dirlbek 2006, AA, Tiznit environs, Sidi Moussa d’Aglou; El Harym and Belqat 2017, AA, Msidira

Goniurellia Hendel, 1927

Goniurellia longicauda Freidberg, 1980

Freidberg 1980, MA, Tizi-s’Tkrine (1700 m), Azrou, AA, Taroudant; Soós 1984b; Freidberg and Kugler 1989; Norrbom et al. 1999; El Harym and Belqat 2017, AA, airport Sidi Ifni, Oued Tisla, Oued Tamanarne, Douar Zaouiet, Oued Tata, Douar Tighrimt, Ksibat Elhdeb, Oued Ziz (Pont Errachidia), Oued Ouarzazate

Goniurellia persignata Freidberg, 1980

Freidberg 1980, EM, Defilia, near Figuig; Soós 1984b; Freidberg and Kugler 1989; Norrbom et al. 1999; Herman and Dirlbek 2006, AA, Tiffoultoute (1146 m); El Harym and Belqat 2017, Rif, Dhar Sbagh Mâasra, AA, Douar Zaouiet, Oued Ouarzazate

Spathulina Rondani, 1856

Spathulina sicula Rondani, 1856

= Spathulina tristis Loew, in Séguy 1930a: 174; Mouna 1998: 87

Séguy 1930a; Mouna 1998; El Harym and Belqat 2017, Rif, Barrage Smir

Sphenella Robineau-Desvoidy, 1830

Sphenella marginata (Fallén, 1814)

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, Rif, Oued Judios (Tanger); Mouna 1998; Cassar et al. 2005, Rif, lagoon Smir; Koçak and Kemal 2013b, HA; El Harym and Belqat 2017, Rif, affluent Tarmast, Dayat Jebel Zemzem, El Malaâb, Oued Maâza (Âachira), Dayat Aïn Jdioui, Oued Maggou, Dayat Afrate

Tephritis Latreille, 1804

Tephritis carmen Hering, 1937

El Harym et al. 2020, Rif, forest house of National Park of Talassemtane

Tephritis dioscurea (Loew, 1856)

Séguy 1930a, MA, El Hajeb; Mouna 1998; El Harym and Belqat 2017

Tephritis divisa (Rondani, 1871)

El Harym and Belqat 2017, Rif, Dayat Amsemlil

Tephritis formosa (Loew, 1844)

Séguy 1930a, HA, Asni; Mouna 1998; El Harym and Belqat 2017, Rif, Oued Abou Bnar, Oued Sidi Ben Saâda, Oued Achekrade, Oued El Kanar

Tephritis leontodontis (De Geer, 1776)

Séguy 1930a (All North Africa); Mouna 1998; El Harym and Belqat 2017

Tephritis matricariae (Loew, 1844)

Séguy 1930a (all North Africa); Mouna 1998; El Harym and Belqat 2017, Rif, affluent Oued Amsemlil, El Haouta, Dayat Jebel Zemzem, Dayat Amsemlil, Oued El Hamma

Tephritis nigricauda (Loew, 1856)

Séguy 1930a, AP, Berrechid; Séguy 1934a; Séguy 1941d, AP, Agadir, Berrechid; Mouna 1998; El Harym and Belqat 2017, Rif, Dayat Jebel Zemzem, Oued Maâza (Tarik El Ouasâa), Aïn El Maounzil, Dayat Tazia, Dayat Amsemlil, Douar Tamakoute

Tephritis postica (Loew, 1844)

Herman and Dirlbek 2006, MA, Volubilis (358 m); El Harym and Belqat 2017, AA, Ksibat Elhdeb, Oued Tinghir

Tephritis praecox (Loew, 1844)

Séguy 1930a, Rif, Oued Judios (Tanger), MA, Tizi-s’Tkrine (Jebel Ahmar, 1700 m); Mouna 1998; Herman and Dirlbek 2006, MA, Ifrane, Azrou National Park (1743 m); El Harym and Belqat 2017, Rif, Dayat El Ânassar, Dayat Amsemlil, affluent Oued Amsemlil, Douar Dacheryène, Douar Taghbaloute, Barrage Nakhla, Oued Sa­hel, Daya Jebel Zemzem, Douar Kitane, Oued El Hamma, Oued Kbir, Aïn el Ma Bared, Aïn El Malaâb, Douar Abou Boubnar (Marabout Sidi Gile), maison forestière, Douar Tizga, Oued Aïn Jdioui (Touaret), Dayat Afrate, Oued Jbara, Aïn El Maounzil, Dayat Tazia, Oued Jnane Niche, Oued Maggou, Aïn Tiouila, Dayat Lemtahane, Lâazaba, Dhar Sbagh Mâasra, El Hajria, Aïn Boharroch, Douar Tamakout, Douar Ouslaf, EM, Oued Béni Ouaklane (Béni Snassen); AP (Essaouira) – MHNNR

Tephritis pulchra (Loew, 1844)

Séguy 1930a; Mouna 1998; El Harym and Belqat 2017

Tephritis simplex (Loew, 1844)

Séguy 1930a; Mouna 1998; El Harym and Belqat 2017; El Harym et al. 2020, Rif, Aïn Soualah, Aïn El Maounzil

Tephritis stictica Loew, 1862

Séguy 1930a, AP, Rabat; Mouna 1998; El Harym and Belqat 2017

Tephritis theryi Séguy, 1930

Séguy 1930a, HA, Marrakech, Asni; Soós 1984b; Mouna 1998; Norrbom et al. 1999; El Harym and Belqat 2017

Tephritis vespertina (Loew, 1844)

El Harym and Belqat 2017, Rif, Dayat Lemtahane, Dhar Sbagh Mâasra

Tephritomyia Hendel, 1927

Tephritomyia lauta (Loew, 1869)

= Acanthiophilus lauta Loew, in Séguy 1930a: 177; Mouna 1998: 87

Séguy 1930a, HA, Tachdirt (Imminen, 2400–2600 m); Freidberg and Kugler 1989; Mouna 1998; Yaran and Kütük 2012; Morgulis et al. 2015, HA, Tizi-n’Tichka; El Harym and Belqat 2017, Rif, Dayat El Birdiyel, Dayat Amsemlil, Lâazaba, AA, Msidira, Oued Ouarzazate

Trupanea Schrank, 1795

Trupanea amoena (Frauenfeld, 1857)

= Trypanea amoena Frauenfeld, in Séguy 1930a: 176; Mouna 1998: 87

Séguy 1930a, MA, Aïn Leuh; Mouna 1998; El Harym and Belqat 2017, Rif, Ksar Rimal, Oued Jnane niche, affluent Tarmast, Oued Martil (Tamouda), Oued Amsa, Oued El Hamma, Oued Boumarouil, Oued Sidi Yahia Aârab, Aïn Tiouila, AA, Oued Massa (Pont Aghbalou), Centre Sidi Ouassay, Avant Sidi Binzarne, Oued Tisla, Douar Tighrimt, Oued Draa (Tahtah), Jnane Makadir, Douar Rggaga, Aït Aissa O Brahim, Oued Draa (Ikhf Mezrou), Isdaoun, Ksibat Elhdeb, Oued Tinghir

Trupanea guimari (Becker, 1908)

El Harym and Belqat 2017, AA, Centre Sidi Ouassay, Msidi­ra, Jnane Makadir, Aït Aissa O Brahim, Ksibat Elhdeb; Norrbom 2004: 06600136 – INHS (AA, 5 km W Ouarzazate)

Trupanea stellata (Fuesslin, 1775)

Séguy 1930a, MA, Timelilt (1900 m); Séguy 1949a, SA, Guelmim; Mouna 1998; El Harym and Belqat 2017, Rif, Mizoghar, Oued Maâza (Tarik El Ouasâa), Dayat Afrate, Aïn El Malâab, Oued Tkarae, AA, Centre Sidi Ouassay; AA (Taliouine) – MHNNR

Terellini

Chaetorellia Hendel, 1927

Chaetorellia conjuncta (Becker, 1912)

El Harym and Belqat 2017, AA, Airport Sidi Ifni, Oued Assa, Oued Sayad, Oued Foum Ziguid (Douar Ouaiftoute), Ksibat Elhdeb, Oued Ziz (Pont Errachidia), Oued Ouarzazate

Chaetorellia hestia Hering, 1937

= Chaetorellia hexachaeta Loew, in Séguy 1930a: 174 [probably a misidentification]

= Orellia hexachaeta Loew, in Séguy 1934a: 135 [misidentification, see White and Macquart 1989: 476]; Mouna 1998: 87

Séguy 1930a, AP, Mogador; Séguy 1934a; Mouna 1998; El Harym and Belqat 2017, AA, Centre Sidi Ouassay

Chaetorellia succinea (Costa, 1844)

El Harym et al. 2020, MA, Douar Oulad Abdoune

Chaetostomella Hendel, 1927

Chaetostomella cylindrica (Robineau-Desvoidy, 1830)

El Harym et al. 2020, Rif, Marabout Douar Halila, Mkhinak, Douar Kitane

Terellia Robineau-Desvoidy, 1830

Terellia colon (Meigen, 1826)

= Orellia colon (Meigen), in Séguy 1930a: 173

Séguy 1930a (common in all North Africa); Mouna 1998; El Harym and Belqat 2017

Terellia fuscicornis (Loew, 1844)

Séguy 1930a (common in all North Africa); Mouna 1998; El Harym and Belqat 2017

Terellia longicauda (Meigen, 1838)

Séguy 1930a, MA, Aïn Leuh (1200–1400 m), HA, Tizi-n’Test, Goundafa (Jebel Imdress, 2000–2450 m); Séguy1934a; Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Mouna 1998; El Harym and Belqat 2017

Terellia luteola (Wiedemann, 1830)

El Harym et al. 2020, Rif, Bakrim, Aforidane, Aïn Siyed

Terellia oasis (Hering, 1938)

El Harym et al. 2020, Rif, Douar Halila

Terellia ptilostemi El Harym et al. 2021

El Harym et al. 2021, Rif, Douar Chourdane (908 m), Aïn Akorian (1610 m), Aïn Elma Sefli (1345 m), Forest house of the Talassemtane National Park (1674 m)

Terellia serratulae (Linnaeus, 1758)

= Tephritis pallens Wiedemann, in Wiedemann 1824: 54

= Trypeta serratula Linnaeus, in Becker and Stein 1913: 94

= Terellia serratulae Linnaeus, in Séguy 1930a: 173

Wiedemann 1824, Rif, Tanger; Becker and Stein 1913, Rif, Tanger; Séguy 1930a (all North Africa); Mouna 1998; Norrbom et al. 1999; El Harym and Belqat 2017, Rif, Dayat Jebel Zemzem, Oued Maâza (Tarik El Ouasâa)

Terellia virens (Loew, 1846)

Séguy 1930a; White 1989, HA, Jebel Ayachi; Mouna 1998; Korneyev et al. 2013, HA, Tizi-n’Talrhemt; El Harym and Belqat 2017, AA, airport Sidi Ifni, Oued Ouarzazate

Trypetinae

Carpomyini

Carpomya Costa, 1854

Carpomya incompleta (Becker, 1903)

El Harym and Belqat 2017, AA, Douar Zaouia

Euleia Walker, 1835

Euleia heraclei (Linnaeus, 1758)

= Acidia heraclei Linnaeus, in Séguy 1953a: 85

Séguy 1953a, MA, Sidi Slimane; Freidberg and Kugler 1989; Koçak and Kemal 2010; El Harym and Belqat 2017, Rif, Oued Boumarouil, Aïn El Âakba Larbaâ

Euleia marmorea (Fabricius, 1805)45

= Philophylla flavescens Fabricius, in Séguy 1930a: 170; Mouna 1998: 87

= Euleia flavescens Fabricius, in Soós 1984b: 88

Séguy 1930a, Rif, Tanger; Zimsen 1964; Soós 1984b; Mouna 1998; Norrbom et al. 1999, Rif, Tanger; El Harym and Belqat 2017

Trypetini

Chetostoma Rondani, 1856

Chetostoma curvinerve Rondani, 1856

El Harym and Belqat 2017, Rif, Oued Kelaâ, Bab el Karn

Acknowledgments

We gratefully acknowledge the assistance and cooperation of Valery Korneyev and the late Amnon Freidberg who contributed to the revision of this family.

ULIDIIDAE

K. Kettani, M.J. Ebejer

Number of species: 13. Expected: 18

Faunistic knowledge of the family in Morocco: good

Otitinae

Ceroxys Macquart, 1835

Ceroxys urticae Linnaeus, 1758

Ebejer et al. 2019, AP, Lower Loukous (6 m), Larache (5 m)

Dorycera Meigen, 1830

Dorycera griseipennis (Becker, 1907)

Soós 1984b

Herina Robineau-Desvoidy, 1830

Herina ghilianii Rondani, 1869

Kameneva 2007, HA, Ansegmir-Tal, W Midelt (1400 m); Ebejer 2015

Herina lacustris (Meigen, 1826)

Kameneva 2007, Rif, Baie de Tanger

Herina oscillans (Meigen, 1826)

= Herina schlueteri Becker, in Becker and Stein 1913: 92; Soós 1984b: 56

Becker and Stein 1913, Rif, Tanger; Soós 1984b; Kameneva 2007, Rif, Tanger

Melieria Robineau-Desvoidy, 1830

Melieria nigritarsis Becker, 1903

Ebejer et al. 2019, AA, Merzouga (714 m)

Otites Latreille, 1804

Otites tangeriana Becker, 1913

= Otites tangeriana Becker, in Becker and Stein 1913, 1918: 92

Becker and Stein 1913, 1918, Rif, Tanger; Soós 1984b, Rif, Tanger

Tetanops Fallén, 1820

Tetanops flavescens Macquart, 1835

Séguy 1930a, Rif, Tanger; Mouna 1998

Ulidiinae

Physiphora Fallén, 1810

Physiphora alceae (Preyssler, 1791)

= Chrysomyza demandata (Fabricius, 1798), in Séguy 1953a: 85; Mouna 1998: 87

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, HA, Goundafa; Séguy 1953a, SA, El Aöun du Draa; Séguy 1949a, AA, Agdz; Mouna 1998; Koçak and Kemal 2010; Kameneva and Korneyev 2016, AA, Tizi-n’Bachkoun (1600 m); Rif (M’Diq farm) – MISR; Rif (Tanger) – MfN; AP (Casablanca) – ZSSM

Physiphora smaragdina (Loew, 1852)

Kameneva and Korneyev 2016, AA, 25 km S Goulmima (100 m) – NHMD

Ulidia Meigen, 1826

Ulidia apicalis (Meigen, 1826)

Séguy 1930a, MA, Meknès, HA, Skoutana; Séguy 1934a; Lyneborg 1969; Soós 1984b; Mouna 1998

Ulidia erythrophthalma Meigen, 1826

Becker and Stein 1913, Rif, Tanger; Séguy 1949a, AA, Foum-el-Hassan; Mouna 1998

Ulidia megacephala Loew, 1845

Soós 1984b; Zaitzev 1984; Koçak and Kemal 2010

Lauxanioidea

CHAMAEMYIIDAE

K. Kettani, M.J. Ebejer

Number of species: 18. Expected: 24

Faunistic knowledge of the family in Morocco: moderate

Chamaemyiinae

Chamaemyia Meigen, 1803

Chamaemyia aridella (Fallén, 1823)

Ebejer 2016a, Rif, Moulay Abdelsalam (1180 m), Issaguen (1620 m); HA (Jebel Ayachi, Mikdane, maison forestière, MR Jaffar, Tizi-n’Zou) – NHMUK

Chamaemyia flavicornis (Strobl, 1902)

Ebejer 2016a, Rif, Martil beach and dunes (on human faeces), MA, Khénifra (17 km NW of Zaida, 1878 m), AA, Errachidia (29 km N of Rich, 1570 m); HA (Asni near Alrene, W Imlil) – NHMUK

Chamaemyia flavipalpis (Haliday, 1838)

= Chamaemyia maritima Zetterstedt, in Mouna 1998: 85

Mouna 1998; Ebejer 2016a, Rif, Ksar Sghir, Oued Araml, Ksar Sghir, AP, Larache, Merja Zerga, Loukous marsh

Chamaemyia herbarum (Robineau-Desvoidy, 1830)

Ebejer 2016a, MA, Khénifra, 17 km SW of Midelt (1940 m), AA, Errachidia, 29 km N of Rich (1570 m); HA (Jebel Ayachi, Mikdane, MR Jaffar, Tizi-n’Zou) – NHMUK

Chamaemyia juncorum (Fallén, 1958)

Ebejer 2016a, MA, Khénifra (17 km SW of Midelt, 1940 m), AA, Errachidia (29 km N of Rich, 1570 m)

Chamaemyia polystigma (Meigen, 1830)

Becker and Stein 1913, Rif, Tanger; Ebejer 2016a, Rif, Moulay Abdelsalam (1180 m), Sidi Yahia Aarab (377 m); HA (Asni) – NHMUK

Melanochthiphila Frey, 1958

Melanochthiphila sp. Ebejer, 2016

Ebejer 2016a, AA, Errachidia (30 km W of Errachidia, 1065 m)

Parochthiphila (Euestelia) Enderlein, 1927

Parochthiphila (Euestelia) coronata (Loew, 1858)

Carles-Tolrá 1993; Mouna 1998; Ebejer 2016a, AP, Larache (Lower Loukous, 2 m); HA (Jebel Ayachi, Mikdane, MR Jaffar) – NHMUK

Parochthiphila frontella (Rondani, 1874)

Ebejer 2016a; HA (Jebel Ayachi, Mikdane, Tizi-n’Zou, MR Jaffar) – NHMUK

Parochthiphila inconstans Becker, 1903

Ebejer 2016a, AA, Ziz river (9.5 km SE of Rich, 1285 m), Errachidia (6 km N of Errachidia, 1010 m)

Parochthiphila (Euestelia) nigripes Strobl, 1900

Ebejer 2016a, Rif, Ksar Sghir, Dardara

Leucopinae

Leucopis Meigen, 1803

Leucopis annulipes Zetterstedt, 1848

Mouna 1998; AP (Maâmora) – MISR

Leucopis griseola (Fallén, 1823)

Mouna 1998; AP (Rabat), MA (Meknès), HA (Marrakech) – MISR

Leucopis formosana Hennig, 1938

Ebejer 2016a, AP, Sidi Smail, Oued Tensift (estuary)

Leucopis glyphinivora Tanasijtshuk, 1958

Ebejer 2016a, AP, Ksar Sghir, Azemmour, Oued Tensift (estuary), AA, Errachidia (30 km W of Errachidia)

Leucopis kerzhneri Tanasijtshuk, 1970

Ebejer 2016a; HA (Jebel Ayachi) – NHMUK

Leucopis palumbii Rondani, 1872

Ebejer 2016a, Rif, Ksar Sghir, Oued Aliane

Lipoleucopis de Meijere, 1928

Lipoleucopis pulchra Raspi, 2008

Ebejer 2016a, AA, Errachidia, Ziz river (12 km S of Rissani)

LAUXANIIDAE

K. Kettani, M.J. Ebejer

Number of species: 27. Expected: 45

Faunistic knowledge of the family in Morocco: good

Homoneurinae

Homoneura Wulp, 1891

Homoneura ericpoli Carles-Tolrá, 1993

Ebejer and Kettani 2019a, Rif, Azilane (1255 m), Adrou (556 m), HA, Lac Tislit (Imlchil, 2254 m)

Homoneura licina Séguy, 1941

Séguy 1941d, AA, Agadir; Mouna 1998; Schacht et al. 2004; Ebejer and Kettani 2019a, Rif, Oued Laou (2 m), AP, Merja Zerga, Sidi Smaine, Safi (estuary of Oued Tensift), Azemmour (El Jadida), Diabat (Essaouira), Larache (5 m)

Homoneura transversa (Wiedemann, 1830)

Wiedemann 1830; Ebejer and Kettani 2019a

Prosopomyia Loew, 1856

Prosopomyia pallida Loew, 1856

Vaillant 1956b, HA, Oukaimeden, Imi-N’Ifri; Carles-Tolrá 1993; Mouna 1998; Ebejer and Kettani 2019a, Rif, Adrou (556 m), Issaguen (1547 m), HA, Jebel Ayachi – NHMUK

Lauxaniinae

Calliopum Strand, 1928

Calliopum oosterbroeki Shatalkin, 2000

Ebejer and Kettani 2019a, Rif, Oued Zarka (Yarghite, 137 m), Moulay Abdelsalam (1180 m), Ketama, Rahbat Amlay (284 m), Ikadjiouen (294 m), Mechkralla (NPT, 981 m), Kharouba (roadside meadow between Chefchaouen and Tétouan, 385 m), Dbani (Béni Selmane, 1046 m), Aïn Ben Ali (Béni Selmane, 1014 m), HA, Lalla Takrkoust (Oued N'fis, 1141 m) – NHMUK

Calliopum tuberculosum (Becker, 1895)

Ebejer and Kettani 2019a, Rif, Oued Zarka (Yarghite, 135 m), Azilane (1255 m), Perdicaris Park (Tanger, 223 m), cascade Chrafate (859 m)

Meiosimyza Hendel, 1925

Meiosimyza (Lyciella) rorida (Fallén, 1820)

= Homoneura rorida Fallén, in Mouna 1998: 85

Mouna 1998; MA (Aïn Leuh) – MISR

Minettia Robineau-Desvoidy, 1830

Minettia aenigmatica Ebejer, 2019

Ebejer 2019a, Rif, Perdicaris Park (223 m), Oued Khmis (Khmis Anjra, 61 m), HA, Mikdane (Jebel Ayachi); Ebejer and Kettani 2019a

Minettia biseriata (Loew, 1847)

Ebejer and Kettani 2019a, Rif, Oued Khmis (Khmis Anjra, 61 m), Jnane Niche (46 m), Oued Kbir (Béni Ratene (PNPB), 157 m)

Minettia cantolraensis Carles-Tolrá, 1998

Ebejer and Kettani 2019a, Rif, Oued Laou, El-Fahsa, Oued Guallet (Bni Boufrah, 942 m), Oued Ouringa (2 m), Maggou (NPT, 962 m), Ksar el-Kbir road to Chefchaouen at bridge near oued Azla (80 m), Oued Siflaou (281 m), Dardara (484 m), Azilane (1255 m), Adrou (556 m), Perdicaris Park (Tanger, 223 m), Bni Bahlou (986 m), El Hamma (936 m), Amaghouse (Oued Laou), Oued Koub (Laghdir, 148 m), EM, Aïn Sfa (Berkane, 638 m), Tafoughalt (Berkane, 788 m), HA, Lalla Takrkoust (Oued N'fis, 1141 m), AA, Taliouine (Taroudant, 1049 m)

Minettia fasciata (Fallén, 1826)

Merz 2004, AP, Rabat; Ebejer and Kettani 2019a, Rif, Oued Khmis (Khmis Anjra, 61 m), Ksar el-Kbir road to Chefchaouen at bridge near oued Azla (80 m), Azilane (1255 m), Adrou (556 m), Issaguen (1547 m), Oued Taida (Al Andalous, 503 m), Perdicaris Park (Tanger, 223 m), El Hamma (936 m), EM, Aïn Sfa (Berkane, 683 m)

Minettia flavipalpis (Loew, 1847)46

= Sapromyza flavipalpis Loew, 1847, in Becker and Stein 1913: 93 (Rif, Tanger), Ebejer and Kettani 2019a: 144 (???)

Minettia flaviventris (Costa, 1844)

Ebejer and Kettani 2019a, Rif, Azilane (1255 m)

Minettia longiseta (Loew, 1847)

Ebejer and Kettani 2019a, Rif, Maggou (NPT, 962 m), Adrou (PNPB, 556 m)

Minettia plumicornis (Fallén, 1820)46

Schacht et al. 2004; Ebejer and Kettani 2019a

Minettia subvittata (Loew, 1847)

Ebejer and Kettani 2019a, Rif, Oued Khmis (Khmis Anjra, 61 m), Azilane (1255 m)

Minettia suillorum (Robineau-Desvoidy, 1830)

= Minettia muricata Becker, 1895, in Mouna 1998: 85

Mouna 1998; Ebejer and Kettani 2019a, Rif, Adrou (556 m), Perdicaris Park (Tanger, 223 m), HA, Mikdane (Jebel Ayachi) – NHMUK

Minettia tabidiventris (Rondani, 1877)

Ebejer and Kettani 2019a, Rif, Adrou (556 m), Oued Kbir (Béni Ratene (PNPB), 157 m), Oued Koub (Laghdir, 148 m), Rahbat Amlay (284 m)

Pachycerina Macquart, 1835

Pachycerina pulchra Loew, 1850

Ebejer and Kettani 2019a, Rif, Oued Zarka (Yarghite, 137 m)

Peplominettia Szilády, 1943

Peplominettia codinai (Hennig, 1951)

Ebejer and Kettani 2019a, Rif, Oued Zarka (Yarghite, 137 m), Azilane (1255 m)

Peplominettia striata Szilády, 1943

Ebejer and Kettani 2019a, Rif, Oued Zarka (Yarghite, 137 m)

Sapromyza Fallén, 1810

Sapromyza apicalis Loew, 1847

Schacht et al. 2004; Ebejer and Kettani 2019a, HA, Jebel Ayachi, Mikdane (stream), Lac Tislit (Imlchil, 2254 m) – NHMUK

Sapromyza gozmanyi Papp, 1981

Ebejer and Kettani 2019a, Rif, Oued Kbir (Béni Ratene (PNPB), 157 m), Perdicaris Park (Tanger, 223 m), Barrage Smir (M’Diq, 27 m)

Sapromyza obscuripennis Loew, 1847

Ebejer and Kettani 2019a, Rif, Jebel Talassemtane (1554 m), Jebel Lakraâ (1541 m)

Sapromyza unizona Hendel, 1908

Ebejer and Kettani 2019a, Rif, Perdicaris Park (Tanger, 223 m), AP, Larache (5 m)

Sapromyza (Sapromyzosoma) laevatrispina Carles-Tolrá, 1992

Ebejer and Kettani 2019a, Rif, Adrou (556 m), Issaguen (maison morestière, 1547 m)

Sapromyza (Sapromyzosoma) parallela Carles-Tolrá, 1992

Ebejer and Kettani 2019a, Rif, Adrou (556 m), Azilane (1255 m), Talassemtane (maison forestière, 1699 m), HA, Jaffar river (Jebel Ayachi, at maison forestière), Mikdane (Jebel Ayachi) – NHMUK

Sciomyzoidea

COELOPIDAE

K. Kettani

Number of species: 1. Expected: 2

Faunistic knowledge of the family in Morocco: moderate

Coelopinae

Coelopa Meigen, 1830

Coelopa pilipes Haliday, 1838

Séguy 1930a, Rif, Tanger; Dakki 1997; Lair 2013

DRYOMYZIDAE

K. Kettani

Number of species: 1. Expected: 1

Faunistic knowledge of the family in Morocco: poor

Dryomyzinae

Dryope Robineau-Desvoidy, 1830

Dryope flaveola (Fabricius, 1794)

Ebejer et al. 2019, Rif, Jebel Lakraâ (Talassemtane, 1596 m)

HELCOMYZIDAE

K. Kettani

Number of species: 1. Expected: 2

Faunistic knowledge of the family in Morocco: poor

Helcomyzinae

Helcomyza Curtis, 1825

Helcomyza ustulata Curtis, 1825

Cassar et al. 2005, Rif, Smir lagoon

SCIOMYZIDAE

K. Kettani, J-C. Vala

Number of species: 25. Expected: 40

Faunistic knowledge of the family in Morocco: good

Sciomyzinae

Sciomyzini

Ditaeniella Sack, 1939

Ditaeniella grisescens (Meigen, 1830)

Vala and Ghamizi 1991, MA, Ras el Ma, AA, Agadir; Kassebeer 1999a, MA, HA; Knutson and Vala 2011

Pherbellia Robineau-Desvoidy, 1830

Pherbellia cinerella (Fallén, 1820)

= Ditaenia cinerella Fallén, in Séguy 1941a: 31

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m), Imi-n’Ouaka (1500 m); Leclercq and Schacht 1987, HA, Ansegmir-Tal, W Midelt, HA, Tizi-n’Test (1900 m); Rozkošný 1987, HA; Vala and Ghamizi 1991, HA, MA; Carles-Tolrá 1993; Kassebeer 1999a, MA, HA; Rif (Talassemtane) – MISR

Pherbellia dorsata (Zetterstedt, 1846)

Kassebeer 1999a, MA; Knutson and Vala 2011

Pherbellia griseola (Fallén, 1820)

Kassebeer 1999a, MA; Knutson and Vala 2011

Pherbellia hermonensis Knutson & Freidberg, 1983

Kassebeer 1999a, HA; Knutson and Vala 2011

Pherbellia nana (Fallén, 1820)

= Pherbellia villiersi Séguy, in Séguy 1941a: 31; Knutson 1981: 336 (new comb.)

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Knutson 1981, HA, Tachdirt; Leclercq and Schacht 1987, HA; Vala and Ghamizi 1989, HA; Kassebeer 1999a, MA, HA; Knutson and Vala 2011; HA – MISR

Tetanocerini

Dichaetophora Rondani, 1868

Dichaetophora obliterata (Fabricius, 1805)

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Leclercq and Schacht 1987; Rozkošný 1987; Vala 1989; Vala and Ghamizi 1991; Kassebeer 1999a

Elgiva Meigen, 1838

Elgiva cucularia (Linnaeus, 1767)

Leclercq and Schacht 1987, HA, Ansegmir-Tal, W Midelt (1400 m); Vala 1989; Kassebeer 1999a

Euthycera Latreille, 1829

Euthycera algira (Macquart, 1849)

Vala and Reidenbach 1982, Rif, Tanger; Vala 1989, Rif, Tanger; Vala and Ghamizi 1991; Kassebeer 1999a, Rif, Tanger

Euthycera soror (Robineau-Desvoidy, 1830)

= Euthycera alaris Vala, 1983, syn. nov.Vala (pers. comm.)

Vala 1989, Rif, Tanger; Vala and Ghamizi 1991, MA; Carles-Tolrá 1993; Kassebeer 1999a, MA, HA

Euthycera stichospila (Czerny, 1909)

= Euthycera leclercqi Vala & Reidenbach, 1982, syn. by Rozkošný (1987)

Czerny and Strobl 1909; Vala and Reidenbach 1982; Rozkošný 1987

Euthycera zelleri (Loew, 1847)

Kassebeer 1999a, Rif, Tanger

Hydromya Robineau-Desvoidy, 1830

Hydromya dorsalis (Fabricius, 1775)

Séguy 1930a, Rif, Tanger; Leclercq and Schacht 1987, HA, Ansegmir-Tal, W Midelt (1400 m), HA, Oukaimeden (2600 m); Dakki 1997, Rif, Ketama; Vala 1989, Rif, Tanger; Vala and Ghamizi 1991, MA, HA; Kassebeer 1999a, MA, HA; Pârvu et al. 2006, AP, Merja Zerga

Ilione Haliday in Curtis, 1837

Ilione albiseta (Scopoli, 1763)

Leclercq and Schacht 1987, HA; Vala 1989; Kassebeer 1999a

Ilione trifaria (Loew, 1820)

Séguy 1941a, HA, Imi-n’Ouaka (1500 m); Vala 1989, MA, Dayat Aoua; Vala and Ghamizi 1991, EM, MA, HA; Carles-Tolrá 1993

Ilione unipunctata (Macquart, 1849)

Leclercq and Schacht 1987, HA; Rozkošný 1987; Vala 1989; Kassebeer 1999a

Oligolimnia Mayer, 1953

Oligolimnia zernyi Mayer, 1953

Rozkošný 1987, HA, Tachdirt; Leclercq and Schacht 1987, HA; Vala 1989, HA; Kassebeer 1999a, HA

Pherbina Robineau-Desvoidy, 1830

Pherbina coryleti (Scopoli, 1763)

Séguy 1930a, Rif, Tanger; Dakki 1997; Pârvu et al. 2006, AP, Merja Zerga; HA (Issougane n’Ouagouns) – MISR

Pherbina mediterranea Mayer, 1953

Leclercq and Schacht 1987, HA, Ansegmir-Tal, W Midelt (1400 m); Rozkošný 1987; Vala 1989; Vala and Ghamizi 1991, AP, Sidi Boughaba, HA, south of Marrakech; Carles-Tolrá 1993; Kassebeer 1999a, MA

 Psacadina Enderlein, 1939

Psacadina disjecta Enderlein, 1939

Verbeke 1964, AA, Tlata Reisana; Leclercq and Schacht 1987, HA, Ansegmir-Tal, W Midelt (1400 m); Rozkošný 1987; Vala 1989; Kassebeer 1999a, MA

Psacadina verbekei Rozkošný, 1975

Vala and Ghamizi 1991, HA, south of Marrakech; Kassebeer 1999a

Sepedon Latreille, 1804

Sepedon hispanica Loew, 1862

Verbeke 1964; Vala 1989, AP, Mohammedia; Vala and Ghamizi 1991, AP, Mohammedia, HA, Oued Tissaout (Kelaâ Sraghna); Kassebeer 1999a

Sepedon sphegea (Fabricius, 1775)

Séguy 1930a Rif, Tanger; Leclercq and Schacht 1987, HA, Ansegmir-Tal, W Midelt (1400 m); Rozkošný 1987; Vala 1989; Vala and Ghamizi 1991, HA, Tamesloht (south of Marrakech), AA, Bou Acheiba (Agadir); Carles-Tolrá 1993; Dakki 1997; Kassebeer 1999a

Sepedon spinipes (Scopoli, 1763)

Leclercq and Schacht 1987, HA, Ansegmir-Tal, W Midelt (1400 m); Vala and Ghamizi 1991, MA, Imouzzer, Tifounassine; Carles-Tolrá 1993; Kassebeer 1999a, MA; Knutson and Vala 2011

Trypetoptera Hendel, 1900

Trypetoptera punctulata (Scopoli, 1763)

Leclercq and Schacht 1987, HA; Vala 1989; Kassebeer 1999a; Knutson and Vala 2011

SEPSIDAE

K. Kettani, J.-P. Haenni

Number of species: 12. Expected: 20

Faunistic knowledge of the family in Morocco: poor

Sepsinae

Saltellini

Saltella Robineau-Desvoidy, 1830

Saltella sphondylii (Schrank, 1803)

Ebejer et al. 2019, Rif, Barrage Smir (145 m), Oued Mhajrate (Ben Karrich, 180 m)

Sepsini

Nemopoda Robineau-Desvoidy, 1830

Nemopoda nitidula (Fallén, 1820)

Mouna 1998 (no locality given)

Sepsis Fallén, 1810

Sepsis biflexuosa Strobl, 1893

Zuska and Pont 1984; Mouna 1998; Pont and Meier 2002; Ozerov 2005; HA (Asni) – NHMUK

Sepsis cynipsea (Linnaeus, 1758)

Séguy 1930a, AP, Rabat, MA, HA, Marrakech; Séguy 1941a, HA, Imi-n’Ouaka (1500 m); Zuska and Pont 1984; Mouna 1998

Sepsis flavimana Meigen, 1826

Ebejer et al. 2019, AA, Ziz river (10 km S of Errachidia, 1008 m)

Sepsis fulgens Meigen, 1826

Séguy 1941d, HA, Tizi-n’Test (2000 m), AA, Agadir; Zuska and Pont 1984; Mouna 1998; Pârvu et al. 2006, MA, Ifrane; Popescu-Mirceni 2011, MA, Ifrane; MA (Ifrane), HA (Asni, near Alrene, Mikdane) – NHMUK

Sepsis lateralis Wiedemann, 1830

Séguy 1941a, HA, Imi-n’Ouaka (1500 m); Zuska and Pont 1984; Mouna 1998; AP (Sale Tropical Garden), HA (Marrakech) – NHMUK

Sepsis punctum (Fabricius, 1794)

Séguy 1941a, HA, Aït Souka (Toubkal); Zuska and Pont 1984; Mouna 1998; Pont and Meier 2002, Rif, Tanger, AP, Rabat; Pârvu et al. 2006, AP, Merja Zerga – MISR; MA (Ifrane), HA (Asni, Mikdane), AA (Souss Massa, Agadir) – NHMUK

Sepsis thoracica (Robineau-Desvoidy, 1830)

Zuska and Pont 1984; Mouna 1998; Pârvu et al. 2006, AP, Merja Zerga; Popescu-Mirceni 2011; MA (near Azrou), HA (Mikdane, Marrakech, Amizmiz) – NHMUK

Sepsis violacea Meigen, 1826

= Sepsis ciliforceps Duda, 1926, in Mouna 1998: 86

Becker and Stein 1913, Rif, Tanger; Zuska and Pont 1984; Mouna 1998; Pârvu et al. 2006, AP, Merja Zerga; Popescu-Mirceni 2011 – MISR

Themira Robineau-Desvoidy, 1830

Themira minor (Haliday, 1833)

Zuska and Pont 1984; Mouna 1998 (no locality given); HA (Mikdane) – NHMUK

Themira paludosa Elberg, 196347

Pârvu et al. 2006, AP, Merja Zerga; Popescu-Mirceni 2011

An additional species, Australosepsis niveipennis (Becker, 1903) is reported from Morocco in major catalogues (Pont and Meier 1984, Ozerov 2005). However this report is based upon a misinterpretation by Zuska (1968) of a name of locality given in Duda (1926b: 2): “Marako” [a locality in Mali, or possibly another in Ethiopia] was wrongly understood for “Marokko”, the German name of Morocco. Both Adrian Pont and Andrey Ozerov have confirmed (pers. comm. 27.xi.15) that they have not seen any Moroccan specimen of A. niveipennis.

The Moroccan records from NHMUK were checked by Adrian Pont to whom we express our grateful thanks for his kind help.

Opomyzoidea

AGROMYZIDAE

K. Kettani, M. Černý

Number of species: 62. Expected: 150

Faunistic knowledge of the family in Morocco: poor

Agromyzinae

Agromyza Fallén, 1810

Agromyza abiens Zetterstedt, 1848

Spencer 1967, HA, Ourika Valley, Marrakech; Černý and Merz 2006

Agromyza albipennis Meigen, 1830

Séguy 1936b; Mouna 1998

Agromyza ambigua Fallén, 1823

= Domomyza ambigua Fallén, in Séguy 1936: 5; Mouna 1998: 84

Séguy 1936b, AP, Rabat; Mouna 1998

Agromyza bicaudata (Hendel, 1920)

Černý and Merz 2006, MA, Azrou, Ifrane

Agromyza frontella (Rondani, 1875)

Černý and Merz 2006, MA, Azrou, Ifrane

Agromyza frontosa (Becker, 1908)

Černý and Merz 2006, MA, Azrou, Ifrane

Agromyza hiemalis Becker, 1908

Spencer 1967, HA, Marrakech; Černý 2019; Černý et al. 2020

Agromyza intermittens (Becker, 1907)

= Phytomyza secalina Hering, 1925, in Maarouf 2003: 42

Maarouf 2003, HA, Chaouia

Agromyza luteitarsis (Rondani, 1875)

Maarouf 2003, HA, Chaouia

Agromyza megalopsis Hering, 1933

Černý et al. 2020, AA, Agadir Id Aissa (western end of gorge and village Amtoudi, 854 m)

Agromyza nana Meigen, 1830

Spencer 1967, 1973, AP, Casablanca; Černý and Merz 2006

Agromyza nigrociliata (Hendel, 1931)

Maarouf 2003, HA, Chaouia

Agromyza rondensis Strobl, 1900

Černý et al. 2020, AA, Agadir Id Aissa (western end of gorge and village Amtoudi, 854 m), River Aoulouz (= Asif Tifnout, 697 m)

Agromyza spenceri Griffiths, 1963

Černý and Merz 2006, HA, Asni; Černý 2013

Melanagromyza Hendel, 1920

Melanagromyza lappae (Loew, 1850)

Mouna 1998; AP (Rabat) – MISR

Melanagromyza verbasci Spencer, 1957

Spencer 1967, HA

Ophiomyia Braschnikov, 1897

Ophiomyia beckeri (Hendel, 1923)

Spencer 1967, HA; Černý and Merz 2006; Černý 2009; Černý and Tschirnhaus 2014

Ophiomyia curvipalpis (Zetterstedt, 1848)

= Ophiomyia proboscidea (Strobl, 1900), in Mouna 1998: 84

Spencer 1967, HA; Mouna 1998; Černý and Merz 2006; Černý and Merz 2007; Černý 2019

Ophiomyia melandryi de Meijere, 1924

Spencer 1967, HA; Černý and Merz 2006; Černý and Merz 2007; Černý 2009

Ophiomyia vimmeri Černý, 1994

Černý and Merz 2006, AP, Maâmora (Rabat); Černý and Merz 2007; Černý 2018

Phytomyzinae

Amauromyza Hendel, 1931

Amauromyza (Amauromyza) morionella (Zetterstedt, 1848)

= Agromyza morionella Zetterstedt, in Becker and Stein 1913: 95

Aulagromyza Enderlein, 1936

Aulagromyza atlantidis (Spencer, 1967)

= Paraphytomyza atlantidis Spencer, 1967, in Mouna 1998: 84

Spencer 1967, HA, Asni; Mouna 1998

Aulagromyza cydoniae (Hendel, 1936)

= Phytagromyza cydoniae Hendel, 1936, in Hendel 1931–1936: 518

Hendel 1931–1936, AP, Rabat

Aulagromyza hamata (Hendel, 1932)*

HA, Asni-Ouirgane

Cerodontha Rondani, 1861

Cerodontha (Cerodontha) denticornis (Panzer, 1806)

Spencer 1967, 1973, HA; Černý and Merz 2006; Černý 2009

Cerodontha (Cerodontha) fulvipes (Meigen, 1830)

Mouna 1998

Cerodontha (Dizygomyza) brisiaca Nowakowski, 1973

Černý and Merz 2006, MA, Azrou, Ifrane

Cerodontha (Icteromyza) capitata (Zetterstedt, 1848)

Černý and Merz 2006, Rif, Chefchaouen

Cerodontha (Icteromyza) rozkosnyi Černý, 2007

Černý 2007, AA, SW Tazenakht (1000 m); Černý 2011

Cerodontha (Poemyza) incisa (Meigen, 1830)

= Dizygomyza incisa Meigen, in Séguy 1936b: 5; Mouna 1998: 84

Séguy 1936b, AP, Rabat; Mouna 1998

Cerodontha (Poemyza) lateralis (Macquart, 1835)

= Dizygomyza lateralis Macquart, in Séguy 1936: 5; Mouna 1998: 84

Séguy 1936b, AP, Rabat; Mouna 1998; Černý et al. 2020

Cerodontha (Poemyza) pygmaea (Meigen, 1830)

= Dizygomyza pygmaea Meigen, in Séguy 1936: 5; Mouna 1998: 84

Séguy 1936b, AP, Rabat; Mouna 1998

Chromatomyia Hardy, 1849

Chromatomyia aprilina (Goureau, 1851)

Griffiths 1974; Spencer 1967, Rif, Tanger (mountains)

Chromatomyia horticola (Goureau, 1851)

= Phytomyza horticola Goureau, in Griffiths 1967: 14

Griffiths 1967, AP, Casablanca; Spencer 1973; Černý 2009

Chromatomyia milii (Kaltenbach, 1864)

= Phytomyza milii Kaltenbach, in Séguy 1936: 5; Mouna 1998: 84

Séguy 1936b, AP, Rabat; Spencer 1967, HA, Ourika Valley; Griffiths 1980; Mouna 1998; Černý and Merz 2006

Chromatomyia syngenesiae Hardy, 1849

= Phytomyza atricornis Meigen, in Kozlowsky and Rungs 1932: 66; Mouna 1998: 84

Kozlowsky and Rungs 1932, AP, Rabat, Kénitra, Casablanca; Mouna 1998; AP (Rabat) – MISR

Liriomyza Mik, 1894

Liriomyza bryoniae (Kaltenbach, 1858)

Spencer 1967, 1973, AP, Casablanca; Ayoub 2002, AA, Souss Massa, Agadir; Černý and Merz 2006

Liriomyza cicerina (Rondani, 1874)

Spencer 1973, HA; Lahmar and Zeouienne 1992; Mouna 1998

Liriomyza congesta (Becker, 1903)

Černý et al. 2020, AA, River bed of Ougni, 0.6 km N Akka N’Ait Sidi and 1.8 km NW Tissint (582 m), River Aoulouz (= Asif Tifnout), bridge of road (697 m), River Oued Draa near hotel, Gardin Oued Tamnougalt (911 m)

Liriomyza huidobrensis (Blanchard, 1926)

Hanafi and Schnitzler 2004, AA, Souss Valley

Liriomyza orbona (Meigen, 1830)

= Agromyza fuscolimbata Strobl, 1900, in Maarouf 2003: 43

= Liriomyza orbonella Hendel, 1931, in Maarouf 2003: 43

Maarouf 2003, HA, Chaouia

Liriomyza pedestris Hendel, 1931

Spencer 1967, Rif, Tanger; Černý and Merz 2006; Černý and Merz 2007; Černý 2019; Černý et al. 2020

Liriomyza pusilla (Meigen, 1830)

= Agromyza pusilla Meigen, in Mouna 1998: 84

Mouna 1998

Liriomyza sonchi Hendel, 1931

Spencer 1967, AP, Casablanca

Liriomyza trifolii (Burgess in Comstock, 1880)

Hanafi and Schnitzler 2004, AA, Souss Valley

Napomyza Westwood, 1840

Napomyza cichorii Spencer, 1966

Černý and Merz 2006, MA, Ifrane; Černý 2009

Napomyza lateralis (Fallén, 1823)

Černý and Merz 2006, MA, Ifrane

Napomyza scrophulariae Spencer, 1966

Černý and Merz 2006, MA, Ifrane; Černý and Merz 2007; Černý 2012, 2013

Phytoliriomyza Hendel, 1931

Phytoliriomyza immoderata Spencer, 1963

Černý and Merz 2006, MA, Azrou, Ifrane; Černý 2009; Černý 2019; Černý et al. 2020

Phytoliriomyza oasis (Becker, 1907)

Spencer 1967, HA; Černý and Merz 2006

Phytomyza Fallén, 1810

Phytomyza conyzae Hendel, 1920

Spencer 1967, Rif, Tanger; Černý 2009

Phytomyza ferulae Hering, 1927

Spencer 1967, Rif, Tanger; Černý 2013; Černý et al. 2020

Phytomyza gymnostoma Loew, 1858

Mouna 1998; AP (Mechra el kettane) – MISR

Phytomyza orobanchia Kaltenbach, 1864

Geipert 1993; Geipert et al. 1994, Rif; Klein 1995; Klein et al. 1995; Boumezzough 1996, MA, Saiss, Rommani; Boughdad et al. 1997; Klein et al. 1999; Kroschel and Klein 1999; Yazough and Klein 1999; Kroschel and Klein 2003

Phytomyza phillyreae Hering in Buhr, 1930

= Phytomyza unedo Séguy, 1953, in Séguy 1953b: 72

Séguy 1953b, AP, Korifla; Spencer 1967, HA, Ourika Valley; Černý and Merz 2006; Černý 2009

Phytomyza ranunculi (Schrank, 1803)

Spencer 1967, HA; Černý and Merz 2006; Černý 2009, 2013

Phytomyza wahlgreni Rydén, 1944

Černý and Merz 2006, MA, Azrou, Ifrane; Černý et al. 2020

Pseudonapomyza Hendel, 1920

Pseudonapomyza atra (Meigen, 1830)

= Phytomyza acuticornis Loew, 1858, in Maarouf 2003: 43

Maarouf 2003, HA, Chaouia

Pseudonapomyza atratula Zlobin, 2003

Černý et al. 2020, AA, 6 km ESE Quijjane, 85 km S Agadir (353 m)

Pseudonapomyza bifida Zlobin, 2003

Černý et al. 2020, AA, Road no. 109/165 from Akna to Taroudant (841 m)

Pseudonapomyza spicata (Malloch, 1914)

Černý et al. 2020, AA, River bed of Ougni, 0.6 km N Akka N’Ait Sidi and 1.8 km NW Tissint (582 m)

Pseudonapomyza spinosa Spencer, 1973

Maarouf 2003, HA, Chaouia

New record for Morocco

Aulagromyza hamata (Hendel, 1932)

High Atlas: Asni-Ouirgane, 31°13'52"N, 8°00'8"W, 1282 m a.s.l., 1♂, 24.iv.2014, river valley, V. Vrabec leg., M. Barták coll. and M. Černý det.

ANTHOMYZIDAE

K. Kettani, M.J. Ebejer

Number of species: 2. Expected: 4

Faunistic knowledge of the family in Morocco: poor

Amygdalops Lamb, 1914

Amygdalops thomasseti Lamb, 1914

Ebejer et al. 2019, Rif, Stehat (0 m)

Anagnota Becker, 1902

Anagnota major Roháček & Freidberg, 1993

Roháček 2006, HA, Marrakech (1000 m)

ASTEIIDAE

K. Kettani, M.J. Ebejer

Number of species: 5. Expected: 10

Faunistic knowledge of the family in Morocco: poor

Asteiinae

Asteia Meigen, 1830

Asteia amoena Meigen, 1830

Mouna 1998; AP (Rabat) – MISR

Asteia ibizana (Enderlein, 1935)

Ebejer et al. 2019, AP, Larache (2 m)

Asteia mahunkai Papp, 1979

Ebejer et al. 2019, AP, Larache (2 m)

Phlebosotera Duda, 1927

Phlebosotera clypeata Freidberg & Carles-Tolrá, 2010

Freidberg and Carles-Tolrá 2010, HA, Jaffar river

Phlebosotera mirabilis Papp, 1972

Ebejer et al. 2019, AA, 12 km S of Rissani (737 m)

AULACIGASTRIDAE

K. Kettani, M.J. Ebejer

Number of species: 1. Expected: 2

Faunistic knowledge of the family in Morocco: poor

Aulacigaster Macquart, 1835

Aulacigaster leucopeza (Meigen, 1830)

Ebejer et al. 2019, Rif, Jebel Lakraâ (Talassemtane, 1541 m)

CLUSIIDAE

K. Kettani, M.J. Ebejer

Number of species: 1. Expected: 3

Faunistic knowledge of the family in Morocco: poor

Clusiodes Coquillett, 1904

Clusioides verticalis (Collin, 1912)

Ebejer et al. 2019, Rif, Amsemlil bog (PNPB, 1067 m)

ODINIIDAE

K. Kettani, M.J. Ebejer

Number of species: 2. Expected: 3

Faunistic knowledge of the family in Morocco: poor

Odiniinae

Odinia Robineau-Desvoidy, 1830

Odinia Boletina (Zetterstedt, 1848)

Séguy 1934a; Mouna 1998; Gaimari and Mathis 2011

Odinia meijerei Collin, 1952

Ebejer et al. 2019, Rif, Adrou (PNPB, 556 m)

OPOMYZIDAE

K. Kettani, M.J. Ebejer

Number of species: 5. Expected: 6

Faunistic knowledge of the family in Morocco: moderate

Geomyza Fallén, 1810

Geomyza apicalis (Meigen, 1830)

Pârvu et al. 2006, MA, Ifrane; Popescu-Mirceni 2011, AP, Merja Zerga

Geomyza combinata (Linnaeus, 1767)

Mouna 1998; AP (Maâmora) – MISR

Geomyza tripunctata (Fallén, 1823)

Maarouf 2003, HA, Chaouia

Opomyza Fallén, 1820

Opomyza florum (Fabricius, 1794)

Maarouf 2003, HA, Chaouia

Opomyza petrei Mesnil, 1934

Ebejer et al. 2019, Rif, Aïn Tissemlal (Azilane, 1255 m)

Carnoidea

CANACIDAE

K. Kettani, L. Munari

Number of species: 15. Expected: 25

Faunistic knowledge of the family in Morocco: poor

Canacinae

Canace Haliday in Curtis, 1837

Canace actites Mathis, 1982

Ebejer et al. 2019, AP, Loukous (2 m)

Canace nasica Haliday, 1839

Dahl 1964, AA, Tamri (north of Agadir); Mouna 1998

Xanthocanace Hendel, 1914

Xanthocanace ranula (Loew, 1874)

Munari 2010, AP, 40 km S Larache; Munari and Mathis 2010; Munari 2011; Munari and Bramuzzo 2018, Rif, Briyech, AP, Azemmour, El Khaoucha, Fedala, Oued Nefifikh, Oued Loukous, 40 km S Larache

Tethininae

Tethina Haliday in Curtis, 1837

Tethina alboguttata (Strobl, 1900)

Freidberg and Beschovski 1996, AP, Aïn Diab, Essaouira, Safi, AA, Agadir, Ouarzazate; Mathis and Munari 1996; Munari 2002, 2004, 2005, 2011; Munari and Mathis 2010; Koçak and Kemal 2010; Munari and Bramuzzo 2018, Rif, Briyech, AP, Agadir, Tamri, Essaouira, Cap Hadid, Larache, Loukous, Safi, Dar Caïd-Hadji

Tethina albosetulosa (Strobl, 1900)

Cassar et al. 2008, Rif, Oued Laou Basin

Tethina flavigenis (Hendel, 1934)

Munari and Bramuzzo 2018, AP, Oued Oum-er-Rbia

Tethina grossipes (Becker, 1908)

Munari 2004, AA, Agadir; Munari and Mathis 2010; Munari and Bramuzzo 2018, AP, Tamri

Tethina incisuralis (Macquart, 1851)

= Tethina pictipes (Becker 1903); Séguy 1941d: 18, Mouna 1998: 87

Munari 1997, AA, Erfoud, Rissani (900 m); Munari 2002, 2010, 2011; Munari and Mathis 2010; Munari and Bramuzzo 2018, AA, Erfoud, Rissani

Tethina longirostris (Loew, 1865)

Cassar et al. 2008, Rif, Oued Laou Basin

Tethina mariae Munari, 1997

Munari 1997, AP, 40 km S Larache; Munari and Báez 2000; Munari 2002, 2004, 2010, 2011; Munari and Mathis 2010 – NHMD (HT ♂)

Tethina pallipes (Loew, 1865)

= Tethina ochracea (Hendel, 1913)

Cassar et al. 2008, Rif, Oued Laou

Tethina pictipennis Freidberg & Beschovski, 1996

Freidberg and Beschovski 1996, AP, 40 km south of Larache; Mathis and Munari 1996, Munari 2002; Munari 2004; Munari and Mathis 2010; Munari 2011, AA, Agadir (Tamri); Munari and Bramuzzo 2018, Rif, Briyech, AP, Tamri, Larache, Loukous – NHMD (HT ♂)

Tethina strobliana (Mercier, 1923)

Munari and Bramuzzo 2018, AP, Oued Abou, Rehouna (Rabat)

Tethina yaromi Freidberg & Beschovski, 1996

Cassar et al. 2005, Rif, Smir lagoon

Tethina  sp. near salinicola Beschovski, 1998

Munari 2004, AA, Agadir, Ouarzazate; Munari 2005; Munari and Bramuzzo 2018, AP, Tamri

CARNIDAE

K. Kettani, M.J. Ebejer

Number of species: 3. Expected: >10

Faunistic knowledge of the family in Morocco: poor

Meoneura Rondani, 1856

Meoneura hungarica Papp, 1977

Ebejer et al. 2019, Rif, Adrou (PNPB, 556 m)

Meoneura prima Becker, 1905

Brake 2011

Meoneura triangularis Collin, 1930

Brake 2011

CHLOROPIDAE48

K. Kettani, M. von Tschirnhaus, M.J. Ebejer

Number of species: 74. Expected: 140

Faunistic knowledge of the family in Morocco: moderate

Chloropinae

Assuania Becker, 1903

Assuania melanoleuca (Séguy, 1949) comb. nov.49

= Chlorops melanoleuca Séguy, in Séguy 1949a: 158

Séguy 1949a, AA, Agdz; Nartshuk 1984; Mouna 1998

Assuania thalhammeri (Strobl, 1893)

Ebejer and Kettani 2016, AA, Oued Ziz (9.5 km SE of Rich, 1285 m)

Camarota Latreille, 1805

Camarota curvipennis (Latreille, 1805)

= Camarota curvipennis (Latreille), in Becker and Stein 1913: 93

Becker and Stein 1913, Rif, Tanger; Nartshuk 1984; Ebejer and Kettani 2016, Rif, Oued Laou

Capnoptera Loew, 1866

Capnoptera pilosa Loew, 1866

Duda 1933; Ebejer and Kettani 2016, Rif, Jebel Lakraâ (Talassemtane, 1288 m), Aïn Jdioui (Tahaddart, 8 m)

Capnoptera scutata (Rossi, 1790)

= Eristalis rufipes Fabricius, in Fabricius 1805: 245

Ebejer and Kettani 2016, AP, Larache (2 m)

Cetema Hendel, 1907

Cetema maroccanum Nartshuk, 1995

= Cetema maroccana Nartshuk, in Nartshuk 1995: 277–280

Nartshuk 1995, HA, Oukaimeden; Ebejer and Kettani 2016

Cetema monticulum Becker, 1910

= Cetema monticula Becker, in Séguy 1941a: 33

= Cetema monticula Rossi, in Mouna 1998: 85

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Mouna 1998; Ebejer and Kettani 2016

Chlorops Meigen, 1803

Chlorops interruptus Meigen, 1830

= Chlorops interrupta Meigen, in Séguy 1953a: 86

Séguy 1953a, AP, Oued Yquem (near Rabat); Ebejer and Kettani 2016, Rif, Jebel Lakraâ (Talassemtane, 1288 m), Oued Aliane (Ksar Sghir, 1 m), Cap Spartel (155 m)

Chlorops limbatus Meigen, 1830

Ebejer and Kettani 2016, Rif, Jebel Moussa (800 m)

Chlorops pumilionis (Bjerkander, 1778)

= Chlorops nasuta Schrank, in Mouna 1998: 85

Mouna 1998; Ebejer and Kettani 2016

Chlorops serenus Loew, 1866

Ebejer and Kettani 2016, MA, 17 km NW of Zaida (Khénifra, 1878 m)

Cryptonevra Lioy, 1864

Cryptonevra flavitarsis (Meigen, 1830)

= Haplegis flavitarsis (Meigen), in Séguy 1949a: 158; Séguy 1953a: 86; Mouna 1998: 85

Séguy 1949a, AA, Agdz; Séguy 1953a, AP, Oued Yquem (near Rabat); Nartshuk 1984; Mouna 1998; Ebejer and Kettani 2016

Eurina Meigen, 1830

Eurina lurida Meigen, 1830

Ebejer and Kettani 2016, Rif, M’Diq, Smir, Kabila beach and dunes

Eutropha Loew, 1866

Eutropha albipilosa (Becker, 1908)

Duda 1930; Deeming and Al-Dhafer 2012, AP, 17 km N of Larache, AA, Azemmour, estuary of Oued Tensift; Ebejer and Kettani 2016, AP, Larache (5 m)

Eutropha fulvifrons (Haliday, 1833)

Séguy 1941d, AA, Agadir; Mouna 1998; Cassar et al. 2005, Rif, Smir lagoon; Ebejer and Kettani 2016, AP, Larache (5 m)

Lagaroceras Becker, 1903

Lagaroceras andalusiaca (Strobl, 1899)

Ebejer and Kettani 2016, Rif, Ksar El Kebir (13 m), Oued Aliane (Ksar Sghir, 1 m), AP, Larache (2 m), AA, Oued Ziz (9.5 km SE of Rich, 1285 m)

Lasiosina Becker, 1910

Lasiosina herpini (Guérin-Méneville, 1843)

= Lasiosina cinctipes Meigen, in Mouna 1998: 85

Ebejer and Kettani 2016, Rif, Smir lagoon

Lasiosina lindbergi (Duda, 1933)

= Steleocerus lindbergi (Duda), in Duda 1933: 142

Duda 1933, MA; Ebejer and Kettani 2016, AA, Oued Ziz (9.5 km SE of Rich, 1285 m)

Meromyza Meigen, 1830

Meromyza athletica Fedoseeva, 1974

= Meromyza variegata Meigen, in Mouna 1998: 85

Mouna 1998; Ebejer and Kettani 2016

Meromyza curvinervis (Zetterstedt, 1848)

= Oxinis curvinervis Latreille, in Mouna 1998: 85

Mouna 1998

Meromyza nigriventris Macquart, 1835

Ebejer and Kettani 2016, MA, 17 km SW of Midelt (Khénifra, 1940 m), AA, 29 km N of Rich (Errachidia, 1570 m)

Meromyza pratorum Meigen, 1830

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Mouna 1998; Ebejer and Kettani 2016

Metopostigma Becker, 1903

Metopostigma sabulona Becker, 1910

Ebejer and Kettani 2016, AA, Oued Ziz (10 km S of Errachidia, 1008 m), Merzouga (714 m)

Platycephala Fallén, 1820

Platycephala scapularum (Becker, 1907)

Nartshuk 1984; Pârvu et al. 2006, AA, Lac Edehby, Ouarzazate; Popescu-Mirceni 2011; Ebejer and Kettani 2016

Pseudopachychaeta Strobl, 1902

Pseudopachychaeta pachycera Strobl, 1902

Ebejer and Kettani 2016, AA, Oued Ziz (9.5 km SE of Rich, 1285 m), Oued Ziz (1052 m)

Thaumatomyia Zenker, 1833

Thaumatomyia elongatula (Becker, 1910)

Ebejer and Kettani 2016, Rif, Beni Maâdene (Oued Martil, 3 m)

Thaumatomyia glabra (Meigen, 1830)

Ebejer and Kettani 2016, Rif, Martil (9 m)

Thaumatomyia notata (Meigen, 1830)

= Chloropisca notata (Meigen), in Séguy 1930a: 179, 1941a: 33; Mouna 1998: 85

Séguy 1930a, Rif, Tanger; Séguy 1941a, HA, Tachdirt (Jebel Toubkal, 2500 m), Canyon Tessaout (M’Goum, 3000–3200 m); Harris et al. 1980: 229, HA, Chichaoua; Mouna 1998; Ebejer and Kettani 2016, Rif, Oued Laou estuary, Oued Amsa (Amsa, 14 m), Jnane Niche (27 m), Oued Maggou (Maggou, 786 m), Marj Khayl (Beni Leit (PNPB), 1088 m), Oued Boumarioul (Aïn Hamra, 560 m), Aïn Tissemlal (Azilane, 1255 m), Oued Bouhya (Bou Ahmed, 19 m), Souk Khemis Anjra (Oued Kbir, 55 m), Zinat (231 m), Oued Zarka (Yarghite, 135 m), Oued Talembote (Usine électrique, 120 m), Oued Kelâa (Akoumi, 400 m), Issaguen (maison forestière Issaguen, 1543 m), Bni Boufrah (94 m), Oued Guallet (Bni Boufrah, 946 m), Oued Tabandoute (Bni Boufrah, 540 m), Oued Taâouniya (Koudiat Ajira, 1536 m), Talankramte (Site sacré Sidi Gneiss (PNPB): 461 m), AP, Larache (5 m)

Thaumatomyia sulcifrons (Becker, 1907)

= Chlorops sulcifrons Becker, in Séguy 1949a: 158

= Chloropisca sulcifrons Zetterstedt, in Mouna 1998: 85

Séguy 1934a; Séguy 1949a, AA, Foum-el-Hassan, Akka, Agdz; Nartshuk 1984; Mouna 1998; Dawah and Abdullah 2006; Ebejer and Kettani 2016

Oscinellinae

Aphanotrigonum Duda, 1932

Aphanotrigonum cinctellum (Zetterstedt, 1848)

Ebejer and Kettani 2016, AP, Larache (5 m)

Aphanotrigonum femorellum Collin, 1946

Ebejer and Kettani 2016, AP, Larache

Aphanotrigonum inerme Collin, 1946

Ebejer and Kettani 2016, Rif, M’Diq (5 m), AP, Larache (Loukous marsh, 2 m)

Aphanotrigonum parahastatum Dely-Draskovits, 1981*

AA

Calamoncosis Enderlein, 1911

Calamoncosis duinensis (Strobl, 1909)

Ebejer and Kettani 2016, AP, Rabat (on Phragmites) – MISR

Conioscinella Duda, 1929

Conioscinella frontella (Fallén, 1810)

Ebejer and Kettani 2016, Rif, Aïn Tissemlal (Azilane, 1255 m)

Conioscinella sordidella (Zetterstedt, 1848)

Ebejer and Kettani 2016, Rif, Aïn Tissemlal (Azilane, 1255 m)

Elachiptera Macquart, 1835

Elachiptera bimaculata (Loew, 1845)

De Lépiney and Mimeur 1932: 110, Rabat; Séguy 1934a: 479; Bléton and Fieuzet 1943; Nartshuk 1984; Mouna 1998; Ebejer and Kettani 2016, Rif, Beni Maâdene (Oued Martil: 3 m), Jnane Niche (46 m), Oued Mhannech (18 m), Souk Khemis Anjra (55 m), Boujdad (7 m); AP (Rabat) – MISR

Elachiptera cornuta (Fallén, 1820)

Mouna 1998: 85

Elachiptera diastema Collin, 1946

Ebejer and Kettani 2016, Rif, Jebel Lakraâ (Talassemtane, 1541 m), Aïn Tissemlal (Azilane, 1255 m), Oued Maggou (Maggou, 786 m), Oued Tiffert (Tiffert, 1230 m), Issaguen (1543 m), AP, Larache (5 m)

Elachiptera graeca Becker, 1910

Séguy 1941, HA, Imi-n’Ouaka (1500 m); Nartshuk 1984; Mouna 1998

Elachiptera megaspis (Loew, 1858)

Nartshuk 1984; Mouna 1998; Ebejer and Kettani 2016, Rif, Dardara (730 m), Aïn Jdioui (Tahaddart, 8 m), Jnane Niche (46 m), Aïn Tissemlal (Azilane, 1255 m), Oued Maggou (Maggou, 786 m); AP (Rabat) – MISR

Elachiptera orizae Séguy, 1949*

AA

Elachiptera rufescens (Walker, 1871)

Deeming and Al-Dhafer 2012, AA, 2 km N Erfoud (818 m); Ebejer and Kettani 2016, AA, 2 km N Erfoud (818 m)

Elachiptera rufifrons Duda, 1932

Ebejer and Kettani 2016, AP, 9 km SE Aïn Chouk (6 m), Larache (5 m)

Elachiptera sarda Nartshuk, 2009

Ebejer and Kettani 2016, AA, 14 km E of Rich (Errachidia, 1278 m)

Elachiptera scrobiculata (Strobl, 1901)

= Elachiptera trapezina (Corti, 1909), in Bléton and Fieuzet 1943: 116

Bléton and Fieuzet 1943; Ebejer and Kettani 2016

Elachiptera strobli (Corti, 1909)

Ebejer and Kettani 2016, Rif, Oued Boumarioul (Aïn Hamra, 560 m), Oued Aliane (Ksar Sghir, 1 m); Cap Spartel (155 m), Oued Sidi Ben Saâda (Laghdir, 242 m)

Epimadiza Becker, 1910

Epimadiza nigrescens Duda, 193350

= Oscinosoma anthracias Séguy, in Séguy 1949a: 158, Mouna 1998: 85

= Oxinosoma anthracias Séguy, in Mouna 1998: 85

= ”Siphonella oscinina (Fallén)”, in Nartshuk 1984: 236

Séguy 1949a, AA, Alnif; Sabrosky 1965: 406; Nartshuk 1984; Mouna 1998; Ebejer and Kettani 2016

Hapleginella Duda, 1933

Hapleginella laevifrons (Loew, 1858)

El Hassani et al. 1986: 8–11, Rif, Nord Occidental, MA

Lasiochaeta Corti, 1909

Lasiochaeta pubescens (Thalhammer, 1898)

= Elachiptera pubescens (Thalhammer), var. rufithorax Duda, 1932 in Duda 1932: 31

= Melanochaeta pubescens Thalh., in De Lépiney and Mimeur 1932: 110

= Melanochaeta pubescens (Thalhammer), in Mouna 1998: 85

De Lépiney and Mimeur 1932, AP, Rabat; Duda 1932: 31, HA; Nartshuk 1984; Mouna 1998; Ebejer and Kettani 2016, Rif, Oued Sidi Yahia Aarab (Sidi Yahia Aarab, 178 m), Beni Maâdene (Oued Martil, 3 m), Oued Mhannech (Tamuda, 18 m), Oued Amsa (Amsa, 14 m), Zinat (231 m), Oued Maggou (Maggou, 786 m), Oued Guallet (Bni Boufrah, 946 m); AP (Rabat) – MISR

Oscinella Becker, 1909

Oscinella cariciphila Collin, 1946

Ebejer and Kettani 2016, Rif, Beni Maâdene (Oued Martil, 3 m), AA, Merzouga (714 m)

Oscinella frit (Linnaeus, 1758)

= Oxinosoma frit Linnaeus, in Mouna 1998: 85

De Lépiney and Mimeur 1932: 109, AP, Rabat; Mouna 1998; Ebejer and Kettani 2016, Rif, Issaguen (1543 m), Aïn Tissemlal (Azilane, 1255 m), Oued Maggou (Maggou, 786 m), Oued Guallet (Bni Boufrah, 946 m), Oued Ametrasse (Ametrasse, 841 m), AP, Lower Loukous saltmarsh (2 m), MA, Khénifra (17 km SW of Midelt, 1940 m), Lac Aguelmane Afennourir (30 km SW of Azrou, 1490 m); AP (Rabat) – MISR

Oscinella nartshukiana Beschovski, 1978

Ebejer and Kettani 2016, Rif, Ksar El Kebir (13 m), AP, Larache

Oscinella nitidigenis (Becker, 1908)

Ebejer and Kettani 2016, AA, 6 km N of Errachidia (1010 m), Oued Ziz (1052 m)

Oscinella nitidissima (Meigen, 1838)

Ebejer and Kettani 2016, Rif, Dardara (730 m), Moulay Abdelsalam (965 m), Cap Spartel (155 m), Oued Laou (dunes, 2 m), AP, Larache (5 m), MA, Lac Aguelmane Afennourir (30 km SW of Azrou, 1490 m)

Oscinella pusilla (Meigen, 1830)

Ebejer and Kettani 2016, AA, Lac Tiffert (4 km W of Merzouga, 702 m)

Oscinella ventricosi Nartshuk, 1955

Ebejer and Kettani 2016, Rif, Oued Boumarioul (Aïn Hamra, 560 m), AP, Larache (5 m)

Oscinella vindicata (Meigen, 1830)

Ebejer and Kettani 2016, AP, Larache (5 m), AA, Merzouga (agriculture under date palms, 714 m)

Oscinimorpha Lioy, 1864

Oscinimorpha arcuata (Duda, 1932)

Ebejer and Kettani 2016, Rif, Oued Guallet (Bni Boufrah, 946 m), Oued Jnane Azaghar (Bni Boufrah, 997 m)

Oscinimorpha longirostris (Loew, 1858)

Ebejer and Kettani 2016, Rif, Ksar El Kebir (13 m)

Oscinimorpha minutissima (Strobl, 1900)

= Siphonella minutissima Strobl, in Becker and Stein 1913: 93

Becker and Stein 1913, Rif, Tanger; Nartshuk 1984; Ebejer and Kettani 2016, Rif, Dardara (484 m)

Oscinimorpha novakii (Strobl, 1893)

= Conioscinella novakii (Strobl), in Duda 1933: 58

Duda 1933; Ebejer and Kettani 2016

Oscinisoma Lioy, 1864

Oscinisoma cognatum (Meigen, 1830)

= Oscinis rufipes Meigen, 1830, with homonym Oscinis rufipes Wiedemann 1830: 580, in Wiedemann 1830: 580, Tanger; synonymy and probable specific identity of homonyms established by Becker 1910: 166, but without considering the similar Oscinisoma gilvipes (Loew, 1858).

Polyodaspis Duda, 193351

Polyodaspis sulcicollis (Meigen, 1838)

= Siphonella sulcicollis (Meigen), in Kroschel and Klein 1999: 138

Kroschel and Klein 1999; Ebejer and Kettani 2016, MA, Khénifra (17 km SW of Midelt, 1940 m)

Pselaphia Becker, 1911

Pselaphia dimidiocera Ebejer & Kettani, 2016

Ebejer and Kettani 2016, Rif, Adrou (Taghzout, 556 m) – NMWC

Rhodesiella Adams, 1905

Rhodesiella fedtshenkoi Nartshuk, 1978

Deeming and Al-Dhafer 2012, SA, Goulimine (Bou Jarif); Ebejer and Kettani 2016, Rif, Smir lagoon, Oued Laou (saltmarsh), Jnane Niche (46 m)

Sabroskyina Beschovski, 1987

Sabroskyina aharonii (Duda, 1933)

Ebejer and Kettani 2016, AA, 2 km N Erfoud (818 m)

Siphunculina Rondani, 1856

Siphunculina ornatifrons (Loew, 1858)

= Microneurum ornatifrons (Loew), in Duda 1933: 98

Becker and Stein 1913, Rif, Tanger; Duda 1933; Nartshuk 1984; Ebejer and Kettani 2016, Rif, Martil (9 m), Oued Mhannech (Tamuda, 18 m), Jnane Niche (27 m), AP, Aïn Chouk 9 km SE (6 m), Larache (5 m)

Trachysiphonella Enderlein, 1936

Trachysiphonella ruficeps (Macquart, 1835)

Ebejer and Kettani 2016, AA, 29 km N of Rich (Errachidia, 1570 m)

Tricimba Lioy, 1864

Tricimba cincta (Meigen, 1830)

Ebejer and Kettani 2016, Rif, Dardara (484 m)

Tricimba heratica Dely-Draskovits, 1983*

AA

Tricimba humeralis (Loew, 1858)

= Tricimba punctifrons Becker, in Becker and Stein 1913: 93

= Tricimba humeralis (Loew), in Séguy 1941d: 18, 1957: 273; Mouna 1998: 85

Becker and Stein 1913, Rif, Tanger; Séguy 1941d, AA, Agadir; Séguy 1957, AA, Agadir; Nartshuk 1984; Mouna 1998; Ebejer and Kettani 2016, Rif, Oued Mhannech (Tamuda, 18 m), Aïn Tissemlal (Azilane, 1255 m), Oued Bouhya (Bou Ahmed, 19 m), AA, Oued Ziz (9.5 km SE of Rich, 1285 m), Oued Ziz (13 km N of Erfoud, 800 m)

Siphonellopsinae

Apotropina Hendel, 1907

Apotropina longepilosa (Strobl, 1893)

Ebejer and Kettani 2016, Rif, Oued Sidi Ben Saâda (Laghdir, 242 m), Oued Kbir (Dardara, 345 m), Oued Siflaou (281 m), Jnane Niche (46 m)

Siphonellopsis Strobl, 1906

Siphonellopsis lacteibasis Strobl, 1906

Séguy 1934a: 482; Nartshuk 1984; Mouna 1998

New records for Morocco

Aphanotrigonum parahastatum Dely-Draskovits, 1981

Anti Atlas: Ougui river, 1.8 km NW Tissint, 29°55'03"N, 7°19'56"W, 582 m, 30.xii.2016, 3♂♂3♀♀, M. von Tschirnhaus det. – ZSM (M. von Tschirnhaus leg.).

Elachiptera orizae Séguy, 1949

Anti Atlas: river Aoulouz up stream of Idrgane, 30°44'11"N, 7°59'13"W, 866 m, 31.xii.2016, 1♂, M. von Tschirnhaus det. – ZSM (M. von Tschirnhaus leg.).

Tricimba heratica Dely-Draskovits, 1983

Anti Atlas: Oued Souss, streamup of Idrgane, 30°44'11"N, 7°59'13"W, 866 m, 31.xii.2016, 1♀ (males are needed to confirm the identification), M. von Tschirnhaus det. – ZSM (M. von Tschirnhaus leg.).

MILICHIIDAE

K. Kettani

Number of species: 8. Expected: 14

Faunistic knowledge of the family in Morocco: moderate

Madizinae

Madizini

Desmometopa Loew, 1866

Desmometopa m-nigrum (Zetterstedt, 1848)

Séguy 1949a, SA, Guelmim; Mouna 1998; Rif (M’Diq), AP (Larache), HA (Tazzarin) – MISR

Desmometopa varipalpis Malloch, 1927

Ebejer et al. 2019, AA, 6 km N of Errachidia (1010 m)

Leptometopa Becker, 1903

Leptometopa rufifrons Becker, 1903

Ebejer et al. 2019, AA, Merzouga (714 m); AA (Merzouga) – MISR

Madiza Fallén, 1810

Madiza glabra Fallén, 1820

Mouna 1998; Ebejer et al. 2019, HA, Anafgou (NPHAO, 2271 m)

Milichiinae

Milichiini

Milichia Meigen, 1830

Milichia albomaculata (Strobl, 1900)

Pont and Singh 1965; Mouna 1998; Brake 2000; Koçak and Kemal 2010; HA – NHMUK

Milichia speciosa Meigen, 1830

Séguy 1930a, HA, Arround (Skoutana)

Milichiella Giglio-Tos, 1895

Milichiella lacteipennis (Loew, 1866)

Brake 2000; Raspi et al. 2009; Rif (M’Diq farm) – MISR

Phyllomyzinae

Phyllomyzini

Phyllomyza Fallén, 1810

Phyllomyza sp. aff. equitans (Hendel, 1919)

Ebejer et al. 2019, AA, Oued Ziz (12 km S of Rissani, 737 m)

Acknowledgement

We gratefully acknowledge the cooperation of Martin J. Ebejer who contributed to the revision of this family.

Sphaeroceroidea

CHYROMYIDAE

K. Kettani, M.J. Ebejer

Number of species: 22. Expected: 30

Faunistic knowledge of the family in Morocco: good

Aphaniosominae

Aphaniosoma Becker, 1903

Aphaniosoma approximatum Becker, 1903

Ebejer 2018, SA, Oued Ougni (0.6 km N of Akka, N’Aït Sidi and 1.8 km NW Tissint, 582 m)

Aphaniosoma claridgei Ebejer, 1995

Ebejer 2016b, Rif, Smir lagoon

Aphaniosoma collini Lyneborg, 1973

Ebejer 2016b, AP, Larache (5 m)

Aphaniosoma forcipatum Ebejer, 1998

Ebejer 2016b, AA, Lac de Tiffert (4 km W of Merzouga, 702 m), Erfoud (30 km N, 894 m), Ziz river (14 km E of Rich, 1278 m)

Aphaniosoma gatti Ebejer, 2016

Ebejer 2016b, AP, El Jadida, Azemmour; AP (El Jadida Azemmour) – MISR

Aphaniosoma melitense Ebejer, 1993

Andrade and Almeida 2010, Rif, Mediterranean coast; Ebejer 2016b, AP, Larache (2 m)

Aphaniosoma nigricauda Ebejer, 1998

Ebejer 2016b, AA, Lac de Tiffert (4 km W of Merzouga, 702 m), Merzouga (714 m), Ziz river (10 km S of Errachidia, 1008 m; 13 km N of Erfoud, 800 m)

Aphaniosoma nigripes Ebejer, 2016

Ebejer 2016b, AA, Ziz river (9.5 km SE of Rich, 1285 m), Erfoud (30 km N, 894 m; AA (Ziz river) – MISR

Aphaniosoma nigrum Ebejer, 1998

Ebejer 2016b, AA, Ziz river (Errachidia, 1052 m; 9.5 km SE of Rich, 1285 m), Erfoud (30 km N, 894 m); AA (Ziz river) – MISR

Aphaniosoma nitidum Ebejer, 2016

Ebejer 2016b, AA, Ziz river (30 km N of Erfoud, 894 m)

Aphaniosoma propinquans Collin, 1949

Ebejer 2016b, Rif, Smir lagoon, Oued Aliane (Ksar Sghir), AP, Larache (2 m); Rif (Oued Aliane) – MISR

Aphaniosoma proximum Ebejer, 1998

Ebejer 2016b, AP, Larache (5 m), Sidi Smail (El Jadida), Azemmour, HA, Oued Tensift

Aphaniosoma quadrinotatum (Becker, 1904)

Ebejer 2018, AA, S of village Sidi R’bat (10 m)

Aphaniosoma rufum Frey, 1935

Ebejer 2018, AA, S of village Sidi R’bat (10 m)

Aphaniosoma soror Ebejer, 2016

Ebejer 2016b, AA, Errachidia (14 km E of Rich, 1278 m)

Aphaniosoma trisetum Ebejer, 2016

Ebejer 2016b, AA, Ziz river (9.5 km SE of Rich, 1285 m, 13 km N of Erfoud, 800 m, Errachidia, 1052 m), Merzouga (714 m); AA (Ziz river) – MISR

Aphaniosoma zizense Ebejer, 2016

Ebejer 2016b, AA, Ziz river (Errachidia, 1052 m; 9.5 km SE of Rich, 1285 m; 10 km S of Errachidia, 1008 m); AA (Ziz river) – MISR

Chyromyinae

Chyromya Robineau-Desvoidy, 1830

Chyromya robusta Hendel, 1931

Ebejer 2018, AA, S of village Sidi R’bat (10 m)

Gymnochiromyia Hendel, 1933

Gymnochiromyia flavella (Zetterstedt, 1848)

Ebejer 2016b, Rif, Ksar El Kebir (13 m)

Gymnochiromyia homobifida Carles-Tolrá, 2001

Ebejer 2016b, Rif, maison forestière de Talassemtane (NPT)

Gymnochiromyia inermis (Collin, 1933)

Ebejer 1998b; Ebejer 2016b

Gymnochiromyia mihalyii Soós, 1979

Ebejer 2016b, Rif, Oued Siflaou (281 m), Aïn Tissemlal (Azilane, 1255 m), AP, Larache (5 m)

Gymnochiromyia tschirnhausi Ebejer, 2018

Ebejer 2018, AA, S of village Sidi R’bat (10 m)

Gymnochiromyia zernyi (Czerny, 1929)

Ebejer 1998b; Ebejer 2016b

HELEOMYZIDAE

K. Kettani, A.J. Woźnica

Number of species: 19. Expected: 30

Faunistic knowledge of the family in Morocco: moderate

Heleomyzinae

Gymnomus Loew, 1863

Gymnomus atlasicus Woźnica, 2011

Woźnica 2011, HA, Tazzeka

Gymnomus caesius (Meigen, 1830)

Ebejer et al. 2019, Rif, Jebel Lakraâ (Talassemtane, 1541 m)

Neoleria Malloch, 1919

Neoleria sp.*

Rif

Heteromyzinae

Tephrochlamys Loew, 1862

Tephrochlamys rufiventris (Meigen, 1830)*

AA

Suilliinae

Suillia Robineau-Desvoidy, 1830

Suillia bicolor (Zetterstedt, 1838)*

Rif

Suillia bistrigata (Meigen, 1830)

Ebejer et al. 2019, Rif, Moulay Abdelsalam (Bouhachem, 1098 m), Jebel Lakraâ (Talassemtane, 1541 m), MA, 6 km S of Azrou (1610 m), 20 km S of Azrou (1720 m)

Suillia humilis (Meigen, 1830)

= Helomyza similis Meigen, in Mouna 1998: 85

Mouna 1998; AP (Maâmora) – MISR

Suillia notata (Meigen, 1830)

Ebejer et al. 2019, Rif, Moulay Abdelsalam (Bouhachem, 1180 m), Jebel Lakraâ (Talassemtane, 1541 m), Oued Kbir (Béni Ratene, 157 m)

Suillia oxyphora (Mik, 1900)*

Rif

Suillia pallida (Fallén, 1820)

Séguy 1941d, HA, Tizi-n’Test (2200 m); Mouna 1998

Suillia tuberiperda (Rondani, 1876)

Ebejer et al. 2019, MA, 6 km S of Azrou (1610 m)

Suillia variegata (Loew, 1862)

= Helomyza variegata Loew, in Becker and Stein 1913: 93

Becker and Stein 1913, Rif, Tanger; Séguy 1953a, AP, Sidi Yahia du Gharb; Mouna 1998

Trixoscelidinae

Trixoscelis Rondani, 1856

Trixoscelis approximata (Loew, 1865)

Cassar et al. 2008, Rif, Smir lagoon; Rif, Tétouan

Trixoscelis baliogastra (Černý, 1909)

Hackman 1970

Trixoscelis canescens (Loew, 1865)

Ebejer et al. 2019, Rif, Aïn Tissemlal (Azilane, 1255 m)

Trixoscelis curvata Carles-Tolrá, 1993

Ebejer et al. 2019, AP, Larache (5 m)

Trixoscelis laeta (Becker, 1907)

Becker 1907; Séguy 1941d, AA, Agadir; Hackman 1970; Mouna 1998; Cassar et al. 2008, Rif, Smir lagoon

Trixoscelis mendizabali Hackman, 1970

Hackman 1970, AA, Aït Melloul (Oued Souss)

Trixoscelis pedestris (Loew, 1865)

Ebejer et al. 2019, Rif, Aïn Jdioui (Tahaddart, 76 m)

New records for Morocco

Neoleria sp.

Rif: Tétouan, Jebel Bouhachem (PNPB), Amsemlil, 35.26234°N, 5.43341°W, 1067 m, 11.xii.15, forest (Pinus pinaster), K. Kettani leg. (1♀), A.J. Woźnica det.

Tephrochlamys rufiventris (Meigen, 1830)

Rif: Tétouan, Jebel Bouhachem (PNPB), Remla, 35.236865, -5.408025, 961 m, 22.iv.18, forest (Pinus pinaster), K. Kettani leg. (1♀), A.J. Woźnica det.

Anti Atlas: Errachidia, 13 km E of Goulmima, 31°44.568N, 4°51.945W, dry stony steppe 1100 m, 3.v.2012, M.J. Ebejer leg. (1♂), A.J. Woźnica det.

Suillia bicolor (Zetterstedt, 1838)

Rif: Talassemtane, Jebel Lakraâ, 35°06.913N, 5°08.034W, 1541 m, 12.vi.2013, meadow in mixed forest, M.J. Ebejer leg. (1♀); Jebel Bouhachem, Taghzout, Adrou, 556 m, 35°22.39N, 5°32.28W, 25.iv.2015, M.J. Ebejer leg. (2♀♀), A.J. Woźnica det.

Suillia oxyphora (Mik, 1900)

Rif: Jebel Talassemtane, 35°07'N, 5°07'W, 1554 m, 13.iv.2009, Fir forest (Abies maroccana), A. Taheri leg. (1♂3♀♀); Azilane, Aïn Tissemlal, 35°11.67N, 5°15.20W, 1255 m, 4.vii.13–13.viii.2013, K. Kettani leg. (1♀), Fir forest (Abies maroccana), A.J. Woźnica det.

SPHAEROCERIDAE

K. Kettani, P. Gatt

Number of species: 67. Expected: 130

Faunistic knowledge of the family in Morocco: poor

Copromyzinae

Borborillus Duda, 1923

Borborillus vitripennis (Meigen, 1830)

Becker and Stein 1913, Rif, Tanger; Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Copromyza Fallén, 1810

Copromyza equina Fallén, 1820

Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Crumomyia Macquart, 1835

Crumomyia glabrifrons (Meigen, 1830)

Gatt et al. 2016, Rif, Tariouma (El Anasser, 1383 m), MA, Lac Aguelmane Sidi Ali (2050 m)

Lotophila Lioy, 1864

Lotophila atra (Meigen, 1830)

Gatt 2006, Rif, Tanger; Marshall et al. 2011

Norrbomia Papp, 1988

Norrbomia hispanica (Duda, 1923)

Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Norrbomia marginatis (Adams, 1905)

Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Norrbomia nilotica (Becker, 1903)

Roháček et al. 2001; Marshall et al. 2011

Norrbomia niveipennis (Duda, 1923)

Roháček et al. 2001; Marshall et al. 2011

Norrbomia somogyii (Papp, 1973)

Roháček et al. 2001; Marshall et al. 2011

Norrbomia sordida (Zetterstedt, 1847)

Gatt et al. 2016, Rif, Oued Guallet (Bni Boufrah, 946 m), AP, Larache, Loukous Marsh

Limosininae

Bifronsina Roháček, 1983

Bifronsina bifrons (Stenhammar, 1855)

Roháček et al. 2001; Marshall et al. 2011

Ceroptera Macquart, 1835

Ceroptera rufitarsis (Meigen, 1830)

= Limosina picta Becker, in Becker and Stein 1913: 94

Becker and Stein 1913, Rif, Tanger; Papp 1984a; Roháček et al. 2001, Rif, Tanger

Chaetopodella Duda, 1920

Chaetopodella scutellaris (Haliday, 1836)

= Leptocera scutellaris (Haliday), in Séguy 1941a: 32

Séguy 1941a, HA, Imi-n’Ouaka (1500 m)

Coproica Rondani, 1861

Coproica digitata (Duda, 1918)

Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Coproica ferruginata (Stenhammar, 1855)

Gatt 2006, Rif, Tétouan, Martil; Marshall et al. 2011

Coproica hirticula Collin, 1956

Gatt et al. 2016, Rif, Dardara (730 m), Oued Laou (30 m), Zarka waterfall (Yarghit, 135 m), Oued Ouara (Beni Zid, 440 m), Oued Jnane Azaghar (Bni Boufrah, 997 m), Oued Mhannech (125 m), Adrou (Taghzout, 556 m), Oued Jnane Niche (Jnane Niche, 27 m)

Coproica hirtula (Rondani, 1880)

Gatt et al. 2016, Rif, Zarka waterfall (Yarghit, 135 m), AA, 14 km N Errachidia (Errachidia, 1214 m)

Coproica lugubris (Haliday, 1836)

Gatt 2006, Rif, Tétouan, M’Diq, Smir, Oued Laou; Marshall et al. 2011

Coproica pusio (Zetterstedt. 1847)

Gatt et al. 2016, Rif, Tamrabete (Oued Laou, 203 m), Smir lagoon (5 m)

Coproica rohaceki Carles-Tolrá, 1990

Gatt et al. 2016, Rif, Oued Amsa (Amsa, 14 m), Oued Moulay Bouchta (Dar Akobaâ, 285 m)

Coproica rufifrons Hayashi, 1991

Gatt et al. 2016, Rif, Issaguen (1543 m)

Coproica vagans (Haliday, 1833)

Gatt et al. 2016, AA, 14 km N Errachidia (1214 m), AA, Oued Ziz (13 km N Erfoud, 800 m), SA, Merzouga (698 m)

Eulimosina ochripes (Meigen, 1830)

Gatt et al. 2016, Rif, Issaguen (1543 m), Ksar el Kebir (20 m), MA, Lac Aguelamane Afenourrir (30 km SW Azrou, 1760 m)

Leptocera Olivier, 1813

Leptocera caenosa (Rondani, 1880)

Gatt et al. 2016, Rif, Dardara (484 m)

Leptocera fontinalis (Fallén, 1826)

Gatt 2006, Rif, Tétouan, M’Diq, Smir; Marshall et al. 2011

Leptocera nigra Olivier, 1813

Munari 1993, MA, Azrou; Roháček et al. 2001; Marshall et al. 2011

Limosina Macquart, 1835

Limosina silvatica (Meigen, 1830)

Gatt et al. 2016, Rif, Issaguen (1543 m), Talassemtane (NPT, 1696 m), MA, 3,5 km S Azrou (1450 m), 17 km NW Zaida (Khénifra, 1878 m)

Minilimosina Roháček, 1983

Minilimosina (Svarciella) vitripennis (Zetterstedt, 1847)

Gatt et al. 2016, MA, 3,5 km S Azrou (1450 m)

Opacifrons Duda, 1918

Opacifrons coxata (Stenhammar, 1855)

Gatt 2006, Rif, Tétouan, M’Diq, Smir, Oued Laou; Marshall et al. 2011

Opacifrons maculifrons (Becker, 1907)

Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Opalimosina Roháček, 1983

Opalimosina (Opalimosina) mirabilis (Collin, 1902)

Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Paralimosina Papp, 1973

Paralimosina fucata (Rondani, 1880)

Gatt et al. 2016, Rif, 5 km W Dardara (730 m), Dardara (484 m), Oued Azla (Mokdassen Oulya, 186 m), Oued Talembote (Talembote, 320 m), Oued El Khizana (El Khizana, 980 m), Jebel Kelaâ (Talassemtane, 1554 m), Adrou (Taghzout, 556 m)

Phthitia Enderlein, 1938

Phthitia (Kimosina) sicana (Munari, 1988)

Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Phthitia (Kimosina) ciliata (Duda, 1918)

Gatt et al. 2016, SA, Bou Jarif (Goulimine)

Phthitia (Kimosina) plumosula (Rondani, 1880)

Gatt et al. 2016, Rif, Jebel Lakraâ (Talassemtane, 1541 m), Jebel Kelaâ (Talassemtane, 1554 m), Zaouiet and Habtiyène (Maggou, 1213 m), Oued Arozane (Bni Moussa, 317 m), Issaguen (1543 m), Zarka waterfall (Yarghit, 135 m), Taghramt (Bine El Ouidane, 276 m)

Phthitia (Kimosina) pteremoides (Papp, 1973)

Gatt et al. 2016, SA, Bou Jarif (Goulimine)

Poecilosomella Duda, 1925

Poecilosomella angulata (Thomson, 1869)

Gatt 2006, Rif, Tétouan, Martil; Marshall et al. 2011

Pseudocollinella Duda, 1924

Pseudocollinella jorlii (Carles-Tolrá, 1990)

Munari 1993, MA, Aguelmane; Roháček et al. 2001; Marshall et al. 2011

Pullimosina Roháček, 1983

Pullimosina (Pullimosina) heteroneura (Haliday, 1836)

Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Pullimosina (Pullimosina) zayensis Marshall, 1986

Roháček et al. 2001; Marshall et al. 2011

Puncticorpus Duda, 1918

Puncticorpus lusitanicum (Richards, 1963)

Papp 1984a; Roháček et al. 2001

Rachispoda Lioy, 1864

Rachispoda acrosticalis (Becker, 1903)

Gatt 2006, Rif, Tétouan, M’Diq, Smir; Marshall et al. 2011

Rachispoda brevior (Roháček, 1991)

Roháček 1991, AP, Oued Bou-Regreg; Munari 1993; Roháček et al. 2001; Marshall et al. 2011

Rachispoda duodecimseta (Papp, 1973)

Roháček 1991, EM, Guercif-Oued Moulouya; Munari 1993; Roháček et al. 2001; Marshall et al. 2011

Rachispoda fuscipennis (Haliday, 1833)

Roháček 1991, EM, Oued Bou-Regreg; Munari 1993; Roháček et al. 2001; Marshall et al. 2011

Rachispoda gel (Papp, 1978)

Gatt 2006, Rif, Tétouan, M’Diq, Smir; Marshall et al. 2011

Rachispoda kabuli (Papp, 1978)

Roháček 1991, MA, Taza, Oued Fès; Munari 1993; Roháček et al. 2001; Marshall et al. 2011

Rachispoda lagura (Roháček, 1991)

Gatt et al. 2016, Rif, Ksar el Kebir (20 m), AA, Oued Ziz (12 km S Rissani, 737 m), Oued Ziz (13 km N Erfoud, 800 m), Oued Ziz (30 km N Erfoud, 894 m)

Rachispoda lutosoidea (Duda, 1938)

Roháček 1991, MA, Azrou, Aguelmane, Oued Sebou; Munari 1993; Roháček et al. 2001; Marshall et al. 2011

Rachispoda modesta (Duda, 1924)

Gatt 2006, Rif, Tétouan, M’Diq, Smir; Marshall et al. 2011

Rachispoda uniseta (Roháček, 1991)

Roháček 1991, MA, Taza, Oued Fès; Munari 1993; Roháček et al. 2001; Marshall et al. 2011; Vaňhara and Rozkošný 1997

Rachispoda varicornis (Strobl, 1900)

Roháček 1991, AP, Oued Bou-Regreg, MA, Azrou; Munari 1993, MA, Oued Sebou; Roháček et al. 2001; Marshall et al. 2011

Spelobia Spuler, 1924

Spelobia baezi (Papp, 1977)

Roháček et al. 2001; Marshall et al. 2011

Spelobia clunipes (Meigen, 1830)

Gatt et al. 2016, Rif, 5 km W Dardara (730 m), Martil (beach and dunes), Oued Mhannech (18 m), Oued Afertane (Afertane, 56 m), Maggou waterfall (Maggou, 786 m), Onsar Akboul (NPB, 1315 m), Oued Ametrasse (Ametrasse, 841 m), Oued Tahaddart (Tahaddart, 87 m), Issaguen (1543 m), Lemtahane (PNPB, 1088 m)

Spelobia hungarica (Villeneuve, 1917)

Gatt 2006, Rif, Tétouan, Martil; Marshall et al. 2011

Spelobia quaesita Roháček, 1983

Roháček et al. 2001; Marshall et al. 2011

Spinilimosina Roháček, 1983

Spinilimosina brevicostata (Duda, 1918)

Roháček et al. 2001; Marshall et al. 2011

Telomerina Roháček, 1983

Telomerina pseudoleucoptera (Duda, 1924)

Gatt et al. 2016, Rif, Maggou waterfall (Maggou, 786 m)

Thoracochaeta Duda, 1918

Thoracochaeta brachystoma (Stenhammar, 1855)

Gatt 2006; Cassar et al. 2008, Rif, Tétouan, M’Diq, Smir; Marshall et al. 2011

Thoracochaeta erectiseta Carles-Tolrá, 1994

Gatt 2006, Rif, Tétouan, M’Diq, Smir, Oued Laou; Marshall et al. 2011

Trachyopella Duda, 1918

Trachyopella (Trachyopella) coprina (Duda, 1918)

Gatt et al. 2016, Rif, Oued Jnane Niche (Jnane Niche, 27 m)

Trachyopella (Trachyopella) melania (Haliday, 1836)

Gatt et al. 2016, Rif, Oued Nwawel (Azla, 57 m), M’Diq (5 m)

Sphaerocerinae

Ischiolepta Lioy, 1864

Ischiolepta pusilla (Fallén, 1820)

Gatt et al. 2016, Rif, Oued Guallet (Bni Boufrah, 946 m), Aïn Tissemlal (Azilane, 1255 m)

Ischiolepta vaporariorum (Haliday, 1836)

Gatt 2006, Rif, Tétouan, Oued Laou; Marshall et al. 2011

Lotobia Lioy, 1864

Lotobia africana (Becker, 1907)

Roháček et al. 2001; Marshall et al. 2011

Lotobia pallidiventris (Meigen, 1830)

Gatt 2006, Rif, Tétouan, M’Diq, Oued Laou; Marshall et al. 2011

Sphaerocera Latreille, 1804

Sphaerocera curvipes Latreille, 1805

Roháček et al. 2001; Marshall et al. 2011

Ephydroidea

BRAULIDAE

K. Kettani

Number of species: 1. Expected: 1

Faunistic knowledge of the family in Morocco: good

Braulinae

Braula Nitzsch, 1818

Braula coeca Nitzsch, 1818

Séguy 1930a; Mouna 1998; Howard et al. 2000

CAMILLIDAE

K. Kettani, M.J. Ebejer

Number of species: 4. Expected: 5

Faunistic knowledge of the family in Morocco: poor

Camilla Haliday in Curtis, 1837

Camilla acutipennis (Loew, 1865)

Ebejer et al. 2019, Rif, Aïn Jdioui (Tahaddart, 76 m), Chrabkha pond (Al Manzla, 58 m)

Camilla flavicauda Duda, 1922

Mouna 1998; Pârvu et al. 2006, AP, Cap Sim; Popescu-Mirceni 2011

Camilla glabra (Fallén, 1823)

Séguy 1941d, HA, Tizi-n’Test; Mouna 1998

Camilla pruinosa Duda, 1934

Ebejer et al. 2019, AP, Larache (5 m)

CRYPTOCHETIDAE

K. Kettani, E.P. Nartshuk

Number of species: 1. Expected: 2

Faunistic knowledge of the family in Morocco: poor

Cryptochetum Rondani, 1875

Cryptochetum (as Cryptochaetum) mimeuri Séguy, 1953

Séguy 1953c, MA, Ifrane; Nartshuk 1979, MA, Ifrane

DIASTATIDAE

K. Kettani, M.J. Ebejer

Number of species: 2. Expected: 3

Faunistic knowledge of the family in Morocco: poor

Campichoetinae

Campichoeta Macquart, 1835

Campichoeta obscuripennis (Meigen, 1830)

Ebejer et al. 2019, HA, Lalla Takrkoust (628 m)

Diastatinae

Diastata Meigen, 1830

Diastata adusta Meigen, 1830

= Diastata unipunctata Zetterstedt, 1847

Pârvu et al. 2006, AP, Merja Zerga; Popescu-Mirceni 2011

DROSOPHILIDAE

K. Kettani, G. Bächli

Number of species: 26. Expected: 30

Faunistic knowledge of the family in Morocco: good

Drosophilinae

Drosophilini

Drosophila Fallén, 1823

Drosophila (Drosophila) busckii Coquillett, 1901

= Drosophila rubrostriata Becker, 1908, in Séguy 1953a: 85

Séguy 1953a, AP, Rabat, Sidi Yahia du Gharb; Prevosti 1974, HA, Asni; Mouna 1998; Chakir et al. 2011, HA, Marrakech – MISR

Drosophila (Drosophila) buzzatii Patterson & Wheeler, 1942

Prevosti 1974, HA, Asni (Admin forest); Mouna 1998; Bächli 2015 (TaxoDros); Rif (Tanger) – ZSM

Drosophila (Drosophila) funebris (Fabricius, 1787)

Prevosti 1974, HA, Asni; Mouna 1998; AP (Maâmora) – MISR

Drosophila (Drosophila) hydei Sturtevant, 1921

Staiger and Gloor 1952, AP, Rabat; Gloor and Satiger 1954, AP, Rabat; Prevosti 1974, HA, Asni (Admin forest); Mouna 1998; Chakir et al. 2011, HA, Marrakech

Drosophila (Drosophila) immigrans Sturtevant, 1921

Mouna 1998; Chakir et al. 2011, HA, Marrakech; Bächli 2015 (TaxoDros); Rif (Tanger) – ZSM

Drosophila (Drosophila) kuntzei Duda, 1924

Mouna 1998; Prevosti 1974, AP, Essaouira, HA, Asni

Drosophila (Drosophila) mercatorum Patterson & Wheeler, 1942

Prevosti 1974, AA, Agadir (Admin forest near Agadir); Mouna 1998

Drosophila (Drosophila) phalerata Meigen, 1830

Prevosti 1974, AP, Essaouira, HA, Asni; Mouna 1998; Bächli 2015 (TaxoDros); Rif (Tanger), MA (Ifrane) – ZSM

Drosophila (Drosophila) repleta Wollaston, 1858

Séguy 1953a, AP, Rabat, MA, Fès

Drosophila (Drosophila) tsigana Burla & Gloor, 1952

Suwito et al. 2014, MA, Ifrane

Drosophila (Sophophora) ambigua Pomini, 1940

Mouna 1998; Bächli 2015 (TaxoDros); Rif (Tanger), MA (Ifrane) – ZSM

Drosophila (Sophophora) melanogaster Meigen, 1830

Medioni 1958; David and Bocquet 1973, HA, Marrakech, AA, Agadir, Ouarzazate, Taroudant, AA, Zagora; Prevosti 1974, AP, Essaouira, HA, Asni (Admin forest); Jousset and Plus 1975; Plus et al. 1975, HA, Marrakech, AA, Agadir, Ouarzazate, Taroudant, AA, Zagora; Prevosti et al. 1975, Rif, Tanger; Allemand and David 1976, AA, Ouarzazate; Ashburner and Lemeunier 1976, HA, Marrakech, AA, Agadir, Taroudant, AA, Zagora; Fleuriet 1976, AA, Zagora; Plus et al. 1976, AA, Ouarzazate, Zagora; Jousset 1976, AA, Ouarzazate; Plus and Scotti 1984, AA, Ouarzazate; Thomas-Orillard 1984, AA, Ouarzazate; Afonso et al. 1985, HA, Asni; Prevosti et al. 1985, AP, Essaouira, AA, Agadir; David et al. 1986, AP, Rabat, Casablanca; Ayala et al. 1989, Rif, Chefchaouen; Boulétreau et al. 1992, AA, Agadir; Costa et al. 1992, AP, Casablanca; Capy et al. 1993, AP, Casablanca, Rabat; Ritchie et al. 1994, AP, Casablanca; Mouna 1998; Bonnivard and Higuet 1999, AA, Agadir; Chakir et al. 2002, 2007, 2008, 2011, HA, Marrakech; Ayrinhac et al. 2004, HA, Marrakech; Catania et al. 2004, HA, Marrakech AA, Agadir; Rohmer et al. 2004, HA, Marrakech; Dieringer et al. 2005, HA, Marrakech, AA, Agadir; Yassin and Orgogozo 2013, HA, Marrakech; Bächli 2015 (TaxoDros) – HNHM, AP (Rabat), HA (Marrakech) –MISR, MA (Ifrane) – ZSM

Drosophila (Sophophora) simulans Sturtevant, 1919

Prevosti 1974, AP, Essaouira, HA, Asni (Admin forest); Baba-Aissa et al. 1988, AP, Rabat; Nigro 1988, AP, Larache; Capy et al. 1990, 1992, 1993, AP, Rabat, MA, Béni Mellal AA, Agadir; Chakrani et al. 1993, AA, Agadir; Mouna 1998; Charlat et al. 2003, AA, Agadir; Biémont et al. 2003, HA, Marrakech; Chakir et al. 2002, 2007, 2008, 2011, HA, Marrakech; Nardon et al. 2005, HA, Marrakech; Yassin and Orgogozo 2013, HA, Marrakech; Bächli 2015 (TaxoDros); AA (Agadir) – ZSM

Drosophila (Sophophora) subobscura Collin, 1936

Götz 1965, Rif, Tanger; Prevosti 1971a, Rif, Tanger; Prevosti 1971b, AP, Essaouira, HA, Asni, AA, Ait-Melloul; Prevosti 1971c; Gonzalez-Duarte et al. 1973, AP, Essaouira, HA, Asni AA, Agadir; Prevosti 1974, AP, Essaouira, HA, Asni (Admin forest); Prevosti et al. 1975, AA, Agadir; Duarte 1976, AP, Essaouira, HA, Asni AA, Agadir; Gonzalez 1976, AP, Essaouira, HA, Asni, AA, Agadir; Prevosti 1978, Rif, Tanger, AP, Essaouira, HA, Asni AA, Agadir; Krimbas and Loukas 1980, AA, Agadir; Cabrera et al. 1983, Rif, Chefchaouen; Larruga et al. 1983, Rif, Chefchaouen; Prevosti et al. 1985, AA, Agadir; Pascual et al. 1986, Rif, Chefchaouen; Latorre et al. 1986, Rif, Chefchaouen; Constanti et al. 1986, Rif, Chefchaouen; Ayala et al. 1989, Rif, Chefchaouen; Afonso et al. 1990, Rif, Chefchaouen; Pascual et al. 1990, Rif, Chefchaouen; Paricio et al. 1991, Rif, Chefchaouen; Menozzi and Krimbas 1992, AA, Agadir; Latorre et al. 1992, Rif, Chefchaouen; Fain et al. 1993, AA, Agadir; Alberola and Frutos 1993; Alberola and Frutos 1996; Pinto et al. 1997, HA, Asni, Marrakech AA, Agadir; Mouna 1998; David et al. 2003, HA, Marrakech; Brehm et al. 2004, AA, Agadir; Nardon et al. 2005, HA, Marrakech; Bächli 2015 (TaxoDros); MA (Azrou) – NHMD

Drosophila (Sophophora) suzukii (Matsumura, 1931)

Landolt et al. 2012, Rif, north-eastern Morocco

Hirtodrosophila Duda, 1924

Hirtodrosophila cameraria (Haliday, 1833)

Ebejer et al. 2019, Rif, Aïn Ras el Ma, ruisseau maison forestière (Talassemtane)

Lordiphosa Basden, 1961

Lordiphosa andalusiaca (Strobl, 1906)

= Lordiphosa forcipata (Collin, 1952), in Hackman 1960: 102

Hackman 1960; Bächli 2015 (TaxoDros); AP (Rabat) – NHMD

Scaptomyza Hardy, 1849

Scaptomyza adusta (Loew, 1862)

Ebejer et al. 2019, Rif, Dardara (730 m), AP, Loukous marsh (2 m)

Scaptomyza flava (Fallén, 1823)

= Scaptomyza flaveola (Meigen, 1830), in Kozlowsky and Rungs 1932: 66

Kozlowsky and Rungs 1932, AP, Rabat

Scaptomyza graminum (Fallén, 1823)

Maarouf 2003, HA, Chaouia; Bächli 2015 (TaxoDros); HA (Asni, Tinerhir) – NHMD

Scaptomyza (Parascaptomyza) pallida (Zetterstedt, 1847)

Ibn Jilali 1988 (agricultural areas); Maarouf 2003, HA, Chaouia; Bächli 2015 (TaxoDros); AP (Essaouira) MHNNR, MA (Azrou) – NHMD

Scaptodrosophila Duda, 1923

Scaptodrosophila rufifrons (Loew, 1873)

Mouna 1998; Bächli 2015 (TaxoDros); Rif (Tanger) – ZSM

Zaprionus Coquillett, 1901

Zaprionus indianus Gupta, 1970

Yassin and David 2010

Steganinae

Gitonini

Gitona Meigen, 1830

Gitona microchaeta Séguy, 1941

Séguy 1941d, AA, Agadir; Bächli 1982, AA, Agadir; Bächli and Rocha Pité 1984, AA, Agadir; Mouna 1998

Phortica Fallén, 1823

Phortica variegata (Fallén, 1823)

Ebejer et al. 2019, Rif, Bab Berred (1433 m), Jebel Lakraâ (Talassemtane, 1541 m)

Steganini

Leucophenga Mik, 1886

Leucophenga maculata (Dufour, 1839)

Séguy 1934b, AP, Port-Liautey (Maâmora)

EPHYDRIDAE

K. Kettani, T. Zatwarnicki

Number of species: 117. Expected: 140

Faunistic knowledge of the family in Morocco: good

Discomyzinae

Discomyzini

Actocetor Becker, 1903

Actocetor indicus (Wiedemann, 1824)

= Actocetor margaritatus (Wiedemann, 1830), in Séguy 1934b: 162, 1953a: 86

Séguy 1934b, Rif, Béni Aross; Séguy 1953a, SA, Tindouf

Discomyza Meigen, 1830

Discomyza incurva (Fallén, 1823)

= Discomyza italica Séguy, 1929, in Vitte 1991: 3; Dakki 1997: 63

Cresson 1939; Vitte 1991, AP, Atlantic coast and Plains; Dakki 1997

Psilopini

Clanoneurum Becker, 1903

Clanoneurum cimiciforme (Haliday, 1855)

Séguy 1941d, AA, Agadir; Dahl 1964; Vitte 1991, AP, Rabat; Dakki 1997

Diasemocera Bezzi, 1895

Diasemocera aequalipes (Becker, 1907)

= Psilopa aequalipes (Becker, 1907), in Ebejer et al. 2019: 147

Zatwarnicki 2018; Ebejer et al. 2019, AA, Lac Tiffert (4 km W of Merzouga, 702 m), Ziz river (13 km N of Erfoud, 800 m)

Diasemocera biskrae (Becker, 1907)

= Psilopa biskrae (Becker, 1907), in Vitte 1991: 32

Vitte 1991, AP, M’Diq; Dakki 1997

Diasemocera composita (Becker, 1903)

= Psilopa composita (Becker, 1903), in Vitte 1991: 32, Dakki 1997: 63

Vitte 1991, AP, Rabat; Dakki 1997

Diasemocera fratella (Becker, 1903)

= Psilopa fratella (Becker, 1903), in Ebejer et al. 2019: 147

Ebejer et al. 2019, AA, Errachidia (1 km N of Tarda, 1023 m), AA, Ziz river (13 km N of Erfoud, 800 m)

Diasemocera glabricula (Fallén, 1813)

= Psilopa nigritella Stenhammar, 1844, in Vitte 1988: 394; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif; Dakki 1997

Diasemocera leucostoma (Meigen, 1830)

= Psilopa leucostoma (Meigen, 1830), in Séguy 1941d: 18

Séguy 1941d, AA, Agadir

Diasemocera maritima (Perris, 1847)

= Psilopa maritima (Perris, 1847), in Cassar et al. 2008: 25

Cassar et al. 2008, Rif, Laou Basin

Diasemocera nana (Loew, 1860)

= Psilopa nana Loew, 1860, in Vitte 1991: 32, Dakki 1997: 63

Vitte 1991, Rif, AP, Atlantic coast; Dakki 1997

Diasemocera rufithorax (Becker, 1903)

= Psilopa rufithorax (Becker, 1903), in Ebejer et al. 2019: 147

Ebejer et al. 2019, AA, Merzouga (714 m)

Psilopa Fallén, 1823

Psilopa clara (Wollaston, 1858)

= Psilopa rutilans Canzoneri & Meneghini, 1972, in Cassar et al. 2005: 69

Cassar et al. 2005, Rif, Smir lagoon; Zatwarnicki 2018, AP, Larache (Lower Loukous), Safi, AA, Sidi Moussa, D’Agion (0–50 m)

Psilopa meneghinii Canzoneri, 1986

Vitte 1991, AP, Atlantic coast; Dakki 1997

Psilopa compta (Meigen, 1830)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997

Psilopa nilotica (Becker, 1903)

Ebejer et al. 2019, AA, Ziz river (10 km S of Errachidia, 1008 m), 1 km N of Tarda (Errachidia, 1023 m), Merzouga (714 m), 2 km N Erfoud (818 m)

Psilopa nitidula (Fallén, 1813)

Becker and Stein 1913, Rif, Tanger; Séguy 1941a, HA, Toubkal; Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif; Zatwarnicki 1991, Rif, Tanger, Tétouan, MA, Ifrane; Dakki 1997

Psilopa obscuripes Loew, 1860

Ebejer et al. 2019, Rif, Oued Azla (near bridge, 80 m), AP, Larache (5 m), Lower Loukous saltmarsh (2 m), MA, Khénifra (28 km S of Timahdit, 2100 m)

Psilopa polita (Macquart, 1835)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif; Dakki 1997

Risini

Achaetorisa Papp, 1980

Achaetorisa brevicornis Papp, 1980

Papp 1980, HA, Ouirgane

Ephydrinae

Dagini

Brachydeutera Leow, 1862

Brachydeutera meridionalis (Rondani, 1856)

= Brachydeutera ibari Ninomyia, 1929, in Ebejer et al. 2019: 147

Ebejer et al. 2019, Rif, Oued Martil (Taboula, 14 m)

Parydrini

Parydra Stenhammar, 1844

Parydra (Chaetoapnaea) fossarum (Haliday, 1833)

Séguy 1930a, AP, Rabat, MA, Meknès, HA, Marrakech; Vitte 1991; Dakki 1997; Vitte 1988, MA, lakes of Middle Atlas; MA (Jebel Khazzane) – MISR

Parydra (Chaetoapnaea) hecate (Haliday, 1833)

= Napaea hecate (Haliday), in Vaillant 1956b: 244

Vaillant 1956b, HA, Imi-N’Ifri

Parydra (Chaetoapnaea) quadripunctata (Meigen, 1830)

Pârvu et al. 2006, AP, Merja Zerga; Pârvu and Zaharia 2007; Popescu-Mirceni 2011, AP, Merja Zerga

Parydra (Paranapaea) pubera Loew, 1860

Dahl 1964, AA, Aït Melloul, Oued Souss; Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997

Parydra (Parydra) aquila (Fallén, 1813)

Vitte 1991, Rif, Ouezzane, Ketama; Dakki 1997

Parydra (Parydra) coarctata (Fallén, 1813)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif, Ksar el Kbir; Dakki 1997

Parydra (Parydra) cognata Loew, 1860

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997

Parydra (Parydra) flavitarsis Dahl, 1964

Vitte 1988, Rif, MA, lakes of Middle Atlas; Vitte 1991, Rif, MA, Fès; Dakki 1997; Gatt and Ebejer 2003; Dakki et al. 2003; Chillasse and Dakki 2004, Rif, MA; Dakki and Himmi 2008, MA, Oued Sebou

Parydra (Parydra) littoralis (Meigen, 1830)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif, Tétouan, Ketama; Dakki 1997

Parydra (Parydra) nigritarsis Strobl, 1893

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif, Ketama; Dakki 1997

Parydra (Parydra) nubecula Becker, 1896

Pârvu et al. 2006, AA, Lac Edehby, Ouarzazate; Popescu-Mirceni 2011, AP, Merja Zerga

Parydra (Parydra) quinquemaculata Becker, 1896

Vitte 1991, AP, Moulay Bousselham; Dakki 1997

Ephydrini

Ephydra Fallén, 1823

Ephydra bivittata Loew, 1860

Vitte 1991; Dakki 1997

Ephydra flavipes (Macquart, 1843)

Vitte 1991, AP, Atlantic coast; Dakki 1997

Ephydra glauca Meigen, 1830

Vitte 1991; Dakki 1997

Ephydra macellaria Egger, 1862

Dahl 1964, AA, Oued Souss; Vitte 1988, MA, lakes of Middle Atlas; Dakki 1997; Vitte 1991, AP, Atlantic coast

Halmopota Haliday, 1856

Halmopota mediterranea Loew, 1860

Dahl 1964, AA, Aït Melloul; Dakki 1997; Vitte 1991, Rif, Asilah, Chefchaouen

Paracoenia Cresson, 1935

Paracoenia fumosa (Stenhammar, 1844)

Vitte 1988, MA, lakes of Middle Atlas; Dakki 1997

Setacera Cresson, 1930

Setacera breviventris (Loew, 1860)

Vitte 1991, Rif, Ksar el Kbir; Dakki 1997

Scatellini

Haloscatella Mathis, 1979

Haloscatella dichaeta (Loew, 1860)

= Scatella dichaeta Loew, 1860, in Vitte 1988: 392, 1991: 26; Dakki 1997: 62

Vitte 1988, MA, Khemisset, Oued Beth, Dayat Aoua; Vitte 1991, AP, Moulay Bousselham; Dakki 1997

Lamproscatella Hendel, 1917

Lamproscatella sibilans (Haliday, 1833)

Ebejer et al. 2019, AA, 1 km N of Tarda (Errachidia, 1023 m)

Limnellia Malloch, 1925

Limnellia quadrata (Fallén, 1813)

Vitte 1991, Rif, Central Rif; Dakki 1997

Philotelma Becker, 1896

Philotelma nigripenne (Meigen, 1830)

= Scatella nigripennis (Meigen, 1830), in Vitte 1988: 391; Dakki 1997: 62

Vitte 1988, MA, lakes of Middle Atlas; Dakki 1997

Scatella Robineau-Desvoidy, 1830

Scatella (Neoscatella) subguttata (Meigen, 1830)

Vitte 1991, AP, Atlantic and Mediterranean coast, Smir lagoon; Dakki 1997; Pârvu et al. 2006, AP, Merja Zerga; Popescu-Mirceni 2011

Scatella (Scatella) ciliata Collin, 1930

Vitte 1991, AP, Moulay Bousselham, Asilah; Dakki 1997

Scatella (Scatella) lacustris (Meigen, 1830)

= Scatella (Scatella) tenuicosta Collin, 1930, in Ebejer et al. 2019: 147

Ebejer et al. 2019, AA, Ziz river (13 km N of Erfoud, 800 m)

Scatella (Scatella) lutosa (Haliday, 1833)

Vitte 1991, AP, Moulay Bousselham; Dakki 1997

Scatella (Scatella) obsoleta Loew, 1861

= Scatella callosicosta Bezzi, 1895, in Vitte 1988: 392, 1991: 26; Dakki 1997: 62

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif, M’Diq; Dakki 1997

Scatella (Scatella) paludum (Meigen, 1830)

Dahl 1964, AP, Oued Korifla; Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997

Scatella (Scatella) rufipes Strobl, 1906

= Scatella rubida Becker, 1907, in Olafsson 1991: 21; Vitte 1991: 26; Dakki 1997: 62

Olafsson 1991, EM, Figuig, Defilia; Vitte 1991, Rif, AP (Atlantic coast); Dakki 1997; Gatt and Ebejer 2003

Scatella (Scatella) stagnalis (Fallén, 1813)

Séguy 1941a, HA, Toubkal; Dahl 1964, AA, Aït Melloul, Oued Souss; Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997

Scatophila Becker, 1896

Scatophila caviceps (Stenhammar, 1844)

Vitte 1988, AP, Rabat, Temara, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997

Scatophila despecta (Haliday, 1839)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, MA, Khemisset, Oued Beth; Dakki 1997

Scatophila farinae Becker, 1903

Zatwarnicki 1987, HA, Vallée de l’Ait Mizane; Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, MA, Taounate; Dakki 1997; Gatt and Ebejer 2003

Scatophila modesta Becker, 1908

Vitte 1991, Rif, Tétouan; Dakki 1997

Scatophila unicornis Czerny, 1900

Ebejer et al. 2019, AA, 14 km E of Rich (Errachidia, 1278 m)

Gymnomyzinae

Discocerinini

Diclasiopa Hendel, 1917

Diclasiopa galactoptera (Becker, 1903)

= Discocerina galactoptera Becker, in Vitte 1988: 394; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Dakki 1997; Kirk-Spriggs and McGregor 2009

Diclasiopa lacteipennis (Loew, 1862)

= Discocerina lacteipennis Loew, 1862, in Vitte 1988: 394, 1991: 32; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Rabat; Vitte 1991, MA, Khemisset, Taounate; Dakki 1997

Diclasiopa niveipennis (Becker, 1896)

= Discocerina niveipennis (Becker, 1896), in Vitte 1988: 394, 1991: 32; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, AP, Moulay Bousselham; Dakki 1997

Discocerina Macquart, 1835

Discocerina obscurella (Fallén, 1813)

Vitte 1988, AP, MA, lakes of Middle Atlas; Vitte 1991, MA, Fès, Taounate; Mathis 1997; Dakki 1997; Wolff et al. 2016

Ditrichophora Cresson, 1924

Ditrichophora calceata (Meigen, 1830)

= Discocerina calceata (Meigen, 1830), in Vitte 1988: 394; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Dakki 1997

Ditrichophora mauritanica (Vitte, 1991)

= Discocerina mauritanica Vitte, 1991, in Vitte 1991: 33

Vitte 1991, Rif, MA, Azrou; Dakki 1997; Chillasse and Dakki 2004, Rif, MA; Dakki and Himmi 2008, MA, Oued Sebou

Gymnoclasiopa Hendel, 1930

Gymnoclasiopa plumosa (Fallén, 1823)

= Discocerina plumosa (Fallén, 1823), in Vitte 1988: 394, 1991: 33; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif; Dakki 1997

Gymnoclasiopa pulchella (Meigen, 1830)

= Discocerina pulchella (Meigen, 1830), in Vitte 1991: 33; Dakki 1997: 63

Vitte 1991, Rif, Ketama, Ouezzane; Dakki 1997

Hecamedoides Hendel, 1917

Hecamedoides glaucellus (Stenhammar, 1844)

= Discocerina glaucella (Stenhammar, 1844), in Vitte 1991: 33

Vitte 1991, Rif, AP

Polytrichophora Cresson, 1924

Polytrichophora duplosetosa (Becker, 1896)

AP (Rabat) – MISR

Gymnomyzini

Athyroglossa Loew, 1860

Athyroglossa (Athyroglossa) glabra (Meigen, 1830)

Vitte 1988, Rif, MA, lakes of Middle Atlas; Dakki 1997

Athyroglossa (Athyroglossa) nudiuscula Loew, 1860

Vitte 1991, Rif; Dakki 1997

Athyroglossa (Parathyroglossa) ordinata Becker, 1896

Vitte 1991, Rif; Vitte 1988, MA, lakes of Middle Atlas; Mathis and Zatwarnicki 1990, MA, Ifrane; Dakki 1997

Chlorichaeta Becker, 1922

Chlorichaeta albipennis (Loew, 1848)

Vitte 1991; Dakki 1997

Mosillus Latreille, 1804

Mosillus subsultans (Fabricius, 1794)

= Gymnopa subsultans Fabricius, in Séguy 1930a: 181

Séguy 1930a, MA, M’Rirt, HA, Imminen (Tachidirt); Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif; Mathis et al. 1993, MA, Ifrane (1650 m), maison forestière (cedar forest: 2700 m), Oued Jaffar (N of source, 0–1500 m), HA, Mikdane (Jebel Ayachi); Dakki 1997; Koçak and Kemal 2010

Hecamedini

Allotrichoma Becker, 1896

Allotrichoma laterale (Loew, 1860)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997

Allotrichoma leotoni Vitte, 1992

Vitte 1992, Rif, Ouezzane, Boured

Allotrichoma quadripectinatum (Becker, 1896)

= Allotrichoma bellicosum Giordani Soika, 1956, in Vitte 1991: 30; Dakki 1997: 63

Vitte 1991, Rif, AP; Dakki 1997

Allotrichoma simplex (Loew, 1861)

= Allotrichoma filiforme Becker, 1896, in Vitte 1988: 393, 1991: 30; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, MA, Khemisset, Oued Sebou; Dakki 1997

Elephantinosoma Becker, 1903

Elephantinosoma chnumi Becker, 1903

Gatt and Ebejer 2003; Kirk-Spriggs and McGregor 2009

Hecamede Haliday, 1837

Hecamede albicans (Meigen, 1830)

Vitte 1991, AP; Dakki 1997; Pârvu et al. 2006, AP, Merja Zerga; Cassar et al. 2008, Rif, Smir Lagoon; Popescu-Mirceni 2011

Lipochaetini

Glenanthe Haliday, 1839

Glenanthe ripicola (Haliday, 1839)

Vitte 1991, AP; Dakki 1997; Cassar et al. 2008, Rif, Laou Basin; Zatwarnicki and Mathis 2011, AA, Tarfaya – HNHM

Homalometopus Becker, 1903

Homalometopus sp.

Vitte 1991

Ochtherini

Ochthera Latreille, 1802

Ochthera manicata (Fabricius, 1794)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997

Ochthera pilimana Becker, 1903

Dakki 1997

Ochthera schembrii Rondani, 1847

= Ochthera mantispa Loew, 1847, in Vitte 1988: 394, 1991: 28; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif, AP; Dakki 1997

Hydrelliinae

Atissini

Asmeringa Becker, 1903

Asmeringa inermis Becker, 1903

Vitte 1991, AP, Rabat; Dakki 1997; Gatt and Ebejer 2003

Atissa Haliday, 1839

Atissa durrenbergensis Loew, 1864

Vitte 1991, AP; Dakki 1997

Atissa hepaticoloris Becker, 1903

Vitte 1991, AP; Dakki 1997; Gatt and Ebejer 2003

Atissa limosina Becker, 1896

Vitte 1991, AP, Rabat, MA, Fès; Dakki 1997

Atissa pygmaea (Haliday, 1839)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997; Pârvu et al. 2006, AA, Lac Edehby, Ouarzazate; Popescu-Mirceni 2011

Ptilomyia Coquillett, 1900

Ptilomyia angustigenis (Becker, 1926)

= Atissa angustigenis Becker, in Vitte 1988: 393, 1991: 30; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997; Gatt and Ebejer 2003

Dryxini

Dryxo Robineau-Desvoidy, 1830

Dryxo ornata (Macquart, 1843)

Mathis and Zatwarnicki 2002, AA, Aoulouz

Hydrelliini

Hydrellia Robineau-Desvoidy, 1830

Hydrellia albifrons (Fallén, 1813)

Vitte 1991, AP, Rif; Dakki 1997

Hydrellia argyrogenis Becker, 1896

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, AP, Atlantic coast and Plains; Dakki 1997

Hydrellia armata Canzoneri & Meneghini, 1976

Vitte 1991, Rif, Ksar el Kbir, MA, Fès; Dakki 1997; Dakki and Himmi 2008, MA, Oued Sebou

Hydrellia atlas Vitte, 1989

Vitte 1989, MA, Dayat Aoua; Dakki 1997; Dakki et al. 2003; Chilasse and Dakki 2004, MA

Hydrellia griseola (Fallén, 1813)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997; Wolff et al. 2016; Rif (Oued Laou) – MISR

Hydrellia maculiventris Becker, 1896

Vitte 1991, Rif, AP, Atlantic coast; Dakki 1997; Gatt and Ebejer 2003

Hydrellia maura Meigen, 1838

= Hydrellia modesta Loew, 1860, in Vitte 1988: 392, 1991: 29; Dakki 1997: 63

Vitte 1988, MA, lakes of Middle Atlas; Zatwarnicki 1988, MA, Ifrane, HA, Vallée de l’Aït Mizane; Vitte 1991; Dakki 1997

Hydrellia nigricans (Stenhammar, 1844)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif; Dakki 1997

Hydrellia obscura (Meigen, 1830)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif; Dakki 1997; Rif (Aïn Jdioui) – MISR

Hydrellia pubescens Becker, 1926

= Hydrellia nasturtii Collin, 1928, in Vitte 1991: 29; Dakki 1997: 63

Vitte 1991, MA, Fès; Dakki 1997; Gatt and Ebejer 2003

Hydrellia ranunculi (Haliday, 1838)

Vitte 1991, AP, Moulay Bousselham; Dakki 1997

Hydrellia rharbia Vitte, 1991

Vitte 1989, AP, Merja Halloufa (near Moulay Bousselham); Dakki 1997; Dakki and Himmi 2008, MA, Oued Sebou

Hydrellia subalbiceps Collin, 1966

Vitte 1991, Rif, Ketama; Vitte 1988, MA, lakes of Middle Atlas; Dakki 1997

Notiphilini

Notiphila Fallén, 1810

Notiphila (Notiphila) annulipes Stenhammar, 1844

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif; Dakki 1997

Notiphila (Notiphila) cinerea Fallén, 1830

Séguy 1930a, MA, Meknès; Séguy 1941a, HA, Imi-n’Ouaka; Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997; Pârvu et al. 2006, AP, Merja Zerga; Kirk-Spriggs and McGregor 2009; Rif (Talassemtane, Aïn Jdioui) – MISR

Notiphila (Notiphila) cogani Canzoneri & Meneghini, 1979

Vitte 1988, MA, lakes of Middle Atlas; Krivosheina 1998; Vitte 1991; Dakki 1997; Rif (Aïn Jdioui) – MISR

Notiphila (Notiphila) dorsata Stenhammar, 1844

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, AP, coastal lake areas; Dakki 1997

Notiphila (Notiphila) maculata Stenhammar, 1844

Vitte 1991, Rif, AP, coastal plains; Vitte 1988, MA, lakes of Middle Atlas; Dakki 1997

Notiphila (Notiphila) riparia Meigen, 1830

Krivosheina 1998, AA, Aït Melloul (Souss); Vitte 1988, MA, lakes of Middle Atlas and reedbeds; Vitte 1991; Dakki 1997

Notiphila (Notiphila) stagnicola (Robineau-Desvoidy, 1830)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991, Rif, AP, coastal plains; Dakki 1997; Pârvu et al. 2006, AA, Lac Edehby, Ouarzazate; Popescu-Mirceni 2011, AA, Ouarzazate

Ilytheinae

Hyadinini

Hyadina Haliday, 1837

Hyadina guttata (Fallén, 1813)

Vitte 1988, MA, lakes of Middle Atlas; Vitte 1991; Dakki 1997

Hyadina pollinosa Oldenberg, 1923

Vitte 1991, MA, Fès; Dakki 1997

Hyadina rufipes (Meigen, 1830)

= Hyadina nitida (Macquart, 1835), in Vitte 1991: 27; Dakki 1997: 63

Vitte 1991, AP, Moulay Bousselham; Dakki 1997

Nostima Coquillett, 1900

Nostima picta (Fallén, 1813)

Vitte 1991, AP; Vitte 1988, MA, lakes of Middle Atlas; Dakki 1997

Pelina Haliday, 1837

Pelina aenea (Fallén, 1813)

Vitte 1991, Rif, Ouezzane; Dakki 1997

Pelina subpunctata Becker, 1896

Ebejer et al. 2019, Rif, Jebel Lakraâ (Talassemtane, 1541 m)

Philygria Stenhammar, 1844

Philygria posticata (Meigen, 1830)

Ebejer et al. 2019, MA, Khénifra (17 km SW of Midelt, 1940 m)

Acknowledgements

We gratefully acknowledge the assistance and cooperation of Martin J. Ebejer who contributed to the revision of this family.

Hippoboscoidea

HIPPOBOSCIDAE

K. Kettani, B. Droz

Number of species: 17. Expected: 25

Faunistic knowledge of the family in Morocco: moderate

Hippoboscinae

Hippoboscini

Crataerina von Olfers, 1816

Crataerina acutipennis Austen, 1926

Mouna 1998: 85

Crataerina pallida (Latreille, 1811)

Maa 1969; Mouna 1998; AA (Tiznit) – MISR

Hippobosca Linnaeus, 1758

Hippobosca camelina Leach, 1817

= Hippobosca dromedarina Speiser, in Séguy 1930a: 184

Séguy 1930a; Bequaert 1939, AP, Rabat, EM, Taourit, HA; Séguy 1953a, AA, Zegdou, SA, Oued Agouidir; Mouna 1998; EM (Oued el Maa) – MISR

Hippobosca equina (Linnaeus, 1758)

Séguy 1930a, AP, Rabat, Settat, Mogador; Séguy 1953a, AA, Inzegane; Maa 1969; Bequaert 1939, MA, Aguelmane, HA, Ijoukak – MISR

Hippobosca fulva Austen, 1912

Bequaert 1939, EM, Taourirt (Ebner), Tendrara (Ebner); Maa 1963

Hippobosca longipennis Fabricius, 1805

= Hippobosca capensis Olfers, in Bequaert 1939: 78

Séguy 1930a, MA, Meknès; Bequaert 1939, HA, Marrakech; Maa 1963, 1969; Mouna 1998; AA (Agdz) – MISR

Hippobosca variegata Megerle, 1803

= Hippobosca maculata Leach, in Séguy 1930a: 184

Séguy 1930a; Moussiaux and Desmecht 2008, HA (south)

Icosta Speiser, 1905

Icosta minor (Bigot, 1858)

= Lynchia minor Bigot, in Mouna 1998: 85

Báez 1978; Mouna 1998

Ornithoica Rondani, 1878

Ornithoica turdi (Olivier in Latreille, 1811)

Maa 1966, EM, Figuig; Maa 1969; Mouna 1998; Droz and Haenni 2011

Ornithomyia Latreille, 1802

Ornithomyia avicularia (Linnaeus, 1758)

Séguy 1930a; Mouna 1998

Ornithomyia fringillina (Curtis, 1836)

Séguy 1930a, MA, Meknès; Mouna 1998

Ornithophila Rondani, 1879

Ornithophila gestroi (Rondani, 1878)

= Ornitheza gestroi Rondani, in Mouna 1998: 85

Mouna 1998; Pape and Thompson 2019

Ornithophila metallica (Schiner, 1864)

= Ornitheza metallica Schiner

Mouna 1998: 85

Pseudolynchia Bequaert, 1926

Pseudolynchia canariensis (Macquart, 1839)

= Pseudolynchia maura Bigot, in Séguy 1930a: 184

Séguy 1930a; Mouna 1998

Stenepteryx Leach, 1817

Stenepteryx hirundinis (Linnaeus, 1758)

= Crataerina hirundinis Linnaeus, in Mouna 1998: 85

Summer 1978, MA, Midelt; Mouna 1998

Lipopteninae

Lipoptena Nitzsch, 1818

Lipoptena capreoli Rondani, 1878

Mouna 1998: 85

Melophagus Latreille, 1802

Melophagus ovinus (Linnaeus, 1758)

= Melanophagus ovinus Linnaeus, in Mouna 1998: 85

Séguy 1930a; Maa 1969; Mouna 1998

NYCTERIBIIDAE

K. Kettani, G. Graciolli

Number of species: 8. Expected: 18

Faunistic knowledge of the family in Morocco: poor

Nycteribiinae

Basilia Miranda-Ribeiro, 1903

Basilia italica Theodor, 1954

Aellen 1955

Nycteribia Latreille, 1796

Nycteribia (Acrocholidia) vexata Westwood, 1835

Aellen 1952; Aellen 1955, AP, Grotte de Sidi Bou Knadel (hosts: Myotis oxygnathus Monticelli, 1885, Miniopterus shreibersii (Kuhl, 1817) and Rhinolophus mehelyi (Matschie, 1901)), MA, Grotte de Ras el Oued (host: Myotis oxygnathus and Miniopterus shreibersii; Aellen 1963; Theodor 1967, MA, Oued Mellah; Mouna 1998

Nycteribia (Nycteribia) latreillei (Leach, 1817)

Séguy 1930a, Rif, Samsa (Tétouan); Aellen 1952; Aellen 1955, AP, Grotte de Sidi Bou Knadel (host: Miniopterus shreibersii (Kuhl, 1817)), MA, Grotte de Ras el Oued (hosts: Myotis oxygnathus Monticelli, 1885 and Miniopterus shreibersii); Aellen 1963; Theodor 1967, AP, Mazagan (host: Myotis myotis (Bourhausen, 1797)); Mouna 1998

Nycteribia (Nycteribia) pedicularia Latreille, 1805

= Listropodia pedicularia Latreille, in Séguy 1930a: 185

Falcoz 1924, Rif, Caverne d’Hercule; Séguy 1930a, Rif, Caverne d’Hercule; Mouna 1998

Nycteribia (Nycteribia) schmidtlii Schiner, 1853

= Listropodia schmidli Schiner, in Séguy 1930a: 186

Falcoz 1924, Rif, Caverne de Samsa (near Tétouan); Séguy 1930a, Rif, Caverne de Samsa, SA; Aellen 1952; Aellen 1955, AP, Grotte de Sidi Bou Knadel (undetermined bat), MA, Grotte de Ras el Ma (host: Rhinolophus ferrumequinum (Schreber, 1774)), Grotte de Ras el Oued (hosts: Miniopterus shreibersii (Kuhl, 1817) and Myotis oxygnathus Monticelli, 1885); Aellen 1963; Theodor 1967 (host: Rhinolophus ferrumequinum); Mouna 1998

Penicillidia Kolenati, 1963

Penicillidia (Penicillidia) conspicua Speiser, 1901

Falcoz 1924, Rif, Caverne d’Hercule, Caverne de Samsa (near Tétouan); Séguy 1930a, Rif, Caverne d’Hercule, Caverne de Samsa; Aellen 1952; Aellen 1955, MA, Grotte de Ras el Oued, AP, Grotte de Sidi Bou Knadel; Aellen 1963; Mouna 1998; Koçak and Kemal 2010

Penicillidia (Penicillidia) dufouri (Westwood, 1835)

Falcoz 1924, Rif, Caverne d’Hercule; Séguy 1930a, Rif, Caverne d’Hercule; Aellen 1952; Aellen 1955, AP, Grotte de Sidi Bou Knadel, MA, Grotte de Ras el Oued, AA, Oulad Teima; Aellen 1963; Theodor 1967, AP, Mazagan (Myotis myotis (Bourhausen, 1797)); Mouna 1998; Koçak and Kemal 2010

Phthiridium Hermann, 1804

Phthiridium biarticulatum Hermann, 1804

= Stylidia biarticulata Herman, in Falcoz 1924: 310; Séguy 1930a: 185

Falcoz 1924, Rif, Caverne de Samsa (near Tétouan); Séguy 1930a, Rif, Caverne de Samsa; Aellen 1952; Aellen 1955, MA, Grotte de Ras el Ma (host: Rhinolophus ferrumequinum (Schreber, 1774)), AA, Oulad Teima (host: Rhinolophus ferrumequinum); Aellen 1963; Theodor 1967 (host: Rhinolophus ferrumequinum); Mouna 1998

STREBLIDAE

K. Kettani, G. Graciolli

Number of species: 2. Expected: 7

Faunistic knowledge of the family in Morocco: poor

Brachytarsininae

Brachytarsina Macquart, 1851

Brachytarsina flavipennis Macquart, 1851

= Nycteribosca kollari Frauenfeld, in Falcóz 1924: 226; Aellen 1955: 100

Falcóz 1924, Rif, caverne d’Hercule, caverne de Samsa, près Tétouan (host: Rhinolophus ferrumequinum (Schreber, 1774); Séguy 1930a, Rif, caverne d’Hercule, caverne de Samsa; Aellen 1955, AP, Grotte de Sidi Bou Knadel (hosts: Rhinolophus mehelyi (Matschie, 1901) and Myotis oxygnathus Monticelli, 1885); Mouna 1998; Koçak and Kemal 2010

Raymondia Frauenfeld, 1855

Raymondia huberi Frauenfeld, 1855

= Raymondia setosa Jobling, 1930

Beaucournu et al. 1985, AA, Assa (Bas Draa) (host: Asellia tridens (E. Geoffroy, 1813))

Muscoidea

ANTHOMYIIDAE

K. Kettani, D.M. Ackland

Number of species: 36. Expected: Many more, especially in the mountains

Faunistic knowledge of the family in Morocco: poor

Anthomyiinae

Adia Robineau-Desvoidy, 1830

Adia cinerella (Fallén, 1825)

= Chortophila cinerella Fallén, in Séguy 1930a: 162

= Hylemyia cinerella Fallén, in Séguy 1941d: 18

Séguy 1941d, AA, Agadir; Séguy 1930a; Mouna 1998; AP (Rabat), HA (Marrakech), AA (Tifnit (south of Agadir)) – MISR

Anthomyia Meigen, 1803

Anthomyia imbrida Rondani, 1866

Séguy 1930a, MA, Meknès; Mouna 1998

Anthomyia liturata (Robineau-Desvoidy, 1830)

= Hylemyia pullula Zetterstedt, in Séguy 1930a: 162

Séguy 1930a, MA, Ras el Ksar (1900 m), Tameghilt (1700–1800 m), Forêt Tiffert (2000–2200 m); Mouna 1998

Anthomyia quinquemaculata Macquart, 1839

Ebejer et al. 2019, Rif, Jebel Lakraâ (Talassemtane, 1541 m), MA, 3.5 km S of Azrou (1450 m)

Anthomyia pluvialis (Linnaeus, 1758)

Séguy 1929b; Séguy 1930a, MA, Meknès; Michelsen 1980, AP, Aïn Diab; Mouna 1998; Ackland 1987, 2001; Pârvu et al. 2006, AA, Foum Zguid (Tata); Popescu-Mirceni 2011 – MISR

Anthomyia procellaris Rondani, 1866

Séguy 1929b; Séguy 1930a, MA, Meknès, Berkane (1350–1400 m), Tlet n’Rhohr; Mouna 1998

Anthomyia tempestatum Wiedemann, 1830

Michelsen and Báez 1985, HA; Ackland 2001; Grabener 2017

Botanophila Lioy, 1864

Botanophila dissecta (Meigen, 1826)

Mouna 1998; MA (Meknès) – MISR

Botanophila varicolor (Meigen, 1826)

Ebejer et al. 2019, MA, Lac Aguelmane Sidi Ali (2052 m)

Delia Robineau-Desvoidy, 1830

Delia antiqua (Meigen, 1826)

Mouna 1998

Delia coarctata (Fallén, 1825)

Mouna 1998

Delia flavibasis (Stein, 1903)

= Hylemyia hordeacea Séguy, in Séguy 1941d: 18

Séguy 1934a, AP, Casablanca; Séguy 1936b, AP, Rabat; Séguy 1941d, AA, Taroudant; Mouna 1998; Ackland 2008

Delia flavogrisea (Ringdahl, 1926)52

Pârvu et al. 2006, AP, Merja Zerga; Pârvu and Zaharia 2007; Popescu-Mirceni 2011

Delia planipalpis (Stein, 1898)

= Chortophila pilipyga Villeneuve, in Mouna 1998: 85

Mouna 1998; AP (Mazagan) – MISR

Delia platura (Meigen, 1826)

= Chortophila cilicrura Rondani, in Séguy 1930a: 162

Séguy 1949a, SA, Goulimine; Mouna 1998; Singh et al. 2014; Grabener 2017

Delia radicum Linnaeus, 1758

= Chortophila brassicae Bouché, in Séguy 1934b: 162, Mouna 1998: 85

Séguy 1934b, AP, Rabat; Mouna 1998; Biron et al. 2000, AP, Rabat; Andreassen 2007 – MISR

Fucellia Robineau-Desvoidy, 1842

Fucellia maritima (Haliday, 1838)

Séguy 1930a, Rif, Agla near Cap Spartel (on Fucus); Cassar et al. 2008, Rif, Smir lagoon; Mouna 1998

Hylemya Robineau-Desvoidy, 1830

Hylemya vagans (Panzer, 1798)

Mouna 1998 – MISR

Leucophora Robineau-Desvoidy, 1830

Leucophora cinerea Robineau-Desvoidy, 1830

Ebejer et al. 2019, MA, 17 km SW of Midelt (Khénifra, 1940 m)

Leucophora dissimilis (Villeneuve, 1920)

Ebejer et al. 2019, MA, 17 km NW of Zaida (Khénifra, 1878 m)

Paregle Schnabl, 1911

Paregle audacula (Harris, 1780)

Pârvu and Zaharia 2007

Paregle pilipes (Stein, 1916)

Mouna 1998

Phorbia Robineau-Desvoidy, 1830

Phorbia fumigata (Meigen, 1826)

= Phorbia securis Tiensuu, in Maarouf et al. 1996: 17

Balachowsky and Mesnil 1935; Bleuton 1938; Jourdan 1938; Maarouf and Chemseddine 1995, AP, Chaouia, Doukkala, Abda; Maarouf and Chemseddine 1995, HA, Chaouia, Doukkala, Abda; Maarouf et al. 1996, AP, Safi, Settat, Sidi El Aydi, Jemaa Riah, Berrechid, Médiouna, Mohammédia, Bouznika, Skhirat, Kénitra, Sidi Allal Tazi, Souk Larbaa du Gharb, MA, Khernisset, Meknès, Douiyat, Fès, Sefrou, Annaceur, Oulad Saïd, El Aounate, Sidi Bennbur, Zemarnra, Chemmaïa, Jemaa, des Shaïm, Tlet Sidi Bouguedra, Khemisset Chaouïa, Béni Mellal, HA, Skhour Rehamna, Ben Guérir, Marrakech, Tamellalet, Kelaâ des Sraghna, Oulad Ayad, Afourér, Azilal, El Borouj, Guisser; Lhaloui et al. 1998

Phorbia sepia (Meigen, 1826)

Bleuton 1938, MA, Fès, Meknès, Taza; Jourdan 1938; Mouna 1998

Subhylemyia Ringdahl, 1933

Subhylemyia longula (Fallén, 1824)

Mouna 1998; AP (Cap Cantin) – MISR

Pegomyinae

Calythea Schnabl in Schnabl and Dziedzicki 1911

Calythea nigricans (Robineau-Desvoidy, 1830)

= Calythea albicincta Fallén, in Séguy 1930a: 161

Séguy 1930a, MA, Meknès, Aïn Leuh; Mouna 1998

Mycophaga Rondani, 1856

Mycophaga testacea (Gimmerthal, 1834)

Ebejer et al. 2019, Rif, Jebel Lakraâ (Talassemtane, 1541 m)

Pegomya Robineau-Desvoidy, 1830

Pegomya betae (Curtis, 1847)

Rungs 1962, AP; Mouna 1998

Pegomya bicolor (Wiedemann, 1817)

Séguy 1934b, AP, Rabat; Koçak and Kemal 2010; Pitkin et al. 2011

Pegomya hyoscyami (Panzer, 1809)

Rungs 1962, AP; Mouna 1998; AP (Rabat) – MISR

Pegomya rufina (Fallén, 1825)

Mouna 1998

Pegomya testacea (De Geer, 1776)

= Pegomya silacea Meigen, in Mouna 1998: 85

Séguy 1930a, MA, Forêt Tiffert (2000–2200 m); Mouna 1998

Pegomya solennis (Meigen, 1826)

= Pegomyia nigritarsis Fallén, in Séguy 1935a: 119

Séguy 1935a, AA, Oued Draa (Taffagount); Rungs 1962; Mouna 1998

Pegomya terminalis Rondani, 1866

Ebejer et al. 2019, Rif, Adrou (556 m), Jebel Lakraâ (Talassemtane, 1541 m)

Pegomya winthemi (Meigen, 1826)

Mouna 1998

Pegoplata Schnabl & Dziedzicki, 1911

Pegoplata annulata (Pandellé, 1899)

= Pegoplata virginea auctt, not Meigen

AP (Rabat) – MISR

FANNIIDAE

K. Kettani, A.C. Pont

Number of species: 10. Expected: 17

Faunistic knowledge of the family in Morocco: poor

Fannia Robineau-Desvoidy, 1830

Fannia canicularis (Linnaeus, 1761)

Becker and Stein 1913, Rif, Tanger; Charrier 1927; Séguy 1930a, HA, Tenfecht; Séguy 1932a; Séguy 1941a; Séguy 1941d, HA; Séguy 1953a, AP, Rabat; Pont 1986a; Mouna 1998; Pont pers. comm., MA, Azrou; AP (Dradek) – MISR

Fannia cothurnata (Loew, 1873)53

Mouna 1998

Fannia krimensis Ringdahl, 1934

Pont 1983, HA, Jebel Ayachi; Pont 1986a; Mouna 1998

Fannia lepida (Wiedemann, 1817)

Pont pers. comm.

Fannia leucosticta (Meigen, 1838)

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA; Pont 1986a; Mouna 1998

Fannia monilis (Haliday, 1838)

Ebejer et al. 2019, Rif, Jebel Lakraâ (Talassemtane, 1541 m), Dardara (484 m)

Fannia norvegica Ringdahl, 1934

Pont 1986a; Pont pers. comm., HA, Jebel Ayachi

Fannia pallidibasis Pont, 1983

Pont 1983, HA, Jebel Ayachi; Mouna 1998

Fannia scalaris (Fabricius, 1794)

Charrier 1927, Rif, Tanger; Séguy 1930a; Séguy 1941a, HA; Séguy 1953a, AP, Rabat; Pont 1986a; Mouna 1998; Rif (Tanger): Caverne d’Hercule (Pont pers. comm.) – MHNP

Fannia sociella (Zetterstedt, 1845)54

Mouna 1998

MUSCIDAE

K. Kettani, A.C. Pont

Number of species: 115. Expected: 140

Faunistic knowledge of the family in Morocco: moderate

Atherigoninae

Atherigona Rondani, 1856

Atherigona humeralis (Wiedemann, 1830)

Pont pers. comm., AP, Casablanca

Atherigona pulla (Wiedemann, 1830)

Pont 1986b; Pont pers. comm., AP, Larache, Rabat, HA, Asni, AA, Agadir, Taroudant

Atherigona soccata Rondani, 1871

= Atherigona varia (Meigen, 1826) (misidentifications of authors) in Séguy 1930a: 159

Bléton and Fieuzet 1943, MA, Fès; Séguy 1953a, AP, Rabat, Sidi Yahia du Gharb; Mouna 1998; Pont 1986b

Atherigona varia (Meigen, 1826)

= Atherigona quadripunctata Rossi, in Séguy 1949a: 159

Séguy 1930a, Rif, Tanger; Séguy 1941d; Séguy 1949a, AA, Akka, Alnif; Bléton and Fieuzet 1943, MA, Meknès, Gharb, Fouarat; Séguy 1953a, AP, Sidi Yahia du Gharb, Rabat; Pont 1986b; Mouna 1998; Pont pers. comm., Rif, Meloussa, Tanger, AP, Larache, Rabat, HA, Jebel Ayachi, AA, Akka

Atherigona (Acritochaeta) yorki Deeming, 1971

Pont 1986b, 1991a; Dike 1990; Pont pers. comm., AP, Rabat

Azeliinae

Azeliini

Azelia Robineau-Desvoidy, 1830

Azelia parva Rondani, 1866

Michelsen pers. comm., Rif, Ouezzane

Hydrotaea Robineau-Desvoidy, 1830

Hydrotaea aenescens (Wiedemann, 1830)

Morocco, first record 1989; Pont et al. 2007

Hydrotaea armipes (Fallén, 1825)

Pont 1986b; Pont pers. comm., MA, Azrou, HA, Jebel Ayachi

Hydrotaea capensis (Wiedemann, 1818)

= Ophyra anthrax (Meigen, 1826), in Séguy 1941d: 20

Séguy 1934b, AP, Chellah; Séguy 1941d, AA, Agadir; Pont 1986b; Hernández-Moreno and Peris 1989, Rif, Tanger; Mouna 1998; Turchetto et al. 2003; Pont pers. comm., HA, Jebel Ayachi; Rif (Environ de Tanger (Pont pers. comm.)) – MHNP

Hydrotaea cinerea Robineau-Desvoidy, 1830

Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Hydrotaea dentipes (Fabricius, 1805)

Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Hydrotaea floccosa Macquart, 1835

= Hydrotaea armipes (Fallén, 1825) (misidentifications of authors) in Charrier 1927: 620

Charrier 1927, Rif, Tanger; Pont 1986b; Pont pers. comm., MA, Azrou, HA, Jebel Ayachi

Hydrotaea glabricula (Fallén, 1825)

Pont 1986b; Pont pers. comm., AP, Forêt Maâmora

Hydrotaea ignava (Harris, 1780)

= Ophyra leucostoma (Wiedemann, 1817), in Séguy 1953a: 87

Séguy 1953a, HA, Tadla; Pont 1986b; Hernández-Moreno and Peris 1989, Rif, Tanger; Pont pers. comm., AP, Casablanca

Hydrotaea pellucens Portschinsky, 1879

Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Hydrotaea tuberculata Rondani, 1866

Michelsen pers. comm., AP, Rabat

Hydrotaea velutina Robineau-Desvoidy, 1830

Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Thricops Rondani, 1856

Thricops simplex (Wiedemann, 1817)

Pont 1986b, 1991b; Pont pers. comm., HA, Jebel Ayachi

Reinwardtiini

Muscina Robineau-Desvoidy, 1830

Muscina levida (Harris, 1780)

= Muscina assimilis (Fallén, 1823)

Mouna 1998 – MHNP (no locality, on Boletus nigrescens (Pont pers. comm.)); AP (Rabat) –MISR

Muscina prolapsa (Harris, 1780)

= Muscina pabulorum (Fallén, 1817)

Mouna 1998; Michelsen pers. comm., AP, Rabat, Larache; AP (Sidi Yahia) – MISR

Muscina stabulans (Fallén, 1817)

Becker and Stein 1913, Rif, Tanger; Charrier 1927, Rif, Tanger; Séguy 1930a, AP, Rabat, Mogador, MA, Meknès, HA, Aguerd El Had, Souss; Séguy 1932, HA, Taroudant; Séguy 1934b, AP, Rabat; Séguy 1953a, AP, Rabat, Salé, Forêt Maâmora, Salé; Pont 1986b; Mouna 1998; Pont pers. comm., AP, Casablanca, MA, El Kebab, HA, Jebel Ayachi; AP (Casablanca, Rabat (Pont pers. comm.)) – MHNP; MISR

Coenosiinae

Coenosiini

Coenosia Meigen, 1826

Coenosia antennata (Zetterstedt, 1849)

Michelsen pers. comm., AP, Larache

Coenosia atra Meigen, 1830

Pont 1986b; Barták et al. 2004; Barták and Kubik 2005; Pont pers. comm., HA, Jebel Ayachi

Coenosia attenuata Stein, 1903

Pont 1986b; Pont pers. comm., EM, Figuig

Coenosia humilis Meigen, 1826

Pont 1986b; Pont pers. comm., EM, Figuig, HA, Imlil, Asni, AA, Agadir

Coenosia mixta Schnabl, 1911

Pont 1986b

Coenosia nevadensis Lyneborg, 1970

Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Coenosia pedella (Fallén, 1825)

= Coenosia decipiens Meigen 1826 (certainly a misidentification) in Charrier 1927: 620

Charrier 1927, Rif, Tanger

Coenosia praetexta Pandellé, 1899

Michelsen pers. comm., AP, Larache, Rabat

Coenosia pumila (Fallén, 1825)55

Pont 1986b; Mouna 1998; Gregor et al. 2002

Coenosia testacea (Robineau-Desvoidy, 1830)

Pont 1986b; Pont pers. comm., MA, Ifrane, HA, Jebel Ayachi

Coenosia tigrina (Fabricius, 1775)

Séguy 1930a, AP, Rabat, MA, Meknès; Pont 1986b; Mouna 1998; Pont pers. comm., MA, Ifrane, HA, Jebel Ayachi; MA (Ifrane) – MISR

Lispocephala Pokorny, 1893

Lispocephala brachialis (Rondani, 1877)

Pont 1986b; Gregor et al. 2002; Barták and Kubik 2005; Pont pers. comm., HA, Jebel Ayachi

Lispocephala mikii (Strobl, 1893)

Pont 1986b; Pont pers. comm., EM, Figuig, HA, Jebel Ayachi, AA, Agadir

Lispocephala ungulata (Rondani, 1866)

Ackland and Pont 1966, HA, Jebel Ayachi; Pont 1986b

Orchisia Rondani, 1877

Orchisia costata (Meigen, 1826)

Charrier 1927, Rif, Tanger; Pont 1986b

Schoenomyza Haliday, 1833

Schoenomyza litorella (Fallén, 1823)

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Pont 1986b; Mouna 1998; Pont pers. comm., EM, Figuig, HA, Asni, Imlil, Jebel Ayachi

Limnophorini

Limnophora Robineau-Desvoidy, 1830

Limnophora bipunctata Stein, 1908

Pont 1986b; Pont pers. comm., EM, Figuig

Limnophora flavitarsis Stein, 1908

Pont pers. comm., EM, Figuig, HA, Jebel Ayachi

Limnophora mediterranea Pont, 2012

Pont pers. comm., HA, Jebel Ayachi

Limnophora obsignata (Rondani, 1866)

Séguy 1930a, MA, Aïn Leuh, HA, Aguerd El Had, Souss (Talekjount); Pont 1986b; Dakki 1997; Mouna 1998; Pont pers. comm., MA, Ifrane, HA, Jebel Ayachi, AA, Agadir

Limnophora olympiae Lyneborg, 1965

Pont 1986b; Pont et al. 2012a; Pont pers. comm., HA, Jebel Ayachi

Limnophora pandellei Séguy, 1923

Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Limnophora pollinifrons Stein, 1916

Pont 1986b; Pont et al. 2012a; Pont pers. comm., MA, Aïn El Orma

Limnophora riparia (Fallén 1824)

= Melanochelia riparia Fallén, in Séguy 1930a: 160

Séguy 1930a, AA, Souss (Tenfecht); Vaillant 1956b, HA, Cascade Siroua, Lac Tamhda (Anremer), Sidi Chamarouch; Pont 1986b; Mouna 1998

Limnophora rufimana (Strobl, 1893)

Séguy 1930a, MA, Aïn Leuh; Pont 1986b; Pont pers. comm., EM, Figuig, HA, Jebel Ayachi

Limnophora tigrina (Am Stein, 1860)

Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Lispe Latreille, 1797

Lispe assimilis Wiedemann, 1824

= Lispe inexpectata Canzoneri & Meneghini, 1966

Canzoneri and Meneghini 1966, MA, Oued Fès (Taza); Pont 1986b; Vikhrev 2012b, AP, Essaouira, HA, Marrakech, SA, Tan-Tan

Lispe apicalis Mik, 1869

Canzoneri and Meneghini 1966, MA, Oued Sebou, EM, Guercif (Oued Moulouya); Canzoneri and Meneghini 1972, MA, Taza, Oued Fès; Pont 1986b; Koçak and Kemal 2010

Lispe bengalensis (Robineau-Desvoidy, 1830)

= Lispe berlandi Séguy, in Séguy 1940: 341

Séguy 1940, AA, Rio de Oro (Oued Eddahab) (type locality of berlandi)

Lispe caesia Meigen, 1826

= Lispe microchaeta Séguy, in Séguy 1940: 342

Séguy 1940, AA, Rio de Oro (Oued Eddahab) (type locality of microchaeta); Canzoneri and Meneghini 1966, AP, Fedhala (Oued Nefifikh), Saline di Sète; Pont 1986b; Mouna 1998; Koçak and Kemal 2010; Vikhrev et al. 2016, AP, Oualidia lagoon, Essaouira, SA, Tan-Tan (salt lagoon); AP (Chellah) – MISR; ZMUM

Lispe candicans Kowarz, 1892

Séguy 1940, AA, Rio de Oro (Villa Cisneros)

Lispe cilitarsis Loew, 1856

Vikhrev 2012b, SA, Tan-Tan province

Lispe draperi Séguy, 1930

Canzoneri and Meneghini 1966, MA, Azrou, Aguelmane, Fès (Oued Sebou); Vikhrev 2011a, AP, Essaouira, HA, Oued N’fis (east of Marrakech)

Lispe halophora Becker, 1903

Vikhrev pers. comm., SA, Tan-Tan province

Lispe kowarzi Becker, 1903

Vikhrev 2012c, AP, Essaouira

Lispe loewi Ringdahl, 1922

= Lispe litorea Fallén, 1825 (misidentification of authors) in Séguy 1930a: 160

Séguy 1930a, AP, saline mud in Mediterranean region; Canzoneri and Meneghini 1966, AP, Fedhala; Pont 1986b; Dakki 1997; Mouna 1998

Lispe marina Becker, 1913

Michelsen pers. comm., AP, Larache

Lispe melaleuca Loew, 1847

Canzoneri and Meneghini 1966, MA, Azrou (Aguelmane); Pont 1986b, 1991b

Lispe modesta Stein, 1913

Vikhrev 2012b, AP, Essaouira, HA, Marrakech, SA, Tan-Tan

Lispe nana Macquart, 1835

Canzoneri and Meneghini 1966, MA, Taza (Oued Fès), Fès (Oued Sebou), Azrou (Aguelmane), EM, Guercif (Oued Moulouya); Pont 1986b; Mouna 1998; Pont pers. comm., MA, Aïn el Orma, EM, Figuig, HA, Jebel Ayachi; Rif (Oued Laou dunes), AP (Rabat) – MISR

Lispe nivalis Wiedemann, 1830

Canzoneri and Meneghini 1966, MA, Taza (Oued Fès); Pont 1986b, 1991; Vikhrev 2012c, AP, Essaouira, HA, Ouarzazate province, SA, Tan-Tan province; Pont pers. comm., MA, Aïn El Orma

Lispe pectinipes Becker, 1903

= Lispa mixticia Séguy, in Séguy 1941d: 19

Séguy 1941d, HA, Taroudant (type locality of mixticia); Pont 1986b; Mouna 1998; Kirk-Spriggs and McGregor 2009; Vikhrev 2011b, AP, Essaouira; Pont pers. comm., HA, Jebel Ayachi

Lispe pygmaea Fallén, 1825

Canzoneri and Meneghini 1966, MA, Azrou (Aguelmane); Pont 1986b; Vikhrev 2012a, AP, Essaouira

Lispe rigida Becker, 1903

Canzoneri and Meneghini 1966, MA, Taza (Oued Fès); Pont 1986b, 1991; Vikhrev 2012c, HA, Ouarzazate

Lispe scalaris Loew, 1847

= Lispe maroccana Canzoneri & Meneghini, 1966 (as scalaris ssp.)

= Lispe persica Becker, 1904 in Kirk-Spriggs and McGregor 2009

Canzoneri and Meneghini 1966, AP, Fedhala, Dielfa (Oued Tadmid), EM, Guercif (Oued Moulouya), MA, Fès (Oued Sebou); Pont 1986b; Kirk-Spriggs and McGregor 2009; Vikhrev 2012a, HA, Ouarzazate province

Lispe tentaculata (De Geer, 1776)

Séguy 1930a, HA, Kasba Taguendaft (Goundafa), Skoutana (Arround); Séguy 1941a, HA, Imi-n’Ouaka (1500 m); Pont 1986b; Dakki 1997; Mouna 1998; Kirk-Spriggs and McGregor 2009; Pont pers. comm., MA, Aïn el Orma, EM, Figuig, HA, Jebel Ayachi; AP (Rabat), HA (Tizi-n’Tichka) – MISR

Muscinae

Muscini

Dasyphora Robineau-Desvoidy, 1830

Dasyphora albofasciata (Macquart, 1839)

= Dasiphora saltuum Rondani, 1862

Pont 1986b; Mouna 1998; Pont pers. comm., HA, Jebel Ayachi

Dasyphora penicillata (Egger, 1856)

Pont 1986b; Koçak and Kemal 2010; Pont pers. comm., HA, Jebel Ayachi

Dasyphora cyanella (Meigen, 1826)

Peris and Llorente 1963, Rif, Tanger; Mouna 1998

Morellia Robineau-Desvoidy, 1830

Morellia asetosa Baranov, 1925

= Morellia simplex (Loew, 1857) (misidentification of authors) in Peris and Llorente 1963, Rif, Tanger; Pont 1986b56

Musca Linnaeus, 1758

Musca autumnalis De Geer, 1776

= Musca corvina Fabricius, 1781 in Séguy 1930a: 156

Charrier 1927, Rif, Tanger; Séguy 1930a, AP, Rabat, MA, Oued Korifla, Meknès; Pont 1986b; Mouna 1998; Pont pers. comm., MA, Azrou, HA, Jebel Ayachi; MA (Volubilis), AA (Tifnit) – MISR

Musca biseta Hough, 1898

Pont 1986b; Pont pers. comm., AP, Temara, MA, Timhadit, Meknès, EM, near Figuig

Musca domestica Linnaeus, 1758

Charrier 1927, Rif, Tanger; Séguy 1930a, 1932, 1934b, AP, Casablanca; Séguy 1934c, AP, Casablanca; Séguy 1941a, HA; Peris and Llorente 1963, Rif, Tanger, Melilla, Bab Taza, El Ajmas, Yebala; Pont 1986b; Mouna 1998; Pârvu et al. 2006, AA, Tiggane Tata; Popescu-Mirceni 2011; Pont pers. comm., AP, Temara, MA, Timhadit, Orionane, Lixus, Meknès, HA, Jebel Ayachi; Grabener 2017; HA (Jebel Tachdirt, 3100 m, Tachdirt (Bords Imminen), 2400–2600 m, Kasba Taguendaft (Goundafa), Andjera (Pont pers. comm.)) – MHNP

Musca larvipara Portschinsky, 1910

Peris and Llorente 1963, Rif, Tanger; Pont 1986b; Pont pers. comm., AP, Forêt Maâmora

Musca osiris Wiedemann, 1830

= Musca vitripennis Meigen, 1826 (misidentifications of authors) in Séguy 1930a: 157

Séguy 1941d, AA, Agadir; Pont 1986b

Musca sorbens Wiedemann, 1830

= Musca angustifrons Thomson, 1869 (misidentification of authors) in Séguy 1930a: 156, 1940: 245, 1953a: 88

Séguy 1930a, MA, Oued Korifla, HA, Talingoult (Goundafa), Souss; Séguy 1940, AA, Rio de Oro (Villa Cisneros); Séguy 1941d, AA, Agadir; Séguy 1949a, AA, from Foum Zguid to Zagora; Saccà 1952; Séguy 1953a, AP, Rabat; Peris and Llorente 1963; Pont 1986b, Rif, Melilla, Tanger, AP, Mogador; Mouna 1998; Koçak and Kemal 2010; Pont pers. comm., MA, Timhadit, HA; Grabener 2017 – MISR

Musca tempestiva Fallén, 1817

Séguy 1941d; Pont 1986b; Pont pers. comm., AP, Forêt Maâmora, HA, Asni

Musca vitripennis Meigen, 1926

= Plaxemyia vitripennis Meigen, 1826 in Becker and Stein 1913: 91

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA, Ras el Ksar, Aïn Leuh, HA, Tinmel (Goundafa), Arround (Skoutana); Séguy 1941d, AA, Agadir; Peris and Llorent 1963, Rif, Tanger, Melilla, AP, Mogador; Pont 1986b; Mouna 1998; Pont pers. comm., HA, Jebel Ayachi; Rif (environs de Tanger, Sart. route de Spartel (Pont pers. comm.)) – MHNP; AP (Cap Cantin, Dradek), MA (Azrou) – MISR

Neomyia Walker, 1859

Neomyia cornicina (Fabricius, 1781)

= Cryptolucilia caesarion (Meigen, 1826) in Séguy 1930a: 156

Becker and Stein 1913, Rif, Tanger; Charrier 1927, Rif, Tanger; Séguy 1930a, Rif, Tanger (Oued Judios), AP, Rabat, MA, M’Rirt, Aïn Leuh, Tizi-s’Tkrine, Forêt Zaers, Forêt Tiffert, HA, Arround (Skoutana), Tachdirt; Séguy 1941d (very common); Peris and Llorente 1963, Rif, Tanger, AP, Mogador, Tzalatza, Reisana, Desembocadura del Lixus; Pont 1986b; Dakki 1997; Mouna 1998; Pont pers. comm., MA, Azrou, HA, Jebel Ayachi – MISR

Neomyia viridescens (Robineau-Desvoidy, 1830)

= Orthellia cornicina (Fabricius, 1781) (misidentifications of authors)

Charrier 1927, Rif, Tanger; Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Polietes Rondani, 1866

Polietes meridionalis Peris & Llorente, 1963

Peris and Llorente 1963, Rif, Tanger; Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Pyrellia Robineau-Desvoidy, 1830

Pyrellia vivida Robineau-Desvoidy, 1830

= Pyrellia cadaverina (Linnaeus, 1758) (misidentifications of authors) in Charrier 1927: 620; Séguy 1930a: 156; Peris and Llorente 1963: 252

= Pyrellia serena (Meigen, 1826) (misidentification of authors) in Charrier 1927: 620

Charrier 1927, Rif, Tanger; Séguy 1930a, MA, Aïn Leuh; Peris and Llorente 1963, Rif, Tanger; Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Stomoxyini

Haematobia Le Peletier & Serville, 1828

Haematobia irritans (Linnaeus, 1758)

= Lyperosia irritans (Linnaeus, 1758) in Séguy 1930a: 157

Séguy 1930a, MA, Meknès; Pont 1986b; Mouna 1998

Stomoxys Geoffroy, 1762

Stomoxys calcitrans (Linnaeus, 1758)

Charrier 1927, Rif, Tanger; Séguy 1930a; Séguy 1941d; Peris 1951, Rif, Tanger; Pont 1986b; Mouna 1998; Pârvu et al. 2006, AA, Tiggane Tata; Dsouli 2009; Popescu-Mirceni 2011; Pont pers. comm., MA, Meknès, Timhadit, Azrou, Sidi Mjber, Neguerett, Tazekka, HA, Jebel Ayachi, El Kebab, AA, Figuig; HA (Haute Réghaya (Pont pers. comm.)) – MHNP; AP (Rabat), MA (Volubilis), HA – MISR

Mydaeinae

Graphomya Robineau-Desvoidy, 1830

Graphomya maculata (Scopoli, 1763)

Séguy 1930a, MA, Forêt Zaers, Aïn Leuh; Pont 1986b; Mouna 1998; Pont pers. comm., HA, Jebel Ayachi; MA (Volubilis) – MISR

Gymnodia Robineau-Desvoidy, 1863

Gymnodia eremophila (Brauer & Bergenstamm, 1894)

Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Gymnodia polystigma (Meigen, 1826)

= Limnophora polystigma (Meigen, 1826)

= Brontaea polystigma (Meigen, 1826)

Mouna 1998; AP (Rabat) – MISR

Gymnodia genurufa (Pandellé, 1899)

Pont 1986b; Pont pers. comm., HA, Jebel Ayachi

Gymnodia tonitrui (Wiedemann, 1824)

= Limnophora tonitrui (Wiedemann, 1824)

= Brontaea tonitrui (Wiedemann, 1824)

Séguy 1949a, AA, Tata; Saccà 1952, AP, Rabat; Pont 1986b; Mouna 1998

Hebecnema Schnabl, 1889

Hebecnema fumosa (Meigen, 1826)

Séguy 1930a, AP, Casablanca; Pont 1986b; Mouna 1998; Pont pers. comm., MA, Azrou, HA, Jebel Ayachi; Rif (Tanger), AP (Mogador (Pont pers. comm.)) – MHNP

Hebecnema nigra (Robineau-Desvoidy, 1830)

= Hebecnema vespertina (Fallén, 1823) (misidentifications of authors) in Séguy 1930a: 159

Séguy 1930a, AP, Casablanca; Pont 1986b; Mouna 1998; Pont pers. comm., HA, Jebel Ayachi

Hebecnema umbratica (Meigen, 1826)

Pont 1986b

Myospila Rondani, 1856

Myospila meditabunda (Fabricius, 1781)

Séguy 1941d, AA, Agadir; Pont 1970; Pont 1986b; Mouna 1998; Pont pers. comm., HA, Jebel Ayachi; AP (Cap Cantin) – MISR

Phaoniinae

Helina Robineau-Desvoidy, 1830

Helina clara (Meigen, 1826)

= Mydaea clara (Meigen, 1826) in Séguy 1930a: 159

Becker and Stein 1913, Rif, Tanger; Charrier 1927, Rif, Tanger; Séguy 1930a, AP, Rabat; Pont 1986b; Mouna 1998; AP (Rabat) – MISR

Helina czernyi Lyneborg, 1970

Michelsen pers. comm., Rif, Chefchaouen, Ouezzane, MA, Azrou, HA, Asni

Helina evecta (Harris, 1780)

= Mydaea lucorum (Fallén, 1823) in Séguy 1930a: 159

Séguy 1930a; Werner 1938, EM, Oudjda-Berguent; Pont 1986b; Mouna 1998; Pont pers. comm., MA, Ifrane, HA, Jebel Ayachi

Helina nevadensis Lyneborg, 1970

Pont 1986b; Pont pers. comm., Rif, Talassemtane, HA, Jebel Ayachi

Helina parcepilosa (Stein, 1907)

Michelsen pers. comm., AP, Rabat, HA, Tinerhir

Helina quadrum (Fabricius, 1805)

= Mydaea quadrum “Fallén” in Séguy 1930a: 159

Séguy 1930a, AP, Rabat; Pont, 1986b; Mouna 1998

Helina reversio (Harris, 1780)

Pont 1986b; Pont pers. comm., MA, Azrou, HA, Jebel Ayachi; MA (Forêt Timelilt, 1650–1900 m (Pont pers. comm.)) – MNHN

Helina richardi Pont, 2012

Pont 2012b, Rif, Ras el Ma, MA, Azrou, HA, Jebel Ayachi

Helina sexmaculata (Preyssler, 1791)

= Mydaea uliginosa (Fallén, 1825)

Mouna 1998; Grabener 2017; MA (Aguelmane Azigza) – MISR

Helina vockerothi Lyneborg, 1970

Michelsen pers. comm., HA, Tizi-n’Test (2100 m), Asni

Phaonia Robineau-Desvoidy, 1830

Phaonia cincta (Zetterstedt, 1846)57

Charrier 1927, Rif, Tanger; Pont 1986b (record queried); Koçak and Kemal 2010

Phaonia errans (Meigen, 1826)

Mouna 1998; Michelsen pers. comm., MA, Azrou; AP (Chellah), MA (Ifrane) – MISR

Phaonia exoleta (Meigen, 1826)

Michelsen pers. comm., AP, Larache

Phaonia mediterranea Hennig, 1963

Pont 1973, HA, Jebel Ayachi; Pont 1986b; Mouna 1998; Gregor et al. 2002

Phaonia rufipalpis (Macquart, 1835)

Michelsen pers. comm., Rif, Ouezzane

Phaonia scutellata (Zetterstedt, 1845)

Michelsen pers. comm., Rif, Ouezzane, HA, Asni, Tizi-n’Test, AA, Aoulouz

Phaonia subventa (Harris, 1780)

Vikhrev and Erofeeva 2018, HA, Oukaimeden (2000 m); Rif (environs de Tanger (Pont pers. comm.)) – MHNP

Phaonia trimaculata (Bouché, 1834)

Séguy 1930a, AP, Casablanca; Séguy 1934b, AP, Maâmora; Séguy 1953a, AP, Port Lyautey, Maâmora, Rabat; Pont 1986b; Dakki 1997; Mouna 1998; Pont pers. comm., AP, Forêt Maâmora, HA, Jebel Ayachi; AP (Rabat) – MISR

Phaonia tuguriorum (Scopoli, 1763)

= Phaonia signata (Meigen, 1826) in Séguy 1930a: 158

Séguy 1930a, MA, Forêt Timlilt; Pont 1986b; Dakki 1997; Mouna 1998; Pont pers. comm., HA, Jebel Ayachi

Phaonia valida (Harris, 1780)

= Phaonia erratica (Fallén, 1825) (misidentifications of authors) in Séguy 1953a: 87

Séguy 1953a, MA, Ifrane; Pont 1986b; Pont pers. comm., HA, Jebel Ayachi; MA (Ifrane (Pont pers. comm.)) – MHNP

Phaonia sp. near szelenyii Mihályi, 1974

HA (Haute Réghaya (Pont pers. comm.)) – MHNP

SCATHOPHAGIDAE

K. Kettani

Number of species: 3. Expected: 4

Faunistic knowledge of the family in Morocco: poor

Scathophaginae

Norellia Robineau-Desvoidy, 1830

Norellia tipularia (Fabricius, 1794)

Ebejer et al. 2019, Rif, Dardara (730 m)

Scathophaga Meigen, 1803

Scathophaga stercoraria (Linnaeus, 1758)

= Scathophaga merdaria Fabricius, 1794, in Séguy 1930a: 163

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, Rif, Tanger, MA, Meknès; Mouna 1998; Koçak and Kemal 2010; AP (Azemour, Oued Yakem, Cap Cantin) – MISR

Scathophaga lutaria (Fabricius, 1794)

Ebejer et al. 2019, Rif, Talassemtane (1554 m), Jebel Lakraâ (1541 m)

Oestroidea

CALLIPHORIDAE

K. Kettani, K. Rognes

Number of species: 8. Expected: 10

Faunistic knowledge of the family in Morocco: moderate

Calliphorinae

Bellardia Robineau-Desvoidy, 1863

Bellardia maroccana (Villeneuve, 1941)

= Onesia maroccana Villeneuve, in Villeneuve 1932: 123

Villeuneuve 1932b; Schumann 1974, AP, Aïn Diab (Casablanca); Verves 2004; Koçak and Kemal 2010

Bellardia mascariensis (Villeneuve, 1926)

Schumann 1974, HA, Marrakech; Verves 2004; Koçak and Kemal 2010

Calliphora Robineau-Desvoidy, 1863

Calliphora vicina Robineau-Desvoidy, 1830

= Calliphora erythrocephala Meigen, in Séguy 1930a: 154

Séguy 1930a; Maurice 1947, Rif, Tanger; AP (Casablanca, Rabat), MA (Azrou, Ifrane) – MNHN, MISR

Calliphora vomitoria (Linnaeus, 1758)

Séguy 1930a; Maurice 1947, Rif, Tanger; Kurahashi 1971; Kurahashi and Magpayo 2000; Mouna 1998; AP (Rabat), HA – MISR, NHMD

Chrysomyinae

Chrysomya Robineau-Desvoidy, 1863

Chrysomya albiceps (Wiedemann, 1819)

Maurice 1947; Séguy 1949a, HA, Alnif; Gonzales-Mora and Peris 1988; Mouna 1998; Verves 2003; Grabener 2017; Dawah et al. 2019; EM (Oujda) – MISR, AP (Rabat) – MNHN

Luciliinae

Lucilia Robineau-Desvoidy, 1863

Lucilia sericata (Meigen, 1824)

Séguy 1930a, Rif, Tanger, EM, Berkane (1350–1400 m), MA, Meknès, Berrechid, HA, Tizi-n’Test, Goundafa (Jebel Imdress, 2000–2450 m); Séguy 1953a, AP, Rabat; Mouna 1998; Grabener 2017; AP (Dradek, Salé), MA (Azrou, Volubilis) – MISR, AP (Temara), MA (Ifrane) – MNHN

Melanomyinae

Melinda Robineau-Desvoidy, 1863

Melinda gentilis (Robineau-Desvoidy, 1830)

MA (Ifrane) – MNHN

Melinda viridicyanea (Robineau-Desvoidy, 1830)

MA (Ifrane) – MNHN

OESTRIDAE

K. Kettani, T. Pape

Number of species: 10. Expected: 12

Faunistic knowledge of the family in Morocco: moderate

Gastrophilinae

Gasterophilus Leach, 1817

Gasterophilus flavipes (Olivier, 1811)

Séguy 1928d, EM, Haute Moulouya; Séguy 1930a, EM, Itzer (Haute Moulouya); Mouna 1998; Li et al. 2019

Gasterophilus haemorrhoidalis (Linnaeus, 1758)

Maurice 1947; Mouna 1998; AP (Rabat) – MISR

Gasterophilus intestinalis (De Geer, 1776)

Séguy 1930a; Mouna 1998; Pandey et al. 1992

Gasterophilus nasalis (Linnaeus, 1758)

= Gasterophilus veterinus Clark, in Mouna 1998: 85

Maurice 1947; Pandey et al. 1992; Mouna 1998

Gasterophilus pecorum (Fabricius, 1794)

Séguy 1930a; Mouna 1998

Hypodermatinae

Hypoderma Rondani, 1856

Hypoderma bovis (Linnaeus, 1758)

Maurice 1947; Mouna 1998

Hypoderma lineatum (De Villers, 1789)

Dakkok et al. 1978, MA, Benslimane; Mouna 1998

Oestrinae

Cephalopina Strand, 1928

Cephalopina titillator (Clark, 1797)

= Cephalopsis titillator Clark, in Séguy 1930a: 139

Séguy 1930a, Sub-SA (camel breeding region); Mouna 1998

Oestrus Linnaeus, 1758

Oestrus ovis Linnaeus, 1761

Séguy 1930a, AP, Rabat (sheep breeding area), HA, Marrakech, Bou Tazzert; Maurice 1947; Mouna 1998 – MISR

Rhinoestrus Brauer, 1886

Rhinoestrus purpureus (Brauer, 1858)

Maurice 1947; Mouna 1998

POLLENIIDAE

K. Kettani, K. Rognes

Number of species: 12. Expected: 15

Faunistic knowledge of the family in Morocco: moderate

Pollenia Fabricius, 1794

Pollenia amentaria (Scopoli, 1763)

Séguy 1949a, AA, Foum-el-Hassan; Séguy 1953a, MA, Sidi Allal Tazi; Mouna 1998; Pârvu et al. 2006, SA, Foum Zghouig; Popescu-Mirceni 2011; Gisondi et al. 2020

Pollenia bicolor Robineau-Desvoidy, 1830

Rognes 1991a, HA, Mikdane (Jebel Ayachi); Rognes 1991b, HA, Mikdane (Jebel Ayachi); Gisondi et al. 2020; HA (Mikdane) – NHMUK

Pollenia contempta Robineau-Desvoidy, 1863

Séguy 1930a, MA, Meknès, from M’Rirt to El Hajeb; Mouna 1998; Rognes 1992

Pollenia haeretica Séguy, 1928

Séguy 1934b, AP, Rabat; Rognes 2010; AP (40 km S Larache) – NHMD

Pollenia ibalia Séguy, 1930

= Pollenia funebris Villeneuve, in Villeneuve 1932: 284

= Pollenia rungsi Séguy, in Séguy 1953a: 88

Séguy 1930a, EM, Berkane, MA, Tlet n’Rhohr (in garden), Douar Ras el Ksar (900 m); Villeneuve 1932, HA, Marrakech; Séguy 1953a, AP, Rabat; Séguy 1941a, HA, cañon Tessaout (M’Goum, 3000–3200 m); Mouna 1998; Rognes 2010; Grabener 2017; Gisondi et al. 2020; HA (Asni) – NHMUK; HA (Ijoukak) – MNHN; HA (15 km SW Tazenakht) – NHMD

Pollenia leclercqiana (Lehrer, 1978)

Rognes 2010, 2011; Gisondi et al. 2020

Pollenia luteovillosa Rognes, 1987

Gisondi et al. 2020, HA, Mikdane (Jbel Ayachi)

Pollenia ponti Rognes, 1991

Rognes 1991b, HA, Jaffar river (Jebel Ayachi); Gisondi et al. 2020; HA (Jebel Ayachi) – NHMUK

Pollenia rudis (Fabricius, 1794)

Séguy 1930a, EM, Itzer (Haute Moulouya), Berkane (1350 m), AP, Casablanca, MA, Meknès, Aharmoumou (1100 m), Tlet n’Rhohr (in garden), Douar Ras el Ksar (1900 m), HA, Tizi-n’Test, Goundafa (Jebel Imdress, 2000–2450 m); Maurice 1947, Rif, Tanger; Rognes 1987, AP, Aïn Diab, Larache, HA, Asni, Jebel Ayachi, Mikdane; Grabener 2017; Gisondi et al. 2020; AP (Casablanca, Rabat), MA (Ifrane), HA (Haute Réghaya) – MNHN

Pollenia ruficrura Rondani, 1862

Rognes 2011; Gisondi et al. 2020

Pollenia stigi (Rognes, 1992)

Rognes 1992, MA, Ifrane, Azrou; Gisondi et al. 2020

Pollenia vagabunda (Meigen, 1826)

= Pollenia hasei Séguy, in Séguy 1928e: 370, Séguy 1930a: 147

Séguy 1928e; Séguy 1930a, AP, Casablanca; Mouna 1998; Rognes 1992; Gisondi et al. 2020

RHINIIDAE

K. Kettani, K. Rognes

Number of species: 17. Expected: ~20

Faunistic knowledge of the family in Morocco: moderate

Cosmina Robineau-Desvoidy, 1830

Cosmina claripennis Robineau-Desvoidy, 1830

= Cosmina bezziana Villeneuve, in Villeneuve 1932a: 285

Villeneuve 1932a, AP, Mogador; Schumann 1986; Mouna 1998; Rognes 2002

Cosmina maroccana Séguy, 1949

Séguy 1949a, AA, Tenfecht, Vallée du Guir, SA, Guelmim; Séguy 1953a, AA, Tenfecht, Vallée du Guir, SA, Guelmim; Mouna 1998; Rognes 2002; Grabener 2017

Cosmina punctulata Robineau-Desvoidy, 1830

Séguy 1930a, AA, Tenfecht (Souss, 1000–1500 m)

Cosmina viridis (Townsend, 1917)

Ebejer et al. 2019, AA, 13 km E of Goulmima (1100 m)

Rhinia Robineau-Desvoidy, 1830

Rhinia apicalis (Wiedemann, 1830)

Séguy 1930a, AP, Rabat, MA, Forêt Zaers; Zumpt 1956; Gonzales-Mora and Peris 1988; Mouna 1998; Verves 2003; Lehrer 2007

Rhyncomya Robineau-Desvoidy, 1830

Rhyncomya callopis (Loew, 1856)58

Séguy 1941d, AA, Agadir; Séguy 1953a, AP, Sidi Ifni, SA, Tindouf, Amguilli Sguelma, Guelta des Zemmours; Schumann 1986; Mouna 1998; Rognes 1992; Dawah et al. 2019; El Hawagry and El-Azab 2019; MA (Tafilalt) – MISR

Rhyncomya columbina (Meigen, 1824)

Peris 1951, Rif, Tanger, MA, Ifrane; Schumann 1986; Gonzalez-Mora and Peris 1988

Rhyncomya cyanescens (Loew, 1844)

= Rhyncomya hemisia Séguy, in Séguy 1930a: 150

Séguy 1930a, EM, Berkane (1350–1400 m); Gonzales-Mora and Peris 1988; Rognes 2002, MA

Rhyncomya desertica (Peris, 1951)

Grabener 2017; El Hawagry and El-Azab 2019

Rhyncomya impavida (Rossi, 1790)

Séguy 1930a, Rif, Tanger, EM, Berkane (1350–1400 m), MA, Forêt Tiffert (2000–2200 m); Schumann 1986; Mouna 1998

Rhyncomya nigripes (Séguy, 1933)

Grabener 2017; El Hawagry and El-Azab 2019

Rhyncomya ursina Séguy 1928

= Rhynchomyia ursina Séguy, in Séguy 1928b: 152

Séguy 1928b, AP, Atlantic coast of SA

Rhyncomya yahavensis Rognes, 2002

Grabener 2017; Ebejer et al. 2019, AA, 30 km W of Errachidia (1065 m)

Rhyncomya zernyana Villeneuve, 1926

Zumpt 1956; Gonzales-Mora and Peris 1988

Stomorhina Rondani, 1861

Stomorhina lunata (Fabricius, 1805)

Séguy 1930a; Peris 1951; Gonzales-Mora and Peris 1988; Mouna 1998; AP (Rabat) – MISR

Villeneuviella Austen, 1914

Villeneuviella icadion (Séguy, 1953)

= Rhynchoestrus icadion Séguy, in Séguy 1953a: 89

Séguy 1953a, SA, Tindouf

Villeneuviella weissi (Séguy, 1926)

= Rhynchoestrus weissi Séguy, in Séguy 1934b: 162

Séguy 1934b, EM, Berkane Zobzit (1100 m); Séguy 1953a, SA, Mader Bergat

RHINOPHORIDAE

K. Kettani, T. Pape

Number of species: 8. Expected: 15

Faunistic knowledge of the family in Morocco: poor

Rhinophorinae

Melanophora Meigen, 1803

Melanophora roralis (Linnaeus, 1758)

Peris 1963, Rif, Tanger, AP, Larache; Mouna 1998

Oplisa Rondani, 1862

Oplisa aterrima (Strobl, 1899)

= Hoplisa aterrima Strobl, in Peris 1963: 602

Peris 1963, Rif, Tanger, Zoco de Taleta, Ketama; Mouna 1998; Cerretti et al. 2020

Paykullia Robineau-Desvoidy, 1830

Paykullia carmela (Peris, 1963)

= Chaetostevenia carmela Peris, in Peris 1963: 606

Peris 1963, Rif, Tanger; Cerretti et al. 2020; Pape and Thompson 2019 – MNCN

Phyto Robineau-Desvoidy, 1830

Phyto atrior (Villeneuve, 1941)

= Styloneuria atrior Villeneuve, in Villeneuve 1941: 122

Villeneuve 1941, AP, Rabat; Cerretti et al. 2020; Pape and Thompson 2019 – IRSNB

Phyto discrepans Pandellé, 1896

Cerretti et al. 2020, Rif, Chefchaouen (600 m), Ouezzane (300 m), MA, 40 km N Fès (1150 m) – NHMD

Phyto melanocephala (Meigen, 1824)

Ebejer et al. 2019, Rif, Barrage Smir (145 m); Cerretti et al. 2020

Stevenia Robineau-Desvoidy, 1830

Stevenia deceptoria (Loew, 1847)

Mulieri et al. 2010; Cerretti et al. 2020

Tricogena Rondani, 1856

Tricogena rubricosa (Meigen, 1824)

= Frauenfeldia rubricosa Meigen, in Peris 1963: 603

Peris 1963, Rif, Tanger; Mouna 1998; Cerretti et al. 2020 – NHMD

SARCOPHAGIDAE

K. Kettani, D. Whitmore, T. Pape

Number of species: 66. Expected: ~150

Faunistic knowledge of the family in Morocco: poor

Miltogramminae

Amobia Robineau-Desvoidy, 1830

Amobia signata (Meigen, 1824)

Pape 1996; Verves 2019

Apodacra Macquart, 1854

Apodacra africana Rohdendorf, 1930

Pape 1996, Rif, Tanger; Verves 2019

Craticulina Pandellé, 1895

Craticulina antachates (Séguy, 1949)

= Apodacra antachates Séguy, in Séguy 1949a: 160

Séguy 1949a, AA, Zagora; Pape 1996, AA, Zagora; Mouna 1998

Craticulina tabaniformis (Fabricius, 1805)

Fabricius 1805, AP, Mogador; Séguy 1930a, AP, Mogador; Séguy 1935a, AP, beach of Rabat; Pape 1996; El Hawagry and El-Azab 2019; Verves 2019

Dolichotachina Villeneuve, 1913

Dolichotachina marginella (Wiedemann, 1930)

Pape 1996; Grabener 2017; El Hawagry and El-Azab 2019; Verves 2019

Macronychia Rondani, 1859

Macronychia lemariei Jacentkovský, 1941*

AP

Macronychia polyodon (Meigen, 1824)

Pape 1996; Verves 2019

Metopia Meigen, 1803

Metopia argyrocephala (Meigen, 1824)

= Metopia leucocephala (Rossi), in Mouna 1998: 86

Mouna 1998

Miltogramma Meigen, 1803

Miltogramma aurifrons Dufour, 1850

Séguy 1930a, AP, Rabat, MA, Meknès; Pape 1996; Mouna 1998; El Hawagry and El-Azab 2019; Verves 2019

Miltogramma germari Meigen, 1824

Séguy 1930a, MA, Meknès, from M’Rirt to El Hajeb, Sidi Taibi; Pape 1996; Mouna 1998; El Hawagry and El-Azab 2019; Verves 2019

Miltogramma maroccana (Séguy, 1941)

= Sphecapatodes maroccana Séguy, in Séguy 1941d: 22

Séguy 1941d, AA, Taroudant; Pape and Szpila 2012

Miltogramma murina Meigen, 1824

Pape 1996; Verves 2019

Miltogramma oestracea (Fallén, 1820)

Ebejer et al. 2019, Rif, Belwazen (M’Diq, 200 m), AP, Lower Loukous saltmarsh (2 m)

Miltogramma rutilans Meigen, 1824

Ebejer et al. 2019, Rif, Oued Mhajrate (Ben Karrich, 180 m)

Miltogramma testaceifrons (Roser, 1840)

= Miltogramma pilitarsis Rondani, in Séguy 1930a: 145; Mouna 1998: 86

Séguy 1930a, MA, Aïn Leuh; Pape 1996; Mouna 1998

Protomiltogramma Townsend, 1916

Protomiltogramma fasciata (Meigen, 1824)

= Setulia fasciata (Meigen), in Mouna 1998: 86

Pape 1996; El Hawagry and El-Azab 2019; Verves 2019

Senotainia Macquart, 1846

Senotainia albifrons (Rondani, 1859)

= Sphecapata albifrons Rondani, in Séguy 1930a: 145

Séguy 1930a

Taxigramma Macquart, 1850

Taxigramma heteroneura (Meigen, 1830)

Pape 1996; El Hawagry and El-Azab 2019; Verves 2019

Taxigramma pluriseta (Pandellé, 1895)

Ebejer et al. 2019, Rif, Oued Mhajrate (Ben Karrich, 180 m), AP, Lower Loukous saltmarsh (2 m)

Paramacronychiinae

Nyctia Robineau-Desvoidy, 1830

Nyctia halterata (Panzer, 1798)

= Musca maura Fabricius, in Fabricius 1805: 302

Fabricius 1805, Rif, Tanger; Pape 1996, Rif, Tanger; Grabener 2017; El Hawagry and El-Azab 2019; Verves 2019

Nyctia lugubris (Macquart, 1834)

Ebejer et al. 2019, AP, Lower Loukous saltmarsh (2 m)

Sarcophila Rondani, 1856

Sarcophila latifrons (Fallén, 1817)59

Séguy 1930a, AP, Maâmora, HA, Skoutana (Arround, 2000–2400 m); Mouna 1998

Wohlfahrtia Brauer & Bergenstamm, 1889

Wohlfahrtia bella (Macquart, 1839)

= Disjunctio bella (Macquart), in Séguy 1930a: 144

Séguy 1930a, MA, Aïn Leuh; Pape 1996; Mouna 1998; Hall et al. 2009; El Hawagry and El-Azab 2019; Verves 2019

Wohlfahrtia indigens Villeneuve, 1928

Pape 1996; El Hawagry and El-Azab 2019; Verves 2019

Wohlfahrtia magnifica (Schiner, 1862)

Delanoë 1922, AP, Doukkala; Séguy 1930a, AP, Maâmora; Séguy 1941a, HA, Tizi-n’Icheden (3000 m); Maurice 1947; Pape 1996; Mouna 1998; Lmimouni et al. 2004; Tliqui et al. 2007; Farkas et al. 2009, Rif, Al Hoceima, Taguidit, Tafensa, EM, Aghbal; Hall et al. 2009; El Hawagry and El-Azab 2019; Verves 2019

Wohlfahrtia nuba (Wiedemann, 1830)

Pape 1996; El Hawagry and El-Azab 2019; Verves 2019

Wohlfahrtia trina (Wiedemann, 1830)

Pape 1996; Mouna 1998; El Hawagry and El-Azab 2019

Sarcophaginae

Blaesoxipha Loew, 1861

Blaesoxipha (Blaesoxipha) lapidosa Pape, 1994

Pape 1996; Grabener 2017; El Hawagry and El-Azab 2019

Blaesoxipha (Blaesoxipha) litoralis (Villeneuve, 1911)

Pape 1996; Verves 2019

Blaesoxipha (Blaesoxipha) pygmaea (Zetterstedt, 1844)

Pape 1996; Verves 2019

Blaesoxipha (Servaisia) rossica Villeneuve, 1912

Pape 1996; Verves 2019

Ravinia Robineau-Desvoidy, 1863

Ravinia pernix (Harris, 1780)

= Gesneriodes disjuncta Séguy, in Séguy 1938: 43

= Sarcophaga striata (Fabricius), in Séguy 1941d: 22, Séguy 1949a: 159

Séguy 1938, HA, Skoutana; Séguy 1941d, AA, Taroudant; Séguy 1949a, AA, Akka; Pape 1996; Grabener 2017; El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga Meigen, 1826

Sarcophaga (Bercaea) africa (Wiedemann, 1824)

Pape 1996; Abkari et al. 1998; Verves 2003, 2019; Grabener 2017; El Hawagry and El-Azab 2019

Sarcophaga (Helicophagella) maculata Meigen, 1835

Pape 1996; El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga (Helicophagella) melanura Meigen, 1826

El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga (Helicophagella) novercoides Böttcher, 1913*

Rif, HA, AA

Sarcophaga (Heteronychia) balanina Pandellé, 1896

Whitmore et al. 2013, AP, Larache; Fendane et al. 2018, AP, Diabat (Essaouira), Sidi Abed (El Jadida), Bir Retma (Casablanca); Verves 2019

Sarcophaga (Heteronychia) cucullans Pandellé, 1896

Séguy 1930a, HA, Maharidja; Mouna 1998

Sarcophaga (Heteronychia) ferox Villeneuve, 1908

Whitmore 2011, Rif, Ouezzane, AP, Larache, MA, Béni Mellal, Afourer, AA, Aoulouz; Whitmore et al. 2013; El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga (Heteronychia) filia Rondani, 1860

Whitmore 2011, MA, Azrou, Timahdit; Whitmore et al. 2013; Verves 2019

Sarcophaga (Heteronychia) javita (Peris, González-Mora & Mingo, 1998)*

AA

Sarcophaga (Heteronychia) longestylata Strobl, 1906

Pape 1996; Whitmore et al. 2013, MA, Ifrane, Azrou; Fendane et al. 2018, AP, Sidi Abed (El Jadida)

Sarcophaga (Heteronychia) minima Rondani, 1862

Whitmore 2011, MA, Azrou, Ifrane, Afourer (Béni Mellal), HA, Ijoukak, Ouirgane (Marrakech), AA, Oulma (Agadir); Whitmore et al. 2013; Fendane et al. 2018, AP, Smimou (Essaouira), El Akarta (Oualidia); Verves 2019

Sarcophaga (Heteronychia) obvia (Povolný, 2004)

Whitmore et al. 2013, MA, Afourer (Béni Mellal), HA, Ait Lekak (Marrakech), S Asni (Imlil, Marrakech), Tagadirt, Quirgane, AA, Oulma Ort (Agadir)

Sarcophaga (Heteronychia) pandellei (Rohdendorf, 1937)

Séguy 1930a, MA, Tizi-s’Tkrine; Mouna 1998; Whitmore et al. 2013, MA, Azrou, Ifrane, Afourer (Béni Mellal)

Sarcophaga (Heteronychia) tangerensis Whitmore, 2011

= Heteronychia (Heteronychia) amica Peris, González-Mora & Mingo, in Peris et al. 1998: 173

Peris et al. 1998, Rif, Tanger; Whitmore 2011, Rif, Tanger

Sarcophaga (Heteronychia) villeneuveana (Enderlein, 1928)

= Pierretia (Bercaea) maroccana Rohdendorf, in Rohdendorf 1937: 325

= Sarcophaga (Heteronychia) penicillata Villeneuve, in Coupland and Barker 2004: 113 (misidentification)

Rohdendorf 1937, MA, Aïn Defali; Coupland and Barker 2004; Whitmore 2009; Fendane et al. 2018, AP, Diabat (Essaouira), Ghabat Tansift (Souiria), Lalla Fatna (Safi), Laatoutate (Safi), El Akarta (Oualidia), Sidi Abed (El Jadida), Bir Retma (Casablanca); Verves 2019

Sarcophaga (Heteronychia) uncicurva Pandellé, 1896

Fendane et al. 2018, AP, Smimou (Essaouira), Diabat (Essaouira), Lalla Fatna (Safi), Laatoutate (Safi), Bir Retma (Casablanca)

Sarcophaga (Liopygia) argyrostoma (Robineau-Desvoidy, 1830)*

HA

Sarcophaga (Liopygia) crassipalpis Macquart, 1839

Pape 1996; Grabener 2017; El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga (Liosarcophaga) aegyptica Salem, 1935

Fendane et al. 2018, AP, Sidi Abed (El Jadida)

Sarcophaga (Liosarcophaga) dux Thomson, 1869*

SA

Sarcophaga (Liosarcophaga) jacobsoni (Rohdendorf, 1937)

Pape 1996; El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga (Liosarcophaga) marshalli Parker, 1923

Fendane et al. 2018, AP, Smimou (Essaouira), Diabat (Essaouira), Ghabat Tansift (Souiria), Laatoutate (Safi); El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga (Liosarcophaga) pharaonis Rohdendorf, 1934

Carles-Tolrá 2002; El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga (Liosarcophaga) tibialis Macquart, 1851

= Sarcophaga beckeri Villeneuve, in Maurice 1947: 57; Mouna 1998: 86

Maurice 1947; Mouna 1998; Fendane et al. 2018, AP, Laatoutate (Safi)

Sarcophaga (Liosarcophaga) teretirostris Pandellé, 1896

= Parasarcophaga decellei Lehrer, in Lehrer 1976: 3

Lehrer 1976, MA, Kandar, Imouzzer; Pape 1996, MA, Kandar, Imouzzer; Verves 2019

Sarcophaga (Liosarcophaga) tuberosa Pandellé, 189660

Mouna 1998

Sarcophaga (Myorhina) nigriventris Meigen, 1826

Pape 1996; Mouna 1998; Fendane et al. 2018, AP, Ghabat Tansift (Souiria), Laatoutate (Safi), El Akarta (Oualidia), Sidi Abed (El Jadida), Bir Retma (Casablanca); Verves 2019

Sarcophaga (Myorhina) soror Rondani, 1860

Fendane et al. 2018, AP, Sidi Abed (El Jadida)

Sarcophaga (Pandelleana) protuberans Pandellé, 1896

Séguy 1949a, AA, Agadir Tissint, SA, Guelmim, Foum-el-Hassan, Tata; Mouna 1998

Sarcophaga (Parasarcophaga) hirtipes Wiedemann, 1830

Pape 1996; Verves 2003; El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga (Sarcophaga) lehmanni Müller, 1922

Pape 1996; Cassar et al. 2005, Rif, Smir lagoon; El Hawagry and El-Azab 2019; Verves 2019

Sarcophaga (Sarcophaga) marcelleclercqi Lehrer, 1975

Lehrer 1975; Pape 1996, MA, Azrou; Verves 2019

Sarcophaga (Thyrsocnema) belgiana (Lehrer, 1976)

Lehrer 1976; Pape 1996

Sarcophaga (Thyrsocnema) sp.61

Ebejer et al. 2019 [as incisilobata, misidentification], Rif, Tahaddart (8 m)

New records for Morocco

Macronychia lemariei Jacentkovský, 1941

Atlantic plain: Rabat, Forêt de Maâmora, 100 m, 25–26.iv.1989, 1♂1♀, Zoological Museum of Copenhagen Expedition (NMHD).

Sarcophaga (Helicophagella) novercoides Böttcher, 1913

Rif: Ouezzane, 300 m, 21–22.iv.1989, 1♂, Zoological Museum of Copenhagen Expedition (NMHD).

High Atlas: Marrakech, Ouirgane, 1000 m, 1–9.iv.1997 Mai, 1♂, C. Kassebeer leg. (NMHD).

Anti Atlas: 30 km NW Aoulouz, 1400 m, 10.iv.1989, 1♂, Zoological Museum of Copenhagen Expedition (NMHD).

Sarcophaga (Heteronychia) javita (Peris, González-Mora & Mingo, 1998)

Anti Atlas: Agadir, S Oulma, 30°31'N, 9°09'W, 200 m, 21.iv.1997, 1♂, C. Kassebeer leg. (NMHD).

Sarcophaga (Liopygia) argyrostoma (Robineau-Desvoidy, 1830)

High Atlas: Marrakech, Tagadirt, Ouirgane, 1000 m, 1.x.1994, 1♀, C. Kassebeer leg. (NMHD).

Sarcophaga (Liosarcophaga) dux Thomson, 1869

Sahara: Erfoud, Rissani area, 900 m, 13–14.iv.1989, Zoological Museum of Copenhagen Expedition (NHMD).

TACHINIDAE

K. Kettani, P. Cerretti, H.-P. Tschorsnig

Number of species: 147. Expected: 200

Faunistic knowledge of the family in Morocco: poor

Dexiinae

Dexiini

Billaea Robineau-Desvoidy, 1830

Billaea lata (Macquart, 1849)

= Rhynchodinera lata Macquart, in Séguy 1930a: 143

Séguy 1930a, MA, Aharmoumou, Camp Boulhout, Sidi Bettache, Aïn Sferguila, Meknès; Mouna 1998; AP (Mehdia) – MISR; AP (Essaouira, 4 km E Ounara), HA (Marrakech, Lakhdar, N Demnate) – PCPT

Estheria Robineau-Desvoidy, 1830

Estheria atripes Villeneuve, 1920

Cerretti and Tschorsnig 2012

Estheria iberica Tschorsnig, 2003*

MA

Estheria nigripes (Villeneuve, 1920)

Cerretti and Tschorsnig 2012; MA (Béni Mellal, El Ksiba), AA (Agadir, Oulma) – PCPT

Estheria picta (Meigen, 1826)62

Moutia 1940

Zeuxia Meigen, 1826

Zeuxia aberrans (Loew, 1847)

= Zeuxia nigripes Meigen, in Séguy 1941d: 23

Brémond 1938; Séguy 1941d, AP, Rabat, MA, Volubilis, AA, Agadir; IOBC-List 11; Mesnil 1980; Mouna 1998; Tschorsnig 2017; AA (10 km NW Aït-Baha) – PCPT

Dufouriini

Dufouria Robineau-Desvoidy, 1830

Dufouria nigrita (Fallén, 1810)

Ebejer et al. 2019, AP, Larache (Lower Loukous saltmarsh, 2 m); MA (Ouzoud) – PCPT

Voriini

Athrycia Robineau-Desvoidy, 1830

Athrycia trepida (Meigen, 1824)*

MA

Cyrtophloeba Rondani, 1856

Cyrtophloeba ruricola (Meigen, 1824)

= Plagia ruricola Meigen, in Séguy 1935a: 120, in Rungs 1940: 14

Séguy 1935a, MA, Ifrane; Rungs 1940, MA (Cédraie); Mouna 1998; HA (Tizi-n’Test), AA (Taroudant) – PCPT

Eriothrix Meigen, 1830

Eriothrix apennina (Rondani, 1862)

Herting and Dely-Draskovits 1993; Koçak and Kemal 2010

Eriothrix rufomaculata (De Geer, 1776)*

MA, HA

Hypovoria Villeneuve, 1913

Hypovoria hilaris Villeneuve, 1912

Séguy 1935a, AP, Oued Beth; Séguy 1953a, AP, Sehoul; Mouna 1998; AA (10 km SE Aït-Ourir) – PCPT

Hypovoria pilibasis (Villeneuve, 1922)

Zeegers 2010; HA (Tizi-n’Test), AA (Taroudant) – PCPT

Kirbya Robineau-Desvoidy, 1830

Kirbya moerens (Meigen, 1830)*

MA

Nanoplagia Villeneuve, 1929

Nanoplagia sinaica (Villeneuve in Hermann & Villeneuve 1909)

Cerretti 2009; Grabener 2017; HA (Marrakech, 8 km N Ouirgane), AA (40 km SW Ouarzazate, 10 km SW Tazenakht, NE Agadir, 12 km W Oulma) – PCPT

Periscepsia Gistel, 1848

Periscepsia meyeri (Villeneuve, 1930)

Ebejer et al. 2019, Rif, Adrou (PNPB, 556 m)

Stomina Robineau-Desvoidy, 1830

Stomina caliendrata (Rondani, 1862)

= Morphomyia caliendrata Rondani, in Séguy 1930a: 143

Séguy 1930a, MA, from M’Rirt to Hajeb, HA, Kasba Taguendaft (Gounfada); Mouna 1998; HA (Massif Toubkal) – PCPT

Thelaira Robineau-Desvoidy, 1830

Thelaira haematodes (Meigen, 1824)63

= Phoenicella haematodes Meigen: Séguy 1930a: 142

Séguy 1930a, HA, Arround; Mouna 1998

Uclesia Girschner, 1901

Uclesia fumipennis Girschner, 1901

Séguy 1934b; Herting and Dely-Draskovits 1993; Mouna 1998; HA (Marrakech) – MISR

Voria Robineau-Desvoidy, 1830

Voria ruralis (Fallén, 1810)

Jourdan 1935c

Wagneria Robineau-Desvoidy, 1830

Wagneria dilatata Kugler, 1977

Kugler 1977

Exoristinae

Acemyini

Ceracia Rondani, 1865

Ceracia mucronifera Rondani, 1865

= Myothyria benoisti (Mesnil), in Mesnil 1959: 20

Mesnil 1959, MA, Forêt Maâmora near Tiflet; Herting and Dely-Draskovits 1993; Cerretti and Ziegler 2004

Blondeliini

Compsilura Bouché, 1834

Compsilura concinnata (Meigen, 1824)

IOBC-list 1 (1956); Hérard and Fraval 1980

Istocheta Rondani, 1859

Istocheta cinerea (Macquart, 1850)

Herting 1960

Istocheta longicornis (Fallén, 1810)

= Latigena longicornis Fallén, in Séguy 1953a: 91

Séguy 1953a, AP, Forêt Zaers

Lomachantha Rondani, 1859

Lomachantha parra Rondani, 1859

Efetov and Tarmann 1999

Robinaldia Herting, 1983

Robinaldia angustata (Villeneuve, 1933)

Herting and Dely-Draskovits 1993; Tschorsnig and Herting 1994

Zaira Robineau-Desvoidy, 1830

Zaira cinerea (Fallén, 1820)

Ebejer et al. 2019, Rif, Aïn Jdioui (Tahaddart, 8 m)

Eryciini

Alsomyia Brauer & Bergenstamm, 1891

Alsomyia olfaciens (Pandellé, 1896)

IOBC-List 12 (1993)

Amphicestonia Villeneuve, 1939

Amphicestonia dispar (Villeneuve, 1922)

Herting and Dely-Draskovits 1993; Cerretti and Ziegler 2004; MA (Ifrane) – PCPT

Aplomyia Robineau-Desvoidy, 1830

Aplomyia confinis (Fallén, 1820)

Ebejer et al. 2019, Rif, Dardara (484 m)

Carcelia Robineau-Desvoidy, 1830

Carcelia dilaticornis Mesnil, 1950

Mesnil 1950

Carcelia iliaca (Ratzeburg, 1840)63

Mouna 1998

Carcelia lucorum (Meigen, 1824)*

HA

Drino Robineau-Desvoidy, 1863

Drino atropivora (Robineau-Desvoidy, 1830)

= Sturmia atropivora Robineau-Desvoidy, in Bléton and Fieuzet 1939: 64

De Lépiney and Mimeur 1932; Bléton and Fieuzet 1939, MA, Fès; Mouna 1998; Tschorsnig 2017; AP (Rabat), MA (Bel Lakssiri) – MISR

Drino galii (Brauer & Bergenstamm, 1891)*

HA, AA

Drino gilva (Hartig, 1838)63

= Sturmia gilva Hartig

Mouna 1998 – MISR (no locality given)

Drino imberbis (Wiedemann, 1830)63

Rungs 1954; Grabener 2017

Drino inconspicua (Meigen, 1830)

Séguy 1935a, AP, Sehoul (Rabat); Bléton and Fieuzet 1939, MA, Dayat Achleff; Mouna 1998; MA (Meknès, Béni Mellal) – MISR

Drino maroccana Mesnil, 1951

De Lépiney 1930; De Lépiney and Mimeur 1932 (probably misidentified as Sturmia inconspicua); Mesnil 1951; Herting and Dely-Draskovits 1993; Ziegler 2011

Drino triplaca Herting, 1979

Herting 1979, AP, Rabat

Drino vicina (Zetterstedt, 1849)

= Sturmia vicina Zetterstedt, 1849

De Lépiney and Mimeur 1932; Bouclier-Maurin 1923; AP (Rabat) – MISR

Gymnophryxe Villeneuve, 1922

Gymnophryxe carthaginiensis (Bischof, 1900)

Mesnil 1956

Nilea Robineau-Desvoidy, 1863

Nilea innoxia Robineau-Desvoidy, 1863

Bléton and Fieuzet 1939

Phryxe Robineau-Desvoidy, 1830

Phryxe caudata (Rondani, 1859)

Biliotti 1956; El Yousfi 1994; IOBC-list 11 (1989)

Phryxe setifacies (Villeneuve, 1910)

IOBC-list 11 (1989); IOBC-list 12 (1993)

Phryxe vulgaris (Fallén, 1810)

Séguy 1953a, MA, Tamrabta (1700 m) – PCPT (HA, Marrakech, Imlil, S Asni)

Ptesiomyia Brauer & Bergenstamm, 1893

Ptesiomyia microstoma Brauer & Bergenstamm, 1893

Séguy 1953a, AP, Rabat; EM (Mte des Béni Snassen, Taforalt), MA (Béni Mellal, Bin-el-Ouidane; Meknès, Ifrane (NPI)), HA (Marrakech, Oukaimeden) – PCPT

Senometopia Macquart, 1834

Senometopia separata (Rondani, 1859)

Hérard and Fraval 1980

Tryphera Meigen, 1838

Tryphera lugubris (Meigen, 1824)

IOBC-List 1 (1956)

Ethillini

Atylomyia Brauer, 1898

Atylomyia albifrons Villeneuve, 1911

= Atylomyia rungsi Mesnil, in Mesnil 1962: 778

Mesnil 1962, AA (near Agadir), Aït Melloul; Herting and Dely-Draskovits 1993

Exoristini

Bessa Robineau-Desvoidy, 1863

Bessa parallela (Meigen, 1824)

Séguy 1935a (probably misidentified as Bessa selecta); Tschorsnig 2017

Chetogena Rondani, 1856

Chetogena filipalpis Rondani, 1859

Ebejer et al. 2019, Rif, Aïn Jdioui (Tahaddart, 8 m); MA (Fès, Sidi Harazem, Ifrane, Forêt de Cèdres) – PCPT

Chetogena mageritensis (Villeneuve & Mesnil, 1936)

Herting and Dely-Draskovits 1993

Chetogena media Rondani, 1859*

MA

Chetogena nigrofasciata (Strobl, 1902)

= Chetogena repanda (Mesnil, 1939), in Herting and Dely-Draskovits 1993 (type locality: Skel): 17

Gheibi et al. 2010

Chetogena obliquata (Fallén, 1810)

De Lépiney and Mimeur 1932; Herting 1960; IOBC-list 12 (1993); HA (Marrakech, Oukaimeden) – PCPT

Exorista Meigen, 1803

Exorista deligata Pandellé, 1896

Mesnil 1946, AP, Sidi Taibi near Kénitra; Herting and Dely-Draskovits 1993; Cerretti and Ziegler 2004; Gheibi et al. 2010

Exorista grandis (Zetterstedt, 1844)

Ebejer et al. 2019, Rif, Dardara (484 m)

Exorista larvarum (Linnaeus, 1758)

Mouna 1998

Exorista nova (Rondani, 1859)

Tschorsnig 2017

Exorista rendina (Herting, 1975)*

AA

Exorista segregata (Rondani, 1859)

Mouna 1998; AA (Agadir, Oulma) – PCPT

Goniini

Anurophylla Villeneuve, 1938

Anurophylla aprica (Villeneuve, 1912)*

MA

Baumhaueria Meigen, 1838

Baumhaueria goniaeformis (Meigen, 1824)

De Lépiney and Mimeur 1932; AP (Maâmora) – MISR; HA (Marrakech, Oukaimeden) – PCPT

Blepharipa Rondani, 1856

Blepharipa pratensis (Meigen, 1824)*

HA

Ceratochaetops Mesnil, 1970

Ceratochaetops triseta (Villeneuve, 1922)

= Ceratochoeta triseta Villeneuve, in Rungs 1940: 15

Rungs 1940, MA (Cédraie); Mouna 1998; MA (Khénifra, El-Herri, Ifrane (NPI), Meknès), HA (Marrakech, 8 km N Ouirgane) – PCPT

Ceromasia Rondani, 1856

Ceromasia rubrifrons (Macquart, 1834)

IOBC-list 11 (1989); IOBC-list 12 (1993); HA (Marrakech, Ouirgane, Tagadirt, S Asni) – PCPT

Clemelis Robineau-Desvoidy, 1863

Clemelis pullata (Meigen, 1824)

IOBC-list 13 (1997)

Gaedia Meigen, 1838

Gaedia connexa Meigen, 1824

Séguy 1953a, EM, Berkane

Gonia Meigen, 1803

Gonia aterrima Tschorsnig, 1991

Tschorsnig 1991

Gonia atra Meigen, 1826

Séguy 1930a, MA, Tizi-n’Bouftene, between Azrou and Ras el Ma, Forêt Azrou, HA, Arround (Skoutana), Jebel Likount, Asni; Mouna 1998; Grabener 2017; MA (Tighassaline, El-Herri, Aïn Leuh-Tagounit, Meknès, Ifrane (NPI)), HA (40 km SW Ouarzazate, Marrakech, Ouirgane, Lakhdar, N Demnate, Oukaimeden), AA (Agadir, Oulma) – PCPT

Gonia bimaculata Wiedemann, 1819

= Gonia cilipeda Rondani, in Séguy 1953a: 91

Séguy 1930a, AP, Rabat, MA, Tizi-s’Tkrine, Tizi-n’Bouftene, between Azrou and Ras el Ma, Forêt Azrou, Berkane, HA, Arround (Skoutana), Jebel Likount, Asni, Ouaouzert (Glaoua); Séguy 1953a, AP, Temara; AP (Rabat) – MISR; HA (S Asni Ouirgane, Marrakech), AA (80 km N Taroudant, Aoulouz), AA (15 km NW Zagora) – PCPT

Gonia capitata (De Geer, 1776)

Mouna 1998; MA (Ifrane) – MISR

Gonia maculipennis Egger, 1862*

MA

Gonia ornata Meigen, 1826

Séguy 1953a, AP, Rabat, MA, Ifrane, SA, Kelaâ M’Goum; MA (Ifrane, Forêt de Cèdres), HA (Oukaimeden (2600 m), Marrakech) – PCPT

Gonia vacua Meigen, 1826*

MA

Pales Robineau-Desvoidy, 1830

Pales pavida (Meigen, 1824)

IOBC-list 1 (1956); Cerretti 2005; MA (Fès, Sidi Harazem) – PCPT

Platymya Robineau-Desvoidy, 1830

Platymya antennata (Brauer & Bergenstamm, 1891)

Efetov and Tarmann 1999

Pseudogonia Brauer and von Bergenstamm, 1889

Pseudogonia rufifrons (Wiedemann, 1830)

De Lépiney and Mimeur 1932; IOBC-list 1 (1956); Mouna 1998; Tschorsnig 2017; Grabener 2017; AA (S Tafraoute, Aït Mansur, Agadir, Oulma) – PCPT

Sturmia Robineau-Desvoidy, 1830

Sturmia bella (Meigen, 1824)

Stefanescu et al. 2012; Tschorsnig 2017

Winthemiini

Nemorilla Rondani, 1856

Nemorilla maculosa (Meigen, 1824)

Kozlovsky and Rungs 1933; Brémond and Rungs 1938 [as N. floralis; probable misidentification]; IOBC list 1 (1956); Mouna 1998; EM (Oujda, Col de Jerada) – PCPT

Phasiinae

Cylindromyiini

Besseria Robineau-Desvoidy, 1830

Besseria lateritia (Meigen, 1824)*

HA

Cylindromyia Meigen, 1803

Cylindromyia bicolor (Olivier, 1812)

Séguy 1930a, AP, Rabat; Mouna 1998

Cylindromyia brassicaria (Fabricius, 1775)

Séguy 1930a, EM, Soufouloud, MA, Aharmoumou, Berrechid, Meknès, Sidi Bettache, Berkane, HA, Tenfecht, Ouaounzert, Marrakech, Asni; Séguy 1935a, AP, Gharb; Séguy 1941a, HA, Jebel Ayachi; Séguy 1949, SA, Guelmim; Dupuis 1963; Mouna 1998; MA (Sefrou) – MISR; AP (Essaouira, 4 km E Ounara), HA (10 km W Chichaoua, Oukaimeden), AA (140 km E Agadir, Aoulouz, Tizi-n’Tichka) – PCPT

Cylindromyia intermedia Meigen, 1824

Becker and Stein 1913, Rif, Tanger; HA (Ouirgane, Imlil, Tizi-n’Test), AA (10 km SE Ouarzazate (oasis), Taroudant) – PCPT

Cylindromyia maroccana Tschorsnig, 1997

Tschorsnig 1997, HA, Ouirgane, Tagadirt (1000 m)

Cylindromyia pilipes Loew, 1844

Becker and Stein 1913, Rif, Tanger; Herting and Dely-Draskovits 1993; Gilasian et al. 2013

Phania Meigen, 1824

Phania albisquama (Villeneuve, 1924)*

MA

Gymnosomatini

Clytiomya Rondani, 1861

Clytiomya continua (Panzer, 1798)

= Clytiomyia dalmatica Robineau-Desvoidy, in Séguy 1935a: 119

Séguy 1935a, AP, Gharb; Mouna 1998; AP (Rabat) – MISR

Clytiomya sola (Rondani, 1861)

Séguy 1935a; Dupuis 1963; MA (Khénifra, Tighassaline) – PCPT

Ectophasia Townsend, 1912

Ectophasia crassipennis (Fabricius, 1794)

= Phasia crassipennis Fabricius, in Séguy 1930a: 141

Séguy 1930a, MA, Aïn Leuh; Mouna 1998

Eliozeta Rondani, 1856

Eliozeta helluo (Fabricius, 1805)

= Clytiomyia helluo Fabricius, in Séguy 1935: 119

Séguy 1930a, MA, Meknès (Aïn Sferguila); Jourdan 1935; Séguy 1935a, AP, Gharb; Thompson 1950; Dupuis 1963; Mouna 1998; Tschorsnig 2017

Gymnosoma Meigen, 1803

Gymnosoma carpocoridis Dupuis, 1961

Dupuis 1963; Herting and Dely-Draskovits 1993

Gymnosoma clavatum (Rohdendorf, 1947)

Dupuis 1963; Grabener 2017; MA (Meknès, Ifrane (NPI)), AA (140 km E Agadir, Aoulouz), AA (80 km S Zagora, Oued Draa, Mhamid) – PCPT

Gymnosoma dolycoridis Dupuis, 1960

Dupuis 1963; MA (Fès, Sidi Harazem) – PCPT

Gymnosoma rotundatum Linnaeus, 175864

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, Rif, Tanger, AP, Mogador, MA, Tizi-s’Tkrine, Sidi Bettache, Moulay Aïn Djemine, HA, Asni; Séguy 1934b; Séguy 1935a; Thompson 1950; Mouna 1998; AP (Mogador), MA (Maghrawa) – MISR; MA (Meknès, Ifrane (NPI)) – PCPT

Gymnosoma rungsi (Mesnil, 1952)

= Rhodogyne rungsi (Mesnil), in Mesnil 1952: 151

Mesnil 1952, AP, Rabat; Dupuis 1963; Herting and Dely-Draskovits 1993

Leucostomatini

Clairvillia Robineau-Desvoidy, 1830

Clairvillia biguttata (Meigen, 1824)*

HA

Dionomelia Kugler, 1978

Dionomelia hennigi Kugler, 1978*

SA

Leucostoma Meigen, 1803

Leucostoma abbreviatum Herting, 1971

Ziegler 2012

Leucostoma crassum Kugler, 1966*

HA

Leucostoma obsidianum (Wiedemann, 1830)

Ebejer et al. 2019, AA, Ziz river (10 km S of Errachidia, 1008 m)

Leucostoma tetraptera (Meigen, 1824)

Dupuis 1953; Ebejer et al. 2019, Rif, Barrage Smir (27 m)

Weberia Robineau-Desvoidy, 1830

Weberia digramma (Meigen, 1824)*

AA

Phasiini

Elomya Robineau-Desvoidy, 1830

Elomya lateralis (Meigen, 1824)

Séguy 1930a, AP, From Zarjoulea to Larache, MA, Berkane; Dupuis 1952; Dupuis 1963; Herting and Dely-Draskovits 1993; Mouna 1998; Cerretti and Ziegler 2004; MA (Khénifra, Tighassaline, Meknès, Ifrane (NPI)), HA (Marrakech, Ouirgane) – PCPT

Phasia Latreille, 1804

Phasia mesnili (Draber-Monko, 1965)

Sun and Marshall 2003, HA; AP (10 km E Essaouira), AA (S Tafraoute, Aït Mansur, S Aït-Baha) – PCPT

Phasia obesa (Fabricius, 1798)

Sun and Marshall 2003, HA, Asni

Phasia pusilla Meigen, 1824

Dupuis 1963; Sun and Marshall 2003, MA

Phasia subcoleoptrata (Linnaeus, 1767)

Dupuis 1963; Herting and Dely-Draskovits 1993; Sun and Marshall 2003, MA; Cerretti and Ziegler 2004

Phasia venturii (Draber-Monko, 1965)

Sun and Marshall 2003, HA, Asni; AP (Essaouira, 4 km E Ounara), AA (11 km NW Taliouine, 10 km SE Aït-Ourir) – PCPT

Trichopodini

Trichopoda Berthold, 1827

Trichopoda pennipes (Fabricius, 1794)

Ebejer et al. 2019, Rif, Tahaddart (8 m)

Xystini

Xysta Meigen, 1824

Xysta holosericea (Fabricius, 1805)*

HA

Tachininae

Graphogastrini

Graphogaster Rondani, 1868

Graphogaster vestita Rondani, 1868*

MA

Phytomyptera Rondani, 1845

Phytomyptera nigrina (Meigen, 1824)

= Phytomyptera nitidiventris Rondani, in Bléton and Fieuzet 1939: 64

Bléton and Fieuzet 1939; Mouna 1998

Leskiini

Aphria Robineau-Desvoidy, 1830

Aphria longirostris (Meigen, 1824)

Ebejer et al. 2019, Rif, Jnane Niche (46 m)

Bithia Robineau-Desvoidy, 1863

Bithia demotica (Egger, 1861)

Tschorsnig and Bläsius 2001; IOBC-list 14 (2005); Tschorsnig 2017

Bithia modesta (Meigen, 1824)

Tschorsnig and Bläsius 2001; Tschorsnig 2017

Linnaemyini + Ernestiini

Gymnochaeta Robineau-Desvoidy, 1830

Gymnochaeta viridis Fallén, 1810

Séguy 1930a, HA, Arround (Skoutana); Mouna 1998

Linnaemya Robineau-Desvoidy, 1830

Linnaemya soror Zimin, 1954*

MA, HA, AA

Loewia Egger, 1856

Loewia setibarba Egger, 1856

Becker and Stein 1913, Rif, Tanger

Panzeria Robineau-Desvoidy, 1830

Panzeria castellana (Strobl, 1906)*

HA

Panzeria nemorum (Meigen, 1824)*

MA

Zophomyia Macquart, 1835

Zophomyia temula (Scopoli, 1763)

Séguy 1930a, AP, Casablanca, MA, Meknès; Mouna 1998; MA (Khénifra, Tighassaline, Meknès, Ifrane (NPI)) – PCPT

Macquartiini

Macquartia Robineau-Desvoidy, 1830

Macquartia chalconota (Meigen, 1824)

Ebejer et al. 2019, Rif, Smir lagoon; HA (Marrakech, Ouirgane, Tizi-n’Test), AA (Taroudant) – PCPT

Macquartia macularis Villeneuve, 1926

Herting and Dely-Draskovits 1993

Macquartia tessellum (Meigen, 1824)

= Macquartia brevicornis Macquart, in Séguy 1941d: 23

Séguy 1941d, MA, Meknès, HA, Tizi-n’Test; Mouna 1998; HA (Imlil, S Asni, Tizi-n’Test), AA (Taroudant) – PCPT

Megaprosopini

Microphthalma Macquart, 1844

Microphthalma europaea Egger, 1860

Ebejer et al. 2019, AA, Ziz river (30 km N of Erfoud, 894 m)

Minthoini

Hyperaea Robineau-Desvoidy, 1863

Hyperaea femoralis (Meigen, 1824)

Herting and Dely-Draskovits 1993

Mintho Robineau-Desvoidy, 1830

Mintho compressa (Fabricius, 1787)

Walker 1849; Herting and Dely-Draskovits 1993

Mintho rufiventris (Fallén, 1817)

= Mintho praeceps (Scopoli, 1763), in Séguy 1930a: 143; Séguy 1953a: 91

Séguy 1930a, AP, Rabat, Casablanca, MA, Meknès; Séguy 1953a, SA, El Aïoun du Draa; Séguy 1949a, AA, Tata; Mouna 1998; Dawah 2011; AP (Rabat, Salé), MA (Meknès) – MISR; MA (Meknès, Ifrane (NPI)), AA (11 km NW Taliouine) – PCPT

Minthodes Brauer & Bergenstamm, 1889

Minthodes numidica Villeneuve, 1932*

AA

Minthodes setifacies Mesnil, 1939

= Minthodes (Myxominthodes) setifacies Mesnil, in Mesnil 1939: 211

Mesnil 1939, MA, Forêt Azrou; Herting and Dely-Draskovits 1993

Plesina Meigen, 1838

Plesina phalerata (Meigen, 1824)

Herting and Dely-Draskovits 1993; Cerretti and Tschorsnig 2008

Pseudomintho Brauer & Bergenstamm, 1889

Pseudomintho diversipes (Strobl, 1889)

Ebejer et al. 2019, Rif, Moulay Abdelsalam (PNPB, 965 m); AP (Essaouira, 4 km E Ounara) – PCPT

Siphonini

Actia Robineau-Desvoidy, 1830

Actia infantula (Zetterstedt, 1844)

Ebejer et al. 2019, Rif, Tanger (Douar Dakchire forest, 320 m)

Peribaea Robineau-Desvoidy, 1863

Peribaea apicalis Robineau-Desvoidy, 1863

Ebejer et al. 2019, Rif, Dardara (484 m)

Peribaea tibialis (Robineau-Desvoidy, 1851)

Draber-Mońko 2011

Siphona Meigen, 1803

Siphona geniculata (De Geer, 1776)65

Séguy 1930a, HA; Mouna 1998; HA (Vallée Oued N’fis) – MISR

Siphona maroccana Cerretti & Tschorsnig, 2007

Cerretti and Tshorsnig 2007, HA, Asif Mellah, W Tizi-n’Tichka

Siphona variata Andersen, 1982

Ebejer et al. 2019, Rif, Sidi Yahia Aârab (377 m), Oued Kbir (PNPB, 157 m)

Tachinini

Germaria Robineau-Desvoidy, 1830

Germaria barbara Mesnil, 1963*

HA

Peleteria Robineau-Desvoidy, 1830

Peleteria ruficornis (Macquart, 1835)*

HA, AA

Tachina Meigen, 1803

Tachina corsicana (Villeneuve, 1931)*

HA, AA

Tachina fera (Linnaeus, 1761)

Becker and Stein 1913, Rif, Tanger; Séguy 1930a, MA, Tizi-s’Tkrine, Forêt Tiffert, Sidi Bettache, HA, Arround (Skoutana), Jebel Likount; Séguy 1953a, MA, Ifrane (1650 m); Mouna 1998; Rif (fir forest of Talassemtane), AP (Dradek) – MISR (MA, Meknès); MA (Ifrane (NPI)), Béni Mellal, Bin-el-Ouidane), HA (Marrakech, Ouirgane) – PCPT

Tachina magnicornis (Zetterstedt, 1844)

Séguy 1930a, MA, Ras el Ksar, Aïn Leuh, Sidi Bettache; Mouna 1998; MA (Béni Mellal, El Ksiba, 5 km N) – PCPT

Tachina praeceps Meigen, 1824

HA, AA

Triarthriini

Lissoglossa Villeneuve, 1912

Lissoglossa bequaerti Villeneuve, 1912

Herting and Dely-Draskovits 1993

New records for Morocco

The data added under the abbreviation “PCPT” (for “personal communication Hans-Peter Tschorsnig”) are based on (unpublished) material which was identified by HPT for several collectors (M. Hauser, M. Hradský, C.F. Kassebeer, U. Koschwitz, J.A.W. Lucas, G. Miksch, H. and T. v. Oorschot, C. Schmid-Egger, M. Schwarz, K. Špatenka, V. Vrabec) during the last ~ 30 years. Usually only a few duplicate specimens were retained in the collection of SMNS. The main part was sent back to the collectors, but the data were noted by HPT on handwritten lists.

Estheria iberica Tschorsnig, 2003

Middle Atlas: Ifrane, National Park of Ifrane, 19.ix.1989, K. Špatenka leg, 1 specimen, PCPT.

Athrycia trepida (Meigen, 1824)

Middle Atlas: Meknès; Ifrane, National Park of Ifrane, 22.v.1995, C. Kassebeer leg., 1 specimen, PCPT.

Eriothrix rufomaculata (De Geer, 1776)

Middle Atlas: Ifrane, Forêt de Cèdres, 29.iv.1999, V. Vrabec leg., 1 specimen, PCPT.

High Atlas: Marrakech, Oukaimeden, 19.v.1995, C. Kassebeer leg., 7 specimens, PCPT.

Kirbya moerens (Meigen, 1830)

Middle Atlas: Ifrane, Forêt de Cèdres, 29.iv.1999, V. Vrabec leg., 1 specimen, PCPT.

Carcelia lucorum (Meigen, 1824)

High Atlas: Marrakech, Imlil, S Asni, 24.iii.1995, C. Kassebeer leg., 1 specimen, PCPT.

Drino galii (Brauer & Bergenstamm, 1891)

High Atlas: Marrakech, Ouirgane, 24.iii.1995, C. Kassebeer leg., 1 specimen, PCPT.

Anti Atlas: 11 km NW Taliouine; Agadir, Ameskroud, 17.v.1995, C. Kassebeer leg., 2 specimens, PCPT.

Chetogena media Rondani, 1859

Middle Atlas: Béni Mellal, El Ksiba, 30.iii.1995, C. Kassebeer leg., 1 specimen, PCPT.

Exorista rendina (Herting, 1975)

Anti Atlas: 11 km NW Taliouine, 15.iii.1997, M. Hauser leg., 1 male in SMNS; 10 km NE Tafraoute, 14.iii.1997, G. Miksch leg., 1 male in SMNS, PCPT.

Anurophylla aprica (Villeneuve, 1912)

Middle Atlas: Béni Mellal, Afourer, 28.iii.1995, C. Kassebeer leg., 1 female in SMNS, PCPT.

Blepharipa pratensis (Meigen, 1824)

High Atlas: Tizi-n’Test, 2000 m, 21.v.1995, M. Hauser leg., 2 specimens, PCPT.

Anti Atlas: Taroudant, PCPT.

Gonia maculipennis Egger, 1862

Middle Atlas: Ifrane, Forêt de Cèdres, 29.iv.1999, V. Vrabec leg., 1 female in SMNS, PCPT.

Gonia vacua Meigen, 1826

Middle Atlas: Meknès; Ifrane, National Park of Ifrane, 29.iii.1995 and 22.v.1995, C. Kassebeerleg., 2 specimens, PCPT.

Besseria lateritia (Meigen, 1824)

High Atlas: SE Asni, Imlil, 23.v.1995, M. Hauser leg., 2 specimens; Marrakech, Lakhdar, N Demnate, 27.iii.1995, C. Kassebeer leg., 1 specimen, PCPT.

Phania albisquama (Villeneuve, 1924)

Middle Atlas: Ifrane, Forêt de Cèdres, 29.iv.1999, V. Vrabec and L. Vrabcová leg., 1 specimen, PCPT.

Clairvillia biguttata (Meigen, 1824)

High Atlas: Marrakech, Lakhdar, N Demnate, 27.iii.1995, C. Kassebeer leg, 1 specimen, PCPT.

Dionomelia hennigi Kugler, 1978

SA: Boujdour, 8.v.1999, V. Vrabec leg, 1 male in SMNS, PCPT.

Leucostoma crassum Kugler, 1966

High Atlas: Tizi-n’Test, pass 23.vi.1996, U. Koschwitz leg., 1 male in SMN, PCPT.

Weberia digramma (Meigen, 1824)

Anti Atlas: 10 km NW Aït-Baha, PCPT.

Xysta holosericea (Fabricius, 1805)

High Atlas: Marrakech, Lakhdar, N Demnate, 27.iii.1995, C. Kassebeer leg, 1 specimen, PCPT.

Linnaemya soror Zimin, 1954

Middle Atlas: Béni Mellal, El Ksiba, 5 km N; Béni Mellal, Afourer; Khénifra, Tighassaline; Meknès; National Park of Ifrane.

High Atlas: Marrakech, Ouirgane; Marrakech, Tagaddirt, S Asni; Marrakech, Lakhdar, N Demnate.

Anti Atlas: 11 km NW Taliouine, all C. Kassebeer leg., 57 specimens (collected between 25.iii. and 23.v.1995), PCPT.

Panzeria castellana (Strobl, 1906)

High Atlas: Marrakech, Ouirgane, 26.iii.1995, C. Kassebeer leg., 1 specimen, PCPT.

Panzeria nemorum (Meigen, 1824)

Middle Atlas: Meknès; National Park of Ifrane, 22.v.1995, C. Kassebeer leg., 1 specimen, PCPT.

Graphogaster vestita Rondani, 1868

Middle Atlas: Ifrane, Forêt de Cèdres, 29.iv.1999, V. Vrabec leg., 1 specimen, PCPT.

Minthodes numidica Villeneuve, 1932

Anti Atlas: S Aït-Baha, PCPT.

Germaria barbara Mesnil, 1963

High Atlas: S Tizi-n’Test, 1900 m, PCPT.

Peleteria ruficornis (Macquart, 1835)

High Atlas: Marrakech, Ouirgane; Marrakech, Tagaddirt, S Asni; Tizi-n’Test.

Anti Atlas: Taroudant, all C. Kassebeer leg., 6 specimens (collected between 28.ix.1994 and 1.iv.1995), PCPT.

Tachina corsicana (Villeneuve, 1931)

High Atlas: Marrakech, Oukaimeden, 19.v.1995, C. Kassebeer leg., 1 specimen; Tizi-n’Test. Anti Atlas: Taroudant, 21.v.1995, M. Hauser leg., 2 specimens, PCPT.

Tachina praeceps Meigen, 1824

High Atlas: Marrakech, Oukaimeden, 2500 m, 27.vi.1987, M. Schwarz leg., 1 specimen; Tizi-n’Test.

Anti Atlas: Taroudant, 29.vi.1987, M. Schwarz leg., 1 specimen, PCPT.

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1  Dicranomyia  majuscula Pierre, is a doubtful species of which the identity is not known. There is only one record, from Pierre 1924a, all other citations come from this record.
2  Tipula ( Emodotipula)  obscuriventris is widespread in the western Palaearctic, but reaches its south-western limits in the the Spanish Pyrenees. Therefore, it is very unlikely that  obscuriventris actually occurs in Morocco, disregarding the references of Oosterbroek and Theowald (1992) and CCW 2020 (which provide Morocco with a question mark for  obscuriventris). Thus far, the only positively identified  Emodotipula species from Morocco and the Rif region for which evidence in the form of specimens is available is T. leo.
3 The following species were considered as Moroccan on the basis of the species represented in the collections of the MISR (Rabat, Morocco): Aspistes berolinensis Meigen, 1818, Apiloscatopse  flavicollis (Meigen, 1818),  Reichertella pulicaria (Loew, 1846). However, none of these species is represented by specimens originating from Morocco. Accordingly, they are excluded from the present checklist.
4  Prosimulium  latimucro (Enderlein, 1925) is a species complex, but only one cytoform has been found in Morocco. It is called  Prosimulium  latimucro cytoform C Kúdela, Kúdelová & Adler, 2017 (Arrondo and de Blas Giral 2017: 16).
5 The original reports of  Prosimulium  rachiliense in Morocco appeared under the name P. rufipes. The chromosomes described by Adler and Belqat (2001), however, indicate that populations in Morocco’s Rif Mountains are not P. rufipes, but are a species closer to P. rachiliense (Belqat et al. 2018; Adler and Şirin 2014).
6  Metacnephia nuragica was listed with faunistic and bibliographical details in Belqat et al. (2011: 48) but its identity is uncertain.
7 The identity of  Thaumalea spinata is not finally clarified. No specimen outside the French Alps was found in Vaillant’s collection.
8 The species of this family are considered as Moroccan on the basis that they are represented by specimens in the collections of the MISR (Mouna 1998). However, it has not been possible to confirm if the specimens present in this collection actually originated from Morocco.
9 Three specimens identified as Rhyphus fenestralis are present in the collection of MISR (Rabat, Morocco). However, genital examination is required for identification of S. fenestralis and the closely related widespread S. cinctus (Fabricius, 1787). Both species could probably occur in Morocco.
10 The following species have also been considered as Moroccan on the basis of specimens present in the collections of the MISR:  Bibio clavipes Meigen, 1818, B. ferruginatus Linnaeus, 1767, B. fulvipes Zetterstedt, 1838, B. hybridus Haliday, 1833, B. lacteipennis Zetterstedt, 1850, B. nigriventris Haliday, 1833, B. pomonae (Fabricius, 1775), B. reticulatus Loew, 1846, B. siculus Loew, 1846, B. varipes Meigen, 1830, B. villosus Meigen, 1818, D. femoratus Meigen, 1804, D. humeralis Zetterstedt, 1850 (Mouna 1998). However, none of these species is represented in this collection by specimens originating from Morocco. Accordingly, they are excluded from the present checklist. An additional, still undescribed, species of  Bibio has been collected in the Rif Mountains (Ketama) in 2004 and 2012 by K. Kettani and will be described elsewhere.
11 The identity of the material of D. bispinosus mentioned in Séguy (1953a) should be checked, as D. clavicornus Skartveit & Kaplan, 1996, a closely related species that may be present in Morocco, has been described from Israel since this publication.
12  Pangonius sobradieli, said to be known from Morocco, was considered to be the same species as P. powelli by Chvála et al. (1972: 90).
13  Vermileo vermileo (Linnaeus, 1758) is a species complex currently undergoing revision (Kehlmaier pers. comm.); additionally, V. vermileo is possibly not a species that occurs in Morocco.
14 The record of  Usia  forcipata is doubtful as this species is probably endemic to Greece (Gibbs 2011).
15  Usia ( Usia)  bicolor is an east Mediterranean species (Gibbs 2014) and the record from Morocco is most likely a misidentification.
16  Thereva  bipunctata has been considered a junior synonym of unica, but these are valid sister species because of differences in the aedeagus between the southern and northern populations (Ebejer pers. comm.).
17  Thereva  brevicornis may be a misidentification of authors.  Thereva  brevicornis is more central European in distribution, and is not found in Spain.
18  Thereva  funebris,  Thereva  graeca, and  Thereva  plebeja are cited only on the basis of the list of Diptera species present in the collections of the MISR published by Mouna (1998). The identifications of this material could not be verified.
19  Thereva  strigata may be a misidentification of authors.
20 The following species records from Morocco are known to the authors by various checklists with no material provided:  Argyra  grata Loew, 1857;  Sybistroma  obscurellum Fallén, 1823;  Tachytrechus  consobrinus (Haliday, 1851);  Medetera  petrophila Kowarz, 1877;  Rhaphium  appendiculatum Zetterstedt, 1849. Their presence in Morocco must be confirmed.
21 Hydrophorus  viridis may be a misidentification from Boumezzough and Vaillant (1986a) as this species is doubtfully present in North Africa. It does not reach Iberia in the west or Greece in the east.
22 Pollet and Stark (2018) have changed the species concept of  Orthoceratium lacustre (Scopoli, 1763). As a result, all previous records of the species from Atlantic countries and almost all records from Africa (except for a female record from Tunisia) have been assigned to O. sabulosum (Becker, 1907). The Moroccan material also belongs to the latter species (Martin Ebejer, pers. comm.).
23 Morocco was most probably erroneously included into the distribution section for  Medetera  flavipes Meigen, 1824 and  Medetera  pallipes (Zetterstedt, 1843) in some papers (e.g. Kazerani et al. 2014). M.  pallipes was recorded in the country by Nourti et al. (2019a). M.  flavipes is not included in the present Catalog.
24 Medetera  truncorum Meigen, 1824 was identified by a female only (Ebejer et al. 2019). However,  Medetera  truncorum is a group of closely related species present in the Mediterranean that can be identified only by fine structures of the male hypopygium (Negrobov and Naglis 2016). Its presence in Morocco must be confirmed.
25 Sciapus  longulus (Fallén, 1823) was identified by females only (Pârvu et al. 2006). Its presence in Morocco must be confirmed, as in most species in this genus from the Mediterranean they can only be distinguished by males (Grichanov and Negrobov 2014).
26 Campsicnemus  curvipes (Fallén, 1823) was recorded by Parent (1924) from Algeciras (Spain) and mentioned by Parent (1927) as collected from “Morocco: Algésiras”.  Argyra  grata Loew, 1857 was recorded by Parent (1927) from Algeria. These species may occur in Morocco.
27 Crossopalpus aenescens (Wiedemann, 1830) was recorded by Séguy (1941d: 12) under the name of  Drapetis aenescens, confusing it with C. aeneus. These two species have often been confused by authors. This is a misidentification, as C. aenescens is an Afrotropical species.
28 Stilpon demnatensis Vaillant, 1956 was indicated as a nomen  nudum by Cumming and Cooper (1992).
29 Becker and Stein’s record of  Tachydromia annulimana is retained, but is considered doubtful as  annulimana is a European species. The record could equally refer to T. umbrarum Haliday, 1833 another European species which has often been misidentified as  annulimana by authors.
30 Diplonevra tangeriana (Becker, 1913) was recorded from Morocco, but fresh specimens are required to ascertain its identity.
31 Microsania raclinae Collart is listed in Mouna 1998, but raclinae is a nomen nudum and the material upon which this citation was given is unknown to us. It cannot be determined whether or not the error is with the species name itself while the genus  Microsania is actually represented in Morocco.
32 The range of  Orthonevra  brevicornis does not extend to Iberia. The identification of the single specimen from Morocco was based on a female in Sahib et al. (2020). The status of this species in Morocco needs to be confirmed.
33 Eristalis  jugorum is not known closer to Morocco than the mountains of northern Spain and Turkey. Its occurrence in high mountains in Morocco is possible, but confirmation of the records would be worthwhile.
34 Eumerus  strigatus is not recorded from Morocco by Claußen and Hauser 1990, or by Claußen 1989b. However, it is listed for Morocco by Claußen 1989b, based on records from earlier authors (Kanervo 1939; Timon-David 1951).  Eumerus strigatus is now known to be a complex of at least four species (E. consimilis, E. narcissi, E. sogdianus, E. strigatus). Any of those four species might be present in Morocco, but E. strigatus is the least likely to be present, given its rarity in the Mediterranean zone of Europe.
35 Revision of the  Merodon constans species group (Vujić et al. 2020) has revealed the existence of cryptic taxa whose status is as yet uncertain. The name  constans can only be applied provisionally to the species recorded from Morocco, until these issues are resolved and the identity of Moroccan specimens can be meaningfully re-assessed.
36 Revision of  Merodon shows that M. aeneus (often also referred to as M. aureus) is confined to the Alps of southern Europe. Two very similar species, M. pumilus and M. unicolor, are known from N. Africa. Moroccan records of M. aeneus almost certainly refer to either M. pumilus or M. unicolor.
37 Merodon arrasus is probably a junior synonym of  Merodon eques Fabricius, 1805 (Ebejer and Bensusan 2010).
38 Revision of the  Merodon avidus complex by Popović et al. (2015) demonstrates the presence of M. ibericus in Morocco, but not M. avidus. Given the confusion between these species, earlier records of M. avidus from Morocco could refer either to  avidus or  ibericus.
39 Platycheirus albimanus is removed from the list for Morocco by Kassebeer 1998, who describes two new Moroccan species closely related to P. albimanus.
40 Eupeodes punctifer is almost confined to Northern Europe, north of the Arctic Circle, and a record from Morocco seems almost impossible. It is not possible to suggest on which species the Moroccan record might be based.
41 Although the faunistic knowledge is good in that there are many species recorded, this does not mean the identifications and taxonomy are all correct (Stuke, in litteris 26.v.2016). Given several recent revisions, especially within the genera  Myopa,  Physocephala, and  Thecophora, most of the Moroccan species listed here are in need of careful reconsideration.
42  Platystoma idia is possibly a synonym of P. meridionale Hendel.
43  Myopites apicatus is probably a synonym of M.  inulaedyssentericae Blot and at least the Moroccan record appears to be the latter species. A revision of  Myopites is in preparation (Korneyev, pers. comm.).
44 Records of  Urophora congrua are possibly based on misidentifications. Confirmation requires re-examination of the specimens.
45  Euleia marmorea may be a synonym of  Euleia heraclei.
46 Minettia flavipalpis and M.  plumicornis are not confirmed for Morocco; see Ebejer and Kettani 2019a.
47  Themira paludosa is a dubious record since T. paludosa seems reliably recorded only from NW Europe (Estonia, Finland, NW Russia).
48 Many species ( Chlorops pumilionis,  Eutropha albipilosa,  Meromyza athletica,  Meromyza curvinervis,  Elachiptera cornuta,  Oscinisoma cognatum) are cited only on the basis of the list of Diptera species present in the collections of the MISR (Rabat, Morocco) published by Mouna (1998). The identification of this material could not be verified. However, the presence of most of these species is possible in Morocco.
49  Assuania melanoleuca comb. nov.: a female type specimen of  Chlorops melanoleucus is a species of  Assuania (Nartshuk, pers. comm.), see also Ebejer and Kettani (2016: 226). It may belong to A. thalhammeri with its three synonyms A. glabra Becker, 1903, A. frontata Becker, 1907, and A. siciliensis Duda, 1933 or to one of the further five  Assuania spp. (Sabrosky 1980: 707) present on the African continent. Dawah and Abdullah (2006: 30) and Deeming and Al-Dhafer (2012: 56, 88) reported an unidentifiable species from the Arabian Peninsula. Until a revision of this genus is made, A. melanoleuca must be treated as a valid taxon.
50  Epimadiza nigrescens: Nartshuk (1984: 236) erroneously sank Oscinosoma [sic! misspelling of Séguy]  anthracias Séguy, 1949 as a junior synonym of Siphonella oscinina (Fallén, 1820), although this synonymy had not been proposed previously. This error was clarified by Ebejer and Kettani (2016: 227) after correspondence with Nartshuk, but the puzzle increased as the latter authors introduced O. anthracias as a new junior synonym of  Polyodaspis ruficornis after personal correspondence of MJE with E.P. Nartshuk who had examined the type of O. anthracias in the year 2001. Both had overlooked the hidden synonymisation of  Oscinisoma anthracias with  Epimadiza nigrescens by Sabrosky (1965: 406), cited by Sabrosky (1980: 699) and by Dawah and Abdullah (2006: 26). Séguy did not mention in his description if he had only one male or more. Sabrosky may have studied a male other than the one studied by Nartshuk. The type(s) of O. anthracias should be examined again. If there is more than one specimen this might explain why two different senior taxa resulted. Here we follow the oldest examination of a type specimen by Sabrosky.
51  Polyodaspis ruficornis (Macquart), a widely distributed species, was reported by Ebejer and Kettani (2016) only on the basis of the literature, but as explained in footnote 38 it must be deleted from the list of Moroccan Diptera as the cited records are based on an earlier misidentification.
52 The record of  Delia flavogrisea needs to be verified. No details or genitalia figures are given. Although it is possible that this species could occur in Morocco, it is not recorded from Spain. Described from northern Sweden, it was recorded by G.C.D. Griffiths from Yukon, northern Canada (1993, Flies of the Nearctic Region Vol. VIII, Part 2, No 10: 1509), where excellent figures of the genitalia are given. It has also been recorded from northern China.
53 We do not have any knowledge of  Fannia cothurnata in Morocco. Identification is difficult, but it could be correct.
54 We do not have any knowledge of  Fannia sociella in Morocco. Its occurrence there seems doubtful.
55  Coenosia pumila requires verification. It is not known from Spain and is distributed in temperate areas of Europe. It may be found in the High Atlas Mts, but specimens from there require confirmation.
56  Morellia  simplex is almost certainly a misidentification of  Morellia  asetosa Baranov, 1925, which replaces it in the south and south-east of the West Palaearctic Region.
57  Phaonia  cincta requires verification. It is known from Spain and from the Middle East, and its occurrence in Morocco is possible.
58 According to Rognes (2002: 38)  Rhyncomya  callopis records may refer to similar species with spines on the surstylus.
59 The records of  Sarcophila latifrons (Fallén) require verification. The genus  Sarcophila is in need of revision and is likely much more diverse in the Mediterranean area than previously thought, so older records of S. latifrons from Morocco must be considered uncertain.
60 This record requires verification, as  Sarcophaga ( Liosarcophaga tuberosa has often been confused with other species in the literature.
61 After checking photographs of the male terminalia of the specimen reported by Ebejer et al. (2019) as  Sarcophaga ( Thyrsocnema incisilobata Pandellé, 1896, we consider that it most likely represents an undescribed species.
62  Estheria picta: the identity of this species is not confirmed; see note in Tschorsnig 2017.
63 The following species  Thelaira haematodes,  Carcelia iliaca,  Drino gilva, and  Drino imberbis, have been considered as Moroccan on the basis of specimens represented in the collection of MISR (Rabat, Morocco) or cited in some old Moroccan references: However, the identifications are not yet confirmed.
64  Gymnosoma rotundatum older identifications of G. rotundatum are not reliable. Possibly they were the true G. rotundatum, but it is more likely that other species of the genus  Gymnosoma were concerned.
65  Siphona geniculata: the identifications of this species in the cited references were made before the revision of the genus by Andersen (1996) and may be mistaken. Although this species could be present in Morocco, this needs to be confirmed.
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