The Whiteflies of the World and Their Host - Keys - Lucid Key Server ...

Version 070606 

Last Revised: June 11, 2007 

THE WHITEFLIES (HEMIPTERA: ALEYRODIDAE) 

OF THE WORLD 

and Their Host Plants and Natural Enemies 

GREGORY A. EVANS 

USDA/Animal Plant Health Inspection Service (APHIS)

2 

Introduction 

Mound & Halsey (1978) listed 1156 species in 126 genera of whiteflies (Aleyrodidae) in 

the latest catalogue of the whiteflies of the world and their host plants and natural enemies. Since 

then, several new genera and species have been described and others have been synonymized 

with previously described taxa. Martin & Mound (2007) recently published a checklist of the 

whiteflies of the world that includes 1556 species in 161 genera belonging to three extant (living) 

subfamilies (Aleurodicinae and Aleyrodinae), and one fossil (non-living) subfamily 

(Bernaeinae). The subfamily Aleurodicinae is primarily New World in distribution and includes 

118 species in 18 genera. The subfamily Aleyrodinae is worldwide in distribution and includes 

1424 species in 148 genera. The subfamily Udamosellinae includes 2 South American species in 

the genus Udamoselis. 

The information included herein was extracted from the literature and includes 

unpublished records of whiteflies and their host plants, distribution and natural enemies taken 

from the FSCA, USNM collections, in addition to records of whitefly parasitoids in my personal 

collection. I have also included records of species intercepted at U.S. ports of entry on shipments 

of imported products, but have listed them separately from the other host and distribution 

records. I wanted to include these records to indicate the possible occurrence of these species in 

the supposed country of origin, but with the understanding that the actual occurrence of these 

species in each country must be confirmed by collections made within each country. Readers are 

also referred to the USDA/Systematic Entomology Laboratory Whitefly Collection website: 

[http://www.sel.barc.usda.gov/whitefly/wfframe.htm] to search the USNM holdings. 

This project began several years ago with the initial support of the National Biological 

Control Institute (USDA/NBCI). Several people have been instrumental in bringing it about. I 

thank: Avas Hamon (FSCA), Steve Nakahara and Louise Russell (USDA/SEL) who identified 

most of the whitefly species in the FSCA and USNM collections; John Dooley (USDA/APHIS), 

who identified many of the whiteflies intercepted at the U.S. ports of entry; Bruce Sutton for his 

help in creating the FILEMAKER PRO search and report forms used in the website; Bob 

Carlson (USDA/SEL) for his help in improving and maintaining the website and the many 

people who have collected whiteflies and their parasitoids throughout the world. I wish to 

especially thank Dr. Anil K. Dubey (Indian Institute of Science) for his comments, corrections 

and additions to the manuscript, and for all of the time and effort that he has put into this work. 

This document was created by merging the records in the database, accessed at 

[http://www.sel.barc.usda.gov:591/1WF/whitefly_catalog.htm], into a word processing 

document, which were then formatted and edited. It was created to compile the information in 

the database/website and make it readily and freely available to researchers and other readers. 

You are free to copy it, use it, and share it with anyone you wish. I will continually be updating 

and editing the information and will post the new versions on my whitefly website, so that 

readers will be able to easily replace the previous version with the updated one. I hope that 

having this information readily available will encourage more systematic research on whiteflies 

and their natural enemies and international cooperation in the study and introduction of natural 

enemies to combat pest species. 

Please send comments, suggestions, additions and corrections to: 

Gregory.A.Evans@usda.gov

Museum, Collections and Depositories 

AEI, American Entomological Institute, Gainesville, Florida, USA 

AMS, Australian Museum, Sydney, Sydney, AUSTRALIA 

AMU, Aligarh Muslim University, Aligarh, INDIA 

ANIC, Australian National Insect Collection, CSIRO, Canberra, AUSTRALIA. 

ANSW, Department of Agriculture, Rydalmer, New South Wales, AUSTRALIA. 

ASCT, The Agricultural Scientific Collections Trust, NSW, Orange, AUSTRALIA. 

BCRI, Biological and Chemical Research Institute, Rydalmer, AUSTRALIA. 

BERL, Berlese Collection, Florence, Italy, ITALY. 

BHU, Humboldt-Universitat Museum, DDR 104, Berlin, GERMANY. 

BMCol, Bink-Moenen Collection, THE NETHERLANDS. 

BMNH, British Museum of Natural History, SW7 5BD London, UK, ENGLAND. 

BPBM, Bernice P. Bishop Museum, Honolulu, Hawaii, USA. 

BTM, Budapest Termeszettudomanyi Muzeum, Allatara, 1088, Budapest, HUNGARY. 

CAESC, Connecticut Agricultural Experiment Station Collection, Connecticut, USA. 

CAS, California Academy of Sciences, San Francisco, California, USA. 

CIRAD, Centre de cooperation Internationale en Recherce Agronomique pour le developpement, 

Montpellier, FRANCE. 

CNC, Canadian National Collection of Insects, Ottawa, CANADA. 

COHIC, Cohic Personal Collection. 

CORSTOM, Centre O.R.S.T.O.M, Noumea Department of Entomology, Noumea, NEW 

CALEDONIA. 

DSIR, Department of Scientific and Industrial Research, Entomology Division, Aukland, NEW 

ZEALAND. 

DIIZ, Department of Invertebrates of the Institute of Zoology of the Academy of Sciences of the 

Georgian, Georgia, USSR. 

DPBS, Department of Parasitology, Institute of Biomedical Services, Sao Paulo University, Sao 

Paolo, BRAZIL. 

DPPE, Department of Plant Pathology and Entomology, Agricultural College of Sweden, 

Uppsala, SWEDEN. 

EDAC, Department of Agriculture, Cairo, Egypt, Cairo, EGYPT. 

EIP, Eberswalde Institute fur Pflanzenschutzforschung, Eberswalde,Germany, Kleinmachnow, 

Eberswalde, GERMANY. 

EMBRAPA, Empresa Brasileira de Pesquisa Agropecuaria, BRAZIL. 

ESCALQ, Escola Superior de Agricultura 'Luis de Querioz', BRAZIL. 

FAU, Fujian Agricultural University, Fujian, CHINA. 

FSCA, Florida State Collection of Arthopods, Division of Plant Industry, PO Box 147100, 

Gainesville, Florida, USA. 

GRF, Gezira Research Farm, Wad Medani, SUDAN. 

HAL, Haliday Collection, Dublin, IRELAND. 

HBL, Hikosan Biological Laboratory, Kyushu University, Fukuoka, JAPAN. 

HDA, Honolulu Department of Agriculture, Honolulu, Hawaii, USA. 

HUZI, Hokkaido University, Zoological Institute, Faculty of Science, Hokkaido, JAPAN. 

IARI, Indian Agricultural Research Institute, New Delhi, INDIA. 

3

IBCK, Institute of Biological Control, Kyushu, JAPAN. 

IBIW, Institute of Biology of Inland Waters, SSR Academy of Science, USSR. 

ICIA, Instituto Canario de Investigaciones Agrarias, La Laguna, Tenerife, SPAIN. 

IDAV, David Collection, INDIA. 

IDEH, Forest Research Institute, Dehra Dun, INDIA. 

IESP, Istituto di Entomologia F. Silvestri, Portici, ITALY. 

IFRI, Forest Research Institute, Dehradun, INDIA. 

IHAY, Hayat Collection, Aligarh, INDIA. 

INBIO, InBio Collection, COSTA RICA. 

INHS, Illinois Natural History Survey, Champaign, IL, USA. 

IPPR, Institute for Plant Protection, Kleinmachnow, GERMANY. 

ISUN, Sundararaj Collection, INDIA. 

ISZAF, Istituto Sperimentale per la Zoologia Agraria, Firenze, ITALY. 

ITZ, Institute for Taxomomic Zoology, Amsterdam, THE NETHERLANDS. 

IZAS, Institute of Zoology, Academia Sinica, CHINA. 

MAGNT, Museum of Art Gallery of the Northern Territory, NT, Darwin, AUSTRALIA. 

MCZ, Museum of Comparative Zoology, at Harvard, Cambridge, MA, USA 

MEJMO, Museo Entomologico Jose M. Osorio, VENEZUELA 

MHN, Museum d'Historie Naturelle, Geneva, SWITZERLAND. 

MNCN, Museo Nacional de Ciencias Naturales, Madrid, SPAIN. 

MNHNP, Museum National d'Histoire Naturelle, Paris, FRANCE. 

MZU, Museo de Zoologia de Universidade de Sao Paolo, BRAZIL. 

NHM, The Natural History Museum, London, UK, ENGLAND. 

NHMZA, Natural History Museum, Zoologische Abteilung, Vienna, AUSTRIA. 

NIAS, National Institute of Environmental Sciences, Tsukuba, JAPAN. 

NRCBC, National Reference Centre for Biological Control, Colima, MEXICO. 

NSM, National Science Museum, Tokya, JAPAN. 

NTU, National Taiwan University, Taipei, TAIWAN. 

NZAC, New Zealand Arthropod Collection, Mt. Albert, Auckland, NEW ZEALAND. 

OUMNH, Oxford University Museum, Hope Entomological Collections, Oxford, ENGLAND. 

PUAL, Department of Entomology, University of Agriculture, Lyallpur, PAKISTAN. 

QDPI, Queensland Department of Primary Industry, Indooropilly, Queensland, AUSTRALIA. 

QMB, Queensland Museum, Brisbane, AUSTRALIA. 

RPS, Rapisarda Personal Collection, ITALY. 

REGU, Regu Personal Collection, INDIA. 

RMNH, Rijks-Museum van Natuurlijke Historie, Leiden, THE NETHERLANDS. 

SIE, Shanghai Institute of Entomology, Shanghai, CHINA. 

SMNS, Staatlichen Museum für Naturkunde, Stuttgart (Ludwigsburg), GERMANY. 

TAMU, Texas A&M University, College Station, TX, USA. 

TARI, Taiwan Agricultural Research Institute, Taichung, TAIWAN. 

TARIT, Taiwan Agricultural Research Institute, Taipei, TAIWAN. 

TUBI, Tottori University, Biological Institute, Faculty of Education, Tottori, JAPAN. 

UAF, Department of Entomology, University of Agriculture, Faisalabad, PAKISTAN. 

UCD, University of California, Davis, CA, USA. 

UCR, University of California, Riverside, CA, USA. 

4

UNLP, Universidad Nacional de La Plata, La Plata, ARGENTINA. 

UNP, Universita degli Studi di Napoli, Portici, ITALY. 

USNM, United States National Museum of Natural History, Washington, DC, USA. 

UTA, University of Tel Aviv, Tel Aviv, ISRAEL. 

WAM, Western Australian Museum, Perth, WA, AUSTRALIA. 

WAU, Wageningen Agricultural University, Wageningen, THE NETHERLANDS. 

WCUA, Waite Campus, University of Adelaide, SA, AUSTRALIA 

ZAHC, Zahradnik Personal Collection, Prague?, Czech Republic. 

ZIN, Zoological Institute of Russian Academy of Sciences, St. Petersburg, RUSSIA. 

ZMUA, Zoologisch Museum, University of Amsterdam, Amsterdam, DENMARK. 

ZSI, Zoological Survey of India, Calcutta, India, Kolkata (Calcutta), INDIA. 

Zoogeographic Distribution of Genera and Species of Aleyrodidae 

I have listed the distribution species as being present in one or more of nine 

zoogeographic regions. For this work, I modified the traditional concept of some of these 

regions, primarily to maintain all of the records from a certain country in the same 

zoogeographic region, with the exception of the former USSR, which in most cases I list as being 

in the Western Palearctic (3) and the records from the Maritime Territory (Southern Primor'ye) 

as being in the Eastern Palearctic (5). The Nearctic region traditionally includes northern Mexico 

and north Florida (USA) with the southern portions being in the Neotropical region. In this work, 

and in the Whitefly website, I list all of the species from Mexico as being in the Neotropical 

region and all of the species recorded from Florida as being in the Nearctic. I divided the 

Palearctic region into the Western Palearctic with includes: Europe, North African countries 

bordering the Mediterranean Sea, most of the former USSR (except the Maritime Territory in the 

Far East), Middle East, Iran, Iraq, Saudi Arabia, Uzbekistan, Kyrgyzstan, Tajikistan, 

Afghanistan, Turkmenistan, and the Azores, Madeira and Canary Islands; the Eastern Palearctic 

includes: includes Eastern Asia, China, Japan, Hong Kong, Korea, Taiwan and the Maritime 

Territory (former USSR). The following numbers and abbreviations are used for the 9 

zoogeographic regions: 

Codes for Zoographical Regions 

1- Nearctic (NA) - USA, Canada, Greenland. 

2- Neotropical (NT) - Mexico, Central and South America, Caribbean Islands. 

3- Western Palearctic (WPA) - includes Europe; North Africa (Mediterranean); 

former USSR, Middle East, Iran, Iraq, Saudi Arabia, Uzbekistan, Kyrgyzstan, Tajikistan, 

Afghanistan, Turkmenistan, and Azores, Madeira and Canary Islands. 

4- Afrotropical (AF) - includes Africa south of the Mediterranean countries, Madagascar and 

Seychelles and Cape Verde Islands. 

5- Eastern Palearctic (EPA) - includes Eastern Asia, China, Japan, Hong Kong, Korea, Taiwan, 

5

Maritime Territory (Southern Primor'ye). 

6- Oriental (O)- India, Pakistan, Philippines, Southeast Asia. 

7- Australasian (AU) - Australia, Indonesia, New Guinea. 

8- Pacific Islands (PI) - New Zealand and South Pacific Islands 

9- (HI) - Hawaii. 

1, 2, 3, 4, 5, 6, 7, 8, 9 - Worldwide. 

Abbreviations for USA States 

AK Alaska RI Rhode Island 

AL Alabama SC South Carolina 

AR Arkansas SD South Dakota 

AZ Arizona TN Tennessee 

CA California TX Texas 

CO Colorado UT Utah 

CT Connecticut VA Virginia 

DC District of Colombia VT Vermont 

DE Delaware WA Washington 

FL Florida WI Wisconsin 

GA Georgia WV West Virginia 

HI Hawaii WY Wyoming 

IA Iowa 

ID Idaho 

IL Illinois 

IN Indiana 

KS Kansas 

KY Kentucky 

LA Louisanna 

MA Massachusetts 

MD Maryland 

ME Maine 

MI Michigan 

MN Minnesota 

MO Missouri 

MS Mississippi 

MT Montana 

NC North Carolina 

ND North Dakota 

NE Nebraska 

NH New Hampshire 

NJ New Jersey 

NM New Mexico 

NV Nevada 

NY New York 

OH Ohio 

OK Oklahoma 

OR Oregon 

PA Pennsylvania 

6

Authors and Major Works on Aleyrodidae by Zoogeographic Region 

1. Nearctic - Whiteflies of Southeastern USA (Hodges and Evans 2005) and works by Russell, 

Nakahara, Quaintance and Baker. 

2. Neotropical - Aleurodicinae (Martin 2004) and Aleyrodinae (Martin 2005a) of Belize; 

Aleyrodidae of South America (Baker & Moles 1923) and works by Bondar, Hempel, 

Martin. 

3. Western Palearctic - The whiteflies (Hemiptera: Aleyrodidae) of Europe and the 

Mediterranean Basin (Martin et al. 2000) and works by Bink-Moenen, Danzig (former 

USSR), Gomez-Menor (Spain), Goux (France); Priesner & Hosny (Egypt); Zahradnik. 

4. Afrotropical - Revision of the African whiteflies (Aleyrodidae) mainly based on a collection 

from Tchad. (Bink-Moenen 1983); works by Takahashi (whiteflies of Madagascar and 

Reunion); Mound 1965; Cohic; Corbett. 

5. Eastern Palearctic - works by Ko, Takahashi (Taiwan), Kuwana (Japan); Young (China); Suh 

(Korea); The whiteflies (Homoptera: Aleyrodidoidea) of the Southern Primor'ye (Soviet 

Far East) (Danzig1966); Takahashi (Japan, Taiwan, other countries). 

6. Oriental - India: works by David; Dubey; Sandararaj; Regu, Jesudasan, Meganathan; Malayan 

Aleurodidae (Corbett); Takahashi (Philippines, southeast Asia); Pakistan (Qureshi 1979); 

Sri Lanka (David 1993). 

7. Australasian - Whiteflies of Australia (Martin 1999); Whiteflies of New Guinea (Martin 

1985). 

8. Pacific Islands - The Aleyrodidae (Hemiptera: Homoptera) of New Caledonia (1961a); New 

Zealand Aleyrodidae (Dumbleton 1957). 

9: Hawaii - Aleyrodidae of Hawaii and Fifi with descriptions of new species (Kotinsky 1907); 

Paulson & Kumashiro 1985. 

Other Resources 

Whitefly Pupae of the World (Dooley 2006) - LUCID KEY 

Key to Pest Species - Martin, J. H. 1987. An identification guide to common whitefly pest 

species of the world (Homoptera: Aleyrodidae). Tropical Pest Management. 33: 298-322. 

7

FAMILY ALEYRODIDAE WESTWOOD 1840 

The family Aleyrodidae (Hemiptera) includes 166 genera and 1551 species in 3 extant (living) 

subfamilies (Aleurodicinae, Aleyrodinae and Udamosellinae), and one fossil (non-living) 

subfamily (Bernaeinae). The identification of genera and species is largely based upon 

characteristics present in the fourth stage nymph, known as the puparium. Most whitefly species 

can be classified into their respective subfamilies with the following key: 

Key to Subfamilies of Aleyrodidae (puparium) 

1. Puparium usually with compound or agglomerate pores present, a claw present at the 

apex of each thoracic leg; the lingula very long, extending past the vasiform orifice with 

two or more pairs of setae at its apex..............................................................Aleurodicinae 

1b. Pupa without compound pores present (simple pores rarely present), thoracic legs with 

adhesive or circular disc at the apices of the legs; lingula usually not long and extending 

past the vasiform orifice and with 1 pair of setae ..............................................Aleyrodinae 

Key to the Subfamilies of Aleyrodidae (adults) 

1. Forewing usually with a forked, central vein (Rs present, R1 and media veins strongly 

developed), forewing of Paraleyrodes with a single vein, males with 3 and females with 

four antennal segments; tarsal paronychium thin and spine-like; females with 4 and males 

with 3 ventral abdominal plates, respectively.................................................Aleurodicinae 

1b. Forewing with a single non-forked central vein (Rs present, R1 usually short or absent), 

tarsal paronychium thick and blade-like; females with 2 and males with 4 ventral 

abdominal plates, respectively...........................................................................Aleyrodinae 

Tribal classification 

1. Sampson (1943) provided a key to the subfamilies and genera of Aleyrodidae and proposed 

their classification into 5 tribes. 

2. Russell (1947) proposed the tribe Trialeurodini, a sixth tribal division. 

3. David (1990) proposed a classification of 35 genera of whiteflies from India (all in the 

subfamily Aleyrodinae) into 12 tribes based on characteristics found in the puparium. 

Plant Viruses Transmitted by Whiteflies 

Readers are referred to Jones (2003) for a summary of the viruses transmitted by whiteflies on 

different plant hosts. 

8

Subfamily Aleurodicinae Quaintance and Baker 1913 

The subfamily Aleurodicinae contains 118 species in 18 genera. Although species in the 

subfamily occur in all regions of the world, the number and diversity of species in the 

Neotropical region is far greater than that of the other regions with 102 species in 14 genera 

represented. Martin (2004) reviewed the aleurodicine fauna of Belize, providing a key to the 

genera and certain genera, generic diagnoses, comments and illustrations of species in the 14 

genera occurring in the Neotropical region. Herein, I have provided a key with these and the 

other 4 remaining genera placed in the subfamily. 

Valid Genera 

Aleurodicus Douglas 1892 

Aleuronudus Hempel 1922 

Austroaleurodicus Tapia 1971 

Azuraleurodicus Martin 1999 

Bakerius Bondar 1923 

Ceraleurodicus Hempel 1922 

Dialeurodicus Cockerell 1902 

Eudialeurodicus Quaintance & Baker 1915 

Lecanoideus Quaintance and Baker 1913 

Leonardius Quaintance & Baker 1913 

Metaleurodicus Quaintance & Baker 1913 

Nealeurodicus Hempel 1923 

Nipaleyrodes Takahashi 1951 

Octaleurodicus Hempel 1922 

Pseudsynaleurodicus Gillespie 2006 

Paraleyrodes Quaintance 1909 

Stenaleyrodes Takahashi 1938 

Synaleurodicus Solomon 1935 

Generic Synonymy 

Cited Genus Valid Genus 

Bondaria Sampson & Drews 1941 Dialeurodicus 

Hexaleurodicus Hempel 1923 Aleuronudus 

Parudamoselis Visnya 1941 Ceraleurodicus 

Pentaleurodicus Bondar 1923 Aleuronudus 

Pseudaleurodicus Hempel 1922 Metaleurodicus 

Quaintancius Bondar 1922 Octaleurodicus 

Radialeurodicus Bondar 1922 Ceraleurodicus 

Generic Transfers 

Septaleurodicus SAMPSON 1943; transferred to Aleyrodinae by Martin 2004 

9

Number and Distribution of Genera and Species of Aleurodicine 

Genus NA NT WPA AF EPA O AU PI HI WW 

Aleurodicus 3 26 2 1 2 8 5 3 2 34 

Aleuronudus 0 9 0 0 0 0 0 0 0 9 

Austroaleurodicus 0 2 0 0 0 0 0 0 0 2 

Azuraleurodicus 0 1 0 0 0 0 0 0 0 1 

Bakerius 0 7 0 0 0 0 0 0 0 7 

Ceraleurodicus 0 9 1* 0 0 0 0 0 0 9 

Dialeurodicus 1 12 0 0 0 0 0 0 0 12 

Eudialeurodicus 0 1 0 0 0 0 0 0 0 1 

Lecanoideus 0 2 1 0 0 0 0 0 0 2 

Leonardius 0 2 0 0 0 0 0 0 0 2 

Metaleurodicus 2 11 0 0 0 0 0 0 0 11 

Nealeurodicus 0 8 0 0 0 0 0 0 0 8 

Nipaleyrodes 0 0 0 0 0 1 1 0 0 1 

Octaleurodicus 0 2 0 0 0 0 0 0 0 2 

Paraleyrodes 6 17 3 2 2 2 0 1 3 17 

Pseudosynaleurodicus 0 0 0 0 0 0 2 0 0 2 

Stenaleyrodes 0 0 0 2 0 0 0 1 0 2 

Synaleurodicus 0 0 0 0 0 0 2 0 0 2 

Total species = 118 12 106 6 5 3 9 10 4 6 122 

Total genera = 18 5 14 6 + 1* 3 2 3 4 3 2 18 

NA = Nearctic; NT = Neotropical; WPA = Western Palearctic; AF = Afrotropical; O = Oriental; AU = Australasian; 

PI = Pacific Islands; 9: Hawaii; WW = Worldwide; * greenhouse only. 

10

Key to the Genera of Aleurodicinae of the World 

1. Compound and agglomerate pores absent ........................................................................2 

1b. Compound and/or agglomerate pores present...................................................................5 

2(1). Claws absent; body long and slender, more than 3x longer than wide; vasiform orifice 

elongate triangular; Old World species on palms (Arecaceae).........................................3 

2b. Claws present; body oval, round or pear-shaped, less than 2x longer than wide; 

vasiform orifice variable; Old or New World species on Proteaceae and plants other 

than palms.. .......................................................................................................................4 

3(2) Dorsolateral margin with about 25 pairs of octagonal pores; 1 O (Malaysia) species on 

Nipa (Arecaceae) .......................................................................................... Nipaleyrodes 

3b. Dorsolateral margin without octagonal pores; submarginal papillae present; 2 AF, 1 PI 

species on Cocos (Arecaceae)...................................................................... Stenaleyrodes 

4(2b) Lingula completely included in vasiform orifice with 2 pairs of setae; 12 NT, 1 NA 

species..........................................................................................................Dialeurodicus 

4b. Lingula not completely included in vasiform orifice, extending beyond posterior 

margin, with 1 pair of long setae, and a shorter second pair of setae sometimes present; 

2 AU species on Proteaceae........................................................... Pseudosynaleurodicus 

5(1b) Lingula apparently without setae; dorsum with 8 pairs of evenly spaced large, round 

pores each containing about 3-5 small pores; 2 AU species on Hakea (Proteaceae) 

.......................................................................................................................Synaleurodicus 

5b. Lingula with 2 pairs of setae; dorsum with compound or agglomerate pores, but not as 

above.................................................................................................................................6 

6(5b) Without cephalic compound pores....................................................................................7 

6b. With a pair of cephalic compound pores ..........................................................................8 

7(6) Abdomen with 4 pairs of submedial pores on A1-4 (= abdominal segments 1-4); 

vasiform orifice triangular; 2 NT species ..................................................Octaleurodicus 

7b. Abdomen with 2 pairs of submedial pores, 1 on A2 and other caudal; vasiform orifice 

semicircular; 1 NT species....................................................................... Eudialeurodicus 

8(6b) Abdomen with only 2 pairs of large compound pores, located submedially on A3 and 

A4; vasiform orifice triangular .........................................................................................9 

8b. Abdomen with more than 2 pairs of compound pores, located submedially and/or 

laterally ...........................................................................................................................10 

9(8) Agglomerate pores present on A5-7 and around the compound pores; 2 NT species 

............................................................................................................................. Leonardius 

9b. Agglomerate pores absent; 5 NT species............................................................. Bakerius 

11

12 

10(8b) Abdomen with at least 1 pair of submedial compound pores present ............................11 

10b. Abdomen with all compound pores in lateral position ...................................................14 

11(10) Posterior 4 pairs of compound pores much larger than anterior 2 pairs; 1 pair of 

cephalothoracic setae present; vasiform orifice semicircular; 16 species, mostly 

Neotropical, but distributed almost worldwide.............................................Paraleyrodes 

11b. Pores and cephalic setae not as above.............................................................................12 

12(11b) Cephalothoracic setae absent; at least some of the abdominal compound pores not 

aligned along a curving arch just inside the lateral margin; 5 NT species 

............................................................................................................. Aleuronudus (in part) 

12b. At 1 pair of cephalothoracic setae present; all abdominal compound pores aligned along 

a curving arch just inside the lateral margin ...................................................................13 

13(12b) Body usually asymmetrical, banana-shaped; margin without long setae; 8 NT species 

....................................................................................................................Ceraleurodicus 

13b. Body symmetrical, oval; margin with long setae; 8 NT species................. Nealeurodicus 

14(10b) At least some abdominal compound pores small (not including caudal pores); 

cephalothoracic setae absent; vasiform orifice triangular...............................................15 

14b. All pores large (except caudal pores in some species); cephalothoracic setae present 

except in Azuraleurodicus); vasiform orifice cordate or semicircular............................16 

15(14) At least some of the abdominal compound pores not aligned along a curving arch just 

inside the lateral margin; stellate pores absent; 5 NT species ......... Aleuronudus (in part) 

15b. Al abdominal compound pores aligned along a curving arch just inside the lateral 

margin; some species with stellate pores; 11 NT, 2 NA species. ............. Metaleurodicus 

16(10b) Body elongate with large, double-rimmed 8-shaped pores; 1 NT species, Belize 

..................................................................................................................... Azuraleurodicus 

16b. Body round or oval without large 8-shaped pores..........................................................17 

17(16b) Cephalic and 6 pairs of abdominal pores similar in size, agglomerate pores present on 

A3-A5; lingula very long, cicatrices present on submedian areas of thorax; 1 NT 

species, Chile ........................................................................................Austroaleurodicus 

17b. Posterior 2 pairs of compound pores smaller (or absent) than other pores, agglomerate 

pores and cicatrices absent..............................................................................................18 

18(17b) If small pores present, then 1 pair located on A8 only (under vasiform orifice ; 

submargin with a broad band of simple pores; margin usually deflexed; 2 NT species 

........................................................................................................................... Lecanoideus 

18b. If small pores present, then 2 pairs located on each of A7 and A8; margin usually not 

deflexed; 34 species, worldwide but with 26 NT species............................... Aleurodicus

Aleurodicus Douglas 1892 

Aleurodicus Douglas 1892: 32. Type species. Aleurodicus anonae Morgan 1892:32, 

synonym of Aleurodicus cocois Curtis 1846: 284 according to Mound and Halsey 1978: 229. 

Comment: This is the largest genus in the Aleurodicinae with 35 described species, primarily 

distributed in the Neotropics but occurring in all of the other regions of the world. 

References: Martin (2004) - generic diagnosis and key to Belize species; Regu & David (1992) - 

key to the Indian species. 

Species Distribution 

Species NA NT WPA AF EPA O AU PI HI Fossil 

aculeatus x x 

antidesmae x x 

antillensis x 

araujoi x 

burmiticus x x 

capiangae x 

cinnamomi x 

coccolobae x 

cocois x x 

destructor x x x x x 

dispersus x x x x x x x x x 

dugesii x x x x 

essigi x 

flavus x 

fucatus x 

guppyii x 

holmesii x x x 

indicus x 

inversus x 

jamaicensis x 

juleikae x 

machili x x x 

magnificus x 

maritimus x x 

13

Species NA NT WPA AF EPA O AU PI HI Fossil 

marmoratus x 

neglectus x 

niveus x 

ornatus x 

pauciporus x 

pulvinatus x 

rugioperculatus x 

talamancensis x 

trinidadensis x 

vinculus x 

wallaceus x 

Total = 35 species 3 27 2+1F 1 3 10 5 3 2 2 

Aleurodicus aculeatus Menge 1856 (fossil species) 

Aleurodicus aculeatus Menge 1856: 18. Adults. Prussia, in amber (?lost). 

Distribution: 3: Prussia. 

Host: in amber (fossil). 

Aleurodicus antidesmae Corbett 1926 

Aleurodicus antidesmae Corbett 1926: 267-268. Holotype. Sri Lanka: Pundaluoya, E. E. Green, 

on Antidesma bunius; Martin 1988: 68. 

Distribution: 6: Sri Lanka; 7: Sulawesi. 

Host: Euphorbiaceae: Antidesma bunias; Myrtaceae. 

Aleurodicus antillensis Dozier 1936 

Aleurodicus antillensis Dozier 1936: 144. Syntypes. Puerto Rico: Rio Piedras, 22.xii.1924, H. L. 

Dozier, on coconut palm [Cocos nucifera] and Calophyllum antillanum, three syntypes 

on Erythrina glauca. 

Distribution: 2: Puerto Rico. 

Host: Arecaceae: Cocos nucifera; Fabaceae: Erythrina glauca; Flacourtiaceae: Casearia 

aculeata; Guttiferae: Calophyllum antillanum; Myrtaceae: Psidium guajava; 

unidentified plant 

Natural enemies: Aphelinidae: Encarsia portoricensis; Eulophidae: Aleuroctonus vittatus. 

Aleurodicus araujoi Sampson & Drews 1941 

Aleurodicus araujoi Sampson & Drews 1941: 145-147. Syntypes. Mexico: Colima State, 

Manzanillo, xii.1925, on Aristolochia sp. 

Distribution: 2: Belize; Mexico. 

Host: Aristolochiaceae: Aristolochia sp.; Asteraceae ; Bignoniaceae: Cydista aequinoctialis; 

14

Combretaceae: Terminalia amazonia; Euphorbiaceae: Codiaeum billbergianus?; 

Piperaceae: Piper ?auritum. 

Aleurodicus burmiticus Cockerell 1919 

Aleurodicus burmiticus Cockerell 1919: 241. Female, in Burmese amber, R. C. J. Swinhoe 

collector. 

Distribution: 6: Burma. 

Host: amber (fossil). 

Aleurodicus capiangae Bondar 1923a 

Aleurodicus capiangae Bondar 1923a: 71-73. Syntypes. Brazil: Bahia, Sao Paolo, G. Bondar, on 

capianga [Vismia brasilensis]. 

Distribution: 2: Anguilla, Brazil, Suriname, Trinidad. 

Host: Clusiaceae: Vismia brasiliensis; Fabaceae: Inga sp.; Musaceae: Musa sapientum; 

Verbenaceae: Citharexylum sp. 

US Port Interceptions: Dominica on unknown plant; St. Lucia on Annona muricata; St. 

Maarten on Aristolochia trilobata and US Virgin Islands on an unknown plant. 

Aleurodicus cinnamomi Takahashi 1951c 

Aleurodicus cinnamomi Takahashi 1951c: 1-2. Syntypes. Malaya: Singapore, 21.xii.1942, on 

Cinnamomum sp, Machilus sp. and Lauraceae plant. 

Distribution: 6: Malaya; Malaysia; Singapore. 

Host: Lauraceae: Cinnamomum sp.; Machilus sp. 

Aleurodicus coccolobae Quaintance & Baker 1913 

Aleurodicus coccolobae Quaintance & Baker 1913: 46-47. Syntype. Mexico: Yucatan, Progreso, 

24 vi.1904, S. Henshaw, on seagrape [Coccoloba uvifera]. 

Distribution: 2: Belize; Brazil; Colombia; Costa Rica; Honduras; Mexico; Panama. 

Host: Anacardiaceae: Schinus terebinthifolius; Arecaceae: Cocos nucifera; Fabaceae: Cassia 

siamea; Lauraceae: Persea americana; Myrtaceae: Psidium guajava; Polygonaceae: 

Coccoloba uvifera; Simaroubaceae: Simarouba? sp. 

US Port Interceptions: Costa Rica on an unknown plant; Honduras on Psidium guajava; 

Mexico on Alpinia purpurata and Lauraceae; Peru on an unknown plant. 

Aleurodicus cocois (Curtis) 1846 

Aleurodicus cocois (Curtis) 1846a: 284-285. Syntypes. Barbados: Sir Robert Schonburgk, on 

coconut [Cocos nucifera]. 

Aleurodicus anonae Morgan 1892: 32. Lectotype (designated by Mound & Halsey 1978:229). 

Guyana?: Demerara, 1891, S. J. McIntire, on Annona muricata; synonymy according to 

Mound and Halsey 1978: 229. 

Note: Quaintance and Baker (1913) synonymized Aleurodicus iridescens Cockerell 1898b: 225 

with Aleurodicus cocois; however Martin (2004) synonymized it with Aleurodicus 

pulvinatus. 

Distribution: 1: USA (FL); 2: Anguilla; Barbados; Brazil; Cuba; Dominican Republic; 

Guatemala; Grenada; Guyana; Honduras; Jamaica; Mexico; Panama, Puerto Rico; Peru; 

15

16 

Suriname, Trinidad; Venezuela; Virgin Islands. 

Host: Anacardiaceae: Anacardium occidentale; Annonaceae: Annona muricata; Annona 

reticulata, Annona squamosa; Arecaceae: Cocos nucifera, Cocos sp., Ptychosperma 

macarthurii, Veitchia merrillii, Washingtonia robusta; Chrysobalanaceae: 

Chrysobalanus icaco, Licania tomentosa; Euphorbiaceae: Hevea brasiliensis; 

Fabaceae: Brya ebenus; Lauraceae: Persea americana; Persea sp.; Moraceae: Ficus 

sp.; Musaceae: Musa sp.; Myrtaceae: Psidium guajava; Piperaceae: Piper sp.; 

Polygonaceae: Coccoloba uvifera; Rubiaceae: Richardia pacifica; Solanaceae: 

Lycopersicon esculentum; Sterculiaceae: Theobroma cacao. 

Natural enemies: Fungi - Cladosporium herbarum; Predators - COLEOPTERA/Coccinellidae: 

Cryptognatha nodiceps, Exoplectra sp., Nephaspis coccois, Nephaspis sp.; Syrphidae: 

Baccha sp; Parasitoids - HYMENOPTERA/Aphelinidae: Encarsia guadeloupae, 

Encarsia nr. meritoria, Encarsiella noyesi, Encarsiella sp, Encyrtidae: Metaphycus sp, 

Eulophidae: Entedononecremnus bimaculata, Signiphoridae: Signiphora sp. (probably 

a hyperparasite) 

US Port Interceptions: Aruba on Cocos nucifera; Dominican Republic on Citrus sp.; Mexico 

on Cocos nucifera, Oncidium cavendishianum, Piper sanctum; Peru on herbs. 

Aleurodicus destructor Mackie 1912 

Aleurodicus destructor Mackie 1912: 142-143. Syntypes. Philippines: vicinity of Guijulngan, on 

coconut palm [Cocos nucifera]. 

Aleurodes albofloccosa Froggatt 1918 :436. Syntypes. Australia: N.S.W, Dungay, Tweed River, 

H. Brooks, on undetermined host; synonymy according to Dumbleton 1956b: 160. 

Distribution: 2: Brazil; 6: Philippines, Malaysia; 7: Australia; Brunei; Celebes; Indonesia; Java; 

New Guinea; Sulawesi; 8: Solomon Islands. 

Host: Annonaceae: Annona squamosa, Annona sp; Arecaceae: Cocos nucifera; Fabaceae: Acacia 

sp.; Lauraceae: Cinnamomum sp.; Liliaceae: Cordyline stricta; Moraceae: Ficus 

microcarpa; Piperaceae: Piper nigrum; Piper sp.; Proteaceae: Banksia sp. 

Natural enemies: Predators - COLEOPTERA/ Coccinellidae: Scymnus sp.; DIPTERA/ 

Drosophilidae: Acletoxenus quadristriatus; NEUROPTERA/Chrysopidae: Chrysopa sp. 

Parasitoids - HYMENOPTERA/Aphelinidae: Encarsia sp., Encarsiella sp.; 

Platygastridae: Aleyroctonus pilosus, Aleyroctonus sp. 

Aleurodicus dispersus Russell 1965 

Aleurodicus dispersus Russell 1965: 49-54. Holotype and paratypes. USA: Florida, Key West, 

12.vi.1964, H. V. Weems, on Cocos nucifera; paratypes on Acalypha hispida, 

16.x.1963, H. V. Weems. 

Distribution: 1: USA (FL; HI); 2: Bahamas, Barbados, Bermuda; Brazil; Colombia, Costa Rica; 

Cuba; Dominica, Ecuador; El Salvador, Guadeloupe, Haiti; Honduras; Jamaica, 

Martinique; Mexico; Panama, Peru; Puerto Rico; Venezuela; Virgin Islands; 3: Canary 

Islands, Iran; Italy; 4: Benin, Cameroun; Congo; Ghana; Madeira; Mauritius; Nigeria; 

Sao Tome; 5: Philippines; 6: Bangladesh, India; Singapore, Sri Lanka; Thailand, 

Vietnam; 7: Australia, Java; 8: American Samoa; Fiji, Guam; Marshall Islands, Palau, 

Saipan; 9: Hawaii. 

Host: Acanthaceae: Barleria parviflora; Blechum pyramidatum, Sanchezia nobilis; Aceraceae:

17 

Acer saccharinum; Amaryllidaceae: Crinum americanum; Anacardiaceae: Mangifera 

indica; Schinus terebinthifolius; Annonaceae: Annona squamosa, Polyalthia longifolia; 

Apocynaceae: Beaumontia grandiflora, Plumeria sp.; Araceae: Anthurium sp., 

Monstera deliciosa, Spathiphyllum cuspidatum, Spathiphyllum sp.; Araliaceae: 

Schefflera arboricola, Schefflera elegantissima; Arecaceae: Chrysalidocarpus lutescens 

(See: Dypsis lutescens); Coccothrinax argentata, Cocos nucifera; Dypsis lutescens, 

Hyophorbe verschaffeltii, Phoenix dactylifera, Pritchardia pacifica, Scheelea 

leandroana; Scindaspis aureus; Trachycarpus fortunei; Begoniaceae: Begonia sp., 

Begnoniaceae: Tecoma stans; Bixaceae: Bixa orellana; Bombacaceae: Quararibea 

cordata; Burseraceae: Bursera simaruba.; Caricaceae: Carica papaya; 

Chrysobalanaceae: Chrysobalanus icaco; Combretaceae: Bucida buceras, Bucida sp.; 

Combretaceae: Conocarpus erectus; Terminalia arjuna; Terminalia catappa; 

Terminalia muelleri; Terminalia sp.; Convolvulaceae: Ipomoea batatas; Cucurbitaceae: 

Coccinia sp.; Dioscoreaceae: Dioscorea sp.; Dryopteridaceae: Bolbitis cladorrhizans; 

Euphorbiaceae: Acalypha hispida; Acalypha sp.; Antidesma bunias; Antidesma 

dallachyanum; Bischofia javanica, Euphorbia heterophylla; Euphorbia pulcherrima; 

Hura crepitans; Jatropha sp.; Macaranga peltata; Mallotus sp.; Manihot glaziovii; 

Fabaceae: Acacia auriculiformis; Arachis hypogaea; Bauhinia purpurea; Bauhinia sp.; 

Bauhinia variegata; Cassia bahamensis; Cassia fistula; Cassia siamea; Cassia sp.; 

Cassia spectabilis; Dalbergia latifolia; Dizygotheca elegantissima; Flemingia 

macrophylla; Inga edulus; Inga laurina; Phaseolus vulgaris; Guttiferae: Calophyllum 

antillanum; Calophyllum inophyllum Guttiferae: Clusia rosea; Labiatae: Coleus sp.; 

Lauraceae: Actinodaphne angustifolia; Cinnamomum malabathrum; Persea americana; 

Lecythidaceae: Barringtonia speciosa; Lobeliaceae: Lobelia excelsa; Magnoliaceae: 

Michelia champaca; Malvaceae: Gossypium hirsutum, Gossypium sp.; Hibiscus 

tiliaceus; Sida acuta, Thespesia populnea; Meliaceae: Melia azedarach; Moraceae: 

Arctocarpus heterophyllus; Artocarpus altilis, Artocarpus heterophyllus; Ficus 

asperrima, Ficus aurea; Ficus benghalensis; Ficus religiosa; Ficus sp.; Morus alba; 

Morus sp.; Musaceae: Heliconia lindsavae; Heliconia sp.; Musa nana; Musa 

sapientum; Musa sp.; Musa sumatrana; Musa x paradisiaca; Ravenala 

madagascariensis, Strelitzia nicolai; Myrtaceae: Eucalyptus sp.; Eucalyptus 

tereticornis; Eugenia foetida, Eugenia uniflora, Melaleuca leucadendra, Psidium 

cattleianum,: Psidium guajava; Psidium littorale; Syzygium cuminii; Oleaceae: 

Osmanthus fragrans; Orchidaceae: Cyrtopodium andersonii, Laelia sp.; Peristeria sp.; 

Sobralia sp.; Plumbaginaceae: Laguncularia racemosa; Poaceae: Saccharum 

officinarum; Polygonaceae: Coccoloba diversifolia, Coccoloba floridana; Coccoloba 

uvifera; Rhizophoraceae: Rhizophora mangle; Rosaceae: Prunus sp.; Rubiaceae: Coffea 

sp.; Ixora parviflora; Rutaceae: Citrus aurantifolia, Citrus aurantium, Citrus limon; 

Citrus sp.; Citrus x paradisi; Santalaceae: Santalum album; Sapindaceae: Melicoccus 

bijugatus; Sapotaceae: Achras sp.; Chrysophyllum cainito; Chrysophyllum oliviforme; 

Manilkara zapota; Solanaceae: Brunfelsia pauciflora; Capsicum sp.; Cestrum diurnum; 

Physalis sp.; Solandra sp.; Solanum blodgettii; Solanum melongena; Solanum sp.; 

Ulmaceae: Celtis pallida; Verbenaceae: Tectona grandis; Vitaceae: Ampelocissus 

latifolia; Leea coccinea. 

Natural enemies:

18 

Parasitoids - HYMENOPTERA/Aphelinidae: Encarsia dispersa, Encarsia guadeloupae, 

Encarsia hispida, Encarsia lahorensis, Encarsia luteola group, Encarsia nigricephala, 

Encarsia sophia, Encarsia sp., Encarsia tabacivora, Encarsiella sp.; Eulophidae: 

Aleuroctonus marki, Aleuroctonus vittatus; Signiphoridae: Signiphora sp. [probably a 

hyperparasite]. 

Predators - COLEOPTERA/Coccinellidae: Axinoscymnus beneficus, Axinoscymnus 

puttarudriahi, Coelophora inequalis, Coelophora pupillata, Cryptolaemus 

montrouzieri, Curinus coeruleus Mulsant, Delphastus pusillus, Harmonia 

octomaculata, Menochilus sexmaculatus, Nephaspis bicolor, Nephaspis oculatus, 

Nephaspis roephei, Nephaspis sp., Olla v-nigrum, Scymnus sp., Serangium 

maculigerum, Nitidulidae: Cybocephalus sp.; DIPTERA/ Drosophilidae: Acletoxenus 

quadristriatus; Syrphidae: Allograpta obliqua, Paragus serratus; 

HYMENOPTERA/Formicidae: Iridomyrmex anceps; NEUROPTERA/Chrysopidae: 

Chrysoperla comanche; Hemerobiidae: Sympherobius barberi. 

US Port Interceptions: American Samoa on Piper sp.; Antigua and Barbuda on Annona 

muricata; Bahamas on Cocos nucifera; Bangladesh on unknown plant; Cambodia on 

Psidium guajava, Psidium sp.; Cameroon on Theobroma cacao; Congo on Ipomoea 

sp.; Costa Rica on Aracaena sp, Curculigo sp.; Dominica on Rosa sp.; Dominican 

Republic on Citrus sp, Psidium guajava; Fiji on Moringa sp., Psidium guajava; 

France on Laurus nobilis; Ghana on unknown plant; Guam on Piper betle, Piper sp.; 

Haiti on Psidium guajava; Hawaii on Abelmoschus manihot, Abelmoschus moschatus, 

Alocasia sp., Alpinia purpurata, Alyxia sp., Apocynaceae, Arundina bambusifolia, 

Arundina sp., Bauhinia purpurea, Bauhinia sp., Bixa orellana, Broussonetia sp., 

Capsicum annuum, Capsicum sp., Carica papaya, Carica sp., Citrus sp., Cocos 

nucifera, Colocasia esculenta, Colocasia sp., Corchorus olitorius, Cordyline fruticosa, 

Cordyline sp., Dimocarpus longan, Ficus sp., Gardenia jasminoides, Gardenia sp., 

Gardenia taitensis, Gossypium hirsutum, Hakea sp., Heckeria sp., Heckeria umbellata, 

Hibiscus manihot, Hibiscus rosa-sinensis, Hibiscus sp., Houttuynia cordata, Jasminum 

multiflorum, Jasminum sp., Lagerstroemia speciosa, Mangifera indica, Mentha 

javanica, Mentha sp., Momordica charantia, Monstera sp., Morinda citrifolia, Morinda 

sp., Moringa citrifolia, Moringa oleifera, Moringa sp., Murraya paniculata, Musa sp., 

Neurolaena lobata, Ocimum sp., Oncidium sp., Pandanus sp., Peperomia sp., Physalis 

peruviana, Piper betle, Piper sp., Plumeria acuminata, Plumeria obtusa, Plumeria 

rubra, Plumeria sp., Polygonatum odoratum, Psidium guajava, Psidium sp., Rosa 

chinensis, Rosa sp., Scaevola sp., Solanum sp., Syzygium malaccense, Vigna sp., Hong 

Kong Psidium guajava; India on Murraya koenigii, Psidium guajava, Psidium sp.; 

Indonesia on unknown plant; Laos on Lansium sp., Psidium guajava, Psidium sp.; 

Liberia on unknown plant; Malaysia on unknown plant; Marshall Islands on 

Hibiscus sp.; Martinique on Alpinia sp.; Mexico on Annona cherimola, Citrus sp., 

Persea americana; Nigeria on Ficus sp., herbs, Manihot esculenta, Musa sp.; 

Northern Mariana Islands on Xanthosoma sagittifolium; Philippines on Annona 

squamosa, Capsicum sp., Crotalaria sp., Eucalyptus sp., Euphorbia pulcherrima, 

Hibiscus rosa-sinensis, Lagerstroemia speciosa, Manihot esculenta, Mentha sp., 

Moringa oleifera, Moringa pterygosperma, Moringa sp., Musa paradisiaca, Musa sp., 

Myristica sp., Orchidaceae; Persea americana, Persea sp., Piper betle, Piper sp.,

19 

Polyathea sp., Psidium guajava, Psidium sp., Xanthosoma sagittifolium; Puerto Rico 

on Capsicum annuum, Citrus sp., Genipa americana, Hibiscus rosa-sinensis, 

Mangifera indica, Morinda citrifolia, Musa acuminata, Musa sp., Persea americana, 

Plumeria sp., Psidium guajava, Ruella sp., Solanum nigrum; Singapore on herbs, 

Pinanga sp.; Spain on Areca catechu; Thailand on Psidium guajava; Tortola on 

Psidium guajava; US Virgin Islands on Pimenta racemosa; Viet Nam on Eucalyptus 

sp., Moringa oleifera, Morus sp., Musa sp., Psidium guajava. 

Aleurodicus dugesii Cockerell 1896 

Aleurodicus dugesii Cockerell 1896a: 302. Syntypes. Mexico: Guanajuato, A. Duges, on 

Hibiscus rosa-sinensis. 

Distribution: 1: USA (AZ; CA; FL; HI; LA; TX); 2: Belize, Costa Rica, El Salvador, Guatemala, 

Mexico, Nicaragua, Venezuela; 6: Pakistan; 9: Hawaii. 

Host: Annonaceae: Annona sp.; Araceae: Spathiphyllum floribundum; Araliaceae: Aralia sp.; 

Begoniaceae: Begonia sp.; Burseraceae: Bursera simaruba; Cannaceae: Canna sp.; 

Canna x generalis; Chenopodiaceae: Chenopodium ambrosioides; Chrysobalanaceae: 

Chrysobalanus icaco; Cucurbitaceae: Cucurbita sp.; Euphorbiaceae: Acalypha 

wilkesiana; Chamaesyce hypericifolia; Geraniaceae: Geranium sp.; Hemerocallidaceae/ 

Liliaceae: Hemerocallis sp.; Labiatae: Coleus scutellarioides; Labiatae: Salvia elegans; 

Lauraceae: Persea sp.; Malvaceae: Hibiscus rosa-sinensis; Hibiscus sp.; Hibiscus 

syriacus; Moraceae: Morus sp.; Musaceae: Heliconia sp.; Polygonaceae: Coccoloba 

belizensis; Rutaceae: Citrus sinensis; Citrus sp.; Sapotaceae: Bumelia laetivirens; 

Solanaceae: Capsicum annuum; Lycianthes hypoleuca; Ulmaceae: Trema micranthum; 

Verbenaceae: Clerodendrum sp.; Viscaceae: Phoradendron leucarpum. 

Natural enemies: Predators - COLEOPTERA/Coccinellidae: Delphastus catalinae; 

Parasitoids - HYMENOPTERA/Aphelinidae: Encarsia guadeloupae, Encarsia hispida, 

Eulophidae: Entedononecremnus krautneri, Pteromalidae: Idioporus affinis. 

US Port Interceptions: Hawaii on Citrus sp., Gossypium sp., Hibiscus sp., Ocimum sp., Plumeria 

sp.; Mexico on Begonia sp., Citrus sinensis, Cocos nucifera, Hedera sp., Heliconia sp., 

Musa sp., Persea americana, Persea sp., Piper sp., Psidium sp.; Pakistan on Citrus sp. 

Aleurodicus essigi Sampson & Drews 1941 

Aleurodicus essigi Sampson & Drews 1941: 147-148. Syntypes. Mexico: Guerrero State, 

Acupulco, on cliffs, iii.1926. 

Distribution: 2: Mexico. 

Host: unidentified shrub. 

Aleurodicus flavus Hempel 1922 

Aleurodicus flavus Hempel 1922: 4-5. Syntypes. Brazil: Bahia, Sao Paulo, on coconut [Cocos 

nucifera]. 

Distribution: 2: Brazil. 

Host: Arecaceae: Cocos nucifera. 

Aleurodicus fucatus Bondar 1923 

Aleurodicus fucatus Bondar 1923a: 74. Syntypes. Brazil: Bahia State, Belmonte, G. Bondar, on

20 

cacauerio [Theobroma cacao], ingazeira [Inga sp.] and embauba [Cecropia palmata]. 


Host: Fabaceae: Inga sp.; Moraceae: Cecropia palmata; Sterculiaceae: Theobroma cacao. 

Aleurodicus guppyii Quaintance & Baker 1913 

Aleurodicus guppyii Quaintance & Baker 1913: 59-60. Syntypes. Trinidad: Port of Spain, 

25.v.1911, F. W. Urich, on Rheedia latiflora. 

Distribution: 2: Trinidad. 

Host: Guttiferae: Rheedia latiflora. 

Aleurodicus holmesii (Maskell) 1895 

Aleurodes holmesii Maskell 1895: 435. Syntypes. Fiji: R. L. Holmes collector, on Psidium sp. 

Aleurodicus malayensis Takahashi 1951c: 2-3. Syntypes. Fiji: R. L. Holmes collector, on 

Psidium sp.; Martin 1985: 312. 

Distribution: 6: Malaysia, Sri Lanka; Thailand; 7: Java; 8: Fiji. 

Host: Myrtaceae: Psidium guajava, Psidium sp.; Sapindaceae: Dimocarpus longon; unidentified 

tree. 

Aleurodicus indicus Regu & David 1992 

Aleurodicus indicus Regu & David 1992: 100. Holotype pupa case, India, Kanyakumari District, 

Munchirai, 15.xii.1989, K. Regu, on Polyalthia suberosa; Regu Collection. 

Distribution: 6: India. 

Host: Annonaceae: Polyalthia suberosa. 

Aleurodicus inversus Martin 2004 

Aleurodicus inversus Martin 2004: 22. Holotype puparium, Belize, CFR, Las Cuevas, 

25.iii.2003, J. Martin, ex. Stizophyllum riparium. 

Distribution: 2: Belize. 

Host: Bignoniaceae: Arrabidaea patellifera; Stizophyllum riparium; Sapindaceae: Paullinia 

pinnata. 

Aleurodicus jamaicensis Cockerell 1902 

Aleurodicus jamaicensis Cockerell 1902a: 280. Syntypes. Jamaica: Kingston, 1893, T. D. A. 

Cockerell, on undetermined host. 

Distribution: 2: Ecuador; Jamaica. 

Host: Arecaceae: Cocos nucifera; Asteraceae: Mikania cordifolia; unidentified plant. 

Aleurodicus juleikae Bondar 1923 

Aleurodicus juleikae Bondar 1923a: 78-80. Syntypes. Brazil: Baia, Sao Paulo, G. Bondar, on 

Phrygilanthus sp.; Costa Lima 1928: 107. 


Host: Loranthaceae: Phrygilanthus sp.; Tiliaceae: Triumfetta semitriloba; Viscaceae: 

Phoradendron platycaulon; unidentified plant. 

Natural enemies: Aphelinidae: Encarsia gallardoi sp. aff.

21 

Aleurodicus machili Takahashi 1931 

Aleurodicus machili Takahashi 1931a: 208-209. Syntypes. Taiwan: Taihoku, x.1930, R. 

Takahashi, on Machilus sp. 

Aleurodicus formosanus Takahashi 1932: 7-8. Syntypes. Taiwan: Taihezan, 21.v1931, R. 

Takahashi, on Machilus sp.; synonymy according to Mound and Halsey 1978: 234. 

Aleurodicus philomenae David 1987: 1247. Holotype puparium, India, Mahabeleshwar, Pune, 

Maharashtra, 23.iii.1987, B.V. David, on Persea macrantha, FIPPT; synonomy 

according to David & Selvakumaran 1990:139. 

Distribution: 3: Iran; 5: Hong Kong, Taiwan; 6: India. 

Host: Labitae: Ajuga gorganica; Lauraceae: Actinodaphne pedicellata; Cinnamomum sp.; Litsea 

travancorica; Machilus sp.; Persea macrantha, Persea zuihoensis, Persea sp.; 

Moraceae: Morus nigra. 

Natural enemies: Aphelinidae: Encarsiella taiwanensis. 

Aleurodicus magnificus Costa Lima 1928 

Aleurodicus magnificus Costa Lima 1928: 129-131. Syntypes. Brazil: S. Lourenco, Sul de 

Minas, 1921, on orange [Citrus sp.]; Martin 2004: 24. 

Distribution: 2: Belize; Brazil; Panama. 

Host: Rutaceae: Citrus sp.; unidentified plant. 

Aleurodicus maritimus Hempel 1923 

Aleurodicus linguosus Bondar 1923a: 76-78. Syntypes. Brazil: Bahia, G. Bondar, on Moquilea 

tomentosa. 

Aleurodicus maritimus Hempel 1923: 1160-1161. Syntypes. Brazil: Sao Paulo State, Sao 

Sebastiao, A. A. Barbiellini, on Psidium sp.; synonymized by Costa Lima 1928: 133. 

Distribution: 2: Belize; Brazil; Guyana; Honduras; Mexico; Nicaragua; Panama; Surinam; 

Trinidad; 6: Malaya; Malaysia. 

Host: Arecaceae: Cocos nucifera; Bignoniaceae: Stizophyllum riparium; Cecropiaceae: Cecropia 

sp.; Chrysobalanaceae: Licania tomentosa; Clusiaceae: Vismia brasiliensis; Clusiaceae: 

Vismia camparaguey; Combretaceae: Combretum laxum; Euphorbiaceae: Croton 

billbergianus; Fabaceae: Cajanus cajan; Melastomataceae: Clidemia octona; 

Melastomataceae: Miconia impetiolaris; Myrtaceae: Psidium guajava; Psidium sp.; 

Rosaceae: Licania tomentosa; Styraceae: Styrax argenteus; Verbenaceae: Petrea 

volubilis. 

Natural enemies: Parasitoids - Aphelinidae: Dirphys mendesi, Encarsia dispersa, Encarsia 

guadeloupae, Encarsia near meritoria, Encarsiella noyes, Encarsiella sp., Encyrtidae: 

Metaphycus sp. 

US Port Interceptions: Mexico on Persea americana, Psidium guajava, Salvia officinalis; 

Trinidad and Tobago on Petrea sp. 

Aleurodicus marmoratus Hempel 1923 

Aleurodicus marmoratus Hempel 1923: 1161-1163. Syntypes. Brazil: Sao Paulo State, Caconde, 

Itatiba and Ypiranga, A. Hempel, on Baccharis genistelloides. 


Host: Asteraceae: Baccharis genistelloides.

Aleurodicus neglectus Quaintance & Baker 1913 

Aleurodicus neglectus Quaintance & Baker 1913 Quaintance & Baker 1913: 63-65. Syntypes. 

Brazil: Para, 1882, A. Koebele, on guava [Psidium guajava]. 

Aleurodicus flumineus Hempel 1918: 211-214. Syntypes. Brazil: Rio de Janeiro State, Pinheiros, 

on Moquilea tomentosa; synonymy according to 1923a: 61. 

Distribution: 2: Barbados; Brazil; Guyana; Trinidad. 

Host: Annonaceae: Annona reticulata, Annona squamosa; Arecaceae: Cocos nucifera; 

Chrysobalanaceae: Licania tomentosa; Moraceae: Cecropia concolor; Cecropia sp.; 

Ficus benghalensis; Myrtaceae: Psidium guajava; Sterculiaceae: Theobroma cacao. 

Natural enemies: Predators - COLEOPTERA/Coccinellidae: Delphastus nebulosus, Nephaspis 

sp., Scymnus sp.; NEUROPTERA/Chrysopidae: Chrysopodes (Neosuarius) collaris 

Parasitoids - HYMENOPTERA/Aphelinidae: Dirphys larensis, Encarsia dispersa, 

Encarsia guadeloupae, Encarsiella noyesi, Encarsiella sp.; Eulophidae: Aleuroctonus 

vittatus, Entedononecremnus sp. 

Aleurodicus niveus Martin 2004 

Aleurodicus niveus Martin 2004: 25. Holotype puparium, Belize, CFR, Las Cuevas forest, on 

Encyclia cochleata, BMNH. 


Host: Orchidaceae: Encyclia cochleata. 

Aleurodicus ornatus Cockerell 1893 

Aleurodicus ornatus Cockerell 1893: 105-106. Syntypes. Jamaica: Kingston, 1893, T. D. A. 

Cockerell, on Capsicum sp., USNM. 

Distribution: 2: Jamaica 

Host: Arecaceae: Cocos nucifera; Solanaceae: Capsicum sp. 

US Port Interceptions: Haiti on Mangifera indica, Stachytarpheta sp. 

Aleurodicus pauciporus Martin 2004 

Aleurodicus pauciporus Martin 2004: 26. Holotype puparium, Belize: CFR, Monkey Tail Track, 

4.iv.2003, J. Martin, on Casearia tremula (Flacourtiaceae), BMNH. 


Host: Aquifoliaceae: Ilex belizensis; Flacourtiaceae: Casearia tremula; Lauraceae: Nectandra sp. 

Aleurodicus pulvinatus (Maskell) 1895 

Aleurodes pulvinatus Maskell 1895: 439-441. Syntypes. Trinidad: F. W. Urich, on Jatropha sp. 

Aleurodicus pulvinatus (Maskell); Cockerell 1898b: 275. 

Aleurodicus iridescens Cockerell 1898b: 225. Syntypes. Mexico: Tabasco, ocean between El 

Faro and San Pedro, 12.vi.1897, C.H.T. Townsend, on "jicaco" [Chrysobalanus icaco]; 

synonymized with A. cocois by Quaintance & Baker 1913: 47; however, Martin 2004: 

413, says that it is synonymous with A. pulvinatus. 

Distribution: 2: Barbados; Belize; Brazil; Costa Rica; Ecuador; Guyana; Mexico; Nevis; 

Panama; St. Kitts; Trinidad; Venezuela. 

Host: Alismataceae: Echinodorus sp.; Araceae: Montrichardia aculeata; Montrichardia 

22

23 

arborescens; Arecaceae: Cocos nucifera; Chrysobalanaceae: Chrysobalanus icaco, 

Licania tomentosa; Clusiaceae: Vismia brasiliensis; Vismia sp.; Combretaceae: 

Terminalia catappa; Euphorbiaceae: Hura crepitans; Jatropha sp.; Fabaceae: Cajanus 

cajan; Lacistataceae: Lacistema sp.; Lauraceae: Nectandra? sp.; Persea americana; 

Moraceae: Ficus sp.; Musaceae: Musa sp.; Myrtaceae: Psidium guajava; Piperaceae: 

Piper nigrum; Polygonaceae: Coccoloba belizensis; Coccoloba uvifera; Rosaceae: 

Licania sp.; Rubiaceae: Coffea canephora, Guettarda combsii, Guettarda sp.; Isertia 

hypoleuca; Sterculiaceae: Theobroma sp.; Verbenaceae: Petrea sp. 

US Port Interceptions (as A. iridescens): Antigua and Barbuda on Annona sp., Spondias dulcis; 

Aruba on Cocos nucifera; Barbados on Annona squamosa; Costa Rica on Schefflera 

arboricola; Dominican Republic on Bauhinia sp., Citrus sp.; Honduras on Psidium 

guajava; Kiribati on Annona sp.; Mexico on Persea americana; Peru on Echinodorus 

radicans, Echinodorus sp.; Puerto Rico on Coccoloba uvifera; St. Kitts and Nevi on 

Annona muricata, Pimenta racemosa; US Virgin Islands on Pimenta racemosa. 

Aleurodicus rugioperculatus Martin 2004 

Aleurodicus rugioperculatus Martin 2004: 29. Holotype puparium, Belize: Belize City, 

11.iii.1996, J. Martin, on Cocos nucifera (Arecaceae), BMNH. 


Host: Arecaceae: Cocos nucifera; Myrtaceae: Psidium guajava. 

US Port Interceptions: Mexico on Cocos nucifera, Laurus nobilis, Persea americana, Psidium 

guajava. 

Aleurodicus talamancensis Martin 2005 

Aleurodicus talamancensis Martin 2005: 843. Holotype puparium, Costa Rica, on Musa 

cultivated variety, Limon Province, Talamanca Canton, 2004, D. Cubillo, BMNH. 

Distribution: 2: Costa Rica. 

Host: Musaceae: Musa sp.; Orchidaceae: Maxillaria sp. 

Aleurodicus trinidadensis Quaintance & Baker 1913 

Aleurodicus trinidadensis Quaintance & Baker 1913: 69. Syntypes. Trinidad: 27.iii.1912, on 

coconut [Cocos nucifera]. 


Host: Arecaceae: Cocos nucifera; Asteraceae: Eupatorium odoratum; Musaceae: Musa 

sapientum. 

Aleurodicus vinculus Martin 2004 

Aleurodicus vinculus Martin 2004: 30. Holotype puparium, Belize: CFR, 20.xi.1994, J. Martin, 

on woody vines, BMNH. 


Host: woody vine. 

Aleurodicus wallaceus Martin 1988 

Aleurodicus wallaceus Martin 1988: 59. Holotype pupal case, Indonesia, Sulawesi Utara, 

23.iii.1985, J. Martin, on Persea americana, BMNH.

Distribution: 7: Indonesia, Sulawesi. 

Host: Lauraceae: Persea americana. 

Aleurodicus sp. 

Natural enemies: Parasitoids: Aphelinidae: Dirphys larensis, Encarsia dispersa, Encarsia 

lanceolata, Encarsia tabacivora, Encarsia variegata, Encarsiella aleurodic, 

Encarsiella ciliata, Encarsiella narroi, Encarsiella sp.; Eulophidae: Aleuroctonus 

vittatus, Entedononecremnus funiculatus, Signiphoridae: Signiphora sp. 

24

Aleuronudus Hempel 1922 

Aleuronudus Hempel 1922: 5. Type-species: Aleuronudus induratus Hempel 1922, by 

monotypy. 

Hexaleurodicus Hempel 1923: 84. Type-species: Hexaleurodicus jaciae Bondar 1923, by 

monotypy; synonymy according to Martin 2004: 32. 

Hexaleurodicus (Drewsia) Sampson 1943: 192-193. Type-species: Hexaleurodicus ferrisi, by 

monotypy; Mound and Halsey 1978: 242; synonymy according to Martin 2004: 32. 

Pentaleurodicus Bondar 1923a: 85-87. Type-species: Aleuronudus induratus Hempel, by 

original designation; synonymy according to Costa Lima 1928: 127. 

Comment: Costa Lima (1936) synonymized Pseudaleurodicus Hempel 1922 with Aleuronudus; 

however, Martin (2004) disagreed, regarding it as a synonym of Metaleurodicus. 


Species NA NT WPA AF EPA O AU PI HI 

acapulcensis x 

bondari x 

induratus x 

jaciae x 

jequiensis x 

manni x 

melzeri x 

Total = 7 species 0 7 0 0 0 0 0 0 0 

Comment: This genus is only known to occur in the Neotropical region (Brazil, Mexico, 

Trinidad). 

Reference: Martin 2004 - Generic description and taxonomy and illustrations of 2 species. 

Aleuronudus acapulcensis (Sampson & Drews) 1941 

Pseudaleurodicus acapulcensis Sampson & Drews 1941: 155-156. Syntypes. Mexico: Guerrero 

State, Acapulco, iii.1926, on woody vine, USNM 

Aleuronudus acapulcensis (Sampson & Drews); Mound and Halsey 1978: 236 

Distribution: 2: Brazil; Mexico. 

Host: Rutaceae: Citrus maxima, Citrus sp. 

Aleuronudus bondari (Costa Lima) 1928 

Aleurodicus bondari Costa Lima 1928: 131-132. Syntypes. Brazil, MZU. 

Aleuronudus bondari (Costa Lima); Costa Lima 1928: 131-132. 


Host: Rutaceae: Citrus sp. 

25

Aleuronudus induratus Hempel 1922 

Aleuronudus induratus Hempel 1922: 5-6. Syntypes. Brazil: Bahia, on Cocos nucifera, MZU; 

Pentaleurodicus induratus (Hempel); Bondar 1923a: 91. 

Aleuronudus induratus Hempel; Costa Lima 1928: 137. 



Aleuronudus jaciae (Bondar) 1923 

Hexaleurodicus jaciae Bondar 1923a: 84-85. Syntypes. Brazil: Bahia, G. Bondar, on Chomelia 

sp., Miconia sp., and Citrus sp., MZU. 

Aleuronudus jaciae (Bondar 1923) 


Host: Melastomataceae: Miconia sp.; Rubiaceae: Chomelia sp.; Rutaceae: Citrus sp. 

Aleuronudus jequiensis (Bondar) 1928 

Metaleurodicus jequiensis Bondar 1928b: 21-23. Syntypes. Brazil: Bahia, Jequie, on unidentified 

Compositae, MZU 


Host: Asteraceae. unidentified plant. 

Aleuronudus manni (Baker) 1923 

Aleurodicus (Metaleurodicus) manni Baker 1923: 253-254. Syntypes. Honduras: Ceiba, iii.1920, 

W. M. Mann, on orange [Citrus sp.], USNM 

Metaleurodicus manni (Baker) 1923; Mound and Halsey 1978: 245 

Hexaleurodicus ferrisi Sampson & Drews 1941: 154-155. Syntypes. Mexico: Oaxaca, Chivela, 

iv.1926, on undetermined Leguminous shrub, UCD; synonymy according to Martin 

2004: 33. 

Distribution: 2: Colombia; Mexico; Trinidad. 

Host: Myrtaceae: Psidium guajava; Passifloraceae: Passiflora edulis, Passiflora sp.; Rutaceae: 

Citrus reticulata, Citrus sp.; unidentified shrub. 

Aleuronudus melzeri (Bondar) 1928 

Metaleurodicus melzeri Bondar 1928b: 19-21. Syntypes. Brazil: Jacaranda, Cannavieiras, on 

climbing Apocynaceae plant, MZU; synonym with Nealeurodicus paulistus according to 

Martin 2004: 54. 

Aleuronudus melzeri (Bondar); Martin & Mound 2007: 51. 


Host: Apocynaceae. 

26

Austroaleurodicus Tapia 1970 

Austroaleurodicus Tapia 1970. Type species - Aleurodicus (Metaleurodicus) pigeanus Baker & 

Moles 1923. 



lomatiae x 

pigeanus x 

Total = 2 species 0 2 0 0 0 0 0 0 0 

Comment: This genus is known from two described species, one from Chile and the other from 

Argentina. 

Reference: Tapia 1970. 

Austroaleurodicus pigeanus (Baker & Moles) 1923 

Aleurodicus (Metaleurodicus) pigeanus Baker & Moles 1923: 622-623. Syntypes. Chile: C. E. 

Porter, on Quilaja saponaria, USNM. 

Metaleurodicus pigeanus (Baker & Moles); Mound and Halsey 1978: 246. 

Austroaleurodicus pigeanus (Baker & Moles); Tapia 1970; Martin 2004: 47. 

Distribution: 2: Chile; Puerto Rico. 

Host: Rosaceae: Quillaja saponaria. 

Natural enemies: Platygastridae: Amitus pigeanus. 

Austroaleurodicus lomatiae Tapia 1970 

Austroaleurodicus lomatiae Tapia 1970: 220. Holotype puparium, Argentina, San Carlos de 

Bariloche, Rio Negro, D. Havrylenko, ex. Lomatia hirsuta. 

Distribution: 2: Argentina. 

Host: Proteaceae: Lomatia hirsuta. 

27

Azuraleurodicus Martin 1999 

Azuraleurodicus Martin 1999: 1547, in Martin & Polaszek 1999. Type species. Azuraleurodicus 

pentarthus Martin 1999, by monotypy. 



pentarthus x 

Total = 1 species 0 1 0 0 0 0 0 0 0 

Comment: The single species of this genus is only known to occur in Belize, Colombia and El 

Salvador. 

Reference: Martin 2004 - Diagnosis and illustration. 

Azuraleurodicus pentarthus Martin 1999 

Azuraleurodicus pentarthus Martin 1999, in Martin & Polaszek 1999: 1547. Holotype pupal 

case. Belize: Cayo District, Chiquibul Forest Reserve, Grano de Oro, 22.xi.1994, J. H. 

Martin, on Guettarda combresii, BMNH. 

Distribution: 2: Belize; Colombia, El Salvador. 

Host: Clusiaceae: Vismia camparguey; Euphorbiaceae: Codiaeum sp.; Fabaceae: Erythrina sp., 

Inga sp, Phaseolus sp.; Rubiaceae: Guettarda combresii. 

Natural enemies: Aphelinidae: Dirphys aphania, Encarsiella pithecura. 

28

Bakerius Bondar 1923 

Bakerius Bondar 1923a: 38-40. Type-species: Bakerius phrygilanthi Bondar 1923, by original 

designation. 



amazonicus x 

attenuatus x 

calmoni x 

conspurcatus x 

glandulosus x 

maculatus x 

phrygilanthi x 

sanguineus x 

sublatus x 

Total = 9 species 0 9 0 0 0 0 0 0 0 

Comment: All of the species that have been described in this genus are from Brazil. However the 

the genus is also known to occur in Colombia and Peru (unpublished). 

Diagnosis - Dorsum with only 2 pairs of abdominal compound pores on submedian area of A3 

and A4, one pair of compound pores on cephalus; 4 pairs of cephalothoracic setae 

present. Differs from Leonardius in that the cephalus of the latter has a pair of 

agglomerate pores. Martin 2004 suggests that they may be synonymous. All of the 

known species have been described from Brazil. The genus has been collected also in 

Colombia and Peru. According to Martin, this genus may possibly be a junior synonym 

of Leonardius. 

Reference: Penny & Arias (1980) - key to adults species. 

Bakerius amazonicus Penny & Arias 1980 

Bakerius amazonicus Penny & Arias 1980: 693. Holotype adult female. Brazil, Manaus, 

Amazonas, 16.x.1979, J. R. Arias, in INPA, Manaus, Brazil. 


Host: unknown, collected in a malaise trap. 

Bakerius attenuatus Bondar 1923 

Bakerius attenuatus Bondar 1923a: 38-40. Syntypes. Brazil: Bahia, G. Bondar, on Chromelia 

oligantha, MZU 


Host: Rubiaceae: Chomelia sp. 

29

Bakerius calmoni Bondar 1928 

Bakerius calmoni Bondar 1928b: 9-12. Syntypes. Brazil: Miguel Calmon, on Loranthaceae plant 

parasitic on coffee [Coffea arabica], MZU. 


Host: Loranthaceae. unidentified plant. 

Bakerius conspurcatus (Enderlein) 1909 

Aleurodicus conspurcatus Enderlein 1909a: 282-284. Syntypes. Brazil: Santa Catharina, 

Luderwalt, on undetermined host. 

Bakerius conspurcatus (Enderlein); Bondar 1923a: 40. 


Host: unidentified plant. 

Bakerius glandulosus Hempel 1938 

Bakerius glandulosus Hempel 1938: 313-314. Syntypes. Brazil: Sao Paulo, on Mikania amara. 


Host: Asteraceae: Mikania amara. 

Bakerius maculatus Penny & Arias 1980 

Bakerius maculatus Penny & Arias 1980: 692. Holotype adult male. Brazil, Manaus, Amazonas, 

11.ix.1979, J. R. Arias, in INPA, Manaus, Brazil. 


Host: unknown, collected in a malaise trap. 

Bakerius phrygilanthi Bondar 1923 

Bakerius phrygilanthi Bondar 1923a: 35-38. Syntypes. Brazil: Bahia, G. Bondar, on 

Phrygilanthus sp., MZU; Biezanko & Freitas 1939: 6; Costa Lima 1968: 108. 


Host: Loranthaceae: Loranthus acutifolium, Phrygilanthus sp.; Rubiaceae: Spermacoce 

verticillata. 

Bakerius sanguineus Bondar 1928 

Bakerius sanguineus Bondar 1928b: 7-9. Syntypes. Brazil: Bahia, G. Bondar, on Borreria 

verticilliata, USNM. 


Host: Rubiaceae: Borreria verticilliata. 

Bakerius sublatus Bondar 1928 

Bakerius sublatus Bondar 1928b: 5-7. Syntypes. Brazil: Bocca de Campo, on unidentified plant, 

MZU. 

Distribution: 2: Brazil. Host: unidentified plant. 

30

Bakerius sp. 

Collected in Peru, Huanuco, Santa Isabel, 22.x.2002, by G. Solis and P. Lozada 

Natural enemies: Encyrtidae: Metaphycus zdeneki. 

In addition, another species of Bakerius was collected in Colombia. 

31

Ceraleurodicus Hempel 1922 

Ceraleurodicus Hempel 1922: 6. Type-species: Ceraleurodicus splendidus Hempel 1922, by 

monotypy, MZU. 

Parudamoselis Visnya 1941a: 4-5. Type species. Parudamoselis kesselyaki, synonymy according 

to Mound & Halsey 1978. 

Radialeurodicus Bondar 1922a: 74. Syntypes. Brazil: Muritiba, G. Bondar, on aracazeiro 

[Psidium araca]; synonymy according to Costa Lima 1928:137. 

Comment: This is a Neotropical genus (Argentina, Brazil, Honduras, Mexico) with 1 species 

reported in a glasshouse in Hungary 

Reference: Martin 2004 - Diagnosis and comments. 



assymetricus x 

duckei x 

hempeli x 

keris x 

neivai x 

splendidus x 

varus x glasshouse 

Total = 7 species 0 7 1* 0 0 0 0 0 0 

Ceraleurodicus assymmetricus (Bondar) 1922 

Radialeurodicus assymmetricus Bondar 1922a: 74-77. Syntypes. Brazil: Bahia, G. Bondar, on 

Cocos nucifera, MZU. 

Ceraleurodicus assymmetricus (Bondar); Costa Lima 1928: 137. 



Ceraleurodicus duckei Penny & Arias 1980 

Ceraleurodicus duckei Penny & Arias 1980a: 903. Holotype adult male. Brazil, Amazonas, 

Reserva Ducke, 26 km. NE of Manaus, 15.vii.1979, J. R. Arias and N. D. Penny, in 

INPA, Manaus, Brazil. 


Host: unknown, collected in light trap 

Ceraleurodicus hempeli Costa Lima 1928 

Ceraleurodicus hempeli Costa Lima 1928: 138. Syntypes assumed to be from Brazil. 


32


Ceraleurodicus keris Martin 2004 

Ceraleurodicus keris Martin 2004: 35. Holotype puparium, Belize: CFR, Las Cuevas, 29.v.2004, 

J. Martin, ex. Laetia thamnia, BMNH. 

Distribution: 2: Belize, Costa Rica, Nicaragua, Panama. 

Host: Asteraceae: Eupatorium sp., Heterocondylus vitalbae; Burseraceae: Protium copal; 

Flacourtiaceae: Lunania parviflora, Laetia thamnia; Melastomataceae: Mouriri 

myrtilloides; Rubiaceae: Pentagonia macrophylla, Uncaria tomentosa. 

Ceraleurodicus neivai (Bondar) 1928 

Radialeurodicus neivai Bondar 1928b: 3-5. Syntypes. Brazil: Bahia State, Boa Nova, on 

unidentified host, MZU. 

Ceraleurodicus neivai (Bondar); Costa Lima 1928: 3-5. 



Ceraleurodicus spendidus Hempel 1922 

Ceraleurodicus splendidus Hempel 1922: 7. Syntypes. Brazil: Bahia, G. Bondar, on coconut 

[Cocos nucifera], MZU. 

Radialeurodicus cinereus 1922a: 78-82. Syntypes. Brazil: Bahia, G. Bondar, on coconut [Cocos 

nucifera], MZU; Costa Lima 1928: 137. 


Host: Arecaceae: Cocos nucifera; Myrtaceae: Psidium guajava. 

Ceraleurodicus varus (Bondar) 1928 

Radialeurodicus varus Bondar 1928b: 1-3. Syntypes. Brazil: Muritiba, G. Bondar, on aracazeiro 

[Psidium araca], USNM. 

Ceraleurodicus varus (Bondar) 1928; Costa Lima 1928: 137. 

Parudamoselis kesselyaki Visnya 1941a: 5-12. Syntypes. Hungary: Budapest, in Botanical 

Garden glasshouse, x.1939-1940, on Cymbidium lowianum and other plants; synonymy 

according to Martin et al. 2000:442, BTM. 

Ceraleurodicus kesselyaki (Visnya); Mound and Halsey 1978: 239. 

Distribution: 2: Brazil; 3: Hungary (in glasshouse). 

Host: Myrtaceae: Psidium araca; Orchidaceae: Cymbidium lowianum. 

33

Dialeurodicus Cockerell 1902 

Aleurodicus (Dialeurodicus) Cockerell 1902a: 280. Type-species: Aleurodicus cockerelli 

Quaintance 1900, by original designation. 

Dialeurodicus Cockerell; full genus Quaintance & Baker 1913: 26. 

Bondaria Sampson & Drews 1941: 149. Type-species: Bondaria radifera Sampson & Drews 

1941, by monotypy; synonomy Martin 2004: 39. 



bondariae x 

caballeroi x 

cockerellii x 

coehli x 

cornutus x 

frontalis x x 

maculatus x 

niger x 

radifera x 

silvestri x 

similis x 

tessellatus x 

Total = 12 species 1 12 0 0 0 0 0 0 0 

Comment: This is a Neotropical genus with 1 species, D. frontalis, known to occur in Florida 

(USA). 

Reference: Martin 2004 - Diagnosis and comments. 

Dialeurodicus bondariae Martin 2004 

Dialeurodicus bondariae Martin 2004: 40. Holotype puparium, Belize, CFR, San Pastor track, 

18.xi.1994, J. Martin on Lauraceae. 


Host: Lauraceae. 

Dialeurodicus caballeroi Martin 2004 

Dialeurodicus caballeroi Martin 2004: 41. Holotype puparium, Belize, CFR, Las Cuevas, 

10.ii.1996, J. Martin, on Persea americana. 

Distribution: 2: Belize; Nicaragua. 

34

Host: Lauraceae: Nectandra? sp, Persea americana. 

Dialeurodicus cockerellii (Quaintance) 1900 

Aleurodicus cockerellii Quaintance 1900: 45-46. Syntypes. Brazil: Sao Paulo State, Campinas, 

30.iii.1898, F. Noak, on leaves of myrtaceous plant, USNM. 

Aleurodicus (Dialeurodicus) cockerellii Quaintance; Cockerell1902a: 280. 

Dialeurodicus cockerellii (Quaintance); Quaintance & Baker 1913: 26. 

Distribution: 2: Brazil 

Host: Myrtaceae: Psidium cattleianum; myrtaceous plant. 

Diagnosis - Hempel (1923) dorsum covered with dense, thick mass of whitefly wax arranged in a 

marginal fringe and a submarginal oval ring on the underside of leaves. 

Dialeurodicus coelhi Bondar 1928 

Dialeurodicus coelhi Bondar 1928b: 12-14. Syntypes. Brazil: Bahia State, Santa Ignez, G. 

Bondar, on Myrtaceous tree, MZU 

Bakerius coelhoi [sic] (Bondar); Costa Lima 1936: 146. 


Host: Myrtaceae. 

Dialeurodicus cornutus Bondar 1923 

Dialeurodicus cornutus Bondar 1923a: 51-52. Syntypes. Brazil: Bahia, G. Bondar, on Miconia 

sp., MZU. 


Host: Melastomataceae: Miconia sp. 

Dialeurodicus frontalis Bondar 1923 

Dialeurodicus frontalis Bondar 1923a: 57-60. Syntypes. Brazil: Bahia State, Belmonte, G. 

Bondar, on Lauraceae tree, USNM. 

Distribution: 1: USA (FL); 2: Bahamas; Brazil. 

Host: Lauraceae: Nectandra coriacea, Ocotea catesbayna, Ocotea coriacea, Persea americana; 

unidentified plant. 

Dialeurodicus maculatus Bondar 1928 

Dialeurodicus maculatus Bondar 1928b: 14-16. Syntypes. Brazil: Bahia State, Cannaieras, G. 

Bondar. 


Host: Malpighiaceae: Byrsonima sp. 

Dialeurodicus niger Bondar 1923 

Dialeurodicus niger Bondar 1923a: 56-57. Syntypes. Brazil: Bahia, G. Bondar, on "aracazeiro" 

[Psidium araca] and Eugenia sp., MZU. 


Host: Myrtaceae: Psidium araca. 

35

Dialeurodicus radifera (Sampson & Drews) 1941 

Bondaria radifera Sampson & Drews 1941: 149. Syntypes. Mexico: Guerrero State, east of 

Acapulco, iii.1926, on undetermined vine, UCD. 

Dialeurodicus radifera (Sampson & Drews); new combination according to Martin 2004: 39. 


Host: undetermined vine. 

Dialeurodicus silvestri (Leonardi) 1910 

Aleurodicus silvestri Leonardi 1910: 320-322. Syntypes. Mexico: Jalapa, on undetermined plant. 

Dialeurodicus silvestri (Leonardi); Quaintance & Baker 1913: 28-30. 

Dialeurodicus tracheiferus Sampson & Drews 1941: 152-153. Syntypes. Mexico: Vera Cruz 

State, santa Lucrecia, iv.1926, on Eugenia sp.; synonymy according to Martin 2004: 43. 


Host: Loranthaceae; Myrtaceae: Eugenia sp. 

Dialeurodicus similis Bondar 1923 

Dialeurodicus similis Bondar 1923a: 49-51. Syntypes. Brazil, Bahia, G. Bondar, on Eugenia sp., 

MZU. 


Host: Myrtaceae: Eugenia sp. 

Dialeurodicus tessellatus Quaintance & Baker 1913 

Dialeurodicus tessellatus Quaintance & Baker 1913: 30-31. Syntypes. Brazil: Ceara, received 

i.1906, F. Richa, on Eugenia mitchelli, USNM. 


Host: Myrtaceae: Eugenia uniflora. 

Diagnosis - Baker and Moles (1921) Puparium dark brown or black with a narrow fringe of wax, 

adult pupa without wax. 

Dialeurodicus sp. 

Parasitoids: Aphelinidae: Encarsia luteola, Encarsia nigricephala, Encarsia protransvena; 

Eretmocerus sp.; Signiphoridae: Signiphora aleyrodis [probably a hyperparasitoid]. 

36

Eudialeurodicus Quaintance & Baker 1915 

Eudialeurodicus Quaintance & Baker 1915b: 369. Type-species: Eudialeurodicus bodkini 

Quaintance and Baker 1915, by monotypy. 



bodkini x 

Total = 1 species 0 1 0 0 0 0 0 0 0 

Diagnosis - According to Quaintance and Baker (1915), the forewing is similar to Dialeurodicus 

with radius, radial sector and media retained; vertex rounded, frons produced beyond 

the vertex; antennae 7-segmented; puparium flat, resembling Dialeurodicus, but with 

one or more pairs of reduced compound pores; vasiform orifice small, lingula included 

and broadly rounded. Adult - forewing with radius, radial sector and media veins, 

vertex rounded, frons produced beyond the vertex, antennae with 7 segments. One 

Neotropical species known from Brazil and Guyana on Fabaceae. 

Eudialeurodicus bodkini Quaintance & Baker 1915 

Eudialeurodicus bodkini Quaintance & Baker 1915b: 369-372. Syntypes. Guyana: Berbice, Rose 

Hall Plantation, 2.iii.1915, G. E. Bodkin, on Erythrina glauca, USNM. 

Distribution: 2: Brazil; Guyana. 

Host: Fabaceae: Erythrina fusca, Inga sp. 

Natural enemies: Aphelinidae: Encarsiella magniclava; Eulophidae: Entedononecremnus unicus. 

Diagnosis and Characteristics 

Field specimen - Body flat, broadly pear-shaped, yellow with a large dark brown area 

on the thorax and on the abdominal region; found along the midrib and larger leaf veins, with 

copious amounts of white wax. 

Slide specimen - Puparium with one pair of compound pores on caudal segment; a pair 

of reduced pores present on third abdominal segment; vasiform orifice small, transversely 

elliptical, operculum filling about half the orifice, transversely elongate with rounded side and 

concave caudal margin; lingula short, broadly rounded, about 1/3 as long as vasiform orifice. 

Adult female forewing white with many groups of small dark dots. 

37

Lecanoideus Quaintance and Baker 1913 

Aleurodicus (Lecanoideus) Quaintance & Baker 1913: 70. Type-species: Aleurodicus 

(Lecanoideus) giganteus, by original designation. 

Lecanoideus Quaintance & Baker; full genus by Costa Lima 1928: 133. 



floccissimus x x 

mirabilis x 

Total = 2 species 0 2 1 0 0 0 0 0 0 

Comment: This is a Neotropical genus with 2 described species, one of which, was described 

from the Canary Islands. 

Reference - Martin (2004) - diagnosis and comments. 

Lecanoideus floccissimus Martin, Hernandez-Suarez and Carnero 1997 

Lecanoideus floccissimus Martin, Hernandez-Suarez and Carnero 1997, in Martin et al. 1997: 

1261. Holotype pupal case. Canary Islands: Tenerife, Santa Cruz de Tenerife, 

26.v.1996, E. Hernandez-Suarez, on Ficus sp., BMNH, NHM. 

Distribution: 2: Ecuador; Panama, Peru, Trinidad; 4: Canary Islands. 

Host: Agavaceae: Yucca aloifolia; Anacardiaceae: Mangifera indica, Schinus terebinthifolius; 

Apocynaceae: Nerium oleander, Acokanthera oblongifolia; Arecaceae: Arecastrum 

romanzoffianum, Elaeis guineensis, Chrysalidocarpus lutescens, Howeia forsteriana, 

Brahea armata, Cocos nucifera, Livistona chinensis, Mascarena verschaffeltii, Phoenix 

dactylifera, Phoenix canariensis, Phoenix roebelinii; Caricaceae: Carica papaya; 

Combretaceae: Terminalia catappa; Dracaenaceae: Dracaena draco; Euphorbiaceae: 

Euphorbia pulcherrima, Acalypha wilkesiana; Fabaceae: Bauhinia variegata; 

Malvaceae: Hibiscus rosa-sinensis, Malvaviscus penduliflorus; Moraceae: Ficus 

macrophylla, Ficus sp., Ficus elastica, Ficus benjamina; Musaceae: Musa acuminata, 

Musa sp., Strelitzia reginae, Strelitzia nicolai; Myrtaceae: Psidium guajava; 

Polygonaceae: Coccoloba uvifera; Proteaceae: Stenocarpus sinuatus; Rutaceae: Citrus 

aurantium; Verbenaceae: Lantana camara. 

Natural enemies: Aphelinidae: Encarsia guadeloupae, Encarsia hispida, Encarsiella noyesi. 

Lecanoideus mirabilis (Cockerell) 1898 

Aleurodes mirabilis Cockerell 1898b: 225. Syntypes. Mexico: Tabasco, Boca del Usumacinta, 

8.vii.1897, C. H. T. Townsend, on "laurel". 

Aleurodicus (Lecanoideus) mirabilis (Cockerell); Quaintance & Baker 1913:72. 

Lecanoideus mirabilis (Cockerell); Mound and Halsey 1978: 243. 

Aleurodicus (Lecanoideus) giganteus Quaintance & Baker 1913: 70. Syntypes. Brazil: 

Pernambuco, 28.xii.1882, A. Koebele, on Ficus sp.; synonymy according to Martin 

2004: 18. 

38

39 

Lecanoideus giganteus (Quaintance & Baker); Costa Lima 1928: 133. 


Host: Annonaceae: Annona cheirimola, Annona muricata, Annona squamosa, Cananga odorata, 

Polyalthia longifolia; Chrysobalanaceae: Licania tomentosa; Euphorbiaceae: Hevea 

brasiliensis; Lauraceae: Laurus nobilis; Rosaceae: Licania tomentosa 

Natural enemies: Predators - NEUROPTERA/Chrysopidae: Chrysoperla externa, 

DIPTERA/Cecidomyidae: Cleodiplosis aleyrodici; Parasitoids - Aphelinidae: Encarsia 

guadeloupae, Encarsia sp.; Encarsiella sp. 

Diagnosis - Puparium light yellow covered with abundant wax, found on the underside of leaves 

along the midrib and major veins.

Leonardius Quaintance & Baker 1913 

Leonardius Quaintance & Baker 1913: 33. Type-species: Aleurodicus lahillei Leonardi 1910, by 

monotypy; Martin 2004: 44. 



kellyae x 

lahillei x 

Total = 2 species 0 2 0 0 0 0 0 0 0 

Comment: This is a Neotropical genus that contains 2 described species. 

Diagnosis -Dorsum with only 2 pairs of abdominal compound pores on submedian area of A3 

and A4, one pair of agglomerate pores on cephalus; 4 pairs of cephalothoracic setae 

present. Differs from Bakerius in that the cephalus of the latter has a pair of compound 

pores. Martin 2004 suggest that they may be synonymous. Adult - forewing rounded, 

usually mottled color with radius, radial sector and media veins, vertex rounded, frons 

produced beyond the vertex, antennae with 7 segments with the third segment the longest 

one. Martin (2004) reported a new species in this genus from Colombia. 

Reference: Martin (2004) - diagnosis and comments. 

Leonardius kellyae Martin 2004 

Leonardius kellyae Martin 2004: 45. Holotype puparium, Belize, CFR, Las Cuevas, 2.viii.2003, 

S. Kelly and A. Polaszek, on Loranthaceae, BMNH. 


Host: Loranthaceae. 

Leonardius lahillei (Leonardi) 1910 

Aleurodicus lahillei Leonardi 1910: 316-320. Syntypes. Argentina: on undetermined plant. 

Leonardius lahillei (Leonardi); Quaintance & Baker 1913: 33; Dozier 1936: 145. 

Leonardius loranthi Bondar 1923a: 44-46. Syntypes. Brazil: Bahia, Belmonte, G. Bondar, on 

"herva de passarinho" [Phrygilanthus sp.], MZU; synonymy Martin 2004: 46. 

Distribution: 2: Argentina; Brazil; Colombia; Puerto Rico. 

Host: unidentified plant, Balanophoraceae: Oryctanthus occidentalis; Loranthaceae: 

Phrygilanthus sp., Steirotis flexicaulis; Rosaceae: Prunus sp.; Viscaceae: Phoradendron 

sp. 

Description according to Hempel: 40. Puparium oval, dark brown, covered with white dust and a 

fringe of white wax around the margin; dorsum with series of 2 large compound glands 

situation on the anterior part of the abdomen - other series composed of 5 pairs of 

agglomerate glands (1 pair on the cephalothorax, 4 pairs on the abdomen posterior to the 

compound glands; margin with 3 rows of papillae-like glands and one row of fine setae; 

vasiform orifice enlarged and long cordate 

40

Metaleurodicus Quaintance & Baker 1913 

Aleurodicus (Metaleurodicus) Quaintance & Baker 1913. Type-species: Aleurodicus minima 

Quaintance 1900, by original designation. 

Metaleurodicus Quaintance and Baker 1913; full genus by Bondar 1923:81. 

Pseudaleurodicus Hempel 1922: 9. Type-species: Pseudaleurodicus bahiensis Hempel, by 

monotypy; synonymized with Aleuronudus according to Costa Lima 1936; however, 

Martin 2004 regards it as a junior synonym of Metaleurodicus. 



arcanus x 

bahiensis x 

cardini x x 

griseus x x 

lacerdae x 

minimus x 

phalaenoides x 

stelliferus x 

tenuis x 

variporus x 

Total = 10 species 2 10 0 0 0 0 0 0 0 

Comment: This is a Neotropical genus with 2 species known to occur in the USA (Florida). 

Diagnosis - Puparium with cephalothoracic submedian setae absent; 4-6 pairs of compound pores 

present, forming a smooth arc on each side of the body; lingula fully included within the 

vasiform orifice; most similar to Aleuronudus but that genus has the compound pores 

offset, not forming a smooth arc. 

Reference: Martin (2004) - diagnosis, comments and key to species of Belize. 

41

(Modified from Martin 2004) 

Key to the Metaleurodicus Species 

1(1b) Abdomen with 4 pairs of compound pores; USA, Bermuda, Cuba, Jamaica, on Psidium, 

Citrus, and other hosts. .................................................................................. cardini (Back) 

1b. Abdomen with 5-6 pairs.......................................................................................................2 

2(1b) Abdomen with 5 pairs of compound pores; Belize on Rubiaceae................arcanus Martin 

2b. Abdomen with 6 pairs of compound pores..........................................................................3 

3(2b). Abdominal compound pores of 2-3 different sizes..............................................................4 

3b. Abdominal compound pores similar in size; caudal pores slightly larger in M. minimus...5 

4(3) Compound pores variable sizes, but C1 smaller than A1-3 pores; Belize on Asteraceae, 

Urticaceae .................................................................................................. variporus Martin 

4b. Compound pores on head, and A1-3 much larger than others, A5,6,7 small; Brazil on 

Cocos, Musa, Artocarpus...................................................................... bahiensis (Hempel) 

5(3b) Caudal compound pore larger than others; lateral margin with row of cylindrical-shaped 

structures; star-shaped pores very large, covering the dorsum, submarginal setae long and 

slender; Puerto Rico on Citrus, Psidium, Cestrum ............................ minimus (Quaintance) 

5b. Caudal pore not larger than others; lateral margin with row of cylindrical-shaped 

structures..............................................................................................................................6 

6(5b) Star-shaped pores exceptionally large, vasiform orifice elongate; Brazil 

...................................................................................................................stelliferus Bondar 

6b. Star-shaped pores not large; vasiform orifice not elongate .................................................7 

7(6b) Abdominal compound pores appearing circular, submarginal setae inset further from 

margin, longer and coarser; faint compound pore-like structures on T2, T3; found on 

upperside of Eugenia leaves in Barbados, Belize, Jamaica, Puerto Rico, USA: FL. 

.................................................................................................................... .griseus (Dozier) 

7b. Abdominal compound pores appearing cylindrical, submarginal setae very small, located 

on extreme outer submargin; Belize on Eugenia? .......................................... tenuis Martin. 

Note: Species: M. lacerdae (Brazil on Annona) and M. phalaenoides (Argentina, Chile on 

Cestrum) not included in key (description, illustrations, reference specimens not available). 

42

43 

Metaleurodicus arcanus Martin 2004 

Metaleurodicus arcanus Martin 2004: 48. Holotype puparium, Belize, CFR, Monkey Tail Track, 

4.iv.2003, J. Martin, ex. Chiococca ?alba, BMNH. 


Host: Rubiaceae: Chiococca alba? 

Metaleurodicus bahiensis (Hempel) 1922 

Pseudaleurodicus bahiensis Hempel 1922: 9-10. Syntypes. Brazil: Bahia, ix.1921, G. Bondar, on 

Cocos nucifera, MZU. 

Aleuronudus bahiensis (Hempel); Bondar 1923a: 91; Costa Lima 1936: 146. 

Metaleurodicus bahiensis (Hempel); Martin 2004: 47. 


Host: Arecaceae: Cocos nucifera; Moraceae: Artocarpus heterophyllus; Musaceae: Musa 

sapientum. 

Metaleurodicus cardini (Back) 1912 

Aleurodicus cardini Back 1912: 148-151. Syntypes. Cuba: Havana and Santiago de las Vegas, 

xi.1910, E. A. Back, on Psidium guajava, USNM. 

Aleurodicus (Metaleurodicus) cardini Back; Quaintance & Baker 1913: 75-77 

Metaleurodicus cardini (Back); Bondar 1923a: 81; Mound and Halsey 1978: 244. 

Distribution: 1: USA (FL; MD); 2: Bermuda; Cuba; Dominican Republic, Haiti, Jamaica; Puerto 

Rico; St. Thomas. 

Host: Annonaceae: Annona cheirimola, Annona sp.; Apocynaceae: Plumeria sp.; Fabaceae: 

Sophora tomentosa; Malpighiaceae: Malpighia urens, Malpighia polytricha, Malpighia 

glabra; Myrtaceae: Eugenia sp., Melaleuca leucadendra, Pimenta dioica; Psidium 

guajava, Psidium cymosum, Psidium littorale, Psidium cattleianum; Punicaceae: Punica 

granatum; Rutaceae: Citrus sp, Citrus sinensis; Sapotaceae: Dipholis salicifolia; 

Sterculiaceae: Guazuma tomentosa; Verbenaceae: Duranta repens, Duranta erecta; 

Citharexylum fruticosum; Zygophyllaceae: Guaiacum officinale. 

Natural enemies: Parasitoids - Aphelinidae: Encarsia sp. (in Cuba); Predators - 

COLEOPTERA/Coccinellidae: Delphastus diversipes; DIPTERA/Syrphidae: Ocyptamus 

parvicornis, Pseudodoros clavatus; NEUROPTERA/Chrysopidae: Chrysopodes 

(Neosuarius) collaris. 

Metaleurodicus griseus (Dozier) 1936 

Aleurodicus griseus Dozier 1936: 143-144. Syntypes. Puerto Rico: along roadside near Punta 

Cangrejos, 19.vii.1925, H. L. Dozier, on Eugenia buxifolia, USNM. 

Metaleurodicus griseus (Dozier); Hamon 1989: 1. 

Distribution: 1: USA (FL) 2: Belize; Barbados; Bermuda; Cuba; Jamaica; Puerto Rico. 

Hosts: Myrtaceae: Eugenia axillaris, Eugenia foetida. 

Natural enemies: NEUROPTERA/Chrysopidae: Ceraeochrysa cincta. 

Metaleurodicus lacerdae (Signoret) 1883 

Aleyrodes lacerdae Signoret 1883: 93. Syntypes. No locality given [assumed to be Brazil based 

on name and address of collector], Antonio de Lacerda, on Annona sylvatica.

Aleurodicus lacerdae (Signoret); Quaintance & Baker 1913: 78 

Metaleurodicus lacerdae (Signoret); Bondar 1923a: 13. 


Host: Annonaceae: Annona sylvatica. 

Metaleurodicus minimus (Quaintance) 1900 

Aleurodicus minima Quaintance 1900: 47-48. Syntypes. Puerto Rico: Bayamon, received 

28.i.1889, A. Busck, on Psidium guajava, USNM. 

Aleurodicus (Metaleurodicus) minimus Quaintance; Quaintance & Baker 1913: 77-78. 

Metaleurodicus minimus (Quaintance); Bondar 1923a: 81. 

Distribution: 2: Honduras; Puerto Rico; Virgin Islands. 

Host: Myrtaceae: Psidium guajava; Rutaceae: Citrus sinensis; Solanaceae: Cestrum diurnum. 

Natural enemies: Aphelinidae: Encarsia hispida. 

Metaleurodicus phalaenoides (Blanchard) 1852 

Aleurodes phalaenoides Blanchard 1852: 319-320. Syntypes. Chile: Santiago, on Cestrum 

parqui. 

Aleurodicus phalaenoides (Blanchard); Quaintance & Baker 1913: 79. 

Metaleurodicus phalaenoides (Blanchard); Bondar 1923a: 13. 

Aleurodes phalaroides Lataste; nomen nudum and synonymy according to Quaintance 1900: 17; 

nomen nudum according to Martin & Mound 2007. 

Distribution: 2: Argentina; Chile. 

Host: Solanaceae: Cestrum parqui. 

Metaleurodicus stelliferus Bondar 1923 

Metaleurodicus stelliferus Bondar 1923a: 82-83. Syntypes. Brazil: Belmonte, G. Bondar, on 

Meliaceae plant, MZU. 


Host: Meliaceae: unidentified plant. 

Metaleurodicus tenuis Martin 2004 

Metaleurodicus tenuis Martin 2004: 49. Holotype puparium, Belize, CFR, Las Cuevas, 

29.iii.2003, J. Martin, on Myrtaceae probably Eugenia, BMNH. 


Host: Myrtaceae: probably Eugenia sp. 

Metaleurodicus variporus Martin 2004 

Metaleurodicus variporus Martin 2004: 51. Holotype puparium, Belize, CFR, San Pastor track, 

28.ii.1996, J. Martin, on Lasianthaea fruticosa, BMNH. 


Host: Asteraceae: Lasianthaea fruticosa; Euphorbiaceae: Acalypha costaricensis; Urticeae: 

Urera sp. 

44

Nealeurodicus Hempel 1923 

Nealeurodicus Hempel 1923: 1170. Type-species: Nealeurodicus paulistus Hempel 1923, by 

monotypy. 



altissimus x 

bakeri x 

fallax x 

ingae x 

moreirai x 

octifer x 

paulistus x 

petiolaris x 

Total = 8 species 0 8 0 0 0 0 0 0 0 

Comment: This is a Neotropical genus with 8 described species. 

Reference - Martin 2004: 54. Key to Nealeurodicus species in Belize. 

Nealeurodicus altissimus (Quaintance) 1900 

Aleurodes altissima Quaintance 1900: 20-21. Syntypes. Mexico: Tabasco State, San Francisco de 

Peal, vii.1897, C. H. T. Townsend, on "palo de gusano", USNM. 

Aleurodicus altissima (Quaintance); Cockerell 1902a: 280. 

Aleurodicus (Metaleurodicus) altissimus (Quaintance); Quaintance & Baker 1913: 73. 

Metaleurodicus altissimus (Quaintance); Bondar 1923a: 81. 

Radialeurodicus altissimus (Quaintance); Baker 1937: 608. 

Ceraleurodicus altissimus (Quaintance); Mound and Halsey 1978: 238 

Nealeurodicus altissimus (Quaintance); Martin 2004: 55. 

Distribution: 2: Belize; Colombia; Costa Rica; Mexico; Nicaragua; Panama; Venezuela. 

Host: Fabaceae: Cassia sp., Canavalia sp., Fabaceae: Inga sp., Loncohcarpus sp., Machaerium 

sp; Verbenaceae: Lippia myriocephala. 

Natural enemies: Aphelinidae: Dirphys encantadora, Dirphys mexicana; Eulophidae: 

Entedononecremnus hansoni. 

Reference: Martin (2004) - diagnosis, comments and key to species of Belize. 

Nealeurodicus bakeri (Bondar) 1923 

Radialeurodicus bakeri Bondar 1923a: 21-24. Syntypes. Brazil: Bahia State, Belmonte, G. 

Bondar, on Cecropia sp., MZU. 

Ceraleurodicus bakeri (Bondar); Costa Lima 1928: 137; Mound & Halsey 1978: 238. 

Nealeurodicus bakeri (Bondar); Martin 2004: 56. 

45

46 

Distribution: 2: Belize; Brazil; Costa Rica; Guatemala; Nicaragua; Panama; Trinidad; Venezuela. 

Host: Fabaceae: Inga sp.; Moraceae: Cecropia sp. 

Nealeurodicus fallax Martin 2004 

Nealeurodicus fallax Martin 2004: 57. Holotype puparium, Belize, CFR, San Pastor Track, 

31.v.2004, J. Martin, on Sebastiania longicuspis, BMNH. 


Host: Euphorbiaceae: Sebastiania longicuspis; Lauraceae: Nectandra nitida?, Persea americana. 

Nealeurodicus ingae (Baker) 1937 

Radialeurodicus altissimus ingae Baker 1937: 608-609. Syntypes. Mexico: Chiapas State, El 

Vergel, 3.vi.1935, A. Dampf, on Inga sp., USNM. 

Ceraleurodicus ingae (Baker); Mound and Halsey 1978: 239. 

Nealeurodicus ingae (Baker); Martin 2004. 

Distribution: 2: Brazil; Mexico. 

Host: Fabaceae: Inga sp., Pithecellobium dulce, Pithecellobium sp.; Lauraceae: Persea borbonia. 

Nealeurodicus moreirai (Costa Lima) 1928 

Ceraleurodicus moreirai Costa Lima 1928: 139. Syntypes. Brazil: Rio de Janeiro, A. F. 

Magarinos Torres, on Annona squamosa, BMNH; Mound & Halsey 1978: 239. 

Nealeurodicus moreirai (Costa Lima); Martin 2004. 


Host: Annonaceae: Annona squamosa; Sapotaceae: Manilkara zapota. 

Note: Mound & Halsey (1978: 239) synonymized Radialeurodicus melzeri Laing 1930 with this 

species; however, Martin 2004: 54 disagreed with the synonymy and regarded R. melzeri 

as a junior synonym of Nealeurodicus paulistis. 

Nealeurodicus octifer (Bondar) 1923 

Radialeurodicus octifer Bondar 1923a: 17-21. Syntypes. Brazil: Bahia, G. Bondar, on Inga sp. 

and Cecropia sp., USNM. 

Ceraleurodicus octifer (Bondar); Costa Lima 1928: 137; Mound & Halsey 1978: 240. 

Nealeurodicus octifer (Bondar); Martin 2004. 


Host: Fabaceae: Inga sp.; Moraceae: Cecropia sp. 

Nealeurodicus paulistis Hempel 1923 

Nealeurodicus paulistus Hempel 1923: 1170-1171. Syntypes. Brazil: Sao Paulo, 15.x.1919, A. 

Hempel, on leaves of cultivate "jaboticaba" [Myrciaria cauliflora], MZU. 


Host: Arecaceae; Myrtaceae: Myrciaria cauliflora. 

Nealeurodicus petiolaris Martin 2004 

Nealeurodicus petiolaris Martin 2004: 59. Holotype puparium, Belize, CFR, Las Cuevas, 

6.iv.2003, J. Martin, on Protium glabrum, BMNH. 

Distribution: 2: Belize.

Host: Burseraceae: Protium glabrum. 

Natural enemies: Aphelinidae: Encarsia sp. 

47

Nipaleyrodes Takahashi 1951 

Nipaleyrodes Takahashi 1951c: 3-4. Type-species: Nipaleyrodes elongata Takahashi 1951, by 

monotypy. 

Species Distribution: 


elongata x x 

Total = 1 species 0 0 0 0 0 1 1 0 

Comment: This genus contains one species known from the Oriental region (Malaysia) and from 

Sulawesi on a palm (Nipa). Its placement in the Aleurodicinae is debatable; it has 

approximately 25 pairs of octagonal pores in a row just inside the lateral margin that 

appears to be “window-like” instead of being compound. The lingula has 1 pair of long 

setae at its apex, and there is possibly another short seta present at its base, but it is not 

clear in the specimens at hand. It is similar to Viennotaleyrodes (Bink-Moenen 1983) for 

having a similar series of pores along the submargin. 

References: Takahashi 1951a; Martin 1988. 

Nipaleyrodes elongata Takahashi 1951 

Nipaleyrodes elongata Takahashi 1951c: 3-4. Syntypes. Malaya: Kuala Selangor, 18.vi.1943, R. 

Takahashi, on "nipah palm" [Nypa sp.], BMNH; Martin 1988: 62. 

Distribution: 6: Malaya; Malaysia; 7: Sulawesi. 

Host: Arecaceae: Nypa sp. 

Diagnosis - Field specimen: puparium pale brown, elongate (2.2 mm long), narrow, flattened; 

Slide specimen - 1 pair of cephalic setae; transverse suture present above mouthparts; 

dorsum with single row of about 25 large octagonal pores on each lateral margin; margin 

not crenate, vasiform orifice triangular, not elevated, and without teeth; operculum much 

wider than long, occupying less than half the orifice; lingula large, knobbed, without long 

setae, and extending to about the posterior end of the vasiform orifice; caudal furrow 

absent; similar to Stenaleyrodes vinsoni but lacks submarginal papillae, has large 

octagonal pores and a shorter vasiform orifice. 

48

Octaleurodicus Hempel 1922 

Octaleurodicus Hempel 1922: 7-8. Type-species: Octaleurodicus nitidus Hempel 1922, by 

monotypy. 

Quaintancius Bondar 1922a: 74. Type-species: Quaintancius rubrus Bondar 1922 [=O. nitidus], 

by monotypy; synonymy according to Costa Lima 1928: 74. 

Comment: This Neotropical genus contains 2 species known from South America on coconut, 

Cocos nucifera (Arecaceae). 

Diagnosis - Hempel (1922) large body, flattened and oval; dorsum irregularly reticulate and 

bears 4 pairs of compound glands; vasiform orifice is cordate; margin is thin and 

ornamented with a row of spatulate plates with their interior edge zig-zag in form. O 

nitidus with 4 pairs of bowl-shaped glands, all of different sizes. 



nididus x 

pulcherrimus x 

Total = 2 species 0 2 0 0 0 0 0 0 0 

Octaleurodicus nitidus Hempel 1922 

Octaleurodicus nitidus Hempel 1922: 8-9. Syntypes. Brazil: Bahia, G. Bondar, on coconut 


Ceraleurodicus nitidus (Hempel); Costa Lima 1928: 137. 

Octaleurodicus nitidus Hempel; Mound & Halsey 1978: 247. 

Quaintancius rubrus Bondar Bondar 1922a: 74. Syntypes. Brazil: Bahia, on coconut, Cocos 

nucifera; synonymy according to Costa Lima 1928: 137. 



Octaleurodicus pulcherrimus (Quaintance & Baker) 1913 

Dialeurodicus pulcherrimus Quaintance & Baker 1913: 31-33. Syntypes. Trinidad: F. W. Urich, 

on coconut [Cocos nucifera], USNM. 

Quaintancius pulcherrimus (Quaintance & Baker); Bondar 1923a: 29. 

Ceraleurodicus pulcherrimus (Quaintance & Baker); Costa Lima 1928: 13. 

Octaleurodicus pulcherrimus (Quaintance & Baker); Mound and Halsey 1978: 247. 

Distribution: 2: Guyana; Trinidad. 


49

Paraleyrodes Quaintance 1909 

Paraleyrodes Quaintance 1909: 169-170. Type species: Aleurodes perseae Quaintance 1900, by 

monotypy. 

Comment: This is a Neotropical genus with all 16 species known from this region. Species 

present in the Old World probably represent introductions of New World species into the 

region. 

Diagnosis - Adult - forewing with radius and short rudiment of media vein, vertex rounded, 

antennae 4-segmented with the third segment the longest one. 

Reference: Martin (2004) - diagnosis, comments 



ancora x 

bondari x x x x x x 

cervus x 

citri x x 

citricolus x x 

crateraformans x 

goyabae x 

minei x x x x x x 

naranjae x 

perplexus x 

perseae x x x 

proximus x 

pseudonaranjae x x x x x 

pulverans x 

singularis x 

triungulae x 

urichii x 

Total = 17 species 6 17 3 2 2 2 0 1 3 

Paraleyrodes ancora Martin 2004 

Paraleyrodes ancora Martin 2004: 63. Holotype puparium, Belize, CFR, Monkey Tail Track, 

22.vi.2002, J. H. Martin, on Asteraceae, possibly Lasianthaea, BMNH. 

Distribution: 2: Belize; Nicaragua. 

50

51 

Host: Asteraceae: Lasianthaea? sp.; Dilleniaceae: Tetracera sp.; Lauraceae: Persea americana; 

Moraceae: Brosimum sp., Trophis? Sp.; Polygonaceae: Coccoloba belizensis. 

Paraleyrodes bondari Peracchi 1971 

Paraleyrodes bondari Peracchi 1971: 146-148. Syntypes. Brazil: Rio de Janeiro, Guanabara, 

3.iii.1967, A. L. Peracchi, on Citrus sp.; Martin 2004: 65. 

Distribution: 1: USA (CA; FL; HI); 2: Belize; Brazil; Honduras; *Puerto Rico; Venezuela; 3: 

Madeira; 4: Mauritius; Reunion; 6: Taiwan, Comosus Islands; 9: Hawaii. 

Host: Arecaceae: Cocos nucifera; Meliaceae: Trichilia ? sp.; Rutaceae: Citrus sp. 

US Port Interceptions: Hawaii on Hibiscus rosa-sinensis; Jamaica on Annona squamosa; 

Mexico on Chamaedorea sp.; Puerto Rico, San Juan, 14.iv.2003, J. Melendez, on Citrus 

sp. 

Paraleyrodes cervus Martin 2004 

Paraleyrodes cervus Martin 2004: 65. Holotype puparium, Belize, CFR, Las Cuevas, 16.ii.1996, 

J. H. Martin, on Bignoniaceae vine, BMNH. 


Host: Bignoniaceae. 

Paraleyrodes citri Bondar 1931 

Paraleyrodes citri Bondar 1931: 24. Syntypes. Brazil: Bahia, G. Bondar, on Citrus aurantium, 

MZU; Meyerdirk et al. 1980. 

Distribution: 1: USA (FL); 2: Brazil, Mexico, Panama. 

Host: Rutaceae: Citrus aurantium L. 

Distribution: 1: USA; 2: Brazil, Mexico, Panama. 

Natural enemies: COLEOPTERA/Coccinellidae: Delphastus pusillus, Nephaspis oculatus. 

US Port Interception: Mexico on Chamaedorea sp. 

Paraleyrodes citricolus Costa Lima 1928 

Paraleyrodes citricolus Costa Lima 1928: 136. Holotype. Brazil?: A. G. Maciel, on Citrus 

aurantium; Martin 2004: 66 

Distribution: 2: Belize; Brazil; Colombia; Costa Rica; Ecuador; Guatemala; Mexico; Panama; 3: 

Azores; Madeira. 

Host: Rutaceae: Citrus aurantium. 

Paraleyrodes crateraformans Bondar 1922 

Paraleyrodes crateraformans Bondar 1922a: 85. Syntypes. Brazil: Bahia, G. Bondar, on 

"coqueiro" [Cocos nucifera], 'cacaoeiro [Theobroma cacao], sapotiseiro [Achrus sapota] 

and other plants, MZU 


Host: Arecaceae: Cocos nucifera; Sapotaceae: Manilkara zapota; Sterculiaceae: Theobroma 

cacao. 

Paraleyrodes goyabae (Goldi) 1886 

Aleurodes goyabae Goeldi 1886. Syntypes. Brazil: Rio de Janeiro, on Psidium guajava and

Laurus persea. 

Paraleyrodes goyabae (Goldi); Bondar 1923a: 93. 

Distribution: 2: Barbados; Brazil. 

Host: Chrysobalanaceae: Licania tomentosa; Lauraceae: Persea gratissima, Laurus persea; 

Myrtaceae: Psidium guajava. 

Natural enemies: Aphelinidae: Encarsia sp. (in Barbados) 

Paraleyrodes minei Iaccarino 1990 

Paraleyrodes minei Iaccarino 1990: 132. Holotype male. Syria: Tartous, 17.viii.1988, on Citrus 

aurantium; Martin 1996: 1856. 

Distribution: 1: USA (CA, FL, TX); 2: Belize, Bermuda, Guatemala, Honduras, Mexico; Puerto 

Rico; 3: Iran; Israel; Lebanon, Morocco, Spain, Syria; Turkey; 4: Benin; 5: Hong Kong; 

9: Hawaii*. 

Host: Annonaceae: Annona glabra; Araceae: Anthurium sp., Syngonium podophyllum; 

Arecaceae: Chamaedorea sp., Ptychosperma elegans; Asteraceae: Lasianthaea fruticosa; 

Euphorbiaceae: Cnidoscolus chayamansa; Lauraceae: Nectandra coriacea, Persea 

americana, Persea borbonia; Myrtaceae: Psidium guajava; Piperaceae: Piper sp.; 

Polygonaceae: Coccoloba diversifolia, Coccoloba uvifera; Rhizophoraceae: Rhizophora 

mangle; Rutaceae: Citrus aurantifolia, Citrus aurantium, Citrus limon, Citrus sinensis, 

Citrus x paradisi. 

Natural enemies: Predators - COLEOPTERA/Coccinellidae: Clitostethus arcuatus, Serangium 

parcesetosum; Parasitoids - HYMENOPTERA/ Aphelinidae Encarsia dominicana Evans, 

Encarsia parvella group, Encarsia sp., Encarsia variegata Howard 1908. 

Comment: Hawaii (new record J. Dooley 2006). 

US Port Interceptions: Hawaii on Cocos nucifera; Jamaica on Annona squamosa; Mexico on 

Citrus sinensis, Citrus sp., Persea americana, Psidium guajava. 

Paraleyrodes naranjae Dozier 1927 

Paraleyrodes naranjae Dozier 1927: 853-855. Syntypes. Puerto Rico: Santurce, 21.xii.1924, H. 

L. Dozier, on "sour-orange" [Citrus sp.], USNM. 


Host: Rutaceae: Citrus aurantium, Citrus aurantiifolia. 

Natural enemies: Aphelinidae: Encarsia variegata; Predators: COLEOPTERA/Coccinellidae: 

Nephaspis oculatus. 

US Port Interception: Bahamas on Cocos nucifera. 

Note: Specimens of this species may have been confused with Paraleyrodes pseudonaranjae 

Martin 2001 in the past. 

Paraleyrodes perplexus Martin 2004 

Paraleyrodes perplexus Martin 2004: 67. Holotype puparium, Belize, CFR, Las Cuevas, 

6.vi.2004, J. Martin; on Piper yucatanensis, BMNH. 


Host: Asteraceae; Flacourtiaceae: Casearia sylvestris; Piperaceae: Piper yucatanense. 

52

53 

Paraleyrodes perseae (Quaintance) 1900 

Aleurodes perseae Quaintance 1900: 32-33. Syntypes. USA: Florida, Fort George, 22.iv.1880, R. 

S. Turner, on Persea carolinensis, USNM. 

Aleurodicus perseae (Quaintance); Cockerell 1903b: 663. 

Paraleyrodes perseae (Quaintance); Quaintance 1909: 170. 

Distribution: 1: USA (FL); 2: Cuba, Mexico, Venezuela; 6: Philippines 

Host: Caprifoliaceae: Viburnum sp.; Ebenaceae: Diospyros sp.; Ericaceae: Lyonia lucida 

Ericaceae: Gaylussacia sp.; Euphorbiaceae: Hevea sp.; Fagaceae: Quercus nigra; 

Lauraceae: Persea borbonia; Persea americana, Persea carolinensis; Magnoliaceae: 

Magnolia sp.; Myrtaceae: Psidium guajava; Rubiacae: Coprosma sp.; Rutaceae: Citrus 

sinensis, Citrus aurantium, Citrus sinensis, Citrus x tangelo; Sterculiaceae: Guazuma 

tomentosa. 

Natural enemies: Fungus - Aschersonia aurantiaca; Parasitoids - Aphelinidae: Encarsia 

variegata Predators - COLEOPTERA/Coccinellidae: Scymnus sp. 

US Port Interception: Mexico on Persea americana. 

Paraleyrodes proximus Teran 1979 

Paraleyrodes proximus Teran 1979: 170. Holotype puparium, Argentina, Tucuman, on Citrus; 

Martin 2004: 69. 

Distribution: 2: Argentina; Belize. 

Host: Arecaceae: Chamaedorea sp.; Meliaceae: Trichilia sp.; Polygonaceae: Coccoloba 

belizensis; Rutaceae: Citrus sp. 

Paraleyrodes pseudonaranjae Martin 2001 

Paraleyrodes pseudonaranjae Martin 2001: 104. Holotype. Hong Kong: New Territories, Tai 

Lung Farm, Sheung Shui, 22.xi.1999, J. Martin, on Citrus grandis, BMNH. 

Distribution: 1: USA (FL, HI); 2: Puerto Rico; 5: Hong Kong; 9: Hawaii. 

Host: Anacardiaceae: Mangifera indica; Annonaceae: Annona glabra; Chrysobalanaceae: 

Licania michauxii; Ericaceae: Rhododendron pulchrum; Euphorbiaceae: Bridelia 

tomentosa, Glochidion zeylanicum; Fabaceae: Lysiloma latisiliquum; Guttiferae: 

Calophyllum inophyllum; Moraceae: Ficus aurea; Polygonaceae: Coccoloba uvifera; 

Rubiaceae: Randia spinosa, Morinda citrifolia; Rutaceae: Citrus x paradisi, Citrus 

sinensis, Citrus grandis, Citrus maxima. 

Paraleyrodes pulverans Bondar 1923 

Paraleyrodes pulverans Bondar 1923a: 97-98. Syntypes. Brazil: Bahia, G. Bondar, on coconut 



Host: Arecaceae: Cocos nucifera; Meliaceae: Guarea trichiodes. 

Paraleyrodes singularis Bondar 1923 

Paraleyrodes singularis Bondar 1923a: 97. Syntypes. Brazil: Bahia, G. Bondar, on "oitiseiro" 

[Licania tomentosa], MZU; Costa Lima 1928: 134. 


Host: Fabaceae: Inga sp.; Myrtaceae: Myrciaria cauliflora; Rosaceae: Licania tomentosa;

Rutaceae: Citrus sp. 

Paraleyrodes triungulae Martin 2004 

Paraleyrodes triungulae Martin 2004: 69. Holotype puparium, Belize, CFR, Guacamallo Bridge, 

25.ii.1996, J. Martin, on Guettarda combsii, BMNH. 


Host: Rubiaceae: Guettarda combsii. 

Paraleyrodes urichii Quaintance & Baker 1913 

Paraleyrodes urichii Quaintance & Baker 1913: 83-84. Syntypes. Trinidad: received on 

25.v.1911, F. W. Urich, on Pithecolobium sp., USNM; Mound and Halsey 1978: 249. 

Distribution: 2: Barbados; Dominican Republic; Trinidad. 

Host: Fabaceae: Caesalpinia sp., Pithecellobium sp.; Flacourtiaceae: Casearia sp.; Myrtaceae: 

Psidium guajava; Rutaceae: Citrus sp.; Verbenaceae: Citharexylum sp. 

Natural enemies: Aphelinidae: Encarsia dispersa, Encarsia meritoria. 

US Port Interceptions: Mexico on Persea americana; Puerto Rico on Citrus sp.; Dominican 

Republic on Citrus sp.; Haiti on Psidium guajava; Puerto Rico on Aglaonema sp.; 

Trinidad and Tobago on Citrus sp.; US Virgin Islands on Ambrosia hispida 

54

Pseudosynaleurodicus Gillespie 2006 

Pseudosynaleurodicus Gillespie 2006: 2. Type species - Pseudosynaleurodicus nigrimarginatus 

Gillespie, 2006, by original designation. 



mayoi x 

nigrimarginatus x 

Total = 2 species 0 0 0 0 0 0 2 0 0 

Comment: This genus is only known to occur on Proteaceae plants in Australia. It is an unusual 

member of the Aleurodicinae in that it lacks compound pores. 

Reference: Gillespie 2006 - Key to species. 

Pseudosynaleurodicus mayoi Gillespie 2006 

Pseudosynaleurodicus mayoi Gillespie 2006: 12. Holotype puparium, Australia, Western 

Australia, Nowergup, Neeragbup National Park, 27.ix.2002, on Banksia petiolaris, 

ASCU. 

Distribution: 7: Australia. 

Host: Proteaceae: Banksia petiolaris. 

Pseudosynaleurodicus nigrimarginatus Gillespie 2006 

Pseudosynaleurodicus nigrimarginatus Gillespie 2006:2. Holotype puparium, Australia, 

Queensland, 21.vii.2004, P.S. Gillespie, on Macademia ternifolia, QM. 


Host: Proteaceae: Macademia ternifolia. 

55

Stenaleyrodes Takahashi 1938 

Stenaleyrodes Takahashi 1938c: 269. Type-species: Stenaleyrodes vinsoni Takahashi 1938, by 

monotypy. 



papillote x 

vinsoni x x 

Total = 2 species 0 0 0 2 0 0 0 1 0 

Comment: This genus contains 2 species occurring on coconut, Cocos nucifera, both of which 

are known from Afrotropical region and one from New Caledonia. Its placement in the 

Aleurodicinae is debatable since the puparium lacks compound pores and the second pair 

of setae on the lingula, and the forewing of the adult does not have the Rs vein branched. 

Reference: Martin & Streito 2003. 

Stenaleyrodes papillote Martin & Streito 2003 

Stenaleyrodes papillote Martin & Streito 2003: 4-14. Holotype puparium, Mayotte Island, Dani, 

on Cocos nucifera, 9.vii.2003, L. Ollivier; MNHN. 

Distribution: 4: Kenya; Mayotte Island; Tanzania. 


Stenaleyrodes vinsoni Takahashi 1938 

Stenaleyrodes vinsoni Takahashi 1938c: 269-271. Syntypes. Reunion: St. Denis, x.1927, J. 

Vinson, on palm, CORSTOM; Mamet 1952: 134. 

Dialeurodicus elongatus Dumbleton 1956a: 129-131. Syntypes. Reunion: St. Denis, x.1927, J. 

Vinson, on palm, CORSTOM; Cohic 1968c: 13; synonymy according to Mound and 

Halsey 1978: 249. 

Distribution: 4: Reunion Island; 8: New Caledonia. 

Host: Arecaceae: Cocos nucifera, palm. 

Natural enemies: COLEOPTERA/Coccinellidae: Clitostethus arcuatus. 

56

Synaleurodicus Solomon 1935 

Synaleurodicus Solomon 1935: 79. Type-species: Synaleurodicus hakeae Solomon 1935, by 

monotypy. 

Diagnosis - Body greatly elevated; dorsum with 8 pairs of evenly spaced agglomerate pores, 

lingula apparently without setae; without longitudinal molting suture; prominent anterior 

and posterior spiracles; antennae with 5-6 segments. Martin (1999) stated that this genus 

has several apparently unique features, and tentatively placed this genus in the 

Aleurodicinae based on the presence of claws on the puparial legs. 

Comment: This species is known from two Australian species occurring on Hakea (Proteaceae in 

Australia. 



hakeae x 

serratus x 

Total = 2 species 0 0 0 0 0 0 2 0 0 

Synaleurodicus hakeae Solomon 1935 

Synaleurodicus hakeae Solomon 1935: 79-83. Syntypes. Australia: Western Australia, Cottesloe, 

district of Perth, prefering dorsal surface of leaves of Hakea prostrata. Lectotype 

designated by Martin 1999:21, BMNH; Mound and Halsey 1978: 250. 


Host: Proteaceae: Hakea pritzelii?, Hakea prostrata. 

Natural enemies: Pteromalidae: Aphobetus moundi (Boucek); Aphelinidae: Encarsia sp. 

Synaleurodicus serratus Martin 1999 

Synaleurodicus serratus Martin 1999: 22. Holotype puparium, Australia, WA, Perth, 

Forrestfield, on Hakea trifucata, 23.xi.1997, P. J. Gullan, BMNH. 


Host: Proteaceae: Hakea trifucata. 

Synaleurodicus sp. 


Natural enemies: Pteromalidae: Aphobetus moundi. 

Reference: Boucek 1988: 365. 

57

SUBFAMILY ALEYRODINAE WESTWOOD 

Aleyrodinae Westwood 1840: 442. 

Diagnosis - Martin (2005) Puparia with lingula much smaller than in Aleurodicinae, with 2 setae, 

with no discernible setae, or with lingula obscured by operculum; puparial legs without 

claws; never with compound on agglomerate pores on the dorsum. 

Number and Distribution of Genera and Species of Aleyrodinae by Zoogeographic Region 1 

1 Worldwide (WW); Nearctic (NA); Neotropical (NT); Western Palearctic (WPA); Afrotropical (AF); Eastern 

Palearctic (EPA); Oriental (O); Australasian (AU); Pacific Islands (PI); Hawaii (HI); Fossil (F); Int = Introduction. 

Genus WW NA NT WPA AF EPA O AU PI HI F 

Acanthaleyrodes 2 0 0 0 0 1 1 0 0 0 0 

Acanthobemisia 2 0 0 0 0 1 1 0 0 0 0 

Acaudaleyrodes 5 0 0 2 4 0 0 0 0 0 0 

Acutaleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Africaleurodes 16 0 0 0 14 0 2 0 0 0 0 

Agrostaleyrodes 1 0 0 0 0 1 0 0 0 0 0 

Aleurocanthus 80 1 2 4 14 12 40 18 9 2 0 

Aleurocerus 10 1 10 0 0 0 0 0 0 0 0 

Aleurochiton 6+ 1 1 4 0 1 0 0 0 0 1 

Aleuroclava 118 3 Int 2 Int 2+1* 3 34 86 5 2 1 0 

Aleurocybotus 4 2 2 0 1 0 0 0 0 1 0 

Aleurocyperus 1 0 0 0 0 1 0 0 0 0 0 

Aleuroduplidens 6 0 0 0 0 0 0 6 0 0 0 

Aleuroglandulus 4 1 4 0 0 0 0 0 0 0 0 

Aleuroinanis 1 0 0 0 0 0 0 1 0 0 0 

Aleurolobus 83 1 1 7 18 19 54 4 0 0 0 

Aleurolonga 1 0 0 0 1 0 0 0 0 0 0 

Aleuromarginatus 14 0 0 1 3 1 4+ 6 0 0 0 

Aleuropapillatus 2 0 0 0 0 0 2 0 0 0 0 

Aleuroparadoxus 10 4 8 0 0 0 0 0 0 0 0 

Aleuroplatus 75 13 17 8* 29 3 13 2 1 0 0 

58


Aleuropleurocelus 12 10 5 0 0 0 0 0 0 0 0 

Aleuroporosus 1 0 0 0 0 0 1 0 0 0 0 

Aleuropteridis 4 0 0 1* 4 0 0 0 0 0 0 

Aleuroputeus 2+ 0 0 0 0 0 2 1 0 0 0 

Aleurothrixus 20 2 16 1 1 3 3 1 1 1 0 

Aleurotithius 2 1 1 0 0 0 0 0 0 0 0 

Aleurotrachelus 83 2 19 4 21 18 24 3 2 1 0 

Aleurotuba 1 1 1 1 0 0 0 0 0 0 0 

Aleurotulus 6+ 1 5 1 1 0 1 2* 0 0 0 

Aleuroviggianus 6 0 0 6 0 0 0 0 0 0 0 

Aleurovitreus 2+ 0 2+ 0 0 0 0 0 0 0 0 

Aleyrodes 33 10 10 7 2 10 2 1 3 1 0 

Aleyrodiella 1 0 0 1 0 0 0 0 0 0 0 

Anomaleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Apobemisia 2 0 0 0 0 2 0 0 0 0 0 

Arachnaleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Asialeyrodes 15 0 0 0 0 3 12 1 1 0 0 

Asterobemisia 9 0 0 7 0 4 0 0 0 0 0 

Asterochiton 8 0 1 1 0 0 1 0 5 0 0 

Axacalia 1 0 0 1 0 0 0 0 0 0 0 

Bellitudo 4 0 4 0 0 0 0 0 0 0 0 

Bemisaleyrodes 4 0 0 0 4 0 0 0 0 0 0 

Bemisia 46 2 6 14 10 12 16 7 4 2 0 

Bemisiella 2 0 0 1 0 2 0 0 0 0 0 

Brazzaleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Bulgarialeurodes 1 0 0 1 0 0 1 0 0 0 0 

Calluneyrodes 1 0 0 1 0 0 0 0 0 0 0 

Chitonaleyrodes 1 0 0 0 0 0 0 1 0 0 0 

59


Cockerelliella 16 0 0 0 0 2 16 1 0 0 0 

Cohicaleyrodes 16 0 0 0 10 0 4 0 0 0 0 

Combesaleyrodes 2 0 0 0 2 0 0 0 0 0 0 

Corbettia 10 0 0 0 10 0 0 0 0 0 0 

Crenidorsum 21 2 12 1 0 2 3 4 1 1 0 

Crescentaleyrodes 2 0 0 0 0 0 2 0 0 0 0 

Cryptolingula 1 0 0 0 0 0 0 1 0 0 0 

Davidiella 1 0 0 0 0 0 1 0 0 0 0 

Dialeurodes 102 3 15 2 8 16 59 5 5 3 0 

Dialeurolobus 2 0 0 2 0 2 1 0 0 0 0 

Dialeurolonga 53 0 1 1 37 2 11 3 0 0 0 

Dialeuronomada 16 0 0 0 0 0 16 0 0 0 0 

Dialeuropora 20 0 0 1 3 8 11 1 1 0 0 

Dialeurotrachelus 1 0 0 0 0 0 1 0 0 0 0 

Disiphon 2 0 2 0 0 0 0 0 0 0 0 

Distinctaleyrodes 1 0 0 0 0 0 1 0 0 0 0 

Dothioia 1 0 0 0 0 0 0 0 1 0 0 

Dumbletoniella 6 0 0 0 0 0 0 5 1 0 0 

Editaaleyrodes 1 0 0 0 0 0 1 0 0 0 0 

Extensaleyrodes 2 0 0 0 2 0 0 0 0 0 0 

Fascaleyrodes 2 0 0 0 2 0 0 0 0 0 0 

Filicaleyrodes 2+ 1 0 1 2 0 0 0 0 0 0 

Fippataleyrodes 6 0 0 0 0 0 6 0 0 0 0 

Gagudjuia 1 0 0 0 0 0 0 1 0 0 0 

Gomenella 3 0 0 0 0 0 0 1 2 0 0 

Harpaleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Hesperaleyrodes 1 0 1 0 0 0 0 0 0 0 0 

Heteraleyrodes 2 0 0 0 0 0 2 0 0 0 0 

60


Heterobemisia 1 0 0 0 0 1 0 0 0 0 0 

Hindaleyrodes 1 0 0 0 0 0 1 0 0 0 0 

Indoaleyrodes 4+ 0 0 0 0 0 1 3+ 0 0 0 

Juglasaleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Keraleyrodes 1 0 0 0 0 0 1 0 0 0 0 

Laingiella 1 0 0 0 0 0 1 0 0 0 0 

Leucopogonella 1 0 0 0 0 0 0 0 4 0 0 

Lipaleyrodes 9+ 0 0 2* 2 1 5 1+ 0 0 0 

Mohanasundaramiella 1 0 0 0 0 0 1 0 0 0 0 

Malayaleyrodes 1 0 0 0 0 0 1 0 0 0 0 

Marginaleyrodes 5 0 0 0 5 0 0 0 0 0 0 

Massilieurodes 16 5 0 3 7 3 1 0 0 0 0 

Metabemisia 3 0 0 1 0 1 1 0 0 0 0 

Metaleyrodes 1 0 0 0 0 0 0 0 1 0 0 

Minutaleyrodes 5 1 1 0 0 1 4 0 1 1 0 

Mixaleyrodes 2 0 0 0 0 2 0 0 0 0 0 

Nealeyrodes 1 0 1 0 0 0 0 0 0 0 0 

Neoaleurodes 1 0 1 0 0 0 0 0 0 0 0 

Neoaleurotrachelus 4 0 0 0 4 0 0 0 0 0 0 

Neomaskellia 3 0 0 1 1 2 2 1 1 0 0 

Neopealius 1 0 0 1 0 1 0 0 0 0 0 

Nigrasialeyrodes 1 0 0 0 0 0 0 1 0 0 0 

Orchamoplatus 14 0 1 1 0 1 1 4 11 1 0 

Orientaleyrodes 2 0 0 0 0 0 1 0 0 0 0 

61


Orstomaleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Papillipes 1 0 0 0 1 0 0 0 0 0 0 

Parabemisia 7 1 1 1 0 3 3 1 2 1 0 

Paraleurolobus 2 1 1 0 0 0 0 0 0 0 0 

Paulianaleyrodes 3 0 0 0 3 0 0 0 0 0 0 

Pealius 43 4 0 4 2 15 23 3 1 1 0 

Pectinaleyrodes 3 0 0 0 3 0 0 0 0 0 0 

Pentaleyrodes 4 0 0 0 0 4 0 0 0 0 0 

Peracchius 1+ 0 1+ 0 0 0 0 0 0 0 0 

Plataleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Pogonaleyrodes 2 0 0 0 2 0 0 0 0 0 0 

Pseudaleurolobus 2 0 1 0 0 0 1 0 0 0 0 

Pseudaleuroplatus 2 0 0 0 0 0 0 2 0 0 0 

Pseudaleyrodes 1 0 1 0 0 0 0 0 0 0 0 

Pseudozaphanera 5 0 0 0 0 0 0 5 0 0 0 

Rabdostigma 3 0 0 0 0 0 2 1 0 0 0 

Ramsesseus 1 0 0 1 0 0 0 0 0 0 0 

Rhachisphora 23 0 0 1 0 10 10 3 1 0 0 

Rosanovia 1 0 0 1 0 0 0 0 0 0 0 

Rugaleyrodes 5 0 0 0 5 0 0 0 0 0 0 

Rusostigma 4 0 0 0 0 3 3 2 0 0 0 

Russellaleyrodes 1 0 0 0 0 0 1 0 0 0 0 

Rutaleyrodes 1 0 0 0 0 0 1 0 0 0 0 

Septaleurodicus 1 0 1 0 0 0 0 0 0 0 0 

Setaleyrodes 6 0 0 0 0 2 3 1 0 0 0 

Shanthiniae 1 0 0 0 0 0 1 0 0 0 0 

Simplaleurodes 1 0 0 1 0 0 0 0 0 0 0 

Singhiella 31 1 1 2 2 12 17 1 1 1 0 

Singhius 4 1 1 0 0 1 4 0 1 1 0 

62


Siphoninus 3 1 1 1 1 1 0 1 0 1 0 

Sphericaleyrodes 2 0 0 0 0 0 1 0 0 0 0 

Tegmaleurodes 1 0 1 0 0 0 0 0 0 0 0 

Tetraleurodes 64 13 20 5 19 3 11 4 5 0 0 

Tetralicia 6 0 0 3 2 0 0 1 0 0 0 

Trialeurodes 67 31 21 8 5 4 9* 6 2 3 0 

Trialeurolonga 1 0 1 0 0 0 0 0 0 0 0 

Trichoaleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Tuberaleyrodes 5 0 0 0 0 3 2 0 0 0 0 

Vasantharajiella 1 0 0 0 0 0 1 0 0 0 0 

Vasdavidius 5 0 0 1 1 4 2 2 0 0 0 

Venezaleurodes 1 0 1 0 0 0 0 0 0 0 0 

Viennotaleyrodes 11 0 0 0 6 0 2 4 0 0 0 

Xenaleyrodes 6+ 0 0 0 0 0 0 6+ 0 0 0 

Xenobemisia 1 0 0 0 1 0 0 0 0 0 0 

Yleyrodes 1 0 0 0 1 0 0 0 0 0 0 

Zaphanera 4 0 0 0 1 0 3 0 0 0 0 

Total Species 1438 124 212 137 284 256 543 132 71 24 1 

Total Genera 150 29 42 41 54 51 73 43 28 18 1 

63

Acanthaleyodes Takahashi 1931 

Acanthaleyodes Takahashi 1931:220. Type species. Acanthaleyrodes callicarpae Takahashi 

1931, by monotypy. 

References: Takahashi 1931c; 1942b; Corbett 1933. 



callicarpae x 

styraci x 

Total = 2 species 0 0 0 0 1 1 0 0 0 

Acanthaleyrodes callicarpae Takahashi 1931 

Acanthaleyrodes callicarpae Takahashi 1931c: 221. Syntypes Taiwan: Shinten, Taihoku, on 

Callicarpa formosana, Mallotus sp, TARI. 

Distribution: 5: China; Japan; Taiwan. 

Hosts: Euphorbiaceae: Mallotus sp.; Rosaceae: Rubus sp.; Verbenaceae: Callicarpa formosana, 

Callicarpa sp.; Vitaceae: Vitis vinifera. 

Acanthaleyrodes styraci Takahashi 1942 

Acanthaleyrodes styraci Takahashi 1942b: 173-174. Holotype. Thailand, Mt. Sutep, 5.iv.1940, 

on Styrax sp, HBL. 

Distribution: 6: Thailand. 

Host: Styracaceae: Styrax sp. 

Species transferred to other genera 

Acanthaleyrodes spiniferosa (Corbett) 1933 to Tuberaleyrodes (Martin & Mound 2007). 

64

Acanthobemisia Takahashi 1935 

Acanthobemisia Takahashi 1935a: 25. Type-species: Acanthobemisia distyllii Takahashi 1935, 

by monotypy and original designation. 



distylii x 

indicus x 

Total = 2 species 0 0 0 0 1 1 0 0 0 

Acanthobemisia distylii Takahashi 1935 

Acanthobemisia distylii Takashashi 1935a: 25-27. Syntypes. Japan, Nagasaki, x.1934, M. 

Watanabe, collector, on Distylium racemosum, TARI. 

Distribution: 5: Japan. 

Host: Hamamelidaceae: Distylium racemosus/m. 

Acanthobemisia indicus Meganathan & David 1994 

Acanthobemisia indicus Meganathan & David 1994: 11. Holotype. India: on Anaphalis aristata, 


Host: Asteraceae: Anaphalis aristata. 

65

Acaudaleyrodes Takahashi 1951 

Acaudaleyrodes Takahashi 1951: 382. Type-species: Acaudaleyrodes pauliani Takahashi 1951, 

by monotypy. 

Reference: Bink-Moenen (1983) - Description and key to African species. 



africanus x 

ebeni x 

pauliani x 

rachipora x x x 

tuberculata x 

Total = 5 species 0 0 2 4 0 0 0 0 0 

Acaudaleyrodes africanus (Dozier) 1934 

Aleurotrachelus africanus Dozier 1934: 190.Syntypes. Zaire: Barumbu, i.1926, J. Ghesquiere, on 

Desmodium sp., BMNH; Bink-Moenen 1983: 21. 

Distribution: 4: Central African Republic; Chad; Congo; Ivory Coast; Nigeria; Zaire. 

Host: Fabaceae: Bauhinia thonningii, Desmodium sp., Pterocarpus lucens. 

Natural enemies: Aschersonia hypocreoidea. 

Acaudaleyrodes ebeni Manzari & Alemansoor 2005 

Acaudaleyrodes ebeni Manzari & Alemansoor 2005: 1. Holotype, Iran, on Ebenus stellata. 

Distribution: 3: Iran. 

Host: Fabaceae: Ebenus stellata. 

Acaudaleyrodes pauliani Takahashi 1951 

Acaudaleyrodes pauliani Takahashi 1951a: 382-384. Syntypes. Madagascar: Maevatanana, 530 km Majunga 

Road, 31.x.1949, R. Paulian, on "Mpanjakabentany". 

Distribution: 4: Madagascar. 

Host: "Mpanjakabentany". 

Acaudaleyrodes rachipora (Singh) 1931 

Aleurotrachelus rachipora Singh 1931: Syntypes. India: Pusa and Dholi (Bihar), Navsari (Baroda), Miani 

(Punjab, on Cassia fistula, Euphorbia pilulifera, Bauhinia sp., Dalbergia sissoo. 

Acaudaleyrodes rachipora (Singh); Russell 1962b: 64. 

Aleurotrachelus citri Priesner & Hosny 1934. Syntypes. Egypt: Behera, on Citrus spp., Punica granatum and 

other plants; synonymy according to Jesudasan & David 1991: 242. 

Acaudaleyrodes citri (Priesner & Hosny); Russell 1962b: 64. 

Aleurotrachelus alhagi Priesner & Hosny 1934. Syntype. Egypt, Minya, Luxor-Karnak, on Alhagi sp.; 

66

67 

Mound 1965c: 119. 

Distribution: 3: Canary Islands, Cyperus; Egypt; Iran; Iraq; Israel; *Jordan, Saudi Arabia; 4: Cameroon; 

Chad; Kenya; *Liberia, Madagascar; Niger; Nigeria; Sierra Leon; South Africa; Sudan; 6: India. 

Host: Anacardiaceae: Rhus sp.; Annonaceae: Hexalobus monopetalus; *Araceae: Anubias sp.; 

Asclepiadaceae: Leptadenia heterophylla; * Burseraceae: Commiphora sp.; Caprifoliaceae: Sambucus 

nigra; Combretaceae: Combretum micranthum, Terminalia laxiflora; Euphorbiaceae: Euphorbia 

pilulifera, Hymenocardia acida; Ricinus communis; Fabaceae: Abrus precatorius,.Acacia arabica, 

Acacia nilotica, Alhagi sp., Bauhinia sp., Cassia auriculata, Cassia fistula, Cassia italica, Cassia 

sieberiana, Ceratonia siliqua, Cercis siliquastrum, Dalbergia sissoo, Delonix elata, Detarium 

microcarpum, Dichrostachys glomerata, Inga dulce, Lonchocarpus laxiflorus, Parkia 

clappertoniana, Prosopis juliflora, Prosopis africana, Prosopis stephaniana, Pterocarpus lucens, 

Tamarindus indica, Tephrosia apollinea, Tephrosia linearis, Lythraceae: Lawsonia inermis; 

Moraceae: Ficus sp., Morus alba, Morus nigra; Myrtaceae: Psidium guajava; Ochnaceae: Ochna 

afzelii; Punicaceae: Punica granatum; Rhamnaceae: Ziziphus spina-christi; Rutaceae: Citrus 

aurantifolia, Citrus sinensis, Citrus limon; Sapindaceae: Dodonaea viscosa; Tiliaceae: Grewia 

similis; Verbenaceae: Vitex simplicifolia; Zygophyllaceae: Balanites aegyptiaca; Entrada africana. 

Natural enemies: Parasitoids - Aphelinidae: Encarsia acaudaleyrodis, Encarsia davidi, Encarsia galilea, 

Encarsia inaron; Encarsia lutea, Encarsia mineoi, Encarsia sophia, Eretmocerus rajasthanicus; 

Eretmocerus roseni, Predators - ACARI/Phytoseiidae: Euseius scutalis. 

New Records: Intercepted from Jordan, on Citrus sp.; Liberia on Anubias sp. [Araceae] and from Israel on 

Citrus sp., Myrtus sp. and Salix sp.; and from Commiphora sp. [Burseraceae] from South Africa. 

References: Singh 1911; Bink-Moenen 1983: 22; Cohic 1969: 4; Jesudasan and David 1991: 242; Mound 

and Halsey 1978: 8. 

Acaudaleyrodes tuberculata Bink-Moenen 1983 

Acaudaleyrodes tuberculata Bink-Moenen 1983: 24. Holotype pupal case, Chad, Bebedjia, 16.xii.1973, FAB 

& RMBM, on Anogeissus leiocarpus, BMNH. 

Distribution: 3: Chad. 

Host: Combretaceae: Anogeissus leiocarpus; Terminalia laxiflora; Euphorbiaceae: Bridelia ferruginea.

Acutaleyrodes Takahashi 1960 

Acutaleyrodes Takahashi 1960b: 146-147. Type-species: Acutaleyrodes palmae Takahashi 1960, by 

monotypy. 

Diagnosis - Puparium with body narrow, pointed at the posterior end, without dorsal papillae or 

tubercles, submarginal area well defined but not separated from dorsal disk by a suture; marginal 

crenulations minute; thoracic tracheal combs present; caudal furrow absent; vasiform orifice rounded, 

operculum wider than long, occupying half the orifice, lingula obscured. 

Comparison - Acutaleyrodes is similar to Stenaleyrodes, also found on palms, in having a very 

elongate narrow body; however the vasiform orifice and the lingula are obscured by the operculum which is 

rounded in Acutaleyrodes; whereas in Stenaleyrodes the vasiform orifice is triangular and the lingula is not 

obscured. 


Genus NA NT WPA AF EPA O AU PI HI 

palmae 0 0 0 1 0 0 0 0 0 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Acutaleyrodes palmae Takahashi 1960 

Acutaleyrodes palmae Takahashi 1960 in Takahashi 1960b: 146-147. Syntypes. Reunion Island: Saint 

Philippe, xii.1955, R. Paulian, on Acanthophoenix rubra. 

Distribution: 4: Reunion Island. 

Host: Arecaceae: Acanthophoenix rubra. 

68

Africaleurodes Dozier 1934 

Africaleurodes Dozier 1934: 187. Type-species: Africaleurodes coffeacola Dozier 1934, by original 

designation. 

Reference: Bink-Moenen (1983) - Description and key to species. 



adami x 

ananthakrishnani x 

balachowskyi x 

capgrasi x 

citri x 

coffeacola x 

fulakariensis x 

hexalobi x 

indicus x 

karwarensis 

loganiaceae x 

martini x 

ochnaceae x 

pauliani x 

simula x 

souliei x 

tetracerae x 

uvariae x 

vrijdaghii x 

Total = 19 species 0 0 0 14 1 4 0 0 0 

Africaleurodes adami Cohic 1968 

Africaleurodes adami Cohic 1968b: 66-69. Syntypes. Congo: Brazzaville, 7.xi.1966, on Dichapetalum 

brazzae. 

Distribution: 4: Congo. 

Host: Dichapetalaceae: Dichapetalum brazzae. 

Africaleurodes ananthakrishnani Dubey & Sundararaj 

69

70 

Africaleurodes ananthakrishnani Dubey and Sundararaj 2006: 35. Holotype. India: 26.iii.2001, A. K. Dubey, 

FRI. 


Host: Unidentified plant. 

Africaleurodes balachowskyi Cohic 1968 

Africaleurodes balachowskyi Cohic 1968b: 69-72. Syntypes. Congo: Brazzaville, Point-Noire, 3.ix.1966, on 

Chyrsobalanus orbicularis. 


Host: Chrysobalanaceae: Chyrsobalanus orbicularis. 

Africaleurodes capgrasi Cohic 1968 

Africaleurodes capgrasi Cohic 1968b: 72-75. Syntypes. Congo: Brazzaville, 19.ii.1966, P. Capgras, on 

Markhamia sessilis, and 8.ii.1967 on Trichilia heudelottii, and on Hymenocardia ulmoides. 


Host: Bignoniaceae: Markhamia sessilis; Euphorbiaceae: Hymenocardia ulmoides; Meliaceae: Trichilia 

heudelottii. 

Africaleurodes citri (Takahashi) 1932 

Aleurolobus citri Takahashi 1932: 34. Syntypes. Taiwan: on Citrus sp., TARI. 

Africaleurodes citri (Takahashi); Martin & Mound 2007: 56. 

Distribution: 5: Taiwan. 

Host: Malpighiaceae: Hiptage madablota; Rutaceae: Citrus sp. 

US Port Interceptions: India on Citrus sp., Murraya koenigii, Murraya sp.; South Korea on Citrus sp.; 

Sri Lanka on Murraya sp.; Taiwan on Citrus reticulata. 

Africaleurodes coffeacola Dozier 1934 

Africaleurodes coffeacola Dozier 1934: 187-188. Syntypes. Zaire, Lodja, iii.1928, J. Ghesquiere, 

on Coffea robusta. 

Africaleurodes lamottei Cohic 1969: 5-9. Holotype female. Ivory Coast, Lamto, 24.xi.1968, on 

Gardenia ternifolia, and on Combretum lamprocarpum; synonymy according to Bink-Moenen 

1983:27. 

Distribution: 4: Aldabra Island; Cameroon; Chad; Ivory Coast, Nigeria; Sierra Leon; Sudan; Zaire. 

Host: Annonaceae: Annona senegalensis, Hexalobus monopetalus; Combretaceae: Combretum 

cinereipetalum, Combretum lamprocarpum, Combretum micranthum, Combretum molle; 

Euphorbiaceae: Hymenocardia acida, Maesobotrya barteri; Fabaceae: Detarium microcarpum, 

Pterocarpus lucens; Loganiaceae: Strychnos spinosa; Rubiaceae: Chomelia niorescens, Coffea 

robusta, Crossopteryx febrifuga, Gardenia ternifolia, Gardenia triacantha; Mitragyna inermis, 

Nauclea latifolia, Tarenna sp. Plectronia sp.; Rhamnaceae: Ziziphus spina-christi; Sapindaceae: 

Allophylus africanus; Sterculiaceae: Cola sp. 

References: Bink-Moenen 1983; Cohic 1969; Dozier 1934. 

Africaleurodes fulakariensis Cohic 1966 

Africaleurodes fulakariensis Cohic 1966b: 17-20. Syntypes. Congo: Brazzaville, Fulakary, 28.iii.1965, on 

Neosloetiopsis kamarunensis.


Host: Moraceae: Neosloetiopsis kamerunensis. 

Africaleurodes hexalobi Bink-Moenen 1983 

Africaleurodes hexalobi Bink-Moenen 1983: 29. Holotype pupal case, Chad, Bentam, 29.xii.1973, FAB & 

RMBM, on Hexalobus monopetalus, BMNH. 


Host: Annonaceae: Hexalobus monopetalus. 

Africaleurodes indicus Regu and David 1993 

Africaleurodes indicus Regu & David 1993: 12. Holotype. India: on Scheichera trjuga. 


Host: Sapindaceae: Schleichera trijuga. 

Africaleurodes karwarensis Dubey and Sundararaj 

Africaleurodes karwarensis Dubey and Sundararaj 2006: 36. Holotype. India: Karwar, 4.ii.2001, A. K. 

Dubey, on Derris trifoliata, FRI. 


Host: Fabaceae: Derris trifoliata. 

Africaleurodes loganiaceae Dozier 1934 

Africaleurodes loganiaceae Dozier 1934: 189. Syntypes. Zaire: Sankuru, i.1928, J. Ghesquiere, on 

Loganiaceae. 

Distribution: 4: Ivory Coast; Nigeria; Zaire. 

Hosts: Icacinaceae: Icacina mannii; Flacourtiaceae: Scottellia coriaceae; Loganiaceae; Rubiaceae: 

Craterispermum caudatum; Sterculiaceae: Cola caricaefolia. 

References: Cohic 1969; Dozier 1934. 

Africaleurodes martini Cohic 1968 

Africaleurodes martini Cohic 1968b: 76-79. Syntypes. Congo: Brazzaville, 28.x.1966, on Cogniauxia 

podolaena. 

Distribution: 4: Congo; Ivory Coast. 

Host: Cucurbitaceae: Cogniauxia podolaena; Rubiaceae: Morinda morindoides. 

Africaleurodes ochnaceae Dozier 1934 

Africaleurodes ochnaceae Dozier 1934: 188-189. Syntypes. Zaire: Kole, 8.i.1928, J. Ghesquiere, on Ochna 

sp, BMNH/USNM. 

Distribution: 4: Nigeria; Zaire. 

Host: Ochnaceae: Ochna sp.; Sterculiaceae: Cola nitida. 

Africaleurodes pauliani Cohic 1968 

Africaleurodes pauliani Cohic 1968a: 4-7. Syntypes. Congo: Brazzaville, 9.xi.1966, on Markhamia sessilis, 


Host: Bignoniaceae: Markhamia sessilis; Ebenaceae: Diospyros heudelotii; Loganiaceae: Strychnos pungens 

71

Africaleurodes simula (Peal) 1903 

Aleurodes simula Peal 1903b: 81. Syntypes. India: on Bombax malabaricum. 

Aleurolobus simula (Peal); Quaintance & Baker 1914: 109. 

Africaleurodes simula (Peal); change in combination, Martin & Mound 2007. 


Host: Bombacaceae: Bombax malabaricum. 

Africaleurodes souliei Ardaillon & Cohic 1970 

Africaleurodes souliei Ardaillon & Cohic 1970: 272-276. Holotype. Ivory Coast: Banco Forest, 2.iv.1970, 

on Drypetes ayleri. 

Distribution: 4: Ivory Coast. 

Host: Apocynaceae: Pleiocarpa mutica; Convolvulaceae: Neuropeltis prevosteoides; Euphorbiaceae: 

Drypetes aylmeri; Pandaceae: Microdesmis puberula; Sterculiaceae: Cola caricaefolia, Cola 

gabonensis. 

Africaleurodes tetracerae Cohic 1966 

Africaleurodes tetracerae Cohic 1966b: 20-23. Syntypes. Congo: Brazzaville, on Tetracera alnifolia. 


Host: Dilleniaceae: Tetracera alnifolia. 

Africaleurodes uvariae Cohic 1968 

Africaleurodes uvariae Cohic 1968a: 7-10. Syntypes. Congo: Brazzaville, 9.xii.1965, on Uvaria scabrida. 


Host: Annonaceae: Uvaria scabrida. 

Africaleurodes vrijdaghii (Ghesquiere) 1934 

Africaleurodes vrijdaghii (Ghesquiere) 1934. Syntypes. Zaire: on coffee [Coffea sp.] 

Distribution: 4: Zaire. 

Host: Rubiaceae: Coffea sp. 

References: Mayne & Ghesquiere 1934: 30; Russell 1962b: 30. 

72

Agrostaleyrodes Ko 2001 

Agrostaleyrodes Ko 2001: 178. Type species. Agrostaleyrodes arcanus Ko 2001, by monotypy 

. 



arcanus x 

Total = 1 species 0 0 0 0 1 0 0 0 0 

Agrostaleyrodes arcanus Ko 2001 

Agrostaleyrodes arcanus Ko 2001 in Ko et al. 2001: 179. Holotype. Taiwan: Taipei City, on Miscanthus 

sinensis, NTU. 


Host: Poaceae: Miscanthus sinensis, Saccharum officinarum. 

Natural enemies: Aphelinidae: Encarsia sophia; Eretmocerus sp. 

73

Aleurocanthus Quaintance & Baker 1914 

Aleurocanthus Quaintance & Baker 1914: 102. Type-species: Aleurodes spinifera Quaintance 1903, by 

original designation. 

Key reference: Dubey & Sundararaj (2005) - key to Indian species. 



arecae x 

ayyari x 

bambusae x 

bangalorensis x 

banksiae x 

brevispinosus x 

calophyllii x 

ceracroceus x 

cheni x 

chiengmaiensis x 

cinnamomi x 

citriperdus x x x 

clitoriae x 

cocois x x x 

corbetti x 

davidi x 

delottoi x x 

dissimilis x 

esakii x x 

eugeniae x 

euphorbiae x 

ficicola x 

firmianae x 

froggatti x 

74


gateri x 

goaensis x 

gordoniae x 

gymnosporiae x 

hibisci x 

hirsutus x 

husaini x x 

imperialis x 

inceratus x x 

indicus x 

ixorae x 

leptadeniae x 

lobulatus x 

longispinus x x 

loyolae x 

luteus x 

mackenziei x 

mangiferae x x 

martini x 

marudamalaiensis x 

mayumbensis x 

multispinosus x 

musae x 

mvoutiensis x 

niger x 

nigricans x x 

nubilans x 

nudus x 

75


palauensis x 

papuanus x 

pendleburyi x x 

piperis x 

platysepali x 

regis x 

rugosa x x x 

russellae x 

satyanarayana x 

serratus x 

seshadrii x 

shillongensis x 

siasmensis x 

singhi x 

spiniferus x x x x x x x x 

spinithorax x 

spinosus x x 

splendens x 

strychnosicola x 

t-signatus x 

terminaliae x 

trispina x 

valenciae x 

valparaiensis x 

vindhyhachali x 

voeltzkowi x 

woglumi x x x x x x x x x 

zizyphi x x 

Total = 79 species 1 2 4 14 12 39 18 9 2 

76

Aleurocanthus arecae David & Manjunatha 2003 

Aleurocanthus arecae David & Manjunatha 2003: 1. Holotype. India: Areca catechu. 


Host: Arecaceae: Areca catechu, Cocos nucifera. 

Aleurocanthus ayyari Regu & David 1993 

Aleurocanthus ayyari Regu & David 1993a. Holotype puparium, India, Tamil Nadu, Kunnathoor, 14.vi.1989, 

K. Regu, on Calophyllum inophyllum, David Collection. 


Host: Guttiferae: Calophyllum inophyllum. 

Aleurocanthus bambusae (Peal) 1903 

Aleurocanthus bambusae (Peal) 1903b: 85-87. Syntypes. India: on bamboo [?Bambusa sp.]. 


Host: Arecaceae: Phoenix dactylifera; Poaceae: Bambusa sp. 

References: Peal 1903b; Quaintance & Baker 1914: 102; Dubey & Sundararaj 2005c. 

Aleurocanthus bangalorensis Dubey & Sundararaj 2005 

Aleurocanthus bangalorensis Dubey & Sundararaj 2005c: 298. Holotype puparium, India, Karnataka, 

Bangalore, 28.viii.2001, A.K. Dubey, on Syzygium jambos, IFRI. 


Host: Myrtaceae: Syzygium jambos. 

Aleurocanthus banksiae (Maskell) 1895 

Aleurocanthus banksiae (Maskell) 1895: 423-424. Lectotype. Australia: Victoria, Melbourne, C. French, on 

Banksia integifolia or Callistemon linearis. 


Host: Myrtaceae: Callistemon linearis; Proteaceae: Banksia integrifolia. 

References: Maskell 1895; Quaintance & Baker 1914: 102; Martin 1999. 

Aleurocanthus brevispinosus Dumbleton 1961 

Aleurocanthus brevispinosus Dumbleton 1961a: 115-116. Holotype. New Caledonia: Dothio?, F. Cohic, on 

an undetermined host. 

Distribution: 8: New Caledonia. 


Aleurocanthus calophyllii (Kotinsky) 1907 

Aleyrodes calophyllii Kotinsky 1907: 98-99. Syntypes. Fiji: Levuka, 30.x.1899, A. Koebele, on Calophyllum 

inophyllum, BMNH; 

Aleurocanthus calophyllii (Kotinsky); Quaintance & Baker 1914: 102. 

Distribution: 8: Fiji. 


77

78 

Aleurocanthus ceracroceus Martin 1999 

Aleurocanthus ceracroceus Martin 1999: 26. Holotype puparium, Australia, Queensland, costal 

creek-bed 3 km north of Cape Tribulation, on Neolitsea sp, 16.xii.1996, J.H. Martin, ANIC 


Host: Lauraceae: Neolitsea sp. 

Aleurocanthus cheni Young 1942 

Aleurocanthus cheni Young 1942:100-101. Syntypes. China: Szechwan Province, Kiangtsing, Pehpei, on 

Citrus sp. 

Distribution: 5: China. 


Aleurocanthus chiengmaiensis Takahashi 1942 

Aleurocanthus chiengmaiensis Takahashi 1942f: 59-60. Syntypes. Thailand: Samkampeng near Chiengmai, 

4.iv.1940, on bamboo [Bambusa? sp.], TARI. 


Host: Poaceae: Bambusa sp. 

Aleurocanthus cinnamomi Takahashi 1913 

Aleurocanthus cinnamomi Takahashi 1913a: page?. Syntypes. Taiwan: Taihoku, Shinten, Urai, Hokuto, 

Sozan, Shikkun near Taihezan, on Cinnamomum camphora and Machilus spp., TARI; Takahashi 

1940a: 26. 

Aleurodes spinosus Kuwana; misidentification; Maki 1915: 31; Shiraki 1913: 107. 

Distribution: 5: Japan; Taiwan. 

Host: Lauraceae: Actinodaphne sp., Cinnamomum camphora; Machilus sp. 

Natural enemies: Aphelinidae: Encarsia bennetti (acc. Huang & Polaszek 1998: 1848); Encarsia lutea (acc. 

Chou et al. 1996b: 198) 

Aleurocanthus citriperdus Quaintance & Baker 1916 

Aleurocanthus citriperdus Quaintance & Baker 1916: 459-463. Syntypes. Java: Buitenzorg, i.1911, R. S. 

Woglum, on orange [Citrus sp.], USNMH. 

Aleurocanthus cameroni Corbett 1935b: 799-800. Syntypes. Malaya: Batu Gajan, Tapah, Serdang, Kuala 

Lumpur, Kuala Phah, on Citrus acida, Citrus aurantium, Citrus limonum, Citrus hystrix; Mound and 

Halsey 1978: 14. 

Distribution: 5: China; Hong Kong; *Japan; *Taiwan; 6: *Bangladesh, *Cambodia, India; Malaya; Malaysia; 

*Philippines; Sri Lanka; Singapore; *Thailand; Vietnam; 7: *Indonesia; Java; Sulawesi; Sumatra 

Host: Rubiaceae: Coffea sp.; Rutaceae: Citrus acida, Citrus aurantium, Citrus limon, Citrus nobilis, Citrus 

sinensis. 

Natural enemies: Parasitoids - HYMENOPTERA/Aphelinidae: Ablerus inquirenda; Encarsia clypealis, 

ncarsia divergens,Encarsia divergens; Encarsia merceti, Encarsia smithi; Eretmocerus serius S; 

Predators - COLEOPTERA./Coccinellidae: Scymnus pallidivestis, Scymnus smithianus; Nitidulidae: 

Cybocephalus sp.; DIPTERA/ Drosophilidae: Acletoxenus indicus; LEPIDOPTERA/ Pyralididae: 

Cryptoblabes gnidiella; NEUROPTERA/ Chrysopidae. 

US Port Interception Records: Bangladesh, Cambodia, China, Hong Kong, Indonesia, Japan, Malaysia, 

Taiwan, Thailand and Vietnam on Citrus sp; Philippines on an undetermined plant; Vietnam on

Areca sp. and Citrus hystrix. 

Aleurocanthus clitoriae Jesudasan & David 1991 

Aleurocanthus clitoriae Jesudasan & David 1991 245. Holotype pupal case. India: on Clitoria ternatea, 

IDAV. 


Host: Fabaceae: Clitoria ternatea. 


Aleurocanthus cocois Corbett 1927 

Aleurocanthus cocois Corbett 1927: 24-25. Syntypes. Malaya: Batu Gajah, 16.viii.1925, on coconut [Cocos 

nucifera]. 

Aleurocanthus canangae Corbett 1935b: 790-791. Syntypes. Malaya: Kuala, Lumpur, on Cananga odorata, 

BMNH; synonymy according to Martin 1985: 313. 

Distribution: 5: Hong Kong; 6: Burma; Cambodia; India; Malaya; Malaysia; Thailand; 7: Sulawesi. 

Host: Annonaceae: Cananga odorata; Arecaceae: Cocos nucifera; Myrtaceae: Eugenia jambos; Lauraceae: 

Machilus sp.; Rutaceae: Citrus sp. 

Distribution: 5: Hong Kong; 6: Burma; Cambodia; India; Malaya; Malaysia; Thailand; 7: Sulawesi. 

References: Singh 1932: 87; Takahashi 1942f: 57. 

Aleurocanthus corbetti Takahashi 1951 

Aleurocanthus corbetti Takahashi 1951c: 7-8. Syntypes. Malaya: Ula Gombak, Selangor, 11.v.1944, on 

undetermined tree, BMNH. 

Distribution: 6: Malaya; Malaysia. 

Host: unidentified tree. 

Aleurocanthus davidi David & Subramaniam 1976 

Aleurocanthus davidi David & Subramaniam 1976: 147-149. Holotype and 14 paratypes. India: Coimbatore, 

3.v.1957, S. K. David, on Ipomoea sp., ZSI. 


Host: Convolvulaceae: Convolvulus arvensis, Ipomoea palmata, Ipomoae sp. 

Aleurocanthus delottoi Cohic 1969 

Aleurocanthus delottoi Cohic 1969: 11-15. Holotype and 15 paratypes. South Africa: Pretoria, 7.viii.1959, A. 

de Villiers, on Citrus sp., BMNH. 

Distribution: 4: Kenya; South Africa; 5: Hong Kong. 

Host: Rutaceae: Citrus sp.; Ulmaceae: Chaetachme aristata. 

Aleurocanthus dissimilis Quaintance & Baker 1917 

Aleurocanthus dissimilis Quaintance & Baker 1917: 342-343. Syntypes. Burma: Mirdon, xii.1910, R. S. 

Woglum, on unknown vine, USNM. 


Host: unknown vine; Arecaceae: Cocos nucifera. 

Aleurocanthus esakii Takahashi 1936 

79

80 

Aleurocanthus esakii Takahashi 1936f: 111-112. Syntypes. Palau Islands: Koror, Arumizu-Arakasao, 

19.ii.1936, T. Esaski, host unknown, TARI; Takahashi 1956: 9; Martin 1985: 314. 

Distribution: 7: Papua New Guinea; 8: Palau Islands. 

Host: Rosaceae: Parinarium glaberrimum; Sapindaceae: Pometia pinnata; unidentified plant. 

Aleurocanthus eugeniae Takahashi 1933 

Aleurocanthus eugeniae Takahashi 1933: 20-21. Syntypes. Taiwan: Garambi near Koshun, 26.v.1932, R. 

Takahashi, on Syzygium sp. (Eugenia sp.), TARI. 



Aleurocanthus euphorbiae Jesudasan & David 1991 

Aleurocanthus euphorbiae Jesudasan & David 1991: 246. Holotype pupal case. India: on Homonoia riparia, 

IDAV. 


Host: Euphorbiaceae: Homonoia riparia. 

Aleurocanthus ficicola David 1993 

Aleurocanthus ficicola David 1993: 11. Holotype. Sri Lanka, Colombo, 29.vi.1986, B. David, on Ficus sp. 

Distribution: 6: Sri Lanka. 

Host: Moraceae: Ficus sp. 

Aleurocanthus firmianae Dubey & Sundararaj 2005 

Aleurocanthus firmianae Dubey & Sundararaj 2005c: 300. Holotype puparium, India, Karnataka, Shimoga, 

29.i.2001, A. K. Dubey, on Firmiana colorata, IFRI. 

Distribution: 6: India 

Host: Sterculiaceae: Firmiana colorata. 

Aleurocanthus froggatti Martin 1999 

Aleurocanthus froggatti Martin 1999: 27. Holotype puparium, Australia, NSW, Sydney, near Gunnedah, on 

Alectryon oleifolium, 13 ix 1902, W.W. Froggatt, (ANIC), ANIC. 


Host: Sapindaceae: Alectryon oleifolium. 

Aleurocanthus gateri Corbett 1927 

Aleurocanthus gateri Corbett 1927: 24. Syntypes. Malaya: Selangor, v.1923, B. A. R. Gater, on coconut 

[Cocos nucifera], BMNH; Takahashi 1942f: 58. 

Aleurocanthus yusopei Corbett 1935b: 792. Syntypes. Malaya: on Cocos nucifera; .synonymy according to 

Martin & Mound 2007. 


Host: Arecaceae: Cocos nucifera, Elaeis guineensis. 

Aleurocanthus goaensis Dubey & Sundararaj 2005 

Aleurocanthus goaensis Dubey & Sundararaj 2005c: 301. Holotype puparium, India, Goa, Kulem, 28.ii.2001,

A. K. Dubey, on Ochlandra talbotti, IFRI. 


Host: Poaceae: Ochlandra talbotti. 

Aleurocanthus gordoniae Takahashi 1941 

Aleurocanthus gordoniae Takahashi 1941c: 392-393. Syntypes. Hong Kong: 8.iii.1940, R. Takahashi, on 

Gordonia sp., TARI 

Distribution: 5: Hong Kong. 

Host: Theaceae: Gordonia sp. 

Aleurocanthus gymnosporiae Jesudasan & David 1991 

Aleurocanthus gymnosporiae Jesudasan & David 1991: 246. Holotype pupal case. India: on Gymnosporia sp, 

IDAV. 


Host: Celastraceae: Gymnosporia sp. 

Aleurocanthus hibisci Corbett 1935 

Aleurocanthus hibisci Corbett 1935b: 798-799. Syntypes. Malaya: Kuala Lumpur, Sepang, on Hibiscus rosasinensis 

and Hibiscus tiliaceus. 

Distribution: 6: Malaysia; *Singapore. 

Host: *Alismataceae: Sagittaria sagittifolia; Malvaceae: Hibiscus rosa-sinensis; Hibiscus tiliaceus. 

US Port Interception Record: Singapore on Sagittaria sagittifolia. 

Aleurocanthus hirsutus (Maskell) 1895 

Aleurodes hirsuta Maskell 1895: 434. Lectotype. Australia: NSW, Sydney, W. W. Froggatt, on 

Acacia longifolia. [designated by Dumbleton 1956:164], ADSIR. 

Aleurocanthus hirsutus (Maskell) 1895: 434; Quaintance & Baker 1914: 102 


Host: Fabaceae: Acacia longifolia. 

Aleurocanthus husaini Corbett 1939 

Aleurocanthus husaini Corbett 1939: 69-70. Syntypes. India: Kulu, Punjab, on Citrus sp.; Martin 2005b: 

8; Husain & Khan 1945. 

Note: Martin (1985: 8) synonymised this species with Aleurocanthus woglumi Ashby, but later (Martin 

2005b) retained it as a valid species. 



Aleurocanthus imperialis Cohic 1968 

Aleurocanthus imperialis Cohic 1968a: 14-17. Syntypes. Congo: Brazzaville, CORSTOM, 22.xii.1965, on 

Uvaria scubrida. 


Host: Annonaceae: Uvaria scabrida; Rubiaceae: Pauridiantha hirtella. 

81

82 

Aleurocanthus inceratus Silvestri 1927 

Aleurocanthus inceratus Silvestri 1927. Syntypes. Vietnam: Tonkin, Annam, on Citrus sp., IESP; 

Abbasii 1975: 6-10; Takahashi 1936e: 455. 

Distribution: 5: China; 6: India; Vietnam. 

Host: Magnoliaceae: Michelia champaca; Rutaceae: Citrus sp. 

Natural enemies: Aphelinidae: Ablerus macrochaeta; Encarsia clypealis, Encarsia opulenta, Encarsia smithi 

Eretmocerus orientalis, Eretmocerus serius, Predators: COLEOPTERA/ Coccinellidae: Scymnus sp. 

Aleurocanthus indicus David & Regu 1989 

Aleurocanthus indicus David & Regu 1989: 275. Holotype puparium, India, Tamil Nadu, 

Padmanabhapuram, 4.viii.1987, K. Regu, on Borreria hispida, David collection; Jesudasan and David 

1991: 247. 


Host: Rubiaceae: Borreria hispida, Ixora sp. 

Aleurocanthus ixorae Jesudasan & David 1991 

Aleurocanthus ixorae Jesudasan and David 1991: 247. Holotype pupal case. India: on Ixora sp. 


Host: Rubiaceae: Ixora sp. 

Aleurocanthus leptadeniae Cohic 1968 

Aleurocanthus leptadeniae Cohic 1968a: 17-21. Syntypes. Chad: Laboratoire de Farcha, Fort-Lamy, 

26.ii.1966, on Leptadenia hastata; Cohic 1969: 16; Mound and Halsey 1978: 17. 

Distribution: 4: Chad; Sudan. 

Host: Asclepiadaceae: Leptadenia hastata, Leptadenia heterophylla; Capparidaceae: Boscia senegalensis; 

Zygophyllaceae: Balanites aegyptiaca. 

Natural enemies: Aphelinidae: Encarsia tremblayi. 

Aleurocanthus lobulatus Jesudasan & David 1991 

Aleurocanthus lobulatus Jesudasan & David 1991: 247. Holotype pupal case. India: on Dodonaea 

viscosa, IDAV; David & Ragupathy 2004: 26; Dubey & Sundararaj 2005c: 303. 


Host: Moraceae: Morus alba; Rhizophoraceae: Rhizophora mucronata; Sapindaceae: Dodonaea viscosa. 

Aleurocanthus longispinus Quaintance & Baker 1917 

Aleurocanthus longispinus Quaintance & Baker 1917: 344. Syntypes. India: Calcutta, x.1910, R. S. 

Woglum, on bamboo [Bambusa? sp.], USNM; Dubey & Sundararaj 2005c: 303; Martin 1988: 

63; Singh 1931: 66; Takahashi 1942f: 58. 

Aleurocanthus lumpurensis Corbett 1935b: 794-795. Syntypes. Malaya: Kuala Lumpur, on 

Bambusa sp.; synonomy according to Martin & Mound 2007. 

Distribution: 6: Burma; Cambodia; India; Malaya; Malaysia; Thailand; 7: Sarawak; Sulawesi. 

Host: Poaceae: Bambusa sp., Bambusa arundinacea. 

Natural enemies: Aphelinidae: Encarsia divergens; Eretmocerus serius.

83 

Aleurocanthus loyolae David & Subramaniam 1976 

Aleurocanthus loyolae David & Subramaniam 1976: 149-150. Holotype and 27 paratypes. India: 

Burliar, Nilgiris, 13.viii.1971, B. V. David, on Streblus sp., ZSI. 


Host: Moraceae: Streblus asper, Streblus sp. 

Aleurocanthus luteus Martin 1985 

Aleurocanthus luteus Martin 1985: 314. Holotype pupal case. New Guinea: on Macaranga sp, 

BMNH. 

Distribution: 7: New Guinea. 

Host: Euphorbiaceae: Macaranga sp. 

Aleurocanthus mackenziei Cohic 1969 

Aleurocanthus mackenziei Cohic 1969: 16-20. Holotype and 21 paratypes. Kenya: Nairobi, 20.viii. 

1958, G. de Lotto, on Chaetachme aristata, BMNH. 

Distribution: 4: Kenya, Mocambique. 

Host: Rutaceae: Citrus limon; Ulmaceae: Chaetachme aristata. 

Aleurocanthus mangiferae Quaintance & Baker 1917 

Aleurocanthus mangiferae Quaintance & Baker 1917: 345-347. Syntypes. India: Bombay, 1909, 

R. S. Woglumi, on mango [Mangifera indica], USNM; Singh 1931; Takahashi 1932: 68; 


Distribution: 5: Taiwan; 6: India; Pakistan. 

Host: Anacardiaceae: Mangifera indica; Fagaceae: Lithocarpus uraiana. 

Aleurocanthus martini David 1993 

Aleurocanthus martini David 1993:12. Holotype puparium. Sri Lanka, Sigiriya, 2.viii.1986, B. David, 

on Sebastiania chamelia; Dubey & Sundararaj 2003. 


Hosts: Arecaceae: Areca catechu; Combretaceae: Terminalia crenulata; Euphorbiaceae: Macaranga peltata, 

Sebastiania chamaelea; Fabaceae: Tamarindus indica; Flacourtiaceae: Homolium zeylanicum; 

Santalaceae: Santalum album; Sapindaceae: Sapindus laurifolia; Sterculiaceae: Pterospermum 

diversifolium; Tiliaceae: Grewia orbiculata; Verbenaceae: Clerodendron viscosum, Tectona grandis. 

Aleurocanthus marudamalaiensis David & Subramaniam 1976 

Aleurocanthus marudamalaiensis David & Subramaniam1976: 1550-151. Holotype and 7 paratypes. India: 

Coimbatore, 22.x.1966, B. V. David, on Ficus bengalensis, ZSI; Dubey & Sundararaj 2005c: 304 


Host: Moraceae: Ficus bengalensis, Ficus benjamina; Tiliaceae: Grewia orbiculata. 

Aleurocanthus mayumbensis Cohic 1966 

Aleurocanthus mayumbensis Cohic 1966a: 23-26. Syntypes. Congo: Brazzaville, Fourastie, 2.ii.1964, on 

Entandrophragma candolle, BMNH; Bink-Moenen 1983: 35. 

Distribution: 4: Congo; Ivory Coast.

84 

Hosts: Dilleniaceae: Tetracera alnifolia; Euphorbiaceae: Protomegabaria stapfiana; Loganiaceae: Strychnos 

pungens; Meliaceae: Entandrophragma candollei, Trichilia heudelottii. 

Aleurocanthus multispinosus Dumbleton 1961 

Aleurocanthus multispinosus Dumbleton 1961a: 116-118. Holotype and paratypes. New 

Caledonia: Montagne des Sources, 3,000 ft, L. J. Dumbleton, on Hibbertia sp., ADSIR 


Host: Dilleniaceae: Hibbertia sp. 

Aleurocanthus musae David & Jesudasan 2002 

Aleurocanthus musae David & Jesudasan 2002: 323. Holotype puparium, Andaman & Nicobar Islands, 

Wandoor, 25.iv.2001, B. David, on Musa sp., IARI. 

Distribution: 6: Andaman and Nicobar Islands. 

Host: Musaceae: Musa sp. 

Aleurocanthus mvoutiensis Cohic 1966 

Aleurocanthus mvoutiensis Cohic 1966a: 26-28. Syntypes. Congo: Brazzaville, 30.v.1964, M'Vouti, BMNH; 


Distribution: 4: Chad; Congo; Ivory Coast. 

Host: Euphorbiaceae: Hymenocardia acida. 

Aleurocanthus niger Corbett 1926 

Aleurocanthus niger Corbett 1926: 276. Syntypes. Sri Lanka: Pundaluoya, E. E. Green, on bamboo 

[Bambusa? sp.], BMNH 



Aleurocanthus nigricans Corbett 1926 

Aleurocanthus nigricans Corbett 1926: 277. Syntypes. Sri Lanka: Pundaluoya, E. E. Green, on Bambusa sp.; 

Martin 1988: 63 

Distribution: 6: Sri Lanka; 7: Sulawesi. 


Aleurocanthus nudus Dumbleton 1961 

Aleurocanthus nudus Dumbleton 1961a: 118. Holotype and paratypes. New Caledonia: Tinip, 4.xi.1954, L. J. 

Dumbleton, on upper surface of Leucopogon sp. leaves, CORSTOM. 


Host: Epacridaceae: Leucopogon sp. 

Aleurocanthus palauensis Kuwana 1931 

Aleurocanthus palauensis Kuwana 1931: 659. Syntypes. Palau Islands: G. Yoshino, on an undetermined 

plant, TARIT; Takahashi 1956: 10. 

Distribution: 8: Palau Islands; Caroline Islands. 

Host: Arecaceae: Cocos nucifera.

Aleurocanthus papuanus Martin 1985 

Aleurocanthus papuanus Martin 1985: 315. Holotype pupal case. New Guinea: on Xanthophyllum 

papuanum, BMNH. 


Host: Rutaceae: Zanthophyllum papuanum. 

Aleurocanthus pendleburyi Corbett 1935 

Aleurocanthus pendleburyi Corbett 1935b: 795-796. Syntypes. Malaya: Pahang, on Syzygium (=Eugenia) sp, 

lost?; Martin 1985: 64. 

Distribution: 6: Malaya; 7: Papau New Guinea; Sulawesi. 

Host: Burseraceae: Canarium sp.; Lauraceae: Persea americana; Myrtaceae: Syzygium sp. (Eugenia sp.); 

Piperaceae species 

Aleurocanthus piperis (Maskell) 1895 

Aleurodes piperis Maskell 1895: 438-439. Syntypes: Sri Lanka: Pundaluoya, E. E. Green, on Piper nigrum?, 

ADSIR. 

Aleurocanthus piperis (Maskell); Quaintance & Baker 1914: 102. 


Host: Piperaceae: Piper nigrum, Piper sp. 

Comment: (nomen novum, need to check if it is a valid species (see David and Manjunatha). 

Aleurocanthus platysepali (Cohic) 1966 

Aleurocanthus platysepali Cohic 1966b: 87. Syntypes. Congo: Brazzaville, 17.vii.1964, on Aframomum 

stipulatum; on Platysepalum vanderystii. 


Host: Fabaceae: Millettia laurentii. 

Aleurocanthus regis Mound 1965 

Aleurocanthus regis Mound 1965c: 124-125. Holotype and 33 paratypes. Nigeria: Ibadan, Moor Plantation, 

i.1961, L. A. Mound, on Combretum sp., BMNH. 

Distribution: 4: Nigeria. 

Host: Combretaceae: Combretum sp. 

Aleurocanthus rugosa Singh 1931 

Aleurocanthus rugosa Singh 1931: 71-73. Syntypes. India: Pusa, on Eugenia jambolana, Piper betle, 

Psidium guajava, Mechelia champaca, ZSI; Takahashi 1952c: 27; David & Subramaniam 1976: 151; 

Martin 1988: 64. 

Distribution: 3: Iran; 6: India; Malaya; Malaysia; 7: Sulawesi. 

Host: Annonaceae: Annona sp., Polyalthia longifolia, Polyalthia pendula; Magnoliaceae: Magnolia 

grandifolia; Michelia champaca; Myrtaceae: Eugenia jambolana, Myrtus communis, Psidium 

guajava; Piperaceae: Piper betle; Rhamnaceae: Ziziphus sp.; Sapindaceae: Dodonaea viscoa. 

US Port Interceptions: Bangladesh and India on Piper betle. 

Aleurocanthus russellae Jesudasan & David 1991 

85

86 

Aleurocanthus russellae Jesudasan & David 1991: 250. Holotype pupal case. India: on Cinnamomum sp., 

IDAV 


Host: Lauraceae: Cinnamomum sp. 

Aleurocanthus satyanarayani Dubey & Sundararaj 2005 

Aleurocanthus satyanarayani Dubey & Sundararaj 2005c: 305. Holotype puparium, India, 

Karnataka, South Kannada, 12. ix.2001, A. K. Dubey, on Tabernmontana heyneana, IFRI. 


Host: Apocynaceae: Tabernaemontana heyneana. 

Aleurocanthus serratus Quaintance & Baker 1917 

Aleurocanthus serratus Quaintance & Baker 1917: 349-351. Syntypes. Java: Buitenzorg, Botanical Gardens, 

I 1911, R. S. Woglumi, on unidentified tree, USNM. 

Distribution: 7: Java. 


Aleurocanthus seshadrii David & Subramaniam 1976 

Aleurocanthus seshadrii David & Subramaniam 1976: 152-153. Holotype and 3 paratypes. India: Mallisera 

Estate near Kongode (Kerala State), 27.ix.1968, B. V. David, on bamboo [Bambusa? sp.], BMNH; 

Dubey & Sundararaj 2005c: 306. 


Host: Poaceae: Bambusa arundinacea, Bambusa sp. 

Aleurocanthus shillongensis Jesudasan & David 1991 

Aleurocanthus shillongensis Jesudasan & David 1991: 251. Holotype pupal case. India: on Quercus sp., 

IDAV 


Host: Fagaceae: Quercus sp. 

Aleurocanthus siasmensis Takahashi 1942 

Aleurocanthus siamensis Takahashi 1942f: 60-61. Syntypes. Thailand: Ubon, 4.v.1940, on undetermined 

tree, HBL. 



Aleurocanthus singhi Jesudasan & David 1991 

Aleurocanthus singhi Jesudasan & David 1991: 251. Holotype pupal case. India: on Phoenix dactylifera, 

IDAV 


Host: Arecaceae: Phoenix dactylifera. 

Note: This species needs to be redescribed since the original description and drawings are not in concurrence. 

Aleurocanthus spiniferus (Quaintance) 1903 

Aleurodes spinifera Quaintance 1903: 63-64. Syntypes. Java: Garolt, 7.xii.1901, C. L. Marlatt, on Citrus sp..

87 

and Rosa sp. (Lectotype designated by Martin 1999: 29), USNM. 

Aleurocanthus spiniferus (Quaintance); Quaintance & Baker 1914: 102; Bink-Moenen 1983: 37. 

Aleurocanthus spiniferus var. intermedia Silvestri 1927: 2. Syntypes. China: Canton, on undetermined plant, 

IESP; synonymy according to Mound and Halsey 1978: 22 

Aleurocanthus citricolus (Newstead) 191: 173. Syntypes. Tanzania: Dar es Salaam, iv.1902, A. Zimmerman, 

on Citrus sp, BHU; synonymy according to Quaintance & Baker 1914: 102. 

Aleurocanthus rosae Singh 1931: 70-71. Syntypes. India: Poona, Pondicherry, on Rosa sp., ZSI; synonymy 

according to Takahashi 1932: 47. 

Distribution: 2: Jamaica; 3: Greece, Iran; 4: Kenya; Mauritius; Tanzania; 5: China; Hong Kong; Japan; 

Philippines; Taiwan; 6: India; Sri Lanka; Malaysia; Thailand; 7: Java; New Guinea; Sumatra; 8: 

Caroline Islands; New Caledonia; 9: Hawaii. 

Host: Annonaceae: Annona reticulata, Annona squamosa; Apocynaceae: Plumeria rubra; Arecaceae: Cocos 

nucifera; Convolvulaceae: Erycibe acutifoliae; Ebenaceae: Diospyros khaki; Elaeocarpaceae: Sloanea 

dasycarpa; Euphorbiaceae: Sapium sebiferum; Fabaceae: Bauhinia championii; Vigna sesquipedalis; 

Flacourtiaceae: Casearia aculeata, Myroxylon japonicum; Hamamelidaceae: Liquidambar 

formosana; Lardizabalaceae: Akebia labata; Lauraceae: Persea zuihoensis; Malvaceae: Hibiscus 

canabinus, Hibiscus sp.; Moraceae: Ficus capensis; Rosaceae: Eriobotrya japonica, Prunus serotina, 

Rosa indica, Rosa sinensis; Rutaceae: Citrus bigaradia, Citrus limon, Citrus sinensis, Citrus sinensis, 

Zanthoxylum nitida, Sabiaceae: Meliosma rigida; Salicaceae: Salix sp.; Vitaceae: Vitis vinifera. 

Natural enemies: Fungus - Aschersonia sp. 

Parasitoids - Aphelinidae Ablerus connectans, Encarsia divergens, Encarsia ishii, Encarsia merceti modesta, 

Encarsia nipponica, Encarsia opulenta, Encarsia smithi, Eretmocerus serius, Eretmocerus silvestrii; 

Platygastridae: Amitus sp. Platygastridae. 

Predators: COLEOPTERA/Coccinellidae: Axinoscymnus beneficus, Catana clauseni, Chilocorus kuwanae, 

Cryptognatha flaviceps, Cryptognatha sp., Cryptolaemus montrouzieri, Delphastus sp., 

Microserangium shikokense, Scymnus near pallidicollis, Scymnus sp., Scymnus (Pullus) hilaris, 

Serangium flavescens, Serangium maculigerum; DIPTERA/Drosophidae: Acletoxenus indicus, 

Acletoxenus sp; NEUROPTERA/Chrysopidae: Chrysopa fulvolineata; Chrysopa sp.; Hemerobiidae: 

Micromus timidus. 

US Port Interceptions: China on Camellia sp. and Pyracantha sp.; Dominican Republic on Citrus sp.; 

Haiti on Citrus sinensis; Hawaii on Citrus reticulata, Citrus sinensis, Citrus sp. and Rosa sp.; 

India on Citrus sp. and Murraya koenigii; Nigeria on Citrus sp.; Philippines on Parthenium sp. and 

Piper sp.; Syrian Arab Republic on Rosa sp.; Taiwan on Pyracantha sp.; Viet Nam on Rosa sp.. 

Aleurocanthus spinithorax Dumbleton 1961 

Aleurocanthus spinithorax Dumbleton 1961a: 118-120. Holotype and paratypes. New Caledonia: Dothio, F. 

Cohic, on undetermined climbing plant, CORSTOM. 


Host: undetermined climbing plant. 

Aleurocanthus spinosus (Kuwana) 1911 

Aleyrodes spinosus Kuwana 1911: 626. Syntypes. Taiwan: 1909, Kuwana, on unidentified plant, TARI. 

Aleurocanthus spinosus (Kuwana); Quaintance & Baker 1914: 102; Capco 1959: 40; Fulmek 1943: 5; 

Takahashi 1931a: 207; Takahashi 1933: 4. 

Distribution: 5: China; Taiwan; 6: Malaya; Malaysia, Philippines.

88 

Host: Annonaceae: Annona sp.; Flacourtiaceae: Scolopia oldhami; Piperaceae: Piper futokadsura; Rubiaceae: 

Gardenia florida; Rutaceae: Citrus sp. 

Natural enemies: Parasitoids - Aphelinidae: Encarsia clypealis ; Predators - COLOPTERA/Coccinellidae: 

Scymnus sp. 

US Port Interceptions: China on Citrus sp; Hawaii on Citrus sinensis; Malaysia on Citrus sp.; Thailand on 

an unknown plant. 

Aleurocanthus splendens David & Subramaniam 1976 

Aleurocanthus splendens David & Subramaniam 1976: 154. Holotype and 9 paratypes. India: Perammur, 

Tiruchirapalli district, Tamil Nadu, 24.iv.1969, B. V. David, on Phoenix humilis, ZSI. 


Host: Arecaceae: Phoenix humilis. 

Aleurocanthus strychnosicola Cohic 1966 

Aleurocanthus strychnosicola Cohic 1966a: 15-17. Syntypes. Congo: Brazzaville, 12.ii.1946, on Strychnos 

pungens; Cohic 1968a: 22. 


Host: Loganiaceae: Strychnos pungens, Strychnos spinosa; Ochnaceae: Ochna gilletiana 

Aleurocanthus t-signatus (Maskell) 1895 

Aleurodes t-signatus Maskell 1895: 443-444. Lectotype. Australia: NSW, Botany near Sydney, W. W. 

Froggatt, on Acacia longifolia, ADSIR. 

Aleurocanthus t-signatus (Maskell); Quaintance & Baker 1917: 353; Dumbleton 1956b: 167; Carver & Reid 

1996. 


Host: Fabaceae: Acacia longifolia, Fabaceae: Acacia sp. 

Natural enemies: Predators - DIPTERA/ Drosophilidae: Acletoxenus formosus; Syrphidae: Eophaerophoria 

sp. 

Aleurocanthus terminaliae Dubey & Sundararaj 2005 

Aleurocanthus terminaliae Dubey & Sundararaj 2005c: 307. Holotype puparium, India, Karnataka, Yellapur, 

11.x.2001, A. K. Dubey, on Terminalia crenulata, IFRI. 


Host: Combretaceae: Anogeissus latifolia, Calycopteris floribunda, Terminalia crenulata; Fabaceae: 

Adenanthera pavonia; Flacourtiaceae: Hydnocarpus petandra; Loganiaceae: Strychnos potatorum; 

Lythraceae: Lagerstroemia microcarpa; Opiliaceae: Cansjera rheedii; Rubiaceae: Catunaregam 

spina; Solanaceae: Lycianthes laevis. 

Aleurocanthus trispina Mound 1965 

Aleurocanthus trispina Mound 1965c: 125-126. Holotype and 5 paratypes. Nigeria: Ibadan, Moor Plantation, 

16.xi.1959, E.A. James, on Combretum micranthum, BMNH; Cohic 1969: 22 

Distribution: 4: Ivory Coast; Nigeria. 

Host: Combretaceae: Combretum micranthum; Meliaceae: Trichilia prieureana; Sapindaceae: Paullinia 

pinnata.

89 

Aleurocanthus valenciae Martin 1999 

Aleurocanthus valenciae Martin 1999: 30. Holotype puparium, Australia, Qld, Mundubbera, on Citrus 

sinensis "Valencia", 21.iii.1996, D. Papacek, NHM. 


Host: Rutaceae: Citrus sinensis. 

Aleurocanthus valparaiensis David & Subramaniam 1976 

Aleurocanthus valparaiensis David & Subramaniam 1976: 154. Holotype and 38 paratypes. India: Valparai, 

6000 ft, 16.iv.1967, B. V. David, on Piper nigrum, ZSI. 


Host: Piperaceae: Piper nigrum. 

Aleurocanthus vindhyhachali Dubey & Sundararaj 2005 

Aleurocanthus vindhyachali Dubey & Sundararaj 2005c: 308. Holotype puparium, India,Goa, Kulem, 

28.ii.2001, A. K. Dubey, on Kandelia candela, IFRI. 


Host: Rhizophoraceae: Kandelia candela. 

Aleurocanthus voeltzkowi (Newstead) 1908 

Aleurodes voeltzkowi Newstead 1908: 12-13. Syntypes. Madagascar: Sainte Marie, viii.1904, on unidentified 

plant, BHU. 

Aleurocanthus voeltzkowi (Newstead); Quaintance & Baker 1914: 102. 



Aleurocanthus woglumi Ashby 1915 

Aleurocanthus woglumi Ashby 1915: 321. Syntypes. Jamaica: on Citrus spp., USNM; Martin 1988: 64. 

Aleurocanthus nagpurensis Raina et al. 

Distribution: Virtually worldwide; 1: USA ( AZ, CA, FL, MS, TX); 2: Mexico, Guatemala, Cuba, Honduras, 

Nicaragua, El Salvador, Panama, Costa Rica, Venezuela, Colombia, Brazil, Argentina, Paraguay, 

Peru Uruguay, Jamaica, Puerto Rico, Virgin Islands, Bahamas, Bermuda; 3: Egypt, England, France, 

Iran, Israel, Italy, Spain, USSR, 4: Uganda; Kenya; South Africa; Seychelles; Tanzania; 5: China, 6: 

India, Philippines, Malaysia, Vietnam, Taiwan Thailand, 7: Australia, Indonesia, 8: New Zealand; 9: 

Hawaii. 

Host: Anacardiaceae Anacardium occidentale; Mangifera indica; Annonaceae Annona cheirimola; Annona 

squamosa; Apocynaceae: Plumeria acutifolia, Tabernaemontana coronaria; Araliaceae: Hedera 

helix; Arecaceae: Elaeis melanococca; Asteraceae: Baccharis sp.; Begoniaceae: Begonia sp.; 

Bignoniaceae Crescentia cujete; Boraginaceae: Cordia alba, Tournefortia hirsutissima; 

Capparidaceae: Capparis pedunculosus, Capparis roxburghii; Caricaceae: Carica papaya; 

Celastraceae: Gymnosporia diversifolia, Kurrimia zeylanica; Euphorbiaceae: Croton sp.; Fabaceae: 

Adinobotrys atropurpureus; Flacourtiaceae: Scolopia oldhami; Hippocrateaceae: Salacia reticulata; 

Lauraceae: Laurus nobilis, Persea americana, Persea gratissima; Loranthaceae: Loranthus sp.; 

Lythraceae: Lagerstroemia indica; Malpighiaceae: Malpighia glabra, Malpighia punicifolia; 

Malvaceae: Hibiscus rosa-sinensis, Hibiscus schizopetalus, Sida cordifolia; Meliaceae: Trichilia 

auranticola, Meliaceae: Trichilia spondiodes; Moraceae: Morus sp.; Musaceae: Musa sapientum,

90 

Musa x paradisiaca; Myrsinaceae: Wallenia laurifolia; Myrtaceae: Syzygium (Eugenia) jambos, 

Syzygium (Eugenia) malaccensis, Syzygium (Eugenia) uniflora, Psidium guajava; Orchidaceae: 

Vanda sp.; Passifloraceae: Passiflora edulis; Polygonaceae: Antigonon leptopus; Punicaceae: Punica 

granatum; Rosaceae: Pyracantha formosana, Pyrus sp.; Rubiaceae: Coffea arabica, Ixora thwaitesii, 

Morinda tinctoria, Citrus acida; Citrus aurantifolia, Citrus aurantifolia, Citrus aurantium, Citrus 

limon, Citrus maxima, Citrus medica, Citrus meyeri, Citrus nobilis, Citrus reticulata, Citrus sinensis, 

Citrus x paradisi, Citrus x tangelo, Clausena lansium, Fortunella crassifolia, Lunasia amara, 

Murraya exotica, Murraya koenigii, Severinia buxifolia, X(Family?) Citrofortunella microcarpa; 

Sapotaceae: Achras sapota, Pouteria mammosa; Solanaceae: Cestrum diurnum, Cestrum nocturnum; 

Sterculiaceae: Guazuma tomentosa; Ulmaceae: Celtis formosana. 

Natural enemies: 

Fungus: Aschersonia aleyrodis. 

Parasitoids: Aphelinidae: Ablerus connectans, Ablerus inquirenda, Ablerus macrochaeta, Cales noacki, 

Encarsia basicincta, Encarsia bennetti, Encarsia clypealis, Encarsia divergens, Encarsia elegans, 

Encarsia ishii, Encarsia merceti, Encarsia narayanani, Encarsia opulenta, Encarsia perplexa, 

Encarsia smithi, Encarsia sp., Encarsia tinctoriae, Encarsia variegata, Eretmocerus breviclavus, 

Eretmocerus serius, Eretmocerus sp.; Encyrtidae: Pseudhomalopoda prima; Platygastridae: Amitus 

hesperidum; Signiphoridae Signiphora sp. 

Predators: ARANEAE/ Araneidae: Gasteracantha cancriformis, Linyphiidae: Linyphia triangularis; 

COLEOPTERA/ Coccinellidae: Azya luteipes, Azya trinitatis, Brumoides suturalis, Catana clauseni, 

Catana parcesetosum, Chilocorus cacti, Chilocorus stigma, Cryptognatha flaviceps, Cryptognatha 

nodiceps, Cryptognatha sp., Cryptolaemus montrouzieri, Cycloneda sanguinea, Decadiomus 

bahamicus, Delphastus argentinicus, Delphastus catalinae, Delphastus diversipes, Delphastus 

nebulosus, Delphastus pallidus, Delphastus pusillus, Delphastus sp., Diomus sp, Hyperaspis 

albicollis, Hyperaspis calderana, Hyperaspis horni, Microserangium shikokense, Nephaspis oculatus, 

Pentilia castanea, Pentilia castanea, Pseudoscymnus quinquepunctata, Scymnus adspersulus, 

Scymnus gorhami, Scymnus gracilis, Scymnus levaillanti Scymnus near pallidicollis, Scymnus 

pallidicollis, Scymnus smithianus, Scymnus thoracius, Scymnus thoracius, Scymnus (Pullus) 

coloratus, Scymnus (Pullus) horni, Serangium parcesetosum, Stethorus sp., Zilus aeneus, Zilus 

cyanescens; Cybocephalidae: Cybocephalus sp.; Staphylinidae: Holoborus pallidicornis; DIPTERA/ 

Cecidomyidae: Lestodiplosis sp.; Dolichopodidae: Condylostylus chrysoprasi, Drosophilidae: 

Acletoxenus formosus, Acletoxenus indicus, Acletoxenus sp; HEMIPTERA/ Anthocoridae: 

Cardiastethus rugicollis; Miridae: Lygus hesperus; Reduviidae: Empicoris rubromaculatus; 

HYMENOPTERA/ Formicidae: Cremastogaster brevispinosa; LEPIDOPTERA/ Pyralididae: 

Cryptoblabes gnidiella; NEUROPTERA/ Chrysopidae Ceraeochrysa sp., Chrysopa sp., Chrysoperla 

carnea, Chrysoperla externa, Malada boninensis, Mallada signata, Nodita sp.; Coniopterygidae: 

Semidalis sp.; Hemerobiidae; THYSANOPTERA, Phlaeothripidae: Aleurodothrips fasciapennis; 

Thripidae: Scolothrips latipennis. 

US Port Interceptions: Antigua and Barbuda on Capsicum sp.; Bangladesh on Citrus sp; Murraya koenigii; 

Cambodia on Murraya koenigii; China on Citrus sp.; Costa Rica on Citrus sp.; Cuba on Citrus 

aurantium; Dominican Republic on Capsicum sp., Citrus aurantium, Citrus sinensis, Citrus sp., 

Prunus cerasus; Ecuador on Citrus sp., Citrus limon, Citrus sinensis; Egypt on Murraya koenigii; 

Fiji on Murraya koenigii; Guatemala on Citrus sp.; Guyana on Citrus sp.; Haiti on Citrus sinensis, 

Citrus sp.; Hawaii on Citrus sinensis, Citrus sp., Piper sp. Plumeria sp.; Honduras on Citrus sp.; 

India on Celosia sp, Citrus sinensis, Citrus sp., Mangifera indica, Murraya koenigii, Murraya

91 

paniculata; Indonesia on Citrus sp.; Italy on Citrus sp.; Jamaica on Citrus sp.; Japan on Citrus sp.; 

Laos on Citrus sp.; Malaysia on Citrus hystrix, Citrus limon, Citrus sp. Murraya koenigii; Mexico 

on Celtis sp., Citrus aurantiifolia, Citrus aurantium, Citrus latifolia, Citrus limettioides, Citrus limon, 

Citrus paradisi, Citrus reticulata, Citrus sinensis, Citrus sp., Laurus nobilis, Mangifera indica, 

Murraya sp., Nardo sp., Persea americana, Plant, Psidium guajava, Psidium sartorianum, Rosa sp. 

Netherlands on Anethum sp.; Nicaragua on Citrus sp.; Pakistan: Murraya koenigii and Murraya 

sp.; Philippines on Citrus sp., Psidium guajava; Puerto Rico on Annona muricata,, Areca sp., 

Citrullus sp., Citrus aurantiifolia, Citrus aurantium, Citrus limon, Citrus paradisi, Citrus sinensis, 

Citrus sp., Cocos nucifera, Ixora sp, Mangifera indica, Pimenta racemosa; Russia on Murraya sp; 

Singapore on Citrus sp., Murraya koenigii and Murraya sp; Spain on Citrus sp.; Sri Lanka on 

Murraya koenigii; St. Lucia on Citrus sp.; Taiwan on Bambusa sp., Psidium guajava; Thailand: 

Citrus limon, Citrus sp., Murraya koenigi.; Trinidad and Tobago on Citrus sp.; US Virgin Islands 

on Citrus auraintiifolia, Citrus sp.; United Arab Emirates on Citrus limon; Viet Nam on Adenium 

sp., Citrus sp., Ochna integerrima, Ochna sp. 

Aleurocanthus zizyphi Priesner & Hosny 1934 

Aleurocanthus zizyphi Priesner & Hosny 1934a: 4. Syntypes. Egypt: Dakhla Oasis, 20.iii.1932, on Ziziphus 

spina christi, EDAC; Bink-Moenen 1983: 38. 

Aleurocanthus aberrans Cohic 1968a: 10-14. Syntypes. Chad: on Gardenia jovis-tonantis, on Cassia siamea; 

Paratypes labeled: Congo: Brazzaville, Centre ORSTOM, 17.iv.1964, on Papilionaceae, BMNH. 

Aleurocanthus hansfordi Corbett 1935c: 240-242. Syntypes. Uganda: Bukalasa, 14.x.1930, C. H. Hansford, 

on Paulinia pinnata, BMNH; Abd-Rabou 1996 

Distribution: 3: Egypt; Iran, Jordan; 4: Chad; Congo; Ivory Coast; Kenya; Nigeria; South Africa; Uganda. 

Host: Asclepiadaceae: Leptadenia sp.; Combretaceae: Combretum paniculatum, Terminalia laxiflora; 

Euphorbiaceae: Alchornea cordifolia, Phyllanthus mullerianus; Fabaceae: Cassia siamea, Dalbergia 

sp., Detarium microcarpum; Flacourtiaceae: Dovyalis caffra, Oncoba spinosa; Moraceae: Ficus 

carica; Myrtaceae: Psidium guajava.; Ochnaceae: Ochna afzelii; Rhamnaceae: Ziziphus spina-christi; 

Rosaceae: Rubus fruticosus; Sapindaceae: Paullinia pinnata; Sapotaceae: Sideroxylon inerme; 


Natural enemies: Parasitoids - Aphelinidae: Encarsia lutea; Eretmocerus breviclavus; Eretmocerus 

diversiciliatus, Eretmocerus nairobii, Eretmocerus nairobii.

Aleurocerus Bondar 1923 

Aleurocerus Bondar 1923a: 156. Type species. Aleurocerus luxuriosus Bondar 1923, by original designation. 

Uraleyodes Sampson & Drews 1941: 180. Type species. Uraleyrodes ceriferus Sampson & Drews 1941, by 

original designation; synonymy according to Mound & Halsey 1978: 27 



ceriferus x 

chiclensis x 

coccolobae x 

colombiae x 

flavomarginatus x 

luxuriosus x 

musae x 

palmae x x 

petiolicola x 

tumidosus x 

Total = 10 species 1 10 0 0 0 0 0 0 0 

Key references: Russell 1986: 151; Martin 2005b: 9. 

Aleurocerus ceriferus (Sampson & Drew) 1941 

Uraleyrodes ceriferus Sampson & Drew 1941: 180. Pupal cases. Mexico: Nayarit, xi.1925, Ferris, on woody 

vine. 

Distribution: 2: Belize, Costa Rica, Honduras, Mexico, Panama. 

Host: Euphorbiaceae: Drypetes sp.; Moraceae: Brosimum sp.; Piperaceae: Piper sp; Vitaceae: Cissus sp.; 

unidentified woody vine. 

Aleurocerus chiclensis Russell 1986 

Aleurocerus chiclensis Russell 1986:154. Holotype. Mexico: intercepted in Brownsville, Texas (USA), 

30.vii.1935, G. A. Pfaffman, on Manikara zapota [chicle], USNM. 

Distribution: 2: Belize, Mexico. 

Host: Burseraceae: Protium sp.; Myrtaceae: Eugenia? sp.; Sapotaceae: Manilkara zapota. 

Aleurocerus coccolobae Russell 1986 

Aleurocerus coccolobae Russell 1986:157. Holotype. Colombia: 18.vii.1935, R. Arbelaez, on Coccoloba 

padiformis, USNM. 

Distribution: 2: Colombia. 

Host: Polygonaceae: Coccoloba padiformis. 

92

Aleurocerus colombiae Russell 1986 

Aleurocerus colombiae Russell 1986: 159. Holotype. Colombia: near Anchicaya, 19.iii.1967, Dieffenbachia? 

sp, USNM. 


Host: Araceae: Dieffenbachia? sp. 

Aleurocerus flavomarginatus Bondar 1923 

Aleurocerus flavomarginatus Bondar 1923a: 161. Syntypes. Brazil: on unidentified plant. 

Distribution: 2: Brazil; Cuba; Guatemala. 

Host: Arecaceae: Chamaedorea sp.; Rubiaceae: Gardenia augusta. 

Aleurocerus musae Russell 1986 

Aleurocerus musae Russell 1986:169. Holotype. Guatemala: Guatemala City (intercepted in San Antonio, 

Texas), on Musa paradisiaca, USNM. 

Distribution: 2: Guatemala. 

Host: Musaceae: Musa paradisiaca. 

US Port Interceptions: Guatemala on Heliconia sp.; Honduras on Musa sp.; Mexico on Alpinia purpurata, 

Zingiber officinale and Zingiber sp. 

Aleurocerus palmae Russell 1986 

Aleurocerus palmae Russell 1986: 172. Holotype. Mexico: intercepted in Dallas, Texas, 28.vi.1966, on 

Chamaedorea elegans, USNM; California PPDR2005: 56. 

Distribution: 1: USA (FL); 2: Belize, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, 

Panama, Peru. 

Host: Arecaceae: Chamaedorea elegans, Chamaedorea erumpens; Desmoncus sp.; Sapotaceae: Pouteria sp. 

US Port Interceptions: Belize on Orchidaceae; Costa Rica on Phoenix roebelenii; Guatemala on Areca sp. 

and Chamaedorea sp.; Mexico on Alpinia purpurata,Chamaecyparis sp., Chamaedorea elegans, 

Chamaedorea sp, Chenopodium sp., Murraya sp. and Chrysalidocarpus sp. 

Aleurocerus petiolicola Russell 1986 

Aleurocerus petiolicola Russell 1986: 176. Holotype. Costa Rica: Las Cruces, on Stromanthe lutea 

[=Calathea lutea], USNM. 

Distribution: 2: Costa Rica; Panama. 

Host: Maranthaceae: Stromanthe lutea. 

Aleurocerus tumidosus Bondar 1923 

Aleurocerus tumidosus Bondar 1923a: 159. Syntypes. Brazil: Bahia, G. Bondar, on "cipo cabcolo" [Davilla 

rugosa]; Russell 1986. 


Host: Dilleniaceae: Davilla rugosa. 

93

Aleurochiton Tullgren 1907 

Aleurochiton Tullgren 1907: 14. Type species: Chermes aceris ovatus Geoffroy, 1762, by designation. 

Aleurochiton (Nealeurochiton) Sampson 1943: 201. Type species. Aleurodes forbesii Ashmead, by 

monotypy; synonymy according to Mound and Halsey 1978: 27. 

Nealeurochiton Zahradnik 1963a: 8. 

Aleurochiton (Nigrinichiton) Zahradnik 1987: 70; synonymy according to Martin & Mound 2007: 9. 

Aleurochiton (Nigrinichiton) Zahradnik; synonymy according to Martin & Mound 2007. 

Host - All of the described species are only known from maple (Acer), except for an undescribed species 

in USA: Florida known from holly (Ilex). 


Species NA NT WPA AF EPA O AU PI HI fossil 

acerinus x 

aceris x 

forbesii x 

orientalis x 

petri x x 

pseudoplatani x 

sp. x 

Total =6+ species 1 1 4 0 1 0 0 0 0 1 

Aleurochiton acerinus Haupt 1934 

Aleurochiton acerina Haupt 1934: 137. Syntypes. On Acer campestre. 

Aleurochiton acerinus Haupt 1934; Mound and Halsey 1978: 28; Martin 1985b: 143; Rapisarda 1982; 

Minelli et al. 1995: 12. 

Distribution: 3: Austria; Czech Republic; England; Germany; Hungary; Iran, Poland; USSR; Yugoslavia. 

Host: Aceraceae: Acer campestre, Acer insigna. 

US Port Interceptions: Germany on Acer sp.; Hungary on Acer campestre, Acer platanoides; United 

Kingdom on Acer insigna, Acer saccharinum and Acer sp. 

Aleurochiton aceris (Modeer) 1778 

Chermes aceris ovatus Geoffroy 1762. Syntypes. Sweden: on Acer platanoides. [rejected trinomial. 

Coccus aceris Modeer 1778: 21. Syntypes. Sweden: on Acer platanoides 

Chermes aceris Geoffroy 1762; homonym of Chermes aceris L. 1758 [Psyllidae]; Danzig 1966: 198. 

Aleurodes aceris Bouche 1851: 109. Syntypes: Germany, on Acer platanoides; synonymy according to 

Zahradnik 1955:44. 

Aleyrodes acerum Kirkaldy 1907: 44. Replacement name for Chermes aceris ovatus Geoffroy 1762; 

synonymized with A. complanatus by Mound 1966: 407. 

Aleurochiton aceris (Modeer); Danzig 1966:198; Zahradnik 1963a: 12; Hulden 1986: 9; Minelli et al. 1995: 

12. 

94

95 

Lecanium complanatum Baerensprung 1849: 169. Syntypes. Germany: Berlin, on Acer platanoides; 

synonymy according to Danzig 1966: 367. 

Aleurochiton complanatus (Baerensprung); Schumacher 1918: 404. 

Distribution: 3: Austria; Czech Republic; Denmark; England; Finland; France; Germany; Hungary; 

Iran, Italy; Netherlands; Poland; Russia; Sweden; USSR; Yugoslavia. 

Host: Aceraceae: Acer cappadocicum, Acer platanoides, Acer tataricum. 

Natural enemies: Aphelinidae: Encarsia aleurochitonis, Encarsia margaritiventris, Encarsia tricolor; 

Eulophidae: Euderomphale secreta; Platygastridae: Amitus aleurodinis, Amitus minervae. 

Aleurochiton forbesii (Ashmead) 1893 

Aleurodes aceris Forbes 1885. Syntypes. USA: Illinois, on Acer sp.; new name nec. 

Aleurodes forbesii Ashmead 1893: 294. Replacement name for Aleurodes aceris Forbes nec. Aleurodes 

aceris (Barensprung). 

Aleurochiton (Nealeurochiton) forbesii (Ashmead); Sampson 1943: 201. 

Aleurochiton forbesii (Ashmead); Mound and Halsey 1978: 28. 

Nealeurochiton forbesii (Ashmead); Zahradnik 1963a: 12. 

Aleurochiton forbesii (Ashmead); combination reviewed by Mound and Halsey 1978: 28. 

Distribution: 1: Canada; USA (DE; FL; GA; IL; MO; NC; NY; VA; WI). 

Host: Aceraceae: Acer dasycarpum, Acer platanoides, Acer rubrum, Acer saccharinum., 

Aceraceae: Acer sp.; Aquifoliaceae: Ilex coriacea, Aquifoliaceae: Ilex glabra. 

Natural enemies: Platygastridae: Amitus aleurodinis Haldeman 1850. 

Aleurochiton orientalis Danzig 1966 

Aleurochiton orientalis Danzig 1966: 367. Holotype. USSR: Maritime Territory, on Acer mono, ZIN 

Distribution: 5: USSR. 

Host: Aceraceae: Acer mono. 

Aleurochiton petri Rietschel 1983 

Aleurochiton petri Rietschel 1983:97-100. Holotype pupal case, Germany, Kiesgrube, fossil, 

Distribution: 3: Germany (fossil). 

Host: fossil. 

Aleurochiton pseudoplatani Visnya 1936 

Aleurochiton pseudoplatani Visnya 1936: 116. Syntypes. Hungary: Koszeg, on Acer pseudoplatanus, BTM 

Nealeurochiton pseudoplatani (Visnya); Zahradnik 1963a: 12. 

Aleurochiton pseudoplatani Visnya ; Danzig 1966: 198 [366]; Mound and Halsey 1978: 29; Martin 2000: 

527 

Distribution: 3: Austria; Czech Republic; Germany; Hungary; Iran, Lebanon; Netherlands; USSR. 

Host: Aceraceae: Acer tataricum, Acer turcomanica, Acer tauricolu; Celasteraceae: Euonymus japonicus 

US Port Interceptions: China on Citrus sp.; France on Acer negundo and Acer sp. 

Aleurochiton sp. 

Aleurochiton sp. Fulmek 1943: 5. - 2: Guyana parasitized by Aphelinidae: Encarsiella magniclava. 

Aleurochiton sp. Girault 1915c: 278. 2: Guyana parasitized by Eulophidae: Entedononecremnus unicus.

Aleuroclava Singh 1931 

Aleuroclava Singh 1931: 90. Type species: Aleuroclava complex Singh 1931, by monotypy. 

Aleuromigda Singh 1931. Nomen nudum, no type species designated. 

Aleurotuberculatus Takahashi 1932: 20. Type species. Aleurotuberculatus gordoniae Takahashi 1932, by 

original designation; synonymy according to Martin 1999: 31. 

Japaneyrodes Zahradnik 1962a: 13. Type species. Aleurotuberculatus trachelospermi Takahashi 1938, by 

original designation; synonymy according to Mound and Halsey 1978: 78. 

Hindaleyrodes Meganathan & David 1994:37. Type species. Hinaleyrodes hindustanicus, by monotypy; 

synonymy according to Martin & Mound 2007: 9. 

Martiniella Jesudasan and David 1990: 7. Type species. Aleurotuberculatus canagae Corbett 1935, by 

original designation; synonymy according to Martin 1999:31; Manzari & Quicke 2006: 2470. 

Taiwanaleyrodes Takahashi 1932: 28. Type species. Taiwanaleyrodes meliosmae Takahashi 1932, by 

monotypy; synonymy according to Manzari & Quicke 2006: 2470. 

Reference: Dubey & Sundararaj 2006a (Key to Indian species); Dubey & Sundararaj 2005c: 241-272. Key to 

Aleuroclava species; Sundararaj & Dubey 2004- Key to Martiniella species. 

Note: Martin 1999: 31 synonymized Martiniella Jesudasan & David 1990 with Aleuroclava, stating that the 

characters used to separate it from Aleuroclava - the very much enlarged, jointed cephalic and first 

abdominal setae were also present in species of Taiwanaleyrodes and Dialeurodes, and that this 

character has been seen to vary among samples. Sundararaj & Dubey 2004: 358 considered 

Martiniella to be a valid genus based upon the its differentiated type of setae. 



afriae x 

angkorensis x 

artocarpi x 

aucubae Int x 

ayyari x 

baccaureae x 

bauhiniae x 

bifurcata x 

bilineata x 

bulbiformi x 

burmanicus x 

calicutensis x 

96


calycopteriseae x 

canangae x 

carpini x 

cinnamomi x 

citri x 

citrifolii x 

complex x 

cordii x 

davidi x 

doddabettaensis x 

doveri x 

dubius x 

elatostemae x 

erythrinae x 

eugeniae x 

euphoriae x 

euryae x 

evanantiae x 

ficicola x 

filamentosa x 

flabellus x 

fletcheri x 

fici x 

goaensis x 

gordoniae x 

grewiae x 

97


guyavae x 

hexcantha x 

hikosanensis x 

hindustanicus x 

jasmini Int Int x x x x x x 

indica x 

kanyakumariensis 

kavalurensis x 

kerala x 

kudremukhensis x 

kuwanai x 

lagerstroemiae x 

lanceolata x 

latus x 

lefroyi x 

lithocarpi x 

longisetosus x 

longispinus x 

louiseae x 

macarangae x 

madhucae x 

magnoliae x 

malloti x 

manii x 

martini x 

maximus x 

98


melastomae x 

meliosmae x 

montanus x 

multipori x x x 

multituberculata x 

murrayae x x 

mysorensis x 

nachiensis x 

nagercoilensis x 

nanjangudensis x 

neolitseae x x x x 

nephelii x 

nigeriae x 

nitidus x x 

orientalis x 

papillata x 

parvus x 

pentatuberculata x 

philomenae x 

phyllanthi x 

piperis x 

pongamiae x x 

porosus x 

psidii x x 

pulcherrima x 

pyracanthi x 

ramachandrani x 

99


regui x 

rhododendri x 

russellae x 

saputarensis x 

selvakumarani x 

sepangensis x 

srilankaensis x 

siamensis x 

similis Int Int Int x 

simplex x 

singhi x 

sivakasiensis x 

stereospermi x 

subindica x 

submarginatus x 

takahashii x x 

tarennae x 

tentactuliformis x 

terminaliae x 

thysanospermi x 

trachelospemi x 

trilineata x 

tripori x 

trivandricus x 

trochodendri x 

ubonensis x 

uraianus x 

100

vernoniae x 


vitexae x 

wrightiae x 

Total = 108 species 3 Int 2 Int 3+ 1Int 3 33 87 5 2 1 

Aleuroclava afriae Sundararaj & David 1995 

Aleuroclava afriae Sundararaj & David 1995: 17. Holotype puparium, India, Rajasthan, Jodhpur, 17.iii.1993, 

R. Sundararaj, on Cordia myxa, Sundararaj coll. (paratypes in USNM, BMNH, IARI). 


Host: Boraginaceae: Cordia myxa. 

Aleuroclava angkorensis (Takahashi) 1942 

Aleurotuberculatus angkorensis Takahashi 1942h: 331. Syntypes. Cambodia: Angkor, 24.iv.1940, on 

undetermined host, HBL. 

Aleurotuberculatus ankorensis [sic] Takahashi; Mound & Halsey 1978: 79. 

Aleuroclava ankorensis (Takahashi); Martin 1999: 31. 

Distribution: 6: Cambodia. 

Host: undetermined host. 

Aleuroclava artocarpi (Corbett) 1935 

Aleurotuberculatus artocarpi Corbett 1935b: 834. Syntypes. Malaya: on Artocarpus sp. 

Aleuroclava artocarpi (Corbett); Martin 1999: 31. 


Host: Moraceae: Artocarpus sp. 

Aleuroclava aucubae (Kuwana) 1911 

Aleyrodes aucubae Kuwana 1911: 625. Syntypes. Japan: on Aucuba japonica, USNM. 

Tetraleurodes aucubae (Kuwana); Quaintance & Baker 1914: 108. 

Aleurotuberculatus aucubae (Kuwana); Takahashi 1932: 20. 

Aleuroclava aucubae (Kuwana); Martin 1999: 31. 

Distribution: 1: USA (CA); 5: China; Japan. 

Host: Aquifoliaceae: Ilex crenata; Araliaceae: Hedera japonica; Caprifoliaceae: Lonicera gracillipes; 

Cornaceae: Aucuba japonica; Flacourtiaceae: Myroxylon japonicum; Juglandaceae: Juglans 

sieboldiana; Lauraceae: Litsea glauca; Moraceae: Ficus erecta, Morus alba; Oleaceae: Ligustrum 

japonicum; Pittosporaceae: Pittosporum tobira 'variegata'; Rosaceae: Kerria japonica; Rosaceae: 

Neillia uekii, Prunus mume, Rhaphiolepis umbellata; Rubiaceae: Paederia sp.; Rutaceae: Citrus sp., 

Phellodendron lavallei, Zanthoxylum piperitum; Theaceae: Eurya japonica; Ulmaceae: Aphananthe 

aspera. 

Natural enemies: Parasitoids - Aphelinidae: Encarsia lutea (ref: Huang & Polaszek 1998: 1913); 

Encarsia sp. (ref: Kuwana 1928: 61). 

Other references: Martin 1999: 31; California PPDR2005: 15; Takahashi & 1952 1952a: 19. 

101

Aleuroclava ayyari (Sundararaj & David) 1993 

Martiniella ayyari Sundararaj & David 1993a: 95. Holotype puparium, India, Tamil Nadu, 

Padmanabhapuram, 4.viii.1987, R. Sundararaj, on Mussaeenda sp., David collection (paratypes in 

USNM, BMNH, IARI). 

Aleuroclava ayyari (Sundararaj & David) 1993; Martin 1999. 


Host: Rubiaceae: Mussaenda sp. 

Aleuroclava baccaureae (Corbett) 1935 

Taiwanaleyrodes baccaureae Corbett 1935b: 838. Syntypes. Malaya: on Baccaurea motleyana. 

Aleuroclava baccaureae (Corbett); Manzari & Quicke 2006: 2470. 


Host: Euphorbiaceae: Baccaurea motleyana. 

Aleuroclava bauhiniae (Corbett) 1935 

Aleurotuberculatus bauhiniae Corbett 1935b: 835. Syntypes. Malaya: on Bauhinia bidentata. 

Aleuroclava bauhiniae (Corbett); Martin 1999: 31. 


Host: Fabaceae: Bauhinia bidentata. 

Aleuroclava bifurcata (Corbett) 1933 

Dialeurodes bifurcata Corbett 1933: 126. Syntypes. Malaya: on Nephelium lappaceum. 

Aleurotuberculatus bifurcata (Corbett); Mound and Halsey 1978: 80. 

Aleuroclava bifurcata (Corbett); Martin 1999: 31. 


Host: Sapindaceae: Nephelium mutabile, Nephelium lappaceum. 

Aleuroclava bilineata Sundararaj & David 1993 

Aleuroclava bilineata Sundararaj & David 1993: 237. Holotype pupal case, India: Tamil Nadu, Kunnathoor, 

25 II 1988, R. Sundararaj, on Cissus sp., IDAV. 

Additional references: Dubey & Sundararaj 2005d: 246. 


Host: Annonaceae: Miliusa tomentosa; Combretaceae: Calycopteris floribunda; Dipterocarpaceae: Vateria 

indica; Lauraceae: Cinnamomum malabratrum; Moraceae: Artocarpus hirsutus; Vitaceae: Cissus sp. 

Aleuroclava burmanicus (Singh) 1938 

Aleurotuberculatus burmanicus Singh 1938: 189. Syntypes. Burma: on Cassia sp., ZSI 

Aleuroclava burmanicus (Singh); Martin 1999: 31. 

Distribution: 6: Burma; Malaya. 

Host: Fabaceae: Cassia sp.; Lauraceae: Cinnamomum sp.; Liliaceae: Smilax sp. 

102

103 

Aleuroclava calicutensis Dubey & Sundararaj 2005 

Aleuroclava calicutensis Dubey & Sundararaj 2005d: 246. Holotype puparium, India, Kerala, Calicut, 

19.iii.2001, A. K. Dubey, on unidentified plant; IFRI. 



Aleuroclava calycopteriseae Dubey & Sundararaj 2005 

Aleuroclava calycopteriseae Dubey & Sundararaj 2005d. Holotype puparium, India, Karnataka, Kudremukh, 

12.vii.2001, A. K. Dubey, on Calycopteris floribunda, IFRI. 


Host: Combretaceae: Calycopteris floribunda. 

Aleuroclava canangae (Corbett) 1935 

Aleurotuberculatus canangae Corbett 1935b: 827. Syntypes. Malaya: on Cananga odorata and Psidium 

guajava, BMNH. 

Aleuroclava canangae (Corbett); Martin 1999: 31. 

Martiniella canangae (Corbett); Jesudasan and David 1990. 


Host: Annonaceae: Cananga odorata; Myrtaceae: Psidium guajava. 

Aleuroclava carpini (Takahashi) 1939 

Taiwanaleyrodes carpini Takahashi 1939a: 78. Syntypes. Taiwan: on Carpinus sp., TARI. 

Taiwanaleyrodes carpini var. yushuniae Takahashi 1939a: 80. Syntypes. Taiwan: on Yushunia randaiensis, 

TARI. 

Aleuroclava carpini (Takahashi); Manzari & Quicke 2006: 2470. 


Host: Betulaceae: Carpinus sp.; Lauraceae: Sassafras randaiensis. 

Natural enemies: Aphelinidae: Encarsia bothrocera, Encarsia perflava. 

Aleuroclava cinnamomi Jesudasan & David 1991 

Aleuroclava cinnamomi Jesudasan and David 1991: 255-256. Holotype. India: Tamil Nadu, Diruvani, 

4.viii.1984, R. W. Alexander Jesudasan, on Cinnamomum malabaricum, TARI. 


Host Lauraceae: Cinnamomum malabaricum. 

Aleuroclava citri Jesudasan & David 1991 

Aleuroclava citri Jesudasan and David 1991: 256-257. Holotype. India: Tamil Nadu, Madras, 8.xii.1972, B. 

V. David, on Citrus sp., IDAV. 


Host: Rutaceae: Citrus sp.

Aleuroclava citrifolii (Corbett) 1935 

Aleurolobus citrifolii Corbett 1935b: 9. Syntypes. Pakistan: on Citrus sp, NHM.; Husain & Khan 1945: 18. 

Aleurotuberculatus citrifolii (Corbett); Mound and Halsey 1978: 81. 

Aleuroclava citrifolii (Corbett); Martin 1999. 

Distribution: 6: India; Pakistan. 

Host: Rutaceae: Citrus sp., Murraya exotica. 

Aleuroclava complex Singh 1931 

Aleuroclava complex Singh 1931: 91. Syntypes. India: on Diospyros montana and Pongamia pinnata 

(=Pongamia glabra), IARI.; Dubey & Sundararaj 2005d. 


Host: Annonaceae: Polyalthia cerasoides; Boraginaceae: Cordia obliqua; Chenopodiaceae: Madhuca 

[=Bassia] latifolia; Ebenaceae: Diospyros montana; Fabaceae: Pongamia pinnata [=glabra]; 

Flacourtiaceae: Hydnocarpus pentandra, Flacourtia montana; Icacinaceae: Nothapodytes 

nimmoniana; Lauraceae: Smilax zeylanica; Loganiaceae: Strychnos nox-vomica; Lythraceae: 

Lawsonia inermis; Malvaceae: Thespesia populnea; Moraceae: Ficus religiosa, Streblus asper; 

Phyllanthaceae: Aporosa lindleyana; Rhamnaceae: Ziziphus rugosa; Rutaceae: Aegle marmelos; 

Verbenaceae: Vitex altissima; Clerodendrum viscosum; Zingiberaceae: Alpinia sp. 

Aleuroclava davidi Qureshi 1982 

Aleuroclava davidi Qureshi 1982. Holotype puparium, Pakistan. 

Distribution: 6: Pakistan. 

Aleuroclava doddabettaensis Dubey & Sundararaj 2005 

Aleuroclava doddabettaensis Dubey & Sundararaj 2005d. Holotype puparium, India, Tamil Nadu, 

Doddabetta, 10.iii.2001, A. K. Dubey, on unidentified bush; IFRI. 


Host: unidentified bush. 

Aleuroclava doveri (Corbett) 1935 

Dialeurodes doveri Corbett 1935b: 774. Syntypes. Malaya: on Musa sapientum. 

Aleuroclava doveri (Corbett); Martin & Mound 2007. 


Host: Musaceae: Musa sapientum. 

Aleuroclava dubius (Bink-Moenen) 1983 

Aleurotuberculatus dubius Bink-Moenen 1983: 81. Holotype pupal case, Chad, Galaba, 2.viii.1973, FAB & 

RMBM, on Terminalia laxiflora; BMNH. 

Aleuroclava dubia (Bink-Moenen); Martin 1999:31. 


Host: Combretaceae: Terminalia laxiflora; Euphorbiaceae: Hymenocardia acida; Rubiaceae: Crossopteryx 

febrifuga, Gardenia triacantha, Mitragyna inermis, Nauclea latifolia. 

104

Aleuroclava elatostemae (Takahashi) 1932 

Aleurotuberculatus elatostemae Takahashi 1932: 23. Holotype. Taiwan: on Elatostema sp., TARI. 

Aleuroclava elatostemae (Takahashi); Martin 1999: 31. 


Host: Urticaceae: Elatostema lineolatum. 

Aleuroclava erythrinae (Corbett) 1935 

Aleurotuberculatus erythrinae Corbett 1935b: 836. Syntypes. Malaya: on Erythrina sp. ? 

Aleuroclava erythrinae (Corbett); Martin 1999: 31. 


Host: Fabaceae: Erythrina sp. 

Aleuroclava eugeniae (Corbett) 1935 

Aleurotuberculatus eugeniae Corbett 1935b: 836. Syntypes. Malaya: on Eugenia aquera 

Aleuroclava eugeniae (Corbett); Martin 1999: 31. 


Host: Myrtaceae: Eugenia aquera. 

Aleuroclava euphoriae (Takahashi) 1942 

Aleurotuberculatus euphoriae Takahashi 1942h: 333. Syntypes. Thailand: on Euphoria longana, TARI. 

Aleuroclava euphoriae (Takahashi); Martin 1999: 31. 


Host: Sapotaceae: Euphoria longana. 

Aleuroclava euryae (Kuwana) 1911 

Aleyrodes euryae Kuwana 1911: 625. Syntypes. Japan: on Eurya ochnacea, USNM. 

Aleuroplatus euryae (Kuwana); Quaintance & Baker 1914: 98. 

Aleurotuberculatus euryae (Kuwana); Takahashi 1938b: 71. 

Aleuroclava euryae (Kuwana); Martin 1999: 31. 


Host: Theaceae: Eurya ochnacea. 

Aleuroclava evanantiae Jesudasan & David 1991 

Aleuroclava evanantiae Jesudasan and David 1991: 259. Holotype. India: Tamil Nadu, Tambaram, 4.xi.1984, 

R. W. Alexander Jesudasan, on Evanantia (=Tabernacoronata) coronaria; Dubey & Sundararaj 

2005d. 


Host: Apocynaceae: Tabernaemontana coronaria; Combretaceae: Calycopteris floribunda, Terminalia 

paniculata; Euphorbiaceae: Croton caudatus; Flacourtiaceae: Flacourtia montana. 

Icacinaceae: Nothapodytes nimmoniana; Rutaceae: Naringi crenulata. 

Aleuroclava fici (Corbett) 1935 

105

106 

Taiwanaleyrodes fici Corbett 1935b: 837. Syntypes. Malaya: on Ficus sp. and Euphorbia pulcherrima. 

Aleuroclava fici (Corbett); Manzari & Quicke 2006: 2470. 


Host: Euphorbiaceae: Euphorbia pulcherrima; Moraceae: Ficus sp. 

Aleuroclava ficicola (Takahashi) 1932 

Aleurotuberculatus ficicola Takahashi 1932: 24. Syntypes. Taiwan: on Ficus sp., TARI. 

Aleuroclava ficicola (Takahashi); Martin 1999: 31. 


Host: Moraceae: Ficus sp.; Myrsinaceae: Maesa japonica; Urticaceae: Boehmeria zollingeriana. 

Natural enemies: Aphelinidae: Encarsia bothrocera, Encarsia cibcensis, Encarsia echinocera, Encarsia 

lutea, Encarsia perflava. 

Aleuroclava filamentosa (Corbett) 1933 

Dialeurodes filamentosa Corbett 1933: 126. Syntypes. Malaya: on Baccaurea motleyana. 

Aleurotuberculatus filamentosa (Corbett); Mound and Halsey 1978: 82. 

Aleuroclava filamentosa (Corbett); Martin 1999: 31. 



Aleuroclava flabellus (Takahashi) 1949 

Aleurotuberculatus flabellus Takahashi 1949: 49. Syntypes. Riouw Islands (south of Singapore): on an 

undetermined tree, HBL. 

Aleuroclava flabellus (Takahashi); Martin 1999: 31. 

Distribution: 8: Riouw (Riau) Islands. 

Host: on undetermined tree. 

Aleuroclava fletcheri (Sundararaj & David) 1992 

Taiwanaleyrodes fletcheri Sundararaj & David 1992a: 18. Holotype puparium, India, Tamil 

Nadu, 3.viii.1987, B. V. David, on Mallotus sp., USNM (NHM, IARI, ZSI). 

Aleuroclava fletcheri (Sundararaj & David); Manzari & Quicke 2006: 2470. 


Host: Euphorbiaceae: Mallotus sp. 

Aleuroclava goaensis Jesudasan & David 1991 

Aleuroclava goaensis Jesudasan and David 1991: 259-260. Holotype. India: Goa, 4.viii.1985, D. Sukumaran, 

on unidentified tree; Dubey & Sundararaj 2005d. 


Host: Anacardiaceae: Holigarna grahamii, Holigarna arnottiana; Araceae: Pothos scandens. 

Dipterocarpaceae: Nothopegia heyneana, Nothopegia racemosa; Euphorbiaceae: Macaranga peltata. 

Aleuroclava gordoniae (Takahashi) 1932

107 

Aleurotuberculatus gordoniae Takahashi 1932: 21. Syntypes. Taiwan: on Photinia [=Pourthiaea] 

bethamiana, Gordonia anomala and other plants, TARI. 

Aleuroclava gordoniae (Takahashi); Martin 1999: 31. 

Distribution: 5: Hong Kong; Taiwan. 

Host: Hamamelidaceae: Liquidambar formosana; Lauraceae: Cinnamomum camphora. 

Rosaceae: Photinia [=Pourthiaea] bethamiana; Theaceae: Gordonia anomala, Gordonia sp. 

Natural enemies: Aphelinidae: Encarsia lutea. 

Aleuroclava grewiae Sundararaj & David 1993 

Aleuroclava grewiae Sundararaj & David 1993: 240. Holotype pupal case. India: Tamil Nadu, Munchirai, 

3.viii.1987, R. Sundararaj, on Grewia sp., IDAV. 


Host: Apocynaceae: Rauvolfia densiflora; Araceae: Pothos scandens; Combretaceae: Calycopteris 

floribunda, Terminalia bellirica; Euphorbiaceae: Croton gibsonianus, Macaranga peltata; 

Fabaceae: Bauhinia phoenicea; Flacourtiaceae: Flacourtia indica, Hydnocarpus pentandra; 

Lauraceae: Smilax zeylanica, Litsea sp.; Loganiaceae: Strychnos wallichiana, Strychnos 

dalzellii; Lythraceae: Lagerstroemia macrocarpa; Melastomataceae: Melastoma 

malabathricum; Moraceae: Streblus asper, Ficus religiosa; Oleaceae: Olea dioica; 

Rhamnaceae: Ziziphus rugosa; Ziziphus xylophyrus; Rubiaceae: Catunaregam spinosa, 

Canthium parviflorum; Rutaceae: Glycosmis pentaphylla; Solanaceae: Solanum 

sisymbrifolium; Tiliaceae: Grewia orientalis, Grewia sp; Urticaceae: Boehmeria macrophylla. 

Aleuroclava guyavae (Takahashi) 1932 

Aleurotuberculatus guyavae Takahashi 1932: 22. Syntypes. Taiwan: on Psidium guajava, TARI. 

Aleuroclava guyavae (Takahashi); Martin 1999: 31. 

Syntypes. Taiwan: on Psidium guajava, TARI 


Host: Myrtaceae: Psidium guajava. 

Aleuroclava hexcantha (Singh) 1940 

Taiwanaleyrodes hexcantha Singh 1940: 194. Holotype. India: Chanda, Nagpur, xii.1937, K. Singh, on 

Bauhinia sp., ZSI. 

Aleurotuberculatus hexcantha (Singh); David 1976: 261-265. 

Aleuroclava hexcantha (Singh); Jesudasan and David 1990: 3. 


Host: Fabaceae: Bauhinia sp.; Moraceae: Ficus sp. 

Aleuroclava hikosanensis (Takahashi) 1938 

Aleurotuberculatus hikosanensis Takahashi 1938b: 71. Syntypes. Japan: on Cinnamonum sp., TARI; Suh et 

al. 2006. 

Japaneyrodes hikosanensis (Takahashi); Zahradnik 1962a: 14. 

Aleuroclava hikosanensis (Takahashi); Martin 1999: 31. 

Distribution: 5: Japan; Korea. 

Host: Aquifoliaceae: Ilex crenata, Ilex pedunculosa; Buxaceae: Buxus microphylla; Ericaceae: Pieris 

japonica; Lauraceae: Cinnamomum sp.; Myricaceae: Myrica sp.; Pittosporaceae: Pittosporum tobira;

Styracaceae: Styrax japonica; Theaceae: Eurya japonica. 

Aleuroclava hindustanicus (Meganathan & David) 1994 

Hindaleyrodes hindustanicus Meganathan & David 1994: 37. Holotype. India: on Glyptopetalum 

zeylandicum. 

Aleuroclava hindustanicus (Meganathan & David); Martin & Mound 2007. 


Host: Crassulaceae: Glyptopetalum zeylandicum. 

Aleuroclava indica (Singh) 1931 

Aleurothrixus indica Singh 1931: 84. Syntypes. India: on Ficus carica and Michelia champaca, BMNH. 

Taiwanaleyrodes indica (Singh) 1931: 84; Takahashi 1935b: 55; Sundararaj & David 1992a: 17. 

Aleuroclava indica (Singh) 1931; Manzari & Quicke 2006: 2470. 

Distribution: 5: Hong Kong; 6: India; Malaya; Malaysia. 

Host: Dilleniaceae: Dillenia indica; Lauraceae: Machilus sp.; Magnoliaceae: Michelia champaca; Moraceae: 

Ficus carica. 

Aleuroclava jasmini (Takahashi) 1932 

Aleurotuberculatus jasmini Takahashi 1932: 26-27. Syntypes. Taiwan: on Jasminum sp., TARI. 

Aleuroclava jasmini (Takahashi); Jesudasan and David 1990: 3. 

Distribution: 1: USA (CA, FL); 2: Paraguay; 3: Egypt; 5: China, Hong Kong, Japan; 6: India, Philippines, 

Singapore, Taiwan, Thailand; 7: Indonesia; 8: Guam; 9: USA: Hawaii. 

Host: Myrsinaceae: Maesa japonica; Rubiaceae: Jasminum sambac, Rubiaceae: Mitragyna sp. 

Rutaceae: Citrus sinensis, Murraya paniculata. 

Natural enemies: Aphelinidae: Encarsia lutea, Encarsia nipponica, Encarsia perflava. 

US Port Interceptions: Hawaii on Jasminum sp.; India on Murraya koenigii; Mexico on Gardenia sp. and 

Murraya sp.; Peru on an unidentified plant; Singapore on Murraya koenigii; Thailand on Ziziphus 

jujuba. 

Aleuroclava kanyakumariensis Sundararaj & David 1993 

Aleuroclava kanyakumariensis Sundararaj & David 1993: 241. Holotype pupal case. India: Tamil Nadu, 

Kunnathoor, 26.x.1988, R. Sundararaj, on Hemidesmus sp. 


Host: Asclepiadaceae: Hemidesmus sp., Flacourtiaceae: Flacourtia indica. 

Aleuroclava kavalurensis Jesudasan & David 1991 

Aleuroclava kavalurensis Jesudasan and David 1991: 262. Holotype puparium, India, Tamil Nadu, Kavalur, 

23.xi.1984, R.W. Alexander Jesudasan, on unidentified tree, IDAV; Dubey & Sundararaj 2005d. 


Host: Combretaceae: Calycopteris floribunda; Phyllanthaceae: Aporosa lindleyana; Rhamnaceae: Ziziphus 

xylophyrus. 

Aleuroclava kudremukhensis Dubey & Sundararaj 2005 

108

109 

Aleuroclava kudremukekhensis Dubey & Sundararaj 2005d: 254. Holotype puparium, India, Karnataka, 

Kudremukh National Park, 10.ix.2001, A. K. Dubey, on Leea indica; IFRI. 


Host: Vitaceae: Leea indica. 

Aleuroclava kuwani (Takahashi) 1934 

Aleurotuberculatus kuwani Takahashi 1934b: 54. Syntypes. Taiwan: on Murraya sp., TARI. 

Aleuroclava kuwani (Takahashi); Martin 1999: 31. 

Distribution: 5: China; Taiwan. 

Host: Lauraceae: Persea borbonia; Rutaceae: Murraya sp. 

Natural enemies: Aphelinidae: Encarsia perplexa. 

Aleuroclava lagerstroemiae (Takahashi) 1934 

Aleurotuberculatus lagerstroemiae Takahashi 1934b: 57. Syntypes. Taiwan: on Lagerstroemia indica, TARI. 

Aleuroclava lagerstroemiae (Takahashi); Martin 1999: 31. 


Host: Lythraceae: Lagerstroemia indica. 

Aleuroclava latus (Takahashi) 1934 

Aleurotuberculatus latus Takahashi 1934b: 47. Syntypes. Taiwan: on Neolitsea acuminatissima and 

Cinnamonum randaiense, TARI. 

Aleuroclava latus (Takahashi); Martin 1999: 31. 


Host: Lauraceae: Cinnamomum randaiense, Neolitsea acuminatissima. 

Aleuroclava lefroyi Sundararaj & David 1993 

Aleuroclava lefroyi Sundararaj & David 1993a: 98. Holotype puparium, India, Maharashtra, Mahableshwar, 

28.iii.1987, B. David, on Elatostemma sp., David Collection. 

Martiniella lefroyi (Sundararaj & David) 


Host: Urticaceae: Elatostema sp. 

Aleuroclava lithocarpi (Takahashi) 1934 

Aleurotuberculatus lithocarpi Takahashi 1934b: 50. Syntypes. Taiwan: on Lithocarpus uraiana, TARI. 

Aleuroclava lithocarpi (Takahashi); Martin 1999: 30 


Host: Fagaceae: Lithocarpus uraiana. 

Aleuroclava longisetosus Jesudasan & David 1991 

Aleuroclava longisetosus Jesudasan and David 1991: 261. Holotype. India: Tamil Nadu, Kolli Hills, 

20.iii.1972, B. V. David, on Neolitsea sp., IDAV; Dubey & Sundararaj 2005d. 


Host: Capparidaceae: Capparis shevaroyensis; Combretaceae: Terminalia paniculata.

110 

Dioscoreaceae: Dioscorea oppositifolia; Lauraceae: Cinnamomum malabatrum, Litsea sp. 

Neolitsea sp., Smilax zeylanica. 

Aleuroclava longispinus (Takahashi) 1934 

Aleurotuberculatus longispinus Takahashi 1934b: 51. Syntypes. Taiwan: on undetermined shrub, TARI. 

Aleuroclava longispinus (Takahashi); Martin 1999: 31. 


Host: on undetermined shrub. 

Aleuroclava louiseae Sundararaj & David 1993 

Aleuroclava louiseae Sundararaj & David 1993: 242-243. Holotype pupal case. India: Tamil Nadu, Cumbum, 

10.i.1988, S. Selvakumaran, on Ficus racemosa, IDAV. 


Host: Moraceae: Ficus racemosa. 

Aleuroclava macarangae (Corbett) 1935 

Aleurotuberculatus macarangae Corbett 1935b: 828. Syntypes. Malaya: on Macaranga sp. 

Martiniella macarangae (Corbett); Jesudasan and David 1990: 7; Sundararaj & Dubey 2004: 358. 

Aleuroclava macarangae (Corbett); Martin 1999: 31. 



Aleuroclava madhucae Jesudasan & David 1991 

Aleuroclava madhucae Jesudasan and David 1991: 262. Holotype. India: Tamil Nadu, Tambaram, 4.xi.1984, 

R. W. Alexander Jesudasan, on Madhuca longifolia, IDAV; Dubey & Sundararaj 2005d: 255. 


Host: Sapotaceae: Madhuca longifolia 

Aleuroclava magnoliae (Takahashi) 1952 

Aleurotuberculatus magnoliae Takahashi 1952: 19. Syntypes. Japan: Tokyo, on Acer palmata and many other 

plants, HBL. 

Aleuroclava magnoliae (Takahashi); Martin 1999: 31; Suh et al. 2006. 


Host: Aceraceae: Acer palmata; Cornaceae: Cornus alba; Magnoliaceae: Magnolia kobus. 

Aleuroclava malloti (Takahashi) 1932 

Aleurotuberculatus malloti Takahashi 1932: 25. Syntypes. Taiwan: on Mallotus sp., TARI 

Aleuroclava malloti (Takahashi); Martin 1999: 31. 


Host: Euphorbiaceae: Mallotus japonicus, Mallotus sp.; Polygonaceae: Polygonum thunbergii. 

Natural enemies: Aphelinidae: Encarsia duorunga, Encarsia lutea, Encarsia perflava, Encarsia synaptocera.

111 

Aleuroclava manii (David) 1978 

Aleurotuberculatus manii David 1978: 133-135. Holotype. India: Keraala, Walayar, 24.iii.1972, B. V. David, 

on Homonoia riparia, IDAV. 

Aleuroclava manii (David); Jesudasan and David 1990: 3. 



Aleuroclava martini Dubey & Sundararaj 2005 

Aleuroclava martini Dubey & Sundararaj 2005d: 256. Holotype puparium, India, Karatataka, Kudremhukh 

National Park, 12.viii.2001, A. K. Dubey, on Celtis philippensis, IFRI. 


Host: Ulmaceae: Celtis philippensis. 

Aleuroclava maximus (Qureshi) 1982 

Aleurotuberculatus maximus Qureshi 1982: 51. Holotype puparium, Pakistan, Karachi, 26.vi.1976, J.I. 

Qureshi, ex. Zizyphus jujuba, FAI. 


Host: Rhamnaceae: Ziziphus [=Zizyphus] jujuba. 

Aleuroclava melastomae (Takahashi) 1934 

Aleurotuberculatus melastomae Takahashi 1934b: 52. Syntypes. Taiwan: on Melastoma candidum, TARI. 

Aleuroclava melastomae (Takahashi); Martin 1999: 31. 


Host: Melastomataceae: Melastoma candidum. 

Natural enemies: Aphelinidae: Encarsia lutea, Encarsia nipponica. 

Aleuroclava meliosmae (Takahashi) 1932 

Taiwanaleyrodes meliosmae Takahashi 1932: 28. Syntypes. Taiwan: on Meliosma rhoifolia and other plants, 

TARI. 

Aleuroclava meliosmae (Takahashi); Manzari & Quicke 2006: 2470. 


Hosts: Aceraceae: Acer kawakamii; Daphniphyllaceae: Daphniphyllum glaucescens; Lauraceae: Machilus sp.; 

Sabiaceae: Meliosma rhoifolia. 

Natural enemies: Encarsia cibcensis; Encarsia lutea, Encarsia perflava. 

Aleuroclava montanus (Takahashi) 1939 

Taiwanaleyrodes montanus Takahashi 1939a: 80. Syntypes. Taiwan: on Lauraceae. 

Aleuroclava montanus (Takahashi); Manzari & Quicke 2006: 2470. 


Host: Lauraceae.

112 

Aleuroclava multipori (Takahashi) 1935 

Aleurotuberculatus multipori Takahashi 1935b: 53. Syntypes. Taiwan: on Oreocnide pedunculata, TARI. 

Aleuroclava multipori (Takahashi); Martin 1999: 31. 

Aleurotuberculatus multipori var. thaiensis Takahashi 1942h: 334. Syntypes. Thailand: on Fagaceae, TARI. 

Distribution: 4: Mauritius; 5: Taiwan; 6: Thailand. 

Host: Caprifoliaceae: Viburnum taiwanianun; Fagaceae; Urticacaeae: Oreocnide pedunculata. 

Natural enemies: Host: Aphelinidae: Encarsia duorunga, Encarsia synaptocera. 

Aleuroclava multituberculata Sundararaj & David 1993 

Aleuroclava multituberculata Sundararaj & David 1993: 243-244. Holotype pupal case. India: Kerala, 

Ambalamedu, 3.vii.1987, R. Sundararaj, on Litsea sp. 


Host: Malvaceae: Hibiscus rosa-sinensis; Lauraceae: Actinodaphne sp., Cinnamomum malabatrum, Litsea sp. 

Aleuroclava murrayae (Singh) 1931 

Aleurotrachelus murrayae Singh 1931: 62. Syntypes. India: on Murraya exotica, ZSI. 

Aleuroclava murrayae (Singh); Martin 1999: 31; Dubey & Sundararaj 2005. 

Aleuromigda deghai Dozier & Baker; nomen nudum, synonymy according to Hussain & Khan 1945: 32; 


Distribution: 5: Taiwan; 6: India. 

Host: Rutaceae: Murraya exotica; Urticaceae: Elatostema lineolatum. 

Natural enemies: Aphelinidae: Encarsia longicauda, Encarsia perflava. 

Aleuroclava mysorensis Jesudasan & David 1991 

Aleuroclava mysorensis Jesudasan and David 1991: 262-263. Holotype. India: Karnataka, Mysore, 

19.ix.1985, B. V. David, on Ailanthus excelsa. 


Host: Simaroubaceae: Ailanthus excelsa. 

Aleuroclava nachiensis (Takahashi) 1963 

Aleurotuberculatus nachiensis Takahashi & 1963 1963: 50. Holotype. Japan: on Lauraceae tree, HBL. 

Aleuroclava nachiensis (Takahashi); Martin 1999: 31. 


Host: Lauraceae tree. 

Aleuroclava nagercoilensis Sundararaj & David 1993 

Aleuroclava nagercoilensis Sundararaj & David 1993: 244-245. Holotype pupal case. India: Tamil Nadu, 

Ulakkai falls, on Cissus sp., R. Sundararaj, IDAV; Dubey & Sundararaj 2005d: 258. 


Host: Vitaceae: Cissus sp. 

Aleuroclava nanjangudensis Jesudasan & David 1991 

Aleuroclava nanjangudensis Jesudasan & David 1991: 263. Holotype. India: Karnataka, Nanjangud, 

1.viii.1984, R. W. Alexander Jesudasan, on unidentified tree.


Host: Flacourtiaceae: Flacourtia indica; unidentified tree. 

Aleuroclava neolitseae (Takahashi) 1934 

Aleurotuberculatus neolitseae Takahashi 1934b: 55. Syntypes. Taiwan: on Neolitsea acuminatissima, TARI. 

Aleuroclava neolitseae (Takahashi); Martin 1999: 31. 

Distribution: 3: Iran; 5: Taiwan; 6: Malaysia; 7: New Guinea; Sarawak; Sulawesi. 

Host: Amaranthaceae: Amaranthus graecizans; Dipterocarpaceae: Anisoptera thurifera polyandra; 

Ebenaceae: Diospyros sp.; Guttiferae: Callophyllum sp.; Lauraceae: Neolitsea acuminatissima; 

Magnoliaceae: Elmerrillea papuana; Moraceae: Ficus carica, Ficus capensis; Myristiaceae: 

Gymnacranthera sp.; Myrtaceae: Myrtella sp., Szygium sp., Syzygium calucob; Ochnaceae: 

Schuurmansia henningsii; Xanthophyllaceae: Xanthophyllum papuanum. 

Aleuroclava nephelii (Corbett) 1935 

Aleurotuberculatus nephelii Corbett 1935b: 831. Syntypes. Malaya: on Nephelium lappaceum and other 

plants. 

Aleuroclava nephelii (Corbett); Martin 1999: 31 


Host: Sapindaceae: Nephelium lappaceum. 

Aleuroclava nigeriae (Mound) 1965 

Aleurotuberculatus nigeriae Mound 1965c: 136. Syntypes. Nigeria: on Psidium guajava, NHM. 

Aleuroclava nigeriae (Mound); Martin 1999: 31 

Aleurotuberculatus caloncobae Cohic 1966a: 28. Syntypes. Congo: on Caloncoba welwitschii; synonymy 

according to Bink-Moenen 1983: 84. 

Aleurotuberculatus kusheriki Mound 1965c: 138. Holotype. Nigeria: on undetermined plant, NHM; synonymy 


Aleurotuberculatus yambiae Gameel 1971: 169. Holotype. Sudan: on Vitex cuneata, GRF; synonymy 


Distribution: 4: Angola; Chad; Congo; Ghana; Ivory Coast; Kenya; Nigeria; Sudan. 

Host: Bignoniaceae: Stereospermum kunthianum, Markhamia sessilis; Burseraceae: Commiphora 

pedunculata; Combretaceae: Combretum micranthum, Combretum confertum, Combretum molle, 

Combretum glutinosum, Anogeissus leiocarpus; Euphorbiaceae: Alchornea cordifolia, Bridelia 

ferruginea, Hymenocardia acida; Fabaceae: Daniellia olivera, Millettia laurentii; Flacourtiaceae: 

Caloncoba welwitschi; Loganiaceae: Nuxia congesta, Strychnos spinosa, Strychnos variabilis; 

Moraceae: Ficus dicranostyla, Ficus verruculosa, Ficus sp., Bosqueiopsis gilletii; Myrtaceae: Psidium 

guajava; Rubiaceae: Crossopteryx febrifuga, Gardenia jovis-tonantis, Gardenia triacantha, 

Sarcocephalus diderrichii, Mitragyna inermis, Nauclea latifolia, Crossopteryx febrifuga, Tarenna sp.; 

Sterculiaceae: Sterculia bequaerti; Ulmaceae: Celtis milbraedii; Verbenaceae: Vitex cuneata. 

US Port Interceptions: Africa on Jasminum grandiflorum; Nigeria on Cola acuminata and Cola sp. 

Aleuroclava nitidus (Singh) 1932 

Dialeurodes nitidus Singh 1932: 84. Syntypes. Burma: on unidentified host, ZSI. 

Taiwanaleyrodes nitidus (Singh); Mound and Halsey 1978: 195. 

Aleurotuberculatus nitidus (Singh); Martin 1988: 69. 

113

114 

Aleuroclava nitidus (Singh); Martin 1999: 31. 

Aleurolobus sandorici Corbett 1935b: 817. Syntypes. Malaya: on Sandoricum indicum; synonomy according 

to Martin & Mound 2007: 11. 

Aleurotuberculatus sandorici (Corbett); Mound and Halsey 1978: 88. 

Aleuroclava sandorici (Corbett); Martin 1999: 31. 

Distribution: 6: Burma, Malaya, Malaysia; Thailand; 7: Sulawesi. 

Host: Magnoliaceae: Michelia champaca; Meliaceae: Sandoricum indicum.Moraceae: Ficus sp. 

Aleuroclava orientalis Jesudasan & David 1991 

Aleuroclava orientalis Jesudasan and David 1991: 29-32. Holotype. India: Tamil Nadu, Coimbatore, 

19.vi.1985, R. W. Alexander Jesudasan, on Holoptelea integrifolia, IDAV. 

Distribution: 6: India; Sri Lanka. 

Host: Ulmaceae: Holoptelea integrifolia. 

Aleuroclava papillata Sundararaj & Dubey 2004 

Martiniella papillata Sundararaj & Dubey 2004: 358. Holotype puparium, Goa, Volpoi, 27.ii.2001, A. K. 

Dubey, on Xeromphis spinosa, IDEH. 

Aleuroclava papillata (Sundararaj & Dubey); Martin 1999. 


Host: Rubiaceae: Xeromphis spinosa. 

Aleuroclava parvus (Singh) 1938 

Aleurotuberculatus parvus Singh 1938: 189. Syntypes. Burma: on Bombax malabaricum, ZSI. 

Aleuroclava parvus (Singh); Martin 1999: 31. 


Host: Bombacaceae: Bombax malabaricum. 

Aleuroclava pentatuberculata Sundararaj & David 1993 

Aleuroclava pentatuberculata Sundararaj & David 1993: 245-246. Holotype pupal case. India: Tamil Nadu, 

Kunnathoor, R. Sundararaj, on unidentified plant, IDAV; David & Ragupathy 2004. 


Host: Moraceae: Morus alba. 

Aleuroclava philomenae Jesudasan & David 1991 

Aleuroclava philomenae Jesudasan and David 1991: 264. Holotype. India: Kerala, Elamthottam, 21.x.1984, B. 

V. David, on Colocasia sp.; IDAV. 


Host: Araceae: Colocasia sp. 

Aleuroclava phyllanthi (Corbett) 1935 

Aleurolobus phyllanthi Corbett 1935b: 818. Holotype. Malaysia: Juala Lumpur (Selangor), on Phyllanthus 

frondosus. 

Aleurotuberculatus phyllanthi (Corbett); Mound & Halsey 1978: 87. 

Aleuroclava phyllanthi (Corbett); Martin 1999: 31

Distribution: 6: Malaysia 

Host: Euphorbiaceae: Phyllanthus frondosus . 

Aleuroclava piperis (Takahashi) 1935 

Aleurotuberculatus piperis Takahashi 1935b: 52. Syntypes. Taiwan: on Piper sp., TARI 

Aleuroclava piperis (Takahashi); Martin 1999: 31. 


Host: Piperaceae: Piper sp. 

Aleuroclava pongamiae Jesudasan & David 1991 

Aleuroclava pongamiae Jesudasan and David 1991: 264-265. Holotype. India: Bihar, Pusa, K. Singh, on 

Pongamia glabra, IDAV. 

Distribution: 6: India; 7: Sulawesi. 

Host: Fabaceae: Pongamia pinnata [=glabra]. 

Aleuroclava porosus (Priesner & Hosny) 1937 

Trialeurodes porosus Priesner & Hosny 1937: 45. Syntypes. Egypt: Assiout, 2.iii.1932, Priesner & Hosny, on 

Zizyphus spina-christi. 

Aleurotuberculatus porosus (Priesner & Hosny); Mound and Halsey 1978: 87. 

Aleuroclava porosus (Priesner & Hosny); Martin 1999: 31. 

Distribution: 3: Egypt; Sudan. 

Host: Rhamnaceae: Ziziphus spina-christi, Ziziphus jujube. 

Aleuroclava psidii (Singh) 1931 

Aleurotrachelus psidii Singh 1931: 61. Syntypes. India: on Psidium guajava, ZSI. 

Aleurotuberculatus psidii (Singh); Takahashi 1932: 20. 

Aleuroclava psidii (Singh); Martin 1999: 31. 

Distribution: 5: China; Taiwan; 6: India. 

Host: Moraceae: Streblus asper; Myrtaceae: Psidium guajava. 


Aleuroclava pulcherrima (Corbett) 1935 

Aleurolobus pulcherrimus Corbett 1935b: 822. Syntypes. Malaya: on Erythina stricta. 

Aleurotuberculatus pulcherrimus (Corbett); Mound and Halsey 1978: 88. 

Aleuroclava pulcherrimus (Corbett); Martin 1999: 31. 


Host: Fabaceae: Erythina stricta. 

Aleuroclava pyracanthi (Takahashi) 1933 

Aleurotuberculatus pyracanthae Takahashi 1933: 12. Syntypes. Taiwan: on Pyracantha koidzumii, TARI. 

Aleuroclava pyracanthae (Takahashi); Martin 1999: 31 


Host: Pyracantha koidzumii. 

115

Aleuroclava ramachandrani Dubey & Sundararaj 2005 

Aleuroclava ramachandrani Dubey & Sundararaj 2005d: 260.Holotype puparium, India, Karnataka, Unachali 

Falls, 19.ii.2001, A. K. Dubey, on Syzygium sp., IFRI. 


Host: Myrtaceae: Syzygium sp. 

Aleuroclava regui Sundararaj & David 1993 

Aleuroclava regui Sundararaj & David 1993: 247-248. Holotype pupal case. India: Maharashtra, Khopoli, 

5.x.1988, K. Regu, on unidentified grass, IDAV. 


Host: Poaceae: unidentified grass. 

Aleuroclava rhododendri (Takahashi) 1935 

Aleurotuberculatus rhododendri Takahashi 1935b: 51. Syntypes. Taiwan: on Rhododendron sp., TARI. 

Aleuroclava rhododendri (Takahashi); Martin 1999: 31. 


Host: Ericaceae: Rhododendron sp. 


Aleuroclava saputarensis Sundararaj & David 1993 

Aleuroclava saputarensis Sundararaj & David 1993: 247-248. Holotype pupal case. India: Maharashtra, 

Saputara, 27.iii.1987, on unidentified plant, IDAV; Dubey & Sudararaj 2005d: 261. 


Host: Apocynaceae: Wrightia tinctoria; Fabaceae: Acacia sp., unidentified plant. 

Aleuroclava selvakumarani Sundararaj & David 1993 

Aleuroclava selvakumarani Sundararaj & David 1993: 248. Holotype pupal case. India: Tamil Nadu, 

Cumbum,, 7.iv.1988, S. Selvakumaran, on Chickrassia velutina, IDAV. 


Host: Cucurbitaceae: Trichosanthes tricuspidata; Flacourtiaceae: Flacourtia montana; Malvaceae: Kydia 

calycina; Meliaceae: Chickrassia velutina, Chukrassia tabalaris; Rhamnaceae: Ziziphus rugosa. 

Aleuroclava sepangensis Martin & Mound 2007 

Taiwanaleyrodes macarangae Corbett 1935b: 840. Syntypes. Malaya: on Macaranga megalophylla. 

Aleuroclava sepangensis Martin & Mound 2007: 56; new name, preoccupied by Aleurotuberculatus 

macarangae Corbett 1935b: 828. 


Host: Euphorbiaceae: Macaranga megalophylla. 

Aleuroclava siamensis (Takahashi) 1942 

Aleurotuberculatus siamensis Takahashi 1942h: 335. Syntypes. Thailand: on unidentified host, TARI. 

Aleuroclava siamensis (Takahashi); Martin 1999: 31. 


116


Aleuroclava similis (Takahashi) 1938 

Aleurotuberculatus similis Takahashi 1938b: 73. Syntypes. Japan: on Ilex crenata, I. peduculosa and Pieris 

japonicum, TARI. 

Japaneyrodes similis (Takahashi); Zahradnik 1962a: 14. 

Aleuroclava similis (Takahashi); Martin 1999: 31. 

Japaneyrodes similis europaeus Zahradnik 1962a: 15. Holotype. Czech Republic: on Vaccinium vitis; Martin 

et al. 2000: 420. 

Aleurotuberculatus similis europaeus (Zahradnik); Mound and Halsey 1978: 89. 

Aleuroclava similis europaeus (Zahradnik); Martin 1999: 31. 

Japaneyrodes similis suborientalis Danzig 1966: 383. Holotype. USSR: southern Maritime Territory, on 

Vaccinum vitis idaea, ZIN; Martin et al. 2000: 420. 

Aleurotuberculatus similis suborientalis (Danzig); Mound and Halsey 1978: 89. 

Aleurochiton vaccinii (Kunow); misidentification Ryberg 1938: 16 (in part). 

Distribution: 1: USA (NY); 2: Cuba; 3: Austria, Czech Republic; Finland; Germany, Iran, Netherlands, 

Norway, Poland, Russia, Sweden; 5: China, Japan. 

Host: Amaranthaceae: Amaranthus retroflexus; Aquifoliaceae: Ilex vomitoria, Ilex crenata, I. peduculosa; 

Ericaceae: Pieris japonicum, Ericaceae: Rhododendron sp., Vaccinium vitis-idaea ssp. minus, 

Ericaceae: Vaccinum vitis idaea; Theaceae: Camellia japonica. 

Natural enemies: Aphelinidae: Eretmocerus diversiciliatus, Eretmocerus zippanguiphagus; Platygastridae: 

Amitus croesus. 

Aleuroclava simplex (Takahashi) 1949 

Aleurotuberculatus simplex Takahashi 1949: 50. Holotype. Riouw Islands (south of Singapore): on 

unidentified plant, HBL. 

Aleuroclava simplex (Takahashi); Martin 1999: 31. 



Aleuroclava singhi Jesudasan & David 1991 

Aleuroclava singhi Jesudasan and David 1991: 265. Holotype. India: Bihar, Pusa, K. Singh, on Diopyros 

montana, IDAV. 


Host: Ebenaceae: Diospyros montana. 

Aleuroclava sivakasiensis Sundararaj & David 1993 

Aleuroclava sivakasiensis Sundararaj & David 1993: 248-249. Holotype pupal case. India: Tamil Nadu, 

Sivakasi, S. Jayaselvan, on unidentified plant, IDAV. 


Host: Euphorbiaceae: Manihot esculenta; Rubiaceae: Jasminum malabaricum; unidentified plant. 

Aleuroclava srilankaensis (David) 1993 

Taiwanaleyrodes srilankaensis David 1993: 29. Holotype. Sri Lanka: on Macaranga sp. 

117

Aleuroclava srilankaensis (David); Manzari & Quicke 2006: 2470 



Aleuroclava stereospermi (Corbett) 1935 

Aleurotuberculatus stereospermi Corbett 1935b: 832. Syntypes. Malaya: on Stereospermum chelonoides. 

Aleuroclava stereospermi (Corbett); Martin 1999: 31. 


Host: Bignoniaceae: Stereospermum chelonoides. 

Aleuroclava subindica Martin & Mound 2007 

Aleuroclava papillata Dubey & Sundararaj 2005d: 258. Holotype puparium, India, Karnataka, Kudremhukh 

National Park, 11 ix 2001, A. K. Dubey, on unidentified plant, IFRI; name preoccupied by Martiniella 

[=Aleuroclava] papillata Sundararaj & Dubey 2004, new name Martin & Mound 2007. 



Aleuroclava submarginatus (Qureshi) 1982 

Aleurotuberculatus submarginatus Qureshi 1982: 53. Holotype puparium, Pakistan, Murree, 20.iv.1973, J.I. 

Qureshi, ex. Ficus carica, FAI.. 

Aleuroclava submarginatus (Qureshi); Martin 1991: 31. 


Host: Moraceae: Ficus carica. 

Aleuroclava takahashii (David & Subramaniam) 1976 

Aleurotuberculatus takahashii David & Subramaniam 1976: 174-175. Holotype. India: Tamil Nadu, Madras, 

19.vii.1971, B. V. David, on Cordia myxa, IDAV. 

Aleuroclava takahashii (David & Subramaniam); Jesudasan and David 1990: 1-13; Dubey & Sundararaj 

2005d. 

Distribution: 3: USSR; 5: China; 6: Burma; India. 

Host: Boraginaceae: Cordia myxa; Euphorbiaceae: Macaranga peltata; Icacinaceae: Mappia sp.; 

Loganiaceae: Strychnos nux-vomica; Moraceae: Streblus asper, Ficus racemosa; Musaceae: Musa sp.; 

Myrtaceae: Syzygium lanceolatum, Syzygium cumini; Rhamnaceae: Ziziphus sp.; Ulmaceae: Zelkova 

sp. 

Other reference: Chou & Yan 1988. 

Aleuroclava tarennae Martin & Mound 2007 

Aleuroclava davidi Dubey & Sundararaj 2005d: 249. Holotype puparium, India, Tamil Nadu, 19.viii.2001, 

AK Dubey, on Tarenna asiatica, IFRI; name preoccupied by Aleuroclava davidi Qureshi 1982; new 

name by Martin & Mound 2007. 

Aleuroclava tarennae Martin & Mound 2007 


Host: Rubiaceae: Tarenna asiatica. 

118

119 

Aleuroclava tentactuliformis (Corbett) 1935 

Aleurotuberculatus tentactuliformis Corbett 1935b: 825. Syntypes. Malaya: on Conocephalus subrinervius. 

Aleuroclava tentactuliformis (Corbett); Martin 1999: 31. 


Host: Moraceae: Conocephalus subrinervius. 

Aleuroclava terminaliae Sundararaj & David 1993 

Aleuroclava terminaliae Sundararaj & David 1993: 250. Holotype pupal case. India: Maharashtra, Saputara, 

26.iii.1987, B. V. David, on Terminalia sp., IDAV. 


Host: Combretaceae: Terminalia sp. 

Aleuroclava thysanospermi (Takahashi) 1934 

Aleurotuberculatus thysanospermi Takahashi 1934b: 56. Holotype. Taiwan: on Thysanospermum diffisum, 

TARI. 

Aleuroclava thysanospermi (Takahashi); Martin 1999: 31. 


Host: Apocynaceae: Thysanospermum diffisum. 

Aleuroclava trachelospemi (Takahashi) 1938 

Aleurotuberculatus trachelospermi Takahashi 1938b: 72. Syntypes. Japan: Trachelospermum asiaticum var. 

intermedium, TARI. 

Japaneyrodes trachelospermi (Takahashi); Zahradnik 1962a: 13. 

Aleuroclava trachelospermi (Takahashi); Martin 1999: 31. 


Host: Apocynaceae: Trachelospermum asiaticum var. intermedium. 

Aleuroclava trilineata Sundararaj & David 1993 

Aleuroclava trilineata Sundararaj & David 1993: 250-251. Holotype pupal case. India: Tamil Nadu, 

Kunnathoor, 27 II 1988, R. Sundararaj, on Streblus asper, IDAV. 


Host: Moraceae: Streblus asper. 

Aleuroclava tripori (Dubey & Sundararaj) 2006 

Taiwanaleyrodes tripori Dubey & Sundararaj 2006: 73. Holotype puparium, India, on unknown plant. 

Aleuroclava tripori (Dubey & Sundararaj); Manzari & Quicke 2006: 2470. 


Host: unknown plant. 

Aleuroclava trivandricus Dubey & Sundararaj 2006 

Aleuroclava trivandricus Dubey & Sundararaj 2006e: 229. Holotype puparium, India, Kerala, 

Trivandrum, 25.V.2001, A. K. Dubey, ex. Terminalia sp., NFICFRI.


Host: Combretaceae: Terminalia sp. 

Aleuroclava trochodendri Takahashi 1957 

Aleuroclava trochodendri Takahashi 1957: 12. Syntypes. Japan: on Trochodendron araliodes, HBL. 


Host: Trochodendraceae: Trochodendron aralioides. 

Aleuroclava ubonensis (Takahashi) 1942 

Aleurotuberculatus ubonensis Takahashi 1942h: 332. Syntypes. Thailand: on undetermined tree. 

Aleuroclava ubonensis (Takahashi); Martin 1999: 31. 


Host: undetermined tree. 

Aleuroclava uraianus (Takahashi) 1932 

Aleurotuberculatus uraianus Takahashi 1932: 27. Syntypes. Taiwan: on Clerodendrum sp., TARI. 

Aleuroclava uraianus (Takahashi); Martin 1999: 31. 


Host: Verbenaceae: Clerodendrum sp. 

Natural enemies: Aphelinidae: Encarsia perflava. 

Aleuroclava vernoniae Meganathan & David 1994 

Aleuroclava vernoniae Meganathan & David 1994: 18. Holotype. India: on Vernonia arborea. 


Host: Asteraceae: Vernonia arborea. 

Aleuroclava vitexae Sundararaj & David 1993 

Aleuroclava vitexae Sundararaj & David 1993: 251-252. Holotype pupal case. India: Tamil Nadu, Ulakkai 

falls, 6.viii.1987, R. Sundararaj, on Vitex sp., IDAV. 


Host: Verbenaceae: Vitex sp. 

Aleuroclava wrightiae Jesudasan & David 1991 

Aleuroclava wrightiae Jesudasan and David 1991: 266. Holotype. India: Tamil Nadu, Madras, 9.xi.1993, R. 

W. Alexander Jesudasan, on Wrightia tinctoria, IDAV. 


Host: Annonaceae: Polyalthia cerasoides; Apocynaceae: Wrightia tinctoria; Campanulaceae: Lobelia 

nicotianaefolia; Icacinaceae: Nothapodytes nimmoniana; Loganiaceae: Strychnos nux-vomica; 

Oleaceae: Olea dioica. 

Synonyms and Transfers 

Aleuroclava cherasensis (Corbett) 1935 - Minutaleyrodes 

Aleuroclava suishanus (Takahashi) year? - Minutaleyrodes 

120

Aleurocybotus Quaintance & Baker 1914 

Aleurocybotus Quaintance & Baker 1914: 101. Type species. Aleurodes graminicola Quaintance 1900, by 

monotypy. 

Reference: Bink-Moenen (1983) originally separated the graminicolus group (=Aleurocybotus) from 

the setiferus group (=Vasdavidius) by having the antennae long and lateral to the prothoracic legs in 

the former versus having the antennae short and mesal to the prothoracic legs in the latter. 



cereus x 

graminicolus x 

occiduus x x x 

Total = 3 species 2 2 0 0 0 0 0 0 1 

Species Transfers 

Aleurocybotus cobarensis Martin 1999 - Vasdavidius 

Aleurocybotus concursus Ko 1998 - Vasdavidius 

Aleurocybotus indicus David & Subramaniam 1976 - Vasdavidius 

Aleurocybotus miscanthus Ko 1998 - Vasdavidius 

Aleurocybotus setiferus Quaintance & Baker 1917 - Vasdavidius 

Host: usually found on grasses (Poaceae) 

References - Russell (2000) - Separation of Aleurocybotus versus Vasdavidius species; Ko et al. (1998) - 

Key to Taiwanese species 

Aleurocybotus cereus Martin 2005 

Aleurocybotus cereus Martin 2005b: 11. Holotype puparium, Belize, CFR, 24.vi.2002, J. H. Martin, on 

Lasiacis ?rugelii (Poaceae), BMNH. 


Host: Poaceae: Lasiacis rugelli? 

Aleurocybotus graminicolus (Quaintance) 1899 

Aleurodes graminicolus Quaintance 1899b: 89. Syntypes. USA: Florida, Lake City, 24.vii.1898, P. H. Rolfs, 

on undetermined grass, USNM. 

Aleurocybotus graminicolus (Quaintance); Quaintance & Baker 1914: 101. 

Distribution: 1: USA (FL). 


Aleurocybotus occiduus Russell 1964 

Aleurocybotus occiduus Russell 1964b: 101. Holotype. USA: California, on Cynodon dactylon, USNM 

Distribution: 1: USA (AZ, CA, FL, HI, TX*); 2: Peru; 9: Hawaii. 

Host: Cyperaceae Cyperus rotundus; Poaceae: Cenchrus echinatus, Chloris sp., Cynodon dactylon, 

Echinochloa crus-galli, Oryza sativa L, Paspalum dilatatum, Paspalum notatum, Setaria italica, 

121

122 

Sorghum halepense, Sorghum vulgare var. sudanese, Stenotaphrum secundatum, Zea mays. 

Natural enemies: Aphelinidae: Encarsia luteola; Eulophidae: Euderomphale hyalina. 

New Record: USA: Texas, 23 FEB 2006, J. Goolsby, on Arundo donax.

Aleurocyperus Ko & Dubey 2007 

Aleurocyperus humus Ko & Dubey 2007: 37-54. Type species – Aleurocyperus humus Ko & 

Dubey 2007, by original designation. 



humus x 

Total= 1 0 0 0 0 1 0 0 0 0 

Aleurocyperus humus Ko and Dubey 2007 

Aleurocyperus humus Ko & Dubey 2007: 37-54. Holotype puparium, Taiwan: Baihe, on Eleusine 

indica, 28.ix.2003, C. C. Ko, NTU. 


Host: Cyperaceae: Cyperus rotundus, Kyllinga brevifolia, Poaceae: Eleusine indica. 

123

Aleuroduplidens Martin 1999 

Aleuroduplidens Martin 1999: 35. Type species: Aleuroduplidens eucalyptifolia Martin 1999, by original 

designation. 



carverae x 

croceata x 

eucalyptifolia x 

santali x 

triangularis x 

wellsae x 

Total = 6 species 0 0 0 0 0 0 6 0 0 

Reference: Martin (1999) - Key to 6 species from Australia. 

Aleuroduplidens carverae Martin 1999 

Aleuroduplidens carverae Martin 1999: 36. Holotype puparium. Australia: on Cerbera manghas, ANIC. 


Host: Apocynaceae: Cerbera manhas. 

Aleuroduplidens croceata (Maskell) 1896 

Aleurodes croceata Maskell 1895: 428. Lectotype. Australia: on Monotoca [=Styphelia] elliptica 

[Epacridaceae], ADSIR. 

Aleurotrachelus croceatus (Maskell); Quaintance & Baker 1914: 103. 

Tetraleurodes croceata (Maskell); Dumbleton 1956b: 177. 

Aleuroduplidens croceata (Maskell) 1896; Martin 1999: 37. 


Host: Epicridaceae: Monotoca elliptica. 

Aleuroduplidens eucalyptifolia Martin 1999 

Aleuroduplidens eucalyptifolia Martin 1999: 38. Holotype puparium, Australia, CT, Molonglo Gorge, on 

Eucalyptus nortonii, 8.xii.1996, J.H. Martin, ANIC. 


Host: Myrtaceae: Eucalyptus nortonii. 

Aleuroduplidens santali Martin 1999 

Aleuroduplidens santali Martin 1999: 39. Holotype puparium. Australia: on Santalum lanceolatum, ANIC. 


124

Host: Santalaceae: Santalum lanceolatum. 

Aleuroduplidens triangularis Martin 1999 

Aleuroduplidens triangularis Martin 1999: 40. Holotype puparium. Australia: on Acaria ?mcdonnelliensis, 

ANIC 


Host: Fabaceae: Acacia ?mcdonnelliensis. 

Aleuroduplidens wellsae Martin 1999 

Aleuroduplidens wellsae Martin 1999: 40. Holotype puparium. Australia: on Acacia ?aneura, ANIC 

Distribution: 7: Australia 

Host: Fabaceae: Acacia ?aneura. 

125

Aleuroglandulus Bondar 1923 

Aleuroglandulus Bondar 1923a: 121. Type species. Aleuroglandulus subtilis Bondar 1923, by monotypy. 

Reference - Russell (1944); Martin (2005) Key to Belize species. 



inanis x 

magnus x 

striatus x 

subtilis x x 

Total 4 species 1 4 0 0 0 0 0 0 0 

Aleuroglandulus inanis Martin 2005 

Aleuroglandulus inanis Martin 2005b: 13. Holotype puparium, Belize, CFR, 5.iii.1996, J. H. Martin, on 

Carica sp., BMNH. 


Host: Caricaceae: Carica sp. 

Aleuroglandulus magnus Russell 1944 

Aleuroglandulus magnus Russell1944a: 4. Holotype. Panama: on Synechantus warscewiszianus, USNM 

Distribution: 2: Panama. 

Host: Arecaceae: Chamaedorea wendleandiana, Synechantus warscewiszianus. 

Aleuroglandulus striatus Sampson & Drews 1941 

Aleuroglandulus striatus Sampson & Drews 1941: 157. Syntypes. Mexico: on unidentified plant, UCD; 

Russell 1944a: 6; Martin 2005b: 15. 

Distribution: 2: Belize, Costa Rica, Guatemala; Honduras; Mexico. 

Host: Flacourtiaceae: Casearia sp., Latia sp; Meliaceae: Trichilia? sp.; Polygonaceae: Coccoloba schiedeana. 

Aleuroglandulus subtilis Bondar 1923 

Aleuroglandulus subtilis Bondar 1923a: 121. Syntypes. Brazil: on Chomelia oligantha. 

Aleuroglandulus striatus Sampson & Drews 1941; misidentification in part; Sampson & Drews 1941: 157. 

Aleuroglandulus emmae Russell 1944a: 5. Holotype. Mexico: Guerrero, on Flacourtiaceae, USNM; synonymy 

according to Martin 2005b: 67. 

Aleuroglandulus malangae Russell 1944a: 5. Holotype. Cuba: on Xanthosoma sp., USNM; synonymy 


Distribution: 1: USA (FL, NC, NV, TX, UT); 2: Bahamas, Brazil, Colombia, Costa Rica, Cuba, Dominica, El 

Salvador, Guatemala, Guadeloupe, Haiti, Honduras, Jamaica, Mexico, Panama, Peru, Puerto Rico, 

Venezuela. 

Host: Araceae Alocasia sp.; Caladium bicolor, Caladium x hortulanum, Colocasia antiquorum, Colocasia 

126

127 

esculentum, Xanthosoma sagittifolium, Xanthosoma undipes; Arecaceae: Chamaedorea wendlandiana, 

Synechanthus warscewiszianus; Euphorbiaceae: Euphorbia heterophylla, Jatropha sp.; Fabaceae: 

Clitoria laurifolia, Erythrina sp.; Galactia acapulcensis, Schizolobium sp.; Lauraceae: Persea 

americana, Persea borbonia; Myrtaceae: Psidium guajava; Rhamnaceae: Gouania lupuloides; 

Rubiaceae: Chomelia oligantha, Gardenia sp. 

Natural enemies: Parasitoids - Aphelinidae: Encarsia formosa, Encarsia hispida, Encarsia portoricensis, 

Encarsia strenua group, Encarsia tabacivora; Eretmocerus sp.; Platygastridae: Amitus aleuroglanduli; 

Predators - Coccinellidae: Delphastus sp. 

US Port Interceptions: Mexico on Caladium sp.; Chamaedorea sp., Fabaceae; Ficus carica, Hexisea bidentata 

Litsea sp.; Antigua and Barbuda on Xanthosoma sp.; Dominica on Colocasia esculenta.

Aleuroinanis Martin 1999 

Aleuroinanis MARTIN 1999: 41. Type species. Aleuroinanis myrtacei Martin 1999, by monotypy 



myrtacei x 

Total = 1 species 0 0 0 0 0 0 1 0 0 

Aleuroinanis myrtacei Martin 1999 

Aleuroinanis myrtacei Martin 1999: 42. Holotype puparium. Australia: on Syzgium sp., ANIC. 



128

Aleurolobus Quaintance & Baker 1914 

Aleurolobus Quaintance & Baker 1914: 108. Type species: Aleurodes marlatti Quaintance 1903, by original 

designation. 

Neoaleurolobus Takahashi 1951c: 5. Type species. Aleurolobus musae Corbett 1935, by monotypy; 

synonymy according to Regu & IS 1993:32; Martin & Mound 2007: 13. 

Rositaleyrodes Meganathan & David 1994: 48. Type species. Aleurolobus opilismeni Takahashi 1931, by 

monotypy; synonymy according to Martin & Mound 2007: 13. 

References: Regu & David 1993 - key to Indian species. 



acanthi x 

antennata x 

azadirachtae x 

azimae x 

barleriae x 

barodensis x x x 

bidentatus x 

burliarensis x 

cassiae x 

cephalidistinctus x 

cissampelosae x 

cohici x 

confusus x 

dalbergiae x 

delamarei x 

delhiensis x 

diacritica x 

diastematus x 

distinctus x 

exceptionalis x 

flavus x 

fouabii x 

129


graminicola x 

greeni x 

gruveli x 

hargreavesi x 

hederae x 

hosurensis x 

indigoferae x 

iteae x 

japonicus x 

jullieni x x 

karunkuliensis x 

lagerstroemia x 

longisetosus x 

luci x 

macarangae x 

madrasensis x 

marlatti x x x x x 

mauritanicus x 

moundi x x 

musae x 

nagercoilensis x 

olivinus x x 

onitshae x 

opilismeni x 

orientalis x 

osmanthi x 

ovalis x 

padappaiensis x 

panvelensis x 

patchily x 

130


philippinensis x x 

piliostigmatos x 

psidii x 

rhachisphora x 

rhododendri x x 

riveae 

russellae x 

sairandhryensis x 

saputarensis 

scolopiae x 

selangorensis x x x 

setigerus x x 

shiiae x 

simula x 

singhi x 

solitarius x 

spinosus x 

styraci x 

subrotundus x x 

sundararaji x 

szechwanensis x 

taeonabae x x 

tassellatus x 

tchadiensis x 

teucrii x 

tuberculatus x x 

valparaiensis x 

vitis x x x 

131


walayarensis x 

wunni x 

sp. x x x x 

Total = 81 species? 1 1 7 16 18 57+2 4 0 0 

Aleurolobus acanthi Takahashi 1936 

Aleurolobus acanthi Takahashi 1936a: 52. Syntypes. SW Africa: Swakopmund, on Acanthus sp., TARI. 

Distribution: 4: Africa; Namibia. 

Host: Acanthaceae: Acanthus sp. 

Aleurolobus antennata Regu & David 1993 

Aleurolobus antennata Regu and David 1993: 17. Holotype. India: on Sapindus sp. 


Host: Sapindaceae: Sapindus sp. 

Aleurolobus azadirachtae Regu & David 1993 

Aleurolobus azadirachtae Regu and David 1993: 18. Holotype. India: on Azadirachta indica. 


Host: Meliaceae: Azadirachta indica. 

Aleurolobus azimae Jesudasan & David 1991 

Aleurolobus azimae Jesudasan and David 1991: 269. Holotype pupal case. India: on Azima tetracantha, 

IDAV. 


Host: Salvadoraceae: Azima tetracantha. 

Aleurolobus barleriae Jesudasan & David 1991 

Aleurolobus barleriae Jesudasan and David 1991: 269. Holotype pupal case. India: on Barleria laxifolia, 

IDAV. 


Host: Acanthaceae: Barleria laxifolia. 

Aleurolobus barodensis (Maskell) 1895 

Aleurodes barodensis Maskell 1895: 424. Syntypes. India: Saccharum officinarum, ADSIR. 

Aleurolobus barodensis (Maskell); Quaintance & Baker 1914: 109. 

Aleurodes longicornis Zehntner 1897a: 381. Syntypes. Java: on sugarcane [Saccharum officinarum], ADSIR; 

Quaintance & Baker 1917: 359. 

Distribution: 5: Philippines; Taiwan; 6: India; Pakistan; Malaysia; Thailand; 7: Java. 

Host: Poaceae: Erianthus aurundanaceum, Miscanthus sp., Saccharum officinarum. 

Natural enemies: Fungus: Aschersonia sp. Parasitoids - Aphelinidae: Ablerus aligarhensis, Ablerus delhiensis 

[probably hyperparasitoids], Encarsia isaaci, Encarsia macroptera, Encarsia muliyali, Encarsia 

132

133 

udaipuriensis; Eulophidae: Euderomphale secunda; Platygastridae: Amitus aleurolobi. 

Predators - NEUROPTERA/ Chrysopidae: Chrysoperla carnea; Coleoptera/Coccinellidae: Chilocorus 

nigritus, Oenopia sauzeti, Coelophora octosignata, Coelophora perrotteti, Coelophora unicolor romani, 

Menochilus sexmaculatus, Brumoides suturalis, Scymnus nubilis, Scymnus ochroderus, Scymnus pallidicollis, 

Serangium parcesetosum, Verania discolor. 

Aleurolobus bidentatus Singh 1940 

Aleurolobus bidentatus Singh 1940: 455. Syntypes. India: on Jasminum sp., ZSI 


Host: Oleaceae: Jasminum sp. 

Aleurolobus burliarensis Jesudasan & David 1991 

Aleurolobus burliarensis Jesudasan and David 1991: 271. Holotype pupal case. India: on unidentified tree, 

IDAV. 


Host: on unidentified tree. 

Aleurolobus cassiae Jesudasan & David 1991 

Aleurolobus cassiae Jesudasan and David 1991: 271. Holotype pupal case. India: on Cassia fistula, IDAV. 


Host: Fabaceae: Cassia fistula. 

Aleurolobus cephalidistinctus Regu & David 1993 

Aleurolobus cephalidistinctus Regu and David 1993: 20. Holotype. India: on Hemidesmus indicus. 


Host: Asclepiadaceae: Hemidesmus indicus. 

Aleurolobus cissampelosae Regu & David 1993 

Aleurolobus cissampelosae Regu and David 1993: 21. Holotype. India: on Cissampelos sp. 


Host: Menispermaceae: Cissampelos sp. 

Aleurolobus cohici Regu & David 1993 

Aleurolobus cohici Regu and David 1993: 22. Holotype. India: on unknown host. 



Aleurolobus confusus David & Subramaniam 1976 

Aleurolobus confusus David & Subramaniam 1976: 160. Holotype. India: on Murraya koenigii, ZSI. 


Host: Rutaceae: Murraya koenigii. 

US Port Interceptions: India on Murraya koenigii, Murraya sp.; Sri Lanka on Citrus sp., Murraya koenigii, 

Murraya sp.

Aleurolobus dalbergiae Dubey & Sundararaj 

Aleurolobus dalbergiae Dubey & Sundararaj 2006: 42-43. Holotype. India: on Dalbergia latifolia, 22.ii.2001, 

A. K. Dubey, FRI. 


Host: Fabaceae: Dalbergia latifolia. 

Aleurolobus delamarei Cohic 1969 

Aleurolobus delamarei Cohic 1969: 26. Holotype. Chad: on Balanites aegyptiaca. 


Host: Zygophyllaceae: Balanites aegyptiaca. 

Aleurolobus delhiensis Regu & David 1993 

Aleurolobus delhiensis Regu and David 1993: 23. Holotype. India: on Kigelia pinnata. 


Host: Bignoniaceae: Kigelia pinnata. 

Aleurolobus diacritica Regu & David 1993 

Aleurolobus diacritica Regu and David 1993: 24. Holotype. India: on unknown host. 


Host: unknown. 

Aleurolobus diastematus Bink-Moenen 1983 

Aleurolobus diastematus Bink-Moenen 1983: 44. Holotype pupal case, Chad, Bebedjia, 23.xi.1973, FAB & 

RMBM, on Guiera senegalensis, BMNH. 


Host: Combretaceae: Guiera senegalensis. 

Aleurolobus distinctus Regu & David 1993 

Aleurolobus distinctus Regu and David 1993: 24. Holotype. India: on Sapindus sp. 


Host: Sapindaceae: Sapindus sp. 

Aleurolobus exceptionalis Regu & David 1993 

Aleurolobus exceptionalis Regu and David 1993: 25. Holotype. India: on Ficus beddomei. 


Host: Moraceae: Ficus beddomei. 

Aleurolobus flavus Quaintance & Baker 1917 

Aleurolobus flavus Quaintance & Baker 1917: 360. Syntypes. Sri Lanka: on unidentified tree, USNM. 


Host: on unidentified tree; Loranthaceae: Loranthus sp. 

134

Aleurolobus fouabii Cohic 1969 

Aleurolobus fouabii Cohic 1969: 32. Holotype. Congo: on Agelaea dewevei, CORSTOM. 


Host: Connaraceae: Agelaea dewevrei; Rubiaceae: Colletoecema dewevrei. 

Aleurolobus graminicola Bink-Moenen 1983 

Aleurolobus graminicola Bink-Moenen 1983: 46. Holotype pupal case, Chad, Baibokoum, 8.xii.1973, FAB & 

RMBM, on Cymbopogon giganteus, BMNH. 


Host: Poaceae: Cymbopogon giganteus. 

Aleurolobus greeni Corbett 1926 

Aleurolobus greeni Corbett 1926: 280. Syntypes. Sri Lanka: Peradeniya, xi.1905, E. E. Green, on unidentified 

shrub, BMNH. 



Aleurolobus gruveli Cohic 1968 

Aleurolobus gruveli Cohic 1968b: 88. Syntypes (21): Cameroon: on Graminae (Poaceae)? 

Distribution: 4: Camaroon; Chad. 

Host: Poaceae: Coelorachis afraurita. 

Aleurolobus hargreavesi Dozier 1934 

Aleurolobus hargreavesi Dozier 1934: 185. Syntypes. Uganda: on Hyparrhenia sp., USNM. 

Distribution: 4: Congo; Nigeria; Uganda. 

Host: Poaeae: Andropogon diplandra, Hyparrhenia sp., Saccharum officinarum. 

Aleurolobus hederae Takahashi 1935 

Aleurolobus hederae Takahashi 1935b: 63. Syntypes. Taiwan; on Hedera formosana, TARI. 


Host: Araliaceae: Hedera formosana. 

Aleurolobus hosurensis Regu & David 1993 

Aleurolobus hosurensis Regu and David 1993: 26. Holotype. India: on an unidentified plant. 



Aleurolobus indigoferae Regu & David 1993 

Aleurolobus indigoferae Regu and David 1993: 27. Holotype. India: on Indigofera oblongifolia. 


Host: Fabaceae: Indigofera oblongifolia. 

135

136 

Aleurolobus iteae Takahashi 1957 

Aleurolobus iteae Takahashi 1957: 13. Syntypes. Japan: on Itea japonica, HBL; Suh et al. 2006. 


Host: Saxifragaceae: Itea japonica; Scrophulariaceae: Melampyrum roseum. 

Aleurolobus japonicus Takahashi 1954 

Aleurolobus japonicus Takahashi 1954a: 3. Syntypes. Japan: on Callicarpa scandens, USNM. 

Distribution: 5: China, Japan. 

Host: Asteraceae: Pertya scandens; Caprifoliaceae: Lonicera sp.; Ranunculaceae: Clematis sp.; Rosaceae: 

Rubus palmatus;Verbenaceae: Callicarpa japonica. 


Aleurolobus jullieni Cohic 1968 

Aleurolobus juillieni Cohic 1968b: 92. Syntypes. Congo: on Clerodendron thomsonae. 

Distribution: 4: Cameroon; Congo. 

Host: Capparidaceae: Buchholzia coriaceae; Verbenaceae: Clerodendrum thomsoniae 

Aleurolobus karunkuliensis Jesudasan & David 1991 

Aleurolobus karunkuliensis Jesudasan and David 1991: 272. Holotype pupal case. India: on unidentified 

shrub, IDAV. 



Aleurolobus lagerstroemia Regu & David 1993 

Aleurolobus lagerstroemia Regu and David 1993: 28. Holotype. India: on Lagerstroemia lanceolata. 


Host: Lythraceae: Lagerstroemia lanceolata. 

Aleurolobus longisetosus Dubey & Sundararaj 

Aleurolobus longisetosus Dubey & Sundararaj 2006. 45. Holotype. India: on Michelia champaca, 27.iv.2001, 

A. K. Dubey, FRI. 


Host: Magnoliaceae: Michelia champaca. 

Aleurolobus luci Cohic 1969 

Aleurolobus luci Cohic 1969: 36. Syntypes. Ivory Coast: on Gardenia ternifolia. 

Distribution: 4: Chad; Ivory Coast. 

Host: Rubiaceae: Gardenia triacantha, Gardenia ternifolia. 

Aleurolobus macarangae Regu & David 1993 

Aleurolobus macarangae Regu and David 1993: 28. Holotype. India: on Macaranga sp. 


Host: Euphorbiaceae: Macaranga sp.

137 

Aleurolobus madrasensis Regu & David 1993 

Aleurolobus madrasensis Regu and David 1993: 29. Holotype. India: on Atalantia monophylla. 


Host: Rutaceae: Atalantia monophylla. 

Aleurolobus marlatti (Quaintance) 1903 

Aleurodes marlatti Quaintance 1903: 61. Lectotype (designated Martin 1999:43). Japan: on orange [Citrus 

sp..], USNM. 

Aleurolobus marlatti (Quaintance); Quaintance & Baker 1914: 109. 

Aleurolobus niloticus Priesner & Hosny 1934a: 1. Syntypes. Egypt: on Zizyphus spina christi, USNM; Bink- 

Moenen 1983: 50; synonymy according to Martin 1999: 43. 

Aleurolobus ravisei Cohic 1968b: 95. Syntypes (2). Congo: on Hymenocardia acida, CORSTOM; 

synonymized with A. niloticus by Bink-Moenen 1983: 50. 

Distribution: 3: Egypt; Iran; Israel; Jordan; Saudi Arabia; 4: Chad; 5: China; Japan; Philippines; Taiwan; 6: 

India; Malaysia; 7: Java. 

Host: Amaranthaceae: Amaranthus bilitoides; Apocynaceae: Nerium indicum; Araceae: Colocasia sp.; 

Araliaceae: Hedera nepalensis; Asclepiadaceae: Leptadenia hastata; Bignoniaceae: Steroespermum 

kunthianum; Bombacaceae: Bombax malabaricum; Boraginaceae: Ehretia aspersa; Capparidaceae: 

Boscia senegalensis, Capparidaceae: Capparis corymbosa; Daphniphyllaceae: Daphniphyllum 

macropodum; Ebenaceae: Diospyros mespiliformis; Hymenocardia acida, Glochidion hongkongensis, 

Mallotus philippensis; Fabaceae: Dalbergia sissoo; Liliaceae: Smilax sp.; Lythraceae: Lawsonia alba, 

Lawsonia inermis; Malvaceae: Abutilon foliosum, Malvaceae: Thespesia populnea; Moraceae: Ficus 

sycomorus, Ficus sp., Morus alba; Myrtaceae: Eucalyptus camalduleis; Oleaceae: Olea cuspidata; 

Punicaceae: Punica granatum; Rhamnaceae: Paliurus spina-christi, Ziziphus jujube, Ziziphus 

hysudrica, Ziziphus mauritiana, Ziziphus spina-christi, Ziziphus sp.; Rosaceae: Rosa indica; Rutaceae: 

Citrus limmeta, Citrus nobilis, Citrus sinensis, Murraya exotica, Murraya paniculata, Murraya 

koenigii; Salvadoraceae: Salvadora persicae; Sapindaceae: Dodonaea viscosa, Schleichera oleosa; 

Ulmaceae: Aphananthe aspera; Verbenaceae: Duranta sp., Verbena officinalis, Gmelina sp.; 


Natural enemies: Parasitoids - Aphelinidae: Encarsia perflava, Encarsia longifasciata, Encarsia davidi, 

Encarsia elegans, Encarsia ancistrocera, Encarsia lutea, Eretmocerus aleurolobi, Eretmocerus 

siphonini, Eretmocerus longiscapus, Eretmocerus parasiphonini, Eretmocerus sp. Predators - 

Coccinellidae: Brumoides suturalis. 

US Port Interceptions: Australia on Eucalyptus sp.; Bangladesh on Citrus limon, Citrus sp., Murraya 

koenigii, Murraya sp.; Cambodia on Murraya koenigii; China on Citrus sp., Murraya koenigii, 

Murraya sp.; Dominican Republic on Murraya koenigii; Egypt on unknown plant; Germany on 

Murraya koenigii; Greece on Citrus limon; Haiti on Citrus sp.; Hong Kong on Clausena lansium; 

India on Alstroemeria sp, Azadirachta indica, Citrus aurantiifolia, Citrus limon, Citrus sinensis, 

Citrus sp., Delphinium sp, Euphorbia lactea, Lawsonia inermis, Liatris sp., Mangifera indica, Mentha 

sp., Murraya koenigii, Murraya paniculata, Murraya sp, Musa acuminata, Musa sp., Ptelea sp., 

Rutaceae; Indonesia on Murraya sp., Rutaceae; Iran on Citrus sp., Mentha sp., Murraya koenigii, 

Murraya sp.; Israel on Alstroemeria sp., Rutaceae; Italy on Murraya koenigii, Jamaica on Murraya 

koenigii; Japan on Murraya sp.; Jordan on Citrus aurantiifolia; Korea (South) on Citrus sp., 

Psidium guajava; Malaysia on Citrus sp., Murraya koenigii, Murraya sp.; Mexico on Citrus sinensis; 

Nigeria on Abelmoschus esculentus, Murraya koenigii; Pakistan on Citrus sp., Murraya koenigii,

138 

Murraya paniculata, Murraya sp.; Philippines Citrus sp., Mangifera indica, Manilkara zapota, 

Murraya sp., Psidium guajava, Psidium sp.; Saudi Arabia on Citrus sp., Murraya koenigii; 

Seychelles on Murraya koenigii; Singapore on Mangifera indica, Murraya koenigii, Murraya sp.; 

South Africa on Murraya koenigii; Sri Lanka on Murraya koenigii, Murraya sp.; Taiwan on Citrus 

reticulata, Murraya koenigii, Murraya sp.; Thailand on Citrus aurantiifolia, Citrus sp., Murraya 

koenigii, Murraya sp.; Trinidad and Tobago on Murraya koenigi; United Arab Emirates on 

Murraya koenigii, Murraya sp.; Venezuela on Citrus sp.; Viet Nam on Eucalyptus sp., Murraya 

koenigii, Murraya sp., Ochna sp., Psidium guajava. 

Aleurolobus mauritanicus Cohic 1969 

Aleurolobus mauritanicus Cohic 1969: 41. Holotype. Mauritania: on Boscia senegalensis. 

Distribution: 4: Chad; Mauritania. 

Host: Capparidaceae: Boscia senegalensis, Ritchiea afzelii. 

Aleurolobus moundi David & Subramaniam 1976 

Aleurolobus moundi David & Subramaniam 1976: 161. Holotype. India: on Bassia sp., ZSI. 

Distribution: 3: Iran; 6: India. 

Host: Chenopodiaceae: Bassia sp. (Madhuca sp.); Euphorbiaceae: Euphorbia glomerifera; Rutaceae: Citrus 

limmeta, Citrus nobilis. 

Aleurolobus musae Corbett 1935 

Aleurolobus musae Corbett 1935b: 820. Holotype. Malaya: on Musa sapientum. 

Neoaleurolobus musae (Corbett); Takahashi 1951c: 6; Mound & Halsey 1978: 171. 

Aleurolobus musae Corbett; Regu & David 1993: 32. 

Distribution: 6: India; Malaya; Malaysia. 


Aleurolobus nagercoilensis Regu & David 1993 

Aleurolobus nagercoilensis Regu and David 1993: 32. Holotype. India: on Barleria cuspidata. 


Host: Acanthaceae: Barleria cuspidata. 

Aleurolobus olivinus (Silvestri) 1911 

Aleurodes olivinus Silvestri 1911: 214. Syntypes. Italy: on Olea sp., IESP. 

Aleurolobus olivinus (Silvestri); Quaintance & Baker 1915a: xi. 

Distribution: 3: Cyperus; Egypt; France; Israel; Italy; Morocco; Spain; 5: China. 

Host: Oleaceae: Olea sp., Phillyrea angustifolia. 

Natural enemies: Aphelinidae: Encarsia arabica, Encarsia elegans, Encarsia olivina; 

Platygastridae: Amitus minervae 

US Port Interceptions: Cyprus on Olea europaea, Olea sp.; Greece on Olea europaea, Olea sp.; Iran on 

Olea europaea; Israel on Olea europaea, Olea sp.; Italy on Olea europaea, Olea sp.; Jordan on Olea 

europaea; Lebanon on Olea europaea; Morocco on Olea europaea; Spain on Olea europaea; Syria 

on unknown plant. 

Aleurolobus onitshae Mound 1965

139 

Aleurolobus onitshae Mound 1965c: 128. Holotype. Nigeria: on Phyllanthus floribundus, BMNH. 

Distribution: 4: Chad; Ivory Coast; Nigeria. 

Host: Euphorbiacae: Flueggea virosa, Phyllanthus floribundus, Securinega virosa. 

Aleurolobus oplismeni Takahashi 1931 

Aleurolobus oplismeni Takahashi 1931b: 261. Syntypes. Taiwan: on Oplismenus compositus. 


Host: Poaceae: Oplismenus compositus. 

Aleurolobus orientalis David & Jesudasan 1988 

Aleurolobus orientalis David & Jesudasan Alexander 1988b. Holotype pupal case. India: on Securinega 

virosa, IDAV. 


Host: Euphorbiacae: Securinega virosa; Sapindaceae: Harpullia pendula. 

Aleurolobus osmanthi Young 1944 

Aleurolobus osmanthi Young 1944: 134. Syntypes. China: on Osmanthus fragrans. 


Host: Oleaceae: Osmanthus fragrans. 

Aleurolobus ovalis Regu & David 1993 

Aleurolobus ovalis Regu and David 1993: 38. Holotype. India: on Carissa sp. 


Host: Apocynaceae: Carissa sp. 

Aleurolobus padappaiensis Regu & David 1993 

Aleurolobus padappaiensis Regu and David 1993: 39. Holotype. India: on Guazuma tomentosa. 


Host: Sterculiaceae: Guazuma tomentosa. 

Aleurolobus panvelensis Regu & David 1993 

Aleurolobus panvelensis Regu and David 1993: 40. Holotype. India: on an unidentified plant. 



Aleurolobus patchily Regu & David 1993 

Aleurolobus patchily Regu and David 1993: 41. Holotype. India: on Bridelia squamosa. 


Host: Asteraceae: Brickellia squamulosa. 

Aleurolobus philippinensis Quaintance & Baker 1917 

Aleurolobus philippinensis Quaintance & Baker 1917: 369. Syntypes. Philippines: on unidentified tree, 

USNM.. 

Distribution: 5: Philippines; Taiwan. 

Host: Bombacaceae: Bombax malabaricum; Myrsinaceae: Ardisia sieboldii; Rubiaceae: Gardenia florida;

Salicaceae: Salix sp. 

Aleurolobus piliostigmatos Bink-Moenen 1983 

Aleurolobus piliostigmatos Bink-Moenen 1983: 55. Holotype pupal case, Chad, Bebedjia, 27.vi.1973, FAB & 

RMBM, on Piliostigma thonningii; BHNM. 


Host: Fabaceae: Piliostigma thonningii. 

Aleurolobus psidii Jesudasan & David 1991 

Aleurolobus psidii Jesudasan and David 1991: 275. Holotype pupal case. India: on Psidium guajava, IDAV. 



Aleurolobus rhachisphora Regu & David 1993 

Aleurolobus rhachisphora Regu and David 1993: 42. Holotype. India: on Loranthus sp. 


Host: Loranthaceae: Loranthus sp. 

Aleurolobus rhododendri Takahashi 1934 

Aleurolobus rhododendri Takahashi 1934b: 62. Syntypes. Taiwan: on Rhododendron sp, TARI. 

Distribution: 5: China; Japan; Taiwan; 6: Thailand. 

Host: Ericaceae: Rhododendron formosanum, Rhododendron sp, Myrtaceae: Psidium guajava. 

Rutaceae: Murraya paniculata. 

Natural enemies: Aphelinidae: Encarsia azimi, Encarsia elegans, Encarsia inaron, Encarsia longifasciata, 

Encarsia lutea. 

Aleurolobus riveae Regu & David 1993 

Aleurolobus riveae Regu and David 1993: 42. Holotype. India: on Rivea hypocrateriformis. 


Host: Convolvulaceae: Rivea hypocrateriformis 

Aleurolobus russellae Regu & David 1993 

Aleurolobus russellae Regu and David 1993: 43. Holotype. India: on an unidentified plant. 



Aleurolobus sairandhryensis Regu & David 1993 

Aleurolobus sairandhryensis Regu and David 1993: 23. Holotype. India: on Acanthopippium sp. 


Host: Caryophyllaceae: Acanthophyllum sp. 

Aleurolobus saputarensis Regu & David 1993 

Aleurolobus saputarensis Regu and David 1993: 45. Holotype. India: unidentified plant. 



140

Aleurolobus scolopiae Takahashi 1933 

Aleurolobus scolopiae Takahashi 1933: 19. Syntypes. Taiwan: on Scolopia oldhami, TARI. 


Host: Solanaceae: Scopolia oldhami. 

Aleurolobus selangorensis Corbett 1935 

Aleurolobus selangorensis Corbett 1935b: 819. Syntypes. Malaya: on unidentified host; Martin 1985: 317. 

Distribution: 3: Iran; 6: Malaya; 7: Papau New Guinea. 

Host: Apocynaceae: Cerbera manghas?; Erythroxylaceae: Erythroxylum sp.; Fabaceae: Dalbergia sp., 

Glycyrrhiza glabra; Naucleaceae: Neonauclea sp.; Punicaceae: Punica granatum; Rhizophoraceae: 

Rhizophora stylosa.; Rutaceae: Citrus limetta, Citrus medica. 

Aleurolobus setigerus Quaintance & Baker 1917 

Aleurolobus setigerus Quaintance & Baker 1917: 372. Lectotype (designated by Martin 1999: 35). Sri Lanka; 

on Harpullia pendula, USNM. 

Distribution: 5: China; Hong Kong; Taiwan; 6: Sri Lanka; Thailand. 

Host: Myrtaceae: Psidium guajava, Rhodomyrtus sp.; Rutaceae: Citrus sp.; Sapindaceae: Harpullia pendula. 

Natural enemies: Aphelinidae: Encarsia longifasciata, Encarsia lutea. 

Aleurolobus shiiae Takahashi 1957 

Aleurolobus shiiae Takahashi 1957: 14. Syntypes. Japan: on Castanopsis [=Shiia] cuspidata, HBL. 


Host: Fagaceae: Castanopsis cuspidata. 

Aleurolobus singhi Regu & David 1993 

Aleurolobus singhi Regu and David 1993: 46. Holotype. India: unidentified plant. 



Aleurolobus solitarius Quaintance & Baker 1917 

Aleurolobus solitarius Quaintance & Baker 1917: 377. Syntypes. USA: Illinois, on Cercis canadensis, 

USNM. 

Distribution: 1: USA (FL, IL, VA). 

Host: Fabaceae: Cercis canadensis. 

Aleurolobus spinosus Jesudasan & David 1991 

Aleurolobus spinosus Jesudasan and David 1991: 276. Holotype pupal case. India: on Semecarpus 

anacardium, IDAV. 


Host: Anacardiaceae: Semecarpus anacardium. 

Aleurolobus styraci Takahashi 1954 

Aleurolobus styraci Takahashi 1954: 4. Syntypes. Japan: on Styrax japonica, HBL. 


141

Host: Styracaceae: Styrax japonicus. 

Aleurolobus subrotundus Silvestri 1927 

Aleurolobus subrotundus Silvestri 1927a: 13. Syntypes. Vietnam and China: on Citrus sp. and Clausenia sp., 

IESP; Takahashi 1942c: 208. 

Distribution: 5: China; 6: Vietnam. 

Host: Rutaceae: Citrus sp., Clausena sp., Murraya sp. 

Natural enemies: Aphelinidae: Encarsia armata, Encarsia strenua 

US Port Interceptions: Australia on Euonymus sp.; China on Euonymus japonicus; Viet Nam on Murraya 

sp. 

Aleurolobus sundararaji Regu & David 1993 

Aleurolobus sundararaji Regu and David 1993: 47. Holotype. India: on Tectona grandis. 


Host: Verbenaceae: Tectona grandis. 

Aleurolobus szechwanensis Young 1942 

Aleurolobus szechwanensis Young 1942: 99. Syntypes. China: on Citrus sp. 



Natural enemies: Aphelinidae: Encarsia fujianensis. 

Aleurolobus taeonabae (Kuwana) 1911 

Aleyrodes taeonabae Kuwana 1911: 623. Syntypes. Japan: on Taonabo japonica, USNM. 

Aleurolobus taeonabae (Kuwana); Quaintance & Baker 1914: 109. 

Aleurolobus chinensis Takahashi 1936e: 453. Syntypes. China: on Cercis chinensis, TARI; synonymy 

according to Takahashi 1954: 2. 

Distribution: 5: China; Japan; 6: India. 

Host: Euphorbiaceae: Mallotus japonicus; Fabaceae: Cercis chinensis; Pittosporaceae: Pittosporum 

tobira; Theaceae: Ternstroemia japonica; Vitaceae: Vitis vinifera. 

Aleurolobus tassellatus Regu & David 1993 

Aleurolobus tassellatus Regu and David 1993: 48. Holotype. India: on Homonoia riparia. 



Aleurolobus tchadiensis Bink-Moenen 1983 

Aleurolobus tchadiensis Bink-Moenen 1983: 57. Holotype pupal case, Chad, Bebedjia, 17.xi.1973, FAB & 

RMBM, on Ritchiea afzelii; BHMN. 


Host: Capparaceae: Ritchiea afzelii, Capparus corymbosa. 

Aleurolobus teucrii Mifsud and Palmeri 1996 

Aleurolobus teucrii Mifsud & Palmeri 1996: 89. Holotype pupal case. Maltese Islands: Gozo, on Teucrium 

fruticans, NHM. 

142

Distribution: 3: Maltese Islands. 

Host: Labiatae: Teucrium fruticans. 

Aleurolobus tuberculatus Regu & David 1993 

Aleurolobus tuberculatus Regu and David 1993: 49. Holotype. India: on Ailanthus excelsa. 

Distribution: 4: Madagascar; 6: Burma; Cambodia; India; Thailand. 


Aleurolobus valparaiensis Jesudasan & David 1991 

Aleurolobus valparaiensis Jesudasan and David 1991: 278. Holotype pupal case. India: on unidentified tree, 

IDAV. 



Aleurolobus vitis Danzig 1966 

Aleurolobus vitis Danzig 1966: 205 [380]. Holotype. USSR: southern Maritime, on Vitis amurensis, ZIN; Suh 

et al. 2006: 

Distribution: 3: Iran; 5: Korea, Maritime Territory (USSR); 6: Malaya; Malaysia. 

Host: Vitaceae: Vitis amurensis, Vitis vinifera. 

Natural enemies: Aphelinidae: Eretmocerus longiscapus. 

Aleurolobus walayarensis Jesudasan & David 1991 

Aleurolobus walayarensis Jesudasan & David 1991: 278. Holotype pupal case. India: Kerala, Walayar, 

28.iii.1973, B. V. David, on unidentified tree. 



Aleurolobus wunni (Ryberg) 1938 

Aleurodes asari Wunn 1926: 28. Syntypes. France: on Asarum europaeum; name preoccupied by Aleurodes 

asari Schrank, 1801. 

Aleurolobus asari (Wunn); Visnya 1941b: 4. 

Aleurodes wunni Ryberg 1938: 28; replacement name for A. asari. 

Aleurolobus wunni (Ryberg); Mound and Halsey 1978: 39. 

Aleurolobus clematidis Goux 1942: 141. Paratypes. France: on Clematis vitalba; synonomy according to 

Zahradnik 1963a: 10. 

Aleurolobus puripennis Ossiannilsson 1944: 188. Syntypes. Sweden: on Symphoricarpus albus; synonomy 

according to Zahradnik 1963a: 10. 

Distribution: 3: Czech Republic; Finland; France; Germany; Hungary; Iran, Italy; Poland; Sweden; USSR; 

Yugoslavia. 

Host: Aristolochiaceae: Asarum europaeum; Caprifoliaceae: Linnaea borealis, Lonicera fragrantissima, 

Symphoricarpos albus, Symphoricarpos racemosus; Rosaceae: Spiraea sp. 

Labiatae: Phlomis sp, Phlomis fruticosa; Ranunculaceae: Clematis vitalba, Cimicifuga daurica; 

Theaceae: Camellia japonica. 

Natural enemies: Aphelinidae: Encarsia maritima, Encarsia lutea, Encarsia olivina, Encarsia lehri, Encarsia 

tricolor; Eretmocerus longicornis; Platygastridae: Amitus longicornis; Platygastridae: Amitus minervae. 

143

US Port Interception: India on Murraya koenigii. 

Aleurolobus sp. 

1. India, Aphelinidae: Encarsia albiscutellum longipalpa (ref: Hayat 1989a: 34). 

2. Kenya, Aphelinidae: Encarsia gerlingi (ref: Huang & Polaszek 1998). 

3. Kenya, on Boraginaceae: Ehretia pitiolaris, Aphelinidae: Encarsia gerlingi (ref: Viggiani 1988). 

4. Japan, on Ericaceae: Rhododendron sp., Aphelinidae: Encarsia longifasciata (ref: Pedata & Polaszek 2003: 

371). 

5. Australia, on Myrtaceae: Eucalyptus bigalerita, Aphelinidae: Encarsia protransvena, Aleurolobus sp. (ref: 

Schmidt et al. 2001: 381). 

6: India, on Myrtaceae: Eugenia jambolana, Aphelinidae: Encarsia septentrionalis (ref: Hayat 1989a: 68). 

7. India, Aphelinidae: Encarsia trivittata (ref: Hayat 1989a: 59). 

8. India, Eulophidae: Pomphale striptipennis (ref: Husain et al. 1983). 

Transfers 

Aleurolobus opilismeni Takahashi 1931 - Rositaleyrodes 

Aleurolobus zeylanicus Corbett 1926 - Orientaleyrodes David 1993 

144

Aleurolonga Mound 1965 

Aleurolonga Mound 1965c: 130. Type species. Aleurolonga cassiae Mound 1965, by monotypy. 



cassiae x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Aleurolonga cassiae Mound 1965 

Aleurolonga cassiae Mound 1965c: 130. Holotype. Nigeria: on Cassia siamea, BMNH; Cohic 1968a: 24; 

Bink-Moenen 1983. 

Distribution: 4: Angola; Chad; Nigeria. 

Host: Fabaceae: Brachystegia sp., Burkea africana, Cassia siamea, Detarium microcarpum, Entada africana. 

145

Aleuromarginatus Corbett 1935 

Aleuromarginatus Corbett 1935c: 246. Type species. Aleuromarginatus tephrosiae Corbett 1935, by 

monotypy. 

Host - only known from legumes (Fabaceae). 



bauhiniae x 

corbettiaformis x 

dalbergiae x 

kallarensis x 

littoralis x 

marginiquus x 

millettiae x 

moundi x 

nemciae x 

nigrus x 

serdangensis x 

shihmenensis x 

tephrosiae x x 

thirumurthiensis x 

sp. x 

Total = 14 species 0 0 1 3 1 4+ 6 0 0 

Aleuromarginatus bauhinae (Corbett ) 1935 

Trialeurodes bauhinae Corbett 1935b: 816. Syntypes. Malaya: on Bauhinia bidentata. 

Aleuromarginatus bauhinae (Corbett ); David 1988. 

Distribution: 6: Malaya; Malaysia, India (new record Dubey et al). 

Host: Fabaceae: Bauhinia bidentata. 

Aleuromarginatus corbettiaformis Martin 1985 

Aleuromarginatus corbettiaformis Martin 1985: 318. Holotype pupal case. New Guinea: on Desmodium 

unbellatum, BMNH. 


Host: Fabaceae: Desmodium umbellatum. 

146

147 

Aleuromarginatus dalbergiae Cohic 1969 

Aleuromarginatus dalbergiae Cohic 1969: 62. Holotype. Ivory Coast: on Dalbergia saxatilis; Bink-Moenen 

1983: 60. 


Host: Fabaceae: Andira inermis; Pterocarpus lucens; Fabaceae: Dalbergia saxatilis; Pterocarpus lucens 

Aleuromarginatus kallarensis David & Subramaniam 1976 

Aleuromarginatus kallarensis David & Subramaniam 1976: 162. Holotype. India: on Pterolobium indicum, 

ZSI 


Host: Fabaceae: Pongamia glabra; Pterolobium indicum. 

Aleuromarginatus littoralis Martin 1985 

Aleuromarginatus littoralis Martin 1985: 319. Holotype pupal case. New Guinea: on ?Derris trifoliata, 

BMNH 


Host: Fabaceae: Derris trifoliata. 

Aleuromarginatus marginiquus Martin 1999 

Aleuromarginatus marginiquus Martin 1999: 45. Holotype puparium. Australia: on Daviesia horrida, ANIC. 


Host: Fabaceae: Daviesia horrida. 

Aleuromarginatus millettiae Cohic 1968 

Aleuromarginatus millettiae Cohic 1968a 28. Syntypes. Congo: on Millettia eetveldeana? Bink-Moenen 1983: 

61. 


Host: Fabaceae: Burkea africana; Cathormion altissimum; Daniellia olivera; Millettia eetveldeana; Newtonia 

glandulifera; Pterocarpus lucens; Tamarindus indica. 

Aleuromarginatus moundi Martin 1999 

Aleuromarginatus moundi Martin 1999: 46. Holotype puparium. Australia: on Acacia ?aneura, ANIC. 

Dstribution: 7: Australia. 

Fabaceae: Acacia aneura. 

Aleuromarginatus nemciae Martin 1999 

Aleuromarginatus nemciae Martin 1999: 46. Holotype puparium. Australia: on Nembia sp, ANIC. 


Host: Nembia sp. 

Aleuromarginatus nigrus Martin 1999 

Aleuromarginatus nigrus Martin 1999: 47. Holotype puparium. Australia: on Oxylobium ilicifolium, ANIC. 


Host: Fabaceae: Oxylobium ilicifolium.

Aleuromarginatus serdangensis Takahashi 1955 

Aleuromarginatus serdangensis Takahashi 1955, in Takahashi 1955b: 223. Syntypes (2). Malaya: on 

undetermined wild tree, BMNH. 


Host: undetermined wild tree. 

Aleuromarginatus shihmensensis Ko 1995 

Aleuromarginatus shihmensensis Ko 1995, in Ko et al. 1995: 893. Holotype puparium, Taiwan, Shihmen, 23 I 

1987, C.C. Ko, ex. Millettia reticulata, NTU. 

Distribution: Taiwan. 

Host: Fabaceae: Millettia reticulata. 

Aleuromarginatus tephrosiae Corbett 1935 

Aleuromarginatus tephrosiae Corbett 1935c: 247. Syntypes. Sierra Leone: on Tephrosia candida, 

Cohic 1969: 61; Bink-Moenen 1983: 62. 

Aleuromarginatus indica Singh 1940: 454. Syntypes. India: on Tephrosia purpurea, ZSI; synonymy 

according to David & Subramaniam 1976: 163. 

Distribution: 3: Egypt; 4: Chad; Congo; Ivory Coast; Nigeria; Sierra Leon; Zaire; 6: India. 

Hosts: Fabaceae: Indigofera sp.; Parkia clappertoniana; Prosopis africana; Tephrosiae linearis; 

Tephrosia purpurea; Tephrosia vogelii; Tephrosia candida. 

Aleuromarginatus thirumurthiensis David 1988 

Aleuromarginatus bauhiniae David 1976: 85. Holotype. India: Thirumurthi Hills, 7.6.1975, B.V. David, on 

Bauhinia racemosa, ZSI 

Aleuromarginatus thirumurthiensis David 1988; name preoccupied by Aleuromarginatus bauhiniae Corbett 

1935. 


Host: Fabaceae: Bauhinia racemosa. 

Fabaceae: Desmodium sp. 

Distribution: 7: Sulawesi 

Reference: Martin 1988: 65 

Aleuromarginatus sp. 

Changes 

Aleuromarginatus bauhiniae David 1976 - Aleuromarginatus thirumurthiensis David 1988, n. name 

148

Aleuropapillatus Regu and David 1993 

Aleuropapillatus Regu and David 1993: 50. Type species. Aleuropapillatus kumariensis Regu & David 1993, 

by original designation. 



gmelinae x 

kumariensis x 

Total = 2 species 0 0 0 0 0 2 0 0 0 

Aleuropapillatus gmelinae (David, Jesudasan & Mathew) 1988 

Aleurolobus gmelinae David, Jesudasan & Mathew 1988: 165. Holotype puparium, India, Kerala State, 

Peechi, 29.x.1986, G. Mathew, on Gmelina arborea, David Collection (paratypes in IARI, USNM, 

BMNH). 

Aleuropapillatus gmelinae (David, Jesudasan & Mathew); Regu & David 1993: 51. 


Host: Verbenaceae: Gmelina arborea. 

Aleuropapillatus kumariensis Regu & David 1993 

Aleuropapillatus kumariensis Regu & David 1993: 50. Holotype puparium, India, on Gmelina asiatica. 


Host: Verbenaceae: Gmelina asiatica. 

149

Aleuroparadoxus Quaintance & Baker 1914 

Aleuroparadoxus Quaintance & Baker 1914: 105. Type species Aleyrodes iridescens Bemis 1904, by 

monotypy. 

References: Russell (1947) revision; Martin 2005. 



arctostaphyli x 

chomeliae x 

gardeniae x x 

ilicicola x x 

iridescens x x 

punctatus 

rhodae x 

sapotae x 

trinidadensis x 

truncatus x 

Total = 10 species 4 8 0 0 0 0 0 0 0 

Aleuroparadoxus arctostaphyli Russell 1947 

Aleuroparadoxus arctostaphyli Russell 1947: 15. Holotype. USA: California, on Artostaphylos sp., USNM 

Distribution: 1: USA (CA, OR). 

Host: Ericaceae: Arctostaphylos manzanita, Arctostaphylos tomentosa, Arctostaphylos viscida, Arbutus 

menziesii. 

Aleuroparadoxus chomeliae Russell 1947 

Aleuroparadoxus chomeliae Russell 1947: 22. Holotype. Panama: on Chomelia spinosa, USNM. 


Host: Rubiaceae: Chomelia spinosa. 

Aleuroparadoxus gardeniae Russell 1947 

Aleuroparadoxus gardeniae Russell 1947: 18. Holotype. Mexico: on Gardenia sp, USNM. 

Distribution: 1: USA (TX); 2: Costa Rica, Cuba, El Salvador, Mexico. 

Host: Ebenaceae: Diospyros sp.; Rubiaceae: Gardenia sp. 

Aleuroparadoxus ilicicola Russell 1947 

Aleuroparadoxus ilicicola Russell 1947: 19. Holotype. USA: Alabama, on Ilex sp., USNM. 

Distribution: 1: USA (AL); 2: Mexico. 

150

Host: Aquifoliaceae: Ilex vomitoria, Aquifoliaceae: Ilex sp. 

Aleuroparadoxus iridescens (Bemis) 1904 

Aleuroparadoxus iridescens (Bemis) 1904. Lectotype (designated by Russell 1947:14). USA: California, on 

Photinia (Heteromeles) arbutifolia, USNM; Quaintance & Baker 1914: 104 

Distribution: 1: USA (CA); 2: Mexico. 

Host: Rubiaceae: Gardenia sp; Ericaceae: Arctostaphylos manzanita; Labiatae: Salvia sp.; Lauraceae: 

Umbellularia californica; Rhamnaceae: Rhamnus californicus, Rhamnus creocea, Rhamnus ilicicola; 

Rosaceae: Heteromeles arbutifolia. 

Aleuroparadoxus punctatus Quaintance & Baker 1917 

Aleuroparadoxus punctatus Quaintance & Baker 1917: 3870. Lectotype (designated by Russell 1947: 35). 

Chile: on Lithraea caustica, USNM; Russell 1947: 35. 

Distribution: 2: Chile. 

Host: Anacardiaceae: Lithrea caustica, Schinus sp. Flacourtiaceae: Azara sp; Euphorbiaceae: Colliguaja 

odorifera; Rosaceae: Quillaja saponaria. 

Aleuroparadoxus rhodae Russell 1947 

Aleuroparadoxus rhodae Russell 1947: 20. Holotype. Mexico: on Gardenia sp, USNM; Martin 2005b: 17 


Host: Myrtaceae: Eugenia? sp.; Rubiaceae: Gardenia sp. 

Aleuroparadoxus sapotae Russell 1947 

Aleuroparadoxus sapotae Russell 1947: 27. Holotype. Belize [British Honduras]: Consejo, Yucatan 

Peninsula, on Achras sapota, USNM; Martin 2005b: 17 


Host: Sapotaceae: Achras sapota, Manilkara sp. 

Aleuroparadoxus trinidadensis Russell 1947 

Aleuroparadoxus trinidadensis Russell 1947: 28. Holotype. Trinidad: on Davilla aspera, USNM. 


Host: Dilleniaceae: Davilla aspera. 

Aleuroparadoxus truncatus Russell 1947 

Aleuroparadoxus truncatus Russell 1947: 30. Holotype. Honduras: Colon, Garunta, on Davilla rugosa, 

USNM; Martin 2005b: 17 

Distribution: 2: Belize, Honduras; Mexico. 

Host: Myrtaceae: Eugenia? sp.; Fabaceae: Lonchocarpus sp.; Dilleniaceae: Davilla matudai, 

Davilla rugosa. 

151

Aleuroplatus Quaintance & Baker 1914 

Aleuroplatus Quaintance & Baker 1914: 98. Type species. Aleurodes quercusaquaticae Quaintance 1900, by 


References: Russell 1944. 



acaciae x 

affinis x 

aguariae x 

akeassii x x 

alcocki 

alpinus x 

anapatsae x 

berbericolus x x 

bignoniae x 

biluminiporus x Int 

bossi x 

cadabae x x 

claricephalus x 

cockerelli x 

cococolus x 

coronatus x 

daitoensis x 

dentatus x 

dorsipallidus x 

dubius x 

elemarae x 

epigaeae x 

evodiae x 

152


fici x 

ficifolii x 

ficusrugosae x 

gelatinosus x x 

graphicus 

hiezi x 

hoyae x 

ilicis x x 

incisus x 

incurvatus x 

insularis x 

joholensis x 

lateralis x 

latus x 

liquidambaris x 

magnoliae x 

malayanus x 

mameti x 

manjakaensis x 

multipori x 

myricae x 

mysorensis x 

neovatus x 

oculiminutus x 

oculireniformis x 

ovatus x 

panamensis x 

pauliani x 

153


pectiniferus x x x x 

periplocae x 

perseaphagus x 

pileae x 

plumosus x x Int 

polystachyae x 

premnae x 

quaintancei x 

quercusaquaticae x 

robinsoni x 

sculpturatus x 

semiplumosus x Int? 

serratus x 

sinepecten x 

stellatus x 

subrotundus x 

translucidus x 

tsibabenae x 

tsimananensis x 

tuberculatus x 

vaccinii Int? 

validus x 

variegatus x 

vinsonioides x x 

weinmanniae x 

Total = 78 species 13 20 8* 29 3 14 2 1 0 

Aleuroplatus acaciae Bink-Moenen 1983 

Aleuroplatus acaciae Bink-Moenen 1983: 63. Holotype pupal case, Egypt, Aswan, Elefantine Island, 

21.i.1977, MAC; ex. Acacia tortilis, BMNH. 

Distribution: 3: Egypt. 

154

Host: Fabaceae: Acacia tortilis. 

Aleuroplatus affinis Takahashi 1961 

Aleuroplatus affinis Takahashi 1961: 332. Holotype. Madagascar: on unidentified plant, MNHNP. 


Host: on unidentified plant. 

Aleuroplatus aguariae Takahashi 1955 

Aleuroplatus agauriae Takahashi 1955a: 425. Syntypes. Madagascar: on Aguaria sp., MNHNP. 


Host: Ericaceae: Agauria sp. 

Aleuroplatus akeassii Cohic 1969 

Aleuroplatus akeassii Cohic 1969: 66. Holotype. Ivory Coast: on Baphia nitida. 


Host: Fabaceae: Baphia nitida. 

Aleuroplatus alcocki (Peal) 1903 

Aleurodes alcocki Peal 1903b: 74. Syntypes. India: Ficus religiosa. 

Aleuroplatus alcocki (Peal); Quaintance & Baker 1914: 98. 


Host: Annonaceae: Polyalthia longifolia, Polyalthia pendula; Moraceae: Ficus indica, Ficus religiosa, Morus 

alba. 

Note: David (1993) synonymized Aleuroplatus ficusgibbosus Corbett with this species; however, Martin 

(1999) did not accept that synonymy since the type material for A. alcocki (Peal) 1903 was not 

available for study; but rather synonymized A. ficusgibbosus with Aleuroplatus pectiniferus 

Quaintance & Baker 1917. 

Aleuroplatus alpinus Takahashi 1955 

Aleuroplatus alpinus Takahashi 1955a: 426. Syntypes. Madagascar: on undetermined host, MNHNP. 



Aleuroplatus anapatsae Takahashi 1951 

Aleuroplatus anapatsae Takahashi 1951a: 376. Syntypes. Madagascar: on "anapatsa-ala", MNHNP. 


Host: on "anapatsa-ala”. 

Aleuroplatus berbericolus Quaintance & Baker 1917 

Aleuroplatus berbericolus Quaintance & Baker 1917: 383. Syntypes. Canada: British Colombia, on Berberis 

aquifolium, USNM. 

Distribution: 1: Canada; USA (FL); 2: Mexico. 

Host: Aquifoliaceae: Ilex sp.; Berberidaceae: Berberis aquifolium, Mahonia sp. 

US Port Interception: Mexico on Eryngium sp. 

155

Aleuroplatus bignoniae Russell 1944 

Aleuroplatus bignoniae Russell 1944: 341. Holotype. USA: Florida, Brooksville, on Bignonia sp, USNM. 


Host: Bignoniaceae: Bignonia sp. 

Aleuroplatus biluminiporus Martin & Malumphy 2002 

Aleuroplatus biluminiporus Martin & Malumphy 2002: 2. Holotype puparium. ex. Colombia, at England, 

Hereford and Worcester, Evesham, Blackminster, on Laurus nobilis, 30.vii.2001, A. Kirby, BMNH. 

Distribution: 2: Colombia; 3: England (introduced). 

Host: Lauraceae: Laurus nobilis. 

Aleuroplatus bossi Takahashi 1936 

Aleuroplatus bossi Takahashi 1936d: 87. Syntypes. South West Africa: Namib, Witport, on Maerua 

angustifolia, TARI. 

Distribution: 4: Africa; Kenya; Nigeria; South Africa; Sudan; Tanzania. 

Host: Asclepiadaceae: Leptadenia heterophylla; Canellaceae: Warburgia ugandensis; Capparaceae: Maerua 

angolensis, Ritchiea afzelii; Liliaceae: Smilax sp.; Meliaceae: Trichilia roka; Polygalaceae: Polygala 

sp. 

Aleuroplatus cadabae Priesner & Hosny 1934 

Aleuroplatus cadabae Priesner & Hosny 1934a: 5. Syntypes. Egypt: on Cadaba rotundifolia, EDAC; Bink- 

Moenen 1983: 67. 

Distribution: 3: Egypt; 4: Ethiopia; Somalia; Sudan. 

Host: Asclepiadaceae: Leptadenia heterophylla; Capparidaceae: Cadaba rotundifolia, Cadaba sp.; 

Salvadoraceae: Salvadora sp. 

Natural enemies: Aphelinidae: Ablerus pumilus, Encarsia abatei, Encarsia aleuroplati, Eretmocerus cadabae, 

Eretmocerus mundus. 

Aleuroplatus claricephalus Takahashi 1940 

Aleuroplatus claricephalus Takahashi 1940b: 44. Syntypes. Mauritius: on undetermined indigenous plant, 

TARI. 

Distribution: 4: Mauritius. 


Aleuroplatus cockerelli (Ihering) 1897 

Aleurodes cockerelli Ihering 1897: 393. Syntypes. Brazil: on Baccharis paucifloscula, BMNH. 

Aleuroplatus cockerelli (Ihering); Quaintance & Baker 1914: 98. 


Host: Asteraceae: Baccharis paucifloscula. 

Aleuroplatus cococolus Quaintance & Baker 1917 

Aleuroplatus cococolus Quaintance & Baker 1917: 385. Syntypes. Trinidad: on Cocos nucifera, USNM; 

Martin 2005b: 18. 

Distribution: 2: Belize, Brazil; Colombia; Costa Rica, Cuba; Nicaragua, Panama; Trinidad. 

156

157 

Host: Arecaceae: Cocos nucifera; Asteraceae; Clusiaceae: Calophyllum sp; Combretaceae: Terminalia sp.; 

Loganiaceae: Strychnos sp.; Moraceae: Bosimum sp.; Myrtaceae: Eugenia michelii, Eugenia uniflora, 

Pimenta sp., Psidium sp.; Polygonaceae: Coccoloba sp.; Rubiaceae: Guettarda sp. 

US Port Interceptions: Dominican Republic on Citrus sp., Murraya sp., Psidium guajava; Guatemala on 

Bromeliaceae: Chamaedorea sp.; Honduras on Bromeliaceae: Nephelium sp.; Mexico on 

Chamaedorea elegans, Chamaedorea sp;, Citrus reticulata, Citrus sinensis, Citrus sp., Ericaceae, 

Laurus nobilis, Mangifera indica, Manilkara zapota, Myrtaceae: Persea americana, Persea sp., 

Pimenta sp., Piper nigrum, Piper sp., Psidium guajava, Strelitzia sp., Zingiber sp.; Puerto Rico on 

Malpighia glabra. 

Aleuroplatus coronatus (Quaintance) 1900 

Aleurodes coronata Quaintance 1900: 22. Syntypes. USA: California, Los Angeles, 5.xii.1887, D. W. 

Coquillett, on Quercus agrifolia, USNM. 

Aleuroplatus coronata (Quaintance); Quaintance & Baker 1914: 98. 

Distribution: 1: USA (AZ, CA, MS, NY); 2: Mexico. 

Host: Ericaceae: Arbutus menziesii; Fagaceae: Castanea sp., Quercus densiflora, Quercus wislizenii, Quercus 

agrifolia Nee var. agrifolia, Quercus chrysolepis; Rhamnaceae: Rhamnus sp., Rhamnus californica; 

Rosaceae: Heteromeles arbutifolia 

Natural enemies: Aphelinidae: Encarsia citrella, Encarsia pergandiella, Eretmocerus haldemani. 

Aleuroplatus daitoensis Takahashi 1940 

Aleuroplatus daitoensis Takahashi 1940d: 328. Syntypes. Loocoo Islands: on Ficus wightiana, TARI. 

Distribution: 8: Loochoo Islands. 

Host: Moraceae: Ficus wightiana. 

Aleuroplatus dentatus Sampson & Drew 1941 

Aleuroplatus dentatus Sampson & Drews 1941: 159. Syntypes. Mexico: Colima State, Manzanillo, on 

Forschammeria sp., UCD. 


Host: Capparidaceae: Forchhammeria sp. 

Aleuroplatus dorsipallidus Martin 1989 

Aleuroplatus dorsipallidus Martin 1988: 66. Holotype pupal case (male), Indonesia,. Sulawesi, Utara, 

15.iv.1985, J. Martin, on Palaquium? Sp., BMNH. 

Distribution: 7: Sulawesi. 

Host: Cunoniaceae: Weinmannia sp.; Moraceae: Taxotrophis ilicifolius; Sapotaceae: Palaquium? sp. 

Aleuroplatus dubius Takahashi 1955 

Aleuroplatus dubius Takahashi 1955a: 428. Syntypes. Madagascar: on undetermined host, MNHNP. 



Aleuroplatus elemarae Mound & Halsey 1978 

Aleuroplatus liquidambaris Russell 1944: 338. Holotype. USA: Lousianna, on Liquidamber styraciflua,

158 

USNM; name preoccupied by A. liquidambaris Takahashi 1941c: 391. 

Aleuroplatus elemarae Mound and Halsey 1978: 47, new name. 

Distribution: 1: USA (DC, FL, GA, LA, MD). 

Host: Ericaceae: Vaccinium sp.; Hamamelidaceae: Liquidambar styraciflua; Magnoliaceae: Magnolia sp.; 

Rosaceae: Pyracantha coccinea. 

Natural enemies: Aphelinidae: Encarsia citrella, Encarsia protransvena. 

Aleuroplatus epigaeae Russell 1944 

Aleuroplatus epigaeae Russell 1944: 340. Holotype. USA: Maryland, Silver Spring, Sligo Park, 6.xi.1943, L. 

Russell, on Epigaea repens, USNM. 

Distribution: 1: Canada; USA (CT, DC, MD, MN, NJ, NY, PA, WI, VA). 

Host: Ericaceae: Epigaea repens, Gaultheria sp., Vaccinium sp.; Lauraceae: Laurus sp. 

Aleuroplatus evodiae Takahashi 1960 

Aleuroplatus evodiae Takahashi 1960a: 141. Syntypes. Reunion Island: on Evodia sp., MNHNP. 


Host: Rutaceae: Evodia sp. 

Aleuroplatus fici Takahashi 1932 

Aleuroplatus fici Takahashi 1932: 31. Syntypes. Taiwan: on Ficus retusa, TARI. 


Host: Moraceae: Ficus retusa, Ficus microcarpa, Ficus swinhoei. 

Aleuroplatus ficifolii Takahashi 1942 

Aleuroplatus ficifolii Takahashi 1942b: 174. Syntypes. Thailand: on Ficus sp. 


Host: Moraceae: Ficus microcarpa, Ficus sp. 

Aleuroplatus ficusrugosae Quaintance & Baker 1917 

Aleuroplatus ficusrugosae Quaintance & Baker 1917: 387. Syntypes. India: on Ficus rugosa, USNM. 


Host: Moraceae: Ficus rugosa. 

Aleuroplatus gelatinosus (Cockerell) 1898 

Aleurodes gelatinosus Cockerell 1898a: 264. Syntypes. USA: New Mexico, on Quercus arizonica, USNM. 

Aleuroplatus gelatinosus (Cockerell); Quaintance & Baker 1914: 98 

Distribution: 1: USA (AZ, CA, GA, NM, NV, OR, TX, UT, WA); 2: Cuba, Mexico. 

Host: Fagaceae: Quercus arizonica, Fagaceae: Quercus agrifolia; Rhamnaceae: Rhamnus californica; 

Rubiaceae: Casasia calophyla. 

Natural enemies: Aphelinidae: Encarsia quercicola. 

Aleuroplatus graphicus Bondar 1923 

Aleuroplatus graphicus Bondar 1923a: 117. Syntypes. Brazil: on Sapotaceae, MZU. 


Host: Sapotaceae.

Aleuroplatus hiezi Cohic 1968 

Aleuroplatus hiezi Cohic 1968b: 103. Syntypes. Congo: on Gaertnera paniculata; Bink-Moenen 1983: 67. 


Host: Apocynaceae: Tabernaemontana crassa, Rauvolfia vomitoria; Celastraceae: Salacia sp. 

Dilleniaceae: Tetracera alnifolia; Rubiaceae: Gaertnera paniculata. 

Aleuroplatus hoyae (Peal) 1903 

Aleurodes hoyae Peal 1903b: 88. Syntypes. India: on Hoya sp. 

Aleuroplatus hoyae (Peal); Quaintance & Baker 1914: 98. 


Host: Asclepiadaceae: Hoya sp. 

Aleuroplatus ilicis Russell 1944 

Aleuroplatus ilicis Russell 1944: 337. Holotype. USA: Maryland, Silver Spring, on Ilex opaca, USNM. 

Distribution: 1: USA (AL, DE, FL, GA, LA, MD, MS, NC, SC, TN, TX, VA, WV); 2: Mexico. 

Host: Aquifoliaceae: Ilex x attenuata, Ilex sp., Ilex vomitoria, Ilex opaca, Ilex opaca, Ilex cassine; Ericaceae: 

Kalmia sp.; Hamamelidaceae: Liquidambar styraciflua; Illiciaceae: Illicium anisatum; Lauraceae: 

Laurus sp.; Nyssaceae: Nyssa sylvatica, Nyssa ogeche; Oleaceae: Osmanthus americanus; Rosaceae: 

Prunus caroliniana; Theaceae: Gordonia lasianthus. 

Natural enemies: Aphelinidae: Encarsia formosa, Encarsia protransvena; Encyrtidae: Metaphycus sp.; 

Platygastridae: Amitus sp. 

Aleuroplatus incisus Quaintance & Baker 1917 

Aleuroplatus incisus Quaintance & Baker 1917: 388. Syntypes. Sri Lanka: on Ostodes zeylanica, USNM. 


Host: Euphorbiaceae: Ostodes zeylanica; Guttiferae: Garcinia specta. 

Aleuroplatus incurvatus Takahashi 1961 

Aleuroplatus incurvatus Takahashi 1961: 333. Syntypes (3). Madagascar: on unidentified plant, MNHNP. 



Aleuroplatus insularis Takahashi 1941 

Aleuroplatus insularis Takahashi 1941b: 355. Syntypes. Mauritius: on Olea sp. 


Host: Oleaceae: Olea sp. 

159

Aleuroplatus joholensis Corbett 1935 

Aleuroplatus joholensis Corbett 1935b: 782. Syntypes. Malaya: on Dillenia sp. 


Host: Dilleniaceae: Dillenia sp. 

Aleuroplatus lateralis Bondar 1923 

Aleuroplatus lateralis Bondar 1923a: 117. Syntypes. Brazil: on Eugenia sp, MZU. 



Aleuroplatus latus Takahashi 1939 

Aleuroplatus latus Takahashi 1939c: 2. Syntypes. Mauritius: on undetermined indigenous tree, TARI. 


Host: on undetermined indigenous tree. 

Aleuroplatus liquidambaris Takahashi 1941 

Aleuroplatus liquidambaris Takahashi 1941c: 391. Syntypes. Hong Kong: on Liquidambar sp. 


Host: Hamamelidaceae: Liquidambar sp. 

Aleuroplatus magnoliae Russell 1944 

Aleuroplatus plumosa Quaintance 1900: 33; misidentification in part. 

Aleuroplatus magnoliae Russell 1944: 337. Holotype. USA: Florida, Lake City, A. L. Quaintance, on 

Magnolia virginiana [=M. glauca], USNM. 


Host: Ericaceae: Vaccinium sp.; Magnoliaceae: Magnolia virginiana, Magnolia sp. 

Aleuroplatus malayanus Takahashi 1955 

Aleuroplatus malayanus Takahashi 1955b: 234. Syntypes. Malaya: on Pueraria javanica stems, BMNH. 


Host: Fabaceae: Pueraria phaseoloides, Pueraria javanica. 

Aleuroplatus mameti Takahashi 1937 

Aleuroplatus mameti Takahashi 1937a: 44. Syntypes. Mauritius: on unidentified tree, MNHNP. 

160



Aleuroplatus manjakaensis Takahashi 1955 

Aleuroplatus manjakaensis Takahashi 1955a: 429. Holotype. Madagascar: on undetermined host, MNHNP. 



Aleuroplatus multipori Takahashi 1940 

Aleuroplatus multipori Takahashi 1940b: 46. Syntypes. Mauritius: on undetermined plant, TARI. 



Aleuroplatus myricae Quaintance & Baker 1917 

Aleuroplatus myricae Quaintance & Baker 1917: 389. Lectotype (designated by Russell 1944: 337). USA: 

Georgia, Griffen, on Myrica pennsylvanica, USNM. 

Distribution: 1: USA (FL, GA, NC, NJ). 

Host: Calycanthaceae: Calycanthus fertilis; Ericaceae: Kalmia sp., Rhododendron nudiflorum, 

Vaccinium sp.; Myricaceae: Myrica pensylvanica; Rosaceae: Prunus serotina. 

Aleuroplatus mysorensis David & Subramaniam 1976 

Aleuroplatus mysorensis David & Subramaniam 1976: 166. Holotype. India: Karnataka, Mysore, 19.IX.1985, 

B. V. David, on Ailantus excelsa, ZSI. 



Aleuroplatus neovatus Takahashi 1961 

Aleuroplatus neovatus Takahashi 1961: 334. Syntypes. Madagascar: on "liane", MNHNP. 



Aleuroplatus oculiminutus Quaintance & Baker 1917 

Aleuroplatus oculiminutus Quaintance & Baker 1917: 390. Syntypes. Trinidad: on Ficus sp., USNM. 

161



Aleuroplatus oculireniformis Quaintance & Baker 1917 

Aleuroplatus oculireniformis Quaintance & Baker 1917: 391. Syntypes. Brazil: on Passiflora sp., USNM. 


Host: Passifloraceae: Passiflora sp. 

Aleuroplatus ovatus Quaintance & Baker 1917 

Aleuroplatus ovatus Quaintance & Baker 1917: 392. Syntypes. USA: Texas, College Station, on Berberis 

trifoliata, USNM. 

Distribution: 1: USA (AL, AZ, FL, NM, TX). 

Host: Aquifoliaceae: Ilex sp.; Berberidaceae: Berberis haemotocarpa, Berberis trifoliata, 

Mahonia trifoliata. 

US Port Interception: Mexico on Mahonia sp. 

Aleuroplatus panamensis Sampson & Drews 1941 

Aleuroplatus panamensis Sampson & Drews 1941: 161. Syntypes. Panama: on Gaultheria sp., UCD. 


Host: Ericaceae: Gaultheria sp. 

Aleuroplatus pauliani Takahashi 1955 

Aleuroplatus pauliani Takahashi 1955a: 432. Syntypes. Madagascar: on undetermined host, MNHNP. 



Aleuroplatus pectiniferus Quaintance & Baker 1917 

Aleuroplatus pectiniferus Quaintance & Baker 1917: 393. Syntypes. India: on Euphorbiaceae, USNM. 

Aleuroplatus buchananiae Jesudasan and David 1991: 282. Holotype pupal case. India: on Buchanania 

lanzan, IDAV; Martin 1999: 48. 

Aleuroplatus cinnamomi Jesudasan and David 1991: 282. Holotype pupal case. India: on Cinnamomum sp., 

IDAV; Martin 1999: 48. 

Aleuroplatus distinctus Jesudasan and David 1991: 283. Holotype pupal case. India: on unidentified tree, 

IDAV; Martin 1999: 48. 

162

163 

Aleuroplatus ficusgibbosae Corbett 1926: 271. Syntypes. Sri Lanka: on Ficus gibbosa, BMNH; Martin 1999: 

48. 

Aleuroplatus pectenserratus Singh 1945: 76. Syntypes. India: on Terminalia sp.; Martin 1999: 48. 

Aleuroplatus walayarensis Jesudasan and David 1991: 285. Holotype pupal case. India: on Casearia 

esculenta, IDAV; Martin 1999: 48. 

Distribution: 3: Iran; 5: China; Taiwan; 6: India; Pakistan; Malaya; Malaysia; Sri Lanka; 7: Australia, Java, 

Sulawesi. 

Host: Anacardiaceae: Buchanania lanzan; Betulaceae: Alnus formosana; Combretaceae: Terminalia sp.; 

Euphorbiaceae: Bischofia javanica, Chrozophora obliqua; Flacourtiaceae: Casearia esculenta; 

Lauraceae: Cinnamomum sp.; Malvaceae: Urena lobata; Moraceae: Morus alba, Ficus gibbosa, Ficus 

capensis, Ficus vasculosa; Myrtaceae: Eugenia jambos; Rutaceae: Evodia sp.; Salicaceae: Salix 

warburgii, Salix sp.; Theaceae: Gordonia anomala. 

Natural enemies: Aphelinidae: Encarsia ishii, Encarsia lutea, Encarsia perplexa, Eretmocerus cadabae. 

Note: David (1993) synonymized Aleuroplatus ficusgibbosus Corbett with this species; however, 

Martin (1999) did not accept that synonymy since the type material for A. alcocki (Peal) 

1903 was not available for study; but rather synonymized A. ficusgibbosus with Aleuroplatus 

pectiniferus Quaintance & Baker 1917. 

Aleuroplatus periplocae (Dozier) 1934 

Aleyrodes periplocae Dozier 1934: 191. Syntypes. Zaire: on Periplaca nigrescens, USNM. 

Aleuroplatus periplocae (Dozier); Mound 1965c: 134; Cohic 1969: 74; Bink-Moenen 1983: 67 


Host: Asclepiadaceae: Periploca nigrescens. 

Natural enemies: Aphelinidae: Eretmocerus sp. 

Aleuroplatus perseaphagus Martin, Aguiar & Pita 1996 

Aleuroplatus perseaphagus Martin, Aguiar & Pita 1996, in Martin et al. 1996: 117. Holotype pupal case, 

Madeira, Funchal, Lombo da Boa Vista, 3.viii.1993, F. Aguiar, on Persea americana, BMNH. 

Distribution: 3: Madeira Islands. 

Host: Lauraceae: Persea americana, Ocotea foetens. 

Aleuroplatus pileae Takahashi 1939 

Aleuroplatus pileae Takahashi 1939c: 3. Syntypes. Mauritius: on Pilea balfouri, TARI. 


Host: Urticaceae: Pilea balfouri.

Aleuroplatus plumosus (Quaintance) 1900 

Aleurodes plumosa Quaintance 1900: 33. Lectotype (designated by Russell 1944:336). USA: Florida, on 

Quercus sp., USNM. 

Tetraleurodes plumosa (Quaintance); Quaintance & Baker 1914: 108. 

Aleuroplatus plumosus (Quaintance); Quaintance & Baker 1917:394. 

Distribution: 1: USA (AL, DC, FL, LA, MA, MD, NJ, WI, WV); 2: Mexico; 3: United Kingdom* 

[interception]. 

Host: Annonaceae: Asimina sp.; Aquifoliaceae: Ilex cassine, Ilex crenata, Ilex glabra, Ilex opaca; Arecaceae: 

Rhapis excelsa; Caprifoliaceae: Viburnum nudum; Ericaceae: Kalmia sp., Lyonia ferruginea, Lyonia 

ferruginea, Lyonia lucida, Lyonia sp., Vaccinium corymbosum, Vaccinium darrowii, Vaccinium 

myrsinites, Vaccinium stamineum; Fagaceae: Quercus sp.; Guttiferae: Hypericum cistifolium, 

Hypericum edisonianum, Hypericum galioides, Hypericum hypericoides, Hypericum myrtifolium, 

Hypericum punctatum, Hypericum stans, Hypericum tetrapetalum; Lauraceae: Persea borbonia, 

Persea carolinensis; Loganiaceae: Gelsemium sempervirens; Magnoliaceae: Magnolia glauca, 

Magnolia grandiflora; Melastomataceae: Rhexia alifanus; Myricaceae: Myrica cerifera, Myrica rubra; 

Oleaceae: Fraxinus caroliniana; Rosaceae: Prunus caroliniana, Pyracantha coccinea; Theaceae: 

Gordonia lasianthus. 

Natural enemies: Platygastridae: Amitus aleurodinis. 

Aleuroplatus polystachyae Takahashi 1955 

Aleuroplatus polystachyae Takahashi 1955a: 433. Syntypes. Madagascar: on Polystachys sp., MNHNP. 


Host: Orchidaceae: Polystachya sp. 

US Port Interception: Madagascar on Oeonia sp. 

Aleuroplatus premnae Corbett 1926 

Aleuroplatus premnae Corbett 1926: 272. Syntypes. Sri Lanka: on Prenna cordifolia, BMNH. 


Host: Verbenaceae: Premna cordifolia. 

Aleuroplatus quaintancei (Peal) 1903 

Aleurodes quaintancei Peal 1903b: 78. Syntypes. India: on Ficus religiosa. 

Aleuroplatus quaintancei (Peal); Quaintance & Baker 1914: 98. 

Host: Moraceae: Ficus religiosa. 

164

Aleuroplatus quercusaquaticae (Quaintance) 1900 

Aleurodes quercus-aquaticae Quaintance 1900: 35. Syntypes. USA: Florida, Lake City, on Quercus aquatica, 

USNM. 

Aleuroplatus quercusaquaticae (Quaintance); Quaintance & Baker 1914: 98. 

Distribution: 1: USA (DC, FL, MD, MS, TX, VA). 

Host: Annonaceae: Asimina incana; Fagaceae: Quercus aquatica, Quercus laurifolia, Quercus sp. 

Aleuroplatus robinsoni Takahashi 1955 

Aleuroplatus robinsoni Takahashi 1955a: 434. Syntypes. Madagascar: on Anthocleista sp., MNHNP. 


Host: Loganiaceae: Anthocleista sp. 

Aleuroplatus sculpturatus Quaintance & Baker 1917 

Aleuroplatus sculpturatus Quaintance & Baker 1917: 396. Syntypes. Panama: on Heliconia sp., USNM. 


Host: Musaceae: Heliconia sp. 

US Port Interceptions: Trinidad and Tobago on Heliconia sp.; Venezuela on Heliconia sp. 

Aleuroplatus semiplumosus Russell 1944 

Aleuroplatus semiplumosus Russell 1944: 336. Holotype. USA: Virginia, Norfolk, on Persea borbonia, 

USNM. 

Distribution: 1: USA (DC, DE, FL, GA, IL, LA, MD, MS, NC, NY, PA, SC, VA, WV); 3: United Kingdom 

Host: Aquifoliaceae: Ilex opaca; Ericaceae: Kalmia latifolia, Rhododendron sp, Vaccinium sp. 

Euphorbiaceae: Codiaeum variegatum; Guttiferae: Hypericum fasciculatum; Lauraceae: Persea 

palustris, Persea borbonia, Persea pubescens, Sassafras albidum; Magnoliaceae: Magnolia 

virginiana; Nyssaceae: Nyssa ogeche, Nyssa sylvatica; Oleaceae: Fraxinus sp.; Rubiaceae: 

Cephalanthus sp. 

Aleuroplatus serratus Takahashi 1955 

Aleuroplatus serratus Takahashi 1955a: 437. Syntypes. Madagascar: on "tambitsy", MNHNP. 


Host: on "tambitsy" 

Aleuroplatus sinepecten Singh 1945 

Aleuroplatus sinepecten Singh 1945: 75. Syntypes. Burma: on undetermined herb. 


165

Host: on undetermined herb. 

Aleuroplatus stellatus (Hempel) 1922 

Aleurotrachelus stellatus Hempel 1922 Hempel 1922: 4. Syntypes. Brazil: on Cocos nucifera. 

Bondar 1923a. 

Aleuroplatus stellatus (Hempel); Martin & Mound 2007. 

Distribution: 2: Brazil; Colombia. 

Host: Arecaceae: Cocos nucifera; Fabaceae: Inga sp.; Rubiaceae: Chomelia oligantha. 

Aleuroplatus subrotundus Takahashi 1938 

Aleuroplatus subrotundus Takahashi 1938d: 261. Syntypes. Mauritius: on undetermined indigenous tree, 

MNHNP. 

Distribution: 4: Mauritius; Reunion Island. 

Host: Myrtaceae: Eugenia jambos. 

Aleuroplatus translucidus Quaintance & Baker 1917 

Aleuroplatus translucidus Quaintance & Baker 1917: 397. Syntypes. Pakistan: Wazirabad and Lahore, 1910 

on Citrus sp., USNM. 

Distribution: 3: Iran; 6: India, Pakistan. 

Host: Rutaceae: Citrus sp.; Murraya sp. 

US Port Interception: India, intercepted at Los Angeles, California, DEC, 19, 1983, J. Floyd, on 

Murraya sp.; India, intercepted at Chicago, Illinois, FEB 8, 1986, B. Yates, on Murraya 

sp; Iran, intercepted at Boston, MA, Sept, 22, 1982, H. Batchelder, on Murraya sp. 

Note: The original record of this species listed Wazirabad and Lahore as being in India; however 

after gaining its independence, these cities became a part of Pakistan. 

Aleuroplatus tsibabenae Takahashi 1955 

Aleuroplatus tsibabenae Takahashi 1955a: 437. Syntypes. Madagascar: on "tsibabena", MNHNP. 


Host: on "tsibabena". 

Aleuroplatus tsimananensis Takahashi 1955 

Aleuroplatus tsimananensis Takahashi 1955a: 440. Syntypes. Madagascar: on unidentified host, MNHNP. 



166

Aleuroplatus tuberculatus Takahashi 1951 

Aleuroplatus tuberculatus Takahashi 1951a: 375. Syntypes. Madagascar: on unidentified host, MNHNP. 


Host: on unidentified. 

Aleuroplatus vaccinii Russell 1944 

Aleuroplatus plumosus (Quaintance); misidentification in part Quaintance & Baker 1917: 395. 

Aleuroplatus vaccinii Russell 1944: 340. Holotype. USA: New Jersey, on Vaccinium corymbosum, USNM 

Distribution: 1: Canada; USA (DC, FL, GA, IN, ME, MD, NJ, NY, NC, SC); 3: England (interception?). 

Host: Aquifoliaceae: Ilex crenata, Ilex sp.; Ericaceae: Gaultheria sp., Gaylussacia baccata, 

Gaylussacia frondosa, Kalmia latifolia, Lyonia lucida, Lyonia sp., Vaccinium ashei, 

Vaccinium corymbosum, Vaccinium darrowii, Vaccinium myrsinites, Vaccinium vacillans; Fabaceae: 

Sesbania exaltata; Lauraceae: Laurus sp.; Loganiaceae: Gelsemium sempervirens; Nyssaceae: Nyssa 

sylvatica; Pyrolaceae: Chimaphila umbellata, Pyrola sp.; Rubiaceae: Randia aculeata. 

Aleuroplatus validus Quaintance & Baker 1917 

Aleuroplatus validus Quaintance & Baker 1917: 398. Syntypes. Jamaica: on undetermined host plant, USNM; 

Vazquez 1999: 143. 

Distribution: 2: Cuba; Jamaica. 

Host: Iridaceae: Espadaea amoena. 

US Port Interceptions: Antigua and Barbuda on Annona muricata; Dominican Republic on Canella 

winteriana, Guatteria blainii; Haiti on Annona sp., Citrus sp., Eucalyptus sp. 

Aleuroplatus variegatus Quaintance & Baker 1917 

Aleuroplatus variegatus Quaintance & Baker 1917: 399. Syntypes. Costa Rica: on Psidium sp., USNM. 

Distribution: 2: Costa Rica. 


Aleuroplatus vinsonioides (Cockerell) 1898 

Aleurodes vinsonioides Cockerell 1898b: 225. Syntypes. Mexico: on leaves of tree having small white 

flowers, USNM. 

Aleuroplatus vinsonioides (Cockerell); Quaintance & Baker 1914: 98; Martin 2005b: 19. 

Distribution: 1: USA (FL, GA, MS); 2: Belize, Costa Rica, Cuba, Guyana, Jamaica, Mexico, Panama 

167

168 

Host: Lauraceae: Nectandra sp., Nectandra coriacea, Nectandra earlei, Ocotea coriacea, Ocotea sp. 

Aleuroplatus weinmanniae Takahashi 1951 

Aleuroplatus weinmanniae Takahashi 1951a: 373. Syntypes. Madagascar: on Weinmannia sp, MNHNP. 


Host: Cunoniaceae: Weinmannia sp. 

Transfers 

Aleuroplatus denticulatus Bondar 1923 to Dialeurodes; Martin 2005: 39

Aleuropleurocelus Drews & Sampson 1956 

Aleuropleurocelus Drews & Sampson 1956: 282. Type species. Aleuropleurocelus laingi Drews & Sampson 

1956, by original designation. 



abnormis x x 

acaudatus x 

ceanothi x 

cecropiae x 

coachellensis x 

granulata x x 

laingi x 

nigrans x x 

oblanceolatus x 

ornatus x 

rotunda x 

sierrae x 

Total = 11 species 10 5 0 0 0 0 0 0 0 

Aleuropleurocelus abnormis (Quaintance) 1900 

Aleyrodes abnormis Quaintance 1900: 17. Syntypes. USA: Florida, on Quercus aquatica, Q. virens and other 

plants, USNM. 

Tetraleurodes abnormis (Quaintance); Quaintance & Baker 1914: 108. 

Aleuropleurocelus abnormis (Quaintance) 1900; Martin 2005b. 

Distribution: 1: USA (AZ, FL, TX); 2: Mexico. 

Host: Aquifoliaceae: Ilex opaca; Fagaceae: Quercus aquatica, Quercus catesbaei, Quercus laurifolia, 

Quercus myrtifolia, Quercus nigra, Quercus virens, Quercus virginiana; Lauraceae: Persea 

americana, Persea borbonia, Persea carolinensis; Magnoliaceae: Magnolia glauca, 

US Port Interceptions: Colombia on Laurus nobilis; Dominican Republic on Lauraceae; El Salvador on 

169

170 

Persea americana; Mexico on Citrus sp., Eucalyptus sp., Laurus nobilis, Mangifera indica, Persea 

americana, Persea sp.; Peru on Laurus nobilis. 

Aleuropleurocelus acaudatus Drews & Sampson 1958 

Aleuropleurocelus acaudatus Drews & Sampson 1958: 125. Syntypes. USA: California, on Artostaphylos sp.. 

Distribution: 1: USA (CA). 

Host: Ericaceae: Arctostaphylos sp. 

Aleuropleurocelus ceanothi (Sampson) 1945 

Tetralicia ceanothi Sampson 1945: 59. Syntypes. USA: California, on Ceanothus cunicatus, UCD. 

Aleuropleurocelus ceanothi (Sampson) 1945; Drews & Sampson 1956: 281 

Distribution: 1: USA (CA, NV). 

Host: Rhamnaceae: Ceanothus cuneatus sp. cuneatus. 

Aleuropleurocelus cecropiae (Bondar) 1923 

Aleurotrachelus cecropiae Bondar 1923a: 143. Syntypes. Brazil: on Cecropia adenops, MZU. 

Aleuropleurocelus cecropiae (Bondar); Martin 2005b: 20. 


Host: Moraceae: Cecropia adenopus. 

Aleuropleurocelus coachellensis Drews & Sampson 1958 

Aleuropleurocelus coachellensis Drews & Sampson 1958: 120. Syntypes. USA: California, on Pluchea 

sericea. 


Host: Asteraceae: Pluchea sericea. 

Aleuropleurocelus granulata (Sampson and Drews) 1941 

Tetalicia granulata Sampson & Drews 1941: 169. Syntypes. Mexico: on undetermined plant. UCD. 

Aleuropleurocelus granulata (Sampson & Drews); Mound and Halsey 1978: 59 

Distribution: 1: USA (TX); 2: Mexico. 

Host: Verbenaceae: Lantana camara, on undetermined plant 

Aleuropleurocelus laingi Drews & Sampson 1956 

Aleuropleurocelus laingi Drews & Sampson 1956: 282. Syntypes. USA: California, on Arctostaphylos sp.


Host: Ericaceae: Arctostaphylos sp.; Labiatae: Salvia apiana. 

Aleuropleurocelus nigrans (Bemis) 1904 

Aleyrodes nigrans Bemis 1904: 522. Syntypes. USA: California, on Clematis ligustifolia and many other 

plants, USNM; Penny 1922: 34. 

Tetraleurodes nigrans (Bemis); Quaintance & Baker 1914: 618. 

Tetralicia nigrans (Bemis); Baker 1937: 281. 

Aleuropleurocelus nigrans (Bemis); Drews & Sampson 1956: 281. 

Distribution: 1: USA (AZ, CA); 2: Mexico. 

Host: Caprifoliaceae: Lonicera involucrata, Symphoricarpos racemosus; Ericaceae: Arctostaphylos 

manzanita, Arbutus menziesii, Arctostaphylos sp.; Fagaceae: Quercus densiflora; Hydrophyllaceae: 

Eriodictyon californicum; Labiatae: Salvia sp., Lauraceae: Umbellularia californica; Ranunculaceae: 

Clematis ligusticifolia; Rhamnaceae: Ceanothus californicus, Rhamnus californica Eschsch sp. 

californica; Rosaceae: Heteromeles arbutifolia, Prunus ilicifolia 

Aleuropleurocelus oblanceolatus Drews & Sampson 1958 

Aleuropleurocelus oblanceolatus Drews & Sampson 1958: 124. Syntypes. USA: California, on Rhamnus 

crocea var ilicifolia. 


Host: Rhamnaceae: Rhamnus crocea Nutt. sp. ilicifolia. 

Aleuropleurocelus ornatus Drews & Sampson 1958 

Aleuropleurocelus ornatus Drews & Sampson 1958: 121. Syntypes. USA: California, on Eriodictyon 

trichocalyx. 


Host: Hydrophyllaceae: Eriodictyon trichocalyx var. trichocalyx. 

Aleuropleurocelus rotunda (J. M. Baker) 1937 

Tetralicia rotunda J. M. Baker 1937: 619. Syntypes. Mexico: unknown host, USNM. 

Aleuropleurocelus rotunda (J. M. Baker); Martin & Mound 2007: 17. 


Host: unknown host. 

Note: This species should be checked to see if it should be placed in Aleuropleurocelus along with the other 

New World species. 

171

Aleuropleurocelus sierrae (Sampson) 1945 

Tetralicia sierrae Sampson 1945: 60. Syntypes. USA: California, on low spreading shrub, UCD. 

Aleuropleurocelus sierrae (Sampson); Drews & Sampson 1956: 60. 


Host: low spreading shrub. 

172

Aleuroporosus Corbett 1935 

Aleuroporosus Corbett 1935b: 844. Type species. Aleuroporosus lumpurensis Corbett 1935, by monotypy. 



lumpurensis x 

Total = 1 species 0 0 0 0 0 1 0 0 0 

Aleuroporosus lumpurensis Corbett 1935 

Aleuroporosus lumpurensis Corbett 1935b: 845. Syntypes. Malaya: on unidentified host, lost? 



173

Aleuropteridis Mound 1961 

Aleuropteridis Mound 1961b: 127. Type species. Aleuropteridis douglasi Mound 1961, by original 

designation [junior synonym of A. filicicola (Newstead). 

Host: on ferns 




eastopi x 

filiciicola Int? x 

hargreavesi x 

jamesi x 

Total = 4 species 0 0 1* 4 0 0 0 0 0 

Aleuropteridis eastopi Mound 1961 

Aleuropteridis eastopi Mound 1961b: 129. Holotype. Ghana: on fern, BMNH; Bink-Moenen 1983: 69. 

Distribution: 4: Ghana; Ivory Coast. 

Host: on fern, Dryopteridaceae: Nephrolepis biserrata. 

Aleuropteridis filicicola (Newstead) 1911 

Aleyrodes filicicola Newstead 1911: 174. Syntypes. Tanzania: on fern, BMNH. 

Aleuropteridis filicicola (Newstead); Mound 1965c: 135; Bink-Moenen 1983: 68. 

Distribution: 3: England; 4: Ivory Coast; Tanzania. 

Host: Polypodiaceae/Pteridaceae: Pteris togoensis; Thelypteridaceae: Cyclosorus dentatus. 

Aleuropteridis hargreavesi Mound 1961 

Aleuropteridis hargreavesi Mound 1961b: 131. Syntypes. Sierra Leone: on bush fern, BMNH; Bink-Moenen 

1983: 68 

Distribution: 4: Sierra Leon. 

Host: on bush fern. 

174

Aleuropteridis jamesi Mound 1961 

Aleuropteridis jamesi Mound 1961b: 129. Syntypes. Nigeria, Ibadan, 29.xii.1960, E. A. James, on Pteris 

togoensis; BMNH; Bink-Moenen 1983: 68. 


Host: Pteridaceae: Pteris togoensis. 

175

Aleuroputeus Corbett 1935 

Aleuroputeus Corbett 1935b: 846. Type species. Aleuroputeus perseae Corbett 1935, by original designation. 



baccaureae x 

perseae x 

perseae? x 

Total = 2+ 0 0 0 0 0 2 1 0 0 

Aleuroputeus baccaureae Corbett 1935 

Aleuroputeus baccaureae Corbett 1935b: 847. Holotype. Malaya: on Baccaurea motleyana, lost? 



Aleuroputeus perseae Corbett 1935 

Aleuroputeus perseae Corbett 1935b: 846. Syntypes. Malaya: on Persea gatissima, lost? 


Host: Lauraceae: Persea gratissima. 

Aleuroputeus perseae? Corbett 1935 

Aleuroputeus perseae? Corbett; Martin 1988: 68. 

Distribution: 7: Java on Myrtaceae: Psidium guajava; 7: Sulawesi on Ulmaceae: Trema orientalis 

176

Aleurothrixus Quaintance & Baker 1914 

Aleurothrixus Quaintance & Baker 1914: 103. Type species. Aleyrodes howardi Quaintance 1907:91, junior 

synonym of Aleurodes floccosa Maskell 1896:432. 

Aleurothrixus (Philodamus) Quaintance & Baker 1917: 404. Type species. Aleyrodes interrogationis Bemis 


Hempelia Sampson & Drews 1941: 166. Type species. Hempelia chivelensis Sampson & Drews 1941, by 

monotypy; synonymy according to Martin 2005b: 20. 



aepim x 

aguiari x 

antidesmae x x x x 

bondari x 

chivelensis x 

floccosus x x x x x x x 

guareae x 

guimaraesi x 

interrogationis x 

lucumai x 

miconiae x 

myrtacei x 

myrtifolii x 

ondinae x 

porteri x 

proximans x 

silvestri x 

similis x 

177


smilaceti x 

solani x 

Total = 20 species 2 16 1 1 3 3 1 1 1 

Aleurothrixus aepim (Goldi) 1886 

Aleurodes aepim Goeldi 1886: 250. Syntypes. Brazil: on Manihot palmata. 

Aleurothrixus aepim (Goldi); Quaintance & Baker 1914: 103. 

Aleurothrixus graneli Blanchard 1918: 344. Syntypes. Argentina: on Ipomoea sp.; synonymy according to 

Costa Lima 1942c: 421. 

Distribution: 2: Argentina; Brazil, Uruguay. 

Host: Asteraceae: Baccharis oxyodenta, Mikania scandens; Convolvulaceae: Ipomoea sp. 

Euphorbiaceae: Manihot esculenta, Manihot palmata, Manihot utilissima; Myrtaceae: Psidium 

guajava; Rubiaceae: Coffea arabica; Rutaceae: Citrus sp. 

Natural enemies: Aphelinidae: Encarsia perplexa Huang & Polaszek 1998, Encarsia porteri (Mercet), 

Encarsia aleurothrixi Evans and Polaszek 1998, Eretmocerus sp., Marietta sp.; Encyrtidae: 

Metaphycus sp. 

Aleurothrixus aguiari Costa Lima 1942 

Aleurothrixus aguiari Costa Lima 1942c: 425. Syntypes. ?Brazil: on Guarea trichiliodes. 


Host: Meliaceae: Guarea trichiliodes; Sapotaceae: Lucuma caimito. 

Aleurothrixus antidesmae Takahashi 1933 

Aleurothrixus antidesmae Takahashi 1933: 13. Syntypes. Taiwan: on Antidesma sp., TARI 

Distribution: 5: Taiwan; 6: Philippines, Thailand; 9: Hawaii. 

Host: Euphorbiaceae: Antidesma sp. 

US Port Interceptions: Hawaii on Anthurium andraeanum, Gardenia jasminoides, Gardenia sp., Ixora sp., 

Morinda citrifolia, Ocimum sp., Piper sp.; Puerto Rico on Ixora sp. 

Aleurothrixus bondari Costa Lima 1942 

Aleurothrixus bondari Costa Lima 1942c: 421. Syntypes. Brazil: on unidentified plant. 


178

Host: on unidentified plant; Tiliaceae: Triumfetta semitriloba. 

Aleurothrixus chivelensis (Sampson & Drews) 1941 

Hempelia chivelensis Sampson & Drews 1941: 166. Syntypes. Mexico: on undetermined tree, UCD. 

Aleurothrixus chivelensis Sampson & Drews; Martin 2005b: 20. 



Aleurothrixus floccosus (Maskell) 1895 

Aleurodes floccosa Maskell 1895: 432. Syntypes. Cuba: on Citrus sp, NZAC 

Aleurothrixus floccosus (Maskell); Quaintance & Baker 1914: 91. 

Aleyrodes horridus Hempel 1899: 394. Syntypes. Brazil: on Psidium guajava. 

Aleurothrixus horridus (Hempel); Quaintance & Baker 1914: 103. 

Aleyrodes howardi Quaintance 1907: 91. Syntypes. Cuba: on Citrus sp., USNM; synonymized by Costa Lima 

1945c: 425. 

Distribution: 1: USA (CA, FL, HI, LA, TX); 2: Antigua, Argentina; Bahamas; Barbados, Bermuda, Bolivia, 

Brazil; Chile; Colombia, Costa Rica; Colombia; Cuba; Dominica, Dominican Republic, Ecuador; El 

Salvador, Guadeloupe; Guatemala, Guinea, Guyana, Haiti; Honduras; Jamaica, Martinique, Mexico, 

Montserrat, Nicaragua, Panama; Paraguay; Puerto Rico; Suriname, Trinidad & Tobago, Uruguay, 

Venezuela, Virgin Islands; 3: Canary Islands, France; Iran, Italy; Madeira, Morocco; Portugal, Sicily, 

Spain; 4: Angola; Congo; Gabon, Guinea, Liberia, Nigeria, Reunion; 5: Japan, Taiwan; 6: India, 

Thailand; 8: Guam, Tahiti. 

Host: Anacardiaceae: Anacardium sp., Mangifera indica, Schinus terebinthifolius, Spondias dulcis, 

Anacardiaceae: Spondias lutea, Spondias mombin; Annonaceae: Annona reticulata, Annona 

squamosa; Apocynaceae: Plumeria rubra; Araceae: Dieffenbachia maculata, Dieffenbachia sp.; 

Asclepiadaceae: Paraquetina nigrescens; Asteraceae: Baccharis genistelloides, Emilia sonchifolia, 

Eupatorium odoratum; Bignoniaceae: Catalpa longissima, Tabebuia glomerata; Boraginaceae: Cordia 

collococca; Burseraceae: Bursera simaruba; Cannaceae: Canna coccinea; Chrysobalanaceae: 

Chrysobalanus icaco; Cyperaceae: Carex sp., Cyperus globosus, Cyperus odoratus, Cyperus ovularis, 

Cyperus rotundus, Cyperus sp.; Cyperus strigosus; Ebenaceae: Diospyros khaki; Fabaceae: Calliandra 

haematocephala; Guttiferae: Calophyllum antillanum; Lauraceae: Persea americana; Liliaceae: 

Gloriosa superba; Malpighiaceae: Malpighia emarginata; Malvaceae: Sida rhombifolia; Meliaceae: 

Khaya nyasica, Swietenia sp.; Musaceae: Musa x paradisiaca; Myricaceae: Myrica cerifera; 

Myrtaceae: Eugenia uniflora, Feijoa sellowiana, Psidium cattleianum, Psidium guajava, Psidium sp.; 

Nyctaginaceae: Bougainvillea sp., Guapira discolor; Polygonaceae: Coccoloba diversifolia, 

Coccoloba uvifera; Rosaceae: Licania tomentosa, Rubus odoratus; Rubiaceae: Coffea arabica; 

Rutaceae: Citrus aurantifolia, Citrus aurantium, Citrus decumana, Citrus limon, Citrus sp., Citrus 

reticulata, Citrus sapota, Citrus sinensis, Citrus x paradisi, Citrus x tangelo, Citrofortunella 

microcarpa; Sapotaceae: Lucuma caimito, Manilkara sp., Manilkara zapota; Solanaceae: Solanum 

melongena; Verbenaceae: Avicennia germinans; Viscaceae: Phoradendron sp.; Zygophyllaceae: 

179

Guaiacum officinale. 

Natural enemies: Fungus - Aegerita webberi, Aschersonia aleyrodis; Predators - Acari/ Phytoseiidae 

Euseius stipulatus, Typhlodromus phialatus; Araneae/Dictynidae: Phantyna segregata; Coleoptera/ 

Coccinellidae: Adalia bipunctata, Adonia variegata, Chilocorus bipustulatus, Clitostethus arcuatus, 

Coelophora pupillata, Cryptolaemus montrouzieri, Delphastus pallidus, Delphastus sp., Exochomus 

sp., Harmonia conglobata, Harmonia sedecimnotata, Nephaspis oculatus, Rhyzobius chrysomeloides, 

Serangium maculigerum; Diptera/ Syrphidae: Allograpta obliqua; Hemiptera/ Lygeidae: Geocoris 

sp.; Nabidae; Reduviidae: Rhinocorus iracundus;.Neuroptera/Chrysopidae: Ceraeochrysa cubana, 

Chrysopa s, Chrysoperla comanche; Coniopterygidae: Coniopteryx vicina, Conwentzia psociformis, 

Semidalis aleyrodiformis; Thysanoptera/ Phlaeothripidae: Haplothrips merrilli. Parasitoids - 

Hymenoptera Aphelinidae: Cales noacki, Dirphys larensis; Encarsia aleurothrixi, Encarsia 

americana, Encarsia basicincta, Encarsia brasiliensis, Encarsia cubensis, Encarsia desantisi, 

Encarsia dispersa, Encarsia dominicana, Encarsia haitiensis, Encarsia hispida, Encarsia luteola, 

Encarsia nigricephala, Encarsia noahi, Encarsia perplexa, Encarsia portoricensis, Encarsia 

quaintancei, Encarsia sophia, Encarsia tabacivora, Encarsia variegata, Encarsiella noyesi, 

Encarsiella sp.; Eretmocerus aleurothrixus, Eretmocerus ampliatus, Eretmocerus comperei, 

Eretmocerus desantisi, Eretmocerus dozieri, Eretmocerus exilis, Eretmocerus gracilis, Eretmocerus 

haldemani, Eretmocerus jimenezi, Eretmocerus longiterebrus, Eretmocerus paulistus, Eretmocerus 

portoricensis; Encyrtidae: Metaphycus angustifrons, Metaphycus sp., Plagiomerus cyaneus; 

Eulophidae: Euderomphale mexicana, Neopomphale aleurothrixi; Platygastridae: Amitus sp., Amitus 

spiniferus; Signiphoridae: Signiphora sp., Signiphora townsendi. 

US Port Interceptions: Colombia on Citrus sp.; Cuba on Citrus sp.; Dominican Republic on Morinda 

citrifolia, Pimenta racemosa; Ecuador on Alpinia purpurata, Alpinia sp., Citrus limon; El Salvador 

on Citrus sp., Mangifera indica; Guatemala on Citrus sp.; Hawaii on Citrus sinensis; India on Citrus 

sp.; Israel on Laurus nobilis; Italy on Citrus reticulata, Citrus sp.; Mexico on Citrus sp., Persea 

americana, Persea sp., Psidium guajava; Panama on unknown plant; Puerto Rico on Bursera 

simaruba, Citrus aurantiifolia, Citrus aurantium, Citrus limon, Citrus sp., Musa sp., Solanum nigrum; 

Singapore on Murraya koenigii, Citrus sp.; St. Lucia on Fabaceae; Viet Nam on unknown plant. 

Aleurothrixus guareae Costa Lima 1942 

Aleurothrixus guareae Costa Lima 1942c: 422. Holotype. Brazil: on Lucuma caimito. 


Host: Meliaceae: Guarea trichiliodes; Myrtaceae: Eugenia tomentosa; Sapotaceae: Achras sapota, 

Lucuma caimito. 

Aleurothrixus guimaraesi Costa Lima 1942 

Aleurothrixus guimaraesi Costa Lima 1942c: 422. Holotype. Brazil: on Achras sapota. 


Host: Sapotaceae: Achras sapota. 

180

Aleurothrixus interrogationis (Bemis) 1904 

Aleyrodes interrogationis Bemis 1904: 510. Syntypes. USA: California, on Ceanothus californicus, USNM. 

Aleurothrixus interrogationis (Bemis); Quaintance & Baker 1914: 104. 

Aleyrodes (Philodamus) interrogationis Bemis; Quaintance & Baker 1917: 402 


Host: Rhamnaceae: Ceanothus californicus, Ceanothus thyrsiflorus. 

Aleurothrixus lucumai Costa Lima 1942 

Aleurothrixus lucumai Costa Lima 1942c: 424. Syntypes. Brazil: on Achras sapota and Lucuma caimito. 


Host: Sapotaceae: Achras sapota, Lucuma caimito. 

Aleurothrixus miconiae Hempel 1923 

Aleurothrixus miconiae Hempel 1923: 1189. Syntypes. Brazil: on Miconia chartacea, MZU. 


Host: Melastomataceae: Miconia chartacea. 

Aleurothrixus myrtacei Bondar 1923 

Aleurothrixus myrtacei Bondar 1923a: 176. Syntypes. Brazil: on Myrtaceae, BMNH. 

Distribution: 2: Barbados, Brazil; Colombia; Cuba, Grenada, Guadeloupe, Guatemala, Honduras, Puerto Rico, 

Suriname, St. Lucia, Trinidad & Tobago, Venezuela. 

Host: Rubiaceae: Gardenia sp, Gardenia augusta; Myrtaceae: Eugenia sp., Psidium guajava. 

US Port Interceptions: Costa Rica on Pimenta racemosa; Dominican Republic on Bucida buceras; 

Guyana on an unidentified plant; Jamaica on Cinnamomum sp.; St. Kitts and Nevi on Pimenta sp.; 

St. Lucia on Laurus nobilis, Ocimum basilicum, Pimenta racemosa; Trinidad and Tobago on 

Annona sp., Laurus sp., Myrtaceae: Pimenta racemosa, Pimenta sp. 

Aleurothrixus ondinae Bondar 1923 

Aleurothrixus ondinae Bondar 1923a: 174. Syntypes. Brazil: on "ondina". 


Host: on "ondina". 

Aleurothrixus porteri Quaintance & Baker 1916 

Aleurothrixus porteri Quaintance & Baker 1916: 466. Syntypes. Chile: on Solanaceae, USNM; Baker & 

181

Moles 1923: 634. 

Distribution: 2: Argentina; Brazil; Chile. 

Host: Anacardiaceae: Lithrea caustica, Lithrea molle, Schinus dependens, Schinus molle; Lauraceae: Persea 

americana; Myrtaceae: Myrtus sp., Myrciaria cauliflora; Rutaceae: Citrus sp.; Solanaceae: Cestrum 

parqui; Verbenaceae: Lippia citriodora. 

Natural enemies: Aphelinidae: Cales noacki, Encarsia hispida. 

Aleurothrixus proximans Bondar 1923 

Aleurothrixus proximans Bondar 1923a: 170. Syntypes. Brazil: on Lauraceae, MZU; Costa Lima 1968: 115. 


Host: Lauraceae: Laurus nobilis, Persea americana. 

Aleurothrixus silvestri Corbett 1935 

Aleurothrixus silvestri Corbett 1935b: 811. Syntypes. Malaya: on unidentified host. 



Aleurothrixus similis Sampson & Drews 1941 

Aleurothrixus similis Sampson & Drews 1941: 161. Syntypes. Mexico: on mistletoe growing on Acacia sp., 

UCD. 



Aleurothrixus smilaceti Takahashi 1934 

Aleurothrixus smilaceti Takahashi 1934b: 63. Syntypes. Taiwan: on Smilax sp., TARI. 


Host: Liliaceae: Smilax china, Smilax sp. 

Natural enemies: Aphelinidae: Encarsia bothrocera, Encarsia perflava. 

Aleurothrixus solani Bondar 1923 

Aleurothrixus solani Bondar 1923a: 173. Syntypes. Brazil: on Solanaceae, MZU. 


Host: Solanaceae. 

182

Aleurotithius Quaintance & Baker 1914 

Aleurotithius Quaintance & Baker 1914: 106. Type species. Aleurotithium timberlakei Quaintance & Baker 



Species 

NA NT WPA AF EPA O AU PI HI 

mexicanus 

timberlakei 

Total = 2 species 

x 

x 

1 1 0 0 0 0 0 0 0 

Aleurotithius mexicanus Russell 1947 

Aleurotithius mexicanus Russell 1947: 42-44. Holotype and paratypes. Mexico: no host information, USNM. 



Aleurotithius timberlakei Quaintance & Baker 1914 

Aleurotithius timberlakei Quaintance & Baker 1914: 106. Lectotype (designated by Russell 1947: 41). USA: 

California, on Eriodictyon crassifolium, USNM 


Host: Hydrophyllaceae: Eriodictyon crassifolium, Eriodictyon tomentosum. 

183

Aleurotrachelus Quaintance & Baker 1914 

Aleurotrachelus Quaintance & Baker 1914: 103. Type species: Aleurodes tracheifer Quaintance 1900, by 


Aleyrodes (Frauenfeldiella) Gomez-Menor 1943: 188. Type species. Aleurodes jelineki Frauenfeld 1867, by 

monotypy; nec. Frauenfeldiella ruebsaamen (Diptera), 1905. 

Frauenfeldiella Gomez-Menor; Sampson & Drews 1956: 694. 

Luederwaldtiana Hempel 1923: 1185. Type species. Luederwaldtiana eriosemae Hempel 1923, by monotypy; 

synonymy according to Martin 2005: 22. 




alpinus x 

ambrensis x 

ampullatus x 

anonae x 

asparagi x 

atratus x x x 

binkae x 

brazzavillense x 

caerulescens x 

camamuensis x 

camelliae x 

chikungensis x 

cinnamomi x 

coimbatorensis x 

corbetti x 

184


debregeasiae x 

distinctus x 

dryandrae x 

duplicatus x 

elatostemae x 

eriosemae x 

erythrinae x 

euphorifoliae x 

fenestellae x 

filamentosus x 

fissistigmae x 

globulariae x 

goaensis x 

granosus x 

gratiosus x 

grewiae x 

hazomiavonae x 

ingafolii x 

ishigakiensis x 

joholensis x 

juiyunensis x 

limbatus x 

longispinus x 

185


lumpurensis x 

machili x 

madagascariensis x 

maesae x 

marginatus x 

mauritiensis x 

mesuae x 

minimus x 

minutus x 

multipapillus x x 

nivetae x 

obscurus x 

orchidicola x 

oriani x 

pandani x 

papilliferus x 

parvus x 

pauliani x 

plectroniae x 

primitus x 

pykarae x 

pyracanthae x 

reunionensis x 

rhamnicola x 

186


rosarius x 

rotundus x 

rubi x 

rubromaculatus x 

selangorensis x 

serratus x 

socialis x 

souliei x 

stypheliae x 

taiwanus x 

tarennae x 

theobromae x 

tracheifer x 

trachoides x x x x x 

tuberculatus x x x 

urticicola x 

vitis x x 

wendlandiae x 

zonatus x x 

Total = 83 species 2 18 4 22 18 24 3 2 1 

Aleurotrachelus alpinus Takahashi 1940 

Aleurotrachelus alpinus Takahashi 1940a: 29. Syntypes. Taiwan: on Rubus sp. 

Distribution: 5: Taiwan 

Host: Rosaceae: Rubus sp. 

187

Aleurotrachelus ambrensis Takahashi & Mamet 1952 

Aleurotrachelus ambrensis Takahashi & Mamet 1952b: 128. Syntypes. Madagascar: on undetermined tree. 



Aleurotrachelus ampullatus Bink-Moenen 1983 

Aleurotrachelus ampullatus Bink-Moenen 1983: 71. Holotype pupal case, Chad, Boro, 6.xii.1973, FAB & 

RMBM, on Detarium microcarpum; BHNM. 


Host: Fabaceae: Detarium microcarpus. 

Aleurotrachelus anonae Corbett 1935 

Aleurotrachelus anonae Corbett 1935b: 802. Syntypes. Malaya: on Annona squamosa, Morus indica and 

Zingiber sp. 

Distribution: 6: Bangladesh, Malaya; Malaysia. 

Host: Annonaceae: Annona squamosa; Moraceae: Morus indica; Zingiberaceae: Zingiber sp. 

US Port Interceptions: Indonesia on Annona sp.; Thailand on Annona sp. 

Aleurotrachelus asparagi (Lewis) 1893 

Aleurodes asparagi Lewis 1893: 88. Syntypes. Natal [South Africa]: on Asparagus sp. 

Tetraleurodes asparagi (Lewis) Quaintance & Baker 1914: 108. 

Aleurotrachelus asparagi (Lewis); Bink-Moenen 1983: 72. 

Distribution: 4: South Africa. 

Host: Liliaceae: Asparagus sp. 

Aleurotrachelus atratus Hempel 1922 

Aleurotrachelus atratus Hempel 1922: 3. Syntypes. Brazil: on Cocos nucifera. 

Distribution: 1: USA (FL); 2: Bahamas; Brazil; Colombia, Guadeloupe, Mexico, Puerto Rico, St. Lucia, 

Trinidad & Tobago, Venezuela, Virgin Islands, West Indies; 4: Uganda, Comoros Islands. 

Host: Arecaceae: Carpentaria acuminata; Chamaedorea erumpens, Cocos nucifera, Ptychosperma elegans, 

Roystonea regia, Syagrus romanzoffiana, Veitchia merrillii; Solanaceae: Solanum melongena L. var. 

esculentum. 

US Port Interceptions: Bahamas on Cocos nucifera; Costa Rica on Livistona sp.; Grenada on Arecaceae; 

Mexico on Persea sp., Piperaceae; Puerto Rico on Areca sp., Cocos nucifera; St. Lucia on Areca sp, 

Cocos nucifera; US Virgin Islands on Cocos nucifera. 

188

Aleurotrachelus binkae Jesudasan & David 1991 

Aleurotrachelus binkae Jesudasan and David 1991: 286. Holotype pupal case. India: on Ecbolium linnearum. 


Host: Cucurbitaceae: Ecballium linnearu. 

Aleurotrachelus brazzavillense Cohic 1968 

Aleurotrachelus brazzavillense Cohic 1968a: 35. Syntypes (15). Congo: on Colletoecema dewevrei; Cohic 

1968b: 113; Bink-Moenen 1983: 73. 


Host: Connaraceae: Agelaea dewevrei; Rubiaceae: Colletoecema dewevrei. 

Aleurotrachelus caerulescens Singh 1931 

Aleurotrachelus caerulescens Singh 1931: 59. Syntypes. India: on Artocarpus integrifolia; 

Takahashi 1935b: 39. 


Host: Araceae: Alocasia macrorrhizos; Cucurbitaceae: Thladiantha dubia; Euphorbiaceae: Bischofia javanica. 

Flacourtiaceae: Scolopia oldhami; Liliaceae: Smilax china; Lythraceae: Lagerstroemia indica; 

Moraceae: Artocarpus integrifolia; Myricaceae: Myrica rubra; Rubiaceae: Gardenia florida; 

Salicaceae: Salix sp.; Symplocaceae: Symplocos laurina; Ulmaceae: Celtis sinensis. 

Aleurotrachelus camamuensis Bondar 1923 

Aleurotrachelus camamuensis Bondar 1923a: 153. Syntypes. Brazil: on unidentified host. 

Distribution: Brazil. 

Host: unidentified plant 

Aleurotrachelus camelliae (Kuwana) 1911 

Aleyrodes camelliae Kuwana 1911: 625. Syntypes. Japan: on Thea japonica. 

Aleurotrachelus camelliae (Kuwana); Quaintance & Baker 1914: 103. 

Distribution: 5: China, Hong Kong; Japan. 

Host: Theaceae: Camellia japonica, Camellia sinensis, Camellia sinensis var. sinensis, Thea japonica. 

Aleurotrachelus chikungensis Mound & Halsey 1978 

Aleurotrachelus parvus Young 1944: 136. Syntypes. China: on unidentified host. [name preoccupied by A. 

parvus (Hempel) 

189

190 

Aleurotrachelus chikungensis Mound and Halsey 1978: 68. Syntypes. China: on unidentified host. [new name 

for A. parvus Young, name preoccupied by A. parvus (Hempel) 



Aleurotrachelus cinnamomi Jesudasan & David 1991 

Aleurotrachelus cinnamomi Jesudasan & David 1991 Jesudasan and David 1991: 287. Holotype pupal case. 

India: on Cinnamomum malabaricum. 


Host: Lauraceae: Cinnamomum malabaricum. 

Aleurotrachelus coimbatorensis David & Subramaniam 1976 

Aleurotrachelus coimbatorensis David & Subramaniam 1976: 168-170. Holotype. India: Coimbatore, 

21.iii.1957, S.K. David, on Jasminum auriculatum. 


Host: Rubiaceae: Jasminum auriculatum. 

Aleurotrachelus corbetti Takahashi 1941 

Aleurotrachelus tuberculatus Corbett 1935b: 804-805. Syntypes. Malaya: Kuala Lumpur, on an 

unidentified host. 

Aleurotrachelus corbetti Takahashi 1941c: 392.[Replacement name for Aleurotrachelus tuberculatus Corbett 

1935b, nec. Aleurotrachelus tuberculatus Singh, 1933: 343. 



Aleurotrachelus debregeasiae Young 1944 

Aleurotrachelus debregeasiae Young 1944: 138-139. Syntypes. China: Szechwan Province, North Hot 

Springs of Pehpei, 7 I 1942, Kao-Keng-Yen, 13.i.1942; Mound and Halsey 1978. 


Host: Urticae: Debregeasia edulis. 

Aleurotrachelus distinctus Hempel 1923 

Aleurotrachelus distinctus Hempel 1923: 1182-1183. Syntypes. Brazil: Blumenau, Sana Catharina, vi.1919, 

H. Luederwaldt, on forest shrub. 

Distribution: 2: Brazil.

Host: Solanaceae: Solanum sp. 

Aleurotrachelus dryandrae Solomon 1935 

Aleurotrachelus dryandrae Solomon 1935: 83-86. Syntypes. Australia: Western Australia, Perth District, 

Crawley, on Dryandra floribunda; Mound and Halsey 1978: 69.; Martin 1999. 


Host: Proteaceae: Banksia attenuata, Banksia grandis, Dryandra floribunda, Dryandra sessilis, Grevillea 

bispinosa, Hakea prostrata, Hakea varia. 

US Port Interceptions: Australia on Banksia hookeriana, Leucadendron sp., Persoonia longiflora, Persoonia 

sp.; Protea sp. 

Aleurotrachelus duplicatus Bink-Moenen 1983 

Aleurotrachelus duplicatus Bink-Moenen 1983: 73. Holotype pupal case, Chad, Bebedjia, 28.xi.1973, FAB & 

RMBM, on Mitragyna inermis, BHNM. 


Host: Rubiaceae: Crossopteryx febrifuga, Mitragyna inermis. 

Aleurotrachelus elatostemae Takahashi 1932 

Aleurotrachelus elatostemae Takahashi 1932: 42. Syntypes. Taiwan: on Elatostema lineolatum.; Takahashi 

1935b: 41. 


Host: Solanaceae: Solanum sp.; Urticaceae: Elatostema edule, Elatostema lineolatum, Oreocnide pedinculata. 

Aleurotrachelus eriosemae (Hempel) 1923 

Luederwaldtiana eriosemae Hempel 1923: 1186. Syntypes. Brazil: on Eriosema heterophyllum. 

Aleurotrachelus eriosemae (Hempel); Martin 2005b: 22. 


Host: Fabaceae: Eriosema heterophyllum. 

Aleurotrachelus erythrinae Corbett 1935 

Aleurotrachelus erythrinae Corbett 1935b: 805. Syntypes. Malaya: on Erythrina stricta. 


Host: Fabaceae: Centrosema plumieri, Erythrina stricta. 

Aleurotrachelus euphorifoliae Young 1944 

191

Aleurotrachelus euphorifoliae Young 1944: 139. Syntypes. China: on Euphoria longana. 


Host: Euphorbiaceae: Euphorbia longana. 

Aleurotrachelus fenestellae Hempel 1923 

Aleurotrachelus fenestellae Hempel 1923: 1183. Syntypes. Brazil: on Baccharis genistelloides. 


Host: Asteraceae: Baccharis genistelloides. 

Aleurotrachelus filamentosus Takahashi 1938 

Aleurotrachelus filamentosus Takahashi 1938a: 27. Syntypes. Mauritius: on undetermined tree; Takahashi 

1939c. 


Host: Sapotaceae: Labourdonnaisia calophylloides; unidentified tree. 

Aleurotrachelus fissistigmae Takahashi 1931 

Aleurotrachelus fissistigmae Takahashi 1931b: 264. Syntypes. Taiwan: on Fissistigma oldhami. 


Host: Annonaceae: Fissistigma oldhami. 

Aleurotrachelus globulariae Goux 1942 

Aleurotrachelus globulariae Goux 1942: 145. Syntypes. France: on Globularia alypum. 

Distribution: 3: France; Iran, Israel. 

Host: Globulariceae: Globularia alypum. 

Aleurotrachelus goaensis Jesudasan & David 1991 

Aleurotrachelus goaensis Jesudasan and David 1991: 288. Holotype pupal case. India: on unidentified tree. 



Aleurotrachelus granosus Bondar 1923 

Aleurotrachelus granosus Bondar 1923a: 155. Syntypes. Brazil: on Theobroma cacao; Costa Lima 1968. 


Host: Musaceae: Musa sp.; Sterculiaceae: Theobroma cacao. 

192

Aleurotrachelus gratiosus Bondar 1923 

Aleurotrachelus gratiosus Bondar 1923a: 146. Syntypes. Brazil: on Lauraceae; Costa Lima 1968. 



Aleurotrachelus grewiae Takahashi 1952 

Aleurotrachelus grewiae Takahashi 1952c: 25. Syntypes. Malaya: on Grewia tomentosa. 


Host: Tiliaceae: Grewia tomentosa. 

Aleurotrachelus hazomiavonae Takahashi 1955 

Aleurotrachelus hazomiavonae Takahashi 1955a: 408. Syntypes. Madagascar: on "hazomiavona" a parasitic 

Loranthaceae 



Aleurotrachelus ingafolii Bondar 1923 

Aleurotrachelus ingafolii Bondar 1923a: 148. Syntypes. Brazil: on Inga sp. 

Distribution: 2: Brazil, Venezuela. 

Host: Fabaceae: Inga sp. 

Aleurotrachelus ishigakiensis (Takahashi) 1933 

Trialeurodes ishigakiensis Takahashi 1933: 21. Syntypes. Japan: on Morus alba. 

Aleurotrachelus ishigakiensis (Takahashi); Takahashi 1951b: 21. 


Host: Araliaceae: Gilibertia trifida, Hedera rhombea; Cornaceae: Cornus sp.; Daphniphyllaceae: 

Daphniphyllum macropodum; Euphorbiaceae: Sapium japonicum; Fabaceae: Pueraria hirsuta. 

Lauraceae: Lindera obtusiloba; Moraceae: Ficus erecta, Morus alba, Morus bombycis; 

Pittosporaceae: Pittosporum tobira, Pittosporum tobira 'variegata' ; Theaceae: Eurya japonica; 

Umbelliferae: Heracleum lanatum. 

US Port Interception: Japan on Eurya japonica. 

Aleurotrachelus joholensis Corbett 1935 

193

Aleurotrachelus joholensis Corbett 1935b: 809. Syntypes. Malaya: on unidentified host. 



Aleurotrachelus juiyunensis Young 1944 

Aleurotrachelus juiyunensis Young 1944: 135. Syntypes. China: on two undetermined plants. 



Aleurotrachelus limbatus (Maskell) 1895 

Aleurodes limbatus Maskell 1895: 436. Lectotype (designated by Dumbleton 1956b:170). Australia: on 

Acacia longifolia. 

Aleurotrachelus limbatus (Maskell); Quaintance & Baker 1914: 103. 


Host: Epacridaceae: Leucopogon juniperus ;Styphelia adscendens; Fabaceae: Acacia longifolia. 

Aleurotrachelus longispinus Corbett 1926 

Aleurotrachelus longispinus Corbett 1926: 277. Syntypes. Sri Lanka: on unidentified host; Singh 1931. 


Host: Rubiaceae: Ixora coccinea, unidentified host. 

Aleurotrachelus lumparensis Corbett 1935 

Aleurotrachelus lumpurensis Corbett 1935b: 807. Syntypes. Malaya: Kuala Lumpur, on Nephelium 



Aleurotrachelus machili Takahashi 1942 

Aleurotrachelus machili Takahashi 1942b: 168. Syntypes. Thailand: on Machilus sp. 


Host: Lauraceae: Machilus sp. 

Aleurotrachelus madagascariensis Takahashi 1955 

Aleurotrachelus madagascariensis Takahashi 1955a: 409. Syntypes. Madagascar: on unidentified host 

194


Host: unidentified host. 

Aleurotrachelus maesae Takahashi 1935 

Aleurotrachelus maesae Takahashi 1935b: 57. Holotype. Taiwan: on Maesa formosana. 

Distribution: 5: China; Hong Kong; Taiwan. 

Host: Myrsinaceae: Maesa formosana. 

Aleurotrachelus marginatus (Newstead) 1911 

Aleurodes marginata Newstead 1911: 172. Syntypes. Tanzania, Sigithal, near Amani, 4.viii.1902, A. 

Zimmerman, on forest tree; BMNH. 

Tetraleurodes marginata (Newstead); Quaintance & Baker 1914: 108. 

Aleurotrachelus marginatus (Newstead); Bink-Moenen 1983: 75. 

Distribution: 4: Tanzania. 

Host: unidentified tree 

Aleurotrachelus mauritiensis Takahashi 1940 

Aleurotrachelus mauritiensis Takahashi 1940b: 43. Syntypes. Mauritius: on undetermined tree. 



Aleurotrachelus mesuae Corbett 1935 

Aleurotrachelus mesuae Corbett 1935b: 803. Syntypes. Malaya: on Mesua ferrea and Adiobotrys 

atroparpureus. 


Host: Fabaceae: Adinobotrys atropurpureus; Guttiferae: Mesua ferrea. 

Aleurotrachelus minimus Young 1944 

Aleurotrachelus minimus Young 1944: 137. Syntypes. China: on undetermined host. 



Aleurotrachelus minutus Takahashi 1952 

Aleurotrachelus minutus Takahashi 1952c: 25. Syntypes. Malaya: on Ficus sp. 

195

196 



Aleurotrachelus multipapillus Singh 1932 

Aleurotrachelus multipapillus Singh 1932: 86. Syntypes. Burma: on Bambusa nana; Martin 1988: 69. 

Distribution: 5: China; 6: Burma; India; Malaysia; Thailand; 7: Sulawesi. 

Host: Poaceae: Bambusa nana. 

Aleurotrachelus nivetae Cohic 1969 

Aleurotrachelus nivetae Cohic 1969: 89. Holotype. Ivory Coast: on Sansevieria trifasciata var laurentii; Bink- 

Moenen 1983: 75. 


Host: Liliaceae: Sansevieria trifasciata, Sansevieria trifasciata var laurentii. 

Aleurotrachelus obscurus Bink-Moenen 1983 

Aleurotrachelus obscurus Bink-Moenen 1983: 76. Holotype pupal case, Chad, Boro, 6.xii.1973, FAB & 

RMBM, on Daniellia olivera; BMNH. 


Host: Fabaceae: Burkea africana, Daniellia olivera, Detarium microcarpum, Prosopsis africana. 

Aleurotrachelus orchidicola Takahashi 1939 

Aleurotrachelus orchidicola Takahashi 1939c: 1. Syntypes. Mauritius: on orchid. 


Host: Orchidaceae: an orchid. 

Aleurotrachelus oriani Martin & Mound 2007 

Aleurotrachelus pauliani Takahashi 1961: 336. Syntypes. Reunion Island: on unidentified host; name 

preoccupied by Aleurotrachelus pauliani Takahashi 1960. 

Aleurotrachelus oriani Martin & Mound 2007 



Aleurotrachelus pandani Takahashi 1951 

Aleurotrachelus pandani Takahashi 1951a: 380. Syntypes. Mauritius: on Pandanus sp. 

Distribution: 4: Mauritius.

Host: Pandanaceae: Pandanus sp. 

Aleurotrachelus papilliferus (Sampson & Drews) 1941 

Tetraleurodes papilliferus Sampson & Drews 1941: 176. Syntypes. Panama: on undetermined tree. 

Aleurotrachelus papilliferus (Sampson & Drews); Nakahara 1995: 136. 



Aleurotrachelus parvus (Hempel) 1899 

Aleurodes parvus Hempel 1899: 295. Syntypes. Brazil: on Maytenus sp. 

Aleurotrachelus parvus (Hempel); Quaintance & Baker 1914: 103. 


Host: Celastraceae: Maytenus sp. 

Aleurotrachelus pauliani Takahashi 1961 

Aleurotrachelus pauliani Takahashi 1960b: 151. Syntypes. Reunion Island: on unidentified host. 



Aleurotrachelus plectroniae Takahashi 1955 

Aleurotrachelus plectroniae Takahashi 1955a: 412. Syntypes. Madagascar: on Plectronia thoursi. 


Host: Rubiaceae: Plectronia thouarsi. 

Aleurotrachelus primitus Young 1944 

Aleurotrachelus primitus Young 1944: 135. Syntypes. China: on undetermined plant. 



Aleurotrachelus pykarae Jesudasan & David 1991 

Aleurotrachelus pykarae Jesudasan and David 1991: 289. Holotype pupal case. India: on Glochidion sp. 


Host: Euphorbiaceae: Glochidion sp. 

197

Aleurotrachelus pyracanthae Takahashi 1935 

Aleurotrachelus pyracanthae Takahashi 1935b: 58. Syntypes. Taiwan: on Pyracantha koidzumi. 


Host: Rosaceae: Pyracantha koidzumi 

Aleurotrachelus reunionensis Takahashi 1960 

Aleurotrachelus reunionensis Takahashi 1960b: 149. Syntypes. Reunion Island: on undetermined plant. 



Aleurotrachelus rhamnicola (Goux) 1940 

Aleyrodes rhamnicola Goux 1940 Goux 1940: -. Syntypes. France; on Rhamnus alaternus. 

Aleurotrachelus rhamnicola (Goux); Martin et al. 1996: 124. 

Aleurotrachelus espunae Gomez-Menor 1945a: 298. Syntypes. Spain: on Berberis sp.; Martin year? 

Distribution: 3: Corsica; France; Greece; Iran, Italy; Madeira; Malta; Morocco; Sicily; Spain. 

Host: Berberidaceae: Berberis integerrima, Berberis sp.; Ericaceae: Arbutus unedo; Fagaceae: Quercus 

calliprinos, Quercus rotundifolia, Quercus sp.; Passifloraceae: Passiflora edulis; Ranunculaceae: 

Clematis vitalba; Rhamnaceae: Rhamnus alaternus; Rosaceae: Rosa sp., Rubus fruticosus, Rubus sp.; 

Vitaceae: Ampelopsis sp. 

Aleurotrachelus rosarius Bondar 1923 

Aleurotrachelus rosarius Bondar 1923a: 150. Syntypes. Brazil: on Psidium guajava. 

Distribution: 2: Brazil, Colombia. 


Aleurotrachelus rotundus Corbett 1935 

Aleurotrachelus rotundus Corbett 1935b: 808. Syntypes. Malaya: on Adinobotrys atropurpureus. 


Host: Fabaceae: Adinobotrys atropurpureus. 

Aleurotrachelus rubi Takahashi 1933 

Aleurotrachelus rubi Takahashi 1933: 16. Syntypes. Taiwan: on Rubus sp. 


Host: Rosaceae: Rubus sp.; Boehmeria nivea. 

198

Aleurotrachelus rubromaculatus Bondar 1923 

Aleurotrachelus rubromaculatus Bondar 1923a: 150. Syntypes. Brazil: on Compositae. 


Host: Asteraceae. 

Aleurotrachelus selangorensis Corbett 1935 

Aleurotrachelus selangorensis Corbett 1935b: 801. Syntypes. Malaya: on Diospyros sp. 


Host: Ebenaceae: Diospyros sp. 

Aleurotrachelus serratus Takahashi 1949 

Aleurotrachelus serratus Takahashi 1949: 52. Syntypes. Rioux (Riau) Islands (south of Singapore): on wild 

palm. 

Distribution: 8: Rioux (Riau) Islands. 

Host: Arecaceae. 

Aleurotrachelus socialis Bondar 1923 

Aleurotrachelus socialis Bondar 1923a: 145. Syntypes. Brazil: on Cecropia sp.; Martin 2005b: 23. 

Distribution - 2: Belize, Brazil; Colombia; Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Panama, 

Venezuela. 

Host: Bignoniaceae: Bignoniaceae: sp.; Clusiaceae: Calophyllum sp.; Dilleniaceae: Tetracera sp.; 

Euphorbiaceae: Manihot caerulescens, Manihot esculenta; Lauraceae: Persea sp.; Moraceae: Cecropia 

sp. ; Myrtaceae: Pimenta sp.; Piperaceae: Piper sp.; Polygonaceae: Coccoloba sp. 

Aleurotrachelus souliei Cohic 1969 

Aleurotrachelus souliei Cohic 1969 Cohic 1969: 94. Holotype. Ivory Coast: on Tabernaemontana crassa; 

Ardaillon & Cohic 1970; Bink-Moenen 1983: 77. 

Distribution: 4: Ivory Coast, Liberia, Nigeria. 

Host: Apocynaceae: Pleiocarpa mutica, Pleiocarpa mutida, Tabernaemontana crassa; Lecythidaceae: 

Napoleona leonensis 

Aleurotrachelus stypheliae (Maskell) 1895 

Aleurodes stypheliae Maskell 1895: 442. Lectotype. Australia: on Styphelia (Monotoca) richei. 

Tetraleurodes stypheliae (Maskell) Quaintance & Baker 1914: 108. Lectotype. Australia: on Styphelia 

199

(Monotoca) richei 

Aleurotrachelus stypheliae (Maskell); Martin 1999: 53. 


Host: Epacridaceae: Leucopogon richei; Styphelia richei. 

Aleurotrachelus taiwanus Takahashi 1932 

Aleurotrachelus taiwanus Takahashi 1932: 45. Syntypes. Taiwan: on Pachyrrhizus erosus. 


Host: Fabaceae: Pachyrhizus erosus. 

Aleurotrachelus tarennae Bink-Moenen 1983 

Aleurotrachelus tarennae Bink-Moenen 1983: 78. Holotype pupal case, Chad, Timberi, 29.xii.1973, FAB & 

RMBM, on Tarenna sp.; BMNH. 


Host: Rubiaceae: Tarenna sp. 

Aleurotrachelus theobromae Bondar 1923 

Aleurotrachelus theobromae Bondar 1923a. Syntypes. Brazil: on Theobroma cacao and Anacardium 

occidentale. 

Distribution: 2: Brazil; Guyana. 

Host: Anacardiaceae: Anacardium occidentale; Sterculiaceae: Theobroma cacao. 

Aleurotrachelus tracheifer (Quaintance) 1900 

Aleurodes tracheifer Quaintance 1900: 38. Syntypes. Mexico: Tabasco, on "escabillo". 

Aleurotrachelus tracheifer (Quaintance); Quaintance & Baker 1914: 39; Martin 2005b. 

Distribution: 2: Belize, Colombia, Mexico, Panama. 

Host: Burseraceae: Protium sp.; Loganiaceae: Strychnos sp.; Moraceae; Myrtaceae: Eugenia sp.; 

Polygonaceae: Coccoloba. 

Aleurotrachelus trachoides (Back) 1912 

Aleyrodes trachoides Back 1912 Back 1912: 151. Syntypes. Cuba: on Solanum seaphorthianum. 

Aleurotrachelus trachoides (Back); Quaintance & Baker 1914: 103. 

Aleurotulus bodkini Baker & Moles 1923: 635. Syntypes. Guyana: on ornamental plant; synonymy according 

to Mound and Halsey 1978: 77. 

200

201 

Distribution: 1: USA (CA, FL, HI, LA, TX); 2: Antigua; Bahamas, Barbados; Brazil, Colombia, Costa Rica, 

Cuba; Dominica, Dominican Republic, El Salvador, Guadeloupe; Guatemala, Guyana; Haiti, 

Honduras, Jamaica; Martinique, Mexico; Nicaragua, Panama, Peru, Puerto Rico; Suriname, Trinidad; 

Venezuela, Virgin Islands; 4: Gambia, Reunion Island; 8: Fiji, Guam, Tahiti; 9: Hawaii. 

Host: Annonaceae: Annona muricata, Annona reticulata; Apocynaceae: Tabernaemontana divaricata; 

Araceae: Syngonium podophyllum, Xanthosoma sagittifolium; Arecaceae: Chamaedorea elegans, 

Chamaedorea sp., Cocos nucifera, Dypsis lutescens, Veitchia merrillii; Asteraceae: Bidens pilosa, 

Mikania cordifolia; Bignoniaceae: Tabebuia glomerata, Tabebuia pallida; Boraginaceae: Cordia 

collococca ; Cannaceae: Canna coccinea; Casuarinaceae: Casuarina sp.; Chrysobalanaceae: Licania 

michauxii; Convolvulaceae: Ipomoea batatas; Ipomoea sp., Merremia sp.; Dioscoreaceae: Dioscorea 

sp.; Fabaceae: Bauhinia divaricata, Canavalia ensiformis, Leucaena sp.; Guttiferae Calophyllum 

antillanum; Hypericum hypericoides; Lauraceae: Persea americana; Malvaceae: Hibiscus elatus; 

Melastomataceae: Miconia magnifolia; Moraceae: Ficus membranaceae, Ficus retusa; Myrsinaceae: 

Ardisia escallonioides; Myrtaceae: Psidium guajava; Phytolaccaceae: Petiveria alliacea; 

Polygonaceae: Coccoloba uvifera; Rubiaceae: Morinda citrifolia, Psychotria nervosa, Randia 

aculeata; Rutaceae: Citrus limon, Cleome sp.; Sapotaceae: Pouteria sapota; Solanaceae: Capsicum 

annuum, Capsicum frutescens, Capsicum sp, Cestrum nocturnum. Datura stramonium, : Lycopersicon 

esculentum, Lycopersicon sp., Nicotiana sp, Solanum melongena, Solanum nigrum, Solanum 

seaforthianum, Solanum sp., Solanum torvum; Sterculiaceae: Theobroma cacao; Verbenaceae: 

Citharexylum sp., Tectona grandis; unidentified plant. 

US Port Interceptions: Dominican Republic on Capsicum sp.; Ecuador on Bromeliaceae, Oncidium sp.; 

Hawaii on Heckeria umbellata, Hibiscus rosa-sinensis, Ixora sp., Jasminum sp., Piper betle, Piper sp., 

Plumeria sp.; Jamaica on Annona muricata, herbs, Mexico on Capsicum sp., Chamaedorea elegans, 

Chamaedorea sp.; Puerto Rico on Capsicum sp., Citrus aurantium, Citrus sp., Mentha sp., Musa sp., 

Piper sp.; St. Lucia on Punica granatum; Trinidad and Tobago on Areca sp.; US Virgin Islands on 

Coccoloba uvifera and herbs. 

Aleurotrachelus tuberculatus Singh 1933 

Aleurotrachelus tuberculatus Singh 1933: 343. Syntypes. Burma: on Ficus sp.; Takahashi 1935b; Takahashi 

1942b. 

Distribution: 4: Madagascar; 5:Hong Kong; Taiwan; 6: Burma; Cambodia; India; Thailand. 

Host: Fabaceae: Centrosema pubescens, Dalbergia sp.; Lauraceae: Lindera oldhami; Moraceae: Ficus sp. 

Aleurotrachelus urticicola Young 1944 

Aleurotrachelus urticicola Young 1944: 138. Syntypes. China: on Urticaceae. 


Host: Urticeae 

Aleurotrachelus vitis Corbett 1935 

Aleurotrachelus vitis Corbett 1935b: 806. Syntypes. Malaya: on Vitex sp. 

Distribution: 3: USSR; 6: Malaya; Malaysia.


Aleurotrachelus wendlandiae Jesudasan & David 1991 

Aleurotrachelus wendlandiae Jesudasan and David 1991: 290. Holotype pupal case. India: on Wendlandia 

thyrosoidea. 


Host: Wendlandia thyrosoidea. 

Aleurotrachelus zonatus Takahashi 1952 

Aleurotrachelus zonatus Takahashi 1952c: 24. Holotype. Malaya: on undetermined host. 

Distribution: 6: Greece; Malaya; Malaysia. 


Transfers, Synonyms and New Names 

Aleurotrachelus cacaorum Bondar 1923 to Tetraleurodes by Nakahara 1995 

Aleurotrachelus cercropiae Bondar 1923 to Aleuropleurocelus by Martin 2005: 20 

Aleurotrachelus espunae Gomez-Menor 1945 = A. rhamnicola 

Aleurotrachelus micheliae Takahashi 1932 to Crenidorsum 

Aleurotrachelus myrtifolii Bondar 1923 to Aleurothrixus; Martin 2005 

Aleurotrachelus parvus Young 1944 to Aleurotrachelus chikungensis Mound & Halsey 1978 

Aleurotrachelus tuberculatus Corbett 1935b to Aleurotrachelus corbetti Takahashi 1941 

Aleurotrachelus turpiniae Takahashi 1932 to Crenidorsum 

202

Aleurotuba Tremblay and Iaccarino 1978 

Aleurotuba Tremblay and Iaccarino 1978: 60. Type species - Aleurodes jelineki Frauenfeld 1867. 



jelinekii x x x 

Total = 1 species 1 1 1 0 0 0 0 0 0 

Aleurotuba jelinekii (Frauenfeld) 1867 

Aleurodes jelinekii Frauenfeld 1867: 799. Syntypes. Italy: on Arbutus unedo. 

Aleurotrachelus jelinekii (Frauenfeld); Fowler 1954: 406. 

Aleyrodes (Frauenfeldiella) jelinekii (Frauenfeld); Gomez-Menor 1943: 188. 

Aleyrodes (Frauenfeldiella) jelinekii form alba; Gomez-Menor 1954: 373. 

Frauenfeldiella jelinekii (Frauenfeld); Sampson & Drews 1956: 694. 

Aleurotuba jelinekii (Frauenfeld) 1867 

Distribution: 1: USA (CA); 2: Colombia; 3: England; France; Germany; Greece; Iran, Italy; Netherlands, 

Portugal, Romania, Spain; Turkey; USSR; Yugoslavia. 

Host: Caprifoliaceae: Viburnum pyramidata, Viburnum rotundifolia, Viburnum tinus; Ericaceae: Arbutus 

unedo; Myrtaceae: Myrtus communis. 

Natural enemies: Aphelinidae: Encarsia aleurotubae, Encarsia margaritiventris, Encarsia tricolor, 

Eretmocerus longicornis; Cales noacki; Platygastridae: Amitus aleurotubae. Predators- Araneidae: 

Meta segmentata; Chamaemyidae: Leucopis bella, Coccinellidae: Clitostethus arcuatus, Menochilus 

sp.; Coniopterygidae: Conwentzia psociformis, Drosophilidae: Acletoxenus formosus; Miridae: 

Campyloneura virgula. 

US Port Interceptions: France on Viburnum sp.; Jamaica on Annona muricata; Spain on Persea americana; 

Netherlands. 

203

Aleurotulus Quaintance & Baker 1914 

Aleurotulus Quaintance & Baker 1914: 101. Type species Aleurodes nephrolepidis Quaintance 1900, by 




anthuricola x x 

arundinacea x x 

laneus x 

mundururu x 

nephrolepidis x x x 

pteridophytae x 

sp. x 

Total = 6+ species 1 5 1 1 0 1 2* 0 0 

Aleurotulus anthuricola Nakahara 1989 

Aleurotulus anthuricola Nakahara 1989: 436. Holotype pupal case, Colombia, intercepted at Miami, FL, 

31.vii.1978, F. D. Matthews, on Anthurium sp., USNM. 

Distribution: 1: USA (intercepted in CA, FL); 2: Barbados, Colombia, Dominica, Guadeloupe, Martinique, 

Trinidad & Tobago, Virgin Islands; 9 Hawaii. 

Host: Araceae: Anthurium andreanum, Anthurium palmarum. Anthurium sp. 

Aleurotulus arundinacea Singh 1931 

Aleurotulus arundinacea Singh 1931: 88. Syntypes. India: on Bambusa arundinacea, ZSI; Martin 1985: 321. 

Distribution: 6: India; 7: Papua New Guinea. 

Host: Poaceae: Bambusa sp., Bambusa arundinacea. 

Aleurotulus laneus Martin 2005 

Aleurotulus laneus Martin 2005b: 26. Holotype puparium, Belize, CFR, 22.III.2003, JH Martin, on Lasiacis 

sp, BMNH. 

204


Host: Poaceae: Lasiacis sp. 

Aleurotulus mundururu Bondar 1923 

Aleurotulus mundururu Bondar 1923a: 131. Syntypes. Brazil: on mundururu [Miconia sp.]. 

Distribution: 2: Belize, Brazil, Costa Rica, Ecuador, Guyana, Trinidad. 

Host: Loganiaceae: Strychnos sp., Melastomataceae: Miconia sp. 

Aleurotulus nephrolepidis (Quaintance) 1900 

Aleurodes nephrolepidis Quaintance 1900: 29. Syntypes. USA: Pennsylvania, on Nephrolepis sp., USNM. 

Aleurotulus nephrolepidis (Quaintance); Quaintance & Baker 1914: 29. 

Aleyrodes extraniens Bemis 1904: 526. Syntypes. USA: California, on Acrostichum capense, USNM; 


Aleurodes filicium Goeldi; misidentification in part; Douglas 1891a: 44. 

Aleuroplatus kewensis Trehan 1938: 183. Lectotype (designated by Mound 1966). England: on Nephrodium 

confluens, BMNH; Mound 1966: 40. 

Distribution: 1: USA (CA, FL, PA); 2: Brazil; 3: England; Hungary; Scotland; Spain; 4: Canary Islands; 

Madagascar. 

Host: Aspidiaceae: Nephrodium confluens, Stenosema aurita; Aspleniaceae: Asplenium cuneatum; 

Blechnaceae: Blechnum occidentale; Blechnum brasiliensis; Dryopteridaceae: Diplazium proliferum, 

Dryopteris flaccida, Nephrolepis sp., Oleandra africana, Oleandra articulata, Polystichum falcatum, 

Tectaria molle; Pteridaceae: Acrostichum capense, Pteris biaurita, Pteris togoensis; Schizaeaceae: 

Anemia sp.; Thelypteridaceae: Cyclosorus dentatus. 

Natural enemies: Aphelinidae: Cales noacki. 

Aleurotulus pteridophytae Martin 1994 

Aleurotulus pteridophytae Martin 1994, in Mound et al. 1994: 1405. Holotype pupal case, Costa Rica, north 

Heredia Province, La Selva, xi.1992, L. A. Mound, on Selaginella eurynota, BMNH. 

Distribution: 2: Colombia; Costa Rica; Ecuador; El Salvador. 

Host: Polypodiaceae: Adiantum sp., Schizaeaceae: Anemia phyllitidis, Selaginellaceae: Selaginella eurynota, 

fern. 

Natural enemies: Aphelinidae: Encarsia sueloderi. 

Aleurotulus sp. 

Collected in 7: Sulawesi on Dryopteridaceae: Nephrolepis sp. [ref: Martin 1988: 70]. 

205

Aleuroviggianus Iaccarino 1982 

Aleuroviggianus Iaccarino 1982: 37. Type species. Aleuroviggianus adrianae Iaccarino 1982, by original 

designation. 



adanaensis x 

adrianae x 

graecus x 

halperini x 

polymorphus x 

zonalus x 

Total = 6 species 0 0 6 0 0 0 0 0 0 

Aleuroviggianus adanaensis Bink-Moenen 1992 

Aleuroviggianus adanaensis Bink-Moenen 1992: 36. Holotype pupal case. Turkey: on Quercus coccifera, 

BMNH. 

Distribution: 3: Egypt; Greece; Israel; Rhodes; Turkey. 

Host: Fagaceae: Quercus coccifera, Quercus sp. 

Distribution: 3: Greece; Israel, Quercus calliprinos, Quercus sp. 

Aleuroviggianus adrianae Iaccarino 1982 

Aleuroviggianus adrianae Iaccarino 1982: 38. Holotype. Italy: Portici, on Quercus ilex, UNP. 

Distribution: 3: France; Greece; Iran, Italy; Spain. 

Host: Fagaceae: Quercus ilex, Quercus rotundifolia, Quercus suber. 

Natural enemies: Platygastridae: Amitus vesuvianus Viggiani & Mazzone 1982; Aphelinidae: Encarsia 

aleuroilicis. 

Aleuroviggianus graecus Bink-Moenen 1992 

Aleuroviggianus graecus Bink-Moenen 1992: 39. Holotype pupal case. Greece: on Quercus coccifera, 

BMNH. 

206

Distribution: 3: Crete; Greece. 

Host: Fagaceae: Quercus coccifera. 

Aleuroviggianus halperini Bink-Moenen 1992 

Aleuroviggianus halperini Bink-Moenen 1992, in Bink-Moenen and Gerling 1991: 14. Holotype. Israel: Mt. 

Meron, ix.1976, R. Neeman, on Quercus calliprinos, BMCol 

Distribution: 3: Crete; Greece; Iran, Rhodes; Turkey. 

Host: Fagaceae: Quercus calliprinos, Quercus coccifera, Quercus ithaburensis, Quercus rotundifolia, 

Quercus sp. 

Aleuroviggianus polymorphus Bink-Moenen 1992 

Aleuroviggianus polymorphus Bink-Moenen 1992: 27. Holotype pupal case. Spain: on Quercus sp, BMNH. 

Aleurodes quercus Signoret; misidentification Gomez-Menor 1945a: 283. 

Distribution: 3: Spain. 

Host: Fagaceae: Quercus ilex, Quercus suber. 

Aleuroviggianus zonalus Bink-Moenen 1992 

Aleuroviggianus zonalus Bink-Moenen 1992: 33. Holotype pupal case. Greece: on Quercus coccifera, 

BMNH. 

Distribution: 3: Crete; Greece; Iran, Rhodes. 

Host: Fagaceae: Quercus coccifera, Quercus ithaburensis, Quercus sp. 

207

Aleurovitreus Martin 2005 

Aleurovitreus Martin 2005b: 28. Type species Aleyrodes insignis Bondar 1923. 

Diagnosis - (From Martin 2005b): In nature - cuticle often pale, but some species with brownish patches, 

cryptic when feeding, silvery and glassy when adults or parasitoids have emerged; On slide - lateral 

margin deflexed and with rounded crenulations, not clearly modified at the tracheal openings; setae - 

cephalic, T2, T3, A8 and caudal setae present, sometimes thoracic setae very reduced, rarely without 

cephalic pair; vasiform orifice cordate, internally smooth, fully occupied by the operculum; A7 not 

reduced medially; geminate pores/porettes regularly distributed over the dorsum; transverse molting 

sutures closely approach the apparent puparial margin. 

Comparison - The puparia of Aleurovitreus are similar to those of Aleuropleurocelus in that both have the 

lateral margin deflexed; however, Aleurovitreus species are pale, or with brown patches, and have the 

cuticle smooth and finer than in Aleuropleurocelus which are black, often granulated and more robust. 



insignis x 

risor x 

spp. x 

Total = 2+ species 0 2+ 0 0 0 0 0 0 0 

Aleurovitreus insignis (Bondar) 1923 

Aleyrodes insignis Bondar 1923a: 127. Syntypes. Brazil: on Persea gratissima, MZU. 

Aleurovitreus insignis (Bondar) 1923; Martin 2005b: 29. 

Distribution: 2: Belize, Brazil, Costa Rica, Venezuela. 

Host: Lauraceae: Persea americana, Persea gratissima. 

Diagnosis: Puparium pale with 3 dark brown patches - head, abdominal base and around the vasiform orifice. 

C1, T2, T3, A8 and caudal setae very long, patch of 4 pores in square arrangement present 

submedially on dorsum at level of leg I. 

Aleurovitreus risor Martin 2005 

Aleurovitreus risor Martin 2005b: 31. Holotype puparium, Belize, CFR, 22.vi.2002, JH Martin, on Piper sp., 

BMNH. 

208

209 


Host: Piperaceae: Piper sp. 

Diagnosis: Puparium color variable - 1 type is pale with 3 dark brown patches - a pair of large round ones on 

thorax and a large one surrounding the vo (appears as a happy-face)- other kind has a spot on the head 

and a pair of two, elliptical patches on the sides of the abdomen. C1, A8 and caudal setae moderately 

long, T2 and T3 absent, represented only by vestigial bases. 

Aleurovitreus spp. 

Seven undescribed species from Brazil (Martin 2005b).

Aleyrodes Latreille 1796 

Aleyrodes Latreielle 1796: 93. Type species. Phalaena (Tinea) proletella Linnaeus 1758:537, by monotypy. 

Conantulus Goux 1987: 65. Type species. Conantulus lacombiensis Goux 1988, by monotypy; synonymy 

according to Martin 1999: 53. 



albescens x 

amnicola x x 

asari x 

asarumis x 

aureocincta x 

baja x 

ciliata x 

crataegi x 

diasemus x 

elevatus x x 

essigi x 

fodiens x 

gossypii x 

hyperici x 

japonica x 

lactea x x 

latus x 

210


lonicerae x x 

millettiae x 

osmaroniae x 

philadelphi x 

proletella x x x x x x 

pruinosus x x 

pyrolae x 

shizuokensis x x x 

singularis x 

sorini x 

spiraeoides x x 

taiheisanus x 

takahashi x 

tinaeoides x 

winterae x 

zygia x 

Total = 34 species 10 9 7 3 10 3 1 3 1 

Aleyrodes albescens Hempel 1923 

Aleyrodes albescens Hempel 1923: 1177. Syntypes. Brazil: on Coffea arabica, MZU. 


Host: Rubiaceae: Coffea arabica. 

Aleyrodes amnicola Bemis 1904 

Aleyrodes amnicola Bemis 1904: 514. Syntypes. USA: California, on Salix laevigata, USNM. 

Distribution: 1: USA (CA); 2: Virgin Islands. 

211

212 

Host: Arecaceae: Washingtonia nuda; Brassicaceae: Brassica oleracea L, Capitata Group; Salicaceae: Salix 

laevigata, Salicaceae: Salix sp. 

Aleyrodes asari (Schrank) 1801 

Coccus asari Schrank 1801: 145. Syntypes. Germany: on Asarum europaeum. 

Aleyrodes asari (Schrank); Lindinger 1932: 223. 

Distribution: 3: Albania; Austria; Czech Republic; Germany; Hungary; Poland. 

Host: Aristolochiaceae: Asarum europaeum. 

Aleyrodes asarumis Shimer 1867 

Aleyrodes asarumis Shimer 1867: 281. Syntypes. USA: Connecticut, on Actaea sp., USNM. 

Aleurodicus asarumis (Shimer); Riley & Howard 1893: 219. 

Aleyrodes asarumis Shimer; Quaintance & Baker 1914: 45. 

Distribution: 1: Canada; USA (CT, IL, PA, TN). 

Host: Aristolochiaceae: Asarum canadense, Asarum sp.; Ranunculaceae: Actaea alba. 

Aleyrodes aureocincta Cockerell 1897 

Aleyrodes aureocincta Cockerell 1897a: 42. Syntypes. USA: New Mexico, on Aquilegia sp., USNM. 

Distribution: 1: USA (IL, NM). 

Host: Ranunculaceae: Aquilegia sp. 

Natural enemies: Eulophidae: Euderomphale flavimedia. 

Aleyrodes baja Sampson 1943 

Aleyrodes baja Sampson 1943: 215. Syntypes. USA: California, on unidentified tree, CAS. 



Aleyrodes ciliata Takahashi 1955 

Aleyrodes ciliata Takahashi 1955a: 399. Syntypes. Madagascar: on "randrompody", MNHNP. 


Host: on "randrompody”. 

Aleyrodes crataegi Kiriukhin 1947 

Aleyrodes crataegi Kiriukhin 1947: 9. Syntypes. Iran: on Crataegus monogyna and Cydonia vulgaris.

Distribution: 3: Iran. 

Host: Rosaceae: Crataegus monogyna, Cydonia vulgaris. 

Aleyrodes diasemus Bemis 1904 

Aleyrodes diasemus Bemis 1904: 516. Syntypes. USA: California, on Symphoricarpus racemosus, USNM; 

Russell 1948: 78. 

Asterochiton diasemus (Bemis); Quaintance & Baker 1914: 105. 

Trialeurodes diasemus (Bemis); Quaintance & Baker 1915a: xi. 

Aleyrodes diasemus Bemis; Russell 1948: 78. 


Host: Caprifoliaceae: Symphoricarpos racemosus; Ericaceae: Symphysia racemosa; Saxifragaceae: Ribes 

glutinosum. 

Aleyrodes elevatus Silvestri 1934 

Aleyrodes elevatus Silvestri 1934: 394. Syntypes. Italy: on Mercurialis annua, IESP. 

Distribution: 3: France, Iran; Italy; Spain, Syria, Turkey. 

Host: Euphorbiaceae: Mercurialis annua; Moraceae: Ficus carica, Ficus religiosa; Tiliaceae: Corchorus 

triocularis; Urticaceae: Urtica dioica. 

Natural enemies: Aphelinidae: Encarsia inaron, Encarsia lahorensis, Encarsia tricolor, Eretmocerus 

ostovani. 

US Port Interception: Yugoslavia on Ficus sp. 

Aleyrodes essigi Penny 1922 

Aleyrodes essigi Penny 1922: 23. Syntypes. USA: California, Mission San Jose, ix.1916, E. Essig, on Ulmus 

sp, USNM. 


Host: Ulmaceae: Ulmus sp. 

Aleyrodes fodiens Maskell 1895 

Aleurodes fodiens Maskell 1895: 433. Lectotype (designated by Dumbleton 1957: 156). New Zealand: on 

Drimys axillaris, ADSIR. 

Dialeurodes fodiens (Maskell); Quaintance & Baker 1914: 97 

Aleyrodes fodiens Maskell; Dumbleton 1957: 155. 

Distribution: 8: New Zealand. 

Host: Winteraceae: Drimys axillaris. 

213

Aleyrodes gossypii (Fitch) 1857 

Aspidiotus gossypii Fitch 1857: 332. Syntypes. China: on Gossypium religiosum. 

Aleyrodes gossypii (Fitch); Ashmead 1895a: 332. 


Host: Malvaceae: Gossypium religiosum. 

Natural enemies: Thripidae: Sericothrips trifasciatus. 

Aleyrodes hyperici Corbett 1926 

Aleyrodes hyperici Corbett 1926: 274. Syntypes. Sri Lanka: on Hypericum sp., BMNH. 


Host: Guttiferae: Hypericum sp. 

Aleyrodes japonica Takahashi 1963 

Aleyrodes japonica Takahashi 1963: 54. Syntypes. Japan: on ?Viola sp., HBL. 


Host: Violaceae: Viola sp.? 

Aleyrodes lactea Zehntner 1897 

Aleurodes lactea Zehntner 1897b: 558. Syntypes. Java: on Saccharum officinarum. 

Aleyrodes lactea Zehntner; Mound and Halsey 1978: 96. 

Distribution: 5: Philippines; 7: Java. 

Host: Poaceae: Saccharum officinarum. 


Aleyrodes latus Hempel 1923 

Aleyrodes latus Hempel 1923: 1178. Syntypes. Brazil: on Baccharis genistelloides, MZU. 


Host: Asteraceae: Baccharis genistelloides. 

Aleyrodes lonicerae Walker 1852 

Aleyrodes lonicerae Walker 1852: 1092. Syntypes. England: on Lonicera periclymenum. 

Aleyrodes borchsenii Danzig 1966: 371. Holotype. USSR: southern Maritime Territory, on Urtica sp., ZIN; 

214

Danzig 2004. 

Aleyrodes fragariae Walker 1852: 1092. Syntypes. England: on Fragaria sp. (strawberry); Ossiannilsson 

1955: 193. 

Conantulus lacombiensis Goux 1987: 65; Martin 1999: 53. 

Aleyrodes menthae Haupt 1934: 139. Syntypes. Germany: on Mentha piperita; Ossiannilsson 1955: 193. 

Aleyrodes spiraeae Douglas 1894b: 73. Syntypes. Germany: on Lonicera xylosteum; Mound 1966: 406. 

Aleyrodes rubi Signoret 1868: 382. Syntypes. France: on Rubus fruticosus; Trehan 1940: 608. 

Distribution: 3: Austria; England; Finland; France; Germany; Hungary; Iran, Israel; Italy; Poland; Portugal, 

Russia, Sweden; Switzerland, Turkey, USSR; Yugoslavia; 5: Korea. 

Host: Balsaminaceae: Impatiens noli-tangere; Campanulaceae: Platycodon grandiflorum; 

Caprifoliaceae: Lonicera periclymenum, Lonicera xylosteum; Ericaceae: Vaccinium myrtillus; 

Fabaceae: Robinia viscosa; Oxalidaceae: Oxalis aceosella, Oxalis acetosella, Oxalis sp.; 

Papaveraceae: Chelidonium majus, Dicentra spectabilis; Rosaceae: Crataegus microphylla, 

Filipendula ulmaria, Fragaria sp., Fragaria vesca, Geum rivale, Prunus dulcis, Rubus chamaemorus; 

Umbelliferae sp., Aegopodium podagraria; Urticaceae: Urtica dioica, Urtica sp.; Violaceae: Viola 

riviniana, Viola sp.. 

Natural enemies: Aphelinidae: Cales noacki, Encarsia formosa, Encarsia inaron, Encarsia lutea, Encarsia 

meritoria, Encarsia tricolor; Eulophidae: Euderomphale chelidonii. 

US Port Interceptions: Belgium on Mentha sp.; Czech Republic on Fragaria sp.; France on Rubus sp.; 

Germany on unknown plant; Italy on Fragaria sp.; Netherlands on Carthamus sp.; United 

Kingdom on Vitis vinifera. 

Aleyrodes millettiae Cohic 1968 

Aleyrodes millettiae Cohic 1968b: 113. Syntypes. Congo: on Millettia laurentii and Vitex madiensis. 

Aleurocybotus millettiae (Cohic); Bink-Moenen 1983: 41. 

Aleyrodes millettiae Cohic; Martin & Mound 2007. 


Host: Fabaceae: Millettia laurentii; Verbenaceae: Vitex madiensis. 

Aleyrodes osmaroniae Sampson 1945 

Aleyrodes osmaroniae Sampson 1945: 58. Syntypes. USA: California, on Osmaronia cerasiformia, UCD. 


Host: Rosaceae: Osmaronia cerasiformia. 

Aleyrodes philadelphi Danzig 1966 

Aleyrodes philadelphi Danzig 1966: 369. Holotype. USSR: Vladivostock, on Philadelphus tenuifolius, ZIN. 


215

216 

Host: Saxifragaceae: Bergenia pacifica, Ribes sp., Philadelphus pacificus, Philadelphus tenuifolius 

Natural enemies: Aphelinidae: Encarsia marinikia, Encarsia tricolor, Encarsia trjapitzini. 

Aleyrodes proletella (Linnaeus) 1758 

Phalaena (Tinea) proletella Linnaeus 1758: 537. Syntypes. Europe: on Brassica sp. 

Aleyrodes brassicae Walker 1852: 1092. Syntypes. England: on cabbage [Brassica sp.]; Haupt 1935: 256. 

Aleyrodes chelidonii Latreille 1807; Walker 1852: 1092 

Aleyrodes euphorbiae Low 1867: 746. Syntypes. Austria: on Euphorbia peplus; Zahradnik 1991: 113. 

Coccus preanthis Schrank 1801: 147. 

Aleyrodes prenanthis (Schrank) 1801; Cockerell 1902a: 281. 

Aleyrodes youngi Hempel 1901: 385. Syntypes. Brazil: on cabbage [Brassica sp.], 

Distribution: 1: USA (intercepted in CA, MA, NY but not known to be established in the USA); 2: Bermuda, 

Brazil, Mexico, Puerto Rico, Virgin Islands; 3: Austria; Azores, Belgium, Canary Islands, Czech 

Republic; Egypt; England; Finland; France; Germany; Hungary; Iran, Italy; Poland; Portugal; Russia; 

Spain; Sweden; Switzerland; USSR; Yugoslavia; 4: Angola; Canary Is; Kenya; Mozambique; Sierra 

Leon, Zimbabwe; 5: Hong Kong; 8: New Zealand. 

Host: Asteraceae: Cichorium intybus, Cichorum juncea; Chondrilla juncea; Emilia sonchifolia, Lactuca 

indica, Lactuca serriola, Lactuca sp., Mycelis muralis, Sonchus arvensis, Sonchus oleraceus, Sonchus 

oleraceus, Sonchus sp.; Brassicaceae: Brassica oleracea, Brassica oleracea Capitata Group, Brassica 

oleracea botrytis, Raphanus raphanistram, Rorippa indica; Campanulaceae: Campanula grandis, 

Campanula persicifolia; Euphorbiaceae: Euphorbia esula; Oxalidaceae: Oxalis sp.; Papaveraceae: 

Chelidonium majus var. laciniatum, Chelidonium majus var. majus, Chelidonium majus var. plenum; 

Ranunculaceae: Celadine sp.; Rutaceae: Citrus sp. 

Natural enemies: Parasitoids- Hymenoptera/Aphelinidae: Encarsia formosa, Encarsia hispida, Encarsia 

inaron, Encarsia japonica, Encarsia lutea, Encarsia melanostoma, Encarsia noahi, Encarsia sophia, 

Encarsia tricolor, Eretmocerus orientalis; Eulophidae: Euderomphale insularis, Euderomphale 

cerris, Euderomphale chelidonii, Euderomphale gomer, Neochrysocharis formosa. Predators - 

Coleoptera/Coccinellidae: Clitostethus arcuatus; Diptera/Cecidomyidae: Phaenobremia aphidivora; 

Drosophilidae: Acletoxenus formosus; Syrphidae: Syrphus auricollis, Syrphus cinctus; 

US Port Interceptions: Azores on unknown plant; Canary Islands on Recordia sp.; France on Brassica 

oleracea; India on Rosa sp.; Israel on Brassica rapa; Italy on Brassica oleracea, Lactuca sativa; 

Mexico on Citrus sinensis; Netherlands on Brassica sp.; Portugal on Brassica oleracea; Romania 

on Artemisia sp.; Spain on Brassica oleracea; St. Kitts and Nevi on Brassica juncea; Zimbabwe on 

Aster sp. 

Aleyrodes pruinosus Bemis 1904 

Aleyrodes pruinosus Bemis 1904: 491. Syntypes. USA: California, on Heteromeles arbutifolia, USNM. 

Aleyrodes pruinosus euphorbium Cockerell 1911: 462. Syntypes. USA: Colorado, on Euphorbia robusta, 

USNM.

Distribution: 1: USA (CA, CO, FL, MA, TX, WA); 2: Mexico. 

Host: Euphorbiaceae: Euphorbia robusta; Rhamnaceae: Ceanothus arbutifolia; Rosaceae: Heteromeles 

arbutifolia. 

Natural enemies: Aphelinidae: Encarsia peltata; Eulophidae: Euderomphale flavimedia. 

Aleyrodes pyrolae Gillette & Baker 1895 

Aleyrodes pryrolae Gillette & Baker 1895: 125. Syntypes. USA: Colorado, on Pyrola rotundifolia, USNM. 

Distribution: 1: USA (CO). 

Host: Pyrolaceae: Pyrola rotundifolia. 

Aleyrodes shizuokensis Kuwana 1911 

Aleyrodes shizuokensis Kuwana 1911: 620. Syntypes. Japan: on Oxalis corniculata, TARI. 

Distribution: 5: Japan; Taiwan; 6: India; 9: Hawaii. 

Host: Asteraceae: Sonchus oleraceus; Euphorbiaceae: Phyllanthus distinctus; Oxalidaceae: Oxalis 

corniculata. 

Reference: David & Jesudasan (1989) - redescription and illustration. 

Aleyrodes singularis Danzig 1964 

Aleyrodes singularis Danzig 1964a: 645. Holotype. USSR: Georgia, on Euphorbia sp., ZIN. 

Distribution: 3: Azores, Canary Islands; Iran, Jordan; USSR. 

Host: Asteraceae: Lactuca serriola, Lactuca sp., Sonchus arvensis; Euphorbiaceae: Euphorbia stygiana, 

Euphorbia pulcherimma, Euphorbia sp.; Primulaceae: Lysimachia nemorum. 

Natural enemies: Aphelinidae: Encarsia estrellae, Encarsia estrellae, Encarsia formosa, Encarsia dichroa, 

Encarsia hispida, Encarsia estrellae, Encarsia tricolor, Encarsia noahi, Encarsia inaron, Eulophidae: 

Euderomphale gomer. 

Aleyrodes sorini Takahashi 1958 

Aleyrodes sorini Takahashi 1958: 63. Syntypes. Japan: on ?Sonchus sp., HBL. 


Host: Asteraceae: Sonchus sp.? 

Aleyrodes spiraeoides Quaintance 1900 

Aleurodes spiraeoides Quaintance 1900. Syntypes. USA: California, on Fuchsia sp., USNM. 

Aleyrodes spiraeoides Quaintance; Mound and Halsey 1978: 102. 

217

218 

Distribution: 1: Canada, USA (AZ, CA, CO, FL, ID, LA, NV, OR, UT, TX); 2: Mexico; Venezuela. 

Host: Asclepiadaceae: Asclepias sp.; Asteraceae: Sonchus oleraceus, Troximon sp.; Caprifoliaceae: Lonicera 

involucrata; Convolvulaceae: Convolvulus sepium, Convolvulus occidentalis; Ericaceae: Gaultheria 

shallon; Gentianaceae: Eustoma sp.; Guttiferae: Hypericum androsamum; Hippocastanaceae: Aesculus 

californica; Iridaceae: Iris sp., Iris cristata; Myrtaceae: Melaleuca hypericifolia; Onagraceae: Fuchsia 

sp.; Plantaginaceae: Plantago major; Rosaceae: Fragaria x ananassa; Rosaceae: Fragaria sp.; 

Opulaster capitatus; Saururaceae: Anemopsis californica; Solanaceae: Solanum tuberosum; 

Solanaceae: Nicotiana glauca, Solanum douglasii, Solanum tuberosum. 

Natural enemies: Parasitoids - Aphelinidae: Encarsia formosa, Encarsia californica, Encarsia peltata; 

Encarsia sp. (in Canada); Eretmocerus californicus; Eulophidae: Euderomphale flavimedia; 

Predators - DIPTERA/Syrphidae: Allograpta obliqua; HEMIPTERA/Lygeidae: Geocoris pallens, 

Geocoris pallens. 

Aleyrodes taiheisanus Takahashi 1939 

Aleyrodes taiheisanus Takahashi 1939a: 77. Syntypes. Taiwan: on Sassafras [=Yushunia] randaiensis, TARI. 


Host: Lauraceae: Sassafras randaiensis. 

Aleyrodes takahashi Ossiannilsson 1966 

Aleyrodes campanulae Takahashi 1963: 53. Japan: on Campanula punctata. 

Aleyrodes takahashii Ossiannilsson 1966: 155. Name preoccupied A. campanulae nec Saalas. Japan: on 

Campanula punctata. 


Host: Campanulaceae: Campanula punctata. 

Aleyrodes tinaeoides Blanchard 1852 

Aleurodes tinaeoides Blanchard 1852: 320. Syntypes. Chile: on unidentified host, BMNH; 

Mound and Halsey 1978: 102 


Host: unknown plant; Solanaceae: Cestrum parqui. 

Aleyrodes winterae Takahashi 1937 

Aleyrodes winterae Takahashi 1937d: 251. Syntypes. New Zealand: on Wintera colorata, TARI. 


Host: Winteraceae: Drimys colorata.

Aleyrodes zygia Danzig 1966 

Aleyrodes zygia Danzig 1966: 370. Holotype. USSR: southern Maritime Territory, on Euphorbia sp., ZIN 


Host: Euphorbiaceae: Euphorbia sp. 

Transfers 

Aleyrodes insignis Bondar 1923 to Aleurovitreus by Martin 2005: 29. 

219

Aleyrodiella Danzig 1966 

Aleyrodiella Danzig 1966: 381 [206]. Type species. Aleyrodiella lamellifera Danzig 1966, by monotypy. 



lamellifera x 

Total = 1 species 0 0 1 0 0 0 0 0 0 

Aleyrodiella lamellifera Danzig 1966 

Aleyrodiella lamellifera Danzig 1966: 381. Holotype. USSR: Vladivostock, Okeanskaya, on Ulmus laciniata, 

ZIN. 


Host: Ulmaceae: Ulmus laciniata. 

220

Anomaleyrodes Takahashi & Mamet 1952 

Anomaleyrodes Takahashi & Mamet 1952b: 129. Type species. Anomaleyrodes palmae Takahashi & Mamet 




palmae x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Anomaleyrodes palmae Takahashi & Mamet 1952 

Anomaleyrodes palmae Takahashi & Mamet 1952b: 130. Syntypes. Madagacar: on palm, MNHNP. 


Host: Arecaceae: a palm. 

221

Apobemisia Takahashi 1954 

Apobemisia Takahashi 1954b: 52. Type species. Bemisia kuwanai Takahashi 1934, by original designation. 



celti x 

kuwanai x 

Total = 2 species 0 0 0 0 2 0 0 0 0 

Apobemisia celti (Takahashi) 1932 

Pealius celti Takahashi 1932: 40. Syntypes. Taiwan: on Celtis sinensis, TARI. 

Apobemisia celti (Takahashi) 1932; Takahashi 1954b: 53. 


Host: Ulmaceae: Celtis sinensis. 

Apobemisia kuwanai (Takahashi) 1934 

Bemisia kuwanai Takahashi 1934b: 60. Syntypes. Taiwan: on Ficus sp., TARI. 

Apobemisia kuwanai (Takahashi); Takahashi 1954b: 52. 


Host: Moraceae: Ficus pumila. 

222

Arachnaleyrodes Bink-Moenen 1983 

Arachnaleyrodes Bink-Moenen 1983: 87. Type species. Arachnaleyrodes insignis Bink-Moenen 1983. 



insignis x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Arachnaleyrodes insignis Bink-Moenen 1983 

Arachnaleyrodes insignis Bink-Moenen 1983: 87. Holotype pupal case, Chad, Bededjia, 8.viii.1973, FAB & 

RMBM, on Combretum hypopilinum; BMNH. 


Host: Combretaceae: Combretum hypopilinum, Combretum glutinosum. 

223

Asialeyrodes Corbett 1935 

Asialeyrodes Corbett 1935b: 841. Type species. Asialeyrodes lumpurensis Corbett 1935, by original 

designation. 

Icfrealeyrodes Dubey & Sundararaj 2007. Type species - Icrealeyrodes indica, by monotypy and 

original designation; synonymy according to Martin & Mound 2007: 56. 

References: Meganathan & Vasantharaj 1994; Dubey & Sundararaj 2005; Ko et al. 1993. 


Species NA 

NT WPA AF EPA O AU PI HI 

corbetti 

dorsidemarcata 

dubius 

elegans 

euphoriae 

indica 

lumpurensis 

lushanensis 

meghalayensis 

maesae 

menoni 

multipori 

papillatus 

selangorensis 

sphaerica 

splendens 

sp. 

Total = 15 species 0 

x 

x 

0 0 0 2 12 1 1 0 

x 

x 

x 

x 

x 

x 

x 

x 

x 

x 

x 

x 

x 

x 

x 

224

Asialeyrodes corbetti Takahashi 1949 

Asialeyrodes corbetti Takahashi 1949: 51. Syntypes. Rioux (Riau) Islands (south of Singapore): on 

unidentified tree, HBL. 

Distribution: 8: Riouw (Riau) Islands 


Asialeyrodes dorsidemarcata (Singh) 1932 

Dialeurodes dorsimarcata Singh 1932: 82. Syntypes. Burma: on an unidentified host. 

Pseudaleurolobus dorsidemarcata (Singh); Takahashi 1934b: 47. 

Pseudaleurolobus dorsimarcata [sic] (Singh); Mound & Halsey 1978: 185 

Asialeyrodes dorsidemarcata (Singh); Martin & Mound 2007. 


Host: unidentified. 

Asialeyrodes dubius Martin & Mound 2007 

Icfrealeyrodes indica Dubey & Sundararaj 2007. Holotype puparium, India, Karnataka, Yana, 17.ii.2001, A. 

K. Dubey, ex. Syzygium sp., IFRI. 

Asialeyrodes dubius Martin & Mound 2007: 22; new name as a result of generic synonymy, A. indica 

preoccupied by Asialeyrodes indica Sundararaj & David 1992. 



Asialeyrodes elegans Meganathan & David 1994 

Asialeyrodes elegans Meganathan & David 1994: 26. Holotype. India: on Sonerilla elegans; Dubey & 

Sundararaj 2005e. 


Host: Lecythidaceae: Sanerilla elegans. 

Asialeyrodes euphoriae Takahashi 1942 

Asialeyrodes euphoriae Takahashi 1942c: 204. Syntypes. Thailand: on Euphoria longana. 

Distribution: 5: China; 6: Thailand. 

Host: Sapotaceae: Euphoria longan. 


225

Asialeyrodes indica Sundararaj & David 1991 

Asialeyrodes indica Sundararaj & David 1992: 415. Holotype. India, Tamil Nadu, Coimbatore, 15.iv.1968, B. 

V. David, on Tabernaemontana [=Ervatomia] coronaria; David & Jesudasan 2002: 325. 

Asialeurodes saklespurensis Regu & David 1993: 44. Holotype puparium, India, Karnataka, Saklespur, 

4.ii.1990, K. Regu, on Olea sp; synonymy according to Dubey & Sundararaj 2005e. 

Distribution: 6: Andaman, India, India, Nicobar Islands. 

Host: Apocynaceae: Ervatomia coronaria, Tabernmontana heyneana; Arliaceae: Scheffera rostrata; 

Capparaceae: Capparis rheedii; Euphorbiaceae: Antidesma sp.; Lytharaceae: Lagerstromea microcarpa; 

Myrtaceae: Syzygium sp., Moraceae: Streblus asper; Oleaceae: Olea dioica; Rubiaceae: Xeromphis spinosa; 

Smilcaceae: Smilax zeylanica; Simplocaceae: Symplocaceae sp. 

Asialeyrodes lumpurensis Corbett 1935 

Asialeyrodes lumpurensis Corbett 1935b: 841. Syntypes. Malaya: on unidentified host, lost? 



Asialeyrodes lushanensis Ko 1993 

Asialeyrodes lushanensis Ko 1993 in Ko et al. 1993. Holotype puparium, Taiwan, Lushanwenchuan, 

6.ix.1986, C. C. Ko, ex. Syzygium buxifolium, NTU. 


Host: Myrtaceae: Syzygium buxifolium. 

Asialeyrodes maesae (Takahashi) 1934 

Pseudaleurolobus maesae Takahashi 1934b: 46; Syntypes. Taiwan: on Maesa formosana, TARI. 

Asialeyrodes maesae (Takahashi); Takahashi 1942c: 207 

Distribution: 5: China; Taiwan. 

Host: Myrsinaceae: Maesa formosana. 

Asialeyrodes meghalayensis Regu & David 199 

Asialeyrodes meghalayensis Regu & David 1991a: 256. Holotype puparium, Meghalaya State, Cherrapunjee, 

21.x.1989, B. V. David, on Ixora brachiata. 


Host: Rubiaceae: Ixora brachiata. 

226

Asialeyrodes menoni Meganathan & David 1994 

Asialeyrodes menoni Meganathan & David 1994: 26. Holotype. India: on Litsea laevigata, - 


Host: Lauraceae: Litsea laevigata. 

Asialeyrodes multipori Takahashi 1942 

Asialeyrodes multipori Takahashi 1942c: 206. Holotype. Thailand: on unidentified host, TARI. 



Asialeyrodes papillatus Regu & David 1991 

Asialeyrodes papillatus Regu & David 1991a: 257. Holotype puparium, Meghalaya State, Cherrapunjee, 

21.x.1989, B. V. David, on Ixora brachiata. 


Host: Rubiaceae: Ixora brachiata. 

Asialeyrodes selangorensis Corbett 1935 

Asialeyrodes selangorensis Corbett 1935b: 842. Syntypes. Malaya: on unidentified host, ? 



Asialeyrodes spherica (Sundararaj & Dubey) 2006 

Rhachisphora spherica Sundararaj & Dubey 2006b. Holotype puparium, India, Karnataka, 22.i.2001, A. K. 

Dubey, on Caesalpinia pulcherimma, IFRI. 

Asialeyrodes spherica (Sundararaj & Dubey); Martin & Mound 2007: 23. 


Host: Fabaceae: Caesalpinia pulcherimma. 

Asialeyrodes splendens Meganathan & David 1994 

Asialeyrodes splendens Meganathan & David 1994: 27. Holotype. India: on Microtropis ramiflora. 


Host: Lamiaceae: Microtropis ramiflora. 

227

228 

Asialeyrodes sp. 1 

Asialeyrodes sp.; Martin 1988: 70 



Asialeyrodes sp. 2 

Indonesia, 8 JUN 2006, B. McCoy, ex. unknown plant parts, possibly Coffea. 

Note: Similar to A. lumpurensis Corbett but has submedial line of reticulations going up the entire body. 

A. euphoriae also shows this line.

Asterobemisia Trehan 1940 

Asterobemisia Trehan 1940: 591. Type species. Aleurodes carpini Koch 1857, by monotypy. [Aleyrodes [sic] 

carpini Koch]. 

Bemisia (Neobemisia) Visnya 1941b: 8. Type species: Bemisia yanagicola Takahashi 1934, by original 

designation; Mound and Halsey 1978: 104. 

Neobemisia Visnya; synonymy according to Zahradnik 1961b: 61. 



atraphaxius x 

carpini x x 

curvata x 

dentata x 

lata x 

obenbergeri x 

paveli x 

salicaria x 

silvatica x 

takahashii x 

trifolii x x 

yangicola x 

Total = 12 species 0 0 9 0 4 1 0 0 0 

Asterobemisia atraphaxius (Danzig) 1969 

Neobemisia atraphaxius Danzig 1969: 874. Holotype. USSR: Kazakhastan, Amolinksk Province, on 

Atraphaxis frutescens, ZIN. 

Asterobemisia atraphaxius (Danzig); Mound and Halsey 1978: 104. 

Distribution: 3: Europe; Iran, Russia; USSR. 

Host: Aceraceae: Acer saccharinum, Acer campestre; Betulaceae: Betula sp., Carpinus betulus, Corylus 

229

230 

avellana L.; Caprifoliaceae: Lonicera fragrantissima, Lonicera nigra; Ericaceae: Vaccinium myrtillus 

L. var. myrtillus; Fabaceae: Robinia pseudoacacia, Castanea sativa; Fagaceae: Quercus robus.; 

Moraceae: Humulus lupulus; Polygonaceae: Atraphaxis spinosa, Atraphaxis frutescens; 

Polygonum convolvulus; Ranunculaceae: Clematis viticalba; Rosaceae: Rubus fruticosus, 

Crataegus sp.; Salicaceae: Salix sp.; Saxifragaceae: Ribes nigrum, Ribes rubrum; Tiliaceae: Tilia 

ulmifolia, Ulmus foliacea. 

Natural enemies: Aphelinidae: Encarsia lutea, Encarsia tricolor, Eretmocerus neobemisia; Eulophidae: 

Euderomphale sp. 

Asterobemisia carpini (Koch) 1857 

Aleurodes carpini Koch 1857: 327. Syntypes. Germany (west): on Carpinus betulus, BMNH 

Asterobemisia carpini (Koch) 1857; Trehan 1940: 593. 

Aleurodes avellanae Signoret 1868: 385. Lectotype (designated by Zahradnik 1961:437). France: on Corylus 

avellana, IESP; Mound and Halsey 1978: 105. 

Aleurochiton avellanae (Signoret); Harrison 1920a: 59; Zahradnik 1956: 44. 

Bemisia (Neobemisia) avellanae (Signoret); Visnya 1941b: 8. 

Aleurodes ribium Douglas 1888: 265. Lectotype. unknown host and locality, BMNH. 

Bemisia (Neobemisia) ribium (Douglas); Visnya 1941b: 9. 

Aleurodes rubicola Douglas 1891b: 200. Lectotype. England: on bramble [Rubus sp.] leaves, BMNH; Trehan 

1939: 266. 

Aleurochiton vaccinii Kunow 1880: 46. Syntypes. Konisberg Prov, on Vaccinium uliginosum, BMNH. 

Distribution: 2: Costa Rica; 3: Austria; Czech Republic; Denmark; England; Finland; France; Germany; 

Hungary; Iran, Italy; Moldavia, Netherlands, Spain; Sweden; USSR, Yugoslavia; 5: Korea, Taiwan. 

Host: Asteraceae: Artemisia carpini, Artemisia princeps; Betulaceae: Corylus avellana, Corylus orientalis; 

Caprifoliaceae: Linnaea borealis; Ericaceae: Vaccinium uliginosum; Fabaceae: Lespedeza bicolor; 

Punicaceae: Punica granatum; Rosaceae: Rosa pimpinellifolia. 

Natural enemies: Aphelinidae: Encarsia asterobemisia, Encarsia coryli, Encarsia inaron, Encarsia 

longicornis, Encarsia sophia, Eretmocerus mundus, Eretmocerus neobemisiae; Eulophidae: 

Euderomphale cerris, Euderomphale sp.; Platygastridae: Amitus longicornis, Isostasius sp. 

US Port Interceptions: France on Acer negundo, Ranunculus sp., Ribes sp., Rosa sp., Rubus sp.; 

Germany on Mahonia sp., Rosa sp., Rubus sp.; Netherlands on Rosa sp.; Philippines on Psidium sp.; 

United Kingdom on Ribes sp. 

Asterobemisia curvata (Qureshi) 1980 

Bemisia curvata Qureschi 1980: 405. Holotype pupal case. Pakistan, Murree, 30.iv.1975, J. I. Qureshi, on 

Aesculus sp., FAI (Pakistan), UAF. 

Asterobemisia curvata (Qureshi); Martin & Mound 2007. 

Distribution: 6: Pakistan.

Host: Hippocastanaceae: Aesculus sp.; Rosaceae: Rosa indica. 

Asterobemisia dentata Danzig 1969 

Asterobemisia dentata Danzig 1969: 873. Holotype. USSR: Tadzikistan, on Morus sp., ZIN 

Distribution: 3: France, Iran, Netherlands, USSR. 

Host: Amaranthaceae: Amaranthus blitoides; Moraceae: Morus alba, Morus sp. 

Asterobemisia lata Danzig 1966 

Asterobemisia lata Danzig 1966: 376. Holotype. USSR: Vladivostock, on Carpinus cordata, ZIN. 

Distribution: 3: Iran, USSR. 

Host: Betulaceae: Carpinus cordata; Fagaceae: Quercus calliprinos, Quercus sp.; Rosaceae: Malus orientalis, 

Malus sp.; Tiliaceae: Corchorus tricularis, Tilia sp. 

Asterobemisia obenbergeri (Zahradnik) 1961 

Neobemisia obenbergeri Zahradnik 1961b: 68. Holotype. Czech Republic: on Thymus sp. 

Asterobemisia obenbergeri (Zahradnik) 1961; Mound and Halsey 1978: 106. 

Distribution: 3: Czech Republic; Yugoslavia. 

Host: Labiatae: Thymus sp. 

Asterobemisia paveli (Zahradnik) 1961 

Neobemisia paveli Zahradnik 1961b: 75. Holotype. Czech Republic: on Euphorbia sp., USNM. 

Asterobemisia paveli (Zahradnik); Mound and Halsey 1978: 107. 

Asterobemisia nigrini Zahradnik 1987a: 350; Martin et al. 2000: 430. 

Distribution: 3: Czech Republic; Germany; Palestine. 

Host: Euphorbiaceae: Euphorbia stepposa, Euphorbia sp. 

Natural enemies: Aphelinidae: Encarsia inaron, Eretmocerus mundus. 

Asterobemisia salicaria (Danzig) 1969 

Bemisia salicaria Danzig 1969: 871. Holotype. USSR: Tadzhikistan, on Salix sp., ZIN; Abd-Rabou et al. 

2005. 

Asterobemisia salicaria (Danzig); Martin & Mound 2007: 23. 

Distribution: 3: Iran, Tadzhikistan, USSR. 

Host: Salicaceae: Populus nigra, Salix alba, Salix sp. 

Natural enemies: Aphelinidae: Encarsia lutea, Encarsia tricolor. 

231

Asterobemisia silvatica (Danzig) 1964 

Bemisia silvatica Danzig 1964a: 638[327]. Holotype. USSR: Caucasian Black Sea coast, on Carpinus sp., 

ZIN. 

Asterobemisia silvatica (Danzig); Martin & Mound 2007: 23. 

Distribution: 3: Iran, Russia; USSR. 

Host: Apocynaceae: Nerium oleander; Betulaceae: Carpinus avellana, Carpinus betulus, 

Carpinus sp.; Ericaceae: Rhododendron flavum; Fabaceae: Acacia sp.; Fagaceae: Quercus 

calliprinos, Quercus sp.; Rhamnaceae: Fragula alnus; Rosaceae: Crataegus sp. 

Natural enemies: Aphelinidae: Encarsia cibcensis; Platygastridae: Amitus longicornis. 

Asterobemisia takahashii Danzig 1966 

Asterobemisia takahashii Danzig 1966: 376. Holotype. USSR: southern Maritime Territory, on Quercus 

mongolica, ZIN. 

Distribution: 5: Maritime Territory (USSR); Korea. 

Host: Fabaceae: Pueraria thunbergiana; Fagaceae: Quercus mongolica. 

Asterobemisia trifolii (Danzig) 1966 

Neobemisia trifolii Danzig 1966: 374 [203].Holotype. USSR: southern Maritime Territory, on Trifolium 

luginaster, ZIN. 

Asterobemisia trifolii (Danzig); Mound and Halsey 1978: 107. 

Distribution: 3: Iran; 5: Martime Territory (USSR). 

Host: Fabaceae: Trifolium lupinaster, Trifolium pratensis. 

Asterobemisia yanagicola (Takahashi) 1934 

Bemisia yanagicola Takahashi 1933: 17. Syntypes. China: on "yanagi" [Salix sp.], TARI. 

Asterobemisia yanagicola (Takahashi); Mound and Halsey 1978: 107. 

Distribution: 5: China; Japan; Taiwan. 

Host: Salicaceae: Salix glandulosa var warburgi, Salix sp. 

232

Asterochiton Maskell 1879 

Asterochiton Maskell 1879: 214. Type species. Asterochiton aureus Maskell 1879, by subsequent designation 

Cockerell 1902: 282. 

Aleyrodes (Asterochiton) Maskell; Cockerell 1902a: 282. 

Dialeurodoides Quaintance & Baker 1914: 98. Type species. Asterochiton aureus Maskell 1879, by original 

designation; synonymized by Quaintance & Baker 1915a: xi. 

Asterochiton Maskell; Quaintance & Baker 1915a: xi. 



aureus x 

auricolor x 

cerata x 

cordiae x 

fagi x 

pittospori x 

simplex x 

Total = 7 species 0 1 0 0 0 1 0 5 0 

Asterochiton aureus Maskell 1879 

Asterochiton aureus Maskell 1879: 216. Lectotype (designated by Dumbleton 1957:149). New Zealand: on 

Melicytus ramiflorus, ADSIR. 

Aleurodes melicyti Maskell 1890b: 174. Lectotype (designated by Dumbleton 1957:149). New Zealand: on 

Melicytus ramiflorus, ADSIR; synonymy by Cockerell 1902a: 281. 


Host: Violaceae: Melicytus ramiflorus. 

Asterochiton auricolor (Bondar) 1923 

Dialeurodoides [sic] auricolor Bondar 1923a: 119. Syntypes. Brazil: on Rubiaceae. 

Asterochiton auricolor (Bondar); Costa Lima 1968: 117. 

Distribution: 2: Brazil, Panama. 

233

Host: Rubiaceae. 

Asterochiton cerata (Maskell) 1895 

Aleurodes cerata Maskell 1895: 425. Lectotype. New Zealand: on Nothofagus menziesii, ADSIR. 

Asterochiton cerata (Maskell); Dumbleton 1957: 149. 


Host: Fagaceae: Nothofagus menziesii. 

Asterochiton cordiae David & Subramaniam 1976 

Asterochiton cordiae David & Subramaniam 1976: 177. Holotype. India: on Cordia myxa, USNM. 


Host: Boraginaceae: Cordia myxa 

Asterochiton fagi (Maskell) 1890 

Aleurodes fagi Maskell 1890a: 175. Lectotype. New Zealand: on Nothofagus menziesii, ADSIR. 

Dialeurodoides fagi (Maskell); Quaintance & Baker 1914: 99. 

Asterochiton fagi (Maskell); Quaintance & Baker 1915a: xi. 


Host: Fagaceae: Nothofagus menziesii. 

Asterochiton pittospori Dumbleton 1957 

Asterochiton pittospori Dumbleton 1957: 151. Holotype. New Zealand: on Pittosporum eugenioides, ADSIR. 


Host: Pittosporaceae: Pittosporum eugenioides. 

Asterochiton simplex (Maskell) 1890 

Aleurodes simplex Maskell 1890b: 175. Lectotype. New Zealand: on unspecified host, ADSIR. 

Dialeurodoides simplex (Maskell); Quaintance & Baker 1914: 99. 

Asterochiton simplex (Maskell); Quaintance & Baker 1915a: xi. 


Host: Pittosporaceae: Pittosporum eugenioides; Rubiaceae: Coprosma lucida. 

Asterochiton sp. 

Aphelinidae: Encarsia strenua (Silvestri) 1927 

234

Axacalia Danzig 1969 

Axacalia Danzig 1969: 876[556]. Type species. Axacalia spiraeanthi Danzig 1969, by monotypy. 



spiraeanthi x 

Total = 1 species 0 0 1 0 0 0 0 0 0 

Axacalia spiraeanthi Danzig 1969 

Axacalia spiraeanthi Danzig 1969: 876[556]. Holotype. Kazakhstan (USSR): Betpakdala, on Spiraeanthus 

schrenkianus, ZIN. 

Distribution: 3: Iran, Kazakhstan (USSR). 

Host: Convolvulaceae: Convolvulus arvensis; Rosaceae: Crataegus pentagyna; Spiraeanthus schrenkianus. 

235

Bellitudo Russell 1943 

Bellitudo Russell 1943: 132. Type species. Bellitudo jamaicae Russell 1943, by original description. 



campae x 

cubae x 

hispaniola x 

jamaicae x 

Total = 4 species 0 4 0 0 0 0 0 0 0 

Bellitudo campae Russell 1943 

Bellitudo campae Russell 1943: 136. Holotype. Jamaica: on Coccoloba uvifera, USNM. 

Distribution: 2: Jamaica. 

Host: Polygonaceae: Coccoloba uvifera. 

Bellitudo cubae Russell 1943 

Bellitudo cubae Russell 1943: 137. Holotype. Cuba: on Coccoloba retusa, USNM. 

Distribution: 2: Cuba. 

Host: Polygonaceae: Coccoloba retusa. 

Bellitudo hispaniola Russell 1943 

Bellitudo hispaniola Russell 1943: 136. Holotype. Haiti: on Coccoloba laurifolia, USNM. 

Distribution: 2: Dominican Republic; Haiti. 

Host: Polygonaceae: Coccoloba diversifolia, Coccoloba laurifolia. 

Bellitudo jamaicae Russell 1943 

Bellitudo jamaicae Russell 1943: 135. Holotype. Jamaica: on Coccoloba longiflora, USNM. 


Host: Polygonaceae: Coccoloba longiflora, Coccoloba uvifera, Coccoloba venosa. 

236

Bemisaleyrodes Cohic 1969 

Bemisaleyrodes Cohic 1969: 103. Type species. Bemisia grjebinei Cohic 1968, by original designation; Bink- 

Moenen 1983: 90. 



balachowskyi x 

brideliae x 

grjebinei x 

pauliani x 

Total = 4 species 0 0 0 4 0 0 0 0 0 

Bemisaleyrodes balachowskyi Cohic 1969 

Bemisaleyrodes balachowskyi Cohic 1969: 103. Holotype. Sao Tome: on Combretaceae. 

Distribution: 4: Sao Tome. 

Host: Combretaceae. 

Bemisaleyrodes brideliae Bink-Moenen 1983 

Bemisaleyrodes brideliae Bink-Moenen 1983: 91. Holotype pupal case, Chad, Boro, 6.xii.1973, FAB & 

RMBM, on Bridelia ferruginea; BMNH. 


Host: Euphorbiaceae: Bridelia ferruginea. 

Bemisaleyrodes grjebinei (Cohic) 1968 

Bemisia grjebinei Cohic 1968b: 117. Syntypes. Congo: on Alchornea cordifolia. 

Bemisaleyrodes grjebinei (Cohic); Cohic 1969: 102; -Bink-Moenen 1983: 92. 

Distribution: 4: Congo; Ivory Coast; Nigeria 

Host: Euphorbiaceae: Alchornea cordifolia, Fabaceae: Detarium microcarpum; Sapindaceae: 

Allophylus africanus, Paullinia pinnata. 

237

Bemisaleyrodes pauliani Cohic 1969 

Bemisaleyrodes pauliani Cohic 1969: 107. Holotype. Ivory Coast: on Ficus lepriuri; 

Bink-Moenen 1983: 94. 

Distribution: 4: Cameroon; Ivory Coast; Nigeria; Sudan; Kenya. 

Host: Moraceae: Ficus lepriuri, Ficus sp;Ulmaceae: Chaetachme aristata. 

238

Bemisia Quaintance & Baker 1914 

Bemisia Quaintance & Baker 1914: 99. Type species. Aleurodes inconspicua Quaintance 1900:28 (junior 

synonym of Aleurodes tabaci Gennadius 1889). 

Cortesiana Goux 1988: 63. Type species. Cortesiana restonicae Goux 1988, by monotypy; synonymy 


Roucasia Goux 1940: 45. Type species. Roucasia ovata Goux 1940, by monotypy; synonymy according to 

Danzig 1964a: 326. 

Reference: Regu & David 1991 - key to Indian species. 



afer x x x x x x 

alni x 

antennata x 

bambusae x 

berbericola x x x 

capitata x x 

caudasculptura x 

centroamericana x 

combreticula x 

confusa x 

cordylinidis x 

decipiens x 

elliptica x 

eoa x 

formosana x x 

giffardi x x x x x x x 

gigantea x 

grossa x 

guieriae x 

239


hirta x 

iole x 

lampangensis x 

lauracea x 

leakii x x 

medinae x 

mesasiatica x 

moringae x 


ovata x 

poinsettiae x 

pongamiae x x x 

porteri x x x 

psiadiae x 

puerariae x 

religiosa x 

shinanoensis x x 

spiraeae x 

spiraeoides x 

subdecipiens x 

sugonjaevi x 

tabaci x x x x x x x x x 

tabaci Q biotype* x x x 

tuberculata x 

Total = 42 species 2 6 12 10 12 15 7 4 2 

Bemisia afer (Priesner & Hosny) 1934 

Dialeurodoides afer Priesner & Hosny 1934b: 6. Syntypes. Egypt, Kom Ombo, 4.vii.1931, on Lawsonia alba, 

USNM. 

Bemisia afer (Priesner & Hosny); Habib & Farag 1970: 8. 

Bemisia (Neobemisia) afra [sic] (Priesner & Hosny); Visnya 1941b: 8. 

Bemisia citricola Gomez-Menor 1945a: 293. Syntypes. Spain, Orihuela, Alicante, on Citrus limonium, Citrus 

240

aurantium, Eucalyptus sp., Morus sp., Cynanchum acutum, Laurus nobilis. 

Bemisia hancocki Corbett 1936: 114. Syntypes. Uganda: 1934, G. Hancock, on cotton [Gossypium sp.]; 

BMNH; synonymy according to Bink-Moenen 1983: 95. 

Bemisia (Neobemisia) hancocki Corbett; Visnya 1941b: 8. 

Distribution: 2: Brazil; 3: Egypt, Iran, Israel; Italy; Sicily; Spain; 4: Cameroon; Chad; Congo; Guinea, Ivory 

Coast; Kenya; Madagascar; Mulawi; Niger; Nigeria; Sierre Leon; South Africa; Sudan; Uganda; Zaire; 

5: China, Korea; 6: India; Pakistan; 7: Australia; New Guinea. 

Host: Anacardiaceae: Lannea shimperi; Annonaceae: Annona senegalensis, Hexalobus monopetalus, Pistacia 

mutica; Apocynaceae: Nerium oleander; Bignoniaceae: Steroespermum kunthianum; Bixaceae: 

Cochlopermum tinctorium; Bombacaceae: Bombax costatum; Burseraceae: Commiphora pedunculata; 

Celastraceae: Maytenus senegalensis, Clethra arborea, Clethra sp.; Caprifoliaceae: Lonicera sp.; 

Combretaceae: Anogeissus leiocarpus, Combretum confertum, Combretum glutinosum, Combretum 

hypopilinum, Combretum molle; Euphorbiaceae: Antidesma venosum, Bridelia ferruginea, 

Hymenocardia acida, Manihot esculenta, Ricinus communis; Fabaceae: Acacia nilotica, 

Adenodolichos paniculatus, Albizia sp., Burkea africana, Cassia siamea, Daniellia olivera, Detarium 

microcarpum, Dichrostachys glomerata, Entada africana, Isoberlinia doka, Neorautanenia 

pseudopachyrhiza, Piliostigma thonningii, Pterocarpus lucens, Robinia pseudoacacia, Terminalia 

laxiflora; Liliaceae: Smilax aspera; Loganiaceae: Strychnos spinosa; Lythraceae: Lawsonia inermis; 

Malvaceae: Gossypium hirsutum, Gossypium sp., Hibiscus asper, Malva parviflora; Moraceae: Ficus 

nitida, Ficus sycamorus, Ficus sycomorus; Myrtaceae: Psidium guajava; Papaveraceae: Bocconia 

frutescens; Rhamnaceae: Ziziphus spina-christi; Rosaceae: Rosa chinensis, Rosa hybrida, Rosa 

multiflora, Rosa sp., Spiraea cantonensis; Rubiaceae: Crossopoteryx febrifuga, Gardenia triacantha, 

Nauclea latifolia; Rutaceae: Citrus aurantium, Citrus limon, Citrus limonia, Citrus sinensis, Clausena 

anisata; Salicaceae: Salix subserrata; Sapindaceae: Allophylus africanus; Paullinia pinnata; 

Solanaceae: Solanum muricatum; Urticaceae: Gesnouinia arborea, Urtica dioica. 

Natural enemies: Parasitoids - Hymenoptera/Aphelinidae: Cales noacki, Encarsia galilea, Encarsia 

levadicola, Encarsia melanostoma, Encarsia noahi, Encarsia pergandiella, Encarsia silvestrii, 

Encarsia sophia, Eretmocerus mundus, Eretmocerus roseni, Eretmocerus sp.; Eulophidae: 

Euderomphale bemisiae, Euderomphale cortinae, Euderomphale gomer; Predators - Coleoptera; 

Coccinellidae: Serangium parcesetosum. 

US Port Interceptions: Ethiopia on Piper sp.; Israel on Vitis sp.; Mexico on Choisya ternata, Citrus sinensis; 

Spain on Laurus nobilis. 

Bemisia alni Takahashi 1957 

Bemisia alni Takahashi 1957: 17. Syntypes (3). Japan: on Alnus sp., HBL. 


Host: Betulaceae: Alnus sp. 

Bemisia antennata Gameel 1968 

Bemisia antennata Gameel 1968: 149. Holotype. Sudan: on Ficus sycamorus.Syntypes (3). Japan: on Alnus 

241

sp., GRF; Bink-Moenen 1983: 98. 

Distribution: 4: Chad; Sudan. 

Host: Boraginaceae: Cordia africana; Fabaceae: Burkea africana, Pithecellobium dulce; Meliaceae: Khaya 

senegalensis; Moraceae: Ficus sycamorus, Ficus dicranostyla, Ficus sycomorus, Ficus sp.; Rubiaceae: 

Nauclea latifolia 

Bemisia bambusae Takahashi 1942 

Bemisia bambusae Takahashi 1942b: 172. Syntypes. Thailand: on bamboo [Bambusa? sp.], TARI. 


Host: Poaceae: Bambusa sp.? 

Bemisia berbericola (Cockerell) 1896 

Aleurodes herbericola Cockerell 1896b: 207. Syntypes. USA: New Mexico, on Berberis sp., USNM. 

Bemisia berbericola (Cockerell); Quaintance & Baker 1914: 100. 

Distribution: 1: USA (AZ, CA, FL, ID, NM, OR, TX); 2: Chile, Guatemala, Mexico; 6: India. 

Host: Asteraceae: Baccharis sp.; Berberidaceae: Berberis sp.; Betulaceae: Carpinus caroliniana. 

Bignoniaceae: Bignonia capreolata; Ericaceae: Gaylussacia frondosa; Euphorbiaceae: Colliguaja sp.; 

Moraceae: Morus sp.; Myricaceae: Myrica cerifera; Rhamnaceae: Ceanothus sp., Colubrina elliptica, 

Rhamnus crocea; Rosaceae: Photinia arbutifolia; Sapotaceae: Bumelia celestrina. 

US Port Interceptions: Netherlands on Gerbera sp. 

Bemisia capitata Regu & David 1991 

Bemisia capitata Regu and David 1991: 78. Holotype pupal case. India, Tamil Nadu, Padappai, 27.v.1989, K. 

Regu, on Rivea hypocrateriformis; Regu Collection. 


Host: Chenopodiaceae: Chenopodium album; Convolvulaceae: Rivea hypocrateriformis; Fabaceae: 

Glycyrrhiza glabra; Vitaceae: Vitis vinifera. 

Bemisia caudasculptura Quaintance & Baker 1937 

Bemisia caudasculptura Quaintance & Baker 1937, in Baker 1937: 614. Syntypes. Mexico: on ash [Fraxinus 

sp.], USNM. 


Host: Oleaceae: Fraxinus sp. 

Bemisia centroamericana Martin 2005 

Bemisia centroamericana Martin 2005b: 33. Holotype puparium, Belize, CFR, 13 II 1996, JH Martin, on 

Bocconia frutescens, BMNH. 

242

Distribution: 2: Belize, El Salvador, Honduras, Mexico. 

Host: Asteraceae: Lasianthaea? sp.; Euphorbiaceae: Manihot esculenta, Poinsettia sp.; Lamiaceae: Origanum 

vulgare; Malvaceae: Hibiscus sp.; Moraceae: Ficus sp.; Myrtaceae: Psidium guajava; Papaveraceae: 

Bocconia frutescens; Sapindaceae: Serjania sp.; Sapotaceae: Calocarpum sp. 

Bemisia combreticula Bink-Moenen 1983 

Bemisia combreticula Bink-Moenen 1983: 98. Holotype pupal case, Chad, Bebedjia, 27.vii.1972, FAB & 

RMBM, on Anogeissus leocarpus; BMNH. 


Host: Combretaceae: Anogeissus leiocarpus. 

Bemisia confusa Danzig 1964 

Bemisia confusa Danzig 1964a: 637. Holotype. USSR: Caucasian Black Sea coast, on Psoralea bituminosa, 

ZIN. 


Host: Fabaceae: Psoralea bituminosa. 

Bemisia cordylinidis Dumbleton 1961 

Bemisia cordylinidis Dumbleton 1961a: 120. Holotype. New Caledonia: on Cordyline sp., ADSIR. 


Host: Liliaceae: Cordyline sp. 

Bemisia decipiens (Maskell) 1895 

Aleurodes decipiens Maskell 1895: 428. Lectotype. Australia: on Styphelia [Monotoca] elliptica, ADSIR. 

Bemisia decipiens (Maskell); Quaintance & Baker 1914: 100. 


Host: Epacridaceae: Monotoca elliptica. 

Bemisia elliptica Takahashi 1960 

Bemisia elliptica Takahashi 1960b: 147. Syntypes. Reunion Island: on unidentified host, MNHNP. 



Bemisia eoa Danzig 1966 

Bemisia eoa Danzig 1966: 374[202]. Holotype. USSR: southern Maritime Territory, on Ulmus propinqua, 

243

ZIN. 

Distribution: 5: Maritime Territory (former USSR). 

Host: Caprifoliaceae: Lonicera sp.; Rosaceae: Sorbaria sorbifolia; Ulmaceae: Ulmus propinqua. 

Bemisia formosana Takahashi 1933 

Bemisia formosana Takahashi 1933: 18. Holotype. Taiwan: on Graminae [Poaceae], TARI. 

Bemisia graminis David & Augustine 1988:33. Holotype pupal case, India, Tamil Nadu, Padappai, 

21.vii.1986, Apluda mutica; David Collection; synonymy according to Martin & Mound 2007. 


Host: Poaceae: Apluda mutica, Echinochloa crusgalli, Echinochloa colonum, Oryza sativa. 

Bemisia giffardi (Kotinsky) 1907 

Aleyrodes giffardi Kotinsky 1907: 94. Syntypes. USA: Hawaii, on Citrus spp, USNM. 

Bemisia giffardi (Kotinsky); Quaintance & Baker 1914: 100. 

Bemisia giffardi hispina Young 1942: 98. Syntypes. China: on Citrus sp.; synonymy according to Mound and 

Halsey 1978: 113. 

Asterobemisia helyi Dumbleton 1956b: 172. Holotype. Australia: on Citrus sp., ADSIR; synonymy according 

to Mound and Halsey 1978: 113. 

Bemisia jasminum David & Subramaniam 1976: 181. Holotype. India: on Jasminum sp., USNM; synonymy 

according to Mound and Halsey 1978: 113. 

Distribution: 3: Iran; 4: Africa; 5: China; Hong Kong Japan; Taiwan; 6: India; Malaysia; Malaysia; Nepal; 

Thailand; Vietnam; 7: Australia, Indonesia; 8: Guam, New Zealand; New Caledonia; 9: Hawaii. 

Host: Boraginaceae: Cordia cordata; Rutaceae: Citrus bigaradia, Citrus decumana, Citrus maxima, Rutaceae: 

Citrus sp.; Oleaceae: Jasminum sambac. 

Natural enemies: Fungus - Aschersonia aleyrodis, Aschersonia andropogonis, Aschersonia goldiana; 

Parasitoids - Aphelinidae: Encarsia strenua. 

US Port Interceptions: Hawaii on Citrus sp.; Hong Kong on Clausena lansium; India on Gardenia 

jasminoides, Jasminum multiflorum, Murraya koenigii; Lebanon on Jasminum sp.; Mexico on Persea 

americana; Sri Lanka on Murraya sp. 

Bemisia gigantea Martin 1999 

Bemisia gigantea Martin 1999: 58. Holotype puparium. Australia: on Elaeocarpus sp., ANIC. 


Host: Elaeocarpaceae: Elaeocarpus sp. 

244

Bemisia grossa Singh 1931 

Bemisia grossa Singh 1931: 82. Syntypes. India: on Eugenia operculata; Dubey et al. 2004. 


Host: Fabaceae: Derris sp.; Myrtaceae: Eugenia operculata. 

Bemisia guieriae Bink-Moenen 1983 

Bemisia guieriae Bink-Moenen 1983: 101. Holotype pupal case, Chad, Bebedjia, 27.viii.1972, FAB & 

RMBM, on Guiera senegalensis; BMNH. 


Host: Combretaceae: Guiera senegalensis. 

Bemisia hirta Bink-Moenen 1983 

Bemisia hirta Bink-Moenen 1983: 103. Holotype pupal case, Chad, Bebedjia,12.vi.1973, FAB & RMBM, on 

Capparis corymbosa; BMNH. 


Host: Capparidaceae: Capparis corymbosa. 

Bemisia iole Danzig 1966 

Bemisia iole Danzig 1966: 374[202]. Holotype. USSR: southern Maritime Territory, on Ulmus propinqua, 

ZIN. 

Distribution: 5: Maritime Territory (USSR). 

Host: Betulaceae: Corylus heterophylla; Ulmaceae: Ulmus propinqua. 

Bemisia lampangensis Takahashi 1942 

Bemisia lampangensis Takahashi 1942b: 171. Holotype. Thailand: on legume [Fabaceae]. 


Host: Fabaceae, a legume. 

Bemisia lauracea Martin, Aguiar & Pita 1996 

Bemisia lauraceae Martin, Aguiar & Pita 1996, in Martin et al. 1996: 113. Holotype pupal case, Madeira, Faja 

da Noguiera, 6.iii.1992, F. Aguiar, on Ocotea foetens (Lauraceae), BMNH. 

Distribution: 3: Madeira. 

Host: Lauraceae: Laurus azorica, Ocotea foetens, Persea indica. 

Natural enemies: Aphelinidae: Encarsia levadicola. 

245

Bemisia leakii (Peal) 1903 

Aleurodes leakii Peal 1903b: 87. Syntypes. India: on Indigofera arrecta and I. tinctoria. 

Bemisia leakii (Peal); Quaintance & Baker 1914: 100. 

Distribution: 6: India; 8: Fiji; Tahiti. 

Host: Fabaceae: Dalbergia sp., Erythrina sp., Indigofera arrecta, Indigofera tinctoria. 

Note: Martin (1985: 321) reported Bemisia leakii? (Peal) from Papua New Guinea on 

Araceae: Colocasia sp. 

Bemisia medinae Gomez-Menor 1954 

Bemisia (Roucasia) medinae Gomez-Menor 1954: 369. Syntypes. Canary Islands: on unknown plant. 

Bemisia medinae Gomez-Menor; Mound and Halsey 1978: 115; Hernandez et al. 2003: 77. 

Distribution: 4: Canary Islands. 

Host: Guttiferae: Hypericum grandifolium. 

Natural enemies: Aphelinidae: Encarsia melanostoma; Eulophidae: Euderomphale gomer. 

Bemisia mesasiatica Danzig 1969 

Bemisia mesasiatica Danzig 1969: 871 [553]. Holotype. USSR: Tadzhikistan, on Spiraea baldshuanica, ZIN. 

Distribution: 3: Iran, Tadzhikistan, USSR. 

Host: Caprifoliaceae: Lonicera korolkovii; Rosaceae: Armeniaca sp., Crataegus microphylla, Crataegus 

pentagyna, Exochorda alberti, Prunus domestica var. insititia, Spiraea baldshuanica; Ulmaceae: 

Ulmus procera. 

Bemisia moringae (David & Subramaniam) 1976 

Asterobemisia moringae David & Subramaniam 1976: 176. Holotype. India: on Moringa oleifera, IDAV. 

Bemisia moringae (David & Subramaniam); Mound and Halsey 1978: 116. 


Host: Moringaceae: Moringa oleifera. 

Bemisia multituberculata Sundararaj & David 1990 

Bemisia multituberculata Sundararaj & David 1990: 113. Holotype puparium, India, Vellimalai, 3.viii.1987, 

R. Sundararaj, on Gmelina sp. 


Host: Verbenaceae: Gmelina sp. 

246

Bemisia ovata (Goux) 1940 

Roucasia ovata Goux 1940: 45. Syntypes. France: on Hedera helix. 

Bemisia ovata (Goux); Danzig 1964a: 326 [637]. 

Distribution: 3: France; Iran, Russia, USSR. 

Host: Araliaceae: Hedera helix; Ericaceae: Rhododendron ponticum; Lauraceae: Laurus nobilis. 

Liliaceae: Smilax excelsa; Moraceae: Morus nigra, Morus sp.; Ranunculaceae: Clematis vitalba; 

Rutaceae: Citrus x paradisi, Citrus sp.; Vitaceae: Vitis sp. 

Natural enemies: Aphelinidae: Eretmocerus mundus. 

Bemisia poinsettiae Hempel 1923 

Bemisia poinsettiae Hempel 1923: 1175. Syntypes. Brazil: on Poinsettia heterophylla, MZU. 

Distribution: 2: Argentina; Brazil. 

Host: Euphorbiaceae: Poinsettia heterophylla. 

Bemisia pongamiae Takahashi 1931 

Bemisia pongamiae Takahashi 1931c: 223. Syntypes. Taiwan: on Pongamia pinnata, TARI; Martin 1985: 

322. 

Distribution: 5: Taiwan; 6: Malaya; Malaysia; 7: Papua New Guinea, Sulawesi. 

Host: Fabaceae: Derris sp, Pongamia pinnata. 

Bemisia porteri Corbett 1935 

Bemisia porteri Corbett 1935b: 786. Syntypes. Malaya: on Quisqualis indica; Takahashi 1955a: 406. 

Distribution: 4: Madagascar; Taiwan; 6: Malaya; Malaysia. 

Host: Combretaceae: Quisqualis indica. 


Bemisia psiadiae Takahashi 1955 

Bemisia psiadiae Takahashi 1955a: 405. Syntypes. Madagascar: on Psiadia sp., MNHNP. 


Host: Asteraceae: Psiadia sp. 

Bemisia puerariae Takahashi 1955 

Bemisia puerariae Takahashi 1955c: 3. Syntypes. Japan: on Pueraria hirsuta, HBL. 

247


Host: Fabaceae: Desmodium sp, Pueraria hirsuta. 

Bemisia religiosa (Peal) 1903 

Aleurodes religiosa Peal 1903b: 67. Syntypes. India: on Ficus bengalensis and F. religiosa. 

Bemisia religiosa (Peal); Quaintance & Baker 1914: 100. 


Host: Moraceae: Ficus bengalensis, Ficus religiosa. 

Bemisia shinanoensis Kuwana 1922 

Bemisia shinanoensis Kuwana 1922: 461. Syntypes. Japan: on cultivated mulberry [Morus sp.]. 

Distribution: 3: Iran; 5: Japan. 

Host: Asteraceae: Pertya scandens; Berberidaceae: Berberis thunbergii; Betulaceae: Corylus avellana; 

Ericaceae: Rhododendron sp.; Fabaceae: Desmodium fallax, Pueraria hirsuta; Lauraceae: Lindera 

umbellatum; Moraceae: Morus sp., Morus alba; Rosaceae: Rosa sp.; Ulmaceae: Aphananthe aspera; 

Ulmus carpinifolia; Verbenaceae: Callicarpa japonica. 

Bemisia spiraeae Young 1944 

Bemisia spiraeae Young 1944: 133. Syntypes. China: on Spiraea cantoniensis. 


Host: Rosaceae: Spiraea cantoniensis. 

Bemisia spiraeoides Mound & Halsey 1978 

Bemisia spireae [sic] Gomez-Menor 1954: 373. Syntypes. Spain: on Spiraea hipericifolia and Rosa sp. 

Bemisia spiraeoides Mound and Halsey 1978: 118, replacement name, B. spiraeae preoccupied by Bemisia 

spiraeae Young 1944. 

Distribution: 3: Spain. 

Host: Rosaceae: Rosa sp., Spiraea hipericifolia 

Bemisia subdecipiens Martin 1999 

Bemisia subdecipiens Martin 1999: 59. Holotype puparium. Australia: on Melaleuca sp., ANIC. 


Host: Myrtaceae: Melaleuca sp. 

248

Bemisia sugonjaevi Danzig 1969 

Bemisia sugonjaevi Danzig 1969: 870[553]. Holotype. USSR: Tadzhikistan, on Trachomitum scabrum, ZIN. 

Distribution: 3: Iran, USSR: Tadzhikistan. 

Host: Apocynaceae: Nerium oleander, Trachomitum scabrum. 

Natural enemies: Aphelinidae: Encarsia mohyuddini. 

Bemisia tabaci (Gennadius) 1889 

Aleurodes tabaci Gennadius 1889: 1-3. Syntypes. Greece: on tobacco [Nicotiana sp.], USNM. 

Bemisia tabaci (Gennadius); Takahashi 1936f: 110. 

Bemisia argentifolii Bellows & Perring 1994; synonymy according to De Barro et al. 2005: 201. 

Bemisia achyranthes Singh 1931: 82. Syntypes. India: on Achyranthes aspera [synonmized with B. 

gossypiperda by Corbett 1935b: 783]. 

Bemisia argentifolii Bellows & Perring 1994. Holotype pupal case. USA: California, xii.1992, stock culture, 

on Phaseolus limensis. 

Bemisia bahiana Bondar 1928: 30. Syntypes. Brazil: on Nicotiana tabacum. 

Bemisia costa-limai Bondar 1928: 27. Syntypes. Brazil: on Euphorbia hirtella. 

Bemisia emiliae Corbett 1926: 273. Syntypes. Sri Lanka: on Emilia sonchifolia. 

Bemisia goldingi Corbett 1935: 249. Syntypes. Nigeria: on cotton [Gossypium sp.]. 

Bemisia gossypiperda Misra & Lamba 1929:1. Syntypes. India: on many plants. 

Bemisia gossypiperda var mosaicivectura Ghesquiere 1934: 30. Syntypes. Zaire: on Jatropha 

multifida and Manihot sp. 

Bemisia hibisci Takahashi 1933: 17. Syntypes. Taiwan: on Hibiscus rosa-sinensis. 

Aleurodes inconspicua Quaintance 1900: 28. Syntypes. USA; Florida, Barlow, on Physalis sp., USNM; 

Russell 1957: 122. 

Bemisia longispina Priesner & Hosny 1934: 6. Syntypes. Egypt: on Psidium guajava. 

Bemisia lonicerae Takahashi 1957: 16. Syntypes. Japan: on Lonicera japonica. 

Bemisia manihotis Frappa 1938: 30. Syntypes. Madagascar: on Manihot sp. 

Bemisia minima Danzig 1964: 638. Holotype. USSR: Caucasian Black Sea coast, on Elsholtzia patrini. 

Bemisia miniscula Danzig 1964: 640. Holotype. USSR: Adzharia, on Cistus salvifolius. 

Bemisia nigeriensis Corbett 1935: 250. Syntypes. Nigeria: on cassava [Manihot sp.]. 

Bemisia rhodesiansis Corbett 1936: 22. Syntypes. Rhodesia: on tobacco [Nicotiana sp.]. 

Bemisia signata Bondar 1928: 29. Syntypes. Brazil: on Nicotiana glauca. 

Bemisia vayssierei Frappa 1939: 255. Syntypes. Madagascar: on tobacco [Nicotiana sp.]. 

Cortesiana restonicae Goux 1988: Holotype. Corsica. 

249

250 

Distribution: Virtually worldwide, 1: USA (AZ, CA, DC, FL, GA, HI, KS, LA, MD, MS, NE, SC, TX, VA); 

2: Argentina, Barbados, Brazil, Chile, Colombia, Cuba, Dominican Republic, Ecuador, El Salvador, 

French Guiana, Guadeloupe, Haiti, Honduras, Grenada, Guatemala, Jamaica, Mexico, Nicaragua, 

Panama, Peru, Puerto Rico, Trinidad & Tobago, Venezuela, Virgin Islands; 3: Afghanistan, Algeria, 

Canary Island, Cyprus, Egypt, England, France, Greece, Iran, Iraq, Israel, Italy, Jordan, Lebanon, 

Libya, Netherlands, Portugal, Romania, Saudi Arabia, Spain, Syria, Turkey, Yemen; 4: Cameroon, 

Central African Republic, Chad, Congo, Ethiopia, Gabon, Gambia, Ghana, Guyana, Ivory Coast, 

Kenya, Liberia, Madagascar, Mauritius, Mozambique, Nigeria, Sierra Leone, South Africa, Sudan, 

Uganda, United Kingdom, Zaire, Zimbabwe; 5: China, Hong Kong, Japan, Korea, Taiwan; 6: India, 

Malaysia, Pakistan, Philippines, Senegal, Seychelles, Singapore, Sri Lanka, Thailand; 7: Australia, 

New Guinea, Sumatra; 8: Caroline Islands, Fiji, Guam, Mariana Islands, Saipan, Tahiti; 9: Hawaii. 

Host: Over 250 host plants in many different families. 


Fungus - Aschersonia aleyrodis, Erynia radicans, Hyphomycetes: Paecilomyces fumosoroseus, Paecilomyces 

sp., Paecilomyces farinosus. 

Parasitoids - Hymenoptera/ Encarsia acaudaleyrodis, Encarsia adusta, Encarsia aleurothrixi, Encarsia 

azimi, Encarsia bimaculata, Encarsia brevivena, Encarsia cibcensis, Encarsia citrella, Encarsia 

davidi, Encarsia desantisi, Encarsia duorunga, Encarsia formosa, Encarsia guadeloupae, Encarsia 

hispida, Encarsia inaron, Encarsia japonica, Encarsia lanceolata, Encarsia longifasciata, Encarsia 

longivalvula, Encarsia lutea, Encarsia luteola, Encarsia melanostoma, Encarsia mineoi, Encarsia 

mohyuddini, Encarsia nigricephala, Encarsia noahi, Encarsia oakeyensis, Encarsia perflava, 

Encarsia pergandiella, Encarsia porteri, Encarsia protransvena, Encarsia pseudocitrella, Encarsia 

quaintancei, Encarsia reticulata, Encarsia smithi, Encarsia sophia, Encarsia synaptocera, Encarsia 

tabacivora, Encarsia tricolor, Encarsia sp., Eretmocerus adustiscutum, Eretmocerus aegypticus, 

Eretmocerus californicus, Eretmocerus eremicus, Eretmocerus hayati, Eretmocerus melanoscutus, 

Eretmocerus mundus, Eretmocerus mundus parthenogenetic form, Eretmocerus nikolskajae, 

Eretmocerus queenslandensis, Eretmocerus rui, Eretmocerus serius, Eretmocerus sp., Eretmocerus 

staufferi, Eretmocerus tejanus; Ceraphronidae: Aphanogmus fumipennis; Eulophidae: Neochrysocharis 

formosa; Platygastridae: Amitus bennetti; Signiphoridae: Signiphora sp. 

Predators - Anystidae: Walzia australica; Acari/Phytoseiidae: Amblyseius chilenensis, Amblyseius limonicus, 

Amblyseius ovalis, Amblyseius sp., Amblyseius swirski, Euseius aleyrodis, Euseius hibisci, Euseius 

metwallyi, Euseius scutalis, Genus? sp, Kampimodromus ricini, Metaseiulus occidentalis, Neoseiulus 

cucumeris, Typhlodromus athiasae, Typhlodromus medanicus, Typhlodromus pelargonicus, 

Typhlodromus sudanicus; Stigmaeidae: Agistemus exsertus; Araneidae: Araneus sp.; Neoscona 

arabesca; Clubionidae; Dictynidae: Dictyna annexa, Dictyna bellans, Phantyna segregata; Episininae; 

Tetragnathidae: Tetragnatha sp.; Theridiidae: Achaearanea sp., Coleosoma acutiventer, Dipoena 

abdita, Theridiidae: Latrodectus sp., Theridion australe, Theridula gonygaster, Theridula opulenta, 

Theridula sp.; Thomisidae: Misimenops sp.; Uloboridae: Philoponella sp.; Coleoptera/Carabidae: 

Callida sp., Cicindellidae: Cincindela viricolis; Coccinellidae: Adonia variegata, Brumoides sp., 

Brumoides suturalis, Chilocorus bipustulatus, Clitostethus arcuatus, Coccinella septempunctata, 

Coccinella sp., Coccinella transversalis, Coccinella undecimpunctata, Coelophora inequalis, 

Coleomegilla cubensis, Coleomegilla cubensis, Coleomegilla maculata, Coleomegilla maculata, 

Coleomegilla sp., Cycloneda sanguinea, Cycloneda sanguinea, Cycloneda sp., Delphastus catalinae, 

Delphastus dejavu, Delphastus pallidus, Delphastus pusillus, Delphastus sp, Diomus notescens,

251 

Eriopis connexa, Eriopis connexa, Exochomus marginipennis, Exochomus quadripustulatus, Genus? 

sp, Harmonia 14-punctata, Harmonia conglobata, Harmonia dimidiata, Harmonia sp., Hippodamia 

convergens, Megalocaria dilatata, Menochilus sexmaculatus, Menochilus sexmaculatus, Menochilus 

sp., Micraspis frenata, Micraspis sp., Nephaspis oculatus, Olla v-nigrum, Procula ferruginea, 

Scymnus sp., Scymnus syriacus, Scymnus variegatus, Serangium cinctum, Serangium parcesetosum, 

Serangium sp., Stethorus jejunus, Stethorus nigripes, Stethorus vagans, Verania quadrimaculata, 

Verania vincta; Melyridae: Collops vittatus, Dicranolaius bellulus; Nitidulidae: Cybocephalus 

binotatus, Cybocephalus sp.; Staphylinidae: Holoborus pallidicornis, Paederus alfierii, Paederus 

fuscipes; Dermaptera/Labiduridae: Doru lineare, Labidura riparia; Diptera/ Cecidomyidae: Genus? 

sp, Phaenobremia aphidivora; Dolichopodidae: Condylostylus sp.; Sciapus psittacinus; Drosophilidae: 

Acletoxenus formosus; Empididae: Drapetis sp., Drapetis subaenescens, Tachidronia annulata; 

Muscidae: Coenosia attenuata; Syrphidae: Allograpta obliqua, Ocyptamus parvicornis, Paragus 

aegyptius, Sphaerophora flavicauda, Syrphus corollae; Hemiptera/Anthocoridae: Cardiastethus 

assimilis, Orius albidipennis, Orius insidiosus, Orius minitus, Orius niger, Orius sp., Orius tristicolor; 

Berytidae: Jalysus wickhami, Geocoris lubra, Lygeidae: Geocoris ochropterus, Geocoris pallens, 

Geocoris punctipes, Geocoris sp.; Miridae: Campylomma diversicornis, Campylomma liebknechti, 

Campylomma nicolasi, Campylomma sp., Campylomma unicolor, Cyrtopeltis sp., Cyrtopeltis modesta, 

Cyrtopeltis tenuis, Cyrtopeltis varians, Deraeocoris indianus, Deraeocoris nebulosus, Deraeocoris 

pallens, Deraeocoris punctulatus, Deraeocoris signatus, Deraeocoris sp., Dicyphus errans, Dicyphus 

tamaninii, Halticus bractatus, Lygus hesperus, Macrolophus caliginosus, Macrolophus costalis, 

Nesidiocoris tenuis, Phytocoris sp.; Nabidae: Nabis alternatus, Nabis sp.; Reduviidae: Coranus 

spiniscutis, Empicoris sp., Sinea confusa, Sinea diadema, Zelus longipes, Zelus renardii, Zelus sp.; 

Neuroptera/Chrysopidae: Ceraeochrysa cubana, Chrysopa exterior, Chrysopa formosa, Chrysopa 

sp., Chrysoperla carnea, Chrysoperla externa, Chrysoperla plorabunda, Chrysoperla rufilabris, 

Chrysoperla sp., Genus? sp, Mallada boninensis, Mallada boninensis, Mallada flavifrons, Nineta 

flava, Pleisiochrysa lacciperda, Plesiochrysa lacciperda, Coniopterygidae: Conwentzia hagani; 

Coniopterygidae: Semidalis sp.; Hemerobidae: Hemerobius hageni, Micromus posticus; Odonata/ 

Coenagrionidae: Enallagma civile; Thysanoptera/ Phlaeothripidae; Thripidae: Franklinothrips 

vespiformis, Scolothrips latipennis, Sericothrips trifasciatus. 

US Port Interceptions: Often intercepted at US ports from different countries and on a wide variety of host 

plants. 

Bemisia tabaci biotype Q 

Distribution: 1: USA (AZ, CA); 3: Egypt, France, Israel, Italy, Morocco, Netherlands, Spain, Portugal; 5: 

China, Korea. 

Bemisia tuberculata Bondar 1923 

Bemisia tuberculata Bondar 1923a: 123. Syntypes. Brazil: on "mandioca" [Manihot aipi], MZU; Evans & 

Serra 2002: 207. 

Distribution: 2: Brazil; Nicaragua; Puerto Rico. 

Host: Euphorbiaceae: Chamaesyce hypericifolia, Manihot esculenta, Manihot aipi; Fabaceae: Erythrina sp.; 

Malvaceae: Gossypium hirsutum.

252 

Natural enemies: Parasitoids - Aphelinidae: Encarsia cubensis, Encarsia hispida, Encarsia sophia; 

Predators - Chrysopidae.

Bemisiella Danzig 1966 

Bemisiella Danzig 1966: 377[204]. Type species. Bemisiella artemisiae Danzig 1966, by original designation. 



artemisiae x x 

lespedezae x x 

Total = 2 species 0 0 2 0 2 0 0 0 0 

Bemisiella artemisiae Danzig 1966 

Bemisiella artemisiae Danzig 1966: 377 [204]. Holotype. USSR: Maritime Territory, on Artemisia sp., ZIN; 

Suh et al. 2006. 

Distribution: 3: Iran; 5: Maritime Territory (USSR); Korea. 

Host: Asteraceae: Artemisia princes, Artemisia vulgaris, Artemisia sp., Chrysanthemum x morifolium; 

Myrtaceae: Eucalpytus gracilis. 

Bemisiella lespedezae Danzig 1966 

Bemisiella lespedezae Danzig 1966: 378[205]. Holotype. USSR: Maritime Territory, on Lespedeza bicolor, 

ZIN. 

Distribution: 3: Iran; 5: Maritime Territory (USSR). 

Host: Fabaceae: Lespedeza bicolor; Polygonaceae: Rumex acetosella. 

253

Brazzaleyrodes Cohic 1966 

Brazzaleyrodes Cohic 1966a: 35. Type species. Brazzaleyrodes eriococciformis Cohic 1966, by monotypy. 



eriococciformis x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Brazzaleyrodes eriococciformis Cohic 1966 

Brazzaleyrodes eriococciformis Cohic 1966a: 35. Syntypes. Congo: on Cassia javanica; Bink-Moenen 1983: 

106. 

Distribution: 4: Congo; Ivory Coast; Nigeria. 

Host: Fabaceae: Albizia lebbeck, Cassia javanica, Peltophorum pterocarpa, Pentaclethra eetveldeana, 

Newtonia glandulifera, Cassia spectabilis. 

254

Bulgarialeurodes Corbett 1936 

Bulgarialeurodes Corbett 1936: 18. Type species: Bulgarialeurodes rosae Corbett 1936 (syn. Aleurodes 

cotesii Maskell 1896), by monotypy. 



cotesii x x 

Total = 1 species 0 0 1 0 0 1 0 0 0 

Bulgarialeurodes cotesii (Maskell) 1895 

Aleurodes cotesii Maskell 1895: 427. Syntypes. Pakistan: on Rosa sp., ADSIR. 

Bulgarialeurodes cotesii (Maskell); Russell 1960a: 30. 

Aleurodes rosae Kiriukhin 1947: 10. Syntypes. Iran: on Rosa spp; synonymy according to Russell 1960a: 30. 

Bulgarialeurodes rosae Corbett 1936: 18. Syntypes. Bulgaria: on Rosa damascena; synonymy according to 

Russell 1960a: 30. 

Distribution: 3: Afganistan; Bulgaria; Iran; Rumania; Turkmenistan; USSR; Yugoslavia; 6: Pakistan. 

Host: Rosaceae: Rosa canina, Rosa hemisphaerica, Rosa lutea, Rosa persica, Rosa sp. 

Natural enemies: Aphelinidae: Encarsia inaron, Encarsia lutea, Encarsia shutovae, Eretmocerus mundus , 

Eretmocerus nikolskajae. 

255

Calluneyrodes Zahradnik 1961 

Calluneyrodes Zahradnik 1961b: 65. Type species. Bemisia callunae Ossiannilsson 1947, by monotypy. 



callunae x 

Total = 1 species 0 0 1 0 0 0 0 0 0 

Calluneyrodes callunae (Ossiannilsson) 1947 

Bemisia callunae Ossiannilsson 1947: 1. Syntypes. Sweden: on Calluna sp., DPPE. 

Calluneyrodes callunae (Ossiannilsson); Zahradnik 1961b: 65. 

Distribution: 3: Czech Republic; Finland; Sweden. 

Host: Ericaceae: Calluna sp., Calluna vulgaris. 

256

Chitonaleyrodes MARTIN 1999 

Chitonaleyrodes Martin 1999: 61. Type species. Chitonaleyrodes canberrensis Martin 1999, by monotypy. 



canberrensis x 

Total = 1 species 0 0 0 0 0 0 1 0 0 

Chitonaleyrodes canberrensis Martin 1999 

Chitonaleyrodes canberrensis Martin 1999: 61. Holotype puparium. Australia: on Callistemom sp., ANIC. 


Host: Myrtaceae: Callistemon sp. 

257

Cockerelliella SUNDARARAJ & DAVID 1992 

Cockerelliella Sundararaj & David 1992: 417. Type species. Cockerelliella indica Sundararaj & David 1992, 


References: Sundararaj & David 1992; Sundararaj (2000) - key to Indian species. 



adinandrae x 

bladhiae x x 

curcumae x 

dehradunensis x 

dioscoreae x 

indica x 

kamardini x 

meghalayensis x 

psidii x x x 

quaintancei x 

rhodamniae x 

rotunda x 

sembilanensis x 

somnathensis x 

splendens x 

zingiberae x 

Total = 17 species 0 0 0 0 2 16 1 0 0 

Cockerelliella adinandrae (Corbett) 1935 

Dialeurodes adinandrae Corbett 1935b: 733. Syntypes. Malaya: on Adinandra dumosa and Ficus sp. 

258

Cockerelliella adinandrae (Corbett); Sundararaj & David 1992: 417. 


Host: Moraceae: Ficus sp.; Theaceae: Adinandra dumosa. 

Cockerelliella bladhiae (Takahashi) 1931 

Dialeurodes bladhiae Takahashi 1931c: 219. Syntypes. Taiwan: on Ardisia [=Bladhia] sieboldii, TARI. 

Cockerelliella bladhiae (Takahashi); Dubey (personal communication) 

Distribution: 5: Hong Kong; Taiwan; 6: Thailand. 

Host: Caprifoliaceae: Viburnum awabucki; Fagaceae: Lithocarpus sp.; Myrsinaceae: Ardisia sieboldii; 

Myrtaceae: Eugenia sp.; Theaceae: Camellia sp., Eurya acuminata. 

Cockerelliella curcumae (Corbett) 1935 

Dialeurodes curcumae Corbett 1935b: 738. Syntypes. Malaya: on Curcuma sp. 

Cockerelliella curcumae (Corbett); Sundararaj & David 1992: 417. 


Host: Zingiberaceae: Curcuma sp. 

Cockerelliella dehradunensis (Jesudasan & David) 1991 

Aleuroclava dehradunensis Jesudasan & David 1991: 258. Holotype. India: Uttar Pradesh, Dehra Dun, 

11ii.1985, R. W. Alexander Jesudasan, on Putaranjiva roxburghii. 

Cockerelliella dehradunensis (Jesudasan & David); Sundararaj 2000: 313. 


Host: Euphorbiaceae: Putaranjiva roxburghii; Flacourtiaceae: Flacourtia montana, Hydnocarpus pentandra; 

Loganiaceae: Strychnos wallichiana. 

Cockerelliella dioscoreae Sundararaj & David 1992 

Cockerelliella dioscoreae Sundararaj & David 1992: 417. Holotype puparium, India, Tamil Nadu, Munchirai, 

3.viii.1987, R. Sundararaj, on Dioscorea alata, David Collection. 


Host: Dioscoreaceae: Dioscorea alata. 

Cockerelliella indica Sundararaj & David 1992 

Cockerelliella indica Sundararaj & David 1992: 417. Holotype. India, Tamil Nadu, Coimbatore, 15.iv.1968, 

B. V. David, on Tabernaemontana [=Ervatomia] coronaria. 


259

Host: Apocynaceae: Tabernaemontana [=Ervatomia] coronaria. 

Cockerelliella kamardini (Corbett) 1935 

Dialeurodes kamardini Corbett 1935b: 735. Syntypes. Malaya; on unidentified host. 

Cockerelliella kamardini (Corbett); Sundararaj & David 1992: 417. 



Cockerelliella meghalayensis Sundararaj & David 1992 

Cockerelliella meghalayensis Sundararaj & David 1992: 420. Holotype puparium, India, Meghalaya, 

Nongkalaw, 1.x.1988, B. V. David, unidentified plant, David Collection. 



Cockerelliella psidii (Corbett) 1935 

Dialeurodes psidii Corbett 1935b: 734. Syntypes. Malaya: on Psidium guajava and other plants. 

Cockerelliella psidii (Corbett); Sundararaj & David 1992: 417. 

Dialeurodes lumpurensis Corbett 1935b: 739. Syntypes. Malaya: on unidentified host, lost; synonymy 


Distribution: 5: China; 6: Malaya; 7: Papau New Guinea; Sarawak. 

Host: Magnoliaceae: Michelia champaca; Moraceae: Artocarpus sp., Conocephalus subtrinervius, Ficus sp.; 

Musaceae: Musa basjoo; Myrtaceae: Psidium guajava; Theaceae: Camellia sp. 


Cockerelliella quaintancei Sundararaj & David 1992 

Cockerelliella quaintancei Sundararaj & David 1992: 421. Holotype puparium, India, Kerala, Ambalamedu, 

30.vii.1987, R. Sundararaj on Ailanthus sp, David Collection. 


Host: Simaroubaceae: Ailanthus sp. 

Cockerelliella rhodamniae (Corbett) 1935 

Dialeurodes rhodamniae Corbett 1935b: 736. Syntypes. Malaya: on Rhodamnia cinerea and other plants. 

Cockerelliella rhodamniae (Corbett); Sundararaj & David 1992: 417. 

Distribution: 6: India; Malaysia. 

260

Host: Anacardiaceae: Gluta sp.; Magnoliaceae: Michelia champaca; Moraceae: Ficus sp. 

Myrtaceae: Eugenia aquea, Eugenia malaccensis, Rhodamnia cinerea; Sterculiaceae: Sterculia laevis. 

Cockerelliella rotunda Regu & David 1993 

Cockerelliella rotunda Regu & David 1993c: 82. Holotype puparium, India, Tamil Nadu, Muchirai, 

3.viii.1987, K. Regu, on Capparis sp., IARI. 


Host: Capparidaceae: Capparis sp. 

Cockerelliella sembilanensis (Corbett) 1935 

Dialeurodes sembilanensis Corbett 1935b: 737. Syntypes. Malaya: on Ficus sp. 

Cockerelliella sembilanensis (Corbett); Sundararaj & David 1992: 417. 


Host: Liliaceae: Smilax wightii; Moraceae: Ficus sp. 

Cockerelliella somnathensis Sundararaj 2000 

Cockerelliella somnathensis Sundararaj 2000: 314. Holotype puparium, Gujarat, Somnath, 5.xii.1996, R. 

Sundararaj, on unidentified climber, David Collection. 


Host: unknown climber. 

Cockerelliella splendens Meganathan & David 1994 

Cockerelliella splendens Meganathan & David 1994: 30. Holotype. India: on Smilax wightii. 


Host: Liliaceae: Smilax wightii. 

Cockerelliella zingiberae Sundararaj & David 1992 

Cockerelliella zingiberae Sundararaj & David 1992: 424. Holotype puparium, India, Tamil Nadu, Manalodia, 

4.viii.1987, R. Sundararaj on Zingiber sp, David Collection. 


Host: Zingiberaceae: Zingiber sp. 

261

Cohicaleyrodes Bink-Moenen 1983 

Cohicaleyrodes Bink-Moenen 1983: 107. Type species. Cohicaleyrodes crossopterygis Bink-Moenen 1983, 





alternans x 

blanzyi x 

crossopterygis x 

descarpentriesi x 

elongatus x 

indicus x 

jesudasani x 

mappiae x 

obscura x 

padminiae x 

pauliani x 

platysepali x 

quadrilongispinae x 

recurvispina x 

saklespurensis x 

uvariae x 

Total = 16 species 0 0 0 10 0 6 0 0 0 

Cohicaleyrodes alternans (Cohic) 1966 

Aleurocanthus alternans Cohic 1966a. Syntypes. Congo: Brazzaville, Centre ORSTOM, 10.v.1964, on 

262

Strychnos variabilis, BMNH. 

Cohicaleyrodes alternans (Cohic); Bink-Moenen 1983: 109. 

Aleurocanthus angolensis Cohic 1966a: 21-23. Holotype. Angola: Portugalia, v.1964, R. Paulian), on 

unidentified host. 

Distribution: 4: Algeria; Chad. 

Host: Combretaceae: Combretum cinereipetalum, Combretum bracteatum, Combretum poggei. 

Connaraceae: Cnestis lescrauwaetii; Euphorbiaceae: Alchornea cordifolia, Bridelia ferruginea, 

Hymenocardia ulmoides, Bridelia micrantha, Sapium cornutum; Fabaceae: Dalbergia sp., Terminalia 

laxiflora, Millettia laurentii; Flacourtiaceae: Caloncoba dusenii; Guttiferae: Harungana 

madagascariensis; Rubiaceae: Gardenia jovis-tonantis, Plectronia arnoldianum; Verbenaceae: 

Clerodendrum polycephalum. 

Cohicaleyrodes blanzyi (Cohic) 1968 

Siphoninus blanzyi Cohic 1968b: 127. Syntypes. Congo: on Bertiera sp. 

Cohicaleyrodes blanzyi (Cohic); Bink-Moenen 1983: 111. 


Host: Rubiaceae: Bertiera sp. 

Cohicaleyrodes crossopterygis Bink-Moenen 1983 

Cohicaleyrodes crossopterygis Bink-Moenen 1983: 107. Holotype pupal case, Chad, Boro, 6.xii.1973, FAB & 

RMBM, on Crossopteryx febrifuga; BMNH. 

Distribution: 4: Africa, Chad. 

Host: Combretaceae: Combretum molle; Rubiaceae: Crossopteryx febrifuga. 

Cohicaleyrodes descarpentriesi (Cohic) 1968 

Aleurocanthus descarpentriesi Cohic 1968b: 81-84. Syntypes. Congo: Brazzaville, 21.vi.1965 and 

11.xii.1965, on Alchornea cordifolia. 

Cohicaleyrodes descarpentriesi (Cohic); Bink-Moenen 1983: 111 


Host: Bignoniaceae: Stereospermum kunthianum, Combretaceae: Combretum dolichopetalum, 

Combretum molle; Euphorbiaceae: Alchornea cordifolia, Verbenaceae: Clerodendrum polycephalum. 

Cohicaleyrodes elongatus Meganathan & David 1994 

Mixaleyrodes elongatus Meganathan & David 1994: 45. Holotype. India: on Begonia subpeltata. 

Cohicaleyrodes elongatus (Meganathan and David): David 2005: page? 

263


Host: Begoniaceae: Begonia subpeltata. 

Cohicaleyrodes indicus (David and Selvakumaran) 1987 

Mixaleyrodes indicus David & Selvakumaran 1987: 654. Holotype pupal case. India: Kerala State, Idukki, 

13.vi.1986, S. Selvakumaran, on Litsea travancorica, IDAV. 


Host: Lauraceae: Litsea travancorica. 

Cohicaleyrodes jesudasani Jesudasan and David 2005 

Cohicaleyrodes indicus Jesudasan and David 1991: 298. Holotype pupal case. India: on Sterculia alata, 

IDAV. 

Cohicaleyrodes jesudasani David 2005: 67; new name, preoccupied by C. indicus (David and Selvakumaran) 

1987. 


Host: Sterculiaceae: Sterculia alata. 

Cohicaleyrodes mappiae Selvakumaran & David 1996 

Cohicaleyrodes mappiae Selvakumaran & David 1996: 61. Holotype female. India. 


Host: 

Cohicaleyrodes obscura Bink-Moenen 1983 

Cohicaleyrodes obscura Bink-Moenen 1983: 111. Holotype female, Chad, Timberi, 29.xii.1973, FAB & 

RMBM, on Tarenna sp; BMNH. 


Host: Rubiaceae: Tarenna sp. 

Cohicaleyrodes padminiae Phillips & Jesudasan 2006 

Cohicaleyrodes padminiae Phillips & Jesudasan 2006, in David et al. 2006: 21. Holotype female. India. 


Host: 

Cohicaleyrodes pauliani (Cohic) 1968 

Aleurocanthus pauliani Cohic 1968b: 84-87. Syntypes. Congo: Brazzaville, Centre ORSTOM, 27.ii.1967, on 

264

Strychnos variabilis. 

Cohicaleyrodes pauliani (Cohic); Bink-Moenen 1983: 113. 


Host: Loganiaceae: Strychnos variabilis. 

Cohicaleyrodes platysepali (Cohic) 1966 

Aleurocanthus platysepali Cohic 1966a: 23-26. Syntypes. Congo: Brazzaville, 17.viii.1964, on Aframomum 

stipulatum; on Platysepalum vanderystii. 

Cohicaleyrodes platysepali (Cohic); Bink-Moenen 1983: 113. 


Host: Fabaceae: Dalbergia kisantuensis, Dalbergia lactea, Platysepalum vanderijstii; Zingiberaceae: 

Aframomum melegueta, Aframomum stipulatum. 

Cohicaleyrodes quadrilongispinae Bink-Moenen 1983 

Cohicaleyrodes quadrilongispinae Bink-Moenen 1983: 113. Holotype pupal case, Chad, Doba, 21.vi.1973, 

FAB & RMBM; on Paullinia pinnata; BMNH. 


Host: Annonaceae: Annona senegalensis, Hexalobus monopetalus; Bignoniaceae: Stereospermum 

kunthianum; Combretaceae: Anogeissus leiocarpus, Combretum molle; Euphorbiaceae: Hymenocardia 

acida; Fabaceae: Adenodolichos paniculatus; Loganiaceae: Strychnos spinosa; Melastomataceae: 

Dissotis sp.; Moraceae: Ficus sp.; Papilionaceae; Rubiaceae: Nauclea latifolia; Sapindaceae: Paullinia 

pinnata; Verbenaceae: Clerodendrum violaceum; Vitaceae: Cissus sp. 

Cohicaleyrodes recurvispina (Cohic) 1966 

Aleurocanthus recurvispinus Cohic 1966a: 17-19. Syntypes. Congo: Brazzaville, Centre ORSTOM, 

21.iii.1964, on Tetracera alnifolia. 

Cohicaleyrodes recurvispina (Cohic); Bink-Moenen 1983: 115. 

Distribution: 4: Chad; Congo; Ivory Coast; Nigeria. 

Host: Annonaceae: Hexalobus monopetalus, Uvaria brazzavillensis, Uvaria scabrida; Dilleniaceae: Tetracera 

alnifolia; Moraceae: Antiaris africana, Ficus capensis; Rubiaceae: Morinda morindoides, Morinda 

confusa. 

Cohicaleyrodes saklespurensis (Regu & David) 1992 

Aleurotrachelus saklespurensis Regu & David 1992b: 135. Holotype pupal case, India, Saklespur, 10.iii.1990. 

K. Regu, on Dimocarpus longan, Regu Collection. 

Cohicaleyrodes saklespurensis (Regu & David); David et al. 2006: 23; Martin & Mound 2007: 27. 

265


Host: Sapindaceae: Dimocarpus longan. 

Cohicaleyrodes uvariae (Cohic) 1968 

Aleurocanthus uvariae Cohic 1968a: 22-24. Syntypes. Congo: Brazzaville, Centre ORSTOM, 9.xii.1965, on 

Uvaria scabrida. 

Cohicaleyrodes uvariae (Cohic); Bink-Moenen 1983: 115. 



266

Combesaleyrodes COHIC 1966 

Combesaleyrodes Cohic 1966b: 35. Type species. Combesaleyrodes bouqueti Cohic 1966, by original 

designation; Bink-Moenen 1983: 116. 



bouqueti x 

tauffiebi x 

Total = 2 species 0 0 0 2 0 0 0 0 0 

Combesaleyrodes bouqueti Cohic 1966 

Combesaleyrodes bouqueti Cohic 1966b: 36. Syntypes. Congo: on Colletocema dewevrei, BMNH; Bink- 

Moenen 1983: 116. 


Host: Rubiaceae: Colletoecema dewevrei. 

Combesaleyrodes tauffiebi Cohic 1966 

Combesaleyrodes tauffiebi Cohic 1966b: 38. Syntypes. Congo: Brazzaville, 12.v.1965, F. Cohic, on Tetracera 

alnifolia, and 25.ii.1965 on Urophyllum hirtellum; BMNH; Bink-Moenen 1983: 116. 


Host: Dilleniaceae: Tetracera alnifolia; Rubiaceae: Urophyllum hirtellum. 

267

Corbettia DOZIER 1934 

Corbettia Dozier 1934: 190. Type species. Corbettia millettiacola Dozier 1934, by monotypy; 





bauhiniae x 

graminis x 

grandis x 

insoberliniae x 

lamottei x 

lonchocarpi x 

millettiacola x 

pauliani x 

tamarindi x 

Total = 9 species 0 0 0 9 0 0 0 0 0 

Corbettia bauhiniae Cohic 1968 

Corbettia bauhiniae Cohic 1968a: 38. Syntypes. Chad: on Piliostigma reticulata and Bauhinia rufescens; 



Host: Fabaceae: Bauhinia rufescens, Piliostigma reticulata, Piliostigma thonninghii. 

Corbettia graminis Mound 1965 

Corbettia graminis Mound 1965c: 144. Holotype. Nigeria: Onitsha, 13.i.1957, V. Eastop, on grass 

[Graminae]; BMNH, USNM; Bink-Moenen 1983: 118. 


Host: Poaceae: a grass. 

268

Corbettia grandis Russell 1960 

Corbettia grandis Russell 1960b: 134. Holotype, Zaire, Eala, 7.ix.1936, J. Ghesquiere, on Millettia sp.; 

MRCB, paratypes in USNM, USNM. 


Host: Fabaceae: Millettia sp., Tephrosia toxicaria. 

Corbettia insoberliniae Bink-Moenen 1983 

Corbettia isoberliniae Bink-Moenen 1983: 119. Holotype pupal case, Chad, Boro, 6.xii.1973, FAB & 

RMBM, on Isoberlinia doka; BMNH. 


Host: Fabaceae: Isoberlinia doka. 

Corbettia lamottei Cohic 1969 

Corbettia lamottei Cohic 1969: 116. Holotype. Ivory Coast: on Piliostigma thonninghii; Bink-Moenen 1983: 

121. 


Host: Fabaceae: Piliostigma reticulatum, Piliostigma thonningii. 

Corbettia lonchocarpi Bink-Moenen 1983 

Corbettia lonchocarpi Bink-Moenen 1983: 121. Holotype pupal case, Chad, Boro, 6.xii.1973, FAB & 

RMBM, on Lonchocarpus laxiflorus; BMNH. 


Host: Fabaceae: Lonchocarpus laxiflorus. 

Corbettia millettiacola Dozier 1934 

Corbettia millettiacola Dozier 1934: 190. Lectotype (designated by Russell 1960b:129). Zaire: Mbanza 

Ngunga, x.1935, J. Ghesquiere on Millettia versicolor; BMNH; Mound 1965c. 

Corbettia baphiae Russell 1960b: 131. Holotype. Zaire: on Baphia sp., USNM; Bink-Moenen 1983: 118. 

Distribution: 4: Ivory Coast; Tanzania; Zaire; synonymy according to Martin & Mound 2007. 

Corbettia indentata Russell 1960b: 131. Holotype. Zaire: on Tephrosia toxicaria, USNM; synonymy 


Distribution: 4: Chad; Congo; Ivory Coast; Nigeria; Zaire. 

Host: Baphia sp., Cassia auriculata, Cassia javanica, Cassia spectabilis, Cassia siamea, Cassia sieberiana, 

Dalbergia kisantuensis, Desmodium lasiocarpus, Lonchocarpus sericeus, Millettia lane-poolei, 

Millettia versicolor, Piliostigma thonningii, Platysepalum vanderystii, Swartzia madagaskariensis, 

269

Tephrosia toxicaria. 

Corbettia pauliani Cohic 1966 

Corbettia pauliani Cohic 1966a: 31. Holotype. Angola: on Leguminosae. 

Distribution: 3: Angola. 

Host: Fabaceae. 

Corbettia tamarindi Takahashi 1951 

Corbettia tamarindi Takahashi 1951a: 365. Syntypes. Madagascar: on Tamarindus sp., MNHNP 

Distribution: 4: Chad; Madagascar. 

Host: Fabaceae: Tamarindus indicus. 

270

Crenidorsum RUSSELL 1945 

Crenidorsum Russell 1945: 55. Type species. Crenidormus tuberculatum Russell 1945, by original 

designation. 

Mohanasundaramiella David, P.M. 2000: 123. Type species - Mohanasundaramiella rubiae David, 2000, by 

monotypy and original designation; synonymy according to Martin & Mound 2007: 27. 

Reference: Russell 1945. 



armatae x 

aroidephagus x x x x x x 

celebes x 

commune x x 

debordae x 

diaphanum x 

differens x 

lasangensis x 

leve x 

magnisetae x 

malpighiae x 

marginale 

micheliae x 

millennium x 

morobensis x 

ornatum x 

271


rubiae x 

russellae x 

stigmaphylli x 

tuberculatum x 

turpiniae x 

Total = 21 species 2 12 1 0 2 3 4 1 1 

Crenidorsum armatae Russell 1945 

Crenidorsum armatae Russell 1945: 60. Holotype. Cuba: on Coccoloba armata, USNM; Vazquez 1999: 146. 


Host: Malpighiaceae: Malpighia glabra; Polygonaceae: Coccoloba armata. 

Crenidorsum aroidephagus Martin and Aguiar 2001 

Crenidorsum aroidephagus Martin et al. 2001: 1. Holotype puparium, Madeira; Martin 2005b: 7. 

Distribution: 1: USA (FL; HI); 2: Barbados, Belize, Brazil; Colombia, Costa Rica, Cuba, Dominica, 

Dominican Republic, Guatemala, Mexico, Panama, Puerto Rico, Trinidad, Venezuela; 3: Madeira 

Island, glass in France and Germany; 6: India, Philippines; 8: Fiji 9: Hawaii. 

Host: Araceae: Anthurium andraeanum, Anthurium affine, Anthurium hookeri, Anthurium sp., 

Monstera deliciosa, Monstera sp., Syngonium podophyllum, Syngonium sp. 

Natural enemies: Aphelinidae: Cales noacki, Encarsia hispida. 

US Port Interceptions: Australia on Philodendron sp.; Costa Rica on Monstera deliciosa. 

Crenidorsum celebes Martin 1988 

Crenidorsum celebes Martin 1988: 71. Holotype pupal case, Indonesia, Sulawesi, Utara, 1.vi.1985, J. Martin, 

on undetermined host, BMNH. 


Host: Cunoniaceae: Weinmannia sp.; Proteaceae; Sapotaceae? 

Crenidorsum commune Russell 1945 

Crenidorsum commune Russell 1945: 60. Holotype. Haiti: on Coccoloba laurifolia, USNM. 

Distribution: 1: USA (FL); 2: Bahamas; Haiti; Nassau. 

Host: Arecaceae: Chamaedorea elegans; Polygonaceae: Coccoloba diversifolia, Coccoloba laurifolia, 

272

273 

Coccoloba krugii Lindau, Coccoloba northropiae, Coccoloba uvifera, Coccoloba sp.; Rhamnaceae: 

Krugiodendron ferreum. 

US Port Interceptions: Bahamas on Coccoloba uvifera. 

Crenidorsum debordae Russell 1945 

Crenidorsum debordae Russell 1945: 64. Holotype. Haiti: on Coccoloba rotundifolia, USNM 

Distribution: 2: Haiti. 

Host: Polygonaceae: Coccoloba rotundifolia. 

Crenidorsum diaphanum Russell 1945 

Crenidorsum diaphanum Russell 1945: 65. Holotype. Haiti: on Coccoloba rotundifolia, USNM. 

Distribution: 2: Haiti. 

Host: Polygonaceae: Coccoloba rotundifolia. 

Crenidorsum differens Russell 1945 

Crenidorsum differens Russell 1945: 63. Holotype. Leeward Islands: Montserrat, on Coccoloba grandiflora, 

USNM. 

Distribution: 2: Leeward Islands. 

Host: Polygonaceae: Coccoloba grandifolia. 

Crenidorsum lasangensis Martin 1985 

Crenidorsum lasangensis Martin 1985: 323. Holotype pupal case. New Guinea: on Musa sp., BMNH. 


Host: Musaceae: Musa sp. 

Crenidorsum leve Russell 1945 

Crenidorsum leve Russell 1945: 62. Holotype. Puerto Rico: on Coccoloba obtusifolia, USNM. 

Distribution: 2: Puerto Rico; Virgin Islands. 

Host: Polygonaceae: Coccoloba krugii Lindau, Coccoloba obtusifolia. 

Crenidorsum magnisetae Russell 1945 

Crenidorsum magnisetae Russell 1945: 63. Holotype. Haiti: on Coccoloba diversifolia, USNM. 

Distribution: 2: Haiti; Jamaica. 

Host: Polygonaceae: Coccoloba diversifolia, Coccoloba revoluta, Coccoloba retusa.

Crenidorsum malpighiae Russell 1945 

Crenidorsum malpighiae Russell 1945: 60. Holotype. Cuba: on Malpighia glabra, USNM. 


Host: Malpighiaceae: Malpighia glabra. 

Crenidorsum marginale Russell 1945 

Crenidorsum marginale Russell 1945: 63. Holotype. Dominican Republic: on Coccoloba pubescens, USNM. 

Distribution: 2: Dominican Republic. 

Host: Polygonaceae: Coccoloba pubescens. 

Crenidorsum micheliae (Takahashi) 1932 

Aleurotrachelus micheliae Takahashi 1932. Syntypes. Taiwan: on Michelia sp., TARI. 

Crenidorsum micheliae (Takahashi); Martin et al. 2001: 1 


Host: Begoniaceae: Begonia taiwanensis; Magnoliaceae: Michelia sp. 

Natural enemies: Aphelinidae: Encarsia bothrocera, Encarsia synaptocera, Encarsia duorunga, Encarsia 

perflava (ref: Chou et al. 1996b: 196). 

Crenidorsum millennium Martin 1999 

Crenidorsum millennium Martin 1999: 64. Holotype puparium. Australia: on Xanthostemom eucalyptoides, 

BMNH. 


Host: Myrtaceae: Xanthostemon eucalyptoides. 

Crenidorsum morobensis Martin 1985 

Crenidorsum morobensis Martin 1985: 324. Holotype pupal case. New Guinea: on Myrtella sp., BMNH. 


Host: Pterigynandraceae: Myurella sp. 

Crenidorsum ornatum Russell 1945 

Crenidorsum ornatum Russell 1945: 62. Holotype. Jamaica: on Coccoloba longifolia, USNM. 


Host: Polygonaceae: Coccoloba laurifolia. 

274

Crenidorsum rubiae (P. M. David) 2000 

Mohanasundaramiella rubiae P.M. David 2000: 123. Holotype puparium, India, Tamil Nadu, Kerala, 

Walayar, 24.ix.1992, M. Mohanasundaram, on Morinda sp.; IARI. 

Crenidorsum rubiae (P. M. David); Martin & Mound 2007. 


Host: Rubiaceae: Morinda sp. 

Crenidorsum russellae David & David 2000 

Crenidorsum russellae David & David 2000: 155. Holotype puparium, India, Tamil Nadu, Madurai, 26.i.1994, 

B. David, on Randia malabarica, IARI (paratypes in BMNH). 


Host: Rubiaceae: Randia malabarica. 

Crenidorsum stigmaphylli Russell 1945 

Crenidorsum stigmaphylli Russell 1945: 64. Holotype. Puerto Rico: on Stigmaphyllon sp., USNM. 

Distribution: 2: Cuba; Puerto Rico. 

Host: Malpighiaceae: Stigmaphyllon sagraeanum. 

US Port Interception: Puerto Rico on unknown plant. 

Crenidorsum tuberculatum Russell 1945 

Crenidorsum tuberculatum Russell 1945: 57. Holotype. Puerto Rico: on Coccoloba obtusifolia, USNM. 


Host: Polygonaceae: Coccoloba obtusifolia. 

Crenidorsum turpiniae (Takahashi) 1932 

Aleurotrachelus turpiniae Takahashi 1932. Syntypes. Taiwan: on Turpinia formosana, TARI. 

Crenidorsum turpiniae (Takahashi); Martin et al. 2001: 1. 


Host: Ebenaceae: Diospyros kaki; Staphyleaceae: Turpinia formosana. 

Crenidorsum sp. 

Natural enemies: Parasitoids - Aphelinidae: Encarsia citrella, Encarsia formosa, Encarsia luteola, Encarsia 

nigricephala, Encarsia protransvena, Encarsia pseudocitrella, Eretmocerus sp.; Signiphoridae: 

Signiphora aleyrodis [a hyperparasitoid]. Predators - Coccinellidae: Delphastus pallidus. 

275

Crescentaleyrodes David & Jesudasan 1987 

Crescentaleyrodes David & Jesudasan 1987: 42. Type species. Tetraleurodes semilunaris Corbett 1926. 



fumipennis x 

monodi x 

paulianae x 

semilunaris x 

veiveriae x 

Total = 5 species 0 1 0 2 0 2 0 0 0 

Distribution: WW (2); Oriental (2). 

Host: Poaceae (grasses). 

Diagnosis: Puparium with crescent-shaped pores on the submargin, pest of lemon grass (Cymbopogon 

flexuous). Re-description of genus (Dubey & Ko 2006). 

Cresentaleyrodes fumipennis (Hempel) 1899 

Aleurodes fumipennis Hempel 1899: 394. Syntypes. Brazil: on grass growing in swampy ground. 

Aleurocanthus fumipennis (Hempel); Baker & Moles 1923: 628; Bondar 1923a. 

Aleurotrachelus fumipennis (Hempel); Mound and Halsey 1978: 70. 

Cresentaleyrodes fumipennis (Hempel); Martin & Mound 2007 

Distribution: 2: Brazil, Colombia, Mexico, Venezuela. 

Host: Poaceae: Andropogon bicorne, unidentified grass. 

Cresentaleyrodes monodi (Cohic) 1969 

Aleurolobus monodi Cohic 1969: 46. Holotype. Chad: on Cymbopogon sp. 

Crescentaleyrodes monodi (Cohic). 


Host: Poaceae: Cymbopogon schoenanthus. 

276

Crescentaleyrodes paulianae (Cohic) 1969 

Aleurolobus paulianae Cohic 1969: 51. Holotype. Ivory Coast: on Sorghastrum sp. 

Crescentaleyrodes paulianae (Cohic). 


Host: Combretaceae: Combretum glutinosum, Combretum hypopilinum; Poaceae: Andropogon diplandra, 

Beckeropsis uniseta, Hyparrhenia rufa, Sorghastrum sp. 

Crescentaleyrodes semilunaris (Corbett) 1926 

Tetraleurodes semilunaris Corbett 1926: 282. Syntypes. Sri Lanka: on citronella grass [Cymbopogon sp.], 

BMNH. 

Aleurolobus semilunaris (Corbett); Bink-Moenen 1983 

Crescentaleyrodes semilunaris (Corbett); David & Jesudasan Alexander 1987: 42; Jesudasan and David 1991: 

299; Dubey & Ko 2006: 71. 

Distribution: 5: China; 6: India, Sri Lanka. 

Host: Poaceae: Cymbopogon sp. 

Crescentaleyrodes vetiveriae Dubey & Ko 2006 

Crescentaleyrodes vetiveriae Dubey & Ko 2006: 71. Holotype puparium, India, Tamil Nadu, 25.vi.2005; 

deposited in ZSI, Kolkata, India. 


Host: Poaceae: Vetiveria zizanioides. 

277

Crytolingula MARTIN & CARVER 1999 

Cryptolingula MARTIN & CARVER 1999, Martin1999: 65. Type species. Cryptolingula perplexa Martin & 

Carver 1999, by monotypy. 



perplexa x 

Total = 1 species 0 0 0 0 0 0 1 0 0 

Crytolingula perplexa Martin & Carver 1999 

Cryptolingula perplexa Martin & Carver 1999, in Martin 1999: 65. Holotype puparium. Australia: on 

unknown host, ANIC. 



278

Davidiella Dubey & Sundararaj 2005 

Davidiella Dubey & Sundararaj 2005: 351. Type species - Davidiella cinnamomi Dubey & Sundararaj 2005, 

by monotypy and original description.. 

Diagnosis: The genus resembles Cockerelliella Sundararaj & David 1992, but differs by the absence of the 

thoracic and caudal tracheal pores and furrows (Dubey & Sundararaj 2005e). 



cinnamomi x 

Total = 1 species 0 0 0 0 0 1 0 0 0 

Davidiella cinnamomi Dubey & Sundararaj 2005 

Davidiella cinnamomi Dubey & Sundararaj 2005e: 352. Holotype puparium, India, Karnataka, Kudremukh 

National Park, 10.ix.2001, A. K. Dubey, on Cinnamomum sulphuratum, FRIDD. 


Host: Lauraceae: Cinnamomum sulphuratum. 

279

DIALEURODES Cockerell 1902 

Aleyrodes (Dialeurodes) Cockerell 1902a: 283. Type species. Aleyrodes citri Riley & Howard 1893, by 

original designation, a synomym of A. citri Ashmead 1885. 

Dialeurodes Cockerell; full genus, Quaintance & Baker 1914: 97. 

Kanakarajiella David & Sundararaj 1993. Type species. Dialeurodes vulgaris Singh 1931, by original 

designation; synonymy according to Martin & Mound 2007: 28. 

Lankaleurodes David 1993: 23. Type species. Dialeurodes radiipuncta Quaintance & Baker 1917, by original 

designation; synonymy according to Martin & Mound 2007: 28. 

Shanthiniae David 2000, in P.M.M. David 2000: 125. Type species - Shanthiniae sheryli David 2000, by 

monotypy and original designation; synonymy according to Martin & Mound 2007: 28. 

Comment: Martin & Mound (2007) tentatively listed species in the following subgenera of Dialeurodes as 

being in the genus Dialeurodes: Dialeurodes (Dialeuronomada) Quaintance & Baker 1917: 51; 

Dialeurodes (Rabdostigma) Quaintance & Baker 1917: 426, Dialeurodes (Gigaleurodes) Quaintance 

& Baker 1917: 426, and Dialeurodes (Dialeuroplata) Quaintance & Baker 1917: 435 

Dialeuronomada Quaintance & Baker; full genus by Sundararaj & David 1991. 

References: Sundararaj & David 1991; Dubey & Sundararaj 2004 - Key to Indian species. 

Comment: Currently, this genus is only known to occur in India. Martin & Mound (2007) placed the species 

in this genus in the genus Dialeurodes. 



abbotabadiensis x 

adinobotris x 

agalmae x 

angulata 

anjumi x 

ara x 

armatus x 

bancoensis x 

bangokana x 

bicornicauda x 

brideliae x 

bladhiae x x 

280


buscki x 

celti x 

cephalidistinctus x 

cerifera x 

chiengsenana x 

cinnamomi x 

cinnamomicola x 

citri x x x x x x x x 

citricola x 

conocephali x 

crescentata x 

cyathispinifera x 

daphniphylli x 

davidi x 

decaspermi x 

delhiensis x 

denticulatus x 

dicksoni x 

didymocarpi x 

distincta x 

doveri x 

drypetesi x 

dubia x 

dumbeaensis x 

egregissima x 

emarginata x 

endospermi x 

evodiae x 

ficicola x 

gardeniae x 

281


gemurohensis x 

glutae x 

greenwoodi x 

heterocera x 

hexpuncta x 

hongkongensis x 

icfreae x 

imperialis x 

indicus x 

joholensis x 

kepongensis x 

kirkaldyi x x x x x x x x x 

kumargiriensis x 

lithocarpi x 

loranthi x 

machilicola x 

maculata x 

maculipennis x 

malabaricus x 

martini x 

maxima x 

mirabilis x 

musae x 

natickis x 

navarroi x 

nigeriae x 

octoplicata x 

ouchii x 

oweni x 

panacis x 

282


papulae x 

pauliani x 

philippinensis x x 

pilahensis x 

platicus x 

pseudocitri x 

psychotriae x 

punctata x 

radiipuncta x 

rangooni x 

razalyi x 

rempangensis x 

rengas x 

reticulosa x 

rotunda x 

saklaspurensis x 

sandorici x 

sepangensis x 

sheryli x 

shintenensis x 

shoreae x 

siemriepensis x 

simmondsi x 

striata x 

struthanthi x 

sundararajani x 

tanakai x 

townsendi x 

tricolor x x 

tuberculosa x 

283


turpiniae x x 

vitis x 

vulgaris x x 

wendlandiae x 

Total = 102 species 3 15 3 8 16 65 5 5 3 

Dialeurodes abbotabadiensis Qureshi 1980 

Dialeurodes abbotabadiensis Qureshi 1980: 62. Holotype puparium, Pakistan, Abbotabad, 28.vi.1973, J.I. 

Qureshi, ex. undetermined plant, FAI. 

Dialeurodes abbotabadiensis: Dubey and Sundararaj, 2006. 62-67. 

Distribution: 6: Pakistan, India. 


Dialeurodes adinobotris Corbett 1935 

Dialeurodes adinobotris Corbett 1935b: 766. Syntypes. Malaya: on Adinobotrys atropurpureus. 



Dialeurodes agalmae Takahashi 1935 

Dialeurodes agalmae Takahashi 1935b: 44. Syntypes. Taiwan: on Agalma [=Schefflera] taiwanianum, TARI. 


Host: Araliaceae: Schefflera octophylla, Schefflera taiwanianum. 

Natural enemies: Aphelinidae: Encarsia cibcensis. 

Dialeurodes angulata Corbett 1935 

Dialeurodes angulata Corbett 1935b: 773. Holotype. Malaya: on unidentified host. 



Dialeurodes ara Corbett 1935 

Dialeurodes ara Corbett 1935b: 755. Syntypes. Malaya: on Ficus sp. 


284


Dialeurodes armatus David & Subramaniam 1976 

Dialeurodes armatus David & Subramaniam 1976: 184. Holotype. India: on Azadirachta indica, ZSI. 


Host: Meliaceae: Azadirachta indica. 

Dialeurodes bancoensis Ardaillon & Cohic 1970 

Dialeurodes bancoensis Ardaillon & Cohic 1970: 277. Holotype. Ivory Coast: on Carpolobia lutea. 


Host: Polygalaceae: Carpolobia lutea. 

Dialeurodes bangokana Takahashi 1942 

Dialeurodes bankokana Takahashi 1942h: 327. Syntypes. Thailand: on unidentified host. 

Distribution: 6: Malaya; Malaysia; Thailand. 


Dialeurodes bicornicauda Martin 1999 

Dialeurodes bicornicauda Martin 1999: 67. Holotype puparium. Australia: on Dysoxylum parasiticum, ANIC. 


Host: Elaeocarpaceae: Dysoxylum parasiticum. 

Dialeurodes brideliae Jesedasan and David 

Dialeurodes brideliae Jesudasan and David 1991: 301-302. 


Host: Euphorbiaceae: Bridelia crenulata. 

Dialeurodes buscki Quaintance & Baker 1917 

Dialeurodes buscki Quaintance & Baker 1917: 428. Syntypes. Puerto Rico: Bayamon, 15.i.1899, A. Busck, on 

''climbing vine", USNM. 


Host: ''climbing vine". 

Natural enemies: Aphelinidae: Encarsia brunnea, Encarsia portoricensis. 

285

Dialeurodes celti Takahashi 1942 

Dialeurodes celti Takahashi 1942g: 307. Syntypes (2). Thailand: on Celtis sp., TARI. 


Host: Ulmaceae: Celtis sp. 

Dialeurodes cephalidistinctus Singh 1932 

Dialeurodes cephalidistinctus Singh 1932: 83. Syntypes. Burma: on Eugenia jambos, ZSI. 

Distribution: 6: Burma; Cambodia. 


Dialeurodes cerifera Quaintance & Baker 1917 

Dialeurodes cerifera Quaintance & Baker 1917: 427. Syntypes. South Africa: on Celastrus buxifolius, 

USNM. 

Distribution: 4: South Africa. 

Host: Celastraceae: Celastrus buxifolius. 

Dialeurodes chiengsenana Takahashi 1942 

Dialeurodes chiengsenana Takahashi 1942e: 276. Syntypes. Thailand: on unidentified host, TARI. 



Dialeurodes cinnamomi Takahashi 1932 

Dialeurodes cinnamomi Takahashi 1932: 13. Syntypes. Taiwan: on Cinnamomum japonicum, TARI. 


Host: Lauraceae: Cinnamomum sp., Cinnamomum japonicum. 

Dialeurodes cinnamomicola Takahashi 1937 

Dialeurodes cinnamomicola Takahashi 1937b: 23. Holotype. China: on Cinnamomum sp. 



Dialeurodes citri (Ashmead) 1885 

Aleyrodes citri Ashmead 1885: 704. Syntypes. USA: Florida, Jacksonville, on Citrus sp., USNM. 

286

Dialeurodes citri (Ashmead); Quaintance & Baker 1916: 469. 

Aleurodes citri Riley & Howard 1893: 219. Syntypes. USA: Florida, Louisanna, on Citrus sp. 

Aleyrodes (Dialeurodes) citri Riley & Howard; Cockerell 1902a: 283. 

Aleurodes citrifolii Foster 1892; Kirkaldy 1907: 49 (nomen nudum). 

Aleurodes eugeniae var. aurantii Maskell 1895: 431. Syntypes. India: on Citrus aurantium, ADSIR; 


Aleyrodes (Dialeurodes) aurantii (Maskell); Cockerell 1902a: 283 

Dialeurodes citri var. hederae Takahashi 1936b: 219. Syntypes. Taiwan; on Hedera formosana, TARI. 

Dialeurodes citri var. kinyana Takahashi 1935b: 43. Holotype. Taiwan: on unidentified plant. 

Aleyrodes kushinasii Sasaki 1908: 55. Syntypes. Japan: on Gardenia sp., TARI; Takahashi 1951b: 19. 

Dialeurodes tuberculatus Takahashi 1932: 9. Syntypes. Taiwan: on Meliosma rhoifolia, TARI; Takahashi 

1958: 66. 

Distribution: 1: USA (AL, AR, AZ, CA, DC, FL, GA, HI, IL, LA, MO, MS, NC, OH, SC, TX, VA); 

2: Bahamas, Bermuda, Cuba, Dominican Republic, El Salvador, Haiti, Honduras, Mexico, Panama, 

Puerto Rico; 3: Afganistan, France, Greece, Iran, Italy, Portugal, Sicily, Turkey; 5: China, Hong Kong, 

Korea; Japan, Macau, Taiwan; 6: India, Pakistan, Philippines, Thailand; 8: Guam; 9: Hawaii 

Host: Anacardiaceae: Cotinus coggyria; Apocynaceae: Allamanda neriifolia, Nerium oleander; 

Aquifoliaceae: Ilex sp., Ilex x attenuata; Araliaceae: Aralia sp., Fatsia sp., Hedera formosana, Hedera 

helix, Hedera helix, Schefflera arboricola; Arecaceae: Sabal megacarpa; Bignoniaceae: Tecoma 

radicans; Boraginaceae: Ehretia sp.; Caprifoliaceae: Lonicera japonica, Viburnum odoratissimum, 

Viburnum sp., Viburnum suspensum, Viburnum tinus; Ebenaceae: Diospyros khaki, Diospyros sp., 

Diospyros virginiana; Ericaceae: Rhododendron sp.; Euphorbiaceae: Bischofia javanica, Glochidion 

hongkongense, Glochidion sp., Ricinus communis; Fabaceae: Glycine max; Fagaceae: Lithocarpus sp, 

Quercus aquatica, Quercus nigra; Hamamelidaceae: Distylium racemosus; Lauraceae: Machilus sp, 

Persea sp.; Liliaceae: Smilax sp.; Lythraceae: Lagerstroemia indica; Magnoliaceae: Magnolia coco, 

Magnolia fuscata, Magnolia sp., Magnolia x soulangiana; Malpighiaceae: Hiptage madablota; 

Hiptage benghalensis; Meliaceae: Melia azedarach; Moraceae: Ficus altissima, Ficus macrophylla, 

Ficus nitida, Maclura aurantiaca; Myrsinaceae: Ardisia humilis, Ardisia sieboldi; Myrtaceae: Eugenia 

jambos, Eugenia sp., Myrtus communis, Myrtus lagerstroemia, Psidium guajava; Nyssaceae: Nyssa 

ogeche; Oleaceae: Chionanthus retusus, Chionanthus virginicus, Forsythia sp., Fraxinus lanceolata, 

Jasminum arborescens, Jasminum frutescens, Jasminum nitidum, Jasminum odoratissimum, Jasminum 

sambac, Ligustrum amurense, Ligustrum amurense, Ligustrum ibota, Ligustrum japonicum, Ligustrum 

lucidum, Ligustrum ovalifolium, Ligustrum sinense, Ligustrum sp., Ligustrum vulgare, Osmanthus 

americanus, Osmanthus fragrans, Osmanthus heterophyllus, Syringa sp., Syringa vulgaris; 

Proteaceae: Helicia sp.; Punicaceae: Punica granatum; Rosaceae: Cerasus sp. Crataegus laevigata, 

Laurocerasus caroliniana, Malus sp., Prunus caroliniana, Prunus caroliniana, Prunus laurocerasus, 

Prunus laurocerasus, Pyracantha coccinea, Pyracantha koidzumii, Pyrus sp., Rubus sp., Rubiaceae: 

Cephalanthus occidentalis, Rubiaceae: Cephalanthus sp., Coffea arabica, Gardenia augusta, 

Gardenia florida, Gardenia jasminoides, Gardenia radicans, Gardenia sp., Hedera helix, Jasminum 

fruticans, Wendlandia glabrata, Rutaceae: Choisya ternata, Citrus aurantifolia, Citrus aurantium, 

Citrus aurantium var formosanus, Citrus bigaradia, Citrus medica, Citrus ponki, Citrus sinensis, 

287

288 

Citrus sp., Citrus x nobilis, Citrus x paradisi, Citrus x tangelo, Fortunella japonica, Poncirus 

trifoliata, Severinia buxifolia, Citrofortunella microcarpa, Zanthoxylum clava-herculis; Sabiaceae: 

Meliosma rigida; Sapindaceae: Sapindus sp. Simaroubaceae: Ailanthus glandulosa; Staphyleaceae: 

Turpinia formosana; Theaceae: Camellia japonica; Theaceae: Camellia sinensis var. sinensis; 

Vitaceae: Ampelopsis tricuspidata. 

Natural enemies: Fungus - Aegerita webberi, Aschersonia aleyrodis, Aschersonia goldiana, Aschersonia 

placenta, Aschersonia tamurai, Beauveria bassiana, Fusarium verticilloides, Paecilomyces 

cinnamomeus, Sporotrichum sp., Verticillum lecanii. 

Parasitoids - HYMENOPTERA/Aphelinidae: Encarsia abundantia, Encarsia armata, Encarsia citrella, 

ncarsia citri, Encarsia citrofila, Encarsia formosa, Encarsia lahorensis, Encarsia longivalvula, 

ncarsia magnivena, Encarsia protransvena, Encarsia sophia, Encarsia sp., Encarsia strenua, 

ncarsia strenua group, Encarsia tabacivora, Encarsia tricolor, Encarsia variegata, Eretmocerus sp., 

Eulophidae: Euderomphale sp. 

Predators - ACARI/Phytoseiidae: Amblyseius eharai, Amblyseius largoensis, Euseius stipulatus, 

Typhlodromus sp.; COLEOPTERA/Coccinellidae: Adonia variegata, Brumoides suturalis, Catana 

chapini, Chilocorus bipustulatus, Chilocorus stigma, Clitostethus arcuatus, Cryptognatha flaviceps, 

Cycloneda sanguinea, Delphastus catalinae, Delphastus pallidus, Delphastus pusillus, Delphastus sp., 

Didion punctatum, Diomus roseicollis, Exochomus quadripustulatus, Menochilus sexmaculatus, 

Nephaspis oculatus, Olla v-nigrum, Scymnus sp., Serangium flavescens, Serangium parcesetosum, 

Stethorus gilvifrons, Verania cardoni; DIPTERA/Dolichopodidae: Condylostylus patibulatus; 

HEMIPTERA/ Lygeidae: Geocoris sp., Campylomma diversicornis, Deraeocoris pallens; 

NEUROPTERA/Chrysopidae: Chrysopa sp., Chrysopa vulgaris, Chrysoperla carnea; 

Coniopterygidae: Conwentzia hagani, Conwentzia psociformis, Semidalis aleyrodiformis; 

Hemerobiidae: Genus? sp.; THYSANOPTERA/ Phlaeothripidae: Aleurodothrips fasciapennis. 

US Port Interceptions: Albania on Citrus sp., Diospyros sp.; Argentina on Gardenia jasminoides; 

Bangladesh on Rutaceae; Bolivia on Citrus sp.; Brazil on Jasminum sp.; Canada on Jasminum sp.; 

China on Desmos chinensis, Gardenia jasminoides; Cuba on Citrus sp.; France on Citrus sinensis; 

Greece on Citrus reticulata, Citrus sinensis; Guinea on Anubias congensis; Haiti on unknown plant; 

Hawaii on unknown plant; India on Citrus sp.; Italy on Citrus limon, Citrus reticulata; Jamaica on 

Aralia sp.; Japan on Citrus reticulata, Citrus sp., Diospyros sp.; Korea (South) on unknown plant; 

Lebanon on Citrus sinensis; Mexico on Citrus sp., Gardenia jasminoides, Gardenia sp.; Nicaragua 

on Citrus sinensis; Philippines on Citrus sp.; Puerto Rico on Citrus sp.; Viet Nam on Citrus sp.; 

Yugoslavia on Citrus reticulata. 

Dialeurodes citricola Young 1942 

Dialeurodes (Gigaleurodes) Young 1942: 96. Syntypes. China: on Citrus tangerina and Citrus sinensis. 

Dialeurodes citricola Young; Mound and Halsey 1978: 135. 


Host: Rutaceae: Citrus reticulata, Citrus sinensis.

Dialeurodes conocephali Corbett 1935 

Dialeurodes conocephali Corbett 1935b: 742. Syntypes. Malaya: on Conocephalus subtrinervius. 


Host: Moraceae: Conocephalus subtrinervius. 

Dialeurodes crescentata Corbett 1935 

Dialeurodes crescentata Corbett 1935b: 778. Syntypes. Malaya: on Cinnamomum sp. 



Dialeurodes cyathispinifera Corbett 1933 

Dialeurodes cyathispinifera Corbett 1933: 121. Syntypes. Malaya: on unidentified plant, BMNH. 



Dialeurodes daphniphylli Takahashi 1932 

Dialeurodes daphniphylli Takahashi 1932: 10. Syntypes. Taiwan: on Daphniphyllum glaucescens, TARI. 


Host: Daphniphyllaceae: Daphniphyllum membranaceum, Daphniphyllum glaucescens 

Dialeurodes davidi Mound & Halsey 1978 

Dialeurodes distinctus David & Subramaniam 1976: 188. Holotype. India: on Elaeodendron glaucum. 

Dialeurodes davidi Mound and Halsey 1978: 136. Replacement name, preoccupied by Dialeurodes distincta 

Corbett 1933. 


Host: Celastraceae: Elaeodendron glaucum. 

Dialeurodes decaspermi Martin 1985 

Dialeurodes decaspermi Martin 1985: 325. Holotype pupal case. New Guinea: on Desacpermum sp., BMNH. 


Host: Myrtaceae: Decaspermum sp. 

Dialeurodes delhiensis David & Sundararaj 1992 

289

290 

Dialeurodes delhiensis David & Sundararaj 1992: 62. Holotype pupal case, India, New Delhi, 24.ii.1986, B. 

David, on Ficus virens, IARI. 


Host: Moraceae: Ficus infectivora, Ficus virens, Ficus sp.. 

Dialeurodes denticulatus (Bondar) 1923 

Aleuroplatus denticulatus Bondar 1923a: 113. Syntypes. Brazil: on Ficus sp, MZU. 

Dialeurodes michoacanensis Sampson & Drews 1941: 165. Syntypes. Mexico: on Ficus sp., UCD; synonymy 


Distribution: 2: Belize, Brazil; Mexico. 

Host: Lauraceae: Persea sp.; Moraceae: Ficus sp. 

Dialeurodes dicksoni Corbett 1935 

Dialeurodes dicksoni Corbett 1935b: 763. Syntypes. Malaya: on Vitex sp. and unidentified plant. 



Dialeurodes didymocarpi Corbett 1935 

Dialeurodes didymocarpi Corbett 1935b: 771. Syntypes. Malaya: on Didymocarpus crinita. 


Host: Gesneriaceae: Didymocarpus crinita. 

Dialeurodes distincta Corbett 1933 

Dialeurodes distincta Corbett 1933: 127. Syntypes. Malaya: on unidentified host, BMNH. 



Dialeurodes dubia Corbett 1935 

Dialeurodes dubia Corbett 1935b: 780. Syntypes. Malaya: on unidentified host. 


Host: unidentified.

Dialeurodes dumbeaensis Dumbleton 1961 

Dialeurodes dumbeaensis Dumbleton 1961a: 122. Holotype. New Caledonia: on ?Homalium sp., CORSTOM. 


Host: Flacourtiaceae: Homalium sp.? 

Dialeurodes egregissima Sampson & Drews 1941 

Dialeurodes egregissima Sampson & Drews 1941: 164. Syntypes. Mexico: on 'higuera" [Ficus involuta], 

UCD. 


Host: Moraceae: Ficus involuta. 

Dialeurodes emarginata (Mound) 1965 

Dialeurolonga emarginata Mound 1965c: 149. Holotype. Nigeria: on Cola cordifolia, BMNH. 

Dialeurodes emarginata (Mound); Bink-Moenen 1983: 128. 


Host: Loganiaceae: Anthocleista vogelii; Sterculiaceae: Cola cordifolia. 

Dialeurodes endospermi Corbett 1935 

Dialeurodes endospermi Corbett 1935b: 745. Syntypes. Malaya: on Endospermum malacense. 


Host: Euphorbiaceae: Endospermum malaccense. 

Dialeurodes evodiae Corbett 1935 

Dialeurodes evodiae Corbett 1935b: 757. Syntypes. Malaya: on Evodia sp. 


Host: Rutaceae: Evodia sp. 

Dialeurodes ficicola Takahashi 1935 

Dialeurodes ficicola Takahashi 1935b: 47. Holotype. Taiwan: on Ficus rigida, TARI. 

Massilieurodes ficicola (Takahashi); Jensen 2001: 292. 

Dialeurodes ficicola Takahashi; Martin & Mound 2007: 36. 


Host: Moraceae: Ficus rigida. 


291

Dialeurodes gardeniae Corbett 1935 

Dialeurodes gardeniae Corbett 1935b: 743. Syntypes. Malaya: on Gardenia florida. 


Host: Rubiaceae: Gardenia florida. 

Dialeurodes gemurohensis Corbett 1935 

Dialeurodes gemurohensis Corbett 1935b: 761. Syntypes. Malaya: on Ficus sp. and Nephelium lappaceum. 


Host: Moraceae: Ficus sp.; Sapindaceae: Nephelium lappaceum. 

Dialeurodes glutae Corbett 1935 

Dialeurodes glutae Corbett 1935b: 759. Syntypes. Malaya: on Gluta sp. 


Host: Anacardiaceae: Gluta sp. 

Dialeurodes greenwoodi Corbett 1936 

Dialeurodes greenwoodi Corbett 1936: 20. Syntypes. Fiji: on Ficus sp., BMNH. 



Dialeurodes heterocera Bondar 1923 

Dialeurodes heterocera Bondar 1923a: 108. Syntypes. Brazil: on Eugenia sp., MZU. 



Dialeurodes hexpuncta Singh 1932 

Dialeurodes hexpuncta Singh 1932: 81. Syntypes. Burma: on undetermined shrub. 


Host: undetermined shrub. 

Dialeurodes hongkongensis Takahashi 1941 

292

293 

Dialeurodes hongkongensis Takahashi 1941b: 351. Syntypes. Hong Kong: on unidentified host, TARI; 

Takahashi 1955b. 

Distribution: 5: China; Hong Kong. 

Host: Liliaceae: Smilax sp. 

Dialeurodes icfreae Sundararaj & Dubey 2003 

Dialeurodes icfreae Sundararaj & Dubey 2003: 294. Holotype puparium, India, Tamil Nadu, Javadu Hills, 

19.iii.2001, A. Dubey, on Santalum album, IDEH. 


Host: Santalaceae: Santalum album. 

Dialeurodes imperialis Bondar 1923 

Dialeurodes imperalis Bondar 1923a: 109. Syntypes (2). Brazil: on unidentified host, MZU. 



Dialeurodes indicus David & Subramaniam 1976 

Dialeurodes indicus David & Subramaniam 1976: 191. Holotype. India: on Eugenia [=Syzygium] 

jambolanum, ZSI. 


Host: Myrtaceae: Eugenia jambolana. 

Dialeurodes joholensis Corbett 1935 

Dialeurodes joholensis Corbett 1935b: 777. Syntypes. Malaya; on unidentified host. 



Dialeurodes kepongensis Corbett 1935 

Dialeurodes kepongensis Corbett 1935b: 756. Syntypes. Malaya; on Eugenia sp. 



Dialeurodes kirkaldyi (Kotinsky) 1907

294 

Aleyrodes kirkaldyi Kotinsky 1907: 95-96. Syntypes. USA: Hawaii, on unidentified trailing shrub, 

Beaumontia grandifolia, Morinda citrifolia and Jasminum grandiflorum, USNM. 

Dialeurodes kirkaldyi (Kotinsky); Quaintance & Baker 1914: 98. 

Dialeurodes yercaudensis Jesudasan and David 1991: 307. Holotype pupal case. India: on Ligustrum walkeri, 

IDAV; synonymy according to Sundararaj & Dubey 2006a. 

Distribution: 1: USA (FL, HI, TX); 2: Bahamas, Barbados, Costa Rica, Cuba; Guyana, Jamaica; Mexico; 

Guyana; Puerto Rico; Trinidad, Virgin Islands; 3: Azores; Egypt; Iran, Israel, Greece, Lebanon; Syria; 

Turkey, UK; 4: Ghana; 5: China; Hong Kong; Japan; Taiwan; 6: Burma; India; Malaysia, Pakistan, 

Philippines, Sri Lanka, Thailand; 7: Australia; 8: Caroline Islands, Cook Islands, Fiji, Guam, Samoa, 

Tahiti; 9: Hawaii. 

Host: Apocynaceae: Allamanda neriifolia, Beaumontia grandiflora, Plumeria acuminata, Plumeria acutifolia, 

Tabernaemontana sp., Trachelospermum jasminoides; Combretaceae: Terminalia sp.; Ebenaceae: 

Diospyros kaki, Ebenaceae: Diospyros sp., Diospyros virginiana; Geraniaceae: Pelargonium sp.; 

Jugandaceae: Juglans regia; Lauraceae: Persea americana; Loganiaceae: Fagraea fragrans; 

Lythraceae: Lagerstroemia indica; Malpighiaceae: Hiptage benghalensis, Hiptage madablota; 

Malvaceae: Malva sylvestris; Oleaceae: Jasminum amplexicaule, Jasminum arabica, Jasminum 

auriculatum, Jasminum biflorum, Jasminum multiflorum, Jasminum nitidum, Jasminum officinale, 

Jasminum sambac, Jasminum volubile, Oleaceae: Jasminum frutiscens, Ligustrum walkeri, Syringa 

sp.?; Rubiaceae: Coffea arabica, Coffea sp., Gardenia tahitiensis, Jasminum sambac, Morinda 

citrifolia, Morinda royoc; Rutaceae: Citrus sinensis, Citrus x paradisi; Verbenaceae: Clerodendrum 

fragrans, Premna integrifolia. 

Natural enemies: Aphelinidae: Encarsia abundantia, Encarsia bothrocera, Encarsia dialeurodis, Encarsia 

lahorensis, Encarsia lutea, Encarsia neoporteri, Encarsia nigricephala, Encarsia perflava, Encarsia 

protransvena, Encarsia sp., Encarsia strenua, Encarsia strenua group, Encarsia tabacivora, 

Eretmocerus portoricensis, Eretmocerus rosei, Eretmocerus sp. 

US Port Interceptions: China on Gardenia jasminoides; Egypt on Murraya paniculata; El Salvador on 

Jasminum sambac; Guinea on Anubias sp.; Hawaii on Jasminum sambac, Moringa sp., Schefflera 

actinophylla, Terminalia catappa; India on Lawsonia sp.; Israel on Jasminum sp.; Jamaica on Aralia 

sp.; Japan on Citrus sp.; Mexico on Choisya sp., Gardenia jasminoides, Gardenia sp., Jasminum sp., 

Persea americana; Philippines on Citrus sp., Gardenia sp.; Puerto Rico on Bougainvillea sp.; 

Tanzania on Jasminum sp.; Thailand on Jasminum sp.; Viet Nam on Jasminum sambac. 

Dialeurodes kumargiriensis Sundararaj & Dubey 2006 

Dialeurodes kumargiriensis Sundararaj & Dubey 2006a. Holotype puparium, India, Kumargiri, Karnataka, 

10.ix.2001, A. K. Dubey, on Strychnos wallichiana, IFRI. 


Host: Loganiaceae: Strychnos wallichiana. 

Dialeurodes lanceolata Takahashi 1949 

Dialeurodes lanceolata Takahashi 1949: 47. Holotype. Riouw (Riau) Islands (south of Singapore): on

undetermined tree, HBL. 



Dialeurodes lithocarpi Takahashi 1931 

Dialeurodes lithocarpi Takahashi 1931c: 218. Syntypes. Taiwan: on Lithocarpus ternaticupula, TARI. 


Host: Fagaceae: Lithocarpus ternaticupula. 

Dialeurodes loranthi Corbett 1926 

Dialeurodes loranthi Corbett 1926: 268. Syntypes. Sri Lanka: on Loranthus sp., BMNH. 



Dialeurodes machilicola Takahashi 1942 

Dialeurodes machilicola Takahashi 1942g: 308. Holotype. Thailand: on Machilus sp. 



Dialeurodes maculata Bondar 1928 

Dialeurodes maculatus Bondar 1928b: 23. Syntypes. Brazil: on Loranthaceae, MZU. 



Dialeurodes maculipennis Bondar 1923 

Dialeurodes maculipennis Bondar 1923a: 104. Syntypes. Brazil: on Ficus sp., MZU. 



Dialeurodes malabaricus Jesudasan and David 

Dialeurodes malabaricus Jesudasan and David 1991: 305. 


Distributon: Lauraceae: Cinnamomum malabaricum. 

295

Dialeurodes martini Jesudasan & David 1991 

Dialeurodes martini Jesudasan and David 1991: 305. Holotype pupal case. India: on Putaranjiva roxburghii, 

IDAV. 


Host: Euphorbiaceae: Putranjiva roxburghii. 

Dialeurodes maxima Quaintance & Baker 1917 

Dialeurodes (Gigaleurodes) maxima Quaintance & Baker 1917: 301. Syntypes. Philippines: on Ficus sp., 

USNM. 

Dialeurodes maxima Quaintance & Baker; Mound & Halsey 1978: 143. 

Distribution: 5: Philippines. 


Dialeurodes mirabilis Takahashi 1942 

Dialeurodes mirabilis Takahashi 1942g: 309. Syntypes. Thailand: on unidentified host. 



Dialeurodes musae Corbett 1935 

Dialeurodes musae Corbett 1935b: 775. Syntypes. Malaya: on Musa sapientum. 



Dialeurodes natickis Baker & Moles 1923 

Dialeurodes natickis Baker & Moles 1923: 641. Syntypes. Chile: on Eugenia luma, USNM. 


Host: Myrtaceae: Eugenia luma. 

Dialeurodes navarroi Bondar 1928 

Dialeurodes navarroi Bondar 1928b: 24. Syntypes. Brazil: on Loranthaceae, MZU. 



Dialeurodes nigeriae Cohic 1966 

296

297 

Dialeurodes (Gigaleurodes) nigeriae Cohic 1966b: 45. Syntype. Nigeria: on unidentified host, BMNH. 

Dialeurodes nigeriae Cohic; Mound and Halsey 1978: 144 


Host: Celastraceae: Salacia nitida, Salacia pyriformis, Salacia senegalensis; Moraceae: Ficus sp. 

Dialeurodes octoplicata Corbett 1935 

Dialeurodes octoplicata Corbett 1935b: 746. Syntypes. Malaya: on Cinnamomum sp. 



Dialeurodes ouchii Takahashi 1937 

Dialeurodes ouchii Takahashi 1937b: 22. Holotype. China: on unidentified host. 



Dialeurodes oweni Singh 1932 

Dialeurodes oweni Singh 1932: 82. Syntypes. Burma: on Eugenia jambos, ZSI. 



Dialeurodes panacis Corbett 1935 

Dialeurodes panacis Corbett 1935b: 741. Syntypes. Malaya; on Panax fruticosum. 


Host: Araliaceae: Panax fruticosum. 

Dialeurodes papulae Singh 1932 

Dialeurodes papulae Singh 1932: 83. Syntypes. Burma: on unidentified tree, ZSI. 



Dialeurodes pauliani Cohic 1966 

Dialeurodes (Gigaleurodes) pauliani Cohic 1966b: 41. Syntypes. Nigeria: on undetermined host, BMNH. 

Dialeurodes pauliani Cohic; Mound and Halsey 1978: 145 

Distribution: 4: Nigeria.


Dialeurodes philippinensis Takahashi 1936 

Dialeurodes (Gigaleurodes) philippinensis Takahashi 1936b: 217. Syntypes. Philippines: on unidentified tree, 

TARI. 

Dialeurodes philippinensis Takahashi; Mound and Halsey 1978: 145. 

Distribution: 5: Philippines; 9: Hawaii. 


Dialeurodes pilahensis Corbett 1935 

Dialeurodes pilahensis Corbett 1935b: 777. Holotype. Malaya: on Eugenia aquea. 


Host: Myrtaceae: Eugenia aquea. 

Dialeurodes platicus Bondar 1923 

Dialeurodes platicus Bondar 1923a: 106. Syntypes. Brazil: on Psidium guajava, MZU. 



Dialeurodes pseudocitri Takahashi 1942 

Dialeurodes pseudocitri Takahashi 1942e: 274. Syntypes. Cambodia: on undetermined tree. 



Dialeurodes psychotriae Dumbleton 1961 

Dialeurodes psychotriae Dumbleton 1961a: 123. Holotype. New Caledonia: on Psychotria deplanchei, 

CORSTOM. 


Host: Rubiaceae: Psychotria deplanchei. 

Dialeurodes punctata Corbett 1933 

Dialeurodes punctata Corbett 1933: 124. Syntypes. Malaya: on unidentified plant, BMNH. 



298

Dialeurodes radiipuncta Quaintance & Baker 1917 

Dialeurodes (Rabdostigma) radiilinealis Quaintance & Baker 1917: 418. Syntypes, Sri Lanka, 10.xi.1913, on 

Memecylon sp., USNM 

Dialeurodes radiipuncta Quaintance & Baker; Mound & Halsey 1978: 147. 

Lankaleurodes radiipuncta (Quaintance & Baker); David 1993: 23. 

Dialeurodes radiipuncta Quaintance & Baker; Martin & Mound 2007. 


Host: Melastomataceae: Memecylon sp. 

Dialeurodes rangooni Singh 1932 

Dialeurodes rangooni Singh 1932: 85. Syntypes. Burma: on Eugenia jambos, ZSI. 



Dialeurodes razalyi Corbett 1935 

Dialeurodes razalyi Corbett 1935b: 769. Syntypes. Malaya: on unidentified host, lost. 



Dialeurodes rempangensis Takahashi 1949 

Dialeurodes rempangensis Takahashi 1949: 49. Syntypes. Riouw (Riau) Islands (south of Singapore): on 

unidentified tree, HBL. 



Dialeurodes rengas Corbett 1935 

Dialeurodes rengas Corbett 1935b: 765. Syntypes. Malaya: on Gluta sp. 



Dialeurodes reticulosa Corbett 1935 

Dialeurodes reticulosa Corbett 1935b: 740. Syntypes. Malaya: on Alyxia forbesii. 


299

Host: Apocynaceae: Alyxia forbesii. 

Dialeurodes rotunda Singh 1931 

Dialeurodes rotunda Singh 1931: 26. Syntypes. India: on Eugenia operculata, ZSI. 


Host: Myrtaceae: Eugenia operculata. 

Dialeurodes saklaspurensis David 1976 

Dialeurodes saklaspurensis David 1976: 261. Holotype pupal case. India: Saklaspur, 28.iii.1969, B.V. David, 

on unidentified tree, IDAV. 



Dialeurodes sandorici Corbett 1935 

Dialeurodes sandorici Corbett 1935b: 770. Syntypes. Malaya: on Sandoricum indicum. 


Host: Meliaceae: Sandoricum indicum. 

Dialeurodes sepangensis Corbett 1935 

Dialeurodes sepangensis Corbett 1935b: 758. Syntypes. Malaya: on Eugenia aquea. 


Host: Myrtaceae: Eugenia aquea. 

Dialeurodes sheryli (P. M. M. David) 2000 

Shanthiniae sheryli David 2000, in P.M.M. David 2000: 125. Holotype puparium, India, Tamil Nadu, 

Karaiyar Dam, 14.iv.1993, P. M. M. David, on unidentified plant, IARI. 

Dialeurodes sheryli (P. M. M. David); Martin & Mound 2007: 31. 



Dialeurodes shintenensis Takahashi 1933 

Dialeurodes (Rhabdostigma) shintenensis Takahashi 1933: 11-12. Syntypes. Taiwan: on Machilus sp., TARI. 

Dialeurodes shintenensis Takahashi; Mound and Halsey 1978: 149. 

300



Dialeurodes shoreae Corbett 1933 

Dialeurodes shoreae Corbett 1933: 124. Syntypes. Malaya: on Shorea glauca, BMNH. 


Host: Dipterocarpaceae: Shorea glauca. 

Dialeurodes siemriepensis Takahashi 1942 

Dialeurodes siemriepensis Takahashi 1942e: 278. Syntypes. Cambodia: on Lauraceae tree. 



Dialeurodes simmondsi Corbett 1927 

Dialeurodes (Gigaleurodes) simmondsi Corbett 1927: 25. Syntypes. Malaya: on Cocos nucifera. 

Dialeurodes simmondsi Corbett; Mound and Halsey 1978: 149. 



Dialeurodes striata Corbett 1935 

Dialeurodes striata Corbett 1935b: 760. Syntypes. Malaya: on Adinobotrys atropurpureus. 



Dialeurodes struthanthi Hempel 1901 

Dialeurodes struthanthi Hempel 1901: 387. Syntypes. Brazil: on Struthantus flexicaulis growing on orange 

(Citrus aurantium). 


Host: Loranthaceae: Steirotis flexicaulis. 

Dialeurodes sundararajani Sundararaj & Dubey 2006 

Dialeurodes sundararajani Sundararaj & Dubey 2006: 62-67. Holotype puparium, India, on Sonerilla 

wallichi; Meganathan & David 1994. 


Host: Unidentified plant (Melastomataceae: Sonerilla wallichi; Ulmaceae: Trema orientalis hosts record prior 

301

to publication of species) 

Dialeurodes tanakai Takahashi 1942 

Dialeurodes tanakai Takahashi 1942g: 302. Holotype. Thailand: on unidentified host. 


Host: unidentified host. 

Dialeurodes townsendi Quaintance & Baker 1917 

Dialeurodes (Dialeuroplata) townsendi Quaintance & Baker 1917: 436. Syntypes. Philippines: on "fern", 

USNM. 

Dialeurodes townsendi Quaintance & Baker; Mound and Halsey 1978: 150. 

Distribution: 5: Philippines. 

Host: a "fern". 

Dialeurodes tricolor Quaintance & Baker 1917 

Dialeurodes tricolor Quaintance & Baker 1917: 419. Syntypes. Brazil: on Myrtaceae, USNM. 



Dialeurodes tuberculosa Corbett 1935 

Dialeurodes tuberculosa Corbett 1935b: 768. Syntypes. Malaya: on Cinnamomum sp. 



Dialeurodes turpiniae (Meganathan & David) 1994 

Kanakarajiella turpiniae Meganathan & David 1994: 40. Holotype. India: on Turpinia malabarica. 

Dialeurodes turpiniae (Meganathan & David); Martin & Mound 2007. 


Host: Staphyleaceae: Turpinia malabarica. 

Dialeurodes vitis Corbett 1935 

Dialeurodes vitis Corbett 1935b: 764. Syntypes. Malaya: on Vitex sp. 



302

Dialeurodes vulgaris Singh 1931 

Dialeurodes vulgaris Singh 1931. Syntypes. India: on Eugenia jambos and Jasminum sambac. 

Kanakarajiella vulgaris (Singh); David & Sundararaj 1993: 294; Jensen 2001: 281. 

Dialeurodes vulgaris Singh; Martin & Mound 2007. 


Host: Asteraceae: Bidens pilosa; Fabaceae: Erythrina listhosperma, Erythrina abyssinica, 

Erythrina lithosperma, Eugenia jambos, Eugenia jambolana; Oleaceae: Jasminum frutescens, 

Jasminum sambac; Rubiaceae: Coffea arabica, Coffea excelsa, Coffea robusta, Jasminum sp., 

Canthium dicococcum; Rubiaceae: Randia malabarica, Randia sp. 

Note: Jensen (1999) treated Gigaleurodes (Quaintance & Baker) as a species group in his cladistical analysis 

of this genus Dialeurodes and its close relatives, in which he included: D. decaspermi, D. 

hongkongensis, D. indica, D. maxima, D. minahassai, D. radiilinealis, D. rotunda, D. saklaspurensis, 

R. eugeniae, R. radiirugosa and R. tristylii. Other authors (Sundararaj & David, 1994) have treated 

Gigaleurodes as a distinct genus. 

Dialeurodes wendlandiae Meganathan & David 1994 

Dialeurodes wendlandiae Meganathan & David 1994: 32. Holotype. India: on Wendladtia thyrsoidea. 


Host: Rubiaceae: Wendlandia thyrosoidea. 

Dialeurolobus Danzig 1964 

Dialeurolobus Danzig 1964a: 634. Type species. Dialeurolobus pulcher Danzig 1964, by monotypy. 



erythrinae x x 

pulcher x x 

rhamni x 

Total = 3 species 0 0 2 0 2 1 0 0 0 

303

Dialeurolobus erythrinae (Corbett) 1935 

Dialeurodes erythrinae Corbett 1935b: 747. Syntypes. Malaya: on Erythrina lithosperma and E. stricta. 

Dialeurolobus erythrinae (Corbett); Martin & Mound 2007: 31. 

Aleuromarginatus erythrinae Selvakumaran & David 1996: 60. 

Distribution: 5: Hong Kong; 6: Malaya; Malaysia. 

Host: Fabaceae: Erythrina stricta, Erythrina lithosperma; Rutaceae: Citrus sp. 

Dialeurolobus pulcher Danzig 1964 

Dialeurolobus pulcher Danzig 1964a: 635. Holotype. USSR: Moldavia, on Crataegus sp., ZIN; Suh et al. 

2006. 

Distribution: 3: Ira, Russia, USSR; 5: Korea. 

Host: Rosaceae: Crataegus pentagyna, Crataegus sp., Rosaceae: Pyracantha coccinea; Rosa damascena; 

Rutaceae: Poncirus trifoliata. 

Dialeurolobus rhamni Bink-Moenen 1992 

Dialeurolobus rhamni Bink-Moenen 1992, in Bink-Moenen and Gerling 1992: 26. Holotype female. Israel: 

Menahem, 15.xi.1969, on Rhamnus palaestina, BMCol. 

Distribution: 3: Iran, Israel. 

Host: Rhamnaceae: Rhamnus palaestina; Ziziphus spina-cristi. 

304

Dialeurolonga Dozier 1928 

Dialeurodes (Dialeurolonga) Dozier 1928: 1001. Type species Dialeurodes elongata, by original designation 

and monotypy 

Dialeurolonga Dozier, full genus by Takahashi 1951: 354. 

Reference: Bink-Moenen 1983: 132. - Key to African species; Sundararaj & Dubey 2006 - Key to 

Indian species. 



africana x 

aguariae x 

ambilaensis x 

angustata x 

aphloiae x 

bambusae x 

bambusicola x 

brevispina x 

davidi x 

elliptica x 

elongata x x x 

erythroxylonis x 

eugeniae x 

fici x 

graminis x 

guettardae x 

hoyti x 

305


kumargiriensis x 

lagerstroemiae x 

lamtoensis x 

lata x 

maculata x x 

malleshwaramensis x 

mameti x 

mauritiensis x 

milloti x 

multipapilla x 

multipori x 


nigra x 

operculobagta x 

paradoxa x 

paucipapillata x 

pauliani x 

perinetensis x 

photiniana x 

phyllarthronis x 

ravensarae x 

rhamni x 

robinsoni x 

rotunda x 

306


rusostigmoides x 

similis x 

simplex x 

strychnosicola x 

subrotunda x 

swainei x 

takahashii x 

tambourissae x 

trialeuroides x 

vendranae x 

307

Total = 52 species 0 1 2 37 2 11 3 0 0 

Dialeurolonga africana (Newstead) 1921 

Aleurodes africana Newstead 1921: 528-529. Syntypes: Nigeria: Ibadan, Moor Plantation, v.1917, C. O. 

Farquharson, on Salacia sp., BMNH. 

Dialeurolonga africana (Newstead); Takahashi 1955a: 393. 


Host: Celastraceae: Salacia sp. 

Natural enemies: Predator - LEPIDOPTERA/Noctuidae: Coccidiphaga scitula. 

Dialeurolonga aguariae Takahashi 1951 

Dialeurolonga agauriae Takahashi 1951a: 354-356. Syntypes. Madagascar: Mt. Tsaratanana, 2200m, x.1949, 

R. Paulian, on Aguaria sp, MNHNP. 


Host: Ericaceae: Agauria sp. 

Dialeurolonga ambilaensis Takahashi 1955 

Dialeurolonga ambilaensis Takahashi 1955a: 323. Holotype. Madagascar: Ambila-Lemaitso, iii.1954, A. 

Robinson, on "Raisonjo", MNHNP. 


Host: "raisonjo". 

Dialeurolonga angustata Takahashi 1961 

Dialeurolonga angustata Takahashi 1961: 323. Holotype. Madagascar: Ambilobe, Ankatoto, iv.1951, R. 

Paulian, on "Fotsy-avadkica", MNHNP. 


Host: "Fotsy-avadkica". 

Dialeurolonga aphloiae Takahashi 1955 

Dialeurolonga aphloiae Takahashi 1955a: 377-378. Holotype. Madagascar: Ankaratra, Manjakatompo, 

2000m, 24.v.1950, R. Paulian and R. Mamet, on Aphloia theaeformis, MNHNP. 


Host: Flacourtiaceae: Aphloia theaeformis. 

Dialeurolonga bambusae Takahashi 1961 

308

309 

Dialeurolonga bambusae Takahashi 1961: 325. Syntypes. Madagascar: on bamboo [?Bambusa sp.], MNHNP. 



Dialeurolonga bambusicola Takahashi 1951 

Trialeurodes bambusicola Takahashi 1951a: 367. Syntypes. Madagascar: on bamboo [?Bambusa sp.], 

MNHNP. 

Dialeurolonga bambusicola (Takahashi); Takahashi & Mamet 1952b: 118 



Dialeurolonga brevispina Takahashi 1951 

Dialeurolonga brevispina Takahashi 1951a: 356. Syntypes. Madagascar: on "maimbovitsika", MNHNP. 


Host: on "maimbovitsika". 

Dialeurolonga davidi Dubey & Sundararaj 2006 

Dialeurolonga davidi Dubey & Sundararaj 2006. Holotype puparium, India, Karnataka, Kudremukh National 

Park, 10.viii.2001, A. K. Dubey, on unidentified plant, IFRI. 



Dialeurolonga elliptica Takahashi 1955 

Dialeurolonga elliptica Takahashi 1955a: 378-379. Holotype. Madagascar: on unidentified host, MNHNP. 


Host: on unidentified host. 

Dialeurolonga elongata Dozier 1928 

Dialeurodes (Dialeurolonga) elongata Dozier 1928: 1001-1002. Syntypes. Pakistan: on Citrus sp., USNM. 

Dialeurolonga elongata Dozier; Takahashi 1951a: 354; Sundararaj & Dubey 2006. 

Distribution: 3: Iran; 5: China; 6: India; Pakistan. 

Host: Apocynaceae: Taberaemontana heyneana; Asteraceae: Calendula arvensis; Rubiaceae: Ixora 

parviflora, Ixora coccinea; Rutaceae: Murraya exotica, Citrus sp.; Sapindaceae: Litchi chinensis. 

Natural enemies: Parasitoids - Aphelinidae: Encarsia aseta; Predators - Chrysopidae: Chrysopa sp.; 

Coccinellidae: Brumoides suturalis, Scymnus sp.

Dialeurolonga erythroxylonis Takahashi 1955 

Dialeurolonga erythroxylonis Takahashi 1955a: 379. Syntypes. Madagascar: on Erythroxylon ampullaceum, 

MNHNP. 


Host: Erythroxylaceae: Erythroxylon ampullaceum. 

Dialeurolonga eugeniae Takahashi 1951 

Dialeurolonga eugeniae Takahashi 1951a: 362. Holotype. Madagascar: on Eugenia sp., HBL. 



Dialeurolonga fici David & Subramaniam 1976 

Dialeurolonga fici David & Subramaniam 1976: 194. Holotype. India: on Ficus religiosa, ZSI. 


Host: Moraceae: Ficus religiosa. 

Natural enemies: Aphelinidae: Eretmocerus dialeurolongae. 

Dialeurolonga graminis (Takahashi) 1951 

Trialeurodes graminis Takahashi 1951a: 370. Syntypes. Madagascar: on Graminae, HBL. 

Dialeurolonga graminis (Takahashi); Takahashi & Mamet 1952b: 118. 


Host: Poaceae, grass. 

Dialeurolonga guettardae Martin 2005 

Dialeurolonga guettardae Martin 2005b: 41. Holotype puparium, Belize, CFR, 5 VI 2002, J. H. Martin, on 

Guettarda combesi, BMNH. 


Host: Rubiaceae: Guettarda combesi. 

Dialeurolonga hoyti Mound 1965 

Dialeurolonga hoyti Mound 1965c: 146. Holotype. Nigeria: on coffee [Coffea sp.], BMNH. 

Dialeurolonga sarcocephali Cohic 1966b: 47. Syntypes. Congo: on Sarcocephalus esculentus; synonymy 


310

311 

Distribution: Angola; Congo; Ivory Coast; Nigeria 

Host: Euphorbiaceae: Hymenocardia acida; Meliaceae: Entandrophragma cylindrium; Meliaceae: 

Entandrophragma utile; Rubiaceae: Coffea sp., Coffea stenophylla, Morinda morindoides, Psychotria 

psychotrichoides, Sarcocephalus esculentus. 

Dialeurolonga kumargiriensis Dubey & Sundararaj 2006 

Dialeurolonga kumargiriensis Dubey & Sundararaj 2006. Holotype puparium, India, Karnataka, Kumargiri, 

1.ix.2001, A. K. Dubey, on Ficus nervosa, IFRI. 


Host: Moraceae: Ficus nervosa. 

Dialeurolonga lagerstroemiae Jesudasan & David 1991 

Dialeurolonga lagerstroemiae Jesudasan and David 1991: 309. Holotype pupal case. India: on Lagerstroemia 

speciosa, IDAV; Sundararaj & Dubey 2006. 


Host: Lythraceae: Lagerstroemia speciosa. 

Dialeurolonga lamtoensis Cohic 1969 

Dialeurolonga lamtoensis Cohic 1969: 123. Holotype. Ivory Coast: on Antiaris africana. 


Host: Euphorbiaceae: Alchornea cordifolia; Moraceae: Antiaris africana. 

Dialeurolonga lata Takahashi 1955 

Dialeurolonga lata Takahashi 1955a: 382. Syntypes. Madagascar: on unidentified host, HBL. 



Dialeurolonga maculata (Singh) 1931 

Aleurotulus maculata Singh 1931: 89. Syntypes. India: on Ficus religiosa. 

Dialeurolonga maculata (Singh); David & Jesudasan 1989a: 371; Sundararaj & Dubey 2006. 


Host: Malvaceae: Abutilon sp.; Moraceae: Ficus religiosa. 

Dialeurolonga malleshwaramensis Sundararaj 2001 

Dialeurolonga malleshwaramensis Sundararaj 2001: 191. Holotype puparium, India, Karnataka, Bangalore,

312 

7.ii.1999, R. Sundararaj, on Polyalthia longifolia, IWST; Sundararaj & Dubey 2006. Redescription by 

Dubey and Ko 2007. 


Host: Annonaceae: Polyalthia longifolia. 

Dialeurolonga mameti Takahashi 1955 

Dialeurolonga mameti Takahashi 1955a: 383. Holotype. Madagascar; on unidentified host, MNHNP. 



Dialeurolonga mauritiensis (Takahashi) 1938 

Dialeurodes mauritiensis Takahashi 1938d: 260. Syntypes. Mauritius: on unidentified host, MNHNP. 

Dialeurolonga mauritiensis (Takahashi); Takahashi 1951a: 354. 



Dialeurolonga milloti Takahashi 1951 

Dialeurolonga milloti Takahashi 1951a: 364. Syntypes. Madagascar: on clove, HBL. 


Host: on clove, Fabaceae: Trifolium sp. 

Dialeurolonga multipapilla Takahashi 1955 

Dialeurolonga multipapilla Takahashi 1955a: 385. Syntypes. Madagascar: on unidentified host, MNHNP. 



Dialeurolonga multipori Dubey & Sundararaj 2006 

Dialeurolonga multipori Dubey & Sundararaj2006. Holotype puparium, India, Karnataka, Yellapur, 1.x.2001, 

A. K. Dubey, on Casearia esculenta, IFRI. 


Host: Flacourtiaceae: Casearia esculenta. 

Dialeurolonga multituberculata Dubey & Sundararaj 2006

313 

Dialeurolonga multituberculata Dubey & Sundararaj 2006. Holotype puparium, India, Karnataka, Yellapur, 

11.x.2001, A. K. Dubey, on Careya arborea, IFRI. 


Host: Lecythidaceae: Careya arborea. 

Dialeurolonga nigra Takahashi & Mamet 1952 

Dialeurolonga nigra Takahashi & Mamet 1952b: 111. Syntypes. Madagascar: on "mahogo-ala", HBL. 


Host: on "mahogo-ala" [possibly cassava, Manihot esculenta (Euphorbiaceae). 

Dialeurolonga operculobagta Martin & Carver 1999 

Dialeurolonga operculobata Martin & Carver 1999, in Martin 1999: 69. Holotype puparium. Australia: on 

Leptospermum saundersoni, ANIC. 


Host: Myrtaceae: Leptospermum saundersoni. 

Dialeurolonga paradoxa Takahashi 1955 

Dialeurolonga paradoxa Takahashi 1955a: 387. Holotype. Madagascar: on "vahimboronandria", MNHNP. 


Host: "vahimboronandria". 

Dialeurolonga paucipapillata Cohic 1969 

Dialeurolonga paucipapillata Cohic 1969: 128. Holotype. Ivory Coast: on Drypetes floribunda. 


Host: Euphorbiaceae: Drypetes floribunda. 

Dialeurolonga pauliani Takahashi 1951 

Dialeurolonga pauliani Takahashi 1951a: 360. Syntypes. Madagascar: on forest climbing plant 

"sandrahidraky", MNHNP. 


Host: forest climbing plant "sandrahidraky"; Asteraceae: Psiadia sp.

Dialeurolonga perinetensis Takahashi & Mamet 1952 

Dialeurolonga perinetensis Takahashi & Mamet 1952b: 114. Holotype. Madagascar: on "kijy", MNHNP. 


Host: on "kijy". [possibly Ochrocarpos orthocladus (Clusiaceae)]. 

Dialeurolonga photiniana (Chen) 1997 

Aleurodicus photiniana Chen 1997: 137. Holotype. China. 

Dialeurolonga photiniana (Chen); Martin & Mound 2007: 33. 


Host: Rosaceae: Photinia sp. 

Dialeurolonga phyllarthronis Takahashi 1955 

Dialeurolonga phyllarthronis Takahashi 1955a: 389. Syntypes. Madagascar: on Phyllarthron bojerianum, 

MNHNP. 


Host: Bignoniaceae: Phyllarthron bojerianum. 

Dialeurolonga pseudocephalidistincta Dubey & Sundararaj 2006 

Dialeurolonga pseudocephalidistincta Dubey & Sundararaj 2006. Holotype puparium, India, Karnataka, 

Kudremukh National Park, 10.viii.2001, A. K. Dubey, on an identified plant, IFRI. 



Dialeurolonga ravensarae Takahashi & Mamet 1952 

Dialeurolonga ravensarae Takahashi & Mamet 1952b: 113. Syntypes. Madagascar: on Ravensara sp., 

MNHNP. 


Host: Lauraceae: Ravensara sp. 

Dialeurolonga rhamni Takahashi 1961 

Dialeurolonga rhamni Takahashi 1961: 326. Syntypes. Madagascar: on Rhamnaceae, MNHNP. 


Host: Rhamnaceae. 

314

Dialeurolonga robinsoni Takahashi & Mamet 1952 

Dialeurolonga robinsoni Takahashi & Mamet 1952b: 118. Syntypes. Madagascar: on "vahinakany", MNHNP. 


Host: "vahinakany". 

Dialeurolonga rotunda Takahashi 1961 

Dialeurolonga rotunda Takahashi 1961: 328. Syntypes. Madagascar: on "fotsy-avadika", MNHNP. 


Host: "fotsy-avadika" [Cracephalum sarcobasis (Asteraceae)] 

Dialeurolonga rusostigmoides Martin 1999 

Dialeurolonga rusostigmoides Martin 1999: 70. Holotype puparium. Australia: on undetermined plant, ANIC. 


Host: undetermined plant. 

Dialeurolonga similis Takahashi 1955 

Dialeurolonga similis Takahashi 1955a: 391. Syntypes. Madagascar: on unidentified host, MNHNP. 



Dialeurolonga simplex Takahashi 1955 

Dialeurolonga simplex Takahashi 1955a: 393. Syntypes. Madagascar: on Aphloia theaeformis, HBL; Martin 

1985a: 461. 

Distribution: 4: Madagascar; Mauritius. 

Host: Flacourtiaceae: Aphloia theaeformis; Rutaceae: Citrus sp. 

Dialeurolonga strychnosicola Cohic 1966 

Dialeurolonga strychnosicola Cohic 1966a: 38. Syntypes. Congo: on Strychnos variabilis, BMNH. 

Distribution: 4: Chad; Congo. 

Host: Chrysobalanaceae: Chrysobalanus orbicularis; Dichapetalaceae: Dichapetalum brazzae. 

Dilleniaceae: Tetracera alnifolia; Fabaceae: Leptoderris nobilis, Pentaclethra eetveldeana. 

Linaceae: Ochthocosmus africanus; Loganiaceae: Strychnos variabilis; Menispermaceae: Triclisia 

gilletii. 

315

Dialeurolonga subrotunda Takahashi 1955 

Dialeurolonga subrotunda Takahashi 1955a: 395. Syntypes. Madagascar: on unidentified host, MNHNP. 



Dialeurolonga swainei Martin 1999 

Dialeurolonga swainei Martin 1999: 70. Holotype puparium. Australia: on Ficus sp., ANIC. 



Dialeurolonga takahashii David & Jesudasan 1989 

Dialeurolonga maculata Takahashi 1951a: 358. Syntypes. Madagascar: on Weinmannia sp., HBL 

Dialeurolonga takahashii David & Jesudasan 1989: 371, name preoccupied by Dialeurolonga maculata 

(Singh) 1931 

Distribution: 4: Madagascar; 6: India. 

Host: Cunoniaceae: Weinmannia sp.; Moraceae: Ficus religiosa. 

Natural enemies: Aphelinidae: Eretmocerus dialeurolongae. 

Dialeurolonga tambourissae Takahashi 1955 

Dialeurolonga tambourissae Takahashi 1955a: 397. Syntypes. Madagascar: on Tambourissa sp., MNHNP. 


Host: Monimiaceae: Tambourissa sp. 

Dialeurolonga trialeuroides Takahashi & Mamet 1952 

Dialeurolonga trialeuroides Takahashi & Mamet 1952b: 116. Syntypes. Madagascar: on unidentified host, 

MNHNP. 



Dialeurolonga vendranae Takahashi 1961 

Dialeurolonga vendranae Takahashi 1961: 330. Syntypes. Madagascar: on bamboo [Bambusa sp.], MNHNP. 


Host: probably Poaceae: Bambusa sp. 

316

Dialeuronomada Quaintance & Baker 1917 

Dialeurodes (Dialeuronomada) Quaintance & Baker 1917: 51. Type species: Dialeurodes (Dialeuronomada) 

dissimilis Quaintance & Baker 1917. 

Dialeuronomada Quaintance & Baker; full genus by Sundararaj & David 1991. 

References: Sundararaj & David 1991; Dubey & Sundararaj 2004 - Key to Indian species. 

Comment: Currently, this genus is only known to occur in India. Martin & Mound (2007) placed the species 

in this genus in the genus Dialeurodes. 



ayyanarensis x 

binkae x 

biventralis x 

canthiae x 

dissimilis x 

gigantica x 

granulata x 

ixorae x 

keralaensis x 

martini x 

nagpurensis x 

palmata x 

remadeviae x 

ruiphaga x 

russellae x 

saklespurensis x 

Total = 16 species 0 0 0 0 0 16 0 0 0 

317

Dialeuronomada ayyanarensis Sundararaj & David 1991 

Dialeuronomada ayyanarensis Sundararaj & David 1991a: 29. Holotype puparium, India, Tamil Nadu, 

Ayyanar falls, 11.ii.1989, K. Regu, on Mimusops sp. 

Dialeurodes ayyanarensis (Sundararaj & David); Martin & Mound 2007: 


Host: Anacardiaceae: Holigarna grahamii; Euphorbiaceae: Bischofia javanica, Phyllanthus emblica; 

Lauraceae: Cinnamomum malabatrum; Rubiaceae: Canthium coromandelicum, Randia rugulosa; 

Rutaceae: Naringi crenulata; Sapotaceae: Mimusops sp. 

Dialeuronomada binkae Sundararaj & David 1991 

Dialeuronomada binkae Sundararaj & David 1991a: 30. Holotype puparium, India, Maharashtra 

Mahableshwar, 28.iii.1987, B.V. David, on Pavetta sp.; Dubey & Sundararaj 2004: 150. 


Host: Lauraceae: Neolitsea zeylanica; Liliaceae: Smilax monophylla; Oleaceae: Olea dioica. 

Oxalidaceae: Oxalis sp.; Rubiaceae: Pavetta sp.; Rutaceae: Atalantia racemosa, Ixora coccinea. 

Dialeuronomada biventralis Sundararaj & David 1991 

Dialeuronomada biventralis Sundararaj & David 1991a: 32. Holotype puparium, India, Tamil Nadu, Sivakasi, 

25.iii.1988, S.Jayaselvan, on Antidesma sp. 


Host: Euphorbiaceae: Antidesma sp.; Rutaceae: Ruta graveolens. 

Dialeuronomada canthiae Sundararaj & David 1991 

Dialeuronomada canthiae Sundararaj & David 1991a: 33. Holotype puparium, India, Tamil Nadu, Nuchirai, 

3.viii.1987, R. Sundararaj, on Canthium dicoccum, in (USDA, BMNH, IARI, ZSI); Dubey & 

Sundararaj 2004: 150. 


Host: Euphorbiaceae: Bischofia javanica; Oleaceae: Olea dioica; Rubiaceae: Canthium dicoccum. 

Dialeuronomada dissimilis (Quaintance & Baker) 1917 

Dialeurodes (Dialeuronomada) dissimilis Quaintance & Baker 1917: 424. Syntypes, India, Saharanpuyr, 

xi.1910, R.S. Woglum, on Phyllanthus sp., USNM. 

Dialeurodes dissimilis Quaintance & Baker; Mound and Halsey 1978: 137. 

Dialeuronomada dissimilis (Quaintance & Baker); Sundararaj & David 1991a: 35; Dubey & Sundararaj 2004: 

151. 

318

319 


Host: Euphorbiaceae: Antidesma menasu, Phyllanthus myrtifolius; Flacourtiaceae: Hydnocarpus wightiana; 

Rubiaceae: Ixora sp., Ixora pavetta, Pavetta sp.; Rutaceae: Ruta graveolens. 

Dialeuronomada gigantica Sundararaj & David 1991 

Dialeuronomada gigantica Sundararaj & David 1991a: 36. Holotype puparium, India, Karnataka, Saklaspur, 

17.v.1967, B. V. David, on unidentified plant, (holotype and paratype in USDA, BMNH, IARI, ZSI). 



Dialeuronomada granulata Sundararaj & David 1991 

Dialeuronomada granulata Sundararaj & David 199a: 38.Holotype puparium, India, Tamil Nadu, Sivakasi, 

25.ii.1988, S. Jayaselvan, on Ixora coccinea, ZSI. 


Host: Rubiaceae: Ixora coccinea. 

Dialeuronomada ixorae (Singh) 1931 

Dialeurodes ixorae Singh 1931: 38. Syntypes. India: on Ixora coccinea, BMNH 

Dialeuronomada ixorae (Singh); Sundararaj & David 1991a: 39; Dubey & Sundararaj 2004: 151. 


Host: Euphorbiaceae: Antidesma acidum; Fabaceae: Flemingia macrophylla; Oleaceae: Olea dioica; 

Phyllanthaceae: Aporosa lindleana; Rubiaceae: Ixora coccinea, Ixora sp., Tarenna asiatica; Rutaceae: 

Archronycha zeylanicum, Atalantia monophylla; Sapotaceae: Mimusops hexandra. 

Natural enemies: Aphelinidae: Encarsia ixorae. 

US Port Interception: Sri Lanka on unknown plant. 

Dialeuronomada keralaensis Meganathan & David 1994 

Dialeuronomada keralaensis Meganathan & David 1994: 35. Holotype puparium, Sispara (Silent Valley), 

29.x.1991, P. Meganathan, on Ilex walkeri, David Collection. 


Aquifoliaceae: Ilex walkeri. 

Dialeuronomada martini Sundararaj & David 1991 

Dialeuronomada martini Sundararaj & David 1991a: 41. Holotype puparium, India, Tamil Nadu, Peachiparai, 

5.viii.1987, R. Sundararaj, on Ixora sp., ZSI.; Dubey & Sundararaj 2004: 152.

320 


Host: Euphorbiaceae: Glochidion zeylanicum; Rubiaceae: Ixora sp.; Rutaceae: Atalantia racemosa. 

Dialeuronomada nagpurensis Sundararaj & David 1991 

Dialeuronomada nagpurensis Sundararaj & David 1991a: 42. Holotype puparium, India, Maharashtra, 

Nagpur, 30.vii.1987, B.V. David, on Ixora sp., ZSI; Dubey & Sundararaj 2004: 152. 


Host: Sapindaceae: Sapindus emarginatus. 

Dialeuronomada palmata Sundararaj & David 1991 

Dialeuronomada palmata Sundararaj & David 1991a: 44. Holotype puparium, India, Tamil Nadu, Munchirai, 

6.viii.1987, R. Sundararaj, on unidentified climber, ZSI. 


Host: unidentified climber. 

Dialeuronomada remadeviae Dubey & Sundararaj 2004 

Dialeuronomada remadeviae Dubey & Sundararaj 2004: 152. Holotype puparium, India, Karnataka, 

18.x.2001, A. K. Dubey, on unidentified plant; deposited in FRI, Dehra, India. 



Dialeuronomada rubiphaga Dubey & Sundararaj 2004 

Dialeuronomada rubiphaga Dubey & Sundararaj 2004: 154. Holotype puparium, India, Goa, Volpoi, 

27.ii.2001, A. K. Dubey, ex. Pavetta sp., IFRI. 


Host: Rubiaceae: Pavetta sp. 

Dialeuronomada russellae Sundararaj & David 1991 

Dialeuronomada russellae Sundararaj & David 1991a: 45. Holotype puparium, India, Tamil Nadu, 

Munchirai, 3.viii.1987, R. Sundararaj, on Olea dioica, ZSI; Dubey & Sundararaj 2004. 


Host: Dipterocarpaceae: Nothopegia sp.; Euphorbiaceae: Bischofia javanica, Phyllanthus emblica; Lauraceae: 

Smilax zeylanica; Loganiaceae: Strychnos dalzellii; Myrtaceae: Syzygium sp.; Oleaceae: Olea dioica; 

Rubiaceae: Psychotria sp., Canthium angustifolium. 

Sapindaceae: Dimocarpus longon; Symplocaceae: Symplocos sp.

Dialeuronomada saklespurensis Regu & David 1993 

Dialeuronomada saklespurensis Regu & David 1993: 83. Holotype puparium, India, Saklespur, Karnataka, 

4.ii.1990, K. Regu, on Dimocarpus longon, in New Delhi. 


Host: Sapindaceae: Dimocarpus longon. 

321

Dialeuropora Quaintance & Baker 1917 

Dialeurodes (Dialeuropora) Quaintance & Baker 1917: 434. Type species. Dialeurodes (Dialeuropora) 

decempunctata Quaintance & Baker 1917, by monotypy. 

Dialeuropora Quaintance & Baker 1917; full genus by Takahashi 1934b: 46. 




bipunctata x 

brideliae x 

centrosemae x 

congoensis x 

decempuncta x x x x x 

hassensanensis x 

heptapora x 

holboelliae x 

indochinensis x 

jendera x 

langsat x 

malayensis x 

mangiferae x 

murrayae x 

papillata x 

photiniana x 

portugaliae x 

pterolobiae x 

silvarum x 

urticata x 

viburni x 

Total = 21 species 0 0 1 3 8 11 1 1 0 

322

Dialeuropora bipunctata (Corbett) 1933 

Dialeurodes bipunctata Corbett 1933: 128. Holotype. Malaya: on Gluta sp. 

Dialeuropora bipunctata (Corbett); Russell 1962b: 65. 



Dialeuropora brideliae (Takahashi) 1932 

Dialeurodes (Dialeuropora) brideliae Takahashi 1932: 15. Syntypes. Taiwan: on Bridelia ovata and Machilus 

sp., TARI. 

Dialeurodes (Dialeuropora) brideliae Takahashi 1934b: 46. 


Host: Euphorbiaceae: Bridelia ovata; Lardizabalaceae: Stauntonia sp.; Lauraceae: Machilus sp. 

Dialeuropora centrosemae (Corbett) 1935 

Dialeurodes centrosemae Corbett 1935b: 750. Syntypes. Malaya: on Centrosema plumieri. 

Dialeuropora centrosemae (Corbett); Russell 1959: 185. 


Host: Fabaceae: Centrosema plumieri. 

Dialeuropora congoensis Cohic 1966 

Dialeuropora congoensis Cohic 1966b: 52. Syntypes. Congo: on Neosloetiopsis kamerunensis. 


Host: Moraceae: Neosloetiopsis kamerunensis. 

Dialeuropora decempuncta (Quaintance & Baker) 1917 

Dialeurodes (Dialeuropora) decempuncta Quaintance & Baker 1917: 434. Syntypes. Sri Lanka: on 

Cinnamomum sp., USNM. 

Dialeuropora decempuncta Quaintance & Baker; Takahashi 1934b: 46. 

Dialeurodes dothioensis Dumbleton 1961a: 121. Holotype. New Caledonia: on unidentified plant, 

CORSTOM; synonymy according to Mound and Halsey 1978: 159. 

Dialeurodes perseae Corbett 1935b: 749. Syntypes. Malaya: on Persea gratissima and other plants, lost; 

synonymy according to Martin 1985: 326. 

Dialeurodes (Dialeuropora) setigerus Takahashi 1932: 14. Syntypes. Taiwan: on Machilus sp, TARI; 

323

synonymy according to Takahashi 1934b: 46; Mound and Halsey 1978: 159 

Distribution: 3: Iran; 5: Taiwan; 6: Cambodia; India; Malaya; Malaysia; Pakistan; Sri Lanka; Thailand; 7: 

New Guinea, Sulawesi; 8: New Caledonia. 

Host: Annonaceae: Annona cheirimola, Annona squamosa; Boraginaceae: Cordia myxa; Burseraceae: 

Canarium sp.; Euphorbiaceae: Breynia sp.; Fabaceae: Acacia longifolia, Dalbergia sissoo; Lauraceae: 

Machilus sp., Persea gratissima, Cinnamomum sp.; Moraceae: Ficus racemosa; Morus alba, Streblus 

asper; Myrtaceae: Eucalyptus camalduleis; Rhamnaceae: Alphitonia sp., Rosaceae: Rosa sp., Rosa 

indica, Prunus sp. 

Natural enemies: Aphelinidae: Encarsia dialeuroporae, Encarsia longivalvula; Predators - Coccinellidae: 

Scymnus sp. 

Dialeuropora hassensanensis Takahashi 1934 

Dialeuropora hassensanensis Takahashi 1934b: 45. Holotype. Taiwan: on Lauraceae, TARI. 



Dialeuropora heptapora Regu & David 1991 

Dialeuropora heptapora Regu & David 1991b: 413. Holotype, India, Ambalamedu, Kerala, 14.xii.1989, K. 

Regu on Hiptage sp.; Regu Collection. 


Host: Malpighiaceae: Hiptage sp. 

Dialeuropora holboelliae Young 1944 

Dialeuropora holboelliae Young 1944: 132. Syntypes. China: on Holboellia sp. 


Host: Lardizabalaceae: Holboellia sp. 

Dialeuropora indochinensis Takahashi 1942 

Dialeuropora indochinensis Takahashi 1942e: 272. Holotype. Cambodia: on unidentified host. 



Dialeuropora jendera (Corbett) 1935 

Dialeurodes jendera Corbett 1935b: 750. Syntypes. Malaya: on Adinobotrys atropurpureus. 

Dialeuropora jendera (Corbett); Russell 1959: 185. 

324

325 


Host: Fabaceae: Adinobotrys atropurpureus 

Note: Specimens of this species or near this species were collected in Hawaii (USNM collection). 

Dialeuropora langsat (Corbett) 1935 

Dialeurodes langsat Corbett 1935b: 752. Syntypes. Malaya: on Lansium domesticum. 

Dialeuropora langsat (Corbett); Russell 1959: 185. 


Host: Meliaceae: Lansium domesticum Correa. 

Dialeuropora malayensis (Corbett) 1935 

Trialeurodes malayensis Corbett 1935b: 812. Syntypes. Malaya: on unidentified host. 

Dialeuropora malayensis (Corbett); Russell 1962b: 65. 



Dialeuropora mangiferae (Corbett) 1935 

Dialeurodes mangiferae Corbett 1935b: 751. Syntypes. Malaya: on Mangifera indica. 

Dialeuropora mangiferae (Corbett); Russell 1959: 186. 


Host: Anacardiaceae: Mangifera indica. 


Note: Martin (1988) reported Dialeuropora mangiferae? (Corbett) 1935 on Magnoliaceae: Elmerillea ovalis 

and Myrtaceae: Syzygium sp. from 7: Sulawesi. 

Dialeuropora murrayae (Takahashi) 1931 

Dialeurodes (Dialeuropora) murrayae Takahashi 1931b: 262. Syntypes. Taiwan: on Murraya sp. 

Dialeuropora murrayae (Takahashi); Russell 1959: 186. 

Distribution: 5: Japan; Taiwan; 6: India (Dubey, personal communication). 

Host: Rutaceae: Murraya sp. 

Dialeuropora papillata Cohic 1966 

Dialeuropora papillata Cohic 1966a: 40. Syntypes. Congo: on Ochthocosmus africanus. 

Dialeuropora cogniauxiae Cohic 1966a: 45. Syntypes. Congo: on Cogniauxia podalaena; synonymy

according to Bink-Moenen 1983:139. 

Dialeuropora ndiria Gameel 1971: 171. Holotype. Sudan: on "Ndii" [Bridelia micrantha]; synonymy 

according to Bink-Moenen 1983:139. 

Dialeuropora platysepali Cohic 1966b: 55. Syntypes. Congo: Brazzaville, on Platysepalum vanderystii; 

synonymy according to Bink-Moenen 1983:139. 

Distribution: 4: Angola; Chad, Congo; Ivory Coast; Nigeria, Sudan; 5: China. 

Host: Annonaceae: Uvaria brazzavillensis, Annonaceae: Uvaria scabrida; Apocynaceae: 

Carpodinus lanceolata; Araceae: Cryptoperma senegalense; Arecaceae: Elaeis guineensis. 

Bignoniaceae: Markhamia sessilis; Chrysobalanaceae: Chrysobalanus orbicularis; Connaraceae: 

Byrsocarpus viridis, Manotes pruinosa; Cogniauxia podalaena; Dilleniaceae: Tetracera alnifolia; 

Euphorbiaceae: Antidesma lanciniatum, Bridelia ferruginea, Bridelia micrantha, Hymenocardia 

ulmoides, Sapium cornutum; Fabaceae: Cassia javanica, Cassia sp., Dalbergia kisantuensis, 

Platysepalum vanderystii; Lardizabalaceae: Akebia trifoliate; Lauraceae: Persea americana, Persea 

grattissima; Lecythidaceae: Napoleona leonensis; Ochthocosmus africanus; Anthocleista inermis; 

Loganiaceae: Strychnos pungens, Strychnos spinosa, Strychnos variabilis; Myrtaceae: Psidium 

guajava, Syzygium brassavillense; Ochnaceae: Ochna gilletiana, Rhabdophyllum welwitschii; 

Passifloraceae: Barteria fistulosa; Rubiaceae: Gaertnera paniculata, Sarcocephalus esculentus, 

Urophyllum hirtellum; Smilaceae: Smilax kraussiana; Sterculiaceae: Cola nitida, Theobroma sp.; 

Zingiberaceae: Aframomum stipulatum. 

Aleurodicus photiniana Chen 1997: 137-143. 

Dialeuropora photiniana (Chen) 

Dialeuropora photiniana (Chen): Dubey and Ko 2006: 391-398. 


Host: Rosaceae: Photinia serrulata. 

Dialeuropora portugaliae Cohic 1966 

Dialeuropora portugaliae Cohic 1966a: 43. Holotype. Angola: on unidentified plant. 

Distribution: 4: Angola; Congo; Ivory Coast. 

Host: Euphorbiaceae: Hymenocardia acida; Flacourtiaceae: Caloncoba dusenii; Icacinaceae: Icacina mannii; 

Loganiaceae: Strychnos variabilis; Meliaceae: Trichilia heudelottii; Myrtaceae: Eugenia leonensis; 

Sapotaceae: Afrosersalisia cerasifera; Zingiberaceae: Aframomum daniellii. 

Dialeuropora pterolobiae David & Subramaniam 1976 

Dialeuropora pterolobiae David & Subramaniam 1976: 197. Holotype. India: on Pterolobium indicum, ZSI. 


Host: Fabaceae: Pterolobium indicum. 

326

Dialeuropora silvarum (Corbett) 1935 

Trialeurodes silvarum Corbett 1935b: 813. Syntypes. Malaya: on unidentified host. 

Dialeuropora silvarum (Corbett); Russell 1962b: 65. 



Dialeuropora urticata Young 1944 

Dialeuropora urticata Young 1944: 131. Syntypes. China: on Urtica sp. 


Host: Urticaceae: Urtica sp. 

Dialeuropora viburni (Takahashi) 1933 

Dialeuropora viburni (Takahashi) 1933: 8. Syntypes. Taiwan: on Viburnum awabucki, TARI 

Dialeuropora viburni (Takahashi); Russell 1959: 186. 


Host: Caprifoliaceae: Viburnum awabucki, Viburnum sp. 

327

Disiphon Russell 1993 

Disiphon Russell 1993: 463. Type species. Asterochiton dubienus Bondar 1923, by designation and 

monotypy, USNM. 

Diagnosis: Puparium with 2 shapes of glands on the dorsum (central glands normal, submarginal glands 

siphon-like, as in Siphoninus); margin dentate; submargin not separated from dorsal disk by suture or 

fold; submedial depressions faint; C1, T2, T3, A8 and caudal setae present; vasiform orifice 

subcircular or cordate with operculum nearly covering the orifice and lingula; lingula spatulate with 

minute spinules and a pair of minute setae; caudal furrow and ridges absent; venter - tracheal fold 

obscure. 



dubienus x 

russellae x 

Total = 2 species 0 2 0 0 0 0 0 0 0 

Disiphon dubienus (Bondar) 1923 

Asterochiton dubienus Bondar 1923a: 179. Holotype. Brazil: Bahia, on Psidium guajava. 

Trialeurodes dubienus (Bondar); Bondar 1928b: 32. 

Disiphon dubienus (Bondar); Russell 1993: 464. 

Distribution: 2: Brazil, Peru, Venezuela. 


Disiphon russellae Martin 2005 

Disiphon russellae Martin 2005b: 43. Holotype puparium, Belize, CFR, 6.xi.1994, J. H. Martin, on Piper sp., 

BMNH. 


Host: Lauraceae: Nectandra sp.; Piperaceae: Piper sp.; Rubiaceae: Chiococca sp. 

328

Distinctaleyrodes Dubey & Sundararaj 2006 

Distinctaleyrodes Dubey & Sundararaj 2006: 36. Type species - Distinctaleyrodes setosus Dubey & 

Sundararaj 2006, by original designation. 



setosus x 

Total = 1 species 0 0 0 0 0 1 0 0 0 

Distinctaleyrodes setosus Dubey and Sundararaj 2006 

Distinctaleyrodes setosus Dubey & Sundararaj 2006: 36. Holotype puparium, India, Karnataka, Kudremukh, 

7.iii.2001, A. K. Dubey, on Derris sp., FRI. 


Host: Fabaceae: Derris sp. 

329

Dothioia Dumbleton 1961 

Dothioia Dumbleton 1961a: 124. Type species. Dothioia bidentatus Dumbleton 1961, by monotypy. 



bidentatus x 

Total = 1 species 0 0 0 0 0 0 0 1 0 

Dothioia bidentatus Dumbleton 1961 

Dothioia bidentatus Dumbleton 1961a: 124. Holotype. New Caledonia: on unidentified tree, CORSTOM. 



330

Dumbletoniella Jesudasan & David 1990 

Dumbletoniella Jesudasan and David 1990: 5. Type species. Aleuroclava eucalypti Dumbleton 1957, by 

monotypy. 

Ryoichiella Jesudasan and David 1990: 9. Type species. Aleuroclava ellipticae Dumbleton 1956, by 

monotypy; Martin 1999: 73. 

References: Jesudasan and David (1990); Martin (1999). 



callistemoni x 

ellipticae x 

eucalypti x 

pittospori x 

rotunda x 

xanthorrhoeae x 

Total = 6 species 0 0 0 0 0 0 5 1 0 

Dumbletoniella callistemoni Martin 1999 

Dumbletoniella callistemoni Martin 1999: 74. Holotype puparium. Australia: on Callistemon shiresii, ANIC. 


Host: Myrtaceae: Callistemon shiresii. 

Dumbletoniella ellipticae (Dumbleton) 1956 

Aleuroclava ellipticae Dumbleton 1956b: 167. Holotype puparium, Australia, NSW, Botany, 1895, Froggatt, 

on Styphelia (Monotoca) elliptica; ANIC. 

Ryoichiella ellipticae (Dumbleton); Jesudasan and David 1990: 9. 

Dumbletoniella ellipticae (Dumbleton); Martin 1999: 73. 


Host: Epicridaceae: Monotoca elliptica; Styphelia (Monotoca) elliptica. 

331

Dumbletoniella eucalypti (Dumbleton) 1957 

Aleuroclava eucalypti Dumbleton 1957: 159. Holotype. New Zealand: on Eucalyptus globulus, ADSIR. 

Dumbletoniella eucalypti (Dumbleton); Jesudasan and David 1990: 5 

Distribution: 8: New Zealand 

Host: Myrtaceae: Eucalyptus globulus. 

Dumbletoniella pittospori Martin & Carver 1999 

Dumbletoniella pittospori Martin & Carver 1999, in Martin 1999: 76. Holotype puparium. Australia: on 

Pittosporum phylliraeoides, ANIC. 


Host: Pittosporaceae: Pittosporum phylliraeoides. 

Dumbletoniella rotunda Martin & Carver 1999 

Dumbletoniella rotunda Martin & Carver 1999, in Martin 1999: 77. Holotype puparium. Australia: on 

Eucalyptus leucoxylon, ANIC. 

Distribution: 7: Australia 

Host: Myrtaceae: Eucalyptus luecoxylon. 

Dumbletoniella xanthorrhoeae Martin 1999 

Dumbletoniella xanthorrhoeae Martin 1999: 78. Holotype puparium. Australia: on Xanthorrhoea australis, 

ANIC. 


Host: Xanthorrhoeaceae: Xanthorrhoea australis. 

332

Editaaleyodes David 2005 

Editaaleyodes David 2005: 317. Type species - Editaaleyrodes indicus David 2005, by monotypy. 



indicus x 

Total = 1 species 0 0 0 0 0 1 0 0 0 

Editaaleyrodes indicus David 2005 

Editaaleyrodes indicus David 2005: 317. Holotype puparium, India, Vandalur, 5.viii.1990, P. Megananathan, 

ex. Dichrostachys cineraeae, IARI. 


Host: Fabaceae: Dichrostachys cineraeae. 

333

Extensaleyrodes Bink-Moenen 1983 

Extensaleyrodes Bink-Moenen 1983. Type species - Extensaleyrodes falcata Bink-Moenen 1983, by 

designation. 



akureensis x 

falcata x 

Total =2 species 0 0 0 2 0 0 0 0 0 

Extensaleyrodes akureensis (Mound) 1965 

Dialeurolonga akureensis Mound 1965c: 148-149. Holotype and 8 paratypes. Nigeria: Akure, i.1957, V. F. 

Eastop, on unidentified tree, BMNH. 



Extensaleyrodes falcata Bink-Moenen 1983 

Extensaleyrodes falcata Bink-Moenen 1983: 141. Holotype female, Chad, Boro, 6.xii.1973, FAB & RMBM, 

ex. Isoberlinia doka, BMNH. 

Distribution: 4: Chad.. 


334

Fascaleyrodes Bink-Moenen 1983 

Fascaleyrodes Bink-Moenen 1983: 143. Type species - Fascaleyrodes rara Bink-Moenen, by monotypy and 





palmae x 

rara x 

Total = 2 species 0 0 0 2 0 0 0 0 0 

Fascaleyrodes palmae (Gameel) 

Tetraleurodes palmae Gameel 1968: 153. Holotype. Sudan: on Elaeis guineensis, GRF 

Fascaleyrodes palmae (Gameel); Bink-Moenen 1983: 146. 

Distribution: 4: Sudan. 

Host: Arecaceae: Elaeis guineensis. 

Fascaleyrodes rara Bink-Moenen 1983 

Fascaleyrodes rara Bink-Moenen 1983: 144. Holotype puparium, Bentam, 29.xii.1973, FAB & RMBM, on 

Hexalobus monopetalus, BMNH. 


Host: Annonaceae: Hexalobus monopetalus; Sapindaceae: Paullinia pinnata. 

335

Filicaleyrodes Takahashi 1962 

Filicaleyrodes Takahashi 1962: 100. Type species. Filicaleyrodes bosseri Takahashi 1962, by monotypy. 



bosseri x 

williamsi x x 

sp. x 

Total = 2+ species 1 0 1 2 0 0 0 0 0 

Filicaleyrodes bosseri Takahashi 1962 

Filicaleyrodes bosseri Takahashi 1962: 101. Syntypes. Madagascar: on undetermined fern, HBL. 


Host: on undetermined fern. 

Filicaleyrodes williamsi (Trehan) 1938 

Trialeurodes williamsi Trehan 1938: 186. Lectotype (designated Mound & Halsey 1978: 163). England: on 

Oleandra africana, BMNH. 

Filicaleyrodes williamsi (Trehan); Mound 1966: 416. 

Distribution: 1: USA (FL); 3: England; Hungary. 

Host: Aspidiaceae: Dryopteris flaccida, Nephrodium confluens, Stenosema aurita; Dryopteridaceae: 

Polystichum falcatum, Oleandra africana, Diplazium proliferum. 


Tanzania: on Pteris usumbarensis, BMNH 

Distribution: 4: Tanzania. 

Host: Pteris usumbarensis. 

Reference: Mound and Halsey 1978: 163. 

Filicaleyrodes sp. 

336

Fippataleyrodes Sundararaj and David 1992 

Fippataleyrodes Sundararaj & David 1992a: 15. Type species. Fippataleyrodes indica Sundararaj & David, 

1992, by designation 

References: Dubey & Sundararaj 2005 - Key to species. 



cinnamomi x 

indica x 

litseae x 

multipori x 

yellapurensis x 

Total = 5 species 0 0 0 0 0 5 0 0 0 

Fippataleyrodes cinnamomi Dubey & Sundararaj 2005 

Fippataleyrodes cinnamomi Dubey & Sundararaj 2005: 204. India: Karnataka, 10.ix.2001, A. K. Dubey, on 

Cinnamomum malabatrum, deposited Dehradun, India. 


Host: Lauraceae: Cinnamomum malabatrum. 

Fippataleyrodes indica Sundararaj & David 1992 

Fippataleyrodes indica Sundararaj & David 1992a: 16. Holotype, India, Tamil Nadu, 10.i.1988, B. V. David, 

on Ficus racemosa, USNM (NHM, IARI, ZSI). 


Host: Dipterocarpaceae: Nothopegia sp.; Lauraceae: Litsea sp.; Moraceae: Ficus racemosa. 

Fippataleyrodes litseae Sundararaj & David 1992 

Fippataleyrodes litseae Sundararaj & David 1992a: 17. Holotype, India, Maharashtra, Mahableshwar, 

28.iii.1987, B. V. David, on Litsea stocksii, USNM (NHM, IARI, ZSI). 


Host: Lauraceae: Litsea stocksii. 

337

Fippataleyrodes multipori Dubey & Sundararaj 2005 

Fippataleyrodes multipori Dubey & Sundararaj 2005: 205. Holotype puparium, India: Karnataka, 11.x.2002, 

A. K. Dubey, on unidentified plant, deposited Dehradun, India. 



Fippataleyrodes yellapurensis Dubey & Sundararaj 2005 

Fippataleyrodes yellapurensis Dubey & Sundararaj 2005: 2-6. Holotype puparium, India, Karnataka, 

11.x.2001, A. K. Dubey, on an unidentified plant, deposited IN-DEH. 



338

Gagudjuia Martin 1999 

Gagudjuia Martin 1999: 79. Type species. Gagudjuia allosyncarpiae Martin 1999, by monotypy. 



allosyncarpiae x 

Total = 1 species 0 0 0 0 0 0 1 0 0 

Gagudjuia allosyncarpiae Martin 1999 

Gagudjuia allosyncarpiae Martin 1999: 80. Holotype puparium. Australia: on Allosyncarpia ternata, ANIC. 


Host: Fabaceae: Alysicarpus ternata. 

339

Gomenella Dumbleton 1961 

Gomenella Dumbleton 1961a: 126. Type species. Gomenella multipora Dumbleton 1961, by original 

designation. 



dryandrae x 

multipora x 

reflexa x 

Total = 3 species 0 0 0 0 0 0 1 2 0 

Gomenella dryandrae (Takahashi) 1950 

Dialeurodes dryandrae Takahashi 1950: 85. Syntypes. Australia: on Dryandra floribunda, BMNH. 

Gomenella dryandrae (Takahashi); Martin 1999: 81. 


Host: Proteaceae: Dryandra floribunda. 

US Port Interceptions: Australia on Dryandra formosa, Dryandra sp. 

Gomenella multipora Dumbleton 1961 

Gomenella multipora Dumbleton 1961a: 126. Holotype. New Caledonia: on unidentified tree, CORSTOM. 



Gomenella reflexa Dumbleton 1961 

Gomenella reflexa Dumbleton 1961a: 127. Holotype. New Caledonia: on Maxwellia sp., CORSTOM. 


Host: Bombaceae: Maxwellia sp. 

340

Harpaleyrodes Bink-Moenen 1983 

Harpaleyrodes Bink-Moenen 1983: 146. Type species - Harpaleyrodes tuberculata Bink-Moenen 1983, by 

monotypy and original description. 



tuberculata x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Harpaleyrodes tuberculata Bink-Moenen 1983 

Harpaleyrodes tuberculata Bink-Moenen 1983: 146. Holotype puparium, Chad, Bebedjia, 17.vi.1973, FAB & 

RMBM, on Combretum hypopilinum, BHNM. 

Distribution: 4: Chad, Sudan. 

Host: Combretaceae: Combretum glutinosum, Combretum hypopilinum. 

341

Hesperaleyrodes Sampson 1943 

Hesperaleyrodes Sampson 1943: 213. Type species. Hesperaleyrodes michoacanensis Sampson 1943, by 

monotypy. 

Comment: This species is only known from Mexico on oak. 



michoacanensis x 

Total = 1 species 0 1 0 0 0 0 0 0 0 

Hesperaleyrodes michoacanensis Sampson 1943 

Hesperaleyrodes michoacanensis Sampson 1943: 213. Holotype. Mexico: on Quercus sp., UCD. 



342

Heteraleyrodes Takahashi 1942 

Heteraleyrodes Takahashi 1942d: 103. Type species. Heteraleyrodes bambusae Takahashi 1942, by 

monotypy. 

Comment: This species in only from south-east Asia on bamboo. 



bambusae x 

bambusicola x 

Total = 2 species 0 0 0 0 0 2 0 0 0 

Heteraleyrodes bambusae Takahashi 1942 

Heteraleyrodes bambusae Takahashi 1942d: 104. Syntypes. Thailand: on bamboo [?Bambusa sp.], TARI. 



Heteraleyrodes bambusicola Takahashi 1951 

Heteraleyrodes bambusicola Takahashi 1951c: 4. Holotype. Malaya: on bamboo [?Bambusa sp.], BMNH. 



343

Heterobemisia Takahashi 1957 

Heterobemisia Takahashi 1957: 19. Type species. Heterobemisia alba Takahashi 1957, by monotypy. 



alba x 

Total = 1 species 0 0 0 0 1 0 0 0 0 

Heterobemisia alba Takahashi 1957 

Heterobemisia alba Takahashi 1957: 19. Syntypes. Japan: on Itea japonica, HBL. 


Host: Saxifragaceae: Itea japonica. 

344

Indoaleyrodes David & Subramaniam 1973 

Indoaleyrodes David & Subramaniam 1973: 199. Type species. Indoaleyrodes pustulatus David & 

Subramaniam 1973, by monotypy. [synonymized with Dialeurodes laos Takahashi 1942h:329 by 

Martin 1985: 327]. 



glochidioni x 

laos x 

pseudoculatus x 

reticulata x 

sp. x 

Total = 4+ species 0 0 0 0 0 1 3+ 0 0 

Indoaleyrodes glochidioni Martin & Carver 1999 

Indoaleyrodes glochidioni Martin & Carver 1999, in Martin 1999: 82. Holotype puparium. Australia: on 

Glochidion lobocarpum, ANIC. 


Host: Euphorbiaceae: Glochidion lobocarpum. 

Indoaleyrodes laos (Takahashi) 1942 

Dialeurodes laos Takahashi 1942h: 329. Syntypes. Thailand: on ?Homonoia sp. I 

Indoaleyrodes laos (Takahashi); Martin 1985: 327. 

Indoaleyrodes pustulatus David & Subramaniam 1973: 199. Syntypes. India: causing pit galls on Morinda 

tinctoria leaves, BMNH; synonymy according to Martin 1985: 327. 

Distribution: 6: India; Thailand. 

Host: Combretaceae: Terminalia catappa; Euphorbiaceae: Homonoia sp.?; Rubiaceae: Morinda tinctoria. 

Indoaleyrodes pseudoculatus Martin 1985 

Indoaleyrodes pseudoculatus Martin 1985: 328. Holotype pupal case. Papua New Guinea: on Syzygium sp., 

BMNH. 

Distribution: 7: Papau New Guinea. 


345

Indoaleyrodes reticulata (Dumbleton) 1961 

Parabemisia reticulata Dumbleton 1961a: 134. Holotype. New Caledonia: on unidentified host, CORSTOM. 

Indoaleyrodes reticulata (Dumbleton); Martin 1985: 328. 



Indoaleyrodes sp. 

Collected in Sulawesi on an unidentified plant (Martin 1988: 79). 

346

Juglasaleyrodes Cohic 1966 

Juglasaleyrodes Cohic 1966b: 58. Type species. Juglasaleyrodes orstomensis Cohic 1966, by monotypy. 

Reference: Bink-Moenen (1983) - description and illustration of genus. 



orstomensis x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Juglasaleyrodes orstomensis Cohic 1966 

Juglasaleyrodes orstomensis Cohic 1966b: 59. Syntypes. Congo: on Triclisia gilletii. 


Host: Menispermaceae: Triclisia gilletii. 

347

Keralaleyrodes Meganathan & David 1994 

Keralaleyrodes Meganathan & David 1994: 41. Type species. Keraleyrodes indicus Meganathan & 

Vasantharaj 1994, by monotypy. 



indicus x 

Total = 1 species 0 0 0 0 0 1 0 0 0 

Keraleyrodes indicus Meganathan & David 1994 

Keralaleyrodes indicus Meganathan & David 1994: 41. Holotype. India: on Cinnamomum malabathrum. 


Host: Lauraceae: Cinnamomum malabathrum. 

348

Laingiella Corbett 1926 

Laingiella Corbett 1926: 283. Type species. Laingiella bambusae Corbett 1926, by monotypy. 



bambusae x 

Total = 1 species 0 0 0 0 0 1 0 0 0 

Laingiella bambusae Corbett 1926 

Laingiella bambusae Corbett 1926: 283. Syntypes. Sri Lanka: on bamboo [?Bambusa sp.],. USNM 



349

Leucopogonella Dumbleton 1961 

Leucopogonella Dumbleton 1961a: 129. Type species. Leucopogonella sinuata Dumbleton 1961, by original 

designation. 

Comment: This genus is only known to occur on Leucopogon in New Caledonia. 



apectenata x 

pallida x 

simila x 

sinuata x 

Total = 4 species 0 0 0 0 0 0 0 4 0 

Leucopogonella apectenata Dumbleton 1961 

Leucopogonella apectenata Dumbleton 1961a: 129. Holotype. New Caledonia: on Leucopogon sp., 

CORSTOM. 



Leucopogonella pallida Dumbleton 1961 

Leucopogonella pallida Dumbleton 1961a: 130. Holotype. New Caledonia: on Leucopogon sp., CORSTOM. 



Leucopogonella simila Dumbleton 1961 

Leucopogonella simila Dumbleton 1961a: 131. Holotype. New Caledonia: on Leucopogon sp., CORSTOM. 



Leucopogonella sinuata Dumbleton 1961 

Leucopogonella sinuata Dumbleton 1961a: 132. Holotype. New Caledonia: on Leucopogon sp., CORSTOM. 



350

Lipaleyrodes Takahashi 1962 

Lipaleyrodes Takahashi 1962: 100. Type species. Lipaleyrodes phyllanthi Takahashi 1962, by monotypy. 

Reference: David & Thenmozhi (1995) - key to Indian species. 



atriplex x x 

breyniae x 

crossandreae x 

emiliae x 

euphorbiae x 

hargreavesi x 

leguminicola x 

phyllanthi x 

vernoniae x 

sp. Canary Island x 

sp. Australia x 

Total = 8+ species 0 0 2* 2 1 5 1+ 0 0 

Lipaleyrodes atriplex (Froggatt) 1911 

Aleyrodes atriplex Froggatt 1911b: 757. Lectotype (designated by Dumbleton 1956b: 171). Australia: on 

Atriplex sp. 

Lipaleyrodes atriplex (Froggatt); Martin 1999: 83 

Distribution: 3: Iran; 7: Australia. 

Host: Chenopodiaceae: Atriplex sp., Chenopodium foliosum, Chenopodium opulilfolium, Chenopodium 

trigonum, Spinacia deracea. 

Natural enemies: Aphelinidae: Encarsia inaron, Encarsia sp., Eretmocerus aleyrodesii. 

351

Lipaleyrodes breyniae (Singh) 1931 

Trialeurodes breyniae Singh 1931: 49. Syntypes. India: Pusa, Bihar, 11.iv.1929, K. Singh, on Breynia 

rhamnoides. 

Lipaleyrodes breyniae (Singh); Mound and Halsey 1978: 167. 


Host: Euphorbiaceae: Breynia rhamnoides. 

Lipaleyrodes crossandreae David & Subramaniam 1976 

Lipaleyrodes crossandreae David & Subramaniam 1976: 201. Holotype. India: Tamil Nadu, Coimbatore, 

15.xi.1966, B.V. David, on Crossandra undulaefolia. 


Host: Acanthaceae: Crossandra undulaefolia; Amaranthaceae: Achyranthes aspera var. aspera. 

Lipaleyrodes emiliae Chen & Ko 2006 

Lipaleyrodes emiliae Chen & Ko 2006: 32. Holotype puparium, Taiwan, Renwu, 6 VII 2003, C. H. Hsieh, ex. 

Emilia sonchifolia, NTU. 


Host: Asteraceae: Emilia sonchifolia. 

Lipaleyrodes euphorbiae David & Subramaniam 1976 

Lipaleyrodes euphorbiae David & Subramaniam 1976: 202. Holotype. India: Tamil, Nadu, Madurai, 

28.i.1967, B. David, on Euphorbia prostrata, USNM; Abd-Rabou et al. 2005. 


Host: Euphorbiaceae: Euphorbia cyanthophora, Euphorbia hirta, Euphorbia prostrata, Phyllanthus acidus; 

Phyllanthus abnormis, Phyllanthus fraternus. 

Natural enemies: Aphelinidae: Ablerus inquirenda Silvestri 1928, Encarsia lipaleyrodis Krishnan & David 

1996, Encarsia tinctoriae Krishnan & David 1996; Eretmocerus flavus Krishnan & David 1996; 

Eretmocerus adustiscutum Krishnan & David 1996; Eretmocerus flavus Krishnan & David 1996; 

Eulophidae: Euderomphale sp. 

Reference: Jeritta & David 1987. 

Lipaleyrodes hargreavesi (Corbett) 1935 

Trialeurodes hargreavesi Corbett 1935c: 243. Syntypes. Sierra Leone: on Lindernia diffusa. 

Lipaleyrodes hargreavesi (Corbett); Mound 1965c: 158. 

Distribution: 4: Sierra Leon. 

Host: Scrophulariaceae: Lindernia diffusa. 

352

Lipaleyrodes leguminicola (Takahashi) 1942 

Bemisia leguminicola Takahashi 1942b: 169. Syntypes. Thailand: on Leguminosae, TARI. 

Aleyrodes leguminicola (Takahashi); Takahashi 1952: 21. 

Lipaleyrodes leguminicola (Takahashi); Martin 1999: 83. 



Lipaleyrodes phyllanthi Takahashi 1962 

Lipaleyrodes phyllanthi Takahashi 1962: 100. Syntypes. Madagascar: on Phyllanthus sp., HBL. 


Host: Euphorbiaceae: Phyllanthus sp. 

Lipaleyrodes vernoniae David & Thenmozhi 1996 

Lipaleyrodes vernoniae David & Thenmozhi 1995: 346. Holotype pupal case, India, Padappai, 21.i.1993, K. 

Thenmozhi, on Vernonia cinerea; IARI. 


Host: Asteraceae: Vernonia cinerea. 

Lipaleyrodes sp. 

Australia, on Euphorbiaceae: Euphorbia sp.; Natural enemies: Coccinellidae: Cryptolaemus montrouzieri 

Mulsant 1853; Carver & Reid 1996: Australia, on Euphorbiaceae: Euphorbia aaron-rossii; 

Coccinellidae: Harmonia conformis (Boisduval); on Euphorbia cyanthophora, Hemerobiidae: 

Micromus subanticus (Walker) [ref: Carver & Reid 1996]. 

Canary Islands; Aphelinidae: Encarsia hispida, Encarsia noahi [ref: Hernandez et al. 2003: 70]. 

Australia, on Euphorbiaceae: Chamaesyce hirta parasitized by Aphelinidae: Encarsia adusta; on 

Convolvulaceae: Convolvulus sp. parasitized by Encarsia inaron [ref: Schmidt et al. 2001: 371]. 

353

Malayaleyrodes Corbett 1935 

Malayaleyrodes Corbett 1935b: 843. Type species. Malayaleyrodes lumpurensis Corbett 1935, by monotypy. 

Comment: Only known to occur on Ficus from south-east Asia. 



lumpurensis x 

Total = 1 species 0 0 0 0 0 1 0 0 0 

Malayaleyrodes lumpurensis Corbett 1935 

Malayaleyrodes lumpurensis Corbett 1935b: 843. Syntypes. Malaya: on Ficus sp., lost? 

Distribution: 6: Cambodia; Malaya; Malaysia. 


354

Marginaleyrodes Takahashi 1961 

Marginaleyrodes Takahashi 1961: 338. Type species. Marginaleyrodes ixorae Takahashi 1961, by monotypy. 

Comment: This genus is only known to occur in Madagascar. 



fanalae x 

fenestrata x 

ixorae x 


tsinjoarivona x 

Total = 5 species 0 0 0 5 0 0 0 0 0 

Marginaleyrodes fanalae (Takahashi) 1951 

Aleurotrachelus fanalae Takahashi 1951a: 379. Syntypes. Madagascar: on "fanala", HBL. 

Marginaleyrodes fanalae (Takahashi); Takahashi 1961: 338. 


Host: on fanala". 

Marginaleyrodes fenestrata (Takahashi) 1955 

Tetraleurodes fenestrata Takahashi 1955a: 416. Syntypes. Madagascar: on unidentified host, MNHNP. 

Marginaleyrodes fenestrata (Takahashi); Takahashi 1961: 338. 



Marginaleyrodes ixorae Takahashi 1961 

Marginaleyrodes ixorae Takahashi 1961: 338. Syntypes. Madagascar: on Ixora sp., MNHNP. 



Marginaleyrodes madagascariensis (Takahashi) 1951 

Aleuroplatus madagascariensis Takahashi 1951a: 378. Syntypes. Madagascar: on "banaraka", MNHNP. 

355

Marginaleyrodes madagascariensis (Takahashi); Takahashi 1961: 338. 


Host: on "banaraka". 

Marginaleyrodes tsinjoarivona (Takahashi) 1955 

Tetraleurodes tsinjoarivona Takahashi 1955a: 424. Syntypes. Madagascar: on unidentified host, MNHNP. 

Marginaleyrodes tsinjoarivona (Takahashi); Takahashi 1961: 338. 



Transfers 

Marginaleyrodes angolensis Cohic 1966 - Rugaleyrodes (Bink-Moenen 1983: 163) 

356

Massilieurodes Goux 1939 

Aleuroplatus (Massilierodes) Goux 1939: 80. Type species. Aleuroplatus (Massileurodes) setiger, by 

monotypy; Mound and Halsey 1978: 43. 

Massileiurodes Goux; Jensen 2001: 292. 



alabamensis x 

americanus x Int 

chittendeni x x 

curiosa x 

euryae x 

fici x 

formosensis x 

homonoiae x 

kirishimensis x 

monticola x 

multipori x x 

myricae x 

rarasana x 

sakaki x 

setiger x 

Total = 16 species 5 0 3 6 4 1 0 0 0 

Massilieurodes alabamensis Jensen 2001 

Massilieurodes alabamensis Jensen 2001: 293. Holotype. USA: Alabama, on Prunus sp., USNM. 

Distribution: 1: USA (AL, GA). 

Host: Aquifoliaceae: Ilex sp.; Betulaceae: Carpinus caroliniana; Ebenaceae: Diospyros virginiana; Ericaceae: 

Gaylussacia frondosa, Kalmia latifolia, Vaccinium darrowii, Vaccinium sp.; Hamamelidaceae: 

Liquidambar styraciflua; Magnoliaceae: Liriodendron tulipifera; Nyssaceae: Nyssa sylvatica, Nyssa 

sp.; Oleaceae: Forsythia sp., Fraxinus caroliniana, Fraxinus profunda; Rosaceae: Prunus sp.; 

Symplocaceae: Symplocos tinctoria. 

357

Massilieurodes americanus Jensen 2001 

Massilieurodes americanus Jensen 2001: 295. Holotype. USA: Maryland, Beltsville, on Diospyros virginiana. 

Distribution: 1: USA (AL, DC, DE, FL, GA, LA, MA, MD, NC, NJ, SC, TN, TX, VA); 3: United Kingdom. 

Host: Aquifoliaceae: Ilex sp.; Betulaceae: Carpinus caroliniana; Ebenaceae: Diospyros virginiana. 

Ericaceae: Gaylussacia frondosa, Kalmia latifolia, Vaccinium darrowii, Vaccinium sp., 

Hamamelidaceae: Liquidambar styraciflua; Magnoliaceae: Liriodendron tulipifera; Nyssaceae: Nyssa 

sp., Nyssa sylvatica; Oleaceae: Forsythia sp, Fraxinus caroliniana, Fraxinus profunda; Symplocaceae: 

Symplocos tinctoria. 

Massilieurodes chittendeni (Laing) 1928 

Dialeurodes chittendeni Laing 1928: 228. Lectotype (designated by Mound & Halsey 1978: 132). England: on 

Rhododendron sp., BMNH. 

Massilieurodes chittendeni (Laing); Jensen 2001: 300. 

Distribution: 1: Canada, USA (DC, MD, NY, PA; TN, WA); 3: Belgium, England; Finland; Italy, 

Netherlands, United Kingdom. 

Host: Ericaceae: Rhododendron sp., Rhododendron maximum. 

Natural enemies: Aphelinidae: Encarsia formosa. 

Massilieurodes curiosa Jensen 2001 

Massilieurodes curiosa Jensen 2001: 301. Holotype. USA: Florida, Fanning Springs, on Viburnum obovatum, 

USNM. 


Host: Caprifoliaceae: Viburnum obovatum; Ericaceae: Lyonia lucida; Rubiaceae: Gardenia augusta. 

Massilieurodes euryae (Takahashi) 1940 

Dialeurodes euryae Takahashi 1940a: 27. Syntypes. Taiwan: on Eurya glaberrima. 

Massilieurodes euryae (Takahashi); Jensen 2001: 292; Suh et al. 2006. 

Distribution: 5: Korea; Taiwan. 

Host: Buxaceae: Buxus microphylla; Theaceae: Eurya glaberrima. 

Massilieurodes fici (Takahashi) 1932 

Dialeurodes (Rhachisphora) fici Takahashi 1932: 19. Syntypes. Taiwan: Sankyaku near Taihoku, on Ficus 

vasculosa, Celtis sinensis and Gagara ailanthoides, TARI. 

Rhachisphora fici (Takahashi); Mound and Halsey 1978: 186. 

358

Massilieurodes fici (Takahashi); Jensen 2001: 292. 

Distribution: 5: Japan; Taiwan; 6: Pakistan. 

Host: Euphorbiaceae: Glochidion hongkongensis, Bischofia javanica; Moraceae: Ficus vasculosa; 

Myrsinaceae: Ardisia sp.; Rutaceae: Zanthoxylum ailanthoidies, Zanthoxylum sp., Citrus sinensis; 

Theaceae: Adinandra sp.; Ulmaceae: Celtis sinensis. 

Natural enemies: Aphelinidae: Encarsia perflava, Encarsia lutea, Encarsia inaron, Encarsia azimi, Encarsia 

ancistrocera. 

Massilieurodes formosensis (Takahashi) 1933 

Dialeurodes (Gigaleurodes) formosensis Takahashi 1933: 1933. Syntypes. Taiwan: on unidentified host, 

TARI. 

Massilieurodes formosensis (Takahashi); Jensen 2001: 292. 

Distribution: 5: Hong Kong; Japan; Taiwan. 

Host: Aquifoliaceae: Ilex crenata; Araliaceae: Schefflera sp.; Caprifoliaceae: Viburnum tinus, 

Viburnum sp.; Ericaceae: Pieris japonica; Euphorbiaceae: Homonoia riparia; Fabaceae: Cassia sp.; 

Magnoliaceae: Magnolia compressa, Magnolia kobus; Myrsinaceae: Maesa sp.; Oleaceae: Fraxinus 

sp.; Rutaceae: Phellodendron amurense; Styracaceae: Styrax obassia, Styrax japonicus; 

Symplocaceae: Symplocos chinensis; Urticaceae: Boehmeria densiflora. 

Natural enemies: Aphelinidae: Encarsia sophia, Encarsia lutea. 

Massilieurodes homonoiae (Jesudasan & David) 1991 

Pealius homonoiae Jesudasan and David 1991: 319. Holotype pupal case. India: on Homonoia riparia, IDAV. 

Massilieurodes homonoiae (Jesudasan & David); Sundararaj & David 2003: 371. 

Dialeurodes (Gigaleurodes) splendens Regu and David 1993. Holotype India, Tamil Nadu, Rajapalayam, 

11.vi.1989, K. Regu, on Homonoia riparia, in IARI.; synonymy according to Sundararaj & David 

2003: 371. 

Massilieurodes splendens (Regu and David); Jensen 2001: 292 


Host: Arecaceae: Phoenix humilis; Euphorbiaceae: Homonoia riparia. 

Massilieurodes kirishimensis (Takahashi) 1963 

Dialeurodes kirishimensis Takahashi 1963: 49. Syntypes. Japan: on Cleyera japonica. 

Massilieurodes kirishimensis (Takahashi); Jensen 2001: 292. 


Host: Theaceae: Cleyera japonica. 

US Port Interception: Mexico on Garcinia sp. 

359

Massilieurodes monticola (Takahashi) 1934 

Dialeurodes monticola Takahashi 1934b: 42. Syntypes. Taiwan: on Daphniphyllum sp., TARI. 

Massilieurodes monticola (Takahashi); Jensen 2001: 292. 


Host: Daphniphyllaceae: Daphniphyllum sp. 

Massilieurodes multipori (Takahashi) 1932 

Dialeurodes multipori Takahashi 1932:42. Syntypes. Taiwan: on Daphniphyllum glaucescens. 

Gigaleurodes multipori (Takahashi); Sundararaj & David 1994a: 328. 

Massilieurodes multipori (Takahashi); Jensen 2001: 292. 


Host: Daphniphyllaceae: Daphniphyllum glaucescens. 

Massilieurodes myricae Jensen 2001 

Massilieurodes myricae Jensen 2001: 305. Holotype pupal case. USA: Florida, Oslo, on Myrica cerifera, 

USNM. 


Host: Myricaceae: Myrica punula, Myrica cerifera. 

Massilieurodes rarasana (Takahashi) 1934 

Dialeurodes rarasana Takahashi 1934b: 4. Syntypes. Taiwan: on ?Lithocarpus sp., TARI 

Massilieurodes rarasana (Takahashi); Jensen 2001: 292. 


Host: Araceae: Symplocarpus sp.; Fagaceae: Lithocarpus sp.? 

Massilieurodes sakaki (Takahashi) 1958 

Dialeurodes sakaki Takahashi 1958: 67. Syntypes. Japan: on Sakakia (Cleyera) ochnacea [=Cleyera japonica 

Thunb.], HBL. 

Massilieurodes sakaki (Takahashi); Jensen 2001: 292 


Host: Theaceae: Cleyera japonica. 

360

Massilieurodes setiger (Goux) 1939 

Aleuroplatus (Massilieurodes) setiger Goux 1939: 81. Syntypes. France: on Viburnum tinus. 

Aleuroplatus setiger Goux; Mound and Halsey 1978: 55. 

Massilieurodes setiger (Goux); Jensen 2001: 292. 

Distribution: 3: France. 

Host: Caprifoliaceae: Viburnum tinus. 

361

Metabemisia Takahashi 1963 

Metabemisia Takahashi & Mamet 1963: 52. Type species. Metabemisia distylii Takahashi 1963, by 

monotypy. 



distylii x 

filicis x 

palawana x 

Total = 3 species 0 0 1 0 1 1 0 0 0 

Metabemisia distylii Takahashi & Mamet 1963 

Metabemisia distylii Takahashi & Mamet 1963: 52. Holotype. Japan: on Distylium racemosum, HBL. 


Host: Hamamelidaceae: Distylium racemosus. 

Metabemisia filicis Mound 1967 

Metabemisia filicis Mound 1967: 32. Holotype. Scotland: on Dryopteris sp., BMNH. 

Distribution: 3: England; Scotland. 

Host: Dryopteridaceae: Dryopteris sp., Nephrolepis sp.; Polypodiaceae: Davallia sp. 

Pteridaceae: Pteris togoensis?; Thelypteridaceae: Cyclosorus dentatus?, ferns. 

Metabemisia palawana Martin 2001 

Metabemisia palawana Martin 2001, in Martin & Camus 2001: 1. Holotype. Philippines (Palawan), on fern. 

Distribution: 6: Philippines (Palawan) 

Host: a fern. 

362

Metaleyrodes Sampson 1943 

Metaleyrodes Sampson 1943: 235. Type species. Aleyrodes oceanica Takahashi 1939, by monotypy. 



oceanica x 

Total = 1 species 0 0 0 0 0 0 0 1 0 

Metaleyrodes oceanica (Takahashi) 1939 

Aleyrodes oceanica Takahashi 1939b: 235. Syntypes. Caroline Islands: on unidentified host, TARI. 

Metaleyrodes oceanica (Takahashi); Sampson 1943: 210. 

Distribution: 8: Caroline Islands. 


363

Minutaleyrodes Jesudasan and David 1990 

Minutaleyrodes Jesudasan and David 1990. Type species Dialeurodes minuta Singh 1931, by original 

designation. 

Reference: Jesudasan and David 1990. 



cherasensis x 

indicus x 

kolliensis x 

minuta Int Int x Int Int 

suishanus x 

Total = 5 species 1 1 0 0 1 4 0 1 1 

Minutaleyrodes cherasensis (Corbett) 1935 

Aleurotuberculatus cherasensis Corbett 1935b: 824. Syntypes. Malaya: on Psidium guajava. 

Aleuroclava cherasensis (Corbett); Martin 1999. 

Minutaleyrodes cherasensis (Corbett); Jesudasan and David 1990. 



Note: This species maybe synonymous of M. minutus; Corbett says that it differs from M. minutus by having 

tessellated dorsum and from A. suishanus by having 3 pairs of tubercles on subdorsum of abdomen. 

Minutaleyrodes indicus Meganathan & David 1994 

Minutaleyrodes indicus Meganathan & David 1994: 17. Holotype. India: on Palaquium ellipticum; Dubey & 

Sundararaj 2005d: 253. 


Host: Caprifoliaceae: Viburnum coriaceum; Sapotaceae: Palaquium ellipticum. 

Minutaleyrodes kolliensis (David) 1977 

Aleurotuberculatus kolliensis David 1977: 89. Holotype pupal case. India: Tamil Nadu, Salem, Dt, Kolli Hills, 

364

20.iii.1972, B.V. David, on Neolitsea sp, IDAV. 

Aleuroclava kolliensis (David); Martin 1999. 

Minutaleyrodes kolliensis (David); Jesudasan and David 1990: 1 


Host: Lauraceae: Neolitsea sp. 

Minutaleyrodes minuta (Singh) 1931 

Dialeurodes minuta Singh 1931: 42. Syntypes. India: Pusa, on Ixora coccinea, IARI. 

Aleuroclava minutus (Singh); Martin 1999: 31. 

Minutaleyrodes minutus (Singh); Jesudasan and David 1990. 

Distribution: 1: USA (FL); 2: Dominican Republic, Guyana, Mexico, Puerto Rico; 6: Burma, India, 

Philippines, Singapore; 8: Guam; 9: Hawaii. 

Host: Fagaceae: Quercus virginiana; Myrtaceae: Eugenia uniflora; Oleaceae: Jasminum sp.; 

Rubiaceae: Ixora coccinea, Gardenia augusta; Rutaceae: Murraya paniculata. 

Natural enemies: Aphelinidae: Eretmocerus sp. [in the Dominican Republic]. 

Minutaleyrodes suishanus (Takahashi) 1934 

Aleurotuberculatus suishanus Takahashi 1934b: 48. Syntypes. Taiwan: on Lauraceae, TARI. 

Aleuroclava suishanus (Takahashi); Martin 1999: 31. 

Minutaleyrodes suishanus (Takahashi), new comb. 



365

Mixaleyrodes Takahashi 1936 

Mixaleyrodes Takahashi 1936c: 150. Type species. Mixaleyrodes polystichi Takahashi 1936, by monotypy. 



polypodicola x 

polystichi x 

Total = 2 species 0 0 0 0 2 1 0 0 0 

Mixaleyrodes polypodicola Takahashi 1963 

Mixaleyrodes polypodicola Takahashi & Mamet 1963: 56. Syntypes. Taiwan: on Polystichopsis aristata, 

HBL. 


Host: Aspidiaceae: Polystichosis aristata; Polypodiaceae: Polystichum sp. 

Mixaleyrodes polystichi Takahashi 1936 

Mixaleyrodes polystichi Takahashi 1936c: 150. Syntypes. Taiwan: on Polystichum sp. 


Host: Polypodiaceae: Polystichum sp. 

Transfers 

Mixaleyrodes indicus David & Selvakumaran 1987 - transferred to Cohicaleyrodes by David 2005:67. 

366

Nealeyrodes Hempel 1923 

Nealeyrodes Hempel 1923: 1179. Type species. Nealeyrodes bonariensis Hempel 1923, by monotypy. 



bonariensis x 

Total = 1 species 0 1 0 0 0 0 0 0 0 

Nealeyrodes bonariensis Hempel 1923 

Nealeyrodes bonariensis Hempel 1923: 1179. Syntypes. Argentina: on Erygium pandanifolium, MZU. 

Distribution: 2: Argentina. 

Host: Umbelliferae: Eryngium pandanifolium. 

367

Neoaleurodes Bondar 1923 

Neoaleurodes Bondar 1923a: 128. Type species. Neoaleurodes clandestinus Bondar 1923, by monotypy. 



clandestinus x 

Total = 1 species 0 1 0 0 0 0 0 0 0 

Neoaleurodes clandestinus Bondar 1923 

Neoaleurodes clandestinus Bondar 1923a: 128. Syntypes. Brazil: on Miconia sp., MZU 


Host: Melastomataceae: Miconia sp. 

368

Neoaleurotrachelus Takahashi & Mamet 1952 

Neoaleurotrachelus Takahashi & Mamet 1952b: 126. Type species. Neoaleurotrachelus aphloiae Takahashi 

& Mamet 1952, by monotypy. 

Jeannelaleyrodes Cohic 1966a: 47. Type species. Jeannelaleyrodes bertilloni Cohic 1966, by monotypy; 

synonymy according to Bink-Moenen 1983. 

Reference: Bink-Moenen (1983) - Description and key to 2 species (syntypes of N. aphloiae in poor 

condition) 



aphloiae x 

bertilloni x 

graberi x 

sudaniensis x 

Total = 4 species 0 0 0 4 0 0 0 0 0 

Neoaleurotrachelus aphloiae Takahashi & Mamet 1952 

Neoaleurotrachelus aphloiae Takahashi & Mamet 1952b: 126. Syntypes. Madagascar, Ambatatomanga, 

x.1949, R. Paulian, on Aphloia theaeformis, MNHNP. 


Host: Flacourtiaceae: Aphloia theaeformis. 

Neoaleurotrachelus bertilloni (Cohic) 1966 

Jeannelaleyrodes bertilloni Cohic 1966a: 47. Syntypes. Congo: on Tetracera alnifolia. 

Neoaleurotrachelus bertilloni (Cohic); Bink-Moenen 1983: 150. 


Host: Bignoniaceae: Markhamia sessilis; Dilleniaceae: Tetracera alnifolia; Euphorbiaceae: Antidesma 

laciniatum; Loganiaceae: Strychnos variabilis; Meliaceae: Entandrophragma utile, Trichilia 

heudelottii; Ochnaceae: Ochna gilletiana; Rubiaceae: Colletoecema dewevrei, Psychotria 

psychotrichoides. 

369

Neoaleurotrachelus graberi (Cohic) 1968 

Jeannelaleyrodes graberi Cohic 1968a: 46. Syntypes. Chad: on Gardenia jovis-tonantis. 

Neoaleurotrachelus graberi (Cohic); Bink-Moenen 1983: 150. 

Distribution: 4: Chad; Ivory Coast; Sudan. 

Host: Bignoniaceae: Steroespermum kunthianum; Lythraceae: Lawsonia inermis; Rhamnaceae: Ziziphus 

spina-christi; Rubiaceae: Gardenia erubescens, Gardenia jovis-tonantis. 

Neoaleurotrachelus sudanensis Gameel 1968 

Orchamoplatus sudanensis Gameel 1968: 151. Holotype. Sudan, Khartoum, Nile Junction Gardens, 5.xi.1961, 

O. I. Gameel; on Cassia fistula, Sudan FRF. 

Neoaleurotrachelus sudanensis Gameel 1968: 151; Martin 1985: 329. 


Host: Fabaceae: Cassia fistula, Cassia occidentalis. 

370

Neomaskellia Quaintance & Baker 1913 

Neomaskellia Quaintance & Baker 1913: 91. Type species. Aleurodes comata Maskell 1895, by monotypy 

[junior synonym of N. bergii (Signoret 1868]. 

Comment: This Old World genus is known to occur on grasses. 



andropogonis x x 

bergii x x x x x x 

Total = 2 species 0 0 1 1 2 2 1 1 0 

Neomaskellia andropogonis Corbett 1926 

Neomaskellia andropogonis Corbett 1926: 278. Syntypes. Sri Lanka: on Andropogon sp., BMNH. 

Neomaskellia hainanensis Chou & Yan 1988: 244. Holotype puparium, China, Hainan, 19.v.1963, I. Chou; 

synonymy according to Martin & Mound 2007: 37. 

Distribution: 5: China; 6: India; Pakistan; Malaya; Malaysia; Sri Lanka. 

Host: Poaceae: Andropogon sp.; Saccharum arundinaceum, Saccharum officinarum, Saccharum sarra, 

Saccharum vulgare. 

Natural enemies: Aphelinidae: Encarsia tristis [ref: Hayat 1981a: 468); Encarsia tristis [ref: Hayat 1989a: 

66]. 

Neomaskellia bergii (Signoret) 1868 

Aleurodes bergii Signoret 1868: 395. Syntypes. Mauritius: on Saccharum officinarum, ADSIR. 

Neomaskellia bergii (Signoret); Quaintance & Baker 1914: 104. 

Aleurodes comata Maskell 1895: 426. Syntypes. Fiji: on Graminae, ADSIR; synonymy according to Martin 

1999: 85. 

Aleurodes sacchari Maskell 1890b: 171. Syntypes. Fiji: on Saccharum officinarum, ADSIR.; synonymy 

according to Quaintance & Baker 1914: 104 

Distribution: 3: Iran; 4: Central African Republic; Congo; Gambia; Mauritius; Senegal; Sierra Leon; South 

Africa; Sudan; Tanzania; Uganda; Zaire; 5: Japan; Taiwan; 6: India; Thailand; 7: Australia; Java; 8: 

Caroline Islands; Fiji; New Zealand; Solomon Islands. 

Host: Poaceae: Bambusa sp.; Cenchrus ciliaris; Panicum maximum; Paspalum conjugatum. 

Pennisetum purpureum; Saccharum officinarum; Setaria chevaliera; Setaria italica, Sorghum bicolor, 

Sorghum caudatum, Sorghum halepense; Sorghum vulgare. 

371

372 

Natural enemies: Parasitoids- Aphelinidae: Encarsia isaaci, Encarsia tristis, Eretmocerus adustiscutum, 

Eretmocerus delhiensis, Eretmocerus mundus, Eretmocerus neomaskelliae. 

Predators - Formicidae: Anoplolepis longipes; Coccinellidae: Coccinella transversalis, Scymnus sp.; 

Nitidulidae: Cybocephalus sp. 

Transfers 

Neomaskellia eucalypti Dumbleton 1956 - Xenaleyrodes (Martin 1985)

Neopealius Takahashi 1954 

Neopealius Takahashi 1954b: 50. Type species: Neopealius rubi Takahashi 1954, by monotypy. 



rubi x x 

Total = 1 species 0 0 1 0 1 0 0 0 0 

Neopealius rubi Takahashi & Mamet 1954 

Neopealius rubi Takahashi 1954b: 51. Syntypes. Japan: on Rubus microphyllus and other plants, HBL. 

Aleyrodes rosae Korobitsin 1967. Holotype. USSR: Southern Crimea, Yalta, on Rosa canina; 

synonymy according to Bink-Moenen 1991: 321; Martin et al. 2000: 434. 

Bemisia rosae Danzig 1969. Holotype. USSR: Hissar Range, on Rosa exae, ZIN. 

Distribution: 3: Iran, Russia, USSR; 5: Japan. 

Host: Fabaceae: Lespedeza buergei; Labiatae: Phlomis sp., Nepeta rugosa; Lauraceae: Lindera obtusiloba, 

Lindera umbellatum; Rosaceae: Crataegus pentagya, Rosa beggeriana, Rosa canina, Rosa exae, Rosa 

hyrcanus, Rubus microphyllus, Rubus palmatus. 

373

Nigrasialeyrodes Martin & Carver 1999 

Nigrasialeyrodes Martin & Carver 1999 in Martin 1999: 86. Type species. Nigrasialeyrodes convexus Martin 

& Carver 1999, by monotypy 



convexus x 

Total = 1 species 0 0 0 0 0 0 1 0 0 

Nigrasialeyrodes convexus Martin & Carver 1999 

Nigrasialeyrodes convexus Martin & Carver 1999, in Martin 1999: 86. Holotype puparium. Australia: on 

Melaleuca quinquenervia, ANIC. 


Host: Myrtaceae: Melaleuca quinquenervia. 

374

Orchamoplatus Russell 1958 

Aleuroplatus (Orchamus) Quaintance & Baker 1917: 400. Type species. Aleuroplatus (Orchamus) 

mammaeferus Quaintance & Baker 1917, by monotypy. 

Orchamus Quaintance & Baker; full genus by Dumbleton 1956a: 131 

Orchamoplatus Russell 1958: 390. Type species. Aleuroplatus (Orchamus) mammaeferus Quaintance & 

Baker 1917, by monotypy.[As replacement name preoccupied by Ochamus stal (Orthoptera) 



caledonicus x 

calophylli x 

citri x 

dentatus x 

dumbletoni x 

incognitus x 

louiserussellae x 

mammaeferus Int x x x x x x 

montanus x 

niuginii x 

noumeae x 

perdentatus x 

plumensis x 

porosus x 

Total = 14 species 0 1 1 0 1 1 4 11 1 

Orchamoplatus caledonicus (Dumbleton) 1956 

Aleuroplatus (Orchamus) [sic] samoanus Laing; misidentification Williams 1944: 100 

375

376 

Orchamus caledonicus Dumbleton 1956a: 133. Holotype. New Caledonia: on Citrus sp., USNM. 

Orchamoplatus caledonicus (Dumbleton); Russell 1958: 400 


Host: Moraceae: Malaisia tortuosa; Rutaceae: Citrus aurantium Citrus reticulata, Citrus medica. 

Ulmaceae: Celtis sp. 

Natural enemies: Aphelinidae: Encarsia neocala. 

Orchamoplatus calophylli Russell 1958 

Orchamoplatus calophylli Russell 1958: 402. Holotype. Tonga: on Calophyllum sp., USNM. 

Distribution: 8: Tonga. 

Host: Guttiferae: Calophyllum sp. 

Orchamoplatus citri (Takahashi) 1940 

Aleuroplatus citri Takahashi 1940e: 381. Syntypes. Australia: on Citrus limon, TARI. 

Orchamoplatus citri (Takahashi); Russell 1958: 397. 


Host: Rutaceae: Citrus limon, Citrus sp. 

Natural enemies: Parasitoids - Aphelinidae: Cales orchamoplati, Encarsia sp.; Predators: Clambidae: 

Clambus sp., Nitidulidae: Cybocephalus aleyrodiphagus. 

Orchamoplatus dentatus (Dumbleton) 1956 

Orchamus dentatus Dumbleton 1956a: 134. Holotype. New Caledonia: on Myrtaceous plant, CORSTOM. 

Orchamoplatus dentatus (Dumbleton); Russell 1958: 403 



Orchamoplatus dumbletoni (Cohic) 1959 

Orchamus dumbletoni Cohic 1959c: 130. Holotype. New Caledonia: on Homalium arboreum, CORSTOM. 

Orchamoplatus dumbletoni (Cohic); Mound and Halsey 1978: 175. 


Host: Flacourtiaceae: Homalium arboreum; Ulmaceae: Celtis sp. 

Orchamoplatus incognitus (Dumbleton) 1956 

Orchamus incognitus Dumbleton 1956a: 136. Holotype. New Caledonia: on unidentified host, CORSTOM.

Orchamoplatus incognitus (Dumbleton); Russell 1958: 399. 


Host: unknown plant 

Orchamoplatus louiserussellae Martin 1999 

Orchamoplatus louiserussellae Martin 1999: 88. Holotype puparium. Australia: on Syzygium sp., ANIC. 



Orchamoplatus mammaeferus (Quaintance & Baker) 1917 

Aleuroplatus (Orchamus) mammaeferus Quaintance & Baker 1917: 400. Syntypes. Java: Buitenzorg, i.1911, 

R. S. Woglum, on Codiaeum variegatum, USNM. 

Orchamoplatus mammaeferus (Quaintance & Baker); Russell 1958: 393. 

Aleuroplatus (Orchamus) samoanus Laing 1927: 43. Syntypes. Samoa: on croton [Codiaeum variegatum], 

BMNH; synonymy according to Russell 1958: 393. 

Distribution: 2: Bahamas; 3: Iran; 5: Japan; 6: Malaysia; Singapore; 7: Australia; Java; New Guinea; 

Sulawesi; 8: Fiji; Samoa; Marques Islands; Tahiti; 9: Hawaii. 

Host: Convolvulaceae: Convolvulus arvensis, Ipomoea purpurea; Euphorbiaceae: Codiaeum variegatum var. 

pictum, Codiaeum sp.; Myrtaceae: Pimenta caryophylla, Pimenta officinalis; Rutaceae: Citrus 

aurantifolia, Citrus medica, Citrus sinensis, Citrus x paradisi. 

Natural enemies: Coccinellidae: Halmus chalybeus, Serangium maculigerum, Nephaspis oculatus, Delphastus 

pusillus. 

US Port Interceptions: American Samoa on Alyxia oliviformis, Alyxia stellata; Cook Islands on Albizia sp. 

Alyxia elliptica, Alyxia oliviformis, Alyxia sp., Alyxia stellata; Fiji on Codiaeum variegatum; French 

Polynesia on Alyxia sp., Alyxia stellata; Guam on Alyxia oliviformis, Alyxia sp., Ixora sp., Piper betle; 

Hawaii on Alyxia oliviformis, Alyxia sp., Alyxia stellata, Citrus sp., Codiaeum sp., Codiaeum 

variegatum, Croton sp., Etingera sp., Heckeria umbellata, Mentha sp., Moringa sp., Pimenta dioica, 

Piper sp., Plumeria sp., Psidium guajava, Hong Kong on Citrus sp.; Japan on Citrus reticulata; 

Malaysia on Codiaeum sp.; Mexico on Psidium sp.; Micronesia on Alyxia sp.; New Zealand on 

Citrus sp.; Codiaeum sp.; Papua New Guinea on Codiaeum sp.; Singapore on Codiaeum sp., 

Codiaeum variegatum; Tahiti on Alyxia oliviformism, Alyxia stellata, Codiaeum sp., Codiaeum 

variegatum, Plumeria sp.; Tonga on Alyxia oliviformis. 

Orchamoplatus montanus (Dumbleton) 1956 

Orchamus montanus Dumbleton 1956a: 138. Holotype. New Caledonia: on undetermined Cunoniaceous 

plant, CORSTOM. 

Orchamoplatus montanus (Dumbleton); Russell 1958: 398 


377

Host: Araliaceae: Myrita sp.; Cunoniaceae; Epacridaceae: Leucopogon dammarifolius. 

Orchamoplatus niuginii Martin 1985 

Orchamoplatus niuginii Martin 1985: 329. Holotype pupal case. Papua New Guinea: on Calophyllum 

inophyllum, BMNH. 

Distribution: 7: Papua New Guinea. 


Orchamoplatus noumeae Russell 1958 

Aleuroplatus (Orchamus) [sic] samoanus Laing; misid Williams 1944 (in part), Williams 1944: 100. 

Orchamoplatus noumeae Russell 1958: 405. Holotype. New Caledonia: on Citrus sp, USNM. 



Orchamoplatus perdentatus Dumbleton 1961 

Orchamoplatus perdentatus Dumbleton 1961a: 133. Holotype. New Caledonia: on unidentified host, 

CORSTOM. 



Orchamoplatus plumensis (Dumbleton) 1956 

Orchamus plumensis Dumbleton 1956a: 138. Holotype. New Caledonia: on unidentified host, CORSTOM. 

Orchamoplatus plumensis (Dumbleton); Russell 1958: 408 



Orchamoplatus porosus (Dumbleton) 1956 

Orchamus porosus Dumbleton 1956a: 140. Holotype. New Caledonia: on ?Mooria artensis, CORSTOM. 

Orchamoplatus porosus (Dumbleton); Russell 1958: 406. 


Host: Myrtaceae: Mooria artensis? 

Orchamoplatus sudaniensis Gameel 1968 

Orchamoplatus sudaniensis Gameel 1968: 151. Holotype. Sudan: on Cassia fistula, GRF; Bink-Moenen 

378

1983: 152. 


Host: Fabaceae: Cassia occidentalis. 

379

Orientaleyrodes David 1993 

Orientaleyrodes David 1993: 25. Type species. Aleurolobus zeylanicus, by original designation. - 



indicus x 

zeylanicus x 

Total = 2 species 0 0 0 0 0 1 0 0 0 

Orientaleyrodes indicus David 1993 

Orientaleyrodes indicus David 1993: 54. Holotype. India: on Capparis sp. 


Host: Capparidaceae: Capparis sp. 

Orientaleyrodes zeylanicus (Corbett) 1926 

Aleurolobus zeylanicus Corbett 1926: 279. Syntypes. Sri Lanka: on unidentified plant, BMNH. 

Orientaleyrodes zeylanicus (Corbett); David 1993. 



380

Orstomaleyrodes Cohic 1966 

Orstomaleyrodes Cohic 1966a: 52. Type species. Aleuroplatus fimbriae Mound 1965, by monotypy. 



fimbriae x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Orstomaleyrodes fimbriae (Mound) 1965 

Aleuroplatus fimbriae Mound 1965c: 134. Holotype. Nigeria: on Cassia siamea, BMNH. 

Orstomaleyrodes fimbriae (Mound); Cohic 1966a: 52. 

Distribution: 4: Angola; Cameroon; Chad; Congo; Ghana; Ivory Coast; Nigeria. 

Host: Bignoniaceae: Tecoma stans; Combretaceae: Combretum bracteatum; Fabaceae: Baphia nitida, Cassia 

siamea, Newtonia glandulifera; Flacourtiaceae: Caloncoba welwitschii; Sapindaceae: Paullinia 

pinnata; Ulmaceae: Celtis milbraedii. 

381

Papillipes Bink-Moenen 1983 

Papillipes Bink-Moenen 1983: 153. Type species - Papillipes spinifera, by monotypy and original 

designation. 



spinifer x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Papillipes spinifer Bink-Moenen 1983 

Papillipes spinifer Bink-Moenen 1983: 154. Holotype puparium, Chad, Bebedjia, 4.i.1973, FAB & RMBM, 

on Daniellia olivera, BMNH. 


Host: Fabaceae: Andira inermis, Daniellia olivera, Dialium guineese, Isoberlinia doka. 

382

Parabemisia Takahashi 1952 

Parabemisia Takahashi & Mamet1952a: 21. Type species. Parabemisia maculata Takahashi 1952, by 




aceris x 

indica x 

jawani x 

lushanensis x 

maculata x x 

myricae x x x x x x 

mymecophila x 

Total = 7 species 1 1 1 0 3 3 1 2 1 

Parabemisia aceris (Takahashi) 1931 

Bemisia aceri Takahashi 1931a: 204. Syntypes. Taiwan: on Acer sp., TARI. 

Parabemisia aceris (Takahashi); Takahashi 1952a: 23. 


Host: Aceraceae: Acer palmatum, Acer sp. 

Parabemisia indica Meganathan & David 1994 

Parabemisia indica Meganathan & David 1994: 47. Holotype. India: on Nothopegia racemosa, - 


Host: Dipterocarpaceae: Nothopegia racemosa. 

Parabemisia jawani Martin 1985 

Parabemisia jawani Martin 1985: 331. Holotype pupal case. Papua New Guinea: on undetermined woody 

host, BMNH. 



383

Parabemisia lushanensis Ko & Luo 1999 

Parabemisia lushanensis Ko & Luo 1999: 19. Holotype puparium, China, Lushan, 27.x.1996, Z.Y. Luo, ex. 

Acer serrulatum, SIE. 


Host: Aceraceae: Acer serrulatum. 

Parabemisia maculata Takahashi & Mamet 1952 

Parabemisia maculata Takahashi 1952a: 22. Syntypes. Japan: on Acer sp., HBL. 

Distribution: 5: Japan; 6: Malaysia. 

Host: Aceraceae: Acer sp.; Rubiaceae: Gardenia florida; Theaceae: Thea sinensis. 

Parabemisia myricae (Kuwana) 1927 

Bemisia myricae Kuwana 1927: 249. Syntypes. Japan: on Myrica rubra, Morus alba, Citrus spp. and other 

plants, TARI. 

Parabemisia myricae (Kuwana); Takahashi 1952a: 24. 

Distribution: 1: USA (CA, FL, HI); 2: Venezuela; 3: Egypt; Iran, Israel; Italy; Morocco; Spain; Turkey; 5: 

China; Japan; Taiwan; 6: India; 9: Hawaii. 

Host: Asteraceae: Lactuca sp.; Cyperaceae: Cyperus papyrus; Ebenaceae: Diospyros khaki; Elaeocarpaceae: 

Elaeocarpus serratus; Ericaceae: Rhododendron sp.; Fagaceae: Quercus serratus; Juglandaceae: 

Engelhardtia roxburghiana; Lauraceae: Cryptocarya sp.,Machilus sp., Persea americana; Moraceae: 

Ficus carica, Morus alba, Morus sp.; Myricaceae: Myrica rubra; Myrsinaceae: Maesa japonica; 

Myrtaceae: Psidium guajava; Olacaceae: Schoepfia schreberi; Polygonaceae: Coccoloba krugii; 

Rosaceae: Prunus mume, Prunus persica, Prunus triflora; Rubiaceae: Chiococca alba, Gardenia 

augusta, Gardenia sp., Gardenia thunbergia; Rutaceae: Citrus aurantifolia, Citrus aurantium, Citrus 

bigaradia, Citrus limon, Citrus sinensis, Citrus x nobilis, Murraya koenigii, Poncirus trifoliata, 

Wendlandia thrysoidea, X Citrofortunella microcarpa; Salicaceae: Salix babylonica. 

Natural enemies: Parasitoids - Hymenoptera/ Aphelinidae: Encarsia azimi, Encarsia collecta, Encarsia 

hispida, Encarsia lutea, Encarsia meritoria, Encarsia perflava, Encarsia protransvena, Encarsia 

pseudocitrella, Encarsia sophia, Encarsia strenua, Eretmocerus debachi, Eretmocerus furuhashii ; 

Platygastridae: Amitus sp. 

Predators - Acari/Phytoseiidae: Amblyseius swirski, Euseius scutalis, Iphiseiodes degenerans, 

Typhlodromus athiasae; Coleoptera/ Coccinellidae: Adonia variegata, Chilocorus bipustulatus, 

Clitostethus arcuatus, Coelophora pupillata, Exochomus quadripustulatus, Hippodamia sp., Oenopia 

conglobata, Scymnus near pallidicollis, Serangium parcesetosum; Coleoptera/ Nitidulidae: 

Cybocephalus binotatus; Diptera/ Syrphidae: Genus? sp.; Hemiptera/ Anthocoridae: Genus? sp.; 

Miridae: Deraeocoris pallens; Neuroptera/Chrysopidae: Chrysoperla carnea; Coniopterygidae: 

Conwentzia hagani, Conwentzia psociformis, Semidalis aleyrodiformis; Hemerobiidae: Sympherobius 

sanctus. 

US Port Interceptions: Bangladesh on Piper betle; China on Magnolia grandiflora, Magnolia sp., Michelia 

384

385 

sp.; Dominican Republic; Egypt on Vitis sp.; El Salvador on Citrus sp.; Greece on Citrus sp.; 

Guyana on Murraya sp.; Hawaii on Gardenia jasminoides, Piper sp.; India on Areca sp., Murraya 

koenigii, Murraya paniculata, Murraya sp.; Indonesia on Psidium sp.; Malaysia on Murraya sp.; 

Mexico on Citrus sp., Gardenia sp., Persea sp., Punica sp.; Nicaragua on Citrus sp.; Pakistan on 

Murraya koenigii; Puerto Rico on Citrus sp.; Singapore on Murraya koenigii; Spain on Vitis sp.; 

Taiwan on Michelia champaca, Michelia sp.; Trinidad and Tobago on Melia azedarach, Meliaceae: 

Murraya sp.; Turkey on Vitis sp.; Viet Nam on Piper betle. 

Parabemisia myrmecophila Martin 1985 

Parabemisia myrmecophila Martin 1985: 332. Holotype pupal case. Papua New Guinea: on Cryptocarya sp., 

BMNH. 


Host: Lauraceae: Cryptocarya sp. 

Parabemisia sp. 

1) Taiwan, ex Urticaceae: Elatostema lineolatum; Aphelinidae: Encarsia duorunga Hayat 1989 (Chou et al. 

1996b) 

2) Taiwan, Aphelinidae: Encarsia synaptocera Huang & Polaszek 1998 (Huang & Polaszek 1998: 1954).

Paraleurolobus Sampson & Drews 1941 

Paraleurolobus Sampson & Drews 1941: 168. Type species. Paraleurolobus imbricatus Sampson & Drews 




chamaedoreae x 

imbricatus x x 

Total = 2 species 1 1 0 0 0 0 0 0 0 

Paraleurolobus chamaedoreae Russell 1994 

Paraleurolobus chamaedoreae Russell 1994: 540. Holotype puparium, intercepted at USA: Texas, San 

Antonio, (origin Mexico) 6.viii.1970, D. Johnston, on Chamaedorea sp., USNM. 

Distribution: 2: Belize, Costa Rica, Mexico, Nicaragua. 

Host: Arecaceae: Chamaedorea sp.; Moraceae. 

US Port Interception: Mexico on Chamaedorea sp. 

Paraleurolobus imbricatus Sampson & Drews 1941 

Paraleurolobus imbricatus Sampson & Drews 1941: 169. Syntypes. Mexico: on undetermined tree, UCD. 

Distribution: 1: USA (FL); 2: Mexico. 

Host: Arecaceae: Chamaedorea sp., unidentified tree. 

386

Paulianaleyrodes Cohic 1966 

Paulianaleyrodes Cohic 1966b: 64. Type species. Paulianaleyrodes splendens Cohic 1966, by original 

designation. 

Reference: Bink-Moenen (1983) - key to species. 



pauliani x 

splendens x 

tetracerae x 

Total = 3 species 0 0 0 3 0 0 0 0 0 

Paulianaleyrodes pauliani Cohic 1966 

Paulianaleyrodes pauliani Cohic 1966b: 69. Holotype. Angola: on unidentified host. 

Distribution: 3: Angola; Congo. 

Host: on unidentified host; Annonaceae: Uvaria scabrida. 

Paulianaleyrodes splendens Cohic 1966 

Paulianaleyrodes splendens Cohic 1966b: 64. Syntypes. Congo: on Dichapetalum brazzae, BMNH. 


Host: Dichapetalaceae: Dichapetalum brazzae. 

Paulianaleyrodes tetracerae Cohic 1966 

Paulianaleyrodes tetracerae Cohic 1966b: 67. Syntypes. Congo: on Tetracera alnifolia, BMNH. 


Host: Cucurbitaceae: Cogniauxia podolaena; Dilleniaceae: Tetracera alnifolia; Guttiferae: Harungana 

madagascariensis. 

387

Pealius Quaintance & Baker 1914 

Pealius Quaintance & Baker 1914: 99. Type species. Aleyrodes maskelli Bemis 1904, by original designation. 

Corbettella Sampson 1943: 207. Type species. Bemisia artocarpi Corbett 1935, by monotypy; Mound and 

Halsey 1978: 179 

Odontaleyrodes Takahashi 1954: 49. Type species. Aleyrodes akebiae Kuwana 1911, by original designation; 

Martin 1999: 91. 

References: Dubey & Sundararaj 2005a - Key to Indian species. 



akebiae x 

amamianus x 

artocarpi x 

azaleae x x x x x x 

bangkokensis x 

bengalensis x 

cambodiensis x 

cinnamomi x 

cryptus x 

damnacanthi x 

elatostemae x 

elongatus x 

euryae x 

ezeigwi x 

fici x 

indicus x 

kalawi x 

kankoensis x 

kelloggi x 

kongosana x 

liquidambari x 

388


longispinus x 

machili x 

maculatus x 

madeirensis x 

maskelli x 

misrae x 

mitakensis x 

mori x x 

nagerkoilensis x 

nilgiriensis x 

polygoni x 

psychotriae x 

quercus x 

rhododendri x x x 

rubi x 

sairandhryensis x 

schimae x x 

setosus x 

spinosus x 

spinus x 

splendens x 

sutepensis x 

tuberculatus x 

walayarensis x 

Total = 45 species 4 0 4 2 15 24 3 1 1 

Pealius akebiae (Kuwana) 1911 

Aleyrodes akebiae Kuwana 1911: 622. Syntypes. Japan: on Akebia quinata, USNM. 

Odontaleyrodes akebiae (Kuwana); Takahashi 1954: 49. 

389

Pealius akebiae (Kuwana); Martin 1999: 91. 


Host: Lardizabalaceae: Akebia labata, Akebia quinata. 

Pealius amamianus Takahashi 1963 

Pealius amamianus Takahashi 1963: 55. Syntypes. Japan: on Quercus sp., HBL. 



Pealius artocarpi (Corbett) 1935 

Bemisia artocarpi Corbett 1935b: 784. Syntypes. Malaya: on Artocarpus sp. 

Pealius artocarpi (Corbett); Mound and Halsey 1978: 179. 


Host: Moraceae: Artocarpus sp. 

Pealius azaleae (Baker & Moles) 1920 

Aleyrodes azaleae Baker & Moles 1920: 81. Syntypes. Belgium: on Azalea sp., USNM. 

Pealius azaleae (Baker & Moles); Takahashi 1954: 50. 

Neopealius nilgiriensis David & Subramaniam 1976: 204. Holotype. India: Tamil Nadu, 3.vii.1969, B.V. 

David, on Rhododendron (=Azalea) indica, ZSI; Martin 1999: 92. 

Distribution: 1: USA (AL, CA, DC, FL, LA, MD, MS, NC, NJ, NY, OH, PA, RI, SC, VA); 3: Belgium; 

England; Iran; Netherlands; Scotland; USSR; 5: Japan; Korea; 6: India; 7: Australia; 8: New Zealand. 

Host: Ericaceae: Azalea pontica, Rhododendron indicum, Rhododendron mucronatum, Rhododendron 

schippenbachii. 

Natural enemies: Aphelinidae: Encarsia gautieri, Encarsia inaron, Encarsia lutea, Encarsia pergandiella, 

Encarsia tricolor. 

Pealius bangkokensis Takahashi 1942 

Pealius bangkokensis Takahashi 1942c: 213. Syntypes. Thailand: on unidentified host. 



Pealius bengalensis (Peal) 1903 

Aleurodes bengalensis Peal 1903b: 71. Syntypes. India: no host information. 

Pealius bengalensis (Peal); Quaintance & Baker 1914: 99. 

390



Pealius cambodiensis Takahashi 1942 

Pealius cambodiensis Takahashi 1942c: 208. Holotype. Cambodia: on unidentified host. 



Pealius cinnamomi David & Sundararaj 1991 

Pealius cinnamomi David & Sundararaj 1991a: 67. Holotype puparium, India, Karnataka, Mercara, 

11.viii.1971, B.V. David, on Cinnamomum sp., IARI.. 



Pealius cryptus Martin 1999 

Pealius cryptus Martin 1999: 92. Holotype puparium. Australia: on Ficus platypodia, ANIC. 


Host: Moraceae: Ficus platypodia. 

Pealius damnacanthi Takahashi 1935 

Pealius damnacanthi Takahashi 1935b: 61. Syntypes. Taiwan: on Damnacanthus indicus var. formosanus, 

TARI. 

Odontaleyrodes damnacanthi (Takahashi); Takahashi 1955d: 12. 

Pealius damnacanthi Takahashi; Martin 1999: 91. 


Host: Rubiaceae: Damnacanthus indicus; Theaceae: Eurya acuminata. 

Pealius elatostemae (Takahashi) 1932 

Trialeurodes elatostemae Takahashi 1932: 39. Syntypes. Taiwan: on Elatostema sp., TARI. 

Pealius elatostemae (Takahashi); Martin & Mound 2007: 40. 


Host: Urticaceae: Elatostema sp. 

391

Pealius elongatus (David, Sundararaj & Regu) 1991 

Odontaleyrodes elongatus David, Sundararaj & Regu 1991, in David et al. 1991: 117. 

Holotype puparium, India, Tamil Nadu, Kunnathoor, 25.i.199, K. Regu, on Ailanthus excelsa, David 

Collection. 

Pealius elongatus (David, Sundararaj & Regu); Martin 1999: 91; Dubey & Sundararaj 2005a: 162 



Pealius euryae (Takahashi) 1955 

Odontaleyrodes euryae Takahashi 1955d: 12. Syntypes. Japan: on Eurya japonica, HBL. 

Pealius euryae (Takahashi); Martin 1999: 91. 


Host: Theaceae: Eurya japonica. 

Pealius ezeigwi Mound 1965 

Pealius ezeigwi Mound 1965c: 151. Holotype. Nigeria: on undetermined plant, BMNH; Cohic 1969. 


Host: Euphorbiaceae: Alchornea cordifolia, Maesobotrya barteri; Sapindaceae: Lecaniodiscus cupanioides, 

Malacantha heudelotiana; Sterculiaceae: Cola nitida. 

Pealius fici Mound 1965 

Pealius fici Mound 1965c: 150. Holotype. Nigeria: on Ficus asperifolia, BMNH; Cohic 1969. 

Distribution: 4: Congo; Ivory Coast; Nigeria; Zaire. 

Host: Moraceae: Antiaris africana, Ficus asperifolia, Ficus capensis. 

Pealius indicus (David) 1972 

Odontaleyrodes indicus David 1972: 309. Holotype. India: Mysore State, Saklaspur, 12.viii.1971, on 

unidentified shrub, BMNH. 

Pealius indicus (David); Martin 1999:91. 


Host: on unidentified shrub. 

Note: not Pealius indicus David & Subramaniam 1976 = Pealius simplex. 

392

Pealius kalawi Singh 1933 

Pealius kalawi Singh 1933: 344. Syntypes. Burma: Singh, on Laurus sp., ZSI. 


Host: Lauraceae: Laurus sp. 

Reference: David & Jesudasan (1987b) - redescription and illustration. 

Pealius kankoensis (Takahashi) 1933 

Trialeurodes kankoensis Takahashi 1933: 15. Syntypes. Taiwan: on Quercus glauca, TARI. 

Pealius kankoensis (Takahashi); Takahashi 1935b: 63. 


Host: Fagaceae: Quercus glauca. 


Pealius kelloggi (Bemis) 1904 

Aleyrodes kelloggi Bemis 1904: 499. Syntypes. USA: California, on Quercus agrifolia, USNM. 

Pealius kelloggi (Bemis); Quaintance & Baker 1914: 99. 


Host: Fagaceae: Quercus agrifolia, Quercus virginiana; Rosaceae: Prunus lyoni, Prunus sp., 

Prunus ilicifolia. 

Natural enemies: Coccinellidae: Delphastus catalinae, Nephaspis oculatus. 

Pealius kongosana (Takahashi) 1955 

Odontaleyrodes kongosana Takahashi 1955d: 11. Syntypes. Japan: on Lindera sp., HBL. 


Host: Lauraceae: Lindera sp. 

Pealius liquidambari (Takahashi) 1932 

Aleyrodes liquidambari Takahashi 1932: 37. Syntypes. Taiwan: on Liquidambar formosana, TARI. 

Pealius liquidambari (Takahashi); Mound and Halsey 1978: 181. 


Host: Hamamelidaceae: Liquidambar formosana. 

393

Pealius longispinus Takahashi 1932 

Pealius longispinus Takahashi 1932: 41. Syntypes. Taiwan: on Ficus beecheyana, TARI. 


Host: Fabaceae: Pueraria lobata; Moraceae: Ficus beecheyana. 

Natural enemies: Aphelinidae: Encarsia sophia. 

Pealius machili Takahashi 1935 

Pealius machili Takahashi 1935b: 62. Syntypes. Taiwan: on Machilus sp., TARI. 



Pealius maculatus Takahashi 1942 

Pealius maculatus Takahashi 1942c: 211. Syntypes. Cambodia: on unidentified tree, TARI. 



Pealius madeirensis Martin, Aguiar & Pita 1996 

Pealius madeirensis Martin, Aguiar & Pita 1996, in Martin et al. 1996: 121. Holotype pupal case, Madeira, 

Faja da Noguiera, 11.ii.1990, A. Aguiar, on Picconia excelsa (Oleaceae), BMNH; Hernandez et al. 

2003: 72 

Distribution: 3: Madeira; Canary Islands. 

Host: Oleaceae: Picconia excelsa. 

Natural enemies: Aphelinidae: Encarsia inaron. 

Pealius maskelli (Bemis) 1904 

Aleyrodes maskelli Bemis 1904: 524. Syntypes. USA: California, on Quercus densiflora, USNM; Sampson & 

Drews 1941: 169. 

Pealius maskelli (Bemis); Quaintance & Baker 1914: 99. 

Distribution: 1: USA (CA); 2: Mexico. 

Host: Fagaceae: Quercus densiflora. 

Pealius misrae Singh 1931 

Pealius misrae Singh 1931: 44. Syntypes. India: Bihar, on Psidium sp. 


394

Host: Myrtaceae: Psidium sp. 

US Port Interceptions: Myanmar on Oroxylum indicum; Philippines on Moringa oleifera, Moringa sp.; 

Viet Nam on unknown plant. 

Pealius mitakensis (Takahashi) 1955 

Odontaleyrodes mitakensis Takahashi 1955d: 12. Syntypes. Japan: on Pertya ovata, HBL. 

Pealius mitakensis (Takahashi); Martin 1999: 91. 


Host: Asteraceae: Pertya ovata. 

Pealius mori (Takahashi) 1932 

Trialeurodes mori Takahashi 1932: 38. Syntypes. Taiwan: on Morus alba and Salix sp., TARI. 

Pealius mori (Takahashi); Mound and Halsey 1978: 182. 

Distribution: 5: Taiwan; 6: Thailand. 

Host: Euphorbiaceae: Glochidion phillpicum; Moraceae: Morus australis, Morus alba; Salicaceae: Salix sp. 

Natural enemies: Aphelinidae: Encarsia cibcensis, Encarsia perflava, Encarsia bothrocera, Encarsia lutea. 

Pealius nagerkoilensis Jesudasan & David 1991 

Pealius nagerkoilensis Jesudasan & David 1991: 320. Holotype pupal case. India: Tamil Nadu, Kanyakumari 

District, Kunnathoor, 16.vi.1989, K. Regu, on Loranthus sp., IDAV. 

Fippataleyrodes nagerkoilensis (Jesudasan & David); Dubey & Sundararaj 2005a: 163. 

Odontaleyrodes artocarpi David, Sundararaj & Regu 1991; Dubey & Sundararaj 2005a: 163. 

Odontaleyrodes selvakumarani David, Sundararaj & Regu 1991: 118. Holotype puparium, India, Tamil Nadu, 

Madurai District, Cumbum, 7.iv.1988, S. Selvakumaran, on Chickrassia velutina, IARI; Dubey & 

Sundararaj 2005a: 163. 


Host: Asteraceae: Ageratum sp., Ageratum conyzoides; Dipterocarpaceae: Madhuca neriifolia; 

Fabaceae: Derris sp, Butea monosperma; Labiatae: Salvia splendens; Loranthaceae: Loranthus sp.; 

Melastomataceae: Osbeckia reticulata; Solanaceae: Cestrum aurantiacum; Thymelaeaceae: Gnidia 

glauca. 

Pealius nilgiriensis (David) 1972 

Odontaleyrodes nilgiriensis David 1972: 310. Holotype. India: Tamil Nadu, Ootacamund, 3.viii.1969, B.V. 

David, on unidentified "twiner", IDAV. 

Pealius nilgiriensis (David); Martin 1999: 91. 

395


Host:on unidentified "twiner". 

Pealius polygoni Takahashi 1934 

Pealius polygoni Takahashi 1934b: 59. Syntypes. Taiwan: on Polygonum sp., TARI; Suh et al. 2006. 

Distribution: 5: Japan; Korea; Taiwan. 

Host: Polygonaceae: Persicaria hydropiper, Polygomum longisetum, Polygonum thunbergii, 

Polygonum sp.; Urticaceae: Boehmeria nivea. 

Natural enemies: Aphelinidae: Encarsia duorunga, Encarsia synaptocera. 

Pealius psychotriae Takahashi 1935 

Pealius psychotriae Takahashi 1935b: 59. Syntypes. Taiwan: on Psychotria serpens, TARI. 


Host: Rubiaceae: Psychotria serpens. 

Pealius quercus (Signoret) 1868 

Aleurodes quercus Signoret 1868: 384. Syntypes. France: on Quercus pedunculata. 

Pealius quercus (Signoret); Trehan 1939: 266. 

Aleurochiton quercus (Signoret); Visnya 1941b: 8. 

Aleyrodes avellanae (Signoret), misidentification Trehan 1940: 597 corrected misidentification, Zahradnik 

1956: 44. 

Distribution: 3: Czech Republic; Denmark; England; Finland; France; Germany; Iran, Ireland; Poland; 

Scotland; Spain; Sweden; Turkmenistan; USSR. 

Host: Betulaceae: Carpinus betulus, Corylus heterophylla, Corylus avellana, Ostrya sativa; 

Fagaceae: Castanea sativa, Fagus silvatica, Quercus persica, Quercus robur, Quercus 

hartwissiana; Tiliaceae: Tilia platyphyllos; Urticaceae: Urtica dioica. 

Natural enemies: Aphelinidae: Encarsia inaron, Encarsia moffsi, Encarsia lutea, Eretmocerus adustiscutum, 

Eretmocerus zippanguiphagus; Platygastridae: Amitus longicornis. 

Pealius rhododendri Takahashi 1935 

Pealius rhododendri Takahashi 1935: 279. Syntypes, imported from Japan into Taiwan, Keelung, iii.1935, on 

Rhododendron sp., TARI. 

Odontaleyrodes rhododendri (Takahashi); Takahashi 1954: 50. 

Pealius rhododendri Takahashi; Martin 1999: 91. 

Distribution: 1: USA (AL, CA, FL, HI, LA, TX); 5: Japan; Taiwan; 9: Hawaii. 

396

397 

Host: Ericaceae: Rhododendron simsii, Rhododendron sp., Rhododendron indicum, Rhododendron 

mucronatum, Rhododendron similis, Rhododendron formosanum. 

Natural enemies: Aphelinidae: Encarsia azimi, Encarsia inaron, Encarsia perflava. 

Pealius rubi Takahashi 1936 

Pealius rubi Takahashi 1936b: 219. Syntypes. Taiwan: on Rubus sp., TARI; Suh et al. 2006. 

Distribution: 5: Japan; Korea; Taiwan. 

Host: Amaranthaceae: Achyranthes bidentata; Anacardiaceae: Rhus sp.; Cornaceae: Cornus kousa; Fabaceae: 

Pueraria hirsuta; Magnoliaceae: Magnolia kobus; Moraceae: Ficus elastica; Polygonaceae: 

Polygonum thunbergii, Polygonum sp.; Rosaceae: Rubus thunbergii, Rubus sp.; Verbenaceae: 

Callicarpa japonica, Clerodendrum trichotomum, Phryma leptostachya. 

Natural enemies: Aphelinidae: Encarsia duorunga, Encarsia lutea, Encarsia synaptocera. 

Pealius sairandhryensis (Meganathan & David) 1994 

Odontaleyrodes sairandhryensis Meganathan & David 1994: 47. Holotype. India: on Oreocnida integrifolia. 

Pealius sairandhryensis (Meganathan & David) 1994; Martin 1999. 


Host: Dipterocarpaceae: Nothopegia racemosa; Urticacae: Oreocnida integrifolia. 

Pealius schimae Takahashi 1950 

Pealius schimae Takahashi 1950: 88. Syntypes. Borneo: on Schima robustum, BMNH; David & 

Subramaniam 1976 

Distribution: 6: India; 7: Borneo. 

Host: Convolvulaceae: Ipomoea sp.; Moraceae: Artocarpus robustum; Theaceae: Schima robustum. 

Pealius setosus Danzig 1964 

Pealius setosus Danzig 1964a: 642. Holotype. USSR: Batumi, on Rubus casius, ZIN. 


Host: Cupressaceae: Juniperus chinensis, Juniperus sp; Rosaceae: Fragaria vesca, Rubus idaeus, Rubus 

oderatus. 

Natural enemies: Aphelinidae: Encarsia lutea, Encarsia tricolor. 

Pealius spinosus David & Jesudasan 1991 

Pealius spinosus Jesudasan and David 1991: 322. Holotype pupal case. India: on unidentified tree, IDAV. 

Distribution: 6: India.


Pealius spinus (Singh) 1931 

Dialeurodes spina Singh 1931: 27. Syntypes. India: on Ficus religiosa. 

Aleuroplatus spinus (Singh); Takahashi 1952: 23; Mound & Halsey 1978: 56. 

Pealius spinus (Singh); Jesudasan and David 1991: 322; Dubey & Sundararaj 2005a: 164. 

Distribution: 6: India; Malaysia; Singapore. 

Host: Moraceae: Ficus arnotiana, Ficus bengalensis, Ficus religiosa. 

Pealius splendens (David, Sundararaj & Regu) 1991 

Odontaleyrodes splendens David, Sundararaj & Regu 1991, in David et al. 1991: 117. Holotype puparium, 

India, Kerala, Emakuzham District, Tirpunithura, 30.vii.1987, R. Sundararaj, on Tectona grandis, 

IARI. 

Pealius splendens (David, Sundararaj & Regu); Dubey & Sundararaj 2005a: 164. 


Host: Verbenaceae: Tectona grandis. 

Pealius sutepensis Takahashi 1942 

Pealius sutepensis Takahashi 1942c: 214. Syntypes. Thailand: on Lauraceae tree, TARI. 



Pealius tuberculatus Takahashi 1942 

Pealius tuberculatus Takahashi 1942c: 210. Syntypes. Thailand: on Celtis sp., TARI 



Pealius walayarensis (Jesudasan & David) 1991 

Aleurolobus walayarensis Jesudasan and David 1991: 278. Holotype pupal case. India: on unidentified tree, 

IDAV. 

Pealius walayarensis (Jesudasan & David); Dubey (personal communication). 



398

Transfers 

Pealius homonoiae Jesudasan & David 1991- to Massileurodes 

399

Pectinaleyrodes Bink-Moenen 1983 

Pectinaleyrodes Bink-Moenen 1983: 158. Type species. Aleuroplatus triclisiae Corbett 1966, by description. 




culcasiae x 

silvaticus x 

triclisiae x 

Total = 3 species 0 0 0 3 0 0 0 0 0 

Pectinaleyrodes culcasiae (Cohic) 1969 

Aleuroplatus culcasiae Cohic 1969: 70. Holotype. Ivory Coast: on Cercestis afzelii. 

Pectinaleyrodes culcasiae (Cohic); Bink-Moenen 1983: 159. 


Host: Annonaceae: Monodora myristica; Araceae: Cercestis afzelii, Culcasia angolensis. 

Connaraceae: Agelaea paradoxa; Sterculiaceae: Cola gabonensis. 

Pectinaleyrodes silvaticus (Cohic) 1969 

Aleuroplatus silvaticus Cohic 1969: 74. Holotype. Ivory Coast: on Culcasia angolensis. 

Pectinaleyrodes silvaticus (Cohic); Bink-Moenen 1983: 159. 


Host: Araceae: Culcasia angolensis. 

Pectinaleyrodes triclisiae (Cohic) 1966 

Aleuroplatus triclisiae Cohic 1966b: 26. Syntypes. Congo: on Triclisia gilletii. 

Pectinaleyrodes triclisiae (Cohic); Bink-Moenen 1983: 159. 

Distribution: 4: Angola; Chad; Congo. 

Host: Annonaceae: Uvaria scabrida; Bignoniaceae: Markhamia sessilis; Dilleniaceae: Tetracera alnifolia; 

Fabaceae: Millettia laurentii; Flacourtiaceae: Caloncoba welwitschii; Meliaceae: Trichilia heudelottii; 

Menispermaceae: Triclisia gilletii; Rubiaceae: Pauridiantha hirtella; Verbenaceae: Clerodendrum 

speciosissimum. 

400

Pentaleyrodes Takahashi 1937 

Pentaleyrodes Takahashi 1937c: 310. Type species. Aleyrodes cinnamomi Takahashi 1932, by monotypy. 



cinnamomi x 

hongkongensis x 

linderae x 

yasumatsui x 

Total = 4 species 0 0 0 0 4 0 0 0 0 

Pentaleyrodes cinnamomi (Takahashi) 1932 

Aleyrodes cinnamomi Takahashi 1932: 36. Syntypes. Taiwan: on Cinnamomum japonicum, Machilus sp. and 

Pheobe formosana, TARI. 

Pentaleyrodes cinnamomi (Takahashi); Takahashi 1937c: 310. 


Host: Lauraceae: Cinnamomum japonicum, Machilus sp., Phoebe formosana. 

Pentaleyrodes hongkongensis Takahashi 1941 

Pentaleyrodes hongkongensis Takahashi 1941c: 389. Syntypes. Hong Kong: on Lauraceae. 



Pentaleyrodes hongkongensis Takahashi 1941 

Pentaleyrodes linderae Chou & Yan 1988: 244. Holotype puparium, China, Dayong, Hunan, 12.x.1985, F. 

Yan, ex. Lindera communis. 


Host: Lauraceae: Lindera communis. 

Pentaleyrodes yasumatsui Takahashi 1939 

Pentaleyrodes yasumatsui Takahashi 1939a: 76. Syntypes. Japan: Litsea glauca, TARI. 


Host: Lauraceae: Lindera sp., Litsea glauca. 

401

Peracchius Lima and Racca-Filho 2005 

Peracchius Lima & Racca-Filho 2005: 39. Type species, Peracchius durantae, by monotypy. 



durantae x 

sp. x 

Total = 1+ species 0 1+ 0 0 0 0 0 0 0 

Peracchius durantae Lima & Racca-Filho 2005 

Peracchius durantae Lima & Racca-Filho 2005: 39. Holotype puparium, Brazil on Duranta repens. 

Distribution: 2: Belize, Brazil. 

Verbenaceae: Duranta repens. 

Peracchius sp. Martin 2005b: 46. 

Distribution: 2: Belize 

Host: Lauraceae: Nectandra sp. 

Peracchius sp. 

402

Plataleyrodes Takahashi & Mamet 1952 

Plataleyrodes Takahashi & Mamet 1952b: 119. Type species. Plataleyrodes anthocleistae Takahashi & 

Mamet 1952, by monotypy. 



anthocleistae x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Plataleyrodes anthocleistae Takahashi & Mamet 1952 

Plataleyrodes anthocleistae Takahashi & Mamet 1952b: 120. Syntypes. Madagascar: on Anthocleista sp., 

MNHNP. 


Host: Loganiaceae: Anthocleista sp. 

403

Pogonaleyrodes Takahashi 1955 

Pogonaleyrodes Takahashi 1955a: 400. Type species. Pogonaleyrodes fastuosa Takahashi 1955, by 

monotypy. 



fastuosa x 

zimmermanni x 

Total = 2 species 0 0 0 2 0 0 0 0 0 

Pogonaleyrodes fastuosa Takahashi 1955 

Pogonaleyrodes fastuosa Takahashi 1955a: 403. Syntypes. Madagascar: on unidentified plant, MNHNP. 


Host: an unidentified plant. 

Pogonaleyrodes zimmermanni (Newstead) 1911 

Aleurodes zimmermanni Newstead 1911: 173. Syntypes. Tanzania: on Acanthaceae, BMNH. 

Pogonaleyrodes zimmermanni (Newstead); Mound 1965c: 154; Cohic 1968b; Bink-Moenen 1983: 160. 

Distribution: 4: Congo; Ivory Coast; Nigeria; Tanzania. 

Host: Rubiaceae: Coffea arabica, Cremaspora triflora, Gardenia jovis-tonantis 

Rutidea syringoides. 

404

Pseudaleurolobus Hempel 1923 

Pseudaleurolobus Hempel 1923: 1190. Type species. Pseudaleurolobus jaboticabae Hempel 1923, by 

monotypy. 



jaboticabae x 

Total = 1 species 0 1 0 0 0 1 0 0 0 

Pseudaleurolobus jaboticabae Hempel 1923 

Pseudaleurolobus jaboticabae Hempel 1923: 1190. Syntypes. Brazil: on Eugenia jaboticabae, MZU. 


Host: Myrtaceae: Eugenia jaboticabae. 

405

Pseudaleuroplatus Martin 1999 

Pseudaleuroplatus Martin 1999: 93. Type species. Pseualeuroplatus kiensis Martin 1999, by original 

designation. 



kiensis x 

litseae x 

Total = 2 species 0 0 0 0 0 0 2 0 0 

Pseudaleuroplatus kiensis Martin 1999 

Pseudaleuroplatus kiensis Martin 1999: 94. Holotype puparium. Australia: on Melaleuca sp., ANIC. 


Host: Myrtaceae: Melaleuca sp. 

Pseudaleuroplatus litseae (Dumbleton) 1956 

Tetraleurodes litseae Dumbleton 1956b: 178. Holotype. Australia: on Litsea dealbata, ADSIR. 

Pseudaleuroplatus litseae (Dumbleton); Martin 1999: 94. 


Host: Lauraceae: Litsea dealbata. 

406

Pseudaleyrodes Hempel 1923 

Pseudaleyrodes Hempel 1923: 1176. Type species. Pseudaleyrodes depressus Hempel 1923, by monotypy. 



depressus x 

Total = 1 species 0 1 0 0 0 0 0 0 0 

Pseudaleyrodes depressus Hempel 1923 

Pseudaleyrodes depressus Hempel 1923: 1176. Syntypes. Brazil: on Maytenus aquifolium, MZU. 


Host: Celastraceae: Maytenus aquifolium. 

407

Pseudozaphanera Manzari 2006 

Pseudozaphanera Manzari 2006, in Manzari & Quicke 2006: 2472. Type species. Aleurodes niger, by original 

description. 

Comment: All of the known species in this genus are found on Acacia (Fabaceae) in Australia. 



niger x 

papyrocarpae x 

rhachisreticulata x 

splendida x 

wariensis x 

Total = 5 species 0 0 0 0 0 0 5 0 0 

Zaphanera niger (Maskell) 1895 

Aleurodes niger Maskell 1895: 437. Lectotype. Australia: on Acacia pycantha, ADSIR. 

Aleyrodes nigra Maskell [sic]; Cockerell 1902a: 281. 

Aleurolobus niger (Maskell); Quaintance & Baker 1914: 109. 

Tetraleurodes niger (Maskell); Dumbleton 1956b: 180; Mound & Halsey 1978: 201. 

Zaphanera nigra (Maskell); Martin 1999: 120. 


Host: Fabaceae: Acacia melanoxylon, Acacia wilhelmiana. 

Zaphanera papyrocarpae Martin 2001 

Zaphanera papyrocarpae Martin 2001, in Bailey et al. 2001: 84. Holotype male puparium, Australia, 

Billakilina Station, 26.iv.2000, J. H. Martin, on Acacia papyrocarpa, ANIC. 


Host: Fabaceae: Acacia papyrocarpa. 

Natural enemies: Encyrtidae: Zarhopaloides anaxenor. 

Zaphanera rhachisreticulata Martin 1999 

Zaphanera rhachisreticulata Martin 1999: 121. Holotype puparium. Australia: on Acacia mackeyana, ANIC. 

408


Host: Fabaceae: Acacia mackeyana. 

Zaphanera splendida Martin 1999 

Zaphanera splendida Martin 1999: 122. Holotype puparium. Australia: on Acacia aprepta, ANIC. 


Host: Fabaceae: Acacia aprepta, Acacia longifolia, Acacia sp. 

Zaphanera wariensis Martin 1999 

Zaphanera wariensis Martin 1999: 122. Holotype puparium. Australia: on Acacia longifolia, ANIC. 


Host: Fabaceae: Acacia longifolia. 

Natural enemies: Aphelinidae: Encarsia prinslooi. 

409

Rabdostigma Quaintance & Baker 1917 

Dialeurodes (Rabdostigma) Quaintance & Baker 1917: 425. Type species. Dialeurodes (Rabdostigma) 

radiilinealis Quaintance & Baker 1917, by original designation. 

Rabdostigma Quaintance & Baker; David & Sundararaj 1993. 

Comment: Treated as Dialeurodes species in Martin & Mound 2007. 



mahableshwarensis x 

minahassai x 

radiilinealis x 

Total = 3 species 0 0 0 0 0 2 1 0 0 

Rabdostigma mahableshwarensis Sundararaj & David 1998 

Rabdostigma mahableshwarensis Sundararaj and David 1998: 326. Holotype. India: Mahableshwar, 

28.iii.1987, B. V. David, on Memcyclon umbellatum, USNM. 


Host: Melastomataceae: Memecylon umbellatum. 

Rabdostigma minahassai Martin 1988 

Dialeurodes (Rabdostigma) minahassai Martin 1988: 74. Holotype pupal case, Indonesia, Sulawesi, Utara, 

13.vi.1985, J. Martin, on Syzygium aromaticum, BMNH. 

Rabdostigma minahassai Martin; 


Host: Myrtaceae: Syzygium aromaticum. 

Rabdostigma radiilinealis (Quaintance & Baker) 1917 

Dialeurodes (Rabdostigma) radiilinealis Quaintance & Baker 1917: 425. Holotype. Sri Lanka: on Mistletoe 

[Loranthaceae]. 

Rabdostigma radiilinealis (Quaintance & Baker); Sundararaj & David 1998: 325. 


Host: Loranthaceae: Arceuthobium sp. 

410

Ramsesseus Zahradnik 1970 

Ramsesseus Zahradnik 1970: 47. Type species. Ramsesseus follioti Zahradnik 1970, by monotypy. 



follioti x 

Total = 1 species 0 0 1 0 0 0 0 0 0 

Ramsesseus follioti Zahradnik 1970 

Ramsesseus follioti Zahradnik 1970: 47. Holotype and paratypes. Egypt: Ramesseium, 10.i.1963, ex. Acacia 

sp.; paratypes in USNM and BMNH, USNM. 

Distribution: 3: Egypt. 

Host: Fabaceae: Acacia nilotica, Acacia sp. 

Natural enemies: Aphelinidae: Encarsia ramsesi. 

411

Rhachisphora Quaintance & Baker 1917 

Dialeurodes (Rhachisphora) Quaintance & Baker 1917: 430. Type species. Dialeurodes (Rhachisphora) 

trilobitoides, by original designation. 

Rhachisphora Quaintance & Baker; full genus by Takahashi 1952: 22 

Greeniella David 1993: 22. Type species. Rhachisphora capitatis Corbett 1926, by monotypy; a junior 

homonym of Greeniella Cockerell 1897, synonymy according to Martin 1999: 95. 

Reference: Sundararaj & Dubey 2006a - key to Indian species 



alishanensis x 

ardisiae x x 

capitatis x 

elongatus x 

fijiensis x 

franksae x 

indica x 

ixorae x 

kallarensis x 

koshunensis x 

machili x 

maesae x 

malayensis x 

oblongata x 

queenslandica x 

reticulata x 

rutherfordi x 

sanhsianensis x 

selangorensis x 

setulosa x x 

412


styraci x 

taiwana x 

trilobitoides x 

Total = 23 species 0 0 0 0 10 10 3 1 0 

Rhachisphora alishanensis Ko 1992 

Rhachisphora alishanensis Ko 1992, in Ko et al. 1992: 251. Holotype puparium, Taiwan, Alishan, 16.iv.1990, 

C. C. Ko, ex. Eurya strigillosa, NTU. 


Host: Theaceae: Eurya strigillosa. 

Rhachisphora ardisiae (Takahashi) 1935 

Dialeurodes ardisiae Takahashi 1935a: 50. Syntypes. Taiwan: on Bladhia sp., TARI. 

Rhachisphora ardisiae (Takahashi); Martin 1985: 333. 

Distribution: 5: Taiwan; 7: Papua New Guinea. 

Host: Myrsinaceae: Ardisia sp. 

Rhachisphora capitatis (Corbett) 1926 

Dialeurodes (Rhachisphora) capitatis Corbett 1926: 270. Syntypes. Sri Lanka: Nuwara Eliya, iii.1911, E. E. 

Green, on unidentified host, BMNH. 

Rhachisphora capitatis (Corbett); Mound and Halsey 1978: 186. 



Rhachisphora elongatus Regu & David 1990 

Rhachisphora elongatus Regu & David 1990: 277. Holotype puparium, Tamil Nadu, Kunnathoor, 24.i.1989, 

K. Regu, on Mimusops elengi, IARI. 


Host: Sapotaceae: Mimusops elengi. 

Rhachisphora fijiensis (Kotinksy) 1907 

Aleyrodes fijensis Kotinsky 1907: 100. Syntypes. Fiji: on pods and leaves of Leguminous plant, USNM. 

413

Dialeurodes fijensis (Kotinsky); Quaintance & Baker 1914: 97. 

Dialeurodes (Rhachisphora) fijensis (Kotinsky); Quaintance & Baker 1917: 431. 

Rhachisphora fijiensis (Kotinksy); Mound and Halsey 1978: 186. 



Rhachisphora franksae Martin 1999 

Rhachisphora franksae Martin 1999: 96. Holotype puparium. Australia: on Fagraea gracilipes, ANIC. 


Host: Loganiaceae: Fagraea gracilipes. 

Rhachisphora indica Sundararaj & David 1991 

Rhachisphora indica Sundararaj & David 1991: 311. Holotype, India, Tamil Nadu, 6.viii.1987, R. Sundararaj, 

on unidentified plant, USNM (NHM, IARI). 



Rhachisphora ixorae Sundararaj & David 1991 

Rhachisphora ixorae Sundararaj & David 1991: 312. Holotype, India, Tamil Nadu, 6.iii.1971, B. V. David, on 

Ixora sp., USNM. 



Rhachisphora kallarensis Jesudasan & David 1991 

Rhachisphora kallarensis Jesudasan and David 1991: 324. Holotype pupal case. India: on unidentified tree, 

IDAV. 



Rhachisphora koshunensis (Takahashi) 1933 

Dialeurodes (Rhachisphora) koshunensis Takahashi 1933: 6. Syntypes. Taiwan: on Cinnamomum zeylanicum, 

TARI. 

Rhachisphora koshunensis (Takahashi); Mound and Halsey 1978: 186. 

Host: Lauraceae: Cinnamomum reticulatum, Cinnamomum zeylanicum, Machilus sp. 

414

Rhachisphora machili (Takahashi) 1932 

Dialeurodes (Rhachisphora) machili Takahashi 1932: 16. Syntypes. Taiwan: Machilus sp., TARI. 

Rhachisphora machili (Takahashi); Mound and Halsey 1978: 187. 



Rhachisphora maesae (Takahashi) 1932 

Dialeurodes (Rhachisphora) maesae Takahashi 1932: 18. Holotype. Taiwan: on Maesa japonica, TARI. 

Rhachisphora maesae (Takahashi); Mound and Halsey 1978: 187. 


Host: Myrsinaceae: Ardisia cornudenta, Maesa japonica. 

Natural enemies: Aphelinidae: Encarsia albiscutellum longipalpa Hayat 1989 

Rhachisphora malayensis (Takahashi) 1952 

Rhachisphora malayensis Takahashi 1952: 22. Holotype. Malaya: on undetermined shrub. 


Host: undetermined shrub. 

Rhachisphora oblongata Ko 1988 

Rhachisphora oblongata Ko 1988, in Ko et al. 1998: 61. Holotype puparium. Taiwan, Pingtung, Kentingkuyen, 

2 III 1994, K. C. Chou, ex. Leea guineensis, NTU. 


Host: Leeaceae: Leea guineensis. 

Rhachisphora queenslandica Martin 1999 

Rhachisphora queenslandica Martin 1999: 97. Holotype puparium. Australia: on Syzygium sp., ANIC. 



Rhachisphora reticulata (Takahashi) 1933 

Dialeurodes (Rhachisphora) reticulata Takahashi 1933: 7. Syntypes. Taiwan: on Machilus sp. 

Rhachisphora reticulata (Takahashi); Mound and Halsey 1978: 187. 



415

Rhachisphora rutherfordi (Quaintance & Baker) 1917 

Dialeurodes (Rhachisphora) rutherfordi Quaintance & Baker 1917: 432. Syntypes. Sri Lanka: on Loranthus 

sp., USNM. 

Rhachisphora rutherfordi (Quaintance & Baker); Mound and Halsey 1978: 187. 

Distribution: 6: India, Sri Lanka. 

Host: Loranthaceae: Loranthus eleasticus, Loranthus sp. 

Reference: David & Regu (1991) - redescription and illustration. 

Rhachisphora selangorensis (Corbett) 1933 

Dialeurodes selangorensis Corbett 1933: 122. Syntypes. Malaya: on Cinnamomum camphora, BMNH. 

Rhachisphora selangorensis (Corbett); Mound and Halsey 1978: 187. 


Host: Lauraceae: Cinnamomum camphora. 

Rhachisphora sanhsianensis Ko 1992 

Rhachisphora sanhsianensis Ko 1992, in Ko et al. 1992: 254. Holotype puparium, Taiwan, Sanhsian, 

22.vi1990, C. C. Ko, ex. Machilus kusanoi, NTU. 


Host: Lauraceae: Machilus kusanoi. 

Rhachisphora setulosa (Corbett) 1926 

Dialeurodes (Rhachisphora) setulosa Corbett 1926: 188. Syntypes. Sri Lanka: on unidentified plant, BMNH. 

Rhachisphora setulosa (Corbett); Mound and Halsey 1978: 188. 

Distribution: 3: Iran; 6: Sri Lanka. 

Host: Ebenaceae: Diospyros khaki; unidentified plant. 

Rhachisphora styraci (Takahashi) 1934 

Dialeurodes (Rhachisphora) styraci Takahashi 1934c: 223. Syntypes. Japan: on Styrax sp., TARI. 

Rhachisphora styraci (Takahashi); Mound and Halsey 1978: 188. 


Host: Styracaceae: Styrax sp. 

US Port Interceptions: Japan on Styrax sp.; Unknown source on Chamaecyparis sp. 

416

Rhachisphora trilobitoides (Quaintance & Baker) 1917 

Dialeurodes (Rhachisphora) trilobitoides Quaintance & Baker 1917: 433. Syntypes. Sri Lanka: on Harpullia 

sp., USNM. 

Rhachisphora trilobitoides (Quaintance & Baker); Takahashi 1952: 22; Sundararaj & David 1991: 315. 

Rhachisphora madhucae Jesudasan & David 1991: 325. Holotype pupal case. India: on Madhuca longifolia, 

IDAV. (Synonymised by Sundararaj and Dubey 2006). 


Host: Boraginaceae: Cordia myxa; Dipterocarpaceae: Madhuga longifolia; Loganiaceae: Strychnos nuxvomica; 

Myrtaceae: Eugenia sp., Eugenia jambos, Eugenia operculata; Rubiaceae: Randia 

malabarica; Sapindaceae: Harpullia sp.; Sapotaceae: Achras sapota, Mimusops elangi, Mimusops 

hexandra. 

417

Rosanovia Danzig 1969 

Rosanovia Danzig 1969: 555. Type species. Rosanovia hulthemiae Danzig 1969, by monotypy. 



hulthemiae x 

Total = 1 species 0 0 1 0 0 0 0 0 0 

Rosanovia hulthemiae Danzig 1969 

Rosanovia hulthemiae Danzig 1969: 556. Holotype. USSR: south Kazakhstan, on Hulthemia berberfolia, 

ZIN. 

Distribution: 3: Iran, Kazakhstan (USSR) 

Host: Rosaceae: Hulthemia berberifolia, Hulthemia persica. 

418

Rugaleyrodes Bink-Moenen 1983 

Rugaleyrodes Bink-Moenen 1983: 162. Type species - Marginaleyrodes angolensis Cohic 1966, by original 

description. 




angolensis x 

bidentata x 

tetracerae x 

villiersi x 

vuattouxi x 

Total = 5 species 0 0 0 5 0 0 0 0 0 

Rugaleyrodes angolensis (Cohic) 1966 

Marginaleyrodes angolensis Cohic 1966a: 49. Holotype. Angola: on unidentified host. 

Rugaleyrodes angolensis (Cohic); Bink-Moenen 1983: 162. 


Host: Icacinaceae: Icacina mannii; Marantaceae: Tachyphrynium violaceum; Ochnaceae: Rhabdophyllum 

welwitschii. 

Rugaleyrodes bidentata Bink-Moenen 1983 

Rugaleyrodes bidentata Bink-Moenen 1983: 165. Holotype puparium, Chad, Caibokoum, 19.xi.1972, ex. 

Hexalobus monopetalus, BMNH. 


Host: Annonaceae: Hexalobus monopetalus. 

Rugaleyrodes tetracerae (Cohic) 1966 

Marginaleyrodes tetracerae Cohic 1966b: 61. Syntypes. Congo: on Tetracera alnifolia, BMNH. 

Rugaleyrodes tetracerae (Cohic); Bink-Moenen 1983: 162. 


Host: Dichapetalaceae: Dichapetalum brazzae; Dilleniaceae: Tetracera alnifolia; Linaceae: Ochthocosmus 

africanus; Loganiaceae: Strychnos usambarensis; Meliaceae: Trichilia heudelottii; Myrtaceae: 

419

Syzygium brazzavillense; Ochnaceae: Ochna gilletiana. 

Rugaleyrodes villiersi (Cohic) 1968 

Aleuroplatus villiersi Cohic 1968b: 109. Syntypes (26). Congo: on Loranthus sp. 

Rugaleyrodes villiersi (Cohic); Bink-Moenen 1983: 162. 



Rugaleyrodes vuattouxi (Cohic) 1969 

Aleuroplatus vuattouxi Cohic 1969: 77. Holotype. Ivory Coast: on Saba senegalensis var glabriflora. 

Rugaleyrodes vuattouxi (Cohic); Bink-Moenen 1983: 162. 


Host: Apocynaceae: Saba senegalensis. 

420

Rusostigma Quaintance & Baker 1917 

Rusostigma Quaintance & Baker 1917: 420. Type species. Dialeurodes (Rusostigma) radiirugosa Quaintance 

& Baker 1917, by original designation. 



eugeniae x x 

radiirugosa x x x 

tokyonis x 

tristylii x x 

Total = 4 species 0 0 0 0 3 3 2 0 0 

Rusostigma eugeniae (Maskell) 1895 

Aleurodes eugeniae Maskell 1895: 430. Syntypes. India: on Eugenia jambolana, USNM. 

Aleyrodes (Dialeurodes) eugeniae (Maskell): Cockerell 1902a: 283. 

Dialeurodes eugeniae (Maskell); Quaintance & Baker 1914: 97. 

Dialeurodes (Rusostigma) eugeniae (Maskell); Quaintance & Baker 1917: 421. 

Rusostigma eugeniae (Maskell); Mound and Halsey 1978: 188 

Distribution: 6: India; 7: Java; Sumatra. 

Host: Myrtaceae: Eugenia jambolana, Eugenia sp. 

Natural enemies: Parasitoids: Aphelinidae: Encarsia strenua; Predators: Coccinellidae: Coccinella sp., 

Menochilus sexmaculatus, Scymnus sp. 

Rusostigma radiirugosa Quaintance & Baker 1917 

Dialeurodes (Rusostigma) radiirugosa Quaintance & Baker 1917: 421. Syntypes. Indonesia: Belitung Isle, on 

Mangifera indica, USNM. 

Rusostigma radiirugosa (Quaintance & Baker); Takahashi & Mamet 1963: 49. 

Distribution: 5: China; Hong Kong; Japan; 6: Cambodia; India; Malaya; Malaysia; Thailand; 7: Indonesia. 

Host: Anacardiaceae: Mangifera indica. 

US Port Interceptions: Indonesia on Magnolia sp., Solanaceae; Singapore on unknown plant. 

Rusostigma tokyonis (Kuwana) 1911 

Aleyrodes tokyonis Kuwana 1911: 622. Syntypes. Japan: on Ilex integra, USNM. 

421

Dialeurodes tokyonis (Kuwana); Quaintance & Baker 1914: 98. 

Dialeurodes (Rusostigma) tokyonis (Kuwana); Quaintance & Baker 1917: 423. 

Rusostigma tokyonis (Kuwana); Takahashi & Mamet 1963: 49. 


Host: Aquifoliaceae: Ilex integra; Theaceae: Cleyera japonica, Eurya ochnacea. 

Rusostigma tristylii (Takahashi) 1935 

Dialeurodes tristylii Takahashi 1935b: 48. Syntypes. Taiwan: on Tristylium ochnaceum, TARI. 

Dialeurodes (Rusostigma) tristylii var. uichancoi (Takahashi) 1936; Takahashi 1936b: 218. 

Syntypes. Taiwan: on Tristylium ochnaceum, TARI. 

Rusostigma tristylii (Takahashi); Takahashi & Mamet 1963: 49. 

Distribution: 5: Japan; Taiwan; 6: Philippines. 

Host: Myrsinaceae: Ardisia sp.; Theaceae: Cleyera ochnaceum, Eurya sp. 

422

Russellaleyrodes David 1973 

Russellaleyrodes David 1973: 557. Type species. Dialeurodes cumiugum Singh 1932, by monotypy. 



cumiugum x 

Total = 1 species 0 0 0 0 0 1 0 0 0 

Russellaleyrodes cumiugum (Singh) 1932 

Dialeurodes cumiugum Singh 1932: 85. Syntypes. Burma: Kalaw, K.S.L, on undetermined herb, ZSI. 

Dialeuropora cumiugum (Singh); Russell 1959: 185. 

Russellaleyrodes cumiugum (Singh); David 1973: 557. 


Host: on undetermined herb. 

423

Rutaleyrodes Dubey & Ko 2007 

Rutaleyrodes Dubey & Ko 2007: in press. Type species – Rutaleyrodes atalantiae Dubey & Ko 

2007, by original designation. 


atalantiae x 

Total= 1 species 0 0 0 0 0 1 0 0 0 

Rutaleyrodes atalantiae Dubey & Ko 2007: in press 

Rutaleyrodes atalantiae Dubey & Ko 2007: in press. Type species – Rutaleyrodes atalantiae 

Dubey & Ko 2007, holotype, India: Chennai, on Atalantia racemosa, 16.v.2004, A. K. 

Dubey, NTU. 


Host: Rutaceae: Atalantiae racemosa. 

424

Septaleurodicus Sampson 1943 

Septaleurodicus Sampson 1943: 189-190. Type-species: Septaleurodes mexicanus Sampson 1943, by 

monotypy. 

Comment: Martin (2004) transferred this genus to Aleyrodinae from Aleurodicinae. 



mexicanus x 

Total = 1 species 0 1 0 0 0 0 0 0 0 

Septaleurodicus mexicanus Sampson 1943 

Septaleurodicus mexicanus Sampson 1943: 190-191. Syntypes. Mexico: Chapultepec Park, 12.vi.1924, A. 

Dampf, on weeds; adult paratypes swept from Senecio salignus, Chapingo, 25.ii.1924 ? 


Host: Asteraceae: Senecio salignus. 

425

Setaleyrodes Takahashi 1931 

Setaleyrodes Takahashi 1931c: 221. Type species. Setaleyrodes mirabilis Takahashi 1931, by monotypy. 



litseae x 

mirabilis x 

quercicola x 

takahashia x 

thretaonai x 

vigintiseta x 

Total = 6 species 0 0 0 0 2 3 1 0 0 

Setaleyrodes litseae David & Sundararaj 1991 

Setaleyrodes litseae David & Sundararaj 1991: 317. Holotype puparium, India, Meghalaya, Shillong, 

29.ix.1988, B. David, on Litsea sp., IARI (paratypes in USNM, BMNH). 


Host: Lauraceae: Litsea sp. 

Setaleyrodes mirabilis Takahashi 1931 

Setaleyrodes mirabilis Takahashi 1931c: 222. Syntypes. Taiwan: Shikikun near Taihezan, 22.v.1931, R. 

Takahashi, on Ficus sp, TARI. 


Host: Menispermaceae: Stephania japonica; Moraceae: Ficus foveolata, Ficus sp. 

Setaleyrodes quercicola Takahashi 1934 

Setaleyrodes quercicola Takahashi 1934b: 65. Syntypes. Taiwan: Kurasu, Kahodai, Tosei-Gun, 6-7.vi.1933, 

R. Takahashi, on Quercus variabilis, TARI. 


Host: Fagaceae: Quercus variabilis. 

426

Setaleyrodes takahashia Singh 1933 

Setaleyrodes takahashia Singh 1933: 344. Syntypes. Burma: Kalaw, xii.1929, K. Singh, ex. Strebulus asper, 

ZSI. 



Reference: David & Jesudasan 1987b - redescription and illustration. 

Setaleyrodes thretaonai David 1981 

Setaleyrodes thretaonai David 1981: 37. Holotype pupal case. India: Kerala, Walayar, 24.iii.1972, B.V. 

David, on Homonoia riparia, IDAV. 



Setaleyrodes vigintiseta Martin 1999 

Setaleyrodes vigintiseta Martin 1999: 99. Holotype puparium. Australia: on Acacia ?aulacocarpa, ANIC. 


Host: Fabaceae: Acacia aulacocarpa? 

427

Simplaleurodes Goux 1945 

Simplaleurodes Goux 1945: 186. Type species. Simplaleurodes hemisphaerica Goux 1945, by monotypy. 



hemisphaerica x 

Total = 1 species 0 0 1 0 0 0 0 0 0 

Simplaleurodes hemisphaerica Goux 1945 

Simplaleurodes hemisphaerica Goux 1945: 186. Holotype and paratypes. France: Marseille, 

Bouches-du-Rhone, Chateau-Neuf-les Martigues, v.1933, on Philyrea media. 

Distribution: 3: France; Italy. 

Host: Oleaceae: Phillyrea media, Phillyrea angustifolia, Phillyrea latifolia. 

Natural enemies: Aphelinidae: Encarsia arabica; Platygastridae: Amitus rugosus, 

428

Singhiella Sampson 1943 

Singhiella Sampson 1943: 211. Type species Trialeurodes bicolor Singh 1931: 50, by original designation and 

monotypy. 

References - Jensen (2001); Martin (2005); Dubey & Sundararaj (2006e) Key to Indian species 



bassiae x 

bauhiniae x 

bicolor x 

brideliae x 

cambodiensis x 

cardamomi x 

chinensis x 

chitinosa x 

citrifolii x x x x x x x 

crenulata x 

delamarei x 

denticulata x 

dioscoreae x 

dipterocarpi x 

elaeagni x 

elbaensis x x 

ficifolii x 

kuraruensis x 

longisetae x 

malabaricus x 

mekonensis x 

melanolepis x 

pallida x 

429


piperis x 

premnae x 

serdangensis x 

simplex x 

subrotunda x 

sutepensis x 

tetrastigmae x 

vanieriae x 

Total = 30 species 1 1 2 2 12 17 1 1 1 

Singhiella bassiae (David and Subramaniam) 1976 

Dialeurodes bassiae David & Subramaniam 1976: 185. Holotype. India: Coimbatore, 7.xi.1966, B. David, on 

on Bassia longifolia, ZSI. 

Kanakarajiella bassiae (David and Subramaniam); David & Sundararaj 1993: 292. 

Singhiella bassiae (David and Subramaniam); Jensen 2001: 307. 


Host: Amaranthaceae (Chenopodiaceae): Bassia longifolia, Bassia latifolia. 

Singhiella bauhiniae Dubey & Sundararaj 2006 

Singhiella bauhiniae Dubey & Sundararaj 2006e: 232. Holotype puparium, India, Karnataka, Kudremukh 

National Park, 7.VIII.2001, A. K. Dubey, ex. Bauhinia phoenicea, IWST, will be transferred to 

NFICFRI. 


Host: Fabaceae: Bauhinia phoenicea. 

Singhiella bicolor (Singh) 1931 

Trialeurodes bicolor Singh 1931: 50. Syntypes. India: Pusa, Bihar, Muzaffarpur, Saharanpur, Miani, Punjab, on 

Eugenia jambos, BMNH. 

Singhiella bicolor (Singh); Sampson 1943: 211; Mound and Halsey 1978; Jensen 2001: 307. 


Host: Fabaceae: Lathyrus sp.; Moraceae: Ficus benghalensis; Myrtaceae: Eugenia jambos, Eugenia jambolana, 

Syzigium jambolanum, Szygium cuminii. 

430

Natural enemies: Aphelinidae: Encarsia septentrionalis. 

Singhiella brideliae (Jesudasan and David) 1991 

Dialeurodes brideliae Jesudasan & David 1991. Holotype puparium, India, Tamil Nadu, Yercaud, 24.xi.1984, 

RW Alexander Jesudasan, on Bridelia crenulata. 

Singhiella brideliae (Jesudasan & David); Jensen 2001: 307. 


Host: Euphorbiaceae: Bridelia crenulata. 

Note: Do not confuse this species with Dialeurodes (Dialeuropora) brideliae Takahashi 1932. 

Singhiella cambodiensis (Takahashi) 1942 

Dialeurodes cambodiensis Takahashi 1942g: 305. Syntypes. Cambodia and Thailand: on unidentified plant. 

Singhiella cambodiensis (Takahashi); Jensen 2001: 307. 

Distribution: 6: Cambodia; Thailand. 


Singhiella cardamomi (David & Subramaniam) 1976 

Dialeurodes cardamomi David & Subramaniam 1976: 187. Holotype. India: on Elettaria cardomomum, IDAV. 

Aleuroclava cardamomi (David & Subramaniam); Jesudasan and David 1990: 3. 

Kanakarajiella cardamomi (David and Subramaniam); David & Sundararaj 1993: 293 

Singhiella cardamomi (David and Subramaniam); Jensen 2001: 307. 


Host: Marantaceae: Maranta arundinacea; Zingiberaceae: Elettaria cardamomum 

Natural enemies: Fungus: Aschersonia placenta. 

Singhiella chinensis (Takahashi) 1941 

Pealius chinensis Takahashi 1941c: 388. Syntypes (3 pupal cases). Hong Kong: R. Takahashi, on unidentified 

tree. 

Aleuroputeus chinensis Takahashi; Mound & Halsey 1978: 61. 

Singhiella chinensis (Takahashi); Jensen 2001: 281. 



Singhiella chitinosa (Takahashi) 1937 

Dialeurodes chitinosa Takahashi 1937b: 21. Syntypes. China: on Cinnamomum sp., TARI. 

431

Singhiella chitinosa (Takahashi); Jensen 2001: 307 



Singhiella citrifolii (Morgan) 1893 

Aleyrodes citrifolii Morgan 1893: 70. Syntypes. USA: Louisiana, on Citrus sp., BMNH. 

Dialeurodes citrifolii (Morgan ); Quaintance & Baker 1914: 97. 

Singhiella citrifolii (Morgan); Jensen 2001: 307. 

Distribution: 1: USA (AL, AR, CA, CO, FL, KY, LA, MD, MS, MO, NC, PA, TX); 2: Argentina, Bahamas; 

Barbados, Bermuda, Brazil; Colombia; Costa Rica; Cuba; Dominican Republic, Grand Cayman Islands; 

Guatemala, Haiti; Honduras; Jamaica; Mexico; Panama, Puerto Rico; Trinidad & Tobago, Venezuela, 

Virgin Islands; 3: Canary Islands, Iran, Lebanon, Madeira, Morocco; 5: China; Hong Kong; 6: India; 

Malaysia, Singapore, Vietnam; 8: Caroline Islands, Mariana; 9: Hawaii. 

Host: Anacardiaceae: Schinus terebinthifolius; Annonaceae: Annona sp.; Araceae: Monstera deliciosa; 

Arecaceae: Chamaedorea sp., Cocos nucifera; Asteraceae: Eupatorium odoratum; Caprifoliaceae: 

Viburnum odoratissimum, Viburnum sp.; Dioscoreaceae: Dioscorea alata, Dioscorea bulbifera, 

Dioscorea sp.; Labiatae: Ocimum sp.; Malpighiaceae: Malpighia glabra; Moraceae: Ficus nitida, Ficus 

sp.; Myrtaceae: Psidium sp.; Oleaceae: Fraxinus sp, Jasminum sp.; Rubiaceae: Cephalanthus sp., 

Gardenia augusta, Gardenia jasminoides, Gardenia sp.; Rutaceae: Citrus aurantifolia, Citrus 

aurantium, Citrus limon, Citrus medica, Citrus nobilis, Citrus reticulata, Citrus sp., Citrus x nobilis, 

Citrus x paradisi, Fortunella crassifolia, Poncirus sp., X Citrofortunella microcarpa, Solanaceae: 

Cestrum nocturnum; Vitaceae: Vitis sp. 

Natural enemies: Fungus - Aegerita webberi, Aschersonia aleyrodis, Aschersonia goldiana, 

Fusarium verticilloides; Predators - ACARI/ Phytoseiidae: Typhlodromalus peregrinus. 

COLEOPTERA/ Coccinellidae: Brumoides sp, Brumoides suturalis, Delphastus catalinae, Delphastus 

pallidus, Delphastus pusillus, Delphastus sp., Nephaspis oculatus, Scymnus sp.; DIPTERA/Syrphidae: 

Ocyptamus fuscipennis; NEUROPTERA/ Chrysopidae: Ceraeochrysa cincta, Ceraeochrysa cubana, 

Ceraeochrysa valida, Chrysopa sp.; Coniopterygidae: Coniopteryx vicina; THYSANOPTERA/ 

Phlaeothripidae: Aleurodothrips fasciapennis; Parasitoids - HYMENOPTERA: Aphelinidae: Encarsia 

lahorensis, Encarsia luteola, Encarsia perstrenua, Encarsia protransvena, Encarsia sophia, Encarsia 

variegata and Encarsia sp. 

US Port Interceptions: Albania on Citrus sp.; Bolivia on Citrus limon; El Salvador on Citrus sp.; 

Guatemala on Citrus sp., Persea americana; Haiti on Citrus sinensis; Italy on Citrus sp.; 

Mexico on Citrullus, Citrus reticulata, Citrus sinensis; Nicaragua on Citrus sinensis; 

Pakistan on Citrus sp.; Panama on Citrus reticulata; Puerto Rico on Citrus sp. 

Singhiella crenulata Qureshi & Qayyum 1969 

Singhiella crenulata Qureschi & Qayyam 1969: 178. Holotype. Pakistan, Lyallpur, on Rosa indica. 

432


Host: Rosaceae: Rosa indica. 

Singhiella delamarei (Cohic) 1968 

Dialeurodes delamarei Cohic 1968a: 43. Syntypes. Congo: on Uvaria scabrida. 

Singhiella delamarei (Cohic); Jensen 2001: 307. 



Singhiella denticulata Qureshi 1979 

Singhiella denticulata Qureshi 1979: 17. Holotype pupal case, Pakistan, Rawalpindi, 13.vi.1973, J. I. Qureshi, 

on Cassia fistula, FAI (Pakistan). 


Host: Fabaceae: Cassia fistula. 

Singhiella dioscoreae (Takahashi) 1934 

Dialeurodes dioscoreae Takahashi 1934b: 43. Syntypes. Taiwan: on Dioscorea rhipogonioides, TARI. 

Singhiella dioscoreae (Takahashi); Jensen 2001: 307. 


Dioscoreaceae: Dioscorea rhipogonioides, Dioscorea sp. 

Natural enemies: Aphelinidae: Encarsia abundantia. 

Singhiella dipterocarpi (Takahashi) 1942 

Dialeurodes dipterocarpi Takahashi 1942g: 304. Syntypes. Thailand: on Dipterocarpus sp., TARI. 

Singhiella dipterocarpi (Takahashi); Jensen 2001: 307. 


Host: Dipterocarpaceae: Dipterocarpus sp. 

Singhiella elaeagni (Takahashi) 1935 

Dialeurodes elaeagni Takahashi 1935b: 45. Syntypes. Taiwan: on Elaeagnus formosana, TARI. 

Singhiella elaeagni (Takahashi); Jensen 2001: 307. 


Host: Elaeagnaceae: Elaeagnus formosana. 

433

Singhiella elbaensis (Priesner and Hosny) 1934 

Dialeurodes (Gigaleurodes) elbaensis Priesner & Hosny 1934b: 10. Syntypes. Egypt: on Ficus salicifolia, 

EDAC; Ardaillon & Cohic 1970. 

Singhiella elbaensis (Priesner and Hosny); Jensen 2001: 307. 

Distribution: 3: Cameroon; Chad; Egypt; Ivory Coast; Niger; Sudan. 

Host: Anacardiaceae: Trichoscypha arborea; Annonaceae: Monodora crispata; Araceae: Cyrtosperma 

senegalense; Chrysobalanaceae: Chrysobalanus orbicularis; Connaraceae: Manotes pruinosa; 

Dilleniaceae: Tetracera alnifolia; Euphorbiaceae: Sapium cornutum; Guttiferae: Harungana 

madagascariensis; Liliaceae: Smilax fraussiana; Moraceae: Ficus sycomorus, Ficus salicifolia; 

Rubiaceae: Morelia senegalensis, Morinda morindoides. 

Singhiella ficifolii (Takahashi) 1942 

Dialeurodes ficifolii Takahashi 1942g: 300. Syntypes. Thailand: on Ficus sp., TARI. 

Massilieurodes ficifolii (Takahashi); Jensen 1999: 292. 

Singhiella ficifolii (Takahashi); Jensen 2001: 307. 



Singhiella kuraruensis (Takahashi) 1933 

Dialeurodes (Rhachisphora) kuraruensis Takahashi 1933: 5. Syntypes. Taiwan: Machilus sp., TARI. 

Rhachisphora kuraruensis (Takahashi); Mound and Halsey 1978: 187. 

Singhiella kuraruensis (Takahashi); Jensen 2001: 307. 



Singhiella longisetae Chou & Yan 1988 

Singhiella longisetae Chou & Yan 1988: 246. Holotype puparium, China, Dayong, Hunan, 14.x.1985, F. Yan, 

ex. Cinnamomum sp. 



Singhiella malabaricus (Jesudasan and David) 1991 

Dialeurodes malabaricus Jesudasan and David 1991: 305. Holotype pupal case. India: on Cinnamomum 

malabaricum, IDAV. 

Singhiella malabaricus (Jesudasan & David); Jensen 2001: 307. 

434


Host: Lauraceae: Cinnamomum malabaricum. 

Singhiella melanolepis Chen & Ko 2007 

Singhiella melanolepis Chen & Ko 2007: 3. Holotype puparium, Taiwan, Chiao-chi, 22.iii.2006, C. Chen and 

Y. Shih, on Melanolepis multiglandulosa, NTU. 


Host: Euphorbiaceae: Melanolepis multiglandulosa. 

Singhiella mekonensis (Takahashi) 1942 

Dialeurodes mekonensis Takahashi 1942g: 301. Syntypes. Thailand: on undetermined plant, TARI. 

Singhiella mekonensis (Takahashi); Jensen 2001: 307. 


Host: on undetermined plant. 

Singhiella pallida (Singh) 1931 

Dialeurodes pallida Singh 1931: 30. Syntypes. India: on Piper betle, ZSI. 

Kanakarajiella pallida (Singh); David & Sundararaj 1993: 294. 

Singhiella pallida (Singh); Jensen 2001: 307. 


Host: Marantaceae: Maranta arundinacea; Piperaceae: Piper betel; Zingiberaceae: Elettaria cardamomum. 

Natural enemies: Predator - DIPTERA/Syrphidae. 

Singhiella piperis (Takahashi) 1934 

Dialeurodes piperis Takahashi 1934b: 44. Syntypes. Taiwan; on Piper sp., TARI. 

Singhiella piperis (Takahashi); Jensen 2001: 307. 


Host: Piperaceae: Piper sp., Piper kasura. 

Natural enemies: Aphelinidae: Encarsia azimi, Encarsia inaron. 

Singhiella premnae Martin 1999 

Singhiella premnae Martin 1999: 100. Holotype puparium. Australia: on Premna serratifolia, ANIC. 


435

Host: Verbenaceae: Premna serratifolia. 

Singhiella serdangensis (Corbett) 1935 

Dialeurodes serdangensis Corbett 1935b: 754. Syntypes. Malaya: on Abelmoschus [=Hibiscus] esculentus. 

Singhiella serdangensis (Corbett); Jensen 2001: 307. 


Host: Malvaceae: Abelmoschus esculentus. 

Singhiella simplex (Singh) 1931 

Aleurocanthus simplex Singh 1931: 69-70. Syntypes. India: Pusa, on Ficus bengalensis, ZSI. 

Pealius simplex (Singh); Jesudasan and David 1991: 321; Dubey & Sundararaj 2005a: 163. 

Singhiella simplex (Singh); Martin & Mound 2007: 43. 

Dialeurodes glomerata Singh 1932: 81. Syntypes. India, on Ficus glomerata, ZSI; synonymy according to 

Jesudasan and David 1991: 321. 

Pealius indicus David & Subramaniam 1976: 206. Holotype. India: on Ficus bengalensis, IDAV; synonymy 

according to Jesudasan and David 1991: 321. 


Host: Ericaceae: Rhododendron indica; Moraceae: Ficus altissima, Ficus bengalensis, Ficus glomerata, Ficus 

benjamina. 

Natural enemies: Aphelinidae: Encarsia tricolor. 

Singhiella subrotunda (Takahashi) 1935 

Dialeurodes subrotunda Takahashi 1935b: 41. Syntypes. Taiwan: on Actinodaphne sp. and Machilus sp., TARI. 

Singhiella subrotunda (Takahashi); Jensen 2001: 307. 


Host: Lauraceae: Actinodaphne sp., Machilus sp. 

Singhiella sutepensis (Takahashi) 1942 

Dialeurodes sutepensis Takahashi 1942e: 275. Syntypes. Thailand and Cambodia: on unidentified host, TARI. 

Singhiella sutepensis (Takahashi); Jensen 2001: 307. 

Distribution: 6: Cambodia; Thailand. 


436

Singhiella tetrastigmae (Takahashi) 1934 

Dialeurodes tetrastigmae Takahashi 1934b: 39. Syntypes. Taiwan: on Tetrastigma umbellata, TARI. 

Singhiella tetrastigmae (Takahashi); Jensen 2001: 307. 


Host: Convolvulaceae: Erycibe henryi; Vitaceae: Tetrastigma formosanum, Tetrastigma umbellata. 

Natural enemies: Aphelinidae: Encarsia abundantia, Encarsia longivalvula, Encarsia magnivena. 

Singhiella vanieriae (Takahashi) 1935 

Dialeurodes vanierieae Takahashi 1935b: 46. Syntypes. Taiwan: on Cudrania [=Vanieria] cochinchinensis, 

TARI. 

Singhiella vanieriae (Takahashi); Jensen 1999: 307. 


Host: Moraceae: Cudrania cochinchinensis. 

437

Singhius Takahashi 1932 

Singhius Takahashi 1932:14. Type species. Aleyrodes hibisci Kotinsky 1907, by monotypy. 

Dialeurodes (Singhius) Takahashi 1932: 14. 

Singhius Takahashi; subgenus elevated to genus by Mound & Halsey 1978: 191. 

Reference: Sundararaj & David (1992b) - key to Indian species. 



ehretiae x 

hibisci x x x x x x 

morindae x 

russellae x 

Total = 4 species 1 1 0 0 1 4 0 1 1 

Singhius ehretiae (Jesudasan & David) 1991 

Aleuroclava ehretiae Jesudasan and David 1991: 258-259. Holotype. India: Tamil Nadu, Tambaram, 4.xi.1984, 

R. W. Alexander Jesudasan, on Ehretia sp. 

Singhius ehretiae (Jesudasan & David); Dubey & Sundararaj 2005d: 251. 


Host: Apocynaceae: Tabernaemontana divaricata; Boraginaceae: Ehretia sp. 

Singhius hibisci (Kotinsky) 1907 

Aleyrodes hibisci Kotinsky 1907: 96. Syntypes. USA: Hawaii, on Paritium tiliaceum and Hibiscus rosasinensis, 

HDA. 

Pealius hibisci (Kotinsky); Quaintance & Baker 1914: 99. 

Singhius hibisci (Kotinsky); 1932:14; Corbett 1935b; Cohic 1959b; Mound and Halsey 1978: 191. 

Dialeurodes fletcheri Singh 1931: 39. Syntypes. India: on Breynia rhamnoides; Takahashi 1942g. 

Distribution: 1: USA (FL, HI, TX); 2: Trinidad & Tobago; 5: China; Hong Kong, Taiwan; 6: India; Cambodia; 

Malaya; Malaysia, Philippines, Thailand; 7: Indonesia, 8: New Caledonia; 9: Hawaii. 

Hosts: Annonaceae: Fissistigma oldhami; Convolvulaceae: Ipomoea batatas; Euphorbiaceae: Baccaurea 

motleyana, Breynia rhamnoides, Bridelia monoica, Glochidion hongkongensis, Macaranga tanarius, 

Sapium sebiferum; Lauraceae: Cinnamomum camphora; Malvaceae: Hibiscus tiliaceus; Hibiscus rosasinensis; 

Moraceae: Ficus elastica; Oleaceae: Forsythia suspensa, Jasminum sp; Rubiaceae: Gardenia 

sp.; Salicaceae: Salix sp.; Solanaceae: Physalis peruviana; Ulmaceae: Celtis sinensis; Vitaceae: Vitis 

438

vinifera. 

Natural enemies: Aphelinidae: Encarsia bothrocera; Encarsia cibcensis; Encarsia lutea; Encarsia perflava, 

Encarsia sophia; Predators - Coccinellidae: Coccinella transversalis 

US Port Interceptions: India on Murraya sp.; Philippines on Mangifera indica; Tahiti on Hibiscus sp.; 

Thailand on Tecoma sp. 

Singhius morindae Sundararaj & David 1992 

Singhius morindae Sundararaj & David 1992: 103. Holotype puparium, India, Tamil Nadu, Vellimalai, 

3.viii.1987, R. Sundararaj, on Morinda tinctoria, David Collection. 


Host: Rubiaceae: Morinda tinctoria. 

Singhius russellae (David & Subramaniam) 1976 

Aleurotuberculatus russellae David & Subramaniam 1976: 172. Holotype. India: on unidentified tree. 

Singhius russellae (David & Subramaniam); Alexander & David 1990: 1. 



439

Siphoninus Silvestri 1915 

Siphoninus Silvestri 1915a: 245. Type species: Siphoninus finitimus Silvestri 1915, regarded by Mound & 

Halsey 1978:191 as synonym of S. phillyreae, by original designation. 



gruveli x 

immaculatus x 

phillyreae x x x x x x x 

Total = 3 species 1 1 1 1 1 0 1 0 1 

Siphoninus gruveli Cohic 1968 

Siphoninus gruveli Cohic 1968a: 50. Syntypes. Chad: on Combretum glutinosum. 

Siphoninus gruveli Cohic; species incertae sedis; Bink-Moenen 1983: 167. 


Host: Combretaceae: Combretum glutinosum. 

Siphoninus immaculatus (Heeger) 1856. 

Aleurodes immaculatus Heeger 1856: 33. Syntypes. Germany?: on Hedera helix. 

Aleurochiton immaculatus (Heeger); Quaintance & Baker 1914: 105. 

Trialeurodes immaculatus (Heeger); Quaintance & Baker 1915a: xi. 

Siphoninus immaculatus (Heeger); Trehan 1940: 601. 

Aleurodes immaculatus Heeger 1856: 33. Syntypes. Germany?: on Hedera helix. 

Siphoninus heegeri Haupt 1935: 259; synonymy according to Zahradnik 1963a: 9. 

Distribution: 3: Austria; Czech Republic; England; Germany; Hungary; Iran, Italy; Sweden; USSR. 

Host: Araliaceae: Hedera canariensis, Hedera helix; Rutaceae: Citrus sp. 

Natural enemies: Aphelinidae: Encarsia elegans, Encarsia inaron; Predators - Tortricidae: Clepsis consimilana 

Coccinellidae: Clitostethus arcuatus. 

Siphoninus phillyreae (Haliday) 1835 

Aleurodes phillyreae Haliday 1835: 119. Syntypes. Ireland: on Phillyrea latifolia, HAL. 

440

Trialeurodes phillyreae (Haliday); Quaintance & Baker 1915a: xi. 

Siphoninus phillyreae (Haliday); Silvestri 1915a: 247. 

Siphoninus phillyreae inequalis Goux 1949: 11. Syntypes. France: on pear [Pyrus sp.]. 

Siphoninus phillyreae mulititubulatus Goux 1949: 11. Syntypes. Corsica: on Olea europea. 

Siphoninus phillyreae mulititubulatus Goux; Mound and Halsey 1978: 192. 

Aleurodes dubia Heeger 1859: 223. Syntypes, Germany?, on Fraxinus sp.; Frauenfeld 1867: 796. 

Aleurochiton dubius (Heeger); Quaintance & Baker 1914: 105. 

Siphoninus dubiosa Haupt 1935: 259; synonymy according to Zahradnik 1963a: 9. 

Aleurodes phylliceae Bouche 1851: 110. Syntypes. Southern Europe: on Phillyrea latifolia? 

Aleurodes phylliceae Bouche 1851; Frauenfeld 1867: 786. 

Asterochiton phillyreae (Haliday); Quaintance & Baker 1914: 105. 

Siphoninus finitimus Silvestri 1915a: 245. Syntypes. Eritrea: on Olea chrysophylla, IESP. 

Siphoninus finitimus Silvestri; Mound and Halsey 1978: 192. 

Siphoninus granati Priesner & Hosny 1932: 1. Syntypes. Egypt: Meadi, 16.viii.1931, Priesner & Hosny, on 

Punica granatum, EDAC 

Siphoninus granati Priesner & Hosny; Mound and Halsey 1978: 192. 

Trialeurodes inaequalis Gautier 1923: 339. Syntypes. France: on Pyrus sp.; synonymy according to Mound and 

Halsey 1978: 192. 

Distribution: 1: USA (CA, GA, HI, NC); 2: Mexico, Peru, Venezuela; 3: Bulgaria; Corsica; Cyprus, England; 

Egypt; Finland; France; Germany; Greece; Hungary, Iran; Ireland; Israel; Italy; Jordan; Spain; Syria; 

USSR; Yugoslavia; 4: Cameroon; Eritrea; Ethiopia; Sudan; Zaire; 5: Taiwan; 7: Australia, Java. 

Host: Araliaceae: Hedera canariensis; Fabaceae: Afzelia sp.; Myrtaceae: Psidium guajava. 

Oleaceae: Fraxinus americana, Fraxinus excelsior, Fraxinus syriacata, Fraxinus sp, Olea chrysophylla, 

Olea europaea, Olea europaea, Phillyrea latifolia; Punicaceae: Punica granatum; Rhamnaceae: 

Rhamnus alaternus, Ziziphus spina-christi; Rosaceae: Crataegus mollis, Crataegus monogyna, 

Crataegus microphylla, Crataegus sp., Cydonia oblonga, Malus sp., Rosaceae: Mespilus sp., Prunus 

persica, Pyrus communis, Pyrus malus, Pyrus sativa, Pyrus sp.; Rutaceae: Citrus aurantifolia, Citrus 

sp.; Ulmaceae: Ulmus carpinifolia. 

Natural enemies: Parasitoids - Hymenoptera: Aphelinidae: Encarsia davidi, Encarsia dichroa, Encarsia galilea, 

Encarsia gautieri, Encarsia hispida, Encarsia inaron, Encarsia near brasiliensis, Encarsia punicae, 

Encarsia siphonini, Encarsia strenua, Eretmocerus siphonini. 

Predators - COLEOPTERA/Coccinellidae: Clitostethus arcuatus, Menochilus sexmaculatus, Menochilus sp, 

Scymnus smithianus; DIPTERA/ Drosophilidae: Acletoxenus formosus, Acletoxenus indicus, 

Acletoxenus sp.; Empididae: Drapetis ghesquierei. 

US Port Interceptions: Italy and Mexico on Punica granatum. 

441

Sphericaleyrodes Selvakumaran and David 1996 

Sphericaleyrodes Selvakumaran and David 1996: 58. Type species - Sphericaleyrodes bambusae Selvakumaran 

and David 1996, by monotypy and original designation. 



bambusae x 

regui x 

Total = 2 species 0 0 0 0 0 2 0 0 0 

Sphericaleyrodes bambusae Selvakumaran and David 1996 

Sphericaleyrodes bambusae Selvakumaran and David 1996: 58. Holotype puparium, India, on Bambusa sp. 



Sphericaleyrodes regui Dubey & Sundararaj 2006 

Sphericaleyrodes regui Dubey & Sundararaj 2006: Holotype puparium, India, Kerala, Calicut, 10.ix.1999, K. 

Regu, ex unidentified grass, FRI. 



442

Tegmaleurodes Martin 2005 

Tegmaleurodes Martin 2005b: 47. Type species Tegmaleurodes lentus Martin, by original description and 

monotypy. 



crustatus x 

integellus x 

lentus x 

Total = 3 species 0 3 0 0 0 0 0 0 0 

Tegmaleurodes crustatus (Bondar) 1928 

Aleuroplatus crustatus Bondar 1928b: 25. Syntypes. Brazil: on Eugenia sp., MZU. 

Tegmaleurodes crustatus (Bondar); Martin 2005b:47. 


Host: Myrtaceae: Eugenia sp, Psidium cymosum. 

Tegmaleurodes integellus (Bondar) 1923 

Aleuroplatus integellus Bondar 1923a: 115. Syntypes. Brazil: on Chomelia oligantha, MZU. 

Tegmaleurodes integellus (Bondar); Martin 2005b: 47. 


Host: Rubiaceae: Chomelia oligantha. 

Tegmaleurodes lentus Martin 2005 

Tegmaleurodes lentus Martin 2005b: 48. Holotype puparium, Belize, CFR, 25.iii.2003, J. H. Martin, 

on Pouteria reticulata, BMNH. 


Host: Lauraceae: Lauraceae sp.; Sapotaceae: Manilkara zapota, Pouteria reticulata. 

443

Tetraleurodes Cockerell 1902 

Aleyrodes (Tetraleurodes) Cockerell 1902a: 283. Type species: Aleyrodes (Tetraleurodes) perileuca 

Cockerell 1902, by original designation. 

Tetraleurodes Cockerell 1902; elevated to full genus by Quaintance & Baker 1914: 107 

Mexicaleyrodes Sampson & Drews 1941: 166. Type species. Mexicaleyrodes contigua Sampson & 

Drews 1941, by monotypy; synonymy according to Martin 2005b: 50. 

References: Nakahara 1995 - Key to Nearctic species; Bink-Moenen 1983 - Key to African species; 

Jesudasan and David 1991; Dubey & Sundararaj 2005 - Key to Indian species. 



acaciae x x x x x 

adabicola x 

andropogoni x 

bambusae x 

banksi x 

bararakae x 

bicolor x 

bidentatus x 

bireflexa x 

burliarensis x 

cacaorum x 

caulicola x 

chivela x 

confusa x x 

contigua x 

corni x 

cruzi x 

dendrocalamae x 

444


dorsirugosa x 

elaeocarpi x 

fici x x 

ghesquierei x 

granulata x 

graminis x 

hederae x 

hirsuta x 

kunnathoorensis x 

leguminicola x x 


malayensis x 

mameti x 

marshalli x 

melanops x 

mexicana x 

mirabilis x 

monnioti x 

mori x x x x x 

moundi x 

neemani x 

oplismeni x 

pauliani x 

perileuca x 

perseae x x 

pluto x 

pringlei x 

pseudacaciae x x 

psidii x 

445


pusana x 

quadratus x 

quercicola x x 

rugosus x 

russellae x 

selachidentata x 

semibarbata x 

simplicior x 

stellata x 

stirlingiae x 

submarginata x 

subrotunda x 

sulcistriatus x 

supraxialis x 

thenmozhiae x 

truncatus x 

tuberculata x 

tuberculosa x 

ursorum x x 

Total = 65 species 13 21 5 19 3 11 4 5 0 

Tetraleurodes acaciae (Quaintance) 1900 

Aleurodes acaciae Quaintance 1900: 19. Syntypes. Mexico: on mesquite [Acacia sp.], USNM. 

Tetraleurodes acaciae (Quaintance); Quaintance & Baker 1914: 108. 

Tetraleurodes herberti Penny 1922: 32. Syntypes. USA: California: Pleasanton, x.1918, F. Herbert, on Robinia 

pseudoacaciae; synonymy according to Nakahara 1995: 108. 

Distribution: 1: USA ( AZ, CA, FL, TX); 2: Belize; Colombia; Costa Rica; Cuba; Dominican Republic; El 

Salvador; Guatemala; Honduras; Jamaica; Mexico; Nicaragua; Panama; Puerto Rico; Trinidad; 

Venezuela; 5: Hong Kong; 6: Philippines; 8: Fiji. 

Host: Anacardiaceae: Rhus sp.; Anacardiaceae: Spondias sp.; Araceae: Symplocarpus sp. 

Arecaceae: Chamaedorea sp.; Bignoniaceae: Bignonia capreolata; Bombacaceae: Pachira sp., 

Pseudobombax ellipticum; Burseraceae: Bursera microphylla; Euphorbiaceae: Chamaesyce sp., 

Cnidoscolus chayamansa, Jatropha gossypifolia; Fabaceae: Acacia baileyana, Acacia collinsii, Acacia 

446

447 

koa, Acacia linearis, Acacia melanoxylon, Acacia pravissima, Acacia scopoides, Acacia sp., 

Aeschynomene sp., Albizia sp., Amorpha californica, Bauhinia fassoglensis Kotschy ex Schweinf, 

Bauhinia mexicana, Bauhinia petersiana, Bauhinia purpurea L, Bauhinia sp., Bauhinia variegata L, 

Bauhinia x blakeana, Bolusanthus speciosus, Brya ebenus, Caesalpinia gilliesii, Caesalpinia mexicana 

Gray, Caesalpinia sp., Calliandra haematocephala, Calliandra sp., Calliandra surinamensis, Cassia 

alata, Cassia aspera, Cassia fasciculata, Cassia fistula, Cassia sp., Centrosema pubescens, Centrosema 

sp., Centrosema virginianum, Ceratonia siliqua, Ceratonia siliqua, Cercis canadensis, Cercis 

siliquastrum, Chamaecrista sp., Cladrastis lutea, Coursetia microphylla, Cytisus scoparius, Cytisus 

sessilifolium, Dalbergia ecastophyllum, Dalbergia sissoo, Desmodium sp., Diphysa robinoides, 

Erythrina abyssinica, Erythrina americana, Erythrina anthocarpa, Erythrina corallodendron, Erythrina 

coralloides, Erythrina crista, Erythrina crista-galli, Erythrina eggersii, Erythrina falcata, Erythrina 

fusca, Erythrina humeana, Erythrina sp., Erythrina tahitensis, Erythrina variegata, Erythrina 

vespertilio, Galactia stricta, Gliricidia sepium, Glycine max, Gymnocladus dioicus, Hardenbergia 

comptoniana, Hardenbergia violacea, Indigofera gerardiana, Inga sp., Inga vera, Kennedia rubicunda, 

Leucaena glauca, Lonchocarpus violaceus, Maackia amurensis, Millettia dura, Mimosa sp., Olneya 

tesota Gray, Pachyrhizus tuberosus, Parkia sp., Phaseolus vulgaris L, Piscidia piscipula (L.) Sarg, 

Pithecellobium dulce, Pithecellobium flexicaule, Pithecellobium sp., Pithecellobium unguis-cati, 

Pongamia pinnata, Prosopis glandulosa, Prosopis pubescens Benth, Robinia hispida L, Robinia hispida 

L. var. kelseyi, Robinia pseudoacacia, Senna alata, Sophora japonica L, Sophora secundiflora, Sophora 

tomentosa, Sophora viciifolia, Tamarindus indica, Templetonia retusa, Tephrosia cinerea, Tephrosia 

heydeana, Wisteria sp.; Geraniaceae: Pelargonium sp.; Moraceae: Ficus sp., Morus sp.; Musaceae: 

Musa sp.; Myricaceae: Myrica sp.; Myrtaceae: Melaleuca sp.; Nyctaginaceae: Bougainvillea sp.; 

Rhamnaceae: Rhamnus californica, Rhamnus californica Eschsch. ssp. californica; Rosaceae: 

Cotoneaster ambiguus, Rosa sp.; Rutaceae: Citrus sinensis; Salicaceae: Populus sp.; Sapindaceae: 

Melicoccus bijugatus; Solanaceae: Capsicum sp.; Umbelliferae: Eryngium yuccifolium; Urticaceae: 

Parietaria sp. 

Natural enemies: Fungus: Aschersonia aleyrodis; Aphelinidae: Encarsia dispersa, Encarsia hispida, Encarsia 

lanceolata, Encarsia nigricephala, Encarsia paracitrella, Encarsia parvella group, Encarsia sp., 

Encarsia tabacivora, Eretmocerus picketti, Eretmocerus sp., Platygastridae: Amitus sp.; Signiphoridae: 

Signiphora aleyrodis (hyperparasitoid), Signiphora sp. 

US Port Interceptions: Dominican Republic on Cajanus cajan; Guatemala on Fabaceae sp.; Jamaica 

Plant; Mexico on Pithecellobium dulce; Philippines on Moringa oleifera. 

Tetraleurodes adabicola Takahashi 1955 

Tetraleurodes adabicola Takahashi 1955a: 414. Syntypes. Madagascar: on Ficus sakalavorum, MNHNP. 


Host: Moraceae: Ficus sakalavorum. 

Tetraleurodes andropogoni (Dozier) 1934 

Aleuroplatus andropogoni Dozier 1934: 185. Syntypes. Zaire: on Andropogon sp., BMNH; Mound & Halsey 

1978: 45. 

Tetraleurodes andropogoni (Dozier); Bink-Moenen 1983: 169.

Distribution: 4: Congo; Ivory Coast; Nigeria; Uganda; Zaire. 

Host: Annonaceae: Annona arenaria; Arecaceae: Elaeis guineensis, Cocos nucifera; Euphorbiaceae: 

Hymenocardia acida; Fabaceae: Cassia javanica; Flacourtiaceae: Caloncoba dusenii; Linaceae: Ochthocosmus 

africanus; Loranthaceae: Loranthus sp.; Poaceae: Andropogon sp.; Rutaceae: Zanthoxylum macrophylla. 

Tetraleurodes bambusae Jesudasan & David 1991 

Tetraleurodes bambusae Jesudasan and David 1991: 332. Holotype pupal case. India: on Bambusa sp., IDAV. 



Tetraleurodes banksi Martin 1999 

Tetraleurodes banksi Martin 1999: 103. Holotype puparium. Australia: on Banksia marginata, ANIC. 


Host: Proteaceae: Banksia marginata. 

Tetraleurodes bararakae Takahashi 1955 

Tetraleurodes bararakae Takahashi 1955a: 415. Holotype. Madagascar: on "bararaka", MNHNP. 


Host: on "bararaka". 

Tetraleurodes bicolor Bink-Moenen 1992 

Tetraleurodes bicolor Bink-Moenen and Gerling 1992: 32. Holotype. Israel: Mt. Meron, xi.1975, R. Neeman, 

on Myrtus communis, BMNH. 


Host: Myrtaceae: Myrtus communis. 

US Port Interceptions: Turkey on Spiraea sp.; United Arab Emirates on Anethum graveolens. 

Tetraleurodes bidentatus Sampson & Drews 1941 

Tetraleurodes bidentatus Sampson & Drews 1941: 172. Holotype pupal cases. Mexico: Oaxaca, Chivela, 

19.iv.1926, G.F Ferris, on undetermined tree, USNM. 



448

Tetraleurodes bireflexa Nakahara 1995 

Tetraleurodes bireflexa Nakahara 1995: 111. Holotype pupal case. USA: California, Imperial County, El 

Centro, 19.viii.1974, ex. mesquite [Prosopis sp.], USNM. 


Host: Fabaceae: Prosopis sp. 

Tetraleurodes burliarensis Jesudasan & David 1991 

Tetraleurodes burliarensis Jesudasan and David 1991: 332. Holotype pupal case. India: on unidentified tree, 

IDAV. 



Tetraleurodes cacaorum (Bondar) 1923 

Aleurotrachelus cacaorum Bondar 1923a: 154. Syntypes. Brazil: on Theobroma cacao, MZU. 

Tetraleurodes cacaorum (Bondar); Nakahara 1995: 135. 

Distribution: 2: Brazil; Colombia, Trinidad & Tobago. 

Host: Sterculiaceae: Theobroma cacao. 

Tetraleurodes caulicola Nakahara 1995 

Tetraleurodes caulicola Nakahara 1995: 113. Holotype pupal cases. USA: California, Riverside County, 

Baneby, Rancho Mirage, 4.v.1960, C. Keifer, on Psorothamnus spinosus (Gray), USNM. 

Distribution: 1: USA (AZ, CA); 2: Mexico. 

Host: Fabaceae: Cassia armata, Dalea sp., Psorothamnus emoryi, Psorothamnus spinosus. 

Tetraleurodes chivela Nakahara 1995 

Tetraleurodes chivela Nakahara 1995: 123. Holotype pupal cases. Mexico: Oaxaca, Chivela, iv.1926, G. F. 

Ferris, paratypes on Chamaedorea sp., USNM. 


Host: Arecaceae: Chamaedorea sp.; Myrtaceae: Eugenia? sp. 

Tetraleurodes confusa Nakahara 1995 

Tetraleurodes confusa Nakahara 1995: 125. Holotype pupal cases. USA: Florida, Palm City, 9.v.1979, E. W. 

Campbell, on Persea sp, USNM; Martin 2005b: 51. 

Distribution: 1: USA (FL, MS); 2: Belize, Puerto Rico. 

Host: Fagaceae: Quercus sp.; Lauraceae: Nectandra coriacea, Persea palustris, Persea borbonia, 

449

Persea sp.; Magnoliaceae: Magnolia virginiana; Theaceae: Gordonia lasianthus. 

Tetraleurodes contigua (Sampson & Drews) 1941 

Mexicaleyrodes contigua Sampson & Drews 1941: 168. Syntypes. Mexico: on undetermined shrub, UCD. 

Tetraleurodes contigua (Sampson & Drews); Martin 2005b: 50. 


Host: on undetermined shrub. 

Tetraleurodes corni (Haldeman) 1850 

Aleurodes corni Haldeman 1850: 109. Syntypes. USA: Pennsylvania, on Cornus sericea, USNM. 

Tetraleurodes corni (Haldeman); Quaintance & Baker 1914: 108. 

Distribution: 1: USA (PA). 

Host: Cornaceae: Cornus sericea; Lauraceae. 

Natural enemies: Aphelinidae: Eretmocerus corni; Platygastridae: Amitus aleurodinis. 

Tetraleurodes cruzi (Cassino) 1991 

Aleurotrachelus cruzi Cassino 1991: 194. Holotype puparium, Brazil, Piracicaba, 25.viii.1975, P. Cassino, on 

Citrus sp., Costa Lima Collection, Rio de Janeiro, Brazil; Cassino & Nascimento 1999: 78. 

Tetraleurodes cruzi (Cassino); Martin & Mound 2007: 44. 



Tetraleurodes dendrocalamae Dubey & Sundararaj 2005 

Tetraleurodes dendrocalamae Dubey & Sundararaj 2005b: 1924. Holotype puparium, India, Kudremukh 

National Park, Karnataka, 10 viii 2001, A. K. Dubey, on Dendrocalamus strictus, deposited FRI India. 


Host: Poaceae: Dendrocalamus strictus. 

Tetraleurodes dorsirugosa Nakahara 1995 

Tetraleurodes dorsirugosa Nakahara 1995: 116. Holotype pupal cases. Colombia: Caldas, Antioquia, ii.1974, R. 

Velaz-Angel, on Caesalpinaceae, USNM. 



450

Tetraleurodes elaeocarpi Takahashi 1950 

Tetraleurodes elaeocarpi Takahashi 1950: 87. Syntypes. Australia: on Elaecarpus reticulatus, BMNH. 


Host: Elaeocarpaceae: Elaeocarpus reticulatus. 

Tetraleurodes fici Quaintance & Baker 1937 

Tetraleurodes fici Quaintance & Baker 1937, in Baker 1937: 616. Syntypes. Mexico: on ?Ficus sp. 

Tetraleurodes nudus Sampson & Drews 1941: 174. Syntypes. Mexico: on Ficus involuta, UCD. 

synonymy according to Nakahara 1995: 135. 

Distribution: 1: USA (FL); 2: Bahamas, Belize, Colombia, Costa Rica, Cuba; Guatemala, Jamaica, Mexico, 

Nicaragua, Trinidad, Venezuela. 

Host: Euphorbiaceae: Cnidoscolus chayamansa; Lauraceae: Nectandra coriacea; Moraceae: Ficus aurea, Ficus 

elastica, Ficus altissima, Ficus benghalensis, Ficus benjamina, Ficus neriifolia, Ficus perforata, Ficus 

involuta; Myrtaceae: Pimenta sp.; Oleaceae: Fraxinus pennsylvanica; Urticaceae: Urera sp.; 

Verbenaceae: Lantana sp. 

US Port Interceptions: Cuba on unknown plant; Guatemala on Croton sp.; Mexico on Citrus sp., 

Cnidoscolus aconitifolius, Cnidoscolus chayamansa, Euphorbiaceae. 

Tetraleurodes ghesquierei Dozier 1934 

Tetraleurodes ghesquierei Dozier 1934: 186. Syntypes. Zaire: on Periplaca nigrescens, BMNH. 

Distribution: 4: Angola; Congo; Ivory Coast; Zaire. 

Host: Arecaceae: Microdesmis puberula Asclepidaceae: Periploca nigrescens; Malpighiaceae: Acridocarpus 

alternifolius. 

Tetraleurodes graminis Takahashi 1934 

Tetraleurodes graminis Takahashi 1934b: 67. Syntypes. Taiwan: on Graminae, ?Imperata sp., TARI. 


Host: Poaceae: Imperata sp.? 

Tetraleurodes granulata Bink-Moenen 1983 

Tetraleurodes granulata Bink-Moenen 1983:170. Holotype puparium, Chad, Bebedjia, 4.vii.1973, FAB & 

RMBM, on Hymenocardia acida, BMNH. 


Host: Annonaceae: Annona senegalensis, Hexalobus monopetalus; Combretaceae: Combretum 

hypopilinum, Combretum micranthum, Combretum molle, Terminalia laxiflora; 

Euphorbiaceae: Hymenocardia acida; Fabaceae: Detarium microcarpum; Loganiaceae: 

451

Strychnos spinosa; Rubiaceae: Crossopteryx febrifuga, Gardenia triacantha, Mitragyna 

inermis, Tarenna sp. 

Tetraleurodes hederae Goux 1939 

Tetraleurodes hederae Goux 1939: 77. Holotype. France: on Hedera helix. 

Distribution: 3: France; Iran, Italy; USSR. 

Host: Araliaceae: Hedera helix, Hedera taurica 

Natural enemies: Aphelinidae: Encarsia inaron, Encarsia tricolor. 

Tetraleurodes hirsuta Takahashi 1955 

Tetraleurodes hirsuta Takahashi 1955a: 419. Syntypes. Madagascar: on unidentified host, MNHNP. 



Tetraleurodes kunnathoorensis Regu & David 1993 

Tetraleurodes kunnathoorensis Regu & David 1993a: 53. Holotype puparium, India, Tamil Nadu, Kunnathoor, 

21.i.1991, K. Regu, on Streblus asper, K. Regu collection. 



Tetraleurodes leguminicola Bink-Moenen 1983 

Tetraleurodes leguminicola Bink-Moenen 1983: 173. Holotype female. Chad, Bebedji, 19.x.1973, on 

Dichrostachys glomerata, BMNH. 

Distribution: 3: Egypt; 4: Chad. 

Host: Fabaceae: Acacia nilotica, Acacia tortilis, Dichrostachys glomerata, Entada africana, 

Parkia clappertoniana. 

Natural enemies: Aphelinidae: Encarsia perconfusa, Encarsia acaudaleyrodis, Encarsia davidi. 

Tetraleurodes madagascariensis Takahashi 1951 

Tetraleurodes madagascariensis Takahashi 1951a: 384. Syntypes. Madagascar: on Chaetacne 

madagascariensis, MNHNP. 


Host: Ulmaceae: Chaetacme madagascariensis. 

452

Tetraleurodes malayensis Mound & Halsey 1978 

Tetraleurodes fici Takahashi 1955b: 232. Syntypes: Malaya: Kepong, Selangor, 9.i.1944, R. Takahashi on Ficus 

sp., BMNH. 

Tetraleurodes malayensis Mound and Halsey 1978: 199. replacement name for T. fici Takahashi 1955b:232, 

nec. Tetraleurodes fici Quaintance & Baker 1937. 



Tetraleurodes mameti Takahashi 1938 

Tetraleurodes mameti Takahashi 1938d: 262. Syntypes. Mauritius: on undetermined indigenous tree, TARI. 


Host: on undetermined indigenous tree. 

Tetraleurodes marshalli Bondar 1928 

Tetraleurodes marshalli Bondar 1928b: 35. Syntypes. Brazil: on Andropogon cicornis, MZU. 


Host: Poaceae: Andropogon cicornis. 

Tetraleurodes melanops (Cockerell) 1902 

Aleyrodes (Tetraleurodes) melanops Cockerell 1902a: 283. Syntypes. USA: California, on Quercus sp., USNM. 

Tetraleurodes melanops (Cockerell); Quaintance & Baker 1914: 108. 



Tetraleurodes mexicana Nakahara 1995 

Tetraleurodes mexicana Nakahara 1995: 117. Holotype pupal cases. Mexico: Sonora, Hermosillo, 19.v.1951, 

Gunderson, on Inga sp., USNM. 


Host: Fabaceae: Inga sp., Pithecellobium dulce, Tamarindus indica; Rutaceae: Ruta sp., Citrus sp. 

Tetraleurodes mirabilis Takahashi 1961 

Tetraleurodes mirabilis Takahashi 1961: 335. Syntypes. Madagascar: on Impatiens sp., MNHNP. 


Host: Balsaminaceae: Impatiens sp. 

453

Tetraleurodes monnioti (Cohic) 1968 

Aleuroplatus monnioti Cohic 1968b: 106. Syntypes (11). Congo: on Heisteria parvifolia. 

Tetraleurodes monnioti (Cohic); Bink-Moenen 1983: 174. 


Host: Olacaceae: Heisteria parvifolia. 

Tetraleurodes mori (Quaintance) 1899 

Aleurodes mori Quaintance 1899a: 1. Syntypes. USA: Florida, Tampa, on mulberry [Morus sp.], USNM. 

Tetraleurodes mori (Quaintance); Quaintance & Baker 1914: 108. 

Aleyrodes dorseyi Kotinsky 1907: 52. Syntypes. USA: California, on Rhamnus crocea, USNM; (Ray Gill, 

personal communication). 

Tetraleurodes dorseyi (Kotinsky); Quaintance & Baker 1914: 108. 

Aleyrodes errans Bemis 1904: 500. Syntypes. USA: California, on Umbellularia californica, USNM; (Ray Gill, 


Tetraleurodes errans (Bemis); Quaintance & Baker 1914: 502. 

Aleyrodes splendens Bemis 1904: 489. Syntypes. USA: California, on Rhamnus californica, USNM; (Ray Gill, 


Tetraleurodes splendens (Bemis); Quaintance & Baker 1914: 108. 

Distribution: 1: USA (AL, AR, AZ, CA, CO, CT, DC, DE, FL, GA, IL, LA, MA, MD, MO, NC, NJ, NM, NY, 

OH, PA, SC, TN, TX, UT, VA, WV); 2: Belize, Brazil; Colombia, Costa Rica, Cuba; El Salvador, 

Guatemala, Honduras; Jamaica; Mexico; Nicaragua, Panama, Puerto Rico, Trinidad & Tobago, 

Venezuela; 3: Iran; 6: Thailand, Vietnam; 8: Marianas Islands. 

Host: Aceraceae: Acer negundo, Acer pseudoplatanus, Acer rubrum, Acer sp.; Anacardiaceae: Rhus copallinum, 

Rhus sp., Toxicodendron radicans; Aquifoliaceae: Ilex opaca, Ilex sp., Ilex vomitoria; Asteraceae: 

Betulaceae: Betula sp., Carpinus caroliniana, Carpinus sp.,Corylus sp., Ostrya virginiana; 

Bignoniaceae: Catalpa sp., Pyrostegia ignea, Stizophyllum sp.; Celastraceae: Euonymus americana; 

Cornaceae: Cornus florida, Cornus foemina, Cornus sanguinea, Cornus sp.; Ebenaceae: Diospyros sp., 

Diospyros virginiana; Ericaceae: Arbutus menziesii, Kalmia latifolia, Vaccinium sp.; Fabaceae: 

Dichrostachys glomerata, Dichrostachys nutans; Fagaceae: Quercus sp.; Hamamelidaceae: 

Liquidambar styraciflua; Hippocastanaceae: Aesculus californica; Lauraceae: Persea borbonia, Persea 

carolinensis, Persea sp., Umbellularia californica; Magnoliaceae: Magnolia macrophylla, Magnolia sp.; 

Moraceae: Ficus sp., Maclura pomifera, Morus alba, Morus nigra, Morus rubra, Morus sp.; Musaceae: 

Musa sp.; Myricaceae: Myrica cerifera; Myrtaceae: Eugenia sp., Psidium guajava; Nyssaceae: Nyssa 

sp.; Oleaceae: Forsythia sp., Fraxinus americana, Fraxinus caroliniana, Fraxinus profunda, Fraxinus 

sp.; Papaveraceae: Argemone mexicana; Plantaginaceae: Platanus occidentalis; Rhamnaceae: Ceanothus 

sp., Rhamnus crocea; Rosaceae: Malus sp.; Rubiaceae: Cephalanthus occidentalis, Pinkneya bracteata; 

Rutaceae: Citrus maxima, Citrus sinensis, Citrus x nobilis, Zanthoxylum clava-herculis; Sapindaceae; 

454

455 

Sapotaceae: Bumelia celestrina, Bumelia lycioides, Bumelia sp.; Saxifragaceae: Itea virginica; Tiliaceae: 

Tilia americana, Tilia caroliniana, Tilia sp.; Ulmaceae: Celtis laevigata, Celtis sp., Ulmus americana, 

Ulmus sp.; Verbenaceae: Callicarpa americana. 

Natural enemies: Parasitoids - HYMENOPTERA/Aphelinidae: Encarsia luteola group, Encarsia parvella 

group, Eretmocerus sp.; Platygastridae: Amitus sculpturatus, Amitus sp. Predators - 

COLEOPTERA/Coccinellidae: Delphastus catalinae, Delphastus dejavu. 

US Port Interceptions: El Salvador on Hamelia sp.; Honduras on Musa sp.; Mexico on Arnica sp., 

Casimiroa edulis, Citrus sinensis, Citrus sp., Cnidoscolus chayamansa, Datura stramonium, Eucalyptus 

sp., Hedera helix, Herbs, Laurus nobilis, Malvaceae: Myrtus communis, Ocimum basilicum, Persea 

americana, Persea sp., Pithecellobium dulce, Pouteria sapota, Prunus domestica, Psidium guajava, 

Psidium sp., Pyrus sp., Ruta sp., Sorbus sp., Spondias sp.; Nigeria on unknown plant. 

Tetraleurodes moundi Cohic 1968 

Tetraleurodes moundi Cohic 1968b: 131. Syntypes. Angola: on Milettia sp, BMNH; Bink-Moenen 1983: 175. 


Host: Connaraceae: Agelaea dewevrei; Dilleniaceae: Tetracera alnifolia; Euphorbiaceae: Hymenocardia acida; 

Fabaceae: Millettia sp; Flacourtiaceae: Caloncoba welwitschii, Caloncoba dusenii; Loganiaceae: 

Strychnos spinosa, Strychnos variabilis, Strychnos pungens; Malpighiaceae: Acridocarpus dewevrei; 

Meliaceae: Entandrophragma utile, Trichilia prieureana; Myrtaceae: Psidium guajava; Ochnaceae: 

Ochna gilletiana; Passifloraceae: Barteria fistulosa; Rubiaceae: Gardenia jovis-tonantis; Ulmaceae: 

Celtis milbraedii. 

Tetraleurodes neemani Bink-Moenen 1992 

Tetraleurodes neemani Bink-Moenen 1992, in Bink-Moenen and Gerling 1992: 34. Holotype. Israel: Nahal 

Yagur, 30.iv.1977, Neeman, on Laurus nobilis, BMCol. 



US Port Interceptions: Greece and Lebanon on Laurus nobilis. 

Tetraleurodes oplismeni Takahashi 1934 

Tetraleurodes oplismeni Takahashi 1934b: 66. Syntypes, Taiwan, Shinten, Urai and Hori, on Oplismenus sp., 

TARI. 


Host: Poaceae: Oplismenus sp. 

Tetraleurodes pauliani Takahashi 1955 

Tetraleurodes pauliani Takahashi 1955a: 420. Syntypes. Madagascar: on ?Feijoa sellowiana, HBL.


Host: Myrtaceae: Feijoa sellowiana? 

Tetraleurodes perileuca (Cockerell) 1902 

Aleyrodes (Tetraleurodes) perileuca Cockerell 1902a: 283. Syntypes. USA: Texas, on Quercus sp., USNM. 

Tetraleurodes perileuca (Cockerell); Quaintance & Baker 1914: 283. 

Distribution: 1: USA (AZ, CA, FL, LA, TX, UT). 

Host: Fagaceae: Quercus agrifolia, Quercus densiflora, Quercus laurifolia, Quercus nigra, Quercus virginiana, 

Quercus wislizenii; Rhamnaceae: Rhamnus sp. 

Natural enemies: Platygastridae: Amitus granulosus, Amitus sp.; Aphelinidae: Encarsia sp. 

US Port Interceptions: El Salvador on Citrus limon; Mexico on Citrus sp., Persea americana, Psidium 

guajava. 

Tetraleurodes perseae Nakahara 1995 

Tetraleurodes perseae Nakahara 1995: 127. Holotype pupal cases. USA: California, San Diego County, San 

Diiego, 5.viii.1982, Ginsky & Blocker, on avocado [Persea americana], USNM. 

Distribution: 1: USA (CA); 2: El Salvador; Mexico. 

Host: Lauraceae: Laurus nobilis, Litsea sp, Persea americana, Umbellularia californica. 

US Port Interceptions: El Salvador and Mexico on Persea americana. 

Tetraleurodes pluto Dumbleton 1956 

Tetraleurodes pluto Dumbleton 1956b: 180. Holotype. Australia: on yellow flowering plant, ADSIR. 


Host: on yellow flowering plant. 

Tetraleurodes pringlei Quaintance & Baker 1937 

Tetraleurodes pringlei Quaintance & Baker 1937, in Baker 1937: 617. Holotype pupal case. Mexico: Oaxaca, 

G. C. G. Pringle, on Caulanthus hispites, USNM; Martin 2005b: 52. 


Host: Brassicaceae: Caulanthus hispites. 

Tetraleurodes pseudacaciae Nakahara 1995 

Tetraleurodes pseudacaciae Nakahara 1995: 119. Holotype puparium, Haiti, Port-Au-Prince, 20 X 1927, G. 

Wolcott, on leaves of Cassia mexicana, USNM. 

Distribution: 1: USA (CA); 2: Dominican Republic, Haiti, Jamaica, Puerto Rico 

456

Host: Fabaceae: Brya ebenus, Cassia mexicana; Zygophyllaceae: Guaiacum officinale. 

Tetraleurodes psidii David 1993 

Tetraleurodes psidii David 1993: 29. Holotype. Sri Lanka: on Psidium guajava. 



Tetraleurodes pusana Takahashi 1950 

Tetraleurodes pusana Takahashi 1950: 86. Syntypes. India: on undetermined grass, BMNH. 

Distribution: 6: India; Malaya; Malaysia. 


Tetraleurodes quadratus Sampson & Drews 1941 

Tetraleurodes quadratus Sampson & Drews 1941: 176. Syntypes. Mexico: on Byrsonima crassifolia, UCD; 

Martin 2005b. 

Distribution: 2: Belize, Costa Rica, Mexico, Nicaragua, Panama, Trinidad. 

Host: Malpighiaceae: Byrsonima crassifolia; Moraceae: Brosimum sp.; Myrtaceae: Pimenta sp. 

US Port Interceptions: Belize on Psidium guajava; Costa Rica on Alpinia purpurata, Pandanus sp., 

Schefflera actinophylla, Schefflera sp.; Guatemala on Citrus sp.; Honduras on dried plant 

Mexico on Citrus sp., Ficus benjamina, Persea americana; Nicaragua on Mangifera indica. 

Tetraleurodes quercicola Nakahara 1995 

Tetraleurodes quercicola Nakahara 1995: 131. Holotype pupal case. USA: Utah, Zion National Park, 

16.iv.1963, G. F. Knowlton, on evergreen oak [Quercus sp.], USNM. 

Distribution: 1: USA (AZ, CA, UT); 2: Mexico. 

Host: Quercus gambelii, Quercus turbinella Greene var. ajoensis. 

Tetraleurodes rugosus Corbett 1926 

Tetraleurodes rugosus Corbett 1926: 281. Syntypes. Sri Lanka: on grass, BMNH. 



457

Tetraleurodes russellae Cohic 1968 

Tetraleurodes russellae Cohic 1968b: 134. Syntypes. Congo: on Allophyllus africanus. 


Host: Annonaceae: Uvaria scabrida; Bignoniaceae: Markhamia sessilis; Combretaceae: Combretum sp.; 

Connaraceae: Manotes pruinosa; Dilleniaceae: Tetracera alnifolia; Euphorbiaceae: Hymenocardia 

acida; Fabaceae: Cassia javanica, Peltophorum pterocarpa; Loganiaceae: Strychnos variabilis; 

Lythraceae: Lagerstroemia indica; Menispermaceae: Triclisia gilletii; Ochnaceae: Ochna gilletiana; 

Passifloraceae: Barteria fistulosa; Rubiaceae: Colletoecema dewevrei, Gardenia jovis-tonantis, 

Pauridiantha hirtella; Sapindaceae: Allophylus africanus; Smilaceae: Smilax kraussiana; Sterculiaceae: 

Triplochiton scleroxylon; Ulmaceae: Celtis milbraedii; Verbenaceae: Clerodendrum sp. 

Tetraleurodes selachidentata Bink-Moenen 1983 

Tetraleurodes selachidentata Bink-Moenen 1983: 176. Holotype puparium, Chad, Bebedjia, 16.xii.1973, FAB 

& RMBM, on Terminalia laxiflora, BMNH. 

Distribution: 4: Chad, Ivory Coast. 

Host: Anacardiaceae: Lannea humilis; Annonaceae: Hexalobus monopetalus; Bignoniaceae: Stereospermum 

kunthianum; Cordia africana; Combretaceae: Anogeissus leiocarpus, Combretum micranthum, 

Combretum molle, Terminalia laxiflora, Terminalia macroptera; Euphorbiaceae: Antidesma venosum, 

Bridelia ferruginea, Hymenocardia acida; Fabaceae: Burkea africana, Detarium microcarpum, 

Piliostigma thonningii; Loganiaceae: Strychnos spinosa; Rhamnaceae: Ziziphus mauritiana; Rubiaceae: 

Crossopteryx febrifuga, Gardenia triacantha; Verbenaceae: Vitex simplicifolia. 

Note: Bink-Moenen (1983) expresses a doubt about the placement of this species in the genus Tetraleurodes 

and listed this species as “species incertae sedis” 

Tetraleurodes semibarbata Takahashi 1955 

Tetraleurodes semibarbata Takahashi 1955a: 422. Holotype. Madagascar: on unidentified host, MNHNP. 



Tetraleurodes simplicior Bink-Moenen 1983 

Tetraleurodes simplicior Bink-Moenen 1983: 178. Holotype puparium, Chad, Doba, 21.vi.1973, FAB & 

RMBM, on Syzygium guineense, BMNH.. 


Host: Myrtaceae: Syzygium guineese; Sapotaceae; Butyrospermum paradoxum. 

Tetraleurodes stellata (Maskell) 1895 

Aleurodes stellata Maskell 1895: 442. Syntypes. Jamaica: on Guaiacum officinale, ADSIR. 

458

Tetraleurodes stellata (Maskell); Quaintance & Baker 1914: 108. 


Host: Zygophyllaceae: Guaiacum officinale. 

Tetraleurodes stirlingiae Martin 1999 

Tetraleurodes stirlingiae Martin 1999: 105. Holotype puparium. Australia: intercepted in USA: California, on 

woody stems of Stirlingia latifolia, USNM. 


Host: Proteaceae: Stirlingia latifolia. 

US Port Interceptions: Australia on Conospermum sp., Leucadendron sp., Persoonia sp., Protea sp., Stirlingia 

sp., Strelitzia sp. 

Tetraleurodes submarginata Dumbleton 1961 

Tetraleurodes submarginata Dumbleton 1961a: 135. Holotype. New Caledonia: on ?Eugenia sp, ADSIR. 


Myrtaceae: Eugenia sp.? 

Tetraleurodes subrotunda Takahashi 1937 

Tetraleurodes subrotunda Takahashi 1937a: 43. Syntypes. Mauritius: on undetermined tree, TARI. 



Tetraleurodes sulcistriatus Martin 1999 

Tetraleurodes sulcistriatus Martin 1999: 106. Holotype puparium. Australia: on Ficus microcarpa, ANIC. 


Host: Moraceae: Ficus microcarpa. 

Tetraleurodes supraxialis Martin 1999 

Tetraleurodes supraxialis Martin 1999: 106. Holotype puparium. Australia: on ?Apocynaceae (shrub), ANIC. 


Host: Apocynaceae. 

Tetraleurodes thenmozhiae Jesudasan & David 1991 

Tetraleurodes thenmozhiae Jesudasan and David 1991: 333. Holotype pupal case. India: on Cinnamomum sp., 

459

IDAV. 



Tetraleurodes truncatus Sampson & Drews 1941 

Tetraleurodes truncatus Sampson & Drews 1941: 178. Syntypes. Mexico: on Psidium guajava, UCD. 

Distribution: 2: Honduras; Mexico. 


Tetraleurodes tuberculosa Nakahara 1995 

Tetraleurodes tuberculosa Nakahara 1995: 121. Holotype pupal cases. USA: Texas, Paso County, 27.vi.1956, J. 

W. Green, on stem of undetermined plant, USNM. 

Distribution: 1: USA (TX). 


Tetraleurodes tuberculata Bink-Moenen 1983 

Tetraleurodes tuberculata Bink-Moenen 1983: 179. Holotype puparium, Chad, Bebedjia, 26.xi.1973, FAB & 

RMBM, on Stereospermum kunthianum, BMNH. 


Host: Annonaceae: Hexalobus monopetalus; Bignoniaceae: Stereospernum kunthianum; Rubiaceae: 

Rothmannia whitfieldii 

Tetraleurodes ursorum (Cockerell) 1910 

Aleyrodes ursorum Cockerell 1910a: 171. Syntypes. USA: Colorado, Boulder, Flagstaff Mountains, 20.iii.1910, 

Cockerell, on Artostaphylos uva-ursi, USNM. 

Tetraleurodes ursorum (Cockerell); Quaintance & Baker 1914: 108. 

Distribution: 1: Canada; USA (AL, AZ, CO, CT, FL, GA, LA, MA, MS, MT, NJ, NM, SC, TX, UT, VA, WA, 

WI); 2: Costa Rica; Jamaica; Mexico; Puerto Rico. 

Host: Aceraceae: Acer barbatum, Acer rubrum, Acer sp.; Anacardiaceae: Schinus sp., Schinus terebinthifolius, 

Spondias mombin; Annonaceae: Annona glabra, Annona reticulata, Annona squamosa, Asimina sp., 

Asimina triloba; Aquifoliaceae: Ilex cassine, Ilex sp.; Arecaceae: Chamaedorea sp.; Asteraceae: Eclipta 

alba, Solidago fistulosa; Berberidaceae: Berberis sp., Mahonia fortunei, Mahonia haematocarpa, 

Mahonia trifoliolata; Caprifoliaceae: Viburnum odoratissimum, Viburnum sp.; Chrysobalanaceae: 

Chrysobalanus sp., Licania michauxii; Ebenaceae: Diospyros sp.; Ericaceae: Arbutus sp., Arctostaphylos 

columbiana, Arctostaphylos uva-ursi, Epigaea repens, Gaultheria shallon, Gaylussacia frondosa, 

Gaylussacia sp., Lyonia ferruginea, Lyonia sp., Rhododendron californicum, Rhododendron sp., 

460

461 

Vaccinium arboreum, Vaccinium ashei, Vaccinium corymbosum, Vaccinium macrocarpon; 

Euphorbiaceae: Cnidoscolus stimulosus, Manihot esculenta, Phyllanthus tenellus; Fabaceae: 

Aeschynomene sp., Cassia aspera, Cassia fasciculata, Dalbergia sp., Pithecellobium flexicaule; 

Fagaceae: Quercus myrtifolia, Quercus nigra, Quercus sp., Quercus virginiana; Guttiferae Calophyllum 

brasiliense, Calophyllum calaba; Hamamelidaceae: Liquidambar styraciflua; Iridaceae: Iris sp.; Labitae 

Conradina grandiflora; Lauraceae: Persea americana, Persea borbonia, Persea palustris, Persea sp.; 

Magnoliaceae: Magnolia grandiflora, Magnolia virginiana; Malvaceae: Abelmoschus esculentus, 

Hibiscus sp.; Moraceae: Ficus aurea, Ficus sp.; Myricaceae: Myrica cerifera, Myrica pensylvanica, 

Myrica rubra, Myrica sp.; Myrtaceae: Eugenia sp., Myrtaceae: Myrtus sp., Psidium cattleianum; 

Polygonaceae: Coccoloba sp.; Polypodiaceae: Platycerium sp.; Rhizophoraceae: Rhizophora mangle; 

Rosaceae: Malus sp,, Pyracantha sp,, Rosa palustris, Rosa sp, Rubus sp., Rubus trivialis; Rubiaceae: 

Cephalanthus occidentalis; Rutaceae: Amyris parviflora, Citrus aurantifolia, Citrus aurantium, Citrus 

limon, Citrus reticulata, Citrus sinensis, Citrus x nobilis, Citrus x paradisi, Zanthoxylum clava-herculis; 

Salicaceae: Salix sp.; Smilaceae: Smilax pumila; Theaceae: Gordonia lasianthus; Tiliaceae: Tilia 

caroliniana; Turneraceae: Piriqueta glabrescens; Ulmaceae: Celtis laevigata, Ulmus parvifolia, Ulmus 

sp.; Zygophyllaceae: Guaiacum officinale. 

Natural enemies: Aphelinidae: Encarsia basicincta, Encarsia citrella, Encarsia hamoni. 

US Port Interceptions: Mexico on Citrus limon, Citrus reticulata, Citrus sinensis, Psidium sp; 

Puerto Rico on Citrus sp.

Tetralicia Harrison 1917 

Tetralicia Harrison 1917a: 60. Type species. Tetralicia ericae Harrison 1917, by monotypy. 



debarroi x 

erianthi x 

ericae x 

graminicola x 

iberiaca x 

tuberculata x 

Total = 6 0 0 3 2 0 0 1 0 0 

Tetralicia debarroi Martin & Carver 1999 

Tetralicia debarroi Martin & Carver 1999, in Martin 1999: 108. Holotype puparium. Australia: on Grevillea 

sp., ANIC. 


Host: Proteaceae: Grevillea sp. 

Tetralicia erianthi Danzig 1969 

Tetralicia erianthi Danzig 1969: 878. Holotype. USSR: Turkmenia [Turkmenistan], on Erianthus purpurascens, 

ZIN. 

Distribution: 3: Iran, Turkmenistan, USSR. 

Host: Poaceae: Erianthus purpurascens, Erianthus ravennae; Rununculaceae: Rununculus arvensis. 

Natural enemies: Aphelinidae: Eretmocerus nikolskajae; Eulophidae: Neochrysocharis formosa. 

Tetralicia ericae Harrison 1917 

Tetralicia ericae Harrison 1917a: 61. Syntypes. England: on Erica tetralix, BMNH; Bink-Moenen 1976. 

Distribution: 3: Austria, Czech Republic, England, Italy, Netherlands, Sweden, Scotland, USSR, Wales. 

Host: Ericaceae: Erica tetralix, Erica carnea. 

462

Tetralicia iberiaca Bink-Moenen 1989 

Tetralicia iberiaca Bink-Moenen 1989: 178. Holotype. Portugal: Monchique, 29.iv.1978, F.A and R. M. Bink, 

on Erica arborea; Spain specimens, on Erica lusitanica, BMNH. 

Distribution: 3: Portugal. 

Host: Ericaceae: Erica lusitanica, Erica arborea. 

Tetralicia graminicola Bink-Moenen 1983 

Tetralicia graminicola Bink-Moenen 1983: 181. Holotype puparium, Chad, Bebedjia, 22.xi.1973, FAB & 

RMBM, on Hyparrhenia rufa, BMNH. 


Host: Poaceae: Hyparrhenia rufa. 

Tetralicia tuberculata Bink-Moenen 1983 

Tetralicia tuberculata Bink-Moenen 1983: 183. Holotype puparium, Chad, Bebedjia, 11.vii.1973, FAB & 

RMBM, on Vitex simplicifolia, BMNH. 


Host: Verbenaceae: Vitex simplicifolia. 

463

Trialeurodes Cockerell 1902 

Aleyrodes (Trialeurodes) Cockerell 1902a: 283. Type species. Aleurodes pergandei Quaintance 1900, by 


Trialeurodes Cockerell; full genus by Quaintance and Baker 1915: xi. 

Aleyrodes (Asterochiton) Maskell; misidentification; Kirkaldy 1907: 43; Quaintance & Baker 1914: 104. 

Aleurodes (Ogivaleurodes) Goux 1948: 31. Types species. Aleurodes lauri, by monotypy; synonymy according 

to Mound & Halsey 1978: 205. 

Gymnaleurodes Sampson & Drews 1940: 29. Types species. Gymnaleurodes bellissima, by monotypy; 

synonymy accoring to Sampson 1943: 209 

Comment - Russell (1948) divided the genus into 6 species groups. Rapisarda (1986) erected the subgenus 

Ericaleyrodes, and included two species, Trialeurodes ericae and T. sardiniae, based on the wedgeshaped 

body shape of the puparium, lack of anterior marginal setae and dorsal waxy rods, and the 

presence of submedian thoracic setae. 



abdita x 

abutiloneus x x x 

amealcensis x 

asplenii x 

bambusae x 

bellissima x 

bemisiae x 

bruneiensis x 

celti x 

chinensis x 

coccolobae x 

colcordae x 

corollis x 

cryptus x 

darwiniensis x 

dicksoniae x 

464


diminutis x 

drewsi x 

dubienus x 

elaphoglossi x 

ericae x 

eriodictyonis x 

euphorbiae x 

fernaldi x 

floridensis x x 

glacialis x 

heucherae x 

hutchingsi x 

intermedia x 

ipomoeae x 

lauri x x 

longispina x 

madroni x 

magnoliae x 

mameti x 

manihoti x 

meggitti x 

merlini x 

mirissimus x 

multipori x 

notata x x 

oblongifoliae x 

packardi x x x 

palaquifolia x 

paucipapilla x 

465


perakensis x 

pergandei x 

phlogis x 

rex x 

ricini x x x 

ruborum x 

sardiniae x 

shawundus x 

similis x x 

tabaci x 

tentaculatus x 

tephrosiae x 

thaiensis x 

unadutus x 

vaporariorum x x x x x x x x x 

varia x 

variabilis x x x x 

vitrinellus x x ?* 

vittata x 

Total = 66 species 31 21 7 4 4 8* 6 2 3 

466

Trialeurodes abdita Martin 2005 

Trialeurodes abdita Martin 2005b: 54. Holotype puparium, Belize, CFR, 29 V 2004, JH Martin, on Lasiacis sp., 

BMNH. 


Host: Poaceae: Lasiacis sp. 

Trialeurodes abutiloneus (Haldeman) 1850 

Aleurodes abutiloneus Haldeman 1850: 108. Syntypes. USA: Pennsylvania, on Sida (Abution) abutilon; 

USNM. 

Asterochiton abutiloneus (Haldeman); Quaintance & Baker 1914: 105. 

Trialeurodes abutiloneus (Haldeman); Quaintance & Baker 1915a: xi. 

Aleyrodes ambrosiae Cockerell 1910b: 370. Syntypes. USA: Colorado, on Ambrosia artemisiaefolia; 

synonymy according toRussell 1948: 71. 

Trialeurodes ambrosiae (Cockerell); Quaintance & Baker 1915a: xi. 

Aleurodes erigerontis Maskell 1895: 429. Syntypes. Mexico: on Erigeron sp.; synonymy according to Russell 

1948: 71. 

Aleyrodes (Trialeurodes) erigerontis Maskell; Cockerell 1902a: 283. 

Asterochiton erigerontis (Maskell); Quaintance & Baker 1914: 105. 

Trialeurodes erigerontis (Maskell); Quaintance & Baker 1915a: xi 

Aleurodes fitchi Quaintance 1900: 24. Syntypes. USA: Mississippi, on cotton [Gossypium sp.]; Quaintance 

1907: 92. 

Aleurodes rolfsii Quaintance 1899: 90. Syntypes. USA: Florida, on geranium [?Pelargonium sp.]; synonymy 

according toRussell 1948: 71. 

Asterochiton rolfsii (Quaintance); Quaintance & Baker 1914: 105. 

Trialeurodes rolfsii (Quaintance); Quaintance & Baker 1915a: xi. 

Distribution: 1: USA (AL, AR, AZ, CA, CO, DC, DE, FL, GA, HI, IL, IN, KS, LA, MD, MI, MO, MS, NJ, 

NM, NY, OK, PA, SC, TN, TX, UT, VA, WA); 2: Brazil; Costa Rica; Cuba; El Salvador; Guatemala; 

Honduras; Jamaica; Mexico; Puerto Rico; 3: Iran; 9: Hawaii. 

Host: Aceraceae: Acer rubrum; Aizoaceae: Mollugo verticillata; Amaranthaceae: Amaranthus sp.; Asteraceae:, 

Ambrosia artemisiifolia L, Ambrosia hispida, Ambrosia sp., Bidens pilosa, Bidens sp., Eclipta alba, 

Emilia fosbergii, Emilia javanica, Emilia sonchifolia, Erigeron sp., Gamolepis chrysanthemoides, 

Gerbera jamesonii, Heliantheae sp., Helianthus sp., Heterotheca grandiflora, Heterotheca subaxillaris, 

Lactuca graminifolia Michaux var. graminifolia, Lactuca sp., Mikania scandens, Pluchea carolinensis, 

Pluchea purpurascens, Pluchea rosea, Pluchea sp., Senecio confusus, Solidago sp., Sonchus oleraceus 

L, Sonchus sp., Taraxacum officinale, Taraxacum sp., Targetes sp., Tithonia rotundifolia, Youngia 

japonica; Brassicaceae: Lepidium virginicum; Campanulaceae: Lobelia cardinalis; Caryophyllaceae: 

Drymaria cordata; Convolvulaceae: Ipomoea sp.; Cucurbitaceae: Cucumis melo, Euphorbiaceae: 

Chamaesyce hirta; Chamaesyce hyssopifolia, Chamaesyce sp., Croton glandulosus, Euphorbia 

467

468 

heterophylla, Euphorbia pulcherrima, Euphorbia sp., Jatropha gossypiifolia, Phyllanthus urinaria; 

Fabaceae: Aeschynomene americana, Aeschynomene sp., Arachis hypogaea, Cercis canadensis L, 

Desmodium geniculata, Glycine max, Indigofera hirsuta Harv, Indigofera sp., Medicago sativa L, 

Sesbania exaltata, Sesbania punicea, Sesbania sp., Sesbania tripetti, Sesbania vesicaria; Fagaceae: 

Quercus sp.; Haloragaceae: Myriophyllum aquaticum; Labiatae Ajuga sp., Monarda didyma; Salvia sp..; 

Lauraceae: Nectandra coriacea, Persea americana; Malvaceae: Abutilon theophrasti, Gossypium 

hirsutum L, Gossypium sp., Hibiscus cannabinus, Hibiscus elatus, Hibiscus esculentus (L.) Moench, 

Hibiscus furcellatus Desr, Hibiscus grandiflorus, Hibiscus moscheutos, Hibiscus moscheutos L, 

Hibiscus mutabilis, Hibiscus mutabilis L, Hibiscus rosa-sinensis, Hibiscus sp., Hibiscus syriacus, Malva 

parviflora, Malva sp., Sida abutilon, Sida acuta, Sida cordifolia, Sida rhombifolia, Sida sp., Urena 

lobata; Moraceae: Fatoua villosa, Fatuoa villosa; Myrtaceae: Melaleuca quinquenervia, Psidium 

guajava; Onagraceae: Ludwigia octovalvis, Ludwigia peruviana; Oxalidaceae: Oxalis corniculata; 

Papaveraceae: Argemone mexicana; Plantaginaceae: Platanus sp.; Poaceae: Sorghum bicolor; 

Polygonaceae: Polygonella fimbriata, Polygonella sp., Polygonum punctatum; Primulaceae: Samolus 

parviflorus; Punicaceae: Punica granatum; Rutaceae: Citrus sp., Poncirus trifoliata, Ruta chalepensis; 

Scrophulariaceae: Bacopa caroliniana; Solanaceae: Capsicum annuum, Lycopersicon esculentum, 

Physalis sp., Solanum americanum, Solanum nigrum, Solanum sp., Solanum villosum; Sterculiaceae: 

Dombeya wallichii; Umbelliferae: Eryngium aromaticum; Urticaceae: Fleurya aestuans, Laportea 

aestuans, Urtica sp.; Verbenaceae: Glandularia maritima, Lantana sp., Verbena scabra. 

Natural enemies: Fungus - Aschersonia aleyrodis; Parasitoids - HYMENOPTERA/ Aphelinidae: Encarsia 

californica, Encarsia formosa, Encarsia hayati, Encarsia hispida, Encarsia luteola, Encarsia meritoria, 

Encarsia n. sp, Encarsia nigricephala, Encarsia parvella group, Encarsia pergandiella, Encarsia 

quaintancei, Encarsia sophia, Encarsia sp, Encarsia strenua group, Encarsia tabacivora, Eretmocerus 

californicus, Eretmocerus haldemani, Eretmocerus sp, Eretmocerus staufferi; Platygastridae: Amitus 

aleurodinis, Amitus bennetti, Amitus fuscipennis, Amitus sp.; Signiphoridae: Signiphora aleyrodis. 

Predators - COLEOPTERA/Coccinellidae: Coccinella novemnotata, Coleomegilla maculata, Coleomegilla 

maculata lengi, Delphastus pusillus, Hippodamia convergens, Olla v-nigrum; DIPTERA/Cecidomyidae: 

Genus? sp.; HEMIPTERA/Anthocoridae: Orius insidiosus, Orius tristicolor; Lygeidae: Geocoris 

punctipes; Miridae: Spanogonicus albofasciatus; Nabidae: Nabis ferus, Nabis sp.; NEUROPTERA 

Chrysopidae: Chrysopa oculata, Chrysoperla plorabunda; THYSANOPTERA/Thripidae: Sericothrips 

trifasciatus 

US Port Interceptions: Mexico on Dendranthema sp., Origanum majorana, Persea americana, Ruta 

graveolens. 

Trialeurodes amealcensis Carapia 2003 

Trialeurodes amealcensis Carapia 2003, in Carapia-Ruiz et al. 2003: 97. Holotype puparium. Mexico: 

Queretaro, 5 km south of Amealco, 21.vii.2001, V. Carapia on Quercus laeta. 


Host: Fagaceae: Quercus laeta.

Trialeurodes asplenii (Maskell) 1890 

Aleurodes asplenii Maskell 1890b: 173. Lectotype. New Zealand: on Asplenium lucidum, ADSIR. 

Asterochiton asplenii (Maskell); Quaintance & Baker 1914: 105. 

Trialeurodes asplenii (Maskell); Quaintance & Baker 1915a: xi. 


Host: Aspleniaceae: Asplenium lucidum. 

Trialeurodes bambusae Takahashi 1943 

Trialeurodes bambusae Takahashi 1943: 29. Syntypes. Thailand: on bamboo [?Bambusa sp.], TARI. 



Trialeurodes bellissima (Sampson & Drews) 1940 

Gymnaleurodes bellissima Sampson & Drews 1940: 29. Holotype. USA: California, on Quercus sp., USNM. 

Trialeurodes bellissima (Sampson & Drews); Sampson 1943: 209. 

Distribution: 1: USA (CA, TX). 

Host: Fagaceae: Quercus sp., Quercus agrifolia. 

Trialeurodes bemisiae Russell 1948 

Aleyrodes tentaculatus Bemis 1904: 494, in part. Holotype. USA: California, Stanford University, 30.i.1902, F. 

E. Bemis, on Quercus agrifolia. 

Trialeurodes bemisae Russell 1948: 494. 


Host: Ericaceae: Arbutus menziesii; Fagaceae: Quercus agrifolia; Rhamnaceae: Ceanothus thyrsiflorus; 

Rosaceae: Photinia arbutifolia. 

Trialeurodes bruneiensis Martin 2001 

Trialeurodes bruneiensis Martin 2001, in Martin & Camus 2001: 4. Holotype puparium. Borneo, Brunei, 

eastern Belait District, near Bukit Bedawan, 30.iii.1989, J. Martin, on fern. 

Distribution: 7: Borneo. 

Host: Thelypteridaceae: fern. 

Trialeurodes celti Takahashi 1943 

Trialeurodes celti Takahashi 1943: 28. Syntypes. Thailand: on Celtis sp., TARI. 

469



Trialeurodes chinensis Takahashi 1955 

Trialeurodes chinensis Takahashi 1955b: 231. Holotype. China: on Solanum sp. 


Host: Solanaceae: Solanum sp. 

Trialeurodes coccolobae Russell 1948 

Trialeurodes coccolobae Russell 1948: 23. Holotype. Mexico: on Coccoloba floribunda, USNM. 


Host: Polygonaceae: Coccoloba floribunda. 

Trialeurodes colcordae Russell 1948 

Trialeurodes colcordae Russell 1948: 75. Holotype. USA: California, on unknown host, USNM. 



Trialeurodes corollis (Penny) 1922 

Asterochiton corollis Penny 1922: 26. Syntypes. USA: California, San Diego County, Pine Hills, x.1917, D. 

Penny, on Arctostaphylos manzanita, CAS. 

Trialeurodes corollis (Penny); Russell 1948: 36. 


Host: Ericaceae: Arctostaphylos manzanita, Arctostaphylos sp. 

Trialeurodes cryptus Martin 2005 

Trialeurodes cryptus Martin 2005b: 56. Holotype puparium, Belize, CFR, 22 XI 1994, JH Martin, on 

Bombacaceae?, BMNH. 


Host: Bombacaceae? 

Trialeurodes darwiniensis Martin 1999 

Trialeurodes darwiniensis Martin 1999: 109. Holotype puparium. Australia: on Flueggea virosa, ANIC. 


470

Host: Euphorbiaceae: Flueggea virosa. 

Trialeurodes dicksoniae Martin 1999 

Trialeurodes dicksoniae Martin 1999: 110. Holotype puparium. Australia: on Dicksonia antarctica, ANIC. 


Host: Dicksoniaceae: Dicksonia antarctica. 

Trialeurodes diminutis (Penny) 1922 

Asterochiton diminutis Penny 1922: 28. Syntypes. USA: California, on Chamaebatia foliolosa, CAS. 


Host: Rosaceae: Chamaebatia foliolosa. 

Trialeurodes drewsi Sampson 1945 

Trialeurodes drewsi Sampson 1945: 62. Holotype. USA: California, on Quercus sp., UCD. 

Aleyrodes gracialis Bemis 1904; misidentification in part; Bemis 1904: 518. 


Host: Fagaceae: Lithocarpus densiflorus, Quercus agrifolia, Quercus dumosa var. elegantula, Quercus sp, 

Quercus ilex, Quercus tomentella. 

Trialeurodes dubienus (Bondar) 1923 

Asterochiton dubienus Bondar 1923a: 179. Holotype. Brazil: Bahia, on Psidium guajava, USNM. 

Trialeurodes dubienus (Bondar); Bondar 1928b: 32. 


Host: Psidium guajava. 

Trialeurodes elaphoglossi Takahashi 1960 

Trialeurodes elaphoglossi Takahashi 1960a: 139. Syntypes. Reunion Island: on Elaphoglossum sp., MNHNP. 


Host: Polypodiaceae: Elaphoglossum sp. 

Trialeurodes ericae Bink-Moenen 1976 

Trialeurodes ericae Bink-Moenen 1976: 17. Holotype. The Netherlands: on Erica tetralix, NHM (Rapisarda 

1986: 497); Bink-Moenen 1989:176. 

Trialeurodes (Ericaleyrodes) ericae Bink-Moenen; Rapisarda 1986: 490. 

471

Distribution: 3: Crete, Corsica, France, Majorca, Netherlands, Spain. 

Host: Ericaceae: Erica arborea, E. australis, E. lusitanica, E. multiflora, E. tetralix, E. umbellata. 

Trialeurodes eriodictyonis Russell 1948 

Trialeurodes eriodictyonis Russell 1948: 76. Holotype. USA: California, San Diego County, Del Mar, 

26.vii.1912, P. H. Timberlake, on Eriodictyon crassifolium, USNM. 

Distribution: 1: USA (CA, FL). 

Host: Hydrophyllaceae: Eriodictyon crassifolium. 

Trialeurodes euphorbiae Russell 1948 

Trialeurodes euphorbiae Russell 1948: 77. Holotype. USA: California, on Euphorbia albomarginata, USNM. 


Host: Euphorbiaceae: Euphorbia albomarginata. 

Trialeurodes fernaldi (Morrill) 1903 

Aleyrodes fernaldi Morrill 1903b: 83. Syntypes. USA: Massachusetts, on strawberry [Fragaria sp.], USNM. 

Trialeurodes fernaldi (Morrill); Russell 1948: 63. 

Distribution: 1: USA (CT, FL, MA). 

Host: Diapensiaceae: Galax urceolata; Rosaceae: Fragaria sp., Spiraea vanhouttei, Spiraea sp. 

Rubiaceae: Cephalanthus occidentalis. 

Natural enemies: Aphelinidae: Encarsia luteola; Platygastridae: Amitus aleurodinis. 

Trialeurodes floridensis (Quaintance) 1900 

Aleurodes floridensis Quaintance 1900: 26. Syntypes. USA: Florida, Lakeland and Punta Gorda, viii.1898, A. 

L. Quaintance, on guava, Psidium guajava, USNM. 

Asterochiton floridensis (Quaintance); Quaintance & Baker 1914: 105. 

Trialeurodes floridensis (Quaintance); Quaintance & Baker 1915a: xi. 

Distribution: 1: USA (DC, FL); 2: Bahamas; Belize, Brazil; Cuba; Guatemala; Haiti, Honduras; Mexico; 

Panama; Puerto Rico; Venezuela; Virgin Islands. 

Host: Anacardiaceae: Mangifera indica; Annonaceae: Annona diversifolia, Annona squamosa; Bignoniaceae: 

Crescentia sp.; Bombacaceae: Bombax malabaricum; Caricaceae: Carica papaya; Chrysobalanaceae: 

Chrysobalanus sp.; Chrysobalanaceae: Licania michauxii; Convolvulaceae: Ipomoea sp.; Dilleniaceae: 

Tetracera scandens, Tetracera volubilis; Ebenaceae: Diospyros sp.; Euphorbiaceae: Chamaesyce 

hyssopifolia; Fabaceae: Acacia nigrescens, Acacia pallens, Albizia moluccana, Bauhinia divaricata, 

Calliandra sp., Cassia sieberana, Lysiloma latisiliquum, Piscidia communis; Lauraceae: Persea 

americana, Persea borbonia, Persea sp.; Lythraceae: Lagerstroemia speciosa ; Malpighiaceae: 

472

473 

Malpighia glabra; Malvaceae: Sida cordifolia; Meliaceae: Melia azedarach; Myrtaceae: Calyptranthes 

pallens, Eucalyptus botryoides, Eucalyptus sp., Psidium guajava, Tristania conferta; Polygonaceae: 

Coccoloba uvifera; Rhamnaceae: Colubrina reclinata; Rosaceae: Licania sp.; Rubiaceae: Casasia 

calophylla, Cephalanthus sp., Randia calophylla, Guettarda sp.; Rutaceae: Citrus sp., Clausena 

lansium, Zanthoxylum clava-herculis; Sapindaceae: Harpullia arborea; Sapotaceae: Achras sapota. 

Natural enemies: Parasitoids - HYMENOPTERA/Aphelinidae: Encarsia hispida, Encarsia meritoria, 

Encarsia nigricephala, Encarsia tabacivora, Encarsia telemachusi; Predators: COLEOPTERA/ 

Coccinellidae: Delphastus pallidus Delphastus sp. 

US Port Interceptions: Ecuador on Lisianthus sp.; Guyana on Persea americana; Italy on Olea sp.; 

Mexico on Abelmoschus esculentus, Citrus aurantiifolia, Citrus sinensis, Citrus sp., Persea 

americana, Persea sp., Psidium guajava, Punica granatum, Ruta graveolens; Puerto Rico on 

Persea americana. 

Trialeurodes glacialis (Bemis) 1904 

Aleyrodes glacialis Bemis 1904: 518. Cotypes. USA: California, Stanford University, on Ceanothus sp., 

USNM. 

Asterochiton glacialis (Bemis); Quaintance & Baker 1914: 105. 

Trialeurodes glacialis (Bemis); Quaintance & Baker 1915a: xi. 

Distribution: 1: USA (CA, UT, WA). 

Host: Caprifoliaceae: Symphoricarpos racemosus, Lonicera interrupta; Fagaceae: Quercus densiflora; Labiatae: 

Salvia mellifera; Ranunculaceae: Clematis ligusticifolia; Rhamnaceae: Ceanothus californicus, 

Ceanothus sp., Rhamnus californica; Rosaceae: Opulaster capitatus, Rubus vitifolius. 

Trialeurodes heucherae Russell 1948 

Trialeurodes heucherae Russell 1948: 68. Holotype. USA: District of Colombia, Washington, Rock Creek Park, 

10.iv.1941, on Heuchera americana, USNM. 

Distribution: 1: USA (DC, PA). 

Host: Saxifragaceae: Heuchera americana. 

Trialeurodes hutchingsi (Bemis) 1904 

Aleyrodes hutchingsi Bemis 1904: 532. Syntypes. USA: California, Yosemite Valie, vii.1902, F. E. Bemis, on 

Arctostaphylos sp., USNM. 

Asterochiton hutchingsi (Bemis); Quaintance & Baker 1914: 105. 

Trialeurodes hutchingsi (Bemis); Quaintance & Baker 1915a: xi 

Distribution: 1: USA (CA, OR). 

Host: Ericaceae: Arctostaphylos sp., Arctostaphylos patula.

Trialeurodes intermedia Russell 1948 

Trialeurodes intermedia Russell 1948: 20. Holotype. USA: California, Mount Low, 8.ix.1912, P. H. 

Timberlake, on Rhamnus crocea, USNM. 

Trialeurodes vitrinellus (Cockerell); misidentification in part; Quaintance & Baker 1916: 470. 

Distribution: 1: USA (AZ, CA, TX). 

Host: Anacardiaceae: Rhus ovata; Fagaceae: Quercus agrifolia; Rhamnaceae: Rhamnus crocea. 

Rutaceae: Amyris texana. 

Trialeurodes ipomoeae Carapia 2003 

Trialeurodes ipomoeae Carapia 2003, in Carapia-Ruiz et al. 2003: 94. Holotype puparium. Mexico: Morelos, 

Chatcatzingo, 13.ii.1993, V. Carapia, on Ipomoea arborescens. 


Host: Convolvulaceae: Ipomoea arborescens. 

Trialeurodes lauri (Signoret) 1882 

Aleurodes lauri Signoret 1882: CLVIII. Syntypes. Greece: Athens (Grenadius), on Laurus nobilis. 

Aleuroparadoxus lauri (Signoret); Silvestri 1934: 399. 

Trialeurodes lauri (Signoret); Russell 1947: 6. 

Aleyrodes (Ogivaleurodes) lauri (Signoret); Goux 1948: 31. 

Ogivaleurodes lauri (Signoret); Goux 1951: 12. 

Trialeurodes lauri (Signoret); Zahradnick 1963b: 232. 

Trialeurodes klemmi Takahashi 1940c: 148. Syntypes. Yugoslavia: Rab, on Laurus nobilis; synonymy 

according to Russell 1947: 6. 

Distribution: 3: Belgium, Cyprus, France; Greece; Iran; Israel; Italy, Lebanon, Luxembourg, Switzerland, 

Turkey, USSR; Yugoslavia; 7: Australia. 

Host: Ericaceae: Arbutus sp.; Lauraceae: Laurus nobilis. 

US Port Interceptions: on Laurus nobilis from Albania, Argentina, Australia, Belgium, Brazil, 

China, Croatia, Cyprus, France, Gambia, Germany, Greece, India, Iran, Italy, Lebanon, 

Malta, Mexico, Netherlands, Russia, Saudi Arabia, Singapore, South Africa, Switzerland, Turkey, 

and Yugoslavia; on Olea sp. from Italy; on Cinnamomum sp. from Spain. 

Trialeurodes longispina Takahashi 1943 

Trialeurodes longispina Takahashi 1943: 30. Syntypes. Thailand: on undetermined host. 


474

Host: undetermined. 

Trialeurodes madroni (Bemis) 1904 

Aleyrodes madroni Bemis 1904: 507. Syntypes. USA: California, on Arbutus menziesii, USNM. 

Asterochiton madroni (Bemis); Quaintance & Baker 1914: 105. 

Trialeurodes madroni (Bemis); Quaintance & Baker 1915a: xi. 


Host: Ericaceae: Arbutus menziesii; Fagaceae: Quercus sp. 

Trialeurodes magnoliae Russell 1948 

Trialeurodes magnoliae Russell 1948: 69. Holotype. USA: Florida, Lake City, 7.xi.1897, A. L. Quaintance, on 

Magnolia virginiana, USNM. 

Distribution: 1: USA (FL, TX). 

Host: Magnoliaceae: Magnolia virginiana. 

Trialeurodes mameti Takahashi 1951 

Trialeurodes mameti Takahashi 1951a: 370. Syntypes (2). Madagascar: on "Mankolody", MNHNP. 


Host: "Mankolody". 

Trialeurodes manihoti (Bondar) 1923 

Asterochiton manihoti Bondar 1923a: 177. Syntypes. Brazil: Bahia, G. Bondar, on Manihot sp., MZU. 

Trialeurodes manihoti (Bondar); Bondar 1923a: 32. 


Host: Euphorbiaceae: Manihot sp. 

Trialeurodes meggitti Singh 1933 

Trialeurodes meggitti Singh 1933: 345. Syntypes. Burma: on Ficus sp, ZSI. 



Trialeurodes merlini (Bemis) 1904 

Aleyrodes merlini Bemis 1904: 512. Syntypes. USA: California, on Arbutus menziesii, USNM. 

Asterochiton merlini (Bemis); Quaintance & Baker 1914: 105. 

475

Trialeurodes merlini (Bemis); Quaintance & Baker 1915a: xi. 

Distribution: 1: Canada; USA (CA, OR, WA). 

Host: Ericaceae: Arctostaphylos glauca, Arctostaphylos manzanita, Arbutus menziesii, Arctostaphylos patula, 

Arctostaphylos pringle Parry ssp. pringlei, Arctostaphylos sp. 

Trialeurodes mirissimus Sampson & Drews 1941 

Trialeurodes mirissimus Sampson & Drews 1941: 179. Syntypes. Mexico: on sapote [?Casimiroa edulis], UCD. 

Distribution: 2: Jamaica; Mexico; St. Kitts; Trinidad. 

Host: Flacourtiaceae:, Casearia hirsuta?; Rutaceae: Casimiroa edulis, Citrus sp.; Verbenaceae: Petrea arborea. 

Trialeurodes multipori Russell 1948 

Trialeurodes multipori Russell 1948: 25. Holotype. USA: Arizona, on Quercus oblongifolia, USNM. 

Distribution: 1: USA (AZ). 

Host: Fagaceae: Quercus oblongifolia. 

Trialeurodes notata Russell 1948 

Trialeurodes notata Russell 1948: 55. Holotype. USA: District of Colombia, Takoma Park, 10.ix.1941, L. M. 

Russell, on Helianthus sp, USNM. 

Distribution: 1: USA (DC, IL, KS, MD, MO, PA). 

Host: Asclepiadaceae: Asclepias syriaca; Asteraceae: Ambrosia trifida, Eupatorium purpureum, Helianthus 

tuberosus, Helianthus sp., Rudbeckia hirta, Solidago altissima, Vernonia missurica; Convolvulaceae: 

Ipomoea pandurata; Labiatae: Mentha arvensis, Mentha canadensis; Solanaceae: Physalis heterophylla, 

Solanum carolinense L. var. carolinense. 

Trialeurodes oblongifoliae Russell 1948 

Trialeurodes oblongifoliae Russell 1948: 26. Holotype. Mexico: on Quercus oblongifolia, USNM. 

Distribution: 1: USA (AZ); 2: Mexico. 

Host: Fagaceae: Quercus oblongifolia, Quercus sp. 

Trialeurodes packardi (Morrill) 1903 

Aleyrodes packardi Morrill 1903a: 25. Syntypes. USA: Massachusetts, on Fragaria sp., USNM. 

Asterochiton packardi (Morrill); Quaintance & Baker 1914: 105. 

Trialeurodes packardi (Morrill); Quaintance & Baker 1915a: xi. 

Aleyrodes coryli Britton 1907: 337. Syntypes. USA: Connecticut, on Corylus americana, USNM; synonymy 

according to Russell 1948: 60 

476

477 

Aleyrodes morrilli Britton 1907: 340. Syntypes. USA: Connecticut, on Impatiens fulca, USNM; synonymy 


Aleyrodes waldeni Britton 1907: 339. Syntypes. USA: Connecticut, on Juglans nigra, USNM; 

synonymy according to Russell 1948: 60. 

Distribution: 1: Canada; USA (AZ, CA, CT, DC, FL, GA, HI, IN, IO, KS, MA, MD, ME, MI, MN, MO, NC, 

NJ, NM, NV, NY, OH, OR, PA, RI, TN, TX, UT, VA, WA, WI); 3: Hungary, Iran; 5: Korea. 

Host: Anacardiaceae: Rhus aromatica; Asteraceae: Ambrosia trifida, Aster laterifolius, Aster novae-angliae, 

Eupatorium rugosum, Helianthus sp., Solidago sp.; Balsaminaceae: Impatiens biflora, Impatiens 

capensis, Impatiens pallida; Betulaceae: Betula sp., Carpinus sp., Corylus americana, Corylus avellana, 

Corylus rostrata, Ostrya sp.; Bignoniaceae: Bignonia radicans, Catalpa speciosa, Caprifoliaceae: 

Lonicera sp., Caprifoliaceae: Symphoricarpos sp., Viburnum acerifolium, Viburnum dilatatum; 

Ebenaceae: Diospyros virginiana; Ericaceae: Arctostaphylos uva-ursi, Rhododendron flavum, 

Rhododendron ponticum, Vaccinium myrtilloides; Fabaceae: Amphicarpaea bracteata, Catalpa sp., 

Cercis canadensis, Desmodium paniculatum; Guttiferae Hypericum sp.; Hydrangeaceae: Philadelphus 

sp.; Juglandaceae: Juglans cinerea, Juglans nigra, Juglans regia; Labiatae: Collinsonia canadensis, 

Mentha arvensis, Mentha canadensis, Mentha citrata, Prunella vulgaris; Malvaceae: Hibiscus 

moscheutos; Moraceae: Maclura pomifera; Oleaceae: Chionanthus virginicus, Fraxinus velutina; 

Punicaceae: Punica granatum; Rhamnaceae: Ceanothus americanus, Rosaceae: Crataegus mollis, 

Fragaria ananassa, Fragaria sp., Fragaria vesca, Fragaria virginiana, Prunus malus, Prunus persica, 

Rosa wichuraiana, Rubus allegheniensis, Rubus nigrobaccus, Rubus odoratus, Rubus procerus, Spiraea 

bumalda, Spiraea vanhouttei; Rutaceae: Zanthoxylum clava-herculis; Saururaceae: Saururus sp.; 

Saxifragaceae: Deutzia sp.; Tilaceae: Tilia americana; Ulmaceae: Celtis sp., Ulmus scabra var pendula; 

Umbelliferae: Cryptotaenia canadensis; Verbenaceae: Callicarpa americana, Verbena urticifolia; 

Violaceae: Viola sp. 

Natural enemies: Aphelinidae: Encarsia luteola, Encarsia pergandiella, Encarsia protransvena, Encarsia 

quaintancei, Eretmocerus corni. 

Trialeurodes palaquifolia Corbett 1935 

Trialeurodes palaquifolia Corbett 1935b: 815. Holotype. Malaya: on Palaquium gutta. 


Host: Sapotaceae: Palaquium gutta. 

Trialeurodes paucipapilla Martin 2005 

Trialeurodes paucipapilla Martin 2005b: 57. Holotype puparium, Belize, CFR, 4.vi.2003, J. H. Martin, on 

Coccoloba ?belizensis, BMNH. 


Host: Polygonaceae: Coccoloba belizensis?

Trialeurodes perakensis Corbett 1935 

Trialeurodes perakensis Corbett 1935b: 814. Holotype. Malaya: on Palaquium gutta. 


Host: Sapotaceae: Palaquium gutta. 

Trialeurodes pergandei (Quaintance) 1900 

Aleurodes pergandei Quaintance 1900: 31. Syntypes. USA: District of Colombia, Washington, on Bignonia 

radicans, USNM. 

Aleyrodes (Trialeurodes) pergandei Quaintance; Cockerell 1902a: 283. 

Asterochiton pergandei (Quaintance); Quaintance & Baker 1914: 105. 

Trialeurodes pergandei (Quaintance); Quaintance & Baker 1915a: xi. 

Distribution: 1: USA (DC, FL, GA, IL, MD, NC, SC, TN, VA). 

Host: Bignoniaceae: Bignonia radicans, Tecoma sp.; Caprifoliaceae: Viburnum dentatum, 

Viburnum opulus; Liliaceae: Smilax sp.; Oeaceae: Chionanthus virginicus, Fraxinus sp. 

Rosaceae: Crataegus oxyacantha, Crataegus stipulosa, Prunus nana, Prunus persica, Pyracantha 

angustifolia, Rosa wichuraiana, Rubus sp., Rubus villosus; Rutaceae: Zanthoxylum americanum; 

Saxifragaceae: Hydrangea sp. 

Trialeurodes phlogis Russell 1993 

Trialeurodes phlogis Russell 1993: 583. Holotype pupa, USA, Virginia, Shenandoah County, SE of Mt. 

Jackson, 12.iv.1992, A. G. Wheeler, on Phlox subulata. 

Distribution: 1: USA (MD, VA). 

Host: Polemoniaceae: Phlox subulata. 

Trialeurodes rex Martin 2001 

Trialeurodes rex Martin 2001, in Martin & Camus 2001: 7. Sulawesi, Utara, Dumoga, Bone NP, forested ridge, 

18.iv.1985, J. Martin, on fern. 


Host: Pteridophyta. 

Trialeurodes ricini (Misra) 1924 

Aleyrodes ricini Misra 1924: 131. Syntypes. India: on Ricinus communis, USNM. 

Trialeurodes ricini (Misra); Singh 1931: 46. 

Trialeurodes rara Singh 1931: 47. Syntypes. India, on Breynia sp; synonymy according to Bink-Moenen 1983: 

185. 

Trialeurodes desmodii Corbett 1935c: 243. Syntypes. Sierra Leone, on Desmodium lasiocarpum, BMNH; 

478

synonymy [with T. rara] according to Mound and Halsey 1978: 217. 

Trialeurodes lubia El Khidir & Khalifa 1962: 47. Holotype. Sudan: on Dolichos lablab; synonymy with T. rara 

according to Mound 1965c: 157. 

Distribution: 3: Iran, Israel; Saudi Arabia, Turkey; 4: Cameroon; Central African Republic, Chad, Ivory Coast, 

Gabon, Madagascar, Nigeria; Sierra Leon, Sudan, Uganda, Zaire, Zimbabwe; 6: Cambodia, India; 

Pakistan; Malaya; Malaysia, Sri Lanka, Thailand. 

Host: Annonaceae: Annona glabra; Convolvulaceae: Ipomoea batatas; Ericaceae: Arbutus sp. 

Euphorbiaceae: Breynia rhamnoides, Breynia sp. Euphorbia sp., Phyllanthus acidus, Phyllanthus sp., 

Ricinus communis; Fabaceae: Dalbergia sissoo, Desmodium lasiocarpus, Dolichos lablab; Malvaceae: 

Gossypium hirsutum; Menispermaceae: Cissampelos owariensis; Myrtaceae: Psidium guajava; 

Pedaliaceae: Sesamum sp.; Piperaceae: Piper umbellatum; Rosaceae: Rosa canina, Rosa sp.; Rubiaceae: 

Gardenia jovis-tonantis, Gardenia ternifolia; Rutaceae: Murraya koenigii; Sapotaceae: Achras sapota; 

Tiliaceae: Corchorus sp. 

Natural enemies: Aphelinidae: Encarsia hispida, Encarsia lahorensis, Encarsia sophia, Eretmocerus 

adustiscutum, Eretmocerus mundus, Eretmocerus trialeurodis. 

US Port Interceptions: Greece on Laurus nobilis; India on Laurus nobilis, Murraya koenigii, Murraya 

sp.; Iran on Murraya koenigii; Mexico on Persea americana; Nigeria on Solanaceae; Pakistan 

on Psidium guajava; Sri Lanka on Murraya koenigii; Turkey on Ficus sp.; Unknown source on 

Capsicum sp. 

Trialeurodes ruborum (Cockerell) 1897 

Aleurodes ruborum Cockerell 1897b: 96. Syntypes. USA: Florida, on Rhus sp., USNM. 

Aleyrodes (Trialeurodes) ruborum Cockerell; Cockerell 1902a: 283. 

Asterochiton ruborum (Cockerell); Quaintance & Baker 1914: 105. 

Trialeurodes ruborum (Cockerell); Quaintance & Baker 1915a: xi. 

Distribution: 1: USA (CA, CO, DC, FL, LA, VA). 

Host: Anacardiaceae: Rhus sp.; Rosaceae: Fragaria sp., Rubus cuneifolius, Rubus glaucus, Rubus trivialis. 

Trialeurodes sardiniae Rapisarda 1986 

Trialeurodes (Ericaleyrodes) sardiniae Rapisarda 1986: 487. Holotype pupal case, Sardegna (Sardinia), Burcei, 

Monte Settefratelli, 19.v.1985, S. Barbagall and C. Rapisarda, on Erica arborea, Rapisarda collection. 

Distribution: 3: Sardinia, Italy. 

Host: Ericaceae: Erica arborea. 

Trialeurodes shawundus Baker & Moles 1923 

Trialeurodes shawundus Baker & Moles 1923: 644. Syntypes. Chile: on unknown host, USNM. 



479

Trialeurodes similis Russell 1948 

Trialeurodes similis Russell 1948: 31. Holotype. USA: Florida, Sanibel Island, summer 1909, E. A. Back, on 

?Galactia sp., USNM. 

Distribution: 1: USA (FL); 2: Belize, Mexico. 

Host: Fabaceae: Galactia sp.; Sterculiaceae: Waltheria sp. 

US Port Interceptions: Mexico on Persea americana. 

Trialeurodes tabaci Bondar 1928 

Trialeurodes tabaci Bondar 1928b: 32. Syntypes. Brazil: on "fumo" [Nicotiana sp.], MZU. 


Host: Solanaceae: Nicotiana sp. 

Trialeurodes tentaculatus (Bemis) 1904 

Aleyrodes tentaculatus Bemis 1904: 494. Syntypes. USA: California, on Quercus agricolia, USNM. 

Asterochiton tentaculatus (Bemis); in part; Quaintance & Baker 1914: 105. 

Trialeurodes tentaculatus (Bemis); Quaintance & Baker 1915a: xi. 


Host: Anacardiaceae: Rhus diversabola; Caprifoliaceae: Lonicera involucrata; Fagaceae: Quercus agrifolia, 

Quercus densiflora; Ranunculaceae: Clematis ligusticifolia; Rosaceae: Opulaster capitatus. 

Trialeurodes tephrosiae Russell 1948 

Trialeurodes tephrosiae Russell 1948: 32. Holotype. Mexico: on Tephrosia arcuata, USNM. 

Distribution: 2: Costa Rica; El Salvador; Mexico; Panama. 

Host: Fabaceae: Tephrosia arcuata, Tephrosia heydeana, Tephrosia langlassei. 

Trialeurodes thaiensis Takahashi 1943 

Trialeurodes thaiensis Takahashi 1943: 31. Syntypes. Thailand: on legume [Fabaceae]. 



Trialeurodes unadutus Baker & Moles 1923 

Trialeurodes unadutus Baker & Moles 1923: 643. Syntypes. Chile: on Drimys winteri, USNM. 


480

Host: Winteraceae: Drimys winteri. 

Trialeurodes vaporariorum (Westwood) 1856 

Aleurodes vaporariorum Westwood 1856: 852. Syntypes. England: on Gonolobus sp., Tecoma velutina, other 

plants, OUMNH. 

Asterochiton vaporariorum (Westwood); Quaintance & Baker 1914: 105. 

Trialeurodes vaporariorum (Westwood); Quaintance & Baker 1915a: xi. 

Aleyrodes glacialis Bemis (misidentification in part); Bemis 1904: 518. 

Asterochiton lecanioides Maskell 1879 (in part); Maskell 1879: 215. Syntypes. New Zealand: on Pittosporum 

eugenioides and Polypodium billardieri, ADSIR; synonymy according to Quaintance & Baker 1914: 

105. 

Trialeurodes mossopi Corbett 1935a: 9. Syntypes. Rhodesia: on Phaseolus vulgaris, BMNH; synonymy 


Aleurodes nicotiana Maskell 1895: 436. Syntypes. Mexico: on Nicotiana tabacum, USNM; synonymy 

according to Quaintance & Baker 1914: 105. 

Asterochiton papillifer Maskell 1890b: 173. Syntypes. New Zealand: on Pittosporum eugenioides, ADSIR; 

synonymy according to Quaintance & Baker 1914: 105. 

Trialeurodes sesbania Corbett 1936: 19. Syntypes. Australia: on Sesbania tripeti, BMNH; synonymy according 

to Russell 1948: 44. 

Aleyrodes sonchi Kotinsky 1907: 97. Syntypes. USA: Hawaii, Honolulu, on Sonchus oleraceus, USNM; 

synonymy according to Baker & Moles 1923: 645. 

Asterochiton sonchi (Kotinsky); Quaintance & Baker 1914: 105. 

Trialeurodes sonchi (Kotinsky); Quaintance & Baker 1914: 105. 

Distribution: Worldwide, 1: Canada, USA (AK, AL, AZ, CA, CO, CT, DC, DE, FL, GA, HI, ID, IN, IL, IO, 

KS, MA, MD, MI, MO, MS, NC, NE, NJ, NM, NY, OH, OR, PA, RI, SC, SD, TX, UT, VA, WA, WI, 

WV); 2: Argentina, Bermuda, Brazil, Chile, Colombia, Costa Rica, Cuba, Dominican Republic, 

Ecuador, El Salvador, Guadeloupe, Guatemala, Honduras, Jamaica, Mexico, Peru, Puerto Rico, 

Venezuela; 3: Azores, Austria, Belgium, Bulgaria, Canary Islands, Denmark, France, Germany, Greece, 

Hungary, Iran, Ireland, Italy, Israel, Jordan, Netherlands, Norway, Poland, Portugal, Spain, Turkey, 

United Kingdom; 4: Ethiopia, Kenya, South Africa, Zimbabwe; 5: Hong Kong, Japan, Korea; 6: 

Bangladesh, India, Philippines, Reunion, Sri Lanka; 7: Australia, Indonesia, New Guinea; 8: New 

Zealand; 9: Hawaii. 

Host: polyphagous (see Mound & Halsey 1978); in addition - Asclepiadaceae: Gonolobus sp.; Asteraceae: 

Ageratum sp., Helianthus annuum, Lactuca serriola, Sonchus oleraceus, Xanthium occidentale; 

Cucurbitaceae: Citrullus colocynthis, Cucumis sativus, Sechium edule; Euphorbiaceae: Euphorbia 

geniculata, Manihot esculenta; Fabaceae: Phaseolus vulgaris; Malvaceae: Hibiscus trionum; 

Solanaceae: Lycopersicon esculentum, Nicotiana glauca, Salpichroa rhomboidea, Solanum melongena. 


Fungus - Acremonium sp., Aphanocladium album, Aschersonia aleyrodis, Aschersonia confluens, Aschersonia 

481

482 

flava, Aschersonia goldiana, Aschersonia papillata, Aschersonia placenta, Aschersonia sp. orange 

Cuba, Aschersonia sp. red Cuba, Aschersonia sp. white Cuba, Aschersonia tamurai, Beauveria 

bassiana, Fusarium verticilliodes, Paecilomyces fumosorseus, Trichothecium roseum, Verticillum 

fusisporum, Verticillum lecanii. 

Parasitoids - HYMENOPTERA/Aphelinidae: Encarsia adusta, Encarsia azimi, Encarsia basicincta, Encarsia 

bimaculata, Encarsia californica, Encarsia costaricensis, Encarsia desantisi, Encarsia formosa, 

Encarsia guadeloupae, Encarsia hispida, Encarsia inaron, Encarsia japonica, Encarsia lutea, Encarsia 

luteola, Encarsia lycopersici, Encarsia melanostoma, Encarsia mineoi, Encarsia nigricephala, Encarsia 

noahi, Encarsia oakeyensis, Encarsia pergandiella, Encarsia porteri, Encarsia protransvena, Encarsia 

quaintancei, Encarsia sophia, Encarsia sp., Encarsia synaptocera, Encarsia tabacivora, Encarsia 

tricolor, Encarsia ustulata, Eretmocerus californicus, Eretmocerus eremicus, Eretmocerus haldemani, 

Eretmocerus mundus, Eretmocerus sp., Eretmocerus warrae; Eulophidae: Euderomphale sp.; 

Platygastridae: Amitus arcturus; Amitus fuscipennis; Pteromalidae: Moranila comperei. 

Predators - ACARI/Anystidae Walzia australica, COLEOPTERA/Coccinellidae: Coccinella transversalis, 

Coelophora inequalis, Diomus notescens, Harmonia conformis, Harmonia octomaculata, Micraspis 

frenata, Serangium nigripes, Serangium sp., Stethorus vagans, Melyridae: Dicranolaius bellulus; 

HEMIPTERA/Lygeidae Geocoris lubra; Miridae: Campylomma liebknechti, Deraeocoris signatus; 

NEUROPTERA/Chrysopidae: Mallada signata. 

US Port Interceptions: Australia on Scaevola sp.; Belgium on Rhododendron sp.; Brazil on Lisianthus sp., 

Solidago sp.; Canada on Artemisia dracunculus, Gerbera sp., Laurus nobilis; Colombia on 

Alstroemeria sp., Aster sp., Dendranthema sp., Mentha sp., Ocimum sp., Origanum sp., Origanum 

vulgare, Rosa sp., Solidago sp., Thymus sp., Thymus vulgaris; Costa Rica on Ananas comosus, 

Duranta sp.; Dominican Republic on Aster sp.; Ecuador on Aster sp., Hypericum sp., Lisianthus 

sp.; El Salvador on Fernaldia pandurata; Ethiopia on plant; France on Melissa sp., Rosa sp.; 

Guatemala on Limonium sp.; Hawaii on Heckeria umbellata, Hibiscus rosa-sinensis, 

Jasminum sp., Moringa oleifera; Japan on Daphne sp., Perilla frutescens; Kenya on Hypericum 

sp., Veronica sp.; Mexico on Abelmoschus esculentus, Alstroemeria sp., Aster ericoides, Aster 

sp., Asteraceae, Bouvardia sp., Callistephus sp., Chenopodium ambrosioides, Chrysanthemum 

sp., Crocosmia sp., Dahlia sp., Dendranthema sp., Dianthus sp., Fuchsia sp., Helichrysum sp., 

Hypericum sp., Limonium sinuatum, Malus sylvestris, Malvastrum sp., Mentha sp., Oenothera 

sp., Origanum sp., Origanum vulgare, Pelargonium sp., Persea americana, Persea sp., Phaseolus 

vulgaris, Physalis ixocarpa, Piper sp., Plant, Rosa sp., Ruta graveolens, Ruta sp., Rutaceae, Solanum 

sp., Solidago sp., Solidaster sp., Thymus sp.; Netherlands on Ageratum sp., Aster sp., Astilbe sp., 

Bouvardia sp., Ceanothus sp., Hypericum sp., Iris sp., Lycopersicon esculentum, Lycopersicon sp., 

Paeonia sp., Phlox sp., Plant, Rosa sp., Veronica sp.; New Zealand on Gentiana sp.; Peru on Ruta 

graveolens; Poland on a plant; United Kingdom on Lavandula sp., Physocarpus sp.; Zimbabwe 

on Aster sp., Hypericum sp. 

Trialeurodes varia Quaintance & Baker 1937 

Trialeurodes varia Quaintance & Baker 1937, in Baker 1937: 620. Syntypes. Mexico: on Rosaceae probably 

Rosa sp, USNM. 

Distribution: 2: Mexico.

Host: Rosaceae: probably Rosa sp.; Saxifragaceae: Philadelphus sp.; Solanaceae: Datura sp. 

Trialeurodes variabilis (Quaintance) 1900 

Aleurodes variabilis Quaintance 1900: 39. Syntypes. USA: Florida, Miami, on Carica papaya, USNM. 

Aleyrodes (Trialeurodes) variabilis Quaintance; Cockerell 1902a: 283. 

Asterochiton variabilis (Quaintance); Quaintance & Baker 1914: 105. 

Trialeurodes variabilis (Quaintance); Quaintance & Baker 1915a: xi. 

Trialeurodes caricae Corbett 1935c: 245. Syntypes. Trinidad: on Carica papaya; synonymy according to 

Russell 1948: 29. 

Aleurodicus (Metaleurodicus) variabilis (Quaintance); Dozier 1936: 145 (in error) 

Distribution: 1: USA (AZ, FL, HI); 2: Belize, Brazil, Colombia; Costa Rica, Cuba; Dominica, Dominican 

Republic, Ecuador, El Salvador, French Giana, Guadeloupe, Guatemala; Haiti, Honduras; Jamaica; 

Mexico; Nicaragua, Panama, Puerto Rico, St. Croix, Trinidad & Tobago, Venezuela, Virgin Islands; 3: 

Iran; 5: Korea; 9: Hawaii. 

Host: Aceraceae: Acer mexicanum; Caricaceae: Carica papaya, Carica sp.; Cyperaceae: Cyperus rotundus; 

Euphorbiaceae: Manihot esculenta, Manihot glaziovii; Polygonaceae: Coccoloba floribunda; Rubiaceae: 

Gardenia sp.; Rutaceae: Citrus aurantifolia, Citrus nobilis, Citrus reticulata, Citrus sinensis, Citrus x 

paradisi. 

Natural enemies: Aphelinidae: Encarsia bellottii, Encarsia hispida, Encarsia pergandiella, Encarsia 

protransvena; Platygastridae: Amitus macgowni; COLEOPTERA/ Coccinellidae: Leis conformis. 

US Port Interceptions: Guatemala on Carica papaya; Mexico on unknown plant. 

Trialeurodes vitrinellus (Cockerell) 1903 

Aleyrodes vitrinellus Cockerell 1903a:241. Syntypes. Mexico: on orange [Citrus sp.] leaves, USNM. 

Asterochiton vitrinellus (Cockerell); Quaintance & Baker 1914: 105. 

Trialeurodes vitrinellus (Cockerell); Quaintance & Baker 1915a: xi. 

Trialeurodes vitrinellus (Cockerell); misidentification of specimens on oak in part by Quaintance & Baker 

1916:470, should be Trialeurodes intermedia Russell according to Mound & Halsey 1978: 225. 

Distribution: 1: USA (UT); 2: Belize, Costa Rica, Mexico; 6: Vietnam? 

Host: Myrtaceae: Psidium sp.; Rutaceae: Citrus sp. 

US Port Interceptions: Greece on Citrus sp.; Mexico on Casimiroa edulis; Citrus limon, Citrus sinensis, Citrus 

sp., Mentha sp., Persea americana, Persea sp., Pithecellobium sp., Psidium guajava, Psidium sp., 

Punica granatum. 

Trialeurodes vittata (Quaintance) 1900 

Aleurodes vittata Quaintance 1900: 42. Syntypes. USA: California, on xerophytic shrub, USNM. 

483

Aleyrodes (Trialeurodes) vittata Quaintance; Cockerell 1902a: 283. 

Asterochiton vittata (Quaintance); Quaintance & Baker 1914: 105. 

Trialeurodes vittata (Quaintance); Quaintance & Baker 1915a: x. 

Aleyrodes glacialis Bemis (misidentification in part); Bemis 1904: 518. 

Aleyrodes wellmanae Bemis 1904: 525. Syntypes. USA: California, on Rhamnus californica, USNM; 

synonymy according to Russell 1948: 65. 

Asterochiton wellmanae (Bemis); Quaintance & Baker 1914: 105. 


Host: Rhamnaceae: Rhamnus californica, Rhamnus californica Eschsch. ssp. californica. 

Rosaceae: Crataegus coccinea; Vitaceae: Vitis vinifera. 

Natural enemies: Aphelinidae: Encarsia californica. 

484

Trialeurolonga Martin 2005 

Trialeurolonga Martin 2005b: 60. Type species Trialeurolonga trifida, by original description and monotypy. 



trifida x 

Total = 1 species 0 1 0 0 0 0 0 0 0 

Trialeurolonga trifida Martin 2005 

Trialeurolonga trifida Martin 2005b: 61. Holotype puparium, Belize, CFR, 1.vi.2004, J. H. Martin, on 

Chiococca alba?, BMNH. 


Host: Rubiaceae: Chiococca alba? 

485

Trichoaleyrodes Takahashi & Mamet 1952 

Trichoaleyrodes Takahashi & Mamet 1952b: 122. Type species. Trichaleyrodes carinata Takahashi & Mamet 




carinata x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Trichoaleyrodes carinata Takahashi & Mamet 1952 

Trichoaleyrodes carinata Takahashi & Mamet 1952b: 123. Syntypes. Madagascar: on "hazon-domoina", 

MNHNP. 


Host: on "hazon-domoina". 

486

Tuberaleyrodes Takahashi 1932 

Tuberaleyrodes Takahashi 1932: 29. Type species. Tuberaleyrodes machili Takahashi 1932, by monotypy. 



bobuae x 

machili x 

neolitseae x 

rambutana x 

spiniferosa x 

Total = 5 species 0 0 0 0 3 2 0 0 0 

Tuberaleyrodes bobuae Takahashi 1934 

Tuberaleyrodes bobuae Takahashi 1934b: 58. Syntypes. Taiwan: on Bobua glauca, TARI. 


Host: Symplocaceae: Symplocos [=Bobua] glauca. 

Tuberaleyrodes machili Takahashi 1932 

Tuberaleyrodes machili Takahashi 1932: 29. Syntypes. Taiwan: on Machilus sp., TARI. 


Host: Lauraceae: Persea japonica, Machilus sp. 

Natural enemies: Aphelinidae: Encarsia lahorensis. 

Tuberaleyrodes neolitseae Young 1944 

Tuberaleyrodes neolitseae Young 1944: 130. Syntypes. China: on Neolitsea aurata var glauca. 


Host: Lauraceae: Neolitsea aurata var. glauca. 

Tuberaleyrodes rambutana Takahashi 1955 

Tuberaleyrodes rambutana Takahashi 1955b: 229. Holotype. Malaya: on Nephelium lappaceum ["Rambutan"], 

TARI. 


487


Tuberaleyrodes spiniferosa (Corbett) 1933 

Acanthaleyrodes spiniferosa (Corbett) 1933: 128. Syntypes. Malaya: Kuala Lumpur, 15.ii.1928, on Diospyros 

sp., BMNH 

Tuberaleyrodes spiniferosa (Corbett); Martin & Mound 2007. 


Host: Ebenaceae: Diospyros sp. 

488

Vasantharajiella David 2000 

Vasantharajiella David 2000, in P.M. David 2000: 127. Type species - Vasantharajiella kalakadensis David 

2000, by monotypy and original designation. 



kalakadensis x 

Total = 1 species 0 0 0 0 0 1 0 0 0 

Vasantharajiella kalakadensis David 2000 

Vasantharajiella kalakadensis David 2000, in P. M. David 2000: 127. Holotype puparium, India, Tamil Nadu, 

Kalakad forest, 5.i.1993, P.M.M. David, on undetermined climber, IARI. 


Host: undetermined climber. 

489

Vasdavidius Russell 2000 

Vasdavidius Russell 2000: 379. Type species. Aleurocybotus indicus David & Subramaniam 1976, by original 

designation. 

References: Key to Taiwanese species [as Aleurocybotus] Ko et al. 1998; Russell 2000. 



cobarensis x 

concursus x 

indicus x x x x 

miscanthus x 

setiferus x x x 

Total = 5 species 0 0 1 1 4 2 2 0 0 

Vasdavidius cobarensis (Martin) 1999 

Aleurocybotus cobarensis Martin 1999: 34. Holotype puparium, Australia, NSW, Cobar, on pasture grass, 1979, 

E. Schicha, ANIC. 

Vasdavidius cobarensis (Martin); Russell 2000: 381. 


Host: Poaceae: on pasture grass. 

Vasdavidius concursus (Ko) 1998 

Aleurocybotus concurus Ko et al. 1998. Holotype. Taiwan, ex. Miscanthus floridulus. 

Vasdavidius concurus (Ko); Russell 2000: 381. 


Host: Poaceae: Miscanthus floridulus. 

Vasdavidius indicus (David & Subramaniam) 1976 

Aleurocybotus indicus David & Subramaniam 1976: 157. Holotype. India: on Chloris barbata, IDAV; Bink- 

Moenen 1983: 40. 

Vasdavidius indicus (David & Subramaniam); Russell 2000: 381. 

Distribution: 3: Iran; 4: Chad; Niger; 5: China; 6: India. 

Host: Poaceae: Chloris barbata, Dactyloctenium aegyptium, Oryza sativa, Oxynanthera abyssinica. 

490

Natural enemies: Aphelinidae: Encarsia japonica, Encarsia sophia. 

Vasdavidius miscanthus (Ko) 1998 

Aleurocybotus miscanthus Ko 1998, in Ko et al. 1998. Holotype. Taiwan: on Miscanthus sp. 

Vasdavidius miscanthus (Ko); Russell 2000: 381. 


Host: Poaceae: Miscanthus sp. 

Vasdavidius setiferus (Quaintance & Baker) 1917 

Aleurocybotus setiferus Quaintance & Baker 1917: 357. Syntypes. Java: on Imperata sp., USNM; Corbett 1926: 

274; David 1993a: 15; Martin 1985: 317. 

Vasdavidius setiferus (Quaintance & Baker); Russell 2000: 381 

Distribution: 5: Hong Kong; Philippines; Taiwan; 6: Malaya; Malaysia; Sri Lanka; Thailand; 7: Australia; Java; 

Papua New Guinea; Sulawesi. 

Host: Pandanaceae: Pandanus sp.; Poaceae: Imperata cylindrica, Imperata arundinacea; Rutaceae: Citrus spp. 

Natural enemies: Aphelinidae: Encarsia persequens. 

491

Venezaleurodes Russell 1967 

Venezaleurodes Russell 1967: 235. Type species. Venezaleurodes pisoniae Russell 1967, by monotypy. 



pisoniae x 

Total = 1 species 0 1 0 0 0 0 0 0 0 

Venezaleurodes pisoniae Russell 1967 

Venezaleurodes pisoniae Russell 1967: 237. Holotype. Venezuela: on Pisonia macranthocarpa, USNM. 

Distribution: 2: Venezuela. 

Host: Nyctaginaceae: Pisonia macranthocarpa. 

492

Viennotaleyrodes Cohic 1968 

Viennotaleyrodes Cohic 1968a: 54. Type species. Viennotaleyrodes bourgini Cohic 1968, by original 

designation. 

Moundiella David 1974a: 43. Type species. Trialeurodes megapapillae Singh 1932, by monotypy and original 

designation; David et al. 1994. 

Brazzaleyrodes (in part). 

Comment: Found almost exclusively on legumes (Fabaceae). 

Reference: Bink-Moenen 1983: 189 - Description of genus, key to species; David et al. 1994 - key to species. 



bergerardi x 

bicolorata x 

bosciae x 

curvisetosus x 

dichrostachi x 

fallax x 

incomptus x 

lacunae x 

megapapillae x x 

nilagiriensis x 

playsepali x 

Total = 11 species 0 0 0 6 0 2 4 0 0 

Viennotaleyrodes bergerardi (Cohic) 1966 

Brazzaleyrodes bergerardi Cohic 1966b: 32. Syntypes (11). Congo: on Platysepalum vanderystii. 

Viennotaleyrodes bergerardi (Cohic); Cohic 1968a: 57. 


Host: Fabaceae: Platysepalum vanderystii. 

Viennotaleyrodes bicolorata Martin 1999 

Viennotaleyrodes bicolorata Martin 1999: 113. Holotype puparium. Australia: on Acacia harpophylla, ANIC. 

493


Host: Fabaceae: Acacia harpophylla. 

Viennotaleyrodes bosciae Bink-Moenen 1983 

Viennotaleyrodes bosciae Bink-Moenen 1983: 192. Holotype puparium, Cameroun, 22 km south of Waza, 

23.i.1966, AJ Leeuwenberg, on Boscia senegalensis, BMNH. 

Distribution: 4: Cameroon. 

Host: Capparidaceae: Boscia senegalensis. 

Viennotaleyrodes curvisetosus Martin 1999 

Viennotaleyrodes curvisetosus Martin 1999: 114. Holotype puparium. Australia: on Acacia pendula, ANIC. 


Host: Fabaceae: Acacia pendula. 

Viennotaleyrodes dichrostachi Bink-Moenen 1983 

Viennotaleyrodes dichrostachi Bink-Moenen 1983: 192. Holotype puparium, Chad, Bebedjia, 19.xi.1973, R. 

Bink-Moenen, on Dichrostachys glomerata, BMNH. 


Host: Fabaceae: Dichrostachys glomerata. 

Viennotaleyrodes fallax Bink-Moenen 1983 

Viennotaleyrodes fallax Bink-Moenen 1983: 193. Holotype puparium, Chad, Boro, 30.xii.1972, R. Bink- 

Moenen, on Burkea africana, BMNH. 


Host: Fabaceae: Fabaceae: Burkea africana, Cassia javanica, Cassia sieberiana, Daniella olivera, Detarium 

microcarpum, Andira inermis. 

Viennotaleyrodes incomptus Martin 1999 

Viennotaleyrodes incomptus Martin 1999: 114. Holotype puparium. Australia: on Acacia harpophylla, ANIC. 


Host: Fabaceae: Acacia harpophylla. 

Viennotaleyrodes lacunae Martin 1999 

Viennotaleyrodes lacunae Martin 1999: 115. Holotype puparium. Australia: on Acacia rubida, ANIC. 


Host: Fabaceae: Acacia rubida. 

494

Viennotaleyrodes megapapillae (Singh) 1932 

Trialeurodes megapapillae Singh 1932: 86. Syntypes. Burma: Syriam, x.1929, K.S. Lamba, on Butea sp., ZSI. 

Moundiella megapapillae (Singh); David 1974a: 43. 

Moundiella indicus Jesudasan and David 1991: 315. Holotype pupal case. India: on Cassia timorensis, IDAV. 

Viennotaleyrodes indicus (Jesudasan & David); David et al. 1994. 

Viennotaleyrodes lafonti Cohic 1968a: 57. Syntypes (18). Congo: on Newtonia glandulifera; Ko et al. 1998: 78. 

Distribution: 4: Congo; 6: Burma, India. 

Host: Fabaceae: Butea sp., Cassia fistula, Cassia timorensis, Newtonia glandulifera, Tamarindus indica. 

Viennotaleyrodes nilagiriensis David & Krishnan 1994 

Viennotaleyrodes nilagiriensis David & Krishnan 1994, in David et al. 1994: 35. Holotype pupal case, India, 

Burliar, 27.x.1991, B. Krishman, on unidentified tree. 



Viennotaleyrodes platysepali (Cohic) 1966 

Brazzaleyrodes platysepali Cohic 1966b: 30. Syntypes. Congo: Brazzaville, on Newtonia glandulifera. 

Viennotaleyrodes platysepali (Cohic); Cohic 1968a: 57. 

Viennotaleyrodes bourgini Cohic 1968a: 54. Holotype. Chad: on Dalbergia sissoo; synonymy according to 



Host: Fabaceae: Dalbergia kisantuensis, Dalbergia sissoo, Platysepalum vanderystii. 

495

Xenaleyrodes Takahashi 1936 

Xenaleyrodes Takahashi 1936f: 113. Types species. Xenaleyrodes artocarpi Takahashi 1936, by monotypy. 



artocarpi x x 

broughae x 

eucalypti x 

fauceregius x 

irianicus x 

timonii x 

sp. x 

Total = 6+ species 0 0 0 0 0 0 6+ 0 0 

Xenaleyrodes artocarpi Takahashi 1936 

Xenaleyrodes artocarpi Takahashi 1936f: 113. Syntypes. Palau Islands: on Artocarpus communis, TARI; 

Martin 1985: 334. 

Distribution: 7: Papua New Guinea; 8: Palau Islands; Caroline Islands. 

Host: Lecythidaceae: Barringtonia racemosa; Moraceae: Artocarpus communis; Verbenaceae: Premna sp. 

Xenaleyrodes broughae Martin 1985 

Xenaleyrodes broughae Martin 1985: 334. Holotype pupal case. Papua New Guinea: on Citrus sp, BMNH. 



Xenaleyrodes eucalypti (Dumbleton) 1956 

Neomaskellia eucalypti Dumbleton 1956b: 175. Holotype. Australia: on Eucalyptus sp., ADSIR. 

Xenaleyrodes eucalypti (Dumbleton); Martin 1985: 334. 


Host: Myrtaceae: Eucalyptus ficifolia, Eucalyptus sp. 

496

Xenaleyrodes fauceregius Martin 1999 

Xenaleyrodes fauceregius Martin 1999: 118. Holotype puparium. Australia: on Acacia ?mcdonnelliensis, ANIC. 


Host: Fabaceae: Acacia lysiphloia, Acacia mcdonnelliensis? 

Xenaleyrodes irianicus Martin 1985 

Xenaleyrodes irianicus Martin 1985: 335. Holotype pupal case. Papua New Guinea: on undetermined tree, 

BMNH. 

Distribution: 7: Irian Jaya; Papua New Guinea. 


Xenaleyrodes timonii Martin 1985 

Xenaleyrodes timonii Martin 1985: 336. Holotype pupal case. Papua New Guinea: on Timonius sp., BMNH. 


Host: Rubiaceae: Timonius sp. 


Xenaleyrodes sp. 

Xenaleyrodes sp. Pedata & Polaszek 2003: 371. 7: Australia, on Myrtaceae: Lophostemon sp. 

Natural enemies: Aphelinidae: Encarsia longifasciata. 

497

Xenobemisia Takahashi 1951 

Xenobemisia Takahashi 1951a: 372. Type species. Xenobemisia coleae Takahashi 1951, by monotypy. 



coleae x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Xenobemisia coleae Takahashi 1951 

Xenobemisia coleae Takahashi 1951a: 372. Syntypes. Madagascar: on Colea telfairii, MNHNP. 


Host: Bignoniaceae: Colea telfairii. 

498

Yleyrodes Bink-Moenen 1983 

Yleyrodes Bink-Moenen 1983: 197. Type species- Yleyrodes isoberliniae Bink-Moenen 1983, by monotypy and 


Comment: According to Bink-Moenen (1983), this genus is similar to Africaleurodes in the separated 

submargin, distribution of the submarginal setae and structure of the vasiform orifice but differs in that 

the moulting sutures reach the lateral margin, the submarginal furrow is entire, the presence of the A1 

setae, the almost straight legs and the connected tracheal fold. 



isoberliniae x 

Total = 1 species 0 0 0 1 0 0 0 0 0 

Yleyrodes isoberliniae Bink-Moenen 1983 

Yleyrodes isoberliniae Bink-Moenen 1983: 197. Holotype puparium, Boro, 6.xii.1973, R. Bink-Moenen & 

FAB, on Isoberlinia doka, BMNH. 



499

Zaphanera Corbett 1926 

Zaphanera Corbett 1926: 282. Type species. Zaphanera cyanotis Corbett 1926, by monotypy. 

Comment: Usually found on legumes (Fabaceae). 



capparis x 

cyanotis x 

indicus x 

publicus x 

Total = 4 species 0 0 0 1 0 3 0 0 0 

Zaphanera capparis Bink-Moenen 1983 

Zaphanera capparis Bink-Moenen 1983: 199. Holotype puparium, Chad, bebedjia, 12.xi.1973, FAB & RMBM, 

on Capparis corymbosa, BMNH. 


Host: Capparidaceae: Capparis corymbosa. 

Zaphanera cyanotis Corbett 1926 

Zaphanera cyanotis Corbett 1926: 282. Syntypes. Sri Lanka: on Cyanotis sp. 

Distribution: 6: Pakistan; Sri Lanka. 

Host: Acanthaceae: Dicliptera roxburghii; Commelinaceae: Cyanotis sp. 

Zaphanera indicus Jesudasan & David 1991 

Zaphanera indicus Jesudasan and David 1991: 337. Holotype pupal case. India: on Oldenlandia corymbosa, 

IDAV. 


Host: Rubiaceae: Oldenlandia corymbosa. 

Zaphanera publicus (Singh) 1938 

Aleuroputeus publicus Singh 1938: 189. Syntypes. India: on Tephrosia purpurea, ZSI. 

Zaphanera publicus (Singh); David 1974b: 115. 


500

Host: Commelinaceae: Commelina sp.; Fabaceae: Tephrosia purpurea, Phaseolus aureus. 

SUBFAMILY UDAMOSELINAE ENDERLEIN 

Udamoselinae Enderlein 1909b: 231. Type genus. Udamoselis Enderlein 1909, by tautonomy. 

Udamoselis Enderlein 1909b: 230. Type species: Udamoselis pigmentaria Enderlein, by monotypy. 

Note: Enderlein (1909b) described these taxa based upon the wing venation of the adult whitefly. 

Solomon (1935) indicated that its wing venation was similar to that of Synaleurodicus and therefore 

should be included in the subfamily Aleurodicinae. Mound & Halsey (1978) stated “ in view of the 

fact that the structure of the pupal case of Udamoselis is unknown, the name is treated here as a 

nomen dubium in order to remove the subfamily from synonymy”. 

Recently, Martin (2007) described another species of this subfamily from Ecuador and 

reviewed and re-established the subfamily, referring to species in this subfamily as "Giant 

whiteflies" for the very large size of the adults. So far, only adults of this genus (and subfamily) 

have been found. 



estrellamarinae x 

pigmentaria prob 

Total = 2 species 0 2 0 0 0 0 0 0 0 

Udamoselis estrellamarinae Martin 2007 

Udamoselis estrellamarinae Martin 2007: 15. Holotype adult male. Ecuador, Manabi Province coast, Briseña, 5 

km north of San Vicente, 10 II 2005, Hernandez-Suarez, ex. Annona prob. muricata, BMNH. 

Distribution: 2: Ecuador. 

Host: Annonaceae: Annona prob. muricata. 

Udamoselis pigmentaria Enderlein 1909 

Udamoselis pigmentaria Enderlein 1909b: 231-233. Type male. Presumably from South America (apparently 

lost), BHU. 

Distribution: probably 2: South America. Host: no host information. 

501

SUBFAMILY BERNAEINAE SHCHERBAKOV 2000 

Bernaeidae Zherikhin 1980: 80, nomen nudum; according to Shcherbakov (2000: 30-31). 

Note: This subfamily is comprised of 5 species belonging to 4 fossil genera. 

Bernaea Schlee 1970 

Bernaea Schlee 1970: 18-29. Type species Bernaea neocomica Schlee 1970, by monotypy, SMNS. 

Bernaea neocomica Schlee 1970 

Bernaea neocomica Schlee 1970: 18-29. Female. Lebanon, in amber, SMNS. 

Distribution: 3: Lebanon. 


502

BURMOSELIS SHCHERBAKOV 2000 

Burmoselis Shcherbakov 2000: 33. Type species - Burmoselis evelynae Shcherbakov 2000, by monotypy and 


Burmoselis evelynae Shcherbakov 2000 

Burmoselis evelynae Shcherbakov 2000: 33. Holotype: Myanmar (Burma), Hukawng Valley [probably Upper 

Cretaceous Period]. 

Distribution: 6: Myanmar. 


503

HEIDEA Schlee 1970 

Heidea Schlee 1970: 9. Type- species - Heidea cretacica Schlee 1970, by monotypy. 

Heidea cretacica Schlee 1970 

Heidea cretacica Schlee 1970: 9-18. Female. Lebanon, in amber, SMNS. 

Distribution: 3: Lebanon. 


Note: Shcherbakov (2000: 30) mentioned that this species was described from Lower Cretaceous Lebanese 

amber. 

504

JULEYRODES SHCHERBAKOV 2000 

Juleyrodes Shcherbakov 2000: 32. Type species - Juleyrodes gilli Shcherbakov 2000, by original designation. 

Juleyrodes gilli Shcherbakov 2000 

Juleyrodes gilli Shcherbakov 2000: 32. Holotype: Mongolia, Ara-Hangayn [Upper Jurassic or Lower 

Cretaceous Period], 

Distribution: 5: Mongolia. 


Juleyrodes visnyai Shcherbakov 2000 

Juleyrodes visnyai Shcherbakov 2000: 33. Holotype: Kazakhstan, Karatau Range, Mikhailovka Upper Jurassic 

Period], 

Distribution: 3: Kazakhstan. 


505

Fossil species 


Aleurodicinae 

Aleurodicus aculeatus 

Aleurodicus burmiticus x 

Aleyrodinae 

Aleurochiton petri 

Bernaeinae 

Bernaea neocomica 

Burmoselis evelynae x 

Heidea cretacica x 

Juleyrodes gilli x 

Juleyrodes visnyai x 

x 

x 

x 

Total = 8 species 0 0 5 0 1 2 0 0 0 

Species Transferred Out of Aleyrodidae to Other Families 

Aleurocanthus palmae Ghesquiere to Aphidae 

Aleurodes palatina Wunn. to Trioza palatina (Wunn.) [Psyllidae] 

Aleyrodes dubia Stephens 1829 to Coniopteryx tineiformis Curtis [Coniopterygidae] 

Cerataphis lataniae Boisduval to Aphidae 

Powellia pellucida Maskell 1879 to Trioza pellucida Maskell [Psyllidae] 

Siphonaleyrodinae Takahashi 1932 to Psyllidae 

Siphonaleyrodes formosanus Takahashi 1932 to Trioza cinnamomi Boselli 1930 [Psyllidae]. 

Nomen dubia 

Aleyrodes campanulae Saalas 1942a: 127. Syntypes, Finland on Campanula grandis; synonymy with Aleyrodes 

proletella, according to Martin 2000; nomen dubium according to Martin & Mound 2007. 

Aleyrodes filicium Goldi 1886: 247. Syntypes. Brazil: on Asplenium cuneatum and other unidentified brazilian 

ferns; nomen dubium according to Martin & Mound 2007 

Aleyrodes fraxini Signoret 1868: 386. Syntypes. France: on Fraxinus sp.; Martin et al. 2000: 442; nomen 

dubium according to Martin & Mound 2007. 

Asterochiton bagnalli Harrison 1920b: 256. Syntypes. England: on Fagus sp.; nomen dubium according to 

Martin & Mound 2007. 

506

Nomen nuda 

Aleurocanthus nubilans (Buckton) 1900; Aleurodes nubilans Buckton 1900:36. Syntypes. India: Backergaunge, 

Court of Ward's Estates, on betel [Piper betle] leaves; Kirkaldy 1907: 63; nomen dubium according to 

David & Manjunatha 2003: 1 and Martin & Mound 2007. 

Aleurodes? bragini Mokrecki 1916; nomen nudum according to Martin & Mound 2007. 

Aleurodes phaseoli Schilling; Ryberg 1938:22; nomen nudum according to Martin & Mound 2007. 

Aleurodes pinicola Kirkaldy 1907; nomen nudum according to Martin & Mound 2007. 

Aleurodidarum HARRISON 1920; nomen nudum according to Martin & Mound 2007. 

Aleyrodes bifasciata Stephens 1829: 367. Syntypes. USA: Colorado, on Ambrosia artemisiaefolia; nomen 

nudum, synonymized by Kirkaldy 1907: 75; nomen nudum according to Martin & Mound 2007. 

Aleurodes phalaroides Lataste; nomen nudum and synonymy according to Quaintance 1900: 17; nomen nudum 

according to Martin & Mound 2007. 

Aleuromigda deghai Dozier & Baker; nomen nudum, synonymy according to Hussain & Khan 1945: 32; 


Aleurycerus chagientios Fulmek 1943: 8; nomen nudum according to Martin & Mound 2007. 

Bemisia oryzae Yong; nomen nudum according to Martin & Mound 2007. 

Tetra-aleurodes citriculus Dozier; Chopra 1928: 51; nomen nudum according to Martin & Mound 2007. 

507

References 

Abbas, H. M, M. S. Khan, and H. Haque. 1955. Black fly of citrus (Aleurocanthus woglumi, Ashby) in Sind and 

its control. Agric. Pakistan. 6:5-23. 

Abbasii, M. 1975. Presence au Maroc d'une espece d'aleurode Aleurothrixus floccosus Maskell. Fruits, Paris. 

30: 173-176. 

Abbasii, M. 1980. Recherche sur deux homopteres fixes des citrus, Aonidiella aurantii Mask. (Homoptera: 

Diaspididae) et Aleurothrixus floccosus Mask. (Homoptera: Aleyrodidae). Les Cahiers Recherche 

Agronomique. 35:1-168. 

Abd- El-Fattah, M. I, A. Hendi, and A. El-Said. 1986. Ecological studies on parasites of the cotton whitefly 

Bemisia tabaci (Genn.) in Egypt. Bulletin de la Societe Royale Entomologie d'Egypte. 0:95-106. 

Abd-El Gawaad, A. A, A. M. El Sayed, F. F. Shalaby, and M. R. Abo El Ghar. 1990. Natural enemies of 

Bemisia tabaci Genn. and their role in suppressing the population density of the pest. Agri. Res. Rev. 68:185- 

195. 

Abd-Rabou, S. 1996. Egyptian Aleyrodidae. Acta Phytopathologica et Entomologica Hungarica. 31(3-4), 1996: 

275-285, illustr. 

Abd-Rabou, S. 1999. Parasitism of Eretmocerus siphonini (Hymenoptera: Aphelinidae) in Egypt. Boll. Lab. 

Ent. agr. Fil. Silvestri. 55: 27-31. 

Abd-Rabou, S. 1999a. Parasitoids attacking the whiteflies (Homoptera: Aleyrodidae) infesting citrus trees in 

Egypt. Boll. Lab. Ent. agr. Fil. Silvestri. 55: 33-38. 

Abd-Rabou, S. and G. A. Evans. 2002. The Eretmocerus (Hymenoptera: Aphelinidae) of Egypt. Mitteilungen 

des Internationalen Entomologischen Vereins. 27(3/4): 115-123. 

Abd-Rabou, S. and H. Ghahari. 2004. A revision of Encarsia (Hymenoptera: Aphelinidae) from Iran. Egyptian 

Journal of Agricultural Research. 82(2): 647-683. 

Abd-Rabou, S, H. Ghahari & G. A. Evans. 2005. Iranian Eretmocerus-species including two new species 

(Hymenoptera: Chalcidoidea: Aphelinidae) parasitoids of whiteflies (Sternorrhyncha: Aleyrodidae). 

Mitteilungen des Internationalen Entomologischen Vereins. 30 (3/4): 157-176. 

Abd-Rabou, S, H. Ghahari, J. Huang & Z. Boucek. 2005. New records of aphelinid and pteromalid wasps 

(Hymenoptera: Chalcidoidea: Aphelinidae: Pteromalidae) from Iran. Egyptian Journal of Agricultural Research. 

508

83(4): 1619-1623. 

Abdel-Fattah et al. 1986. Ecological studies on parasites of the cotton whitefly Bemisia tabaci (Genn.) in Egypt. 

Bulletin de la Societe Royale Entomologie d'Egypte. 0:95-106. 

Abdelrahman, A. A. 1986. The potential of natural enemies of the cotton whitefly in Sudan Gezira. Insect Sci. 

Applic. 7:69-73. 

Abdelrahman, A. A. and B. Munir. 1989. Sudanese experience in integrated pest management of cotton. Insect 

Sci. Applic. 10:787-794. 

Abou-Awad, B. A, and S. A. Elsawi. 1993. Biology and life table of the predacious mite, Agistemus exsertus 

Gonz. (Acari: Stigmaeidae). Anzeiger Fuer Schaedlingskunde Pflanzenschutz Umweltschutz. 66:101-103. 

Agarwal, M. M. 1964. Studies of forms of Aphelinidae (Hymenoptera - Chalcidoidea) collected at Aligarh 

(India). I. Proceedings of the Indian Academy of Sciences (B). 59:263-286. 

Agekyan, N. G. 1965. Conwentzia psociformis Curt. (Neuroptera: Coniopterygidae) a little known 

entomophage in the Georgian Republic. Entomol. Rev. 44(1):84-88. 

Agekyan, N. G. 1977. Clitostethus arcuatus Rossi (Coleoptera: Coccinellidae), a predator of the citrus whitefly 

in Adzharia. Entomol. Rev. 56:22-23. 

Agekyan, N. G. 1978. A little known entomophagous insect Semidalis aleyrodiformis Stephens (Neuroptera: 

Coniopterygidae) in Adzharia. Entomol. Rev. 57:348-350. 

Al-Hitty, A, A. K. Abbass, and A. S. A. Ali. 1988. Development and survival of Deraecoris pallens Reut. 

(Heteroptera, Miridae) on Bemisia tabaci Gen. and Aphis gossypii. Glv. J. Agri. Water Resource Res, Plant 

Prod. 7:115-123. 

Albajes, R, R. Gabarra, C. Castane, O. Alomar, J. Arno, J. Bellavista, M. Marti, J. Moliner, and M. Ramirez. 

1994. Implementation of an IPM program for spring tomatoes in Mediterranean greenhouses. IOBC/WPRS 

Bull. 17(5):14-21. 

Albert, R, and H. Schneller. 1991. Biological pest control on ornamental plants I. Experiences with 

Trialeurodes vaporariorum and Bemisia tabaci. Gartnerborse und Gartenwelt. 91:10-15. 

509

510 

Alomar, O, and R. Albajes. 1992. Management of the facultative predator Dicyphus tamaninii Wagner in field 

tomatoes. In Proceedings of workshop "Use of facultative zoo-phytophagous bugs in IMP" XIX International 

Congress of Entomology, Beijing, China, June 28 - July 4 1992. pp. 305. 

Alomar, O, C. Castane, R. Gabarra, and R. Albajes. 1990. Mirid bugs: another strategy for IPM on 

Mediterranean vegetable crops. IOBC/WPRS Bull. 13:6-9. 

Alvarez, P, L. Alfonseca, A. Abud, A. Villar, R. Rowland, E. Marcano, J. Borbon, and L. Garrido. 1993. Las 

moscas blancas in La Republica Dominicana. In L. Hilje and O. Arboleda [eds.], Serie Tecnica, Informe 

Tecnico No. 205: Las Moscas Blancas (Homoptera: Aleyrodidae) en America Central Y El Caribe. CATIE, 

Turrialba, Costa Rica. pp. 34-37. 

Anderson, P. K, Martin, J. H, Hernandez, P. & A. Lagnaoui. 2001. Bemisia afer sens. lat. (Homoptera: 

Aleyrodidae) outbreak in the Americas. Florida Entomologist. 84 (2): 316-317. 

Annecke, D.P. and H. P. Insley. 1970. New and little-known species of Azotus Howard, Ablerus Howard and 

Physcus Howard (Hym, Aphelinidae) from Africa and Mauritius. Bulletin of Entomological Research. 

60(2):237-251. 

Antadze, A. I, and T. V. Timofeyeva. 1976. A predator of the citrus whitefly. Zashchita Rastenii (Moskva). 

10:46. 

Ardaillon, J. B. and F.Cohic. 1970. Notes sur quelques aleurodes de la foret dedu Banco (Cote d'Ivoire). Annals 

Univ. Abidjan (E). 3: 269-283. 

Ardon, M. L, R. A. Cuellar, and J. I. Henriquez. 1992. Reconocimento de Enemigos Naturales y Hospederos de 

Moscas Blancas (Homoptera: Aleyrodidae) en Tres Zonas de la Cuenca del Lago de Iopango. Inginiero 

Agronimo Thesis, Universidad de El Salvador.

511 

Arnal, E, A. Chavez & F. Ramos. 2003. Parasitoides registrados en moscas blanca (Homoptera: Aleyrodidae) 

colectados en Venezuela durante el periodo 1991-2000. Entomotropica. 18(3): 183-191. 

Arzone, A. 1976. Indagini su Trialeurodes vaporariorum ed Encarsia tricolor in pica aria. Inf. Fitopatol. 26:5- 

10. 

Arzone, A, A. Alma, and L. Tavella. 1990. Ruolo dei Miridi (Rhynchota: Heteroptera) nella limitazione di 

Trialeurodes vaporariorum Westwood (Homoptera: Aleyrodidae). Boll. Zool. agr. Bachic. 22:43-52. 

Ashby, S. F. 1915. Notes on diseases of cultivated crops observed in 1913-1914. Bull. Dep. Agric. Jamaica. 2: 

299-327. 

Ashmead, W. H. 1880. Orange insects: a treatise on the injurious and beneficial insects found in the orange 

trees in Florida. 1-xv,. 1-78, plates I-IV. Jacksonvill. 

Ashmead, W. H. 1885. The orange Aleurodes (Aleurodes citri n. sp.). Florida Dispatch. 2(42): 704. 

Ashmead, W. H. 1887. Studies on the North American Proctotrupidae with descriptions of new species from 

Florida. (Part 1.). Entomologica Americana. 3:97-100. 

Ashmead, W. H. 1893. Monograph of the North American Proctotrypidae. Amitus Haldeman. Bull. U.S. 

National Museum. 45: 292-294. 

Ashmead, W. H. 1893a. Notes on cotton insects found in Mississippi. Insect Life. 7: 27. 

Ashmead, W. H. 1895. Notes on cotton insects found in Mississippi. Insect Life Wash. 7: 323.

Ashmead, W. H. 1900. Genera of the Encyrtinae. Subfamily III - Signiphorinae. Proceedings of the U. S. 


Ashmead, W. H. 1900a. On the genera of the Chalcid-flies belonging to the subfamily Encyrtinae. Proc. U.S. 


Ashmead, W. H. 1904. Descriptions of new genera and species of Hymenoptera from the Philippine Islands. 

Proceedings of the United States National Museum. 28:127-158. 

Ashmead, W. H. 1904a. New generic names in the Chalcidoidea. Proceedings of the Entomological Society of 

Washington. 6:126. 

Avidzba, N. S. 1983. Biology of citrus whitefly and its integrated management. Proceedings of 10th 

International Congress of Plant Protection, vol. 3, Brighton, England, 20-25 Nov. 1983. 

Azim, M.N. and S. A. Shafee. 1980. Indian species of the genus Trichaporus Foerster (Hymenoptera: 

Aphelinidae). Journal of the Bombay Natural History Society. 76(2):335-338. 

Babrikova, T. 1979. Study of the biology of Chrysopa perla. Ras. nauki (Plant Sci.). 16:95-100. 

Back, E. A. 1909. A new enemy of the Florida orange (Aleyrodes howardi Quaintance). Journal of Economic 

Entomology. 2: 448-449. 

Back, E. A. 1912. Notes on Cuban whiteflies with descriptions of two new species. Canadian Entomologist. 44: 

145-153. 

512

513 

Baerensprung, F.V. 1849. Beobachtungen uber einige einheimische Arten aus der Familie der Coccinen. In - 

Alton & Burmeister. Zig Zool. Zoot. Palaeoz. Leipzig 1848-49, 212 pp. 165-176 p. 

Baerensprung, F.V. 1973. Oko-faunistische Untersuchungen an Mottenschildlausen (Homoptera: Aleyrodina) in 

der Umgebung von Jena/Thuringen. Wiss.Z. Friedrich Schiller-Univ. Jena. 22: 507-517. 

Bailey, P. T, Martin, J. H, Noyes, J. S. & A. D. Austin. 2001. Taxonomy and biology of a new species of 

Zaphanera (Hemiptera: Aleyrodidae) and its association with the widespread death of western myall trees, 

Acacia papyrocarpa, near Roxby Downs, South Australia. Transactions of the Royal Society of South 

Australia. 125 (2): 83: 96. 

Baker, A. C. 1923. An undescribed orange pest from Honduras. Journal of Agricultural Research. 25: 253-254. 

Baker, A. C. and M. L. Moles. 1920. A new species of Aleyrodidae found on Azalea (Hom.). Proceedings of the 

Entomological Society of Washington. 22: 81-83. 

Baker, A. C. and M. L. Moles. 1923. The Aleyrodidae of South America with descriptions of four new Chilean 

species. Revista Chil. Hist. Nat. 25: 609-648. 

Baker, J. M. 1937. Notes on some Mexican Aleyrodidae. An. Inst. Biol. Univ. Mex. 8: 599-629. 

Banks, N. 1911. Descriptions of new species of North American neuropteroid insects. Transactions of the 

American Entomological Society. 37: 355-360. 

Barbagallo, S, I. Longo, C. Patti, and C. Rapisarda. 1992. Efficiency of biological control against citrus 

whiteflies of Italy. Boll. Zool. Agr. Bachic. 24:121-135.

Barnadas, I. 1994. Avaluacio de la capacitat depredadora dels mirids Dicyphus tamaninii I Macrolophus 

caliginosus sobre l'aleirodid Bemisia tabaci biotype B. Projecte final de carrera d'enginyeria agronoma, 

Universitat de Lleida, Lleida, Italy. 

Barnes, H. F. 1930. Gall midges (Cecidomyidae) as enemies of the Tingidae, Psyllidae, Aleyrodidae and 

Coccidae. Bulletin of Entomological Research. 21: 319-329. 

Basha, A, A. A. Yousef and E. M. Mostafa. 2002. Morphology and biology of Euseius metwallyi n. sp. (Acari: 

Gamasida: Phytoseiidae). Acarologia. 42: 29-37. 

Bathon, V. H, and J. Pietrzik. 1986. On the food consumption of Clitostethus arcuatus (Rossi) (Col, 

Coccinellidae), a predator of Aleurodes proletella L. (Hom, Aleyrodidae). Journal of Applied Entomology. 

102:321-326. 

Bathon, V. H, and J. Pietrzik. 1987. Zur Biologie des Bogen-Marienkagers, Clitostethus arcuatus (Rossi) (Col, 

Coccinellidae), einem Vertilger der Kohomottenlaus Aleurodes proletella (Linne) (Hom, Aleyrodidae). 

Mitteilungen der Deutschen Gesellschaft fur Allgemeine und Angewandte Entomologie. 5:204-205. 

Becker, H, J. Corliss, J. De Quarto, M. Gerrietts, D. Senet, D. Stanly, and M. Wood. 1992. Get the whitefly 

swatters. Agric. Res. (Washington). 49:4-13. 

Bedford, E. C. G, E. D. Thomas. 1965. Biological control of citrus blackfly, Aleurocanthus woglumi (Ashby) 

(Homoptera, Aleyrodidae) in South Africa. J. Entomol. Soc. S. Africa. 28, 117-132. 

Beingolea, O. G. 1980. Cotton protection through integrated control. Consultancy report TCP/SYR/001 ed. 

FAO, Rome. 

514

Bellows, T. S, Jr, T. D. Paine, and D. Gerling. 1992. Development, survival, longevity, and fecundity of 

Clitostethus arcuatus (Coleoptera: Coccinellidae) on Siphoninus phillyreae (Homoptera: Aleyrodidae) in the 

laboratory. Environmental Entomology. 21:659-663. 

Bellows, T. S, Jr, T. D. Paine, J. R. Gould, L. G. Bezark, and J. C. Ball. 1992. Biological control of ash 

whitefly: a success in progress. California Agriculture. 46:24-28. 

Bellows, T. S, Perring, T. M, Gill, R. J. and D. H. Headrick. 1994. Description of a new species of Bemisia 

(Homoptera: Aleyrodidae). Annals of the Entomological Society of America. 87: 195-206. 

Bemis, F. E. 1904. The aleyrodids or mealy-winged flies of California with reference to other American species. 

Proceedings of the U. S. National Museum. 27: 471-537. 

Benuzzi, M, and M. Mosti. 1994. I miridi predatori di aleyrodidi. Inf. Fitopatol. 44:25-30. 

Berger, E. W. 1909. Whitefly studies in 1908. Bull. Fla. agric. Exp Stn. 97: 43-71. 

Berger, E. W. 1910. Whitefly control. Bulletin of the Florida Agricultural Experiment Station, Gainesville. 113: 

5-28. 

Berger, E. W. 1920. Natural enemies of scale insects and whiteflies of Florida. Qt. Bull. State Plant Board 

Florida. 5:141-154. 

Biezanko, C. M. and R. G. Freitas. 1939. Catalogo dos insetos encontados em Pelotas e seus arredores Fasciculo 

II Homopteros. Bolm Esc Agron Vet Eliseu Maciel. 26: 6. 

515

516 

Bink-Moenen, R. M. 1976. A new whitefly of Erica tetralis: Tetraleurodes ericae sp. n. (Homoptera: 

Aleyrodidae). Bull Ber Amst. 36: 17-19. 

Bink-Moenen, R. M. 1983. Revision of the African whiteflies (Aleyrodidae) mainly based on a collection from 

Tchad. Monografieen van de Nederlandse Entomologische Vereniiging, AmsterdamMonografieen van de 

Nederlandse Entomologische Vereniiging, Amsterdam. No. 10, 211 pp. 

Bink-Moenen, R. M. 1989. A new species and new records of European whiteflies (Homoptera: Aleyrodidae) 

from heathers (Erica spp.). Entomologist's Gazette. 40: 173-181. 

Bink-Moenen, R. M. 1991. Comparison between Neopealius rubi and Bemisia tabaci in Europe (Homoptera: 

Aleyrodidae). Entomologische Berichten. 51: 29-37. 

Bink-Moenen, R. M. 1992. Whitefly from Mediterranean evergreen oaks (Homoptera: Aleyrodidae). Systematic 


Bink-Moenen, R. M and D. Gerling. 1991. Aleyrodidae of Israel. Bollettino Del Laboratorio Di Entomologia 

Agraria Filippo Silvestri. 47 (1990): 3-49. 

Blanchard, E. 1852. Fauna Chilena - Aleurodidas. Historia fisica y politica de Chile Zoologica. 471 pp. (318- 

320). 

Blanchard, E. E. 1918. Una nueva especied de 'Aleurothrixus (Homoptera: Aleyrodidae). Physis B. Aires. 4: 

244-347. 

Blanchard, E. E. 1936. Apuntes sobre calcidoideos argentinos, nuevos y conocidos. Revista de la Sociedad 

Entomologica Argentina. 8: 7-32.

Blanchard, E. E. 1937. Informaciones tecnicas. Boletin Informativo de la Direccion de Sanidad Vegetal. 1:25- 

32. 

Blanchard, E. E. 1940. Division de Zoologia Agricola. Boletin Informativo de la Direccion de Sanidad Vegetal. 

10:24-30. 

Bock, I. R. 1982. Drosophilidae of Australia V. Remaining genera and synopsis (Insecta: Diptera). Australian 

Journal of Zoology. 89:1-164. 

Bodkin, G. E. 1919. Notes on the Coleoptera of British Guiana. Entomologist's Monthly Magazine. Sept.:210- 

216. 

Bogran, C. E, J. J. Obrycki and R. D. Cave. 1998. Assessment of biological control of Bemisia tabaci 

(Homoptera: Aleyrodidae) on common bean in Honduras. Florida Entomologist. 81(3): 384-394. 

Bondar, G. 1922. Insectos damninhos e molestias do coquerio- no Brasil. Bahia. 113 pp. 

Bondar, G. 1923. Aleyrodideos do Brasil. Bahia. 183 pp. 

Bondar, G. 1928. Aleyrodideos do Brasil (2a contribuicao). Boletim Lab. Path. veg. Est. Bahia. 5: 1-37. 

Bondar, G. 1928a. Molestia nos cafezaes da Bahia. Correio Agricola. 6:82-85. 

Bondar, G. 1929. Uma provaco da lavoura cafeeira na Bahia. Correio Agricola. 6: 252-256. 

517

Bondar, G. 1931. Uma nova praga das laranjeiras. O Campo. 2(5): 24. 

Borah, D. C, and P. S. Rai. 1989. 1989. Potentality of Ambyseius ovalis (Acari: Phytoseiidae) as a biological 

control agent on Bemisia tabaci (Homoptera: Aleyrodidae). In G. P. ChannaBasavanna and C. A. Viraktamath 

[eds.], Progress in acarology, vol. 2. Oxford and IBH Publishing CO. Pvt. LTD, New Delhi. pp. 375-379. 

Boscan de Martínez, N, B. J. Terán, and F. Geraud. 1982. Enemigos naturales de la mosca prieta do los citricos 

Aleurocanthus woglumy [woglumi] (Homoptera: Aleyrodidae) en Venezuela. Agron. Tropic. (Maracay). 

29:453-458. 

Boucek, Z. 1988. Australasian Chalcidoidea (Hymenoptera). A Biosystematic revision of genera of fourteen 

families, with a reclassification of species, CAB International, Wallingford. -. 832. 

Bouche, J. Fr. 1851. Beschreibung zwei neuer Arten der Gattung Aleurodes. Stettin. Ent. Zig. 12: 108-110. 

Bounfour, M. 1985. Biology and ecology of Euseius scutalis (Athias-Henriot) (Acarina: Phytosiidae). M.Sc. 

Thesis, Univ. of California, Riverside. -. 

Brazzle, J. R, K. M. Heinz, M. P. Parrella, and C. H. Pickett. 1994. Field evaluations of Delpahstus pusillus for 

control of Bemisia tabaci infesting upland cotton. In T. J. Henneberry, N. C. Toscano, R. M. Faust, and J. R. 

Coppedge [eds.], Sweetpotato whitefly: 1994 supplement to the five-year national research and action plan. 

USDA/ARS. -. 

Breene, R. G, D. A. Dean, and W. Quarles. 1994. Predators of sweetpotato whitefly. IPM Prac. 16:1-9. 

Breene, R. G, R. L. Meagher, D. Nordlund, and Yin-T. Wang. 1992. Biological control of Bemisia tabaci 

(Homoptera: Aleyrodidae) in a greenhouse using Chrysoperla rufilabris (Neuroptera: Chrysopidae). Biological 

518

Control. 2:9-14. 

Brethes, J. 1914. Les ennemis de la Diaspis pentagona dans la Republique Argentine. Nunquam otiosus, Buenos 

Aires. 1: 1-16. 

Brethes, J. 1916. Description de deux hymenopteres chiliens. Annales de Zoologia Aplicada. 3: 24-27. 

Brethes, J. 1916a. Hymenopteres parasites de l'Amerique Meridionale. An. Mus. Nac. Hist. Nat. Buenos Aires. 

27: 401-430. 

Britton, W. E. 1905. Some new or little known Aleyrodidae from Connecticut - I. Entomological News. 16: 65- 

67. 

Britton, W. E. 1906. Some new or little known Aleyrodidae from Connecticut - II. Entomological News 17: 

127-130. 

Britton, W. E. 1907. Some new or little known Aleyrodidae from Connecticut - III. Entomological News. 18: 

337-342. 

Britton, W. E. 1923. Family Aleyrodidae. Bull. Conn. St. Geol. Nat. Hist. 34: 335-345. 

Broadley, R, and M. Thomas. [eds.]. 1995. The Good Bug Book. Beneficial insects and mites commercially 

available in Australia for biological pest control. Australian Biological Control Inc, Richmond. -. 

Brooks, S. J. and P. C. Barnard. 1990. The green lacewings of the world: a generic review (Neuroptera: 

Chrysopidae). Bulletin of the British Museum (Natural History) (Entomology). 59(2): 117-286. 

519

Browning, H. W, and J. L. Stimac. 1994. Classical biological control of whiteflies on citrus. In D. Rosen, F. D. 

Bennett, and J. L. Capinera [eds.], Pest management in the subtropics: biological control- a Florida perspective. 

Intercept, Andover, U.K. pp. 79-100. 

Bruner, S. C. 1931. Informe del departament de entomologia y fitopatologia, ejercico de 1929 a 1930. Estacion 

Exper. Agron, Santiago de las Vargas, Cuba. -. 

Brzezinski, K. 1982. Report of investigations on the morphology, biology, and ecology of the heteropteran 

Macrolophus costalis (Fieb.) [sic.] (Heteroptera, Miridae) and its predation in relation to greenhouse whitefly 

(Trialeurodes vaporariorum) Westwood. Materialy. 22/23:285-292. 

Buckton, J. 1900. Description of a new species of Aleurodes destructive to betel. Indian Museum Notes. 5: 36. 

Buren, W. F, and W. H. Whitcomb. 1979. Ants of citrus. Proceedings of International Congress of Citrus 

Culture. 

Butler, C. G. 1938. A further contribution to the ecology of Aleurodes brassicae Walk. (Hemiptera). 

Proceedings of the Royal Entomological Society of London (A). 13:161-172. 

Butler, G. D. 1965. Spanogonicus albofasciatus as an insect and mite predator (Hemiptera: Miridae). Journal of 

the Kansas Entomological Society. 38:70-75. 

Butler, G. D, and T. J. Henneberry. 1988. Laboratory studies of Chrysoperla carnea (Stephens) predation on 

Bemisia tabaci (Gennadius). Southwestern Entomol. 13(3):165-170. 

520

521 

CIBC Pakistan Station 1983. 1983. Studies on potential biological control agents of whiteflies in Pakistan. 

Mar.1979-Feb.1982. Unpublished report. Ravalpindi, Pakistan. 

Caballero, R. 1993. Moscas blancas neotropicales (Homoptera: Aleyrodidae): hospedantes, distribucion, 

enemigos naturales e importancia economica. In L. Hilje and O. Arboleda [eds.], Serie Tpcnica, Informe 

Tpcnico No. 205: Las Moscas Blancas (Homoptera: Aleyrodidae) en Amprica Central Y El Caribe. CATIE, 

Turrialba, Costa Rica,. pp. 10-15. 

Calabro, M. 1994. I Miridi predatori: loro biologia e possibili applicazioni in lotta biologica-integrata esperienze 

maturate in Sicilia. In Atti Convengno , Cagliari, 30 gennaio 1992. 

pp. 29-42. 

California PPDR. 2005. California Plant Pest and Disease Report July 2002-July 2005. California Plant Pest & 

Disease Report. 21(1): 1-78. 

Cameron, P. 1912. Descriptions of new genera and species of parasitic Hymenoptera taken at Kuching, 

Sarawak, Borneo, by Mr John Hewitt, B.A. (parts 1-6). Societas Entomologica, Frankfurt. 

Cameron, P. 1912a. On a collection of parasitic Hymenoptera (chiefly bred) made by Mr. Walter W. Froggatt, 

F.L.S. in New South Wales, descriptions of new genera and species. Part iii. Proceedings of the Linnean Society 

of New South Wales. 37:172-216. 

Capco, S. R. 1959. A list of plant pests of the Philippines with special reference to field crops, fruit trees and 

vegetalies. Philippine Journal of Agriculture. 22: 1-80. 

Carapia-Ruiz, V. E. H. Gonzalez-Hernandez, J. Romero-Napoles, L. D. Ortega-Arenas & S. D. Koch. 2003). 

Descripcion de dos nuevas especies de Trialeurodes (Cockerell) (Homoptera: Aleyrodidae) y una clave para las

especies de Mexico. Acta Zoologica Mexicana 90: 93-101. 

Carayon, J. 1961. Queleques remarque sur les Hemipteres: leur importance commune insects auxiliares et les 

possibiltps de lur utilisation dans la lutte biologique. Entomophaga. 6:133-141. 

Carvalho, M. B. de, M. Aquino, M. Oliveira. 1972. Considerations on the biological control of Aleurodicus 

cocois (Curtis, 1846) (Homoptera: Aleyrodidae, cashew whitefly. Anais do Instituto de Ciencias Biologicas. 2: 

25-30. 

Carver, M. and L. A. Reid. 1996. Aleyrodidae (Hemiptera: Sternorrhyncha) of Australia. Systematic catalogue, 

host-plant spectra, distribution, natural enemies and biological control. Technical Paper, Division of 

Entomology, Commonwealth Scientific and Industrial Research Organization, Canberra. 37: 55 pp. 

Cassino, P. C. R. and F. N. Nascimento. 1999. Aleirodideos (Homoptera: Aleyrodidae) em plantas citricas no 

Brasil: distribuicao e identificacao. Anais da la Sociedade Entomologica do Brasil. 28(1): 75-83. 

Castineiras, A. 1995. Natural enemies of Bemisia tabaci (Homoptera: Aleyrodidae) in Cuba. Florida 

Entomologist. 78(3): 538-540. 

Chandra, J, and P. N. Avasthy. 1978. First record of Cybocephalus sp. predaceous on whitefly, Neomaskellia 

bergii Signoret. Indian J. Entomol. 40:445-446. 

Chavez Torres, H. A. 1974. Early season population of whiteflies on citrus and ornamentals in north Central 

Florida. M.S. Thesis, Univ. of Florida. 

Chavez, H.A. 1996. Una nueva especie de Dirphys (Hymenoptera: Aphelinidae). Boletin de Entomologia 

Venezolana. 11: 11-17. 

522

523 

Chen, C. and C. Ko. 2006. Lipaleyrodes emiliae, a new species of whitefly (Hemiptera: Aleyrodidae) from 

Taiwan and Hong Kong. Zootaxa. 1331: 31-54. 

Chen, Y. and C. Ko. 2007. Singhiella melanolepis, a new species of whitefly (Hemiptera: Aleyrodidae) from 

Taiwan with remarks on the genus Singhiella Sampson. Zootaxa 1390: 1-16. 

Cherry, R, and R. V. Dowell. 1979. Predators of the citrus blackfly (Hom.: Aleyrodidae). Entomophaga. 

24:385-391. 

Chien, C. C. and S. C. Chui. 1986. Biology of the citrus spiny blackfly Aleurocanthus spiniferus. Journal of 

Agricultural Research of China. 35: 222-229. 

Chou, I. and F. Yan. 1988. New species and new records of Aleyrodidae (Homoptera) from China. 

Entomotaxonomia 10: 243-247. 

Chou, K. C, Y. S. Su, L. Y. Chou, and C. C. Ko. 1996. New records of Aphelinidae (Hymenoptera) from 

Taiwan. Journal of Agricultural Research of China. 45(2): 195-202. 

Chou, K.C. and L.Y. Chou. 1994. A new species of Encarsiella from Taiwan. Journal of Agricultural Research 

of China. 43: 469-472. 

Chou, L. Y, Y. S. Su, K. C. Chou and C. C. Ko. 1996. Two new species of Encarsia (Hymenoptera: 

Aphelinidae) from Taiwan. Plant Protection Bulletin (Taichung). 38(2), June 1996: 137-142, illustr. 

Clausen, C. P. 1934. The natural enemies of Aleyrodidae in tropical Asia. Phil. J. Sci. 53:253-265.

Clausen, C. P. 1940. Entomophagous Insects. McGraw Hill, New York. -. 

Clausen, C. P. 1958. Biological control of insect pests. Ann. Rev. Entomol. 3:291-310. 

Clausen, C. P, and P. A. Berry. 1932. The citrus blackfly in Asia and the importation of its natural enemies into 

tropical America. Tech. Bull. 320. USDA, Washington, D.C. 

Cockerell, T. D. A. 1893. A third species of Aleurodicus. Entomologist's Monthly Magazine. 29: 105-106. 

Cockerell, T. D. A. 1896. A Mexican Aleurodicus. Canadian Entomologist. 28: 302. 

Cockerell, T. D. A. 1896a. New species of insects taken on a trip from the Mesilla Valley to the Sacramento 

Mts, New Mexico. Aleurodidae. Journal of the New York Entomological Society. 4: 207. 

Cockerell, T. D. A. 1897. A new Aleurodes found on Aquilegia. Journal of the New York Entomological 

Society. 5: 42. 

Cockerell, T. D. A. 1897a. A new Aleurodes on Rubus from Florida. Journal of the New York Entomological 

Society. 5: 96-97. 

Cockerell, T. D. A. 1898. A new Aleurodes on oak. Canadian Entomologist. 30: 264. 

Cockerell, T. D. A. 1898a. Three new Aleurodidae from Mexico. Psyche, Cambridge. 8: 225-226. 

Cockerell, T. D. A. 1900. Economic entomology in Arizona. The marked mealy-wing. Sci. Gossip. 6: 366-367. 

524

Cockerell, T. D. A. 1902. The classification of the Aleyrodidae. Proceedings of the Academy of Natural 

Sciences of Philadelphia. 54: 279-283. 

Cockerell, T. D. A. 1903. Aleyrodes (Trialeurodes) vitrinellus Ckll. Entomological News. 14: 241. 

Cockerell, T. D. A. 1903a. The whitefly (Aleyrodes citri) and its allies, In Gossard, H. A. Whitefly (Aleyrodes 

citri). Bull. Fla. agric. Exp Stn. 67: 599-666. 

Cockerell, T. D. A. 1910. A new Aleyrodes on Ambrosia. Canadian Entomologist. 42: 370-371. 

Cockerell, T. D. A. 1910a. A new Aleyrodes on bearberry. Canadian Entomologist. 42: 171-172. 

Cockerell, T. D. A. 1911. An Aleyrodes on Euphorbia and its parasite. Entomological News. 22: 462-464. 

Cockerell, T. D. A. 1919. Insects in Burmese amber. Entomologist. 52: 241. 

Cohen, A. C, D. Nelson, J. Buckner, and D. Brummett. 1996. Nutrient rewards for predators of whiteflies: 

Generalists' requirements, In T. J. Henneberry, N. C. Toscano, R. M. Faust, and J. R. Coppedge [eds.], 

Silverleaf whitefly: 1996 supplement to the 5-year national research and action plan,. USDA/ARS 1996-01. pp. 

28. 

Cohen, A. C, L. Smith, and D. Brummett. 1997. Development of an artificial diet for Chrysoperla carnea. In T. 

J. Henneyberry [ed.], Silverleaf whitefly: 1997 supplement to the five-year national research and action plan: 

progress review, technology transfer, and new research and action plan, USDA/ARS 1997-02. pp. 137. 

525

526 

Cohen, A. C, L. Smith, and D. Brummett. 1997a. Measurement of nutrient rewards for predators consuming 

whiteflies: sterols from Bemisia argentifolii, pp. 138. In T. J. Henneyberry [ed.], Silverleaf whitefly: 1997 

supplement to the five-year national research and action plan: progress report. 

Cohen, A. C, R. T. Staten, and D. Brummett. 1995. Generalist and specialist predators: how prey profitability 

and nutrient reward influence the two strategies or whiteflies as "junkfood". Proceedings Beltwide Cotton 

Conferences (Addendum) 4-7 January 1995, San Antonio, TX. National Cotton Council, Memphis, TN. pp. 71- 

72. 

Cohen, A. C, and D. N. Byrne. 1992. Geocoris punctipes as a predator of Bemisia tabaci: a laboratory 

evaluation. Entomol. Exp. Appl. 64:195-202. 

Cohic, F. 1959. Aleyrodidae actuellement connus de Nouvelle-Caledonie et dependances. Agron. trop. Nogent. 

14: 242-243. 

Cohic, F. 1959a. Contribution a l'etude des aleurodes de Nouvelle-Caledonie [Hom.] Orchamus dumbleton n. 

sp. Bull. Soc. Ent. Fr. 64: 130-136. 

Cohic, F. 1963. Contribution a l'etude des aleurodes africains (2e Note). Cahiers de l'Office de la Recherche 

Scientifique et Technique Outre-Mer (Serie Biologie). 2: 13-72. 

Cohic, F. 1966. Contribution a l'etude des aleurodes africains (1e Note). Cahiers de l'Office de la Recherche 


Cohic, F. 1966a. Contribution a l'etude des aleurodes africains (2e Note). Cahiers de l'Office de la Recherche 


Cohic, F. 1966b. Contribution a l'etude des aleurodes africains (3e Note). Cahiers de l'Office de la Recherche


Cohic, F. 1968. A propos de Dialeurodicus elongatus Dumbl. et de Stenaleyrodes vinsoni Tak. Aleurodes 

parasities des palmiers. Annls. Univ. Abidjan (E). 1: 9-17. 

Cohic, F. 1968a. Contribution a l'etude des aleurodes africains (1e Note). Cahiers de l'Office de la Recherche 


Cohic, F. 1968b. Contribution a l'etude des aleurodes africains (3e Note). Cahiers de l'Office de la Recherche 


Cohic, F. 1968c. Contribution a l'etude des aleurodes africains (4e Note). Cahiers de l'Office de la Recherche 


Cohic, F. 1969. Contribution a l'etude des aleurodes africains (5e Note). Annls. Univ. Abidjan (E). 2: 1-156. 

Cole, F. R. 1926. New facts about the citrus aphid and its natural enemies. Proc. Fla. Hort. Soc. 39:156-158. 

Colombo, M, and F. R. Eordegh. 1991. Ritrovamento de Coenosia attentuata attivo predatore di aleirodidi, in 

colture protette in Liguria e Lombardia. Inf. Agrario. -. 10:187-189. 

Compere, H. 1931. A revision of the species of Coccophagus a genus of hymenopterous coccid-inhabiting 

parasites. Proceedings of the United States National Museum. 78: 1-132. 

Compere, H. 1936. Notes on the classification of the Aphelinidae with descriptions of new species. University 

of California Publications in Entomology. 6(12):277-322. 

527

Compere, H. 1947. A report on a collection of Encyrtidae with description of new genera and species. 

University of California Publications in Entomology. 8 (1): 1-24. 

Compere, H. 1961. The red scale and its insect enemies. Hilgardia. 31: 173-278. 

Compere, H. and D. P. Annecke. 1961. Descriptions of parasitic Hymenoptera and comments (Hymenopt.: 

Aphelinidae, Encyrtidae, Eulophidae). Journal of the Entomological Society of Southern Africa. 24:17-71. 

Corbett, G. H. 1926. Contribution towards our knowledge of the Aleyrodidae of Ceylon. Bulletin of 

Entomological Research. 16: 267-284. 

Corbett, G. H. 1927. Three new aleyrodids on coconuts in Malaya. Malay. Agriculture Journal. 15: 24-25. 

Corbett, G. H. 1933. Aleurodidae of Malaya. Stylops. 2: 121-129. 

Corbett, G. H. 1935. Malayan Aleurodidae. Journal of the Federated Malay States Museum. 17: 722-852. 

Corbett, G. H. 1935a. On new Aleurodidae (Hem.). Annals and Magazine of Natural History (10). 16: 240-252. 

Corbett, G. H. 1935b. Three new aleurodids (Hem.). Stylops. 4: 8-10. 

Corbett, G. H. 1936. New Aleurodidae (Hem.). Proceedings of the Royal Entomological Society of London (B). 

5: 18-22. 

528

529 

Corbett, G. H. 1939. A new species of Aleurodidae from India. Indian Journal of Entomology. 1: 69-70. 

Corbett, G. H. 1939a. New Aleurodidae (Hem.). Proceedings of the Royal Entomological Society of London 

(B). 5: 18-22. 

Costa Lima, A. Da. 1928. Contribuicao as estudio dos aleyrodideos da subfamilia Aleurodicinae. Supplto 

Mems. Inst. Oswaldo Cruz. 4: 128-140. 

Costa Lima, A. Da. 1936. Terceiro catalogo dos insectos que vivem nas plantas do Brasil. Rio de Janeiro. 460 

pp. 

Costa Lima, A. Da. 1942c. Sobre aleirodideos do genero 'Aleurothrixus' (Homoptera). Revista Brasileira de 

Biologia. 2:419-426. 

Costa Lima, A. M. da. 1968. Quarto catalogo dos insectos que vivem nas plantas do Brasil seus parasitos e 

predadores, 4th ed. Ministrio da Agricultura, Rio de Janeiro, Brasil. part 2, 1st section. -. -. 

Curtis, J. 1846. Entomology: Aleyrodes cocois (the cocoa-nut Aleyrodes). Gardener's Chronicles. 1846:284- 

285. 

Danzig, E. M. 1962. The whiteflies (Homoptera: Aleyrodidae) of Leningrad environments). [In Russian]. Trudy 

Zoologicheskogo Instituta. Akademiya Nauk SSSR. Leningrad. 31: 13-21. 

Danzig, E. M. 1964. Order Homoptera. Suborder Aleyrodinea. In. Keys to the insects of the European USSR, 

Bei-Bienko ed. 1: 482-489. 

Danzig, E. M. 1964a. The whiteflies (Homoptera: Aleyrodidoidea) of the Caucasus. [In Russian].

Entomologicheskoe Obozrenie. 43: 633-646. 

Danzig, E. M. 1966. The whiteflies (Homoptera: Aleyrodidoidea) of the Southern Primor'ye (Soviet Far East). 

Entomologicheskoe Obozrenie. 45(2): 197-209. 

Danzig, E. M. 1969. On the fauna of the whiteflies (Homoptera: Aleyrodidoidea) of Soviet Central Asia and 

Kazakhstan [In Russian]. Ent. Rev. Wash. 48 (4): 552-559. 

Danzig, E. M. 2004. Aleyrodes borchsenii Danzig, 1966 is a junior synonym of A. lonicerae Walker, 1852 

(Homoptera: Aleyrodidae). Zoological Institute. Pages? 

Darwish, Y. A, and A. I. Farghal. 1990. Evaluation of certain pesticides activity against the cotton whitefly, 

Bemisia tabaci, and associated natural enemies on cotton plants under field conditions in Assiut. Assiut J. Agri. 

Sci. 21:331-339. 

David, B. V. 1972. Two new species of Odontaleyrodes Takahashi (Homoptera: Aleyrodidae) from India. 

Oriental Insects. 6: 309-312. 

David, B. V. 1973. Description of a new genus Russellaleyrodes (Homoptera: Aleyrodidae) for Dialeuropora 

cumiugum (Singh, 1932) from Burma. Madras Agricultural Journal. 60: 557-558. 

David, B. V. 1974. A new host for the aleyrodid, Zaphanera publicus (Singh). Science & Culture. 40: 115. 

David, B. V. 1974a. Description of a new genus, Moundiella, for Trialeurodes megapapillae Singh, 1932 

(Homoptera: Aleyrodidae). Oriental Insects. 8: 43-45. 

530

David, B. V. 1976. A new species of the genus Aleuromarginatus Corbett (Aleyrodidae: Hemiptera) from India. 

Entomon. 1: 85-86. 

David, B. V. 1977. A new species of Aleurotuberculatus Takahashi and redescription of Aleurotuberculatus 

minutus (Singh) (Aleyrodidae: Hemiptera). Entomon. 2(1): 89-92. 

David, B. V. 1978. On a new species of Aleurotuberculatus (Hemiptera: Aleyrodidae) from India with a key to 

Indian species. Oriental Insects. 12(1): 133-135. 

David, B. V. 1981. A new species of Setaleyrodes (Homoptera: Aleyrodidae) from India. Colemania. 1(1): 37- 

38. 

David, B. V. 1988. Aleuromarginatus bauhiniae (Corbett) comb. nov. and A. thirumurthiensis nom. nov. 

(Aleyrodidae: Homoptera). Journal of the Bombay Natural History Society. 85(2): 445. 

David, B. V. 1990. Key to tribes of whiteflies (Aleyrodidae: Homoptera) of India. Journal of Insect Science. 3 

(1): 13-17. 

David, B. V. 1993. The whitefly of Sri Lanka (Homoptera: Aleyrodidae). Frederick Institute of Plant Protection 

& Technology, Entomology Series. 1993: 1-32, illustr. 

David, B. V. 1993a. Biodiversity in whiteflies (Aleyrodidae: Homoptera). Hexapoda. 5(2): 165-171. 

David, B. V. 2005. Cohicaleyrodes indicus (David and Selvakumaran) comb. n. and Cohicaleyrodes jesudasani 

nomen novum. Entomon. 30(1): 67-68. 

531

532 

David, B. V. 2005a. Editaaleyrodes indicus, a new genus and species of whitefly (Hemiptera: Aleyrodidae) 

from India. Entomon. 30(4): 317-320. 

David, B. V. and A. W. Augustine. 1988. A new whitefly Bemisia graminis sp. nov. (Aleyrodidae: Homoptera) 

from India. Entomon. 13: 33-35. 

David, P. M. M. and B. V. David. 2000. Occurrence of a new whitefly species of the Neotropical genus 

Crenidorsum Russell (Homoptera: Aleyrodidae). Entomon. 25 (2): 155-158. 

David, B. V. and R.W. A. Jesudasan. 1987. Description of a new genus Cresentaleyrodes for Aleurolobus 

semilunaris (Corbett) (Aleyrodidae: Homoptera). Current Science. 56 (1): 42-44. 

David, B.V. and R.W.A. Jesudasan 1987a. Redescription of the whitefly Aleyrodes shizuokensis Kuwana 

(Aleyrodidae: Homoptera). Journal of the Bombay Natural History Society. 86 (2): 260-261. 

David, B. V. and R.W. A. Jesudasan. 1988. On two new species of aleyrodids from both India and Sri Lanka. 

Entomon. 13(1): 29-32. 

David, B. V. and R.W. A. Jesudasan. 1989. Dialeurolonga maculata (Singh) comb. nov. and Dialeurolonga 

takahashii nom. nov. for Dialeurolonga maculata Takahashi (Aleyrodidae: Homoptera) from Madagascar. 

Entomon. 14 (3): 371. 

David, B. V. and R. W. A. Jesudasan. 2002. A new species of Aleurocanthus and Asialeyrodes indica 

Sundararaj & David (Aleyrodidae: Homoptera) from Andaman & Nicobar Islands. Entomon. 27 (3): 323-325 

David, B. V., R. W. A. Jesudasan & G. Mathew. 1988. Description of a new species of the genus Aleurolobus 

Quaintance & Baker (1914) (Aleyrodidae: Homoptera). Journal of the Bombay Natural History Society. 85 (1):

165-167. 

David, B. V., B. Krishnan & K. Thenmozhi. 1994. A new species of Viennotaleyrodes Cohic (Aleyrodidae: 

Homoptera) from India. Hexapoda. 6(1): 33-38. 

David, B. V. and M. Manjunatha. 2003. A new species of Aleurocanthus Quaintance & Baker (Homoptera: 

Aleyrodidae) from Areca catechu in India, with comments on the status of Aleurodes nubilans Buckton. 

Zootaxa. 173: 1-4. 

David, B. V. and E. Ragupathy. 2004. Whiteflies (Homoptera: Aleyrodidae) of mulberry, Morus alba, in India. 

Pestology. 28 (10): 24-33. 

David, B.V. and K. Regu. 1989. A new whitefly Aleurocanthus indicus sp. nov. (Aleyrodidae: Homoptera) from 

India. Entomon. 14 (3): 275-276. 

David, B.V. and K. Regu. 1991. A new record and description of Rhachiphora rutherfordi (Quaintance & 

Baker) (Homoptera: Aleyrodidae) from India. Journal of Insect Science 4(1): 69: 70. 

David, B. V. and K. Regu. 1995. Aleurodicus dispersus Russell (Aleyrodidae: Homoptera) a whitefly pest, new 

to India. Pestology. 19(3): 5-7. 

David, B. V. and S. Selvakumaran. 1987. A new species of whitefly Mixaleyrodes indicus sp. nov. from India. 

Journal of the Bombay Natural History Society. 84(3): 654-565. 

David, B. V. and S. Selvakumaran. 1990. On the occurrence of the whitefly Aleurodicus machilis Takahashi 

(Aleurodicinae, Aleyrodidae, Homoptera) in India. Entomon. 15(1-2): 139. 

533

David, B. V. and T. R. Subramaniam. 1976. Studies on some India Aleyrodidae. Records of the Zoological 

Survey of India. 70: 133-233. 

David, B. V. and R. Sundararaj. 1991. On a new species of Setaleyrodes Takahashi (Aleyrodidae: Homoptera) 

from India. Entomon. 16 (4): 317-318. 

David, B. V. and R. Sundararaj 1991a. A new species of the whitefly genus Pealius Quaintance and Baker 

(Aleyrodidae: Homoptera) from India. Journal of Insect Science. 4(1): 67-68. 

David, B. V. and R. Sundararaj. 1992. Dialeurodes delhiensis sp. nov. (Aleyrodidae: Homoptera) - a new 

species of whitefly from India. Journal of Insect Science. 5(1): 62-63. 

David, B. V. and R. Sundararaj. 1993. Studies on Dialeurodini (Aleyrodidae: Homoptera) of India: 

Kanakarajiella gen. nov. Journal of Entomological Research (New Dehli). 17 (4): 289-295. 

David, B. V, R. Sundararaj & K. Regu. 1991. On the four species of Odontaleyrodes Takahashi (Aleyrodidae: 

Homoptera) with a key to the Indian species. Journal of Insect Science. 4(2): 117-119. 

David, B. V. and K. Thenmozhi. 1995. On the characteristics of pupal case, adult and egg of Indian species of 

Lipaleyrodes Takahashi (Aleyrodidae: Homoptera) with description of a new species. Journal of the Bombay 

Natural History Society. 92(3), December 1995. 

David, P. M. 2000. Three new genera of whiteflies Mohanasundaramiella, Shanthiniae and Vasantharajiella 

(Aleyrodidae: Homoptera) from India. Journal of the Bombay Natural History Society. 97(1): 123-129. 

534

535 

De Barro, P. and M. Coombs. 2006. Release of Eretmocerus hayati in Australia. ESWN website. 

De Barro, P. J., J. W. H. Trueman and D. R. Frohlich. 2005. Bemisia argentifolli is a race of B. tabaci 

(Hemiptera: Aleyrodidae): the molecular genetic differentiation of B. tabaci populations around the world. 

Bulletin of Entomological Research. 95: 193-203. 

De Barro, P.J., F. Driver, I. Naumann, S. Schmidt, G. M. Clarke and J. Curran. 2000. Descriptions of three 

species of Eretmocerus Haldeman parasitizing Bemisia tabaci and Trialeurodes vaporariorum in Australia 

based on morphological and molecular data. Australian Journal of Entomology. 39:259-269. 

De Santis, L. 1937. Una nueva especie de Himenoptero del genero Amitus de la Republica Argentina. Revista 

Argentina de Agronomia. 4(2): 115-118. 

De Santis, L. 1948. Estudio monografico de los Afelinidos de La República Argentina (Hymenoptera, 

Chalcidoidea). Revista del Museo de La Plata (Nueva Serie) (Zoologia). 5:23-280. 

De Santis, L. 1948a. Adiciones a la fauna Argentina de Afelinidos (Hymenoptera: Chalcidoidea). Notas del 

Museo de La Plata, Buenos Aires. 13(101):43-48. 

De Santis, L. 1957. Adiciones a la fauna Argentina de Afelinidos. III. (Hymenoptera: Chalcidoidea). Notas del 

Museo de La Plata, Buenos Aires (Zoologia). 19(172):101-106. 

De Santis, L. 1964. Encirtidos de la Republica de Argentina (Hymenoptera: Chalcidoidea). An. Com. Inv. Cien. 

Prov. Buenos Aires Gober. 4: 9-422. 

De Santis, L. 1979. Additions to the Argentine aphelinid fauna VIII. (Insecta, Hymenoptera). (in Spanish). 

Revista de la Sociedad Entomologica Argentina. 38(1/4):133-136.

De Santis, L. 1979a. Catalogo de los himenopteros calcidoideos de america al sur de los estados unidos. 

Publicacien Especial Comisien de Investigaciones Cientificas Provincia de Buenos Airies488pp. -. 

De Santis, L. 1981. Sobre dos especes de Encarsia (Hymenoptera, Aphelinidae) del Brasil parasitoides de 

Bemisia tabaci (Hymenoptera, Aleyrodidae). Revista Brasileira de Entomologia. 25:37-39. 

DeBach, P. and M. Rose. 1981. A new genus and species of Aphelinidae with some synonymies, a rediagnosis 

of Aspidiotiphagus and a key to the pentamerous and heteromerous Prospaltellinae (Hymenoptera: 

Chalcidoidea: Aphelinidae). Proceedings of the Entomological Society of Washington. 83:658-679. 

Dean, D. E. 1994. Predaceous arthropods of the sweetpotato whitefly, Bemisia tabaci (Gennadius), on tomatoes 

in Florida. Ph.D. Dissertation, Univ. of Florida. -. 

Dietz, H. F, and J. Zetek. 1920. The black fly of citrus and other subtropical plants. USDA Bulletin 885. USDA, 

Washington D.C. -. 

Dobreanu, E. and C. Manolache. 1969. Homoptera: Aleyrodidoidea. Superfamilia Aleyrodinae. Fauna Repub. 

pop. rom. Insecta. 8 (5): 152 pp. 

Donia, A. R. A., E. M. Helal, M. M. Abd El Hamid, and E. A. Zakzouk. 1995. Mass rearing and field evaluation 

of released predaceous mite, Euseius scutalis (Athias-Henriot) on the citrus whitefly, Aleurotrachelus citri 

(Priesner and Hosny). Alexandria J. Agric. Res. 40:209-219. 

Dooley, J. W. 2006. Whitefly Pupae of the World. www.lucidcentral.org/keys/v3/whitefly/Homepage.htm. 

536

537 

Douglas, J. W. 1878. Note on the genus Aleurodes. Entomologist's Monthly Magazine. 14: 230-232. 

Douglas, J. W. 1884. Aleurodes immaculata Heeger. Entomologist's Monthly Magazine. 20: 215. 

Douglas, J. W. 1888. Description of new species of Aleurodes. Aleurodes ribium. Entomologist's Monthly 

Magazine. 24: 265-267. 

Douglas, J. W. 1891. A new species of Aleurodes?. Entomologist's Monthly Magazine. 27: 200. 

Douglas, J. W. 1891a. On a Brazilian species of Aleurodes found in England. Aleurodes filicium. 

Entomologist's Monthly Magazine. 27: 44. 

Douglas, J. W. 1892. Footnote to p. 32. A new genus and species of Aleurodidae. Entomologist's Monthly 

Magazine. 28: 29-33. 

Douglas, J. W. 1894. A new species of Aleurodes. Aleurodes spiraeae. Entomologist's Monthly Magazine. 30: 

732-74. 

Dowell, R. V, and R. H. Cherry. 1981. Survey traps for parasitoids, and coccinellid predators of the citrus 

blackfly, Aleurocanthus woglumi. Entomol. Exp. Appl. -. 29:356-362. 

Dozier, H. L. 1926. Some new Porto Rican scale parasites (Hymenoptera: Encyrtidae). Proceedings of the 


Dozier, H. L. 1926a. Annual report of the division of entomology. Annual Rept. Insular Expt. Sta. Rio Piedras. 

Insular Expt. Sta. Rio Piedras, San Juan. pp. 115-124.

Dozier, H. L. 1927. An undescribed whitefly attacking Citrus in Puerto Rico. Journal of Agricultural Research. 

36: 853-855. 

Dozier, H. L. 1928. Two new aleyrodid (Citrus) pests from India and the South Pacific. Journal of Agricultural 

Research. 36: 1001-1005. 

Dozier, H. L. 1932. The identity of certain whitefly parasites of the genus Eretmocerus Hald, with descriptions 

of new species (Hymenoptera: Aphelininae). Proceedings of the Entomological Society of Washington. 34: 112- 

118. 

Dozier, H. L. 1932a. Two undescribed chalcid parasites of the woolly whitefly Aleurothrixus floccosus 

(Maskell) from Haiti. Proceedings of the Entomological Society of Washington. 34: 118-122. 

Dozier, H. L. 1933. Miscellaneous notes and descriptions of chalcidoid parasites (Hymenoptera). Proceedings 

of the Entomological Society of Washington. 35(6):85-100. 

Dozier, H. L. 1934. Descriptions of new genera and species of African Aleyrodidae. Annals and Magazine of 

Natural History (10). 14: 184-192. 

Dozier, H. L. 1936. A revised annotated check-list of the insects of Puerto Rico. In Wolcott, G. N. Insectae 

Borinquenses. Journal of Agriculture of the University of Puerto Rico. 20: 143-146. 

Dozier, H. L. 1937. Descriptions of miscellaneous chalcidoid parasites from Puerto Rico (Hymenoptera). 

Journal of Agriculture of the University of Puerto Rico. 21:121-135. 

538

539 

Drews, E. A. and W. W. Sampson. 1956. Tetralicia and a new related genus Aleuropleurocelus (Homoptera: 

Aleyrodidae). Annals of the Entomological Society of America. 49: 280-283. 

Drews, E. A. and W. W. Sampson. 1958. California aleyrodids of the genus Aleuropleurocelus. Annals of the 

Entomological Society of America. 51: 120-125. 

Dubey, A. K and C. Ko. 2006. Toward an understanding of host plant associations of whiteflies (Hemiptera: 

Aleyrodidae): an evolutionary approach. Formosan Entomol. 26: 197-201. 

Dubey, A. K. and C. Ko. 2006a A new species of whitefly Crescentaleyrodes vetiveriae (Hemiptera: 

Aleyrodidae) from India. Formosan Entomol. 26: 69-75. 

Dubey, A. K. and C. C. Ko. 2007. Ruraleyrodes atalantiae, a new genus and species from India. Current 

Science. (in press).Dubey, A. K, K. Regu, and R. Sundararaj. 2004. Aleyrodidae (Hemiptera: Aleyrodidae) 

fauna of the Lakshadweep, India. Entomon. 29 (3): 279-286. 

Dubey, A. K. and R. Sundararaj. 2002. Development and survival of the spiralling whitefly, Aleurodicus 

dispersus Russell (Homptera: Aleyrodidae) on some tree species. Journal of Applied Zoological Research. 

13(2/3): 185-187. 

Dubey, A. K. and R. Sundararaj. 2003. New record of Aleurocanthus martini David, Homoptera: Aleyrodidae, 

from India. Journal of the Bombay Natural History Society. 102(1): 131. 

Dubey, A. K. and R. Sundararaj. 2004. A review of the genus Dialeuronomada Quaintance & Baker 

(Hemiptera: Aleyrodidae) with descriptions of two new species. Formosan Entomol. 24: 147-157. 

Dubey, A. K. and R. Sundararaj. 2004a. Host range of the spiralling whitefly Aleurodicus dispersus Russell

540 

(Aleyrodidae: Homoptera) in western ghats of south India. Indian Journal of Forestry. 27(1): 63-65. 

Dubey, A. K. and R. Sundararaj. 2005. A review of the genus Aleuroclava Singh (Hemiptera: Aleyrodidae) with 

descriptions of eight new species from India. Oriental Insects. 39: 241-272. 

Dubey, A. K. and R. Sundararaj. 2005a. A taxonomic study of the genus Pealius Quantance & Baker 

(Homoptera: Aleyrodidae) in India. Journal of the Bombay Natural History Society. 102(2): 162-165. 

Dubey, A. K. and R. Sundararaj. 2005b. Description of a new species of the genus Tetraleurodes Cockerell 

(Hemiptera: Aleyrodidae) with a key to the Indian species. Zoos' Print Journal. 20(7): 1924-1926. 

Dubey, A. K. and R. Sundararaj. 2005c. Three new species of Fippataleyrodes Sundararaj and David 

(Aleyrodidae: Hemiptera) from western gnats of south India. Journal of the Bombay Natural History Society. 

102(2): 204-207. 

Dubey, A. K. and R. Sundararaj. 2005d. Whitefly species of the genus Aleurocanthus Quaintance & Baker 

(Hemiptera: Aleyrodidae) from India, with descriptions of six new species. Oriental Insects. 39: 295-321. 

Dubey, A, K. and R. Sundararaj. 2006. On the genus Asialeyrodes Corbett (Hemiptera: Aleyrodidae) 

of India with a key to Indian species. Journal of the Bombay Natural History Society. 103: 117-119. 

Dubey, A, K. and R. Sundararaj. 2006. Distinctaleyrodes setosus Dubey & Sundararaj (Sternorrhyncha: 

Aleyrodidae), a new whitefly genus and species from India. Zootaxa. 1154: 35-39. 

Dubey, A. K. & R. Sundararaj. 2006a. A new whitefly species of the genus Taiwanaleyrodes Takahashi 

(Homoptera: Aleyrodidae) from western Ghats of South India. Entomon. 31(1): 73-76.

Dubey, A. K. & R. Sundararaj 2006b. Key to whiteflies of the tribe Aleurolobini (Hemiptera: Aleyrodidae) of 

India with description of five new species. Oriental Insects. 40: 33-60. 

Dubey, A. K. & R. Sundararaj 2006c. Description of five new species of whitefly genus Dialeurolonga 

(Hemiptera: Aleyrodidae) from India. Oriental Insects. 40: 159-170. 

Dubey, A. K. & R. Sundararaj 2006d. Icfrealeyrodes indica, a new genus and species of whitefly (Hemiptera: 

Aleyrodidae) from India. Entomon 32(2): 125-128. 

Dubey, A. K. & R. Sundararaj 2006e. Two new aleyrodids (Hemiptera: Aleyrodidae) from India. Entomon 

31(3): 229-235. 

Duda, O. 1935. Drosophilidae, In Linder,E. [ed.], Die Fliegen der Plparktischen Region. Schweizerbartsche 

Verlagsbuchhandlung G.H.B.H. Stuttgart. pp. 58. 

Dumbleton, L. J. 1956. New Aleyrodidae (Hemiptera: Homoptera) from New Caledonia. Proceedings of the 

Royal Entomological Society of London (B). 25: 129-141. 

Dumbleton, L. J. 1956a. The Australian Aleyrodidae (Hemiptera: Homoptera). Proceedings of the Linnean 

Society of New South Wales. 81: 159-183. 

Dumbleton, L. J. 1957. New Zealand Aleyrodidae (Hemiptera: Homoptera). Pacif. Sci. 11: 141-160. 

Dumbleton, L. J. 1961. Aleyrodidae (Hemiptera: Homoptera) from the South Pacific. New Zealand Journal of 

Science. 4: 770-774. 

541

542 

Dumbleton, L. J. 1961a. The Aleyrodidae (Hemiptera: Homoptera) of New Caledonia. Pacif. Sci. 15: 114-136. 

Dumbleton, L. J. 1990. The Aleyrodidae (Hemiptera: Homoptera) of New Caledonia. Pacif. Sci. 15: 114-136. 

Dysart, R. J. 1966. Natural enemies of the banded-wing whitefly, Trialeurodes abutilonea (Hemiptera: 

Aleyrodidae). Annals of the Entomological Society of America. 59: 28-33. 

Dysart, R. J. 1966. Natural enemies of the bandedwing whitefly Trialeurodes abutilonea (Hemiptera: 

Aleyrodidae). Entomol. Soc. Amer. Misc. Pub. -. 59:28-33. 

Ehler, L. E, and R. van den Bosch. 1974. An analysis of the natural biological control of Trichoplusia ni 

(Lepidoptera: Noctuidae) on cotton in California. Can. Entomol. 106:1067-1073. 

Ekbom, B. S. 1981. Efficiency of the predatory Anthocoris nemorum (Het.: Anthocoridae) against the 

greenhouse whitefly, Trialeurodes vaporariorum (Hom.: Aleyrodidae). J. Appl. Entomol. 92:26-34. 

El Badry, E. A. 1967. Three new species of phytoseiid mites preying on the cotton whitefly, Bemisia tabaci in 

the Sudan (Acarina: Phytoseiidae). The Entomologist. 100:106-111. 

El Badry, E. A. 1968. Biological studies on Amblyseius aleyrodis a predator of cotton whitefly (Acarina, 

Phytoseiidae). Entomophaga. 13:323-329. 

El Helaly, M. S, A. Y. El Shazli, and F. H. El Gayar. 1971. Biological studies on Bemisia tabaci, Gennadius 

(Homoptera: Aleyrodidae) in Egypt. Z. Angewandte Entomol. 69:48-55. 

El Khidir, E. and A. Khalifa. 1962. A new aleyrodid from the Sudan. Proceedings of the Royal Entomological

Society of London (B). 31: 47-51. 

El Sayed, A. M. 1981. Ecological and control studies on the whitefly, Bemisia tabaci (Genn.) attacking tomato 

plants. M.Sc. Thesis, Faculty of Agriculture, Menoufia Univ. 

El-Banhawy, E. M, and M. E. El-Bagoury. 1991. Biological studies of the predacious mite Typhlodromus 

pelargonicus a predator of the two spotted spider mite Tetranychus urticae on cucumber plants (Acari: 

Phytoseiidae: Tetranychidae). Entomophaga. 36:587-591. 

Elizondo, J. M, and J. R. Quezada. 1990. Identification y evaluacion do los enemigos naturales de la mosca 

prieta citricolas de Costa Rica. Turrialba. 40:190-197. 

Enderlein, G. 1909. Aleurodicus conspurcatus, eine neue Aleurodide aus Sud-Brasilien. Stettin. Ent. Zig. 70: 

282-284. 

Erdos. 1961. Annales Historico-Naturales Musei Nationalis Hungarici. 53: 482. 

Erdos, J. 1966. Nonnullae Eulophidae novae Hungaricae (Hymenoptera: Chalcidoidea). Annales Historico- 

Naturales Musei Nationalis Hungarici. 58: 395-420. 

Escobar, J. C. 1983. Dinamica de poblacion y control natural de Bemisia tabaci Genn. en el cultivo del algodon. 

Departamento de Proteccion Vegetal, Facultad de Ciencias Agronomicas, Universidad de El Salvador, San 

Salvador. 

Evans, G. A and A. Polaszek. 1997. Additions to the Encarsia parasitoids (Hymenoptera: Aphelinidae) of the 

Bemisia tabaci complex (Hemiptera: Aleyrodidae). Bulletin of Entomological Research. 87:563-571. 

543

Evans, G. A. and A. Polaszek. 1998. The Encarsia cubensis species-group (Hymenoptera: Aphelinidae). 

Proceedings of the Entomological Society of Washington. 100(2): 222-233. 

Evans, G. A. and C. Serra. 2002. Whitefly parasitoids of Hispaniola. Journal of Hymenoptera Research. 11(2): 

197-212. 

Evans, G. A. and S. Abd-Rabou. 2005. Two new species, and additional records, of aphelinids from Egypt 

(Hymenoptera: Aphelinidae). Zootaxa. 838: 1-7. 

Evans, G. and J. A. Castillo. 1998. Parasites of Aleurotrachelus socialis (Homoptera: Aleyrodidae) from 

Colombia including descriptions of two new species (Hymenoptera: Aphelinidae: Platygasteridae). Florida 


Evans, G.A. 1997. A new Encarsia (Hymenoptera: Aphelinidae) species reared from the Bemisia tabaci 

complex (Homoptera: Aleyrodidae). Florida Entomologist. 80(1):24-27. 

Evans, G.A. and C. L. Angulo. 1996. A new species of Encarsia (Hymenoptera: Aphelinidae) from Costa Rica. 

(in English with Spanish summary). Florida Entomologist. 79(4):582-586. 

Evans, G.A. and F. D. Bennett. 1996. A new Eretmocerus (Hymenoptera: Aphelinidae) species reared from 

Dialeurodes kirkaldyi (Homoptera: Aleyrodidae). (in English with Spanish summary). Florida Entomologist. 

79(4):579-582. 

Evans, G.A. and P. A. Pedata. 1997. Parasitoids of Comstockiella sabalis (Homoptera: Diaspididae) in Florida 

and description of a new species of the genus Coccobius (Hymenoptera: Aphelinidae). Florida Entomologist. 

80(3):328-334. 

544

Fabricius. 1798. Suplementum entomologisicae systematicae. Hafniae. 

Fabricius, J. C. 1798. Suplementum entomologisicae systematicae. Hafniae. 

Felt, E. P. 1914. List of zoophagous Itonididae. J. Econ. Entomol. 7:458-459. 

Felt, E. P. 1922. A new cecidomyiid parasite of the white fly. Proc. U.S. Nat. Mus. 61:1-2. 

Ferragut, F, F. Garcia-Mari, J. Costa-Comelles, and R. Laborda. 1987. Influence of food and temperature on 

development and oviposition of Euseius stipulatus and Typhlodromus phialatus (Acari: Phytoseiidae). Exper. 

Appl. Acarol. 3:317-330. 

Ferriere, Ch. 1965. Hymenoptera Aphelinidae d'Europe et du Bassin Mediterraneen. Faune de l'Europe et du 

Bassin Mediterraneen, Paris. 206 pp. 

Fishpool, L. D. C, and C. Burban. 1994. Bemisia tabaci: the whitefly vector of African cassava mosaic 

geminivirus. Trop. Sci. 34:55-72. 

Fitch, A. 1857. Third report on the noxious, beneficial and other insects of the state of New York. Transactions 

of the New York State Agriculture Society. 16: 332. 

Fitzpatrick, G. E, R. H. Cherry, and R. V. Dowell. 1978. Short-term effects of three insecticides on predators 

and parasites of the citrus blackfly. Environ. Entomol. 7:553-555. 

Fitzpatrick, G. E, R. H. Cherry, and R. V. Dowell. 1979. Effects of Florida citrus control practices on the citrus 

545

546 

blackfly (Homoptera: Aleyrodidae) and its associated natural enemies. Can. Entomol. 111:731-734. 

Foerster, A. 1878. Kleine Monographien parasitischer Hymenopteren. Verhandlungennaturhistorischen 

Vereines der preussischen Rheinlande und Westfalens. 35: 43-82. 

Forbes. S.A. 1884. Rep. Ill. St. Ent. 13: (footnote), 205. Rep. Ill. St. Ent. 205 p. 

Forbes. S.A. 1885. Aleurodes aceris n. s. Order Hemiptera, Family Aleurodidae. Rep. Ill. St. Ent. 14: 110. 

Fouts, R. M. 1924. Revision of North American wasps of the subfamily Platygasterinae. Proceedings of the 

United States National Museum. 63(15): 8-10. 

Fowler, V. W. 1954. Notes on some pests observed in the course of advisory work at Wisley during 1953. 

Journal of the Royal Horticultural Society. 79: 405-408. 

Franklin, H. J. 1908. On a collection of thysanopterous insects from Barbados and St. Vincent Islands. Proc. 

U.S. Nat. Mus. 33:727-729. 

Franklin, H. J. 1909. On Thysanoptera. Entomological News. 20:228-231. 

Fransen, J. J. 1994. Bemisia tabaci in the Netherlands - Here to Stay. Pest. Sci. 42:129-134. 

Frappa, Cl. 1938. Description de Bemisia manihotis, n. sp. (Hem:Hom. Aleyrodidae) nuisible au manioc a 

Madagascar. Bulletin de la Societe Entomologie de France. 43: 30-32. 

Frappa, Cl. 1939. Note sur une nouvelle espece d'aleurode muisible aux plantations de tabac de la Tsiribihina. 

Bull. econ. trimest. Madagascar. 16: 254-259.

Frauenfeld, G. R. 1867. Zoologiche Miscellen XIII. Ueber Aleurodes und Thrips, vorzuglich im Warmhouse. 

Verhandlungen der Zoologische-Botanishcen Gesellschaft in Wien. 17: 793-799. 

Froggatt, W. W. 1911. A new pest of salt-bush whitefly (Aleurodicus atriplex n.sp.). Agric. Gaz. N. S. W. 22: 

757-758. 

Froggatt, W. W. 1918. Notes on 'Snow-flies', with the description of a new species (Aleurodicus albofloccosa). 

Agric. Gaz. N. S. W. 29: 434-436. 

Fulmek, L. 1943. Wirtsindex der Aleyrodiden-und Cocciden-parasiten. Ent. Beih. Berl.-Dahlem. 10:1-100. 

Fulmek, L. 1959. Die Feindinsekten der Mottenschildlpuse (Parasiten und Rpuber der Aleyrodidae-Homoptera). 

Pismo Entomol. 29:107-130. 

Gahan, A. B. 1924. Some new parasitic Hymenoptera with notes on several described forms. Proceedings of the 

United States National Museum. 65: 1-23. 

Gahan, A. B. 1927. Miscellaneous descriptions of new parasitic Hymenoptera with some synonymical notes. 

Proceedings of the United States National Museum. 71: 1-39. 

Gahan, A. B. 1931. A new species of Encarsia from Cuba (Hymenoptera: Aphelininae). Proceedings of the 

Entomological Society of Washington. 33:121-122. 

Gahan, A. B. 1932. Miscellaneous descriptions and notes on parasitic Hymenoptera. Annals of the 

Entomological Society of America. 25(4):736-757. 

547

Ghahari, H. J. Huang and Z. Wang. 2005. Biology of a thelytokous biotype of Eretmocerus mundus 

(Hymenoptera: Aphelinidae) on Bemisia tabaci (Homoptera: Aleyrodidae). Insect Science 12: 461-465. 

Gameel, O. I. 1968. Three new species of Aleyrodidae from the Sudan (Hemiptera: Homoptera). Proceedings of 

the Royal Entomological Society of London (B). 37: 149-155. 

Gameel, O. I. 1969. Studies on whitefly parasites Encarsia lutea Masi and Eretmocerus mundus Mercet 

(Hymenoptera: Aphelinidae). Revue de Zoologie et de Botanique Africaines. 79: 65-77. 

Gameel, O. I. 1971. New Aleyrodidae (Hemiptera: Homoptera). Revue de Zoologie et de Botanique Africaines. 

84: 169-174. 

Gameel, O. I. 1971a. The whitefly eggs and first larval stages as prey for certain Phytoseiid mites. Rev. Zoo. 

Bot. Africa. 84:79-82. 

Gargani, E. 1990. Osservazioni su Dialeurodes citri (Ashmead) (Homoptera: Aleyrodidae) e soui antagonisti in 

Toscana. Redia. 73:7-16. 

Gautier, C. 1922. Un Aleurode parasite du Poirier et du Frene, Trialeurodes inaequalis n. sp. (Hemiptera, 

Aleurodidae). Ann. Soc. Ent. Fr. 91:337-350. 

Gautier, C. 1923. Un aleurodes parasitre du poirier et du trene Trialeurodes inaequalis n. sp. (Hem. 

Aleurodidae). Annls. Soc. Ent. Fr. 91: 337-350. 

Gennadius, P. 1889. Disease of tobacco plantations in the Trikonia. The aleurodid of tobacco [In Greek]. 

548

Ellenike Georgia. 5: 1-3. 

Geoffroy, E. L. 1762. Histoire abregee des insectes des insectes qui se trouvent aux environs de Paris. 

Entomologia Parisiensis. 544 pp. 

Geoffroy, E. L. 1795. in Fourcroy, A. L. Entomologia Parisiensis. 544 pp. 

Gerling, D. 1966. Studies with whitefly parasites of southern California. II. Eretmocerus californicus Howard 

(Hymenoptera: Aphelinidae). Canadian Entomologist. 98(12):1316-1329. 

Gerling, D. 1970. Comments on Eretmocerus serius Silvestri (Hymenoptera: Aphelinidae) with a description of 

two new species. Bollettino del Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 27: 79-88. 

Gerling, D. 1970. Two African species of Eretmocerus Haldeman (Hymenoptera: Aphelinidae). J. Ent. Soc. sth. 

Afr. 33: 325-329. 

Gerling, D. 1972. Notes on three species of Eretmocerus Haldeman occurring in Israel with a description of a 

new species. Entomologische Berichten, Amsterdam. 32(8):156-161. 

Gerling, D. 1985. Parasitoids attacking Bemisia tabaci (Hom.: Aleyrodidae) in Eastern Africa. Entomophaga. 

30: 163-165. 

Gerling, D. 1986. Natural enemies of Bemisia tabaci, biological characteristics and potential as biological 

control agents: a review. Agriculture, Ecosystems & Environment. 17: 99-110. 

Gerling, D. 1990. Natural enemies of whiteflies: predators and parasites. In D. Gerling [ed.], Whiteflies: their 

549

ionomics, pest status, and management. Intercept, Wimborne, UK. pp. 147-186. 

Gerling, D. 1996. Status of Bemisia tabaci in the Mediterranean countries: Opportunities for biological control. 

Biol. Control. 6:11-22. 

Gerling, D, V. Kravchenko, and M. Guershon. 1997. Deraeocoris pallens Reuter, Orius niger Wolf and 

Campylomma unicolor Poppius (Heteroptera) as predators of Bemisia nymphs. In T. J. Henneyberry [ed.], 

Silverleaf whitefly: 1997 supplement to the five-year national research and action plan: progress review, 

technology transfer, and new research and action plan, USDA/ARS 1997-02. pp. 140. 

Gerling, D, V. Kravchenko, and M. Lazare. 1997. Dynamics of common green lacewing (Neuroptera: 

Chrysopidae) in Israeli cotton fields in relation to whitefly (Homoptera: Aleyrodidae) populations. Environ. 

Entomol. 26:815-827. 

Gerling, D, S. Kyamanywa & J. Legg. 2006. Some new whitefly pests in Africa. Second European Whitefly 

Symposium, Croatia.. 

Gerling, D. and T. Rivnay. 1984. A new species of Encarsia (Hym.: Aphelinidae) parasitizing Bemisia tabaci 

(Hom.: Aleyrodidae). Entomophaga. 29: 439-444. 

Gerling, D, and T. U. Stern. 1993. Biology of Delpahstus pusillus: the influence of density on fitness. Bull. 

OILB/SROP. 16:39-42. 

Gill, R.J. 1990. The morphology of whiteflies. Whiteflies: Their Bionomics, Pest Status, and Management, ed. 

by D. Gerling, Intercept Ltd, Andover. p. 13-46. 

Gill, R.J. 1996. Jasmine Whitefly. California Plant Pest & Disease Report. 15: 156-157. 

550

Gillespie, D. R. 1985. Endemic Aleyrodidae (Homoptera) and their parasites (Hymenoptera) on southern 

Vancouver Island, British Colombia. Journal of the Entomological Society of British Colombia 82: 12-13. 

Gillespie, P.S. 2006. A new genus and two new species of whiteflies from Australia (Hemiptera: Aleyrodidae). 

Zootaxa. 1306: 1-23. 

Gillette, C. P. and C. F. Baker. 1895. A preliminary list of the Hemiptera of Colorado. 

Ginting, C. U, A. Diamin, and L. Sidauruk. 1992. Some biological aspects of Megalocaria dilatata F. (Col.: 

Coccinellidae) a potential coconut leaf aphid predator. Oleagineux. 47:569-572. 

Girault, A. A. 1908. Encarsia versicolor species novum, and eulophid parasite of the greenhouse whitefly, 

Aleyrodes vaporariorum Westwood. Psyche. 15: 53-57. 

Girault, A. A. 1910. Synonymic and descriptive notes on the chalcidoid family Mymaridae. Journal of the New 

York Entomological Society. 18: 233-259. 

Girault, A.A. 1911. The chalcidoid parasites of the coccid Kermes pubescens Bogue, with description of two 

new genera and three new species of Encyrtidae from Illinois. Canadian Entomologist. 43: 168-178. 

Girault, A. A. 1913. Australian Hymenoptera Chalcidoidea - IV. The family Eulophidae with descriptions of 

new genera and species. Memoirs of the Queensland Museum. 2: 140-296. 

Girault, A. A. 1914. Descriptions of new chalcid-flies. Proceedings of the Entomological Society of 

Washington. 16: 109-119. 

551

Girault, A. A. 1915. Australian Hymenoptera Chalcidoidea - VII. The family Encyrtidae with descriptions of 

new genera and species. Memoirs of the Queensland Museum. 4: 1-184. 

Girault, A. A. 1915a. Some new chalcidoid Hymenoptera form North and South America. Annals of the 

Entomological Society of America. 8: 272-278. 

Girault, A. A. 1916. A new genus of omphaline eulaphid [sic] chalcis-flies from Maryland. Canadian 

Entomologist. 48: 410. 

Girault, A. A. 1917. Some new Australian chalcid-flies, mostly of the family Encyrtidae (Hymenoptera). 

Insecutor Inscitiae Menstruus. 5: 29-37. 

Girault, A. A. 1920. New syrphoid, cynipoid and chalcidoid Hymenoptera. Proc. U.S. National Museum. 58: 

177-216. 

Girault, A. A. 1921. Miscellaneous species of Chalcid-flies from Australia. Inscitiae Menstruus. 9: 186-191. 

Girault, A. A. 1930. New Pests from Australia IX. 2 pp. 

Girault, A. A. 1932. New lower Hymenoptera from Australia and India (Privately publ, 6 pp. ). in: Gordh et al. 

(1979). Memoirs of the American Entomological Institute. 28: 293-298. 

Girault, A. A. 1932a. New pests from Australia. Brisbane, private publication. 10:5. 

Girault, A. A. and A. P. Dodd. 1915. The cane grubs of Australia. Bulletin Bur. Sug. Exp. Stations, Queensland 

Division of Entomology. 2: 1-60. 

552

Goeldi, E. A. 1886. Beitrage zur Kenntnis der kleinen und kleinsen Gliederthierwelt Brasiliens. II Neue 

brasilianische Aleurodes-Arten. Mitteilungen der Schweizerischen Entomologischen Gesellschaft. 7: 241-250. 

Goeze, J. A. E. 1777. Entomologische Beitrage zu des Reitter Linne. Ausgabe der Natursystems. Leipizig. 1: 

736 pp. 

Gold, C. S, and M. A. Altieri. 1989. The effects of intercropping and mixed varieties of predators and 

parasitoids of cassava whiteflies (Hemiptera: Aleyrodidae) in Columbia. Bull. Entomol. Res. 79:115-121. 

Gold, C. S, M. A. Altieri, and A. C. Bellotti. 1989. Cassava intercropping and pest incidence: a review 

illustrated with a case study from Colombia. Trop. Pest Mgt. 35:339-344. 

Gold, C. S, M. A. Altieri, and A. C. Bellotti. 1991. Survivorship of the cassava whiteflies Aleurotrachelus 

socialis and Trialeurodes variabilis (Homoptera: Aleyrodidae) under different cropping systems in Colombia. 

Crop Protection. 10:305-309. 

Golfari, L. 1937. Contributi alla conoscenza dell'entomofauna del Pero (Pirus communis L.). Boll. Istitu. 

Entomol. Univers. Studi Bologna. 9:206-249. 

Gomez, J. and O. Garcia. 2000. A new species of Encarsia (Hymenoptera: Aphelinidae), a parasitoid of 

whitefly Aleurodicus sp. (Homoptera: Aleyrodidae) in Mexico. Pan pacific Entomologist. 76(1), January, 2000: 

49 51, illustr."Notes on some species of Encarsia (Hymenoptera: Aphelinidae). 

Gomez-Menor, J. 1943. Contribucion al conocimiento de los aleyrodidos de Espana (Hem. Homoptera), 1a 

nota. Eos. Revista Espanola di Entomologia. Madrid. 19: 173-209. 

Gomez-Menor, J. 1945. Contribucion al conocimiento de los aleyrodidos de Espana (Hem. Homoptera) 

553

554 

Variabilidad en las especies espanolos y descripcion de dos nuevas, 2a nota. Eos. Revista Espanola di 

Entomologia. Madrid. 20: 277-308. 

Gomez-Menor, J. 1945a. Aleurodidos de interes agricola. Boln. Patol. veg. Ent. agric. 13: 161-198. 

Gomez-Menor, J. 1953. Algunos insectos como pequenos enemigos: los aleurodidos. Boln. Patol. veg. Ent. 

agric. 2: 27-55. 

Gomez-Menor, J. 1954. Aleurodidos de Espana, islas Canarias y Africa occidental. Eos. Revista Espanola di 

Entomologia. Madrid. 30: 363-377. 

Gonzalez-Olazo, E. V. 1987. Los neuropteros asociados con los cultivos citricos de la provincia de Tucuman y 

descripcion de una nueva especie de Nomerobius (Hemerobiidae). CIRPON, Rev. Invest. 5:37-54. 

Gordon, R. D. 1972. A review of the genus Nephaspis Casey and a comparison with the genus Clitostethus 

Weise (Coleoptera: Coccinellidae). Revista Agric. Piracicaba. 47:145-154. 

Gordon, R. D. 1982. Two species of Nephaspis Casey (Coleoptera: Coccinellidae) from Trinidad and Colombia. 

Proc. Entomol. Soc. Wash. 84:332-336. 

Gordon, R. D. 1985. The Coccinellidae (Coleoptera) of America north of Mexico. J. New York Entomol. Soc. 

93:1-912. 

Goula, M, and O. Alomar. 1994. Miridos (Heteroptera: Miridae) de interes en el control integrado de plagas en 

el tomate, guia para su identification. Boletin de Sanidad Vegetal, Plagas. 20:131-143.

555 

Goux, L. 1939. Contribution a l'etude des aleurodes (Hem. Aleyrodidae) de la France. I. Description d'un sous- 

genre et de deux especes nouveaux. Bulletin de la Societe Linneenne de Provence. 12(1938): 77-82. 

Goux, L. 1940. Contribution a l'etude des aleurodes (Hem. Aleyrodidae) de la France. I. Description de deux 

especes nouvelles de Marseille. Bulletin de la Societe Entomologie de France. 45: 45-48. 

Goux, L. 1942. Contribution a l'etude des aleurodes (Hem. Aleyrodidae) de la France. III. Description d'un 

Aleurolobus et d'un Aleurotrachelus nouveaux. Bulletin du Museum d'Histoire Naturelle de Marseille. 2: 141- 

148. 

Goux, L. 1945. Contribution a l'etude des aleurodes (Hem. Aleyrodidae) de la France. IV. L'etude 

morphologique et biologique d'une espece nouvelle constituant un genre nouveau. Bulletin du Museum 

d'Histoire Naturelle de Marseille. 5: 186-197. 

Goux, L. 1948. Contribution a l'etude des aleurodes (Hem. Aleyrodidae) de la France. V. L'aleurode du laurier 

sauce (Laurus nobilis L.). Ann. Soc. Sc. Nat. Toulon. 2: 30-34. 

Goux, L. 1949. Contribution a l'etude des aleurodes (Hem. Aleyrodidae) de la France. VI. Le genre Siphoninus 

Silvestri. Bull mens. Soc. Linn. Lyon N. S. 18: 7-12. 

Goux, L. 1987. Aleurodes de France - VII. Description de deux especes nouvelles constituant des genres 

nouveaux. Bulletin de la Societe Linneenne de Provence. 39: 63-66. 

Goux, L. 1988. Aleurodes de France - VII. Description de deux especes nouvelles constituant des genres 

nouveaux. Bulletin de la Societe Linneenne de Provence. 39: 63-66. 

Gowdey 1928. Report of government entomologist. Ann. Rev. Entomol. 16:527.

Gowdey, C. C. 1921. The citrus Black Fly (Aleurocanthus woglumi, Ashly). Entomol. Circ. 3. Jamaica Dept. 

Agric, Kingston. 3. 

Gowdey, C. C. 1923. The white flies (Aleurodidae) of Jamaica. Jamaica Dep. Agr. Entomol. Bull. 3:1-6. 

Gowdey, C. C. 1925. Report of government entomologist. Ann. Rev. Entomol. 14:147-148. 

Gowdey, C. C. 1928. Report of government entomologist. Ann. Rev. Entomol. 16:527. 

Graham, M.V.R. 1976. The British species of Aphelinus with notes and descriptions of other European 

Aphelinidae (Hymenotpera). Systematic Entomology. 1: 123-146. 

Graham, M.W.R. de V. 1986. Madeira insects (chiefly Hymenoptera) including additions to the list and 

descriptions of three new species of Chalcidoidea. Bulletin do Museu Municipal do Funchal. 38: 28-42. 

Grandi, G. 1914. Descrizione di un nuovo Coccinellidae africano. Boll. Lab. Zool. Agrar. 1914:165-178. 

Gravena, S, M. G. C. Churata-Masca, J. Arai, and A. Raga. 1984. Manejo integrado da mosca blanca Bemisia 

tabaci (Gennadius, 1889) em cultivares de tomateiro de crecimento determinado visando reducao de virose do 

mosaico dourado. Anais Soc. Entomol. Brasil. 13:35-45. 

Griffiths, J. T, and W. L. Thompson. 1957. Insects and mites found on Florida citrus. Florida Agr. Exp Sta. 

Bull. 591:33-39. 

Grissell, E. E. 1979. The Prospaltella of Florida (Hymenoptera: Aphelinidae). Florida Department 

556

557 

ofAgriculture, Conservation Service Division of Plant Industry Entomology Circular. 203: 4 pp. 

Guerrieri, E. and J. S. Noyes. 2000. Revision of European species of genus Metaphycus Mercet (Hymenoptera: 

Chalcidoidea: Encyrtidae), parasitoids of scale insects (Homoptera: Coccoidea). Systematic Entomology. 25: 

147-222. 

Habib, A. and F. A. Farag. 1970. Studies on nine common aleurodids of Egypt. Bulletin de la Societe 

Entomologique d'Egypte. 54: 1-41. 

Hafez, M, M. F. S. Tawfik, K. T. Awadallah, and A. A. Sarhan. 1979. Natural enemies of the cotton whitefly, 

Bemisia tabaci (Genn.), in the world and in Egypt. Bull. Soc. Ent. Egypte. 62:9-13. 

Hagler, J. R, and S. E. Naranjo. 1996. Evaluating the feeding behavior of commercial and indigenous 

Hippodamia convergens on sweet potato whitefly: A laboratory and field study. In T. J. Henneberry, N. C. 

Toscano, R. M. Faust, and J. R. Coppedge [eds.], Silverleaf whitefly: 1996 supplement to the 5-year national 

research and action plan, USDA/ARS 1996-01. pp. 115. 

Hagler, J. R. and S. E. Naranjo. 1994. Determining the frequency of heteropteran predation on sweetpotato 

whitefly and pink bollworm using multiple ELISAs. Entomol. Exp. Appl. 72:59-66. -. 

Hagler, J. R. and S. E. Naranjo. 1994a. Qualitative survey of two coleopteran predators of Bemisia tabaci 

(Homoptera: Aleyrodidae) and Pectinophora gossypiella (Lepidoptera: Gelechiidae) using a multiple prey gut 

content ELISA. Environ. Entomol. 23:193-197. 

Haldeman, S. S. 1850. On four new species of Hemiptera of the genera Ploiaria, Chermes and Aleurodes and 

two new Hymenoptera, parasitic in the last named genus. American Journal of Science. (2)9: 108-111.

Haliday, A. H. 1835. Aleyrodes phillyreae. Entomological Magazine. 2: 119-120. 

Hamon, A. B. 1989. Metaleurodicus cardini in Florida (Homoptera: Aleyrodidae: Aleurodicinae). Entomology 

Circular ed, Florida Department of Agriculture, Gainesville. 316; 2 pp. 

Hansson, C. and J. LaSalle. 2003. Revision of the Neotropical species of the tribe Euderomphalini 

(Hymenoptera: Eulophidae). Journal of Natural History. 37: 697-778. 

Harrison, J. W. H. 1917. A new species and genus of Aleurodidae from Durham. Vasculum. 3: 60-62. 

Harrison, J. W. H. 1917a. New and rare British Aleurodidae. Entomologist. 53: 255-257. 

Harrison, J. W. H. 1920. Notes and records. Aleyrodidae. Vasculum. 6: 59. 

Haupt, H. 1934. Neues uber die Homoptera-Aleurodina. Deutsche Entomologische Zeitschrift. 1934: 127-141. 

Haupt, H. 1935. Schmetterlings -od. Mottenlause, Aleurodina. Dt. ent. Z. 1934: 127-141. 

Hayat, M. 1972. The species of Eretmocerus Haldeman, 1850 (Hymenoptera: Aphelinidae) from India. 

Entomophaga. 17: 99-106. 

Hayat, M. 1974. On some Indian species of Aphelinidae, with a description of a new Coccophagus 

(Hymenoptera; Chalcidoidea). Journal of Natural History. 8:179-186. 

Hayat, M. 1976. Two new species of Aphelinidae parasitic on Acaudaleyrodes rhachipora from India. 

Entomophaga. 21(2):157-162. 

558

Hayat, M. 1979. Notes on some Indian species of Azotus Howard and Coccophagoides Girault, with records of 

Mesidia Foerster and Prococcophagus Silvestri (Hym.: Aphelinidae). Journal of Natural History. 13:185-193. 

Hayat, M. 1981a. Taxonomic notes on some Oriental Aphelinidae with some new records (Hym.: 

Chalcidoidea). Oriental Insects. 14: 461-472. 

Hayat, M. 1983. The genera of Aphelinidae (Hymenoptera) of the World. Systematic Entomology. 8:63-102. 

Hayat, M. 1984. Encarsia bennetti sp. n. (Hymenoptera: Aphelinidae) from India, associated with the citrus 

black fly, Aleurocanthus woglumi (Homoptera: Aleyrodid. Bulletin of Entomological Research. 74: 399-401. 

Hayat, M. 1986. Family Aphelinidae. (In: Subba Rao, B.R.; Hayat, M. (Eds) - The Chalcidoidea (Insecta: 

Hymenoptera) of India and the adjacent countries. Part II, pp. 143-171). Oriental Insects. 20:143-171,. 

Hayat, M. 1989a. A revision of the species of Encarsia Foerster (Hymenoptera: Aphelinidae) from India and the 

adjacent countries. Oriental Insects. 23: 1-131. 

Hayat, M. 1989b. Descriptions of two new species of Encarsia (Hymenoptera: Aphelinidae) from Saudi Arabia 

and the Solomon Islands. Oriental Insects. 23:163-167. 

Hayat, M. 1989c. Notes on some species of Coccophagoides, Dirphys and Encarsia (Hymenoptera: 

Aphelinidae). Oriental Insects. 23:286-290. 

Hayat, M. 1994. Notes on some genera of the Aphelinidae (Hymenoptera: Chalcidoidea), with comments on the 

classification of the family. Oriental Insects. 28:81-96. 

559

560 

Hayat, M. 1998. Aphelinidae of India (Hymenoptera: Chalcidoidea): A taxonomic revision. Memoirs on 

Entomology, International. 13, 416 pp. 

Hayat, M. and A. Polaszek. 1992. Encarsia aseta new species (Hymenoptera: Aphelinidae) a parasitoid of 

Dialeurolonga elongata Dozier (Homoptera; Aleyrodidae) from India. Oriental Insects. 26:107-109. 

Hayat, M. and S. B. Zeya. 1992. Description of a new species of Pomphale (Hymenoptera: Eulophidae). 

Hexapoda. 4(2):183-186. 

Hayat, M. and S. B. Zeya. 1993. Records and descriptions of some Indian Aphelinidae (Hymenoptera: 

Chalcidoidea). Hexapoda. 5(1):57-66. 

Heeger, E. 1856. Beitrage zur Naturgeschichte der Insecten. Naturgeschichte der Aleurodes immaculata Steph. 

Sitzungberichte der Kaiserlichen Akademie der Wissenschaften. Mathematische-naturwissenschaftliche klasse. 

Wien. 18: 33-36. 

Heeger, E. 1859. Beitrage zur Naturgeschichte der Insecten. Naturgeschichte der Aleyrodes dubia Stephens. 

Sitzungberichte der Kaiserlichen Akademie der Wissenschaften. Mathematische-naturwissenschaftliche klasse. 

Wien. 24: 223-226. 

Heinz, K. M, J. R. Brazzle, C. H. Pickett, E. T. Natwick, J. M. Nelson, and M. P. Parrella. 1994. Predatory 

beetle may suppress silverleaf whitefly. Calif. Agri. 48:35-40. 

Heinz, K. M, and J. M. Nelson. 1996. Interspecific interactions among natural enemies of Bemisia in an 

inundative biological control program. Biol. Control. 6:384-393. 

Heinz, K. M, and M. P. Parrella. 1994a. Biological control of Bemisia argentifolli (Homoptera: Aleyrodidae)

561 

infesting Euphorbia pulcherrima: evaluations of releases of Encarsia luteola (Hymenoptera: Aphelinidae) and 

Delphastus pusillus (Coleoptera: Coccinellidae). Environ. Entomol. 23:1346-1353. 

Heinz, K. M, and M. P. Parrella. 1994b. Poinsettia (Euphorbia pulcherrima Wild. ex Koltz.) Cultivar-mediated 

differences in performance of five natural enemies of Bemisia argentifolii Bellows and Perring, n. sp. 

(Homoptera: Aleyrodidae). Biol. Control. 4:305-318. 

Heinz, K. M, and F. G. Zalom. 1996. Performance of the predator Delpahstus pusillus on Bemisia resistant and 

susceptible tomato lines. Entomol. Exp. Appl. 81:345-352. 

Hempel, A. 1899. Descriptions of three new species of Aleurodidae from Brazil. Psyche, Cambridge. 8: 394- 

395. 

Hempel, A. 1901. A preliminary report on some new Brazilian Hemiptera. Annals and Magazine of Natural 

History (7). 8: 383-391. 

Hempel, A. 1904. Notas sobre dois inimigos da laranjeria. Familia Aleurodidae. Bolm. Agric. S. Paulo. 5: 10- 

21. 

Hempel, A. 1918. Descripcao de uma nova especie de Aleurodidae. Revista Mus. Paul. 10: 211-214. 

Hempel, A. 1922. Algumas especies novas de Hemipteros da familia Aleyrodidae. Notas prelim. Mus. Paul. 

2(1): 3-10. 

Hempel, A. 1923. Hemipteros novos ou pouco conhecidos da familia Aleyrodidae. Notas prelim. Mus. Paul. 13: 

1121-1157; English 13: 1158-1191.

562 

Hempel, A. 1938. Uma nova especie de Aleyrodidae (Hemiptera). Archos Inst. Biol. S. Paulo. -. 

Heraty, J. M.and A. Polaszek. 2000. Morphometric analysis and descriptions of selected species in the Encarsia 

strenua group (Hymenoptera: Aphelinidae). Journal of Hymenoptera Research. 9(1), April, 2000: 142 169, 

Illustr. 

Hernandez-Suarez, E, A. Carnero, A. Aguiar, G. Prinsloo, J. LaSalle, and A. Polaszek. 2003. Parasitoids of 

whiteflies (Hymenoptera: Aphelinidae; Hemiptera: Aleyrodidae) from Macaronesian archipelagos of the Canary 

Islands, Madeira and the Azores. Systematics and Biodiversity. 1(1): 55-108. 

Heyer, W, M. L. C. Lok, and B. Cruz. 1989. The population dynamics of the cotton whitefly Bemisia tabaci 

Genn. in bean fields in the Republic of Cuba. Archiv fur Phytopathologie und Pflanzenschutz. 25:473-479. 

Hoda, F. M, G. A. Ibrahim, H. A. Taha, and M. E. El Naggar. 1986. Mites associated with pea-nut plant in 

Egypt. Bull. Soc. Ent. Egypte 66:107-111. -. 

Hodges, G. and G.A. Evans. 2005. An identification guide to the whiteflies (Hemiptera: Aleyrodidae) of the 

southeastern United States. Florida Entomologist. 88 (4): 518-534. 

Hoelmer, K. A. 1994. Evaluation of indigenous and exotic natural enemies of Bemisia in southwestern desert 

agroecosystems. In T. J. Henneberry, N. C. Toscano, R. M. Faust, and J. R. Coppedge [eds.], Sweetpotato 

whitefly: 1994 Supplement to the five-year national research and action plan. USDA/ARS, Springfield, VA. pp. 

91. 

Hoelmer, K. A, and L. S. Osborne. 1990. Biological control of sweetpotato whitefly in Florida with predators 

and parasitoids. In R. K. Yokomi, R. K, K. R. Narayanan, and D. J. Schuster [eds.], Proceedings Workshop on 

the Sweetpotato Whitefly Mediated Vegetable Disorders in Florida. Homestead, FL. Feb. 1990 IFAS, U. FL, 

Gainesville.

Hoelmer, K. A, L. S. Osborne, and R. K. Yokomi. 1993. Reproduction and feeding behavior of Delphastus 

pusillus (Coleoptera: Coccinellidae), predator of Bemisia tabaci (Homoptera: Aleyrodidae). Environ. Entomol. 

86(2):322-329. 

Hoelmer, K, and J. Hagler. 1996. Preliminary biological studies of Semidalis sp, a native neuropteran predator 

of Bemisia. In T. J. Henneberry, N. C. Toscano, R. M. Faust, and J. R. Coppedge [eds.], Silverleaf whitefly: 

1996 supplement to the 5-year national research and action plan, USDA/ARS 1996-01. pp. 117. 

Howard, L. O. 1881. Part III. Report on the parasites of the coccidae in the collection of this department. 

Annual Report of the Commissioner of Agriculture for the year 1880. 350-373 pp. 

Howard, L. O. 1894. The hymenopterous parasites of the California red scale. Insect Life. 6:227-236. 

Howard, L. O. 1894a. Two parasites of important scale-insects. Insect Life. 7:5-8. 

Howard, L. O. 1895. Revision of the Aphelinidae of North America. United States Department of Agriculture 

Division of Entomology Technical Series. 1: 1-44. 

Howard, L. O. 1898. On some parasites of Coccidae with descriptions of two new genera of Aphelininae. 

Proceedings of the Entomological Society of Washington. 4:133-139. 

Howard, L. O. 1907. New genera and species of Aphelininae with a revised table of genera. Technical Series, 

Bureau of Entomology, United States Department of Agriculture. 12(4):69-88. 

Howard, L. O. 1908. A key to the species of Prospaltella, with table of hosts, and descriptions of four new 

species. Annals of the Entomological Society of America. 1: 281-284. 

563

Howard, L. O. 1908a. On two new species of parasites of Aleyrodidae. Proceedings of the Entomological 

Society of Washington. 10:63-65. 

Howard, L. O. 1911. A note on the Indian enemies of Aleyrodes citri R. and H, with description of a new 

species of Prospaltella. Journal of Economic Entomology. 4:130-132. 

Howard, L. O. 1914. Concerning some Aphelininae. Proceedings of the Entomological Society of Washington. 

16:79-85. 

Huang, J. and A. Polaszek. 1996. The species of Encarsiella Hayat (Hymenoptera: Aphelinidae) from China. 

Journal of Natural History. 30:1649-1659. 

Huang, J. and A. Polaszek 1998. A revision of the Chinese species of Encarsia Foerster (Hymenoptera: 

Aphelinidae): parasitoids of whiteflies, scale insects and aphids (Hemiptera. Journal of Natural History. 32: 

1825-1966. 

Huang, Jian. 1994. Systematic Studies on Aphelinidae of China. Chongquing Publ. House, Chongquing. 348 pp. 

Hulden, L. 1986. The whiteflies (Homoptera, Aleyrodidae) and their parasites in Finland. Notulae 

Entomologicae. 66: 1-40. 

Hunter, C. D. 1994. Suppliers of beneficial organisms in North America, PM 94-03 ed. Calif. Environ. Prot. 

Agency, Sacramento, Calif. -. 

Husain, M. A. 1928. Annual report of entomologist of the government Punjab, Lyallpur, for the year 1927-28. 

564

565 

In Department of Agriculture [eds.], Report of the Department of Agriculture Punjab 1927-28, Part 2 ed. 

Department of Agriculture, Lahore, Punjab. (Ann. Rev. Entomol. (1930) 18:27-29). pp. 55-79. 

Hussain, M. A.and A. W. Khan. 1945. The citrus Aleyrodidae (Homoptera) in Punjab and their control. 

Memoirs of the Entomological Society of India. 1: 1-41. 

Husain, M. A, and K. N. Terhan. 1933. Observations on the life-history bionomics and control of the white-fly 

of cotton (Bemisia gossypiperda M. and L.). Indian J. Agri. Sci. 3:701-753. 

Husain, T. and M. M. Agarwal. 1982. Studies on some aphelinid (Hym.: Chalcidoidea) parasites of hemipterous 

insects of India. Journal of the Bombay Natural History Society. 79(1):155-162. 

Husain, T, A. Rauf and P. P. Kudeshia. 1983. Pomphale striptipennis gen. et sp. nov. (Hymenoptera: 

Eulophidae). Journal of Entomological Research (New Dehli). 7: 112-114. 

Hutson, J. C. 1918. Some insect pests in Cuba. Agr. News West Indies. 17:186-187. -. 

Iaccarino, F. M. 1990. Descrizione di Paraleyrodes minei n.sp. (Homoptera: Aleyrodidae), nuovo aleirodide 

degli agrumi, in Siria. Bollettino del Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 46: 131- 

149. 

Iaccarino, F. M. 1982. Descrizione di Aleuroviggianus adrianae, n. gen, n. sp. (Homoptera: Aleyrodidae). 

Bollettino Del Laboratorio Di Entomologia Agraria Filippo Silvestri. 39: 37-45. 

Ihering, H. 1897. Os piolhois vegetaes (Phytophthires) do Brazil. Revista Mus. Paul. 2: 385-420.

566 

Inayatullah, C. 1984. Sugar-cane aleyrodids, Aleurolobus barodensis (Maskell) and Neomaskellia andropogonis 

Corbett (Hom.: Aleyrodidae), and their natural enemies in Pakistan. Insect Sci. Applic. 5:279-282. 

Inayatullah, C, and M. A. Ghani. 1987. Incidence of cotton whitefly and its natural enemies in Pakistan. 

Pakistan Journal of Agricultural Research. 8:179-183. 

Inayatullah, C, and A. A. Goraya. 1980. Studies on potential biological control agents of whiteflies in Pakistan. 

Commonwealth Institute of Biological Control, Rawalpindi, Pakistan. Ann. Rept. CIBC, Pakistan station, 

Rawalpindi. 

Ishii, T. 1938. Descriptions of six new species of Aphelinidae with a revised table of genera. Kontyu. 12: 27-32. 

Jenkins, C. F. H, and D. G. Shedley. 1953. The citrus white fly (Aleuroplatus citri Tak.). J. Dep. Agric. W. 

Aust. 3 2:49-55. 

Jensen, A. S. 1999. Cladistics of a sampling of the world's diversity of whiteflies of the genus Dialeurodes 

Cockerell (Hemiptera: Aleyrodidae). Annals of the Entomological Society of America. 92: 359-369. 

Jensen, A. S. 2001. A cladistic analysis of Dialeurodes, Massilieurodes and Singhiella, with notes and keys to 

the Nearctic species and desciptions of four new Massilieurodes species (Homoptera: Aleyrodidae). Systematic 


Jesudasan, R.W.A. and B. V. David. 1990. Revision of two whitefly genera, Aleuroclava Singh and 

Aleurotuberculatus Takahashi (Homoptera: Aleyrodidae). Frederick Institute of Plant Protection & Technology, 

Entomology Series. 2: 1-13. 

Jesudasan, R.W.A. and B. V. David. 1991. Taxonomic studies of Indian Aleyrodidae (Insecta: Homoptera).


Jesudasan, R.W.A. and B. V. David. 1999. Revision of two whitefly genera, Aleuroclava Singh and 

Aleurotuberculatus Takahashi (Homoptera: Aleyrodidae). Frederick Institute of Plant Protection & Technology, 

Entomology Series. 2: 1-13. 

Jesudasan, R. W. A, C. Kandaswami & B. V. David. 1984. A new pupal parasite, Eretmocerus corni Haldeman 

(Aphelinidae: Hymenoptera) on Dialeurolonga fici David and Subramaniam (Aleyrodidae: Homoptera). 

Entomon. 9: 283-284. 

Jones, D. R. 2003. Plant viruses transmitted by whiteflies. European Journal of Plant Pathology 109: 

195-219. 

Jones, W. A, P. Silvia, and G. Snodgrass. 1997. Biology of Dereocoris nebulosus (Heteroptera: Miridae) on 

Bemisia. In T. Hennebery, N. C. Toscano, T. M. Perring, and R. M. Faust [eds.], Silverleaf whitefly: 1997 

supplement to the five-year research and action plan. USDA/ARS 1997-02. pp. 151. 

Joshi, B. C, and D. N. Yadav. 1990. Biology and feeding potential of Mallada boninensis (Okamoto), a 

chrysopid predator of whitefly Bemisia tabaci Gennadius. J. Bio. Cont. 4:18-21. 

Kairo, M, F.L. Vyjayanthi, G.V. Pollard and R. Hector. 2001. Biological control of coconut whitefly, 

Aleurodicus pulvinatus, in Nevis. Biocontrol News and Information. 22(2): 45N-50N. 

Kajita, H. 1982. Predation by adult Orius sauteri Poppius (Hemiptera: Anthocoridae) on the greenhouse 

whitefly, Trialeurodes vaporariorum (Westwood) (Homoptera: Aleyrodidae). Appl. Entomol. Zoo. 17:424-425. 

567

568 

Kajita, H, I. M. Samudra, and A. Naito. 1991. Discovery of the spiraling whitefly Aleurodicus dispersus Russell 

(Homoptera: Aleyrodidae) from Indonesia, with notes on its host plants and natural enemies. Appl. Entomol. 

Zoo. 26(3):397-400. 

Kalshoven, L. G. E. 1981. The Pests of Crops in Indonesia. P.T. Ichtiar Baru-Van Hoeve, Jakarta. (Revised and 

translated by P.A. van der Laan). 

Kamiya, H. 1963. On a new genus and species of Coccinellidae (Coleoptera) feeding on whiteflies. Mushi. 

37:127-130. 

Kapadia, M. N, and S. N. Puri. 1989. Seasonal incidence of natural enemies of Bemisia tabaci (Gennadius). 

India J. Ecol. 2 16:164-168. 

Kapadia, M. N, and S. N. Puri. 1992. Biology of Serangium paracesetosum Sicard as a predator of cotton 

whitefly. J. Maharashtra Agri. Univ. 17:162-163. 

Kapadia, M. N, and S. N. Puri. 1992a. Development of Chrysoperla carnea reared on aphids and whitefly. J. 

Maharashtra Agri. Univ. 17:163-164. 

Kapadia, M. N, and S. N. Puri. 1993. New records of natural enemies of castor whitefly Trialeurodes ricini 

(Misra). J. Maharashtra Agri. Univ. 18:339. 

Kapur, A. P. 1940. A note on the ladybird beetle (Coccinellidae) predating on the cane whitefly (Aleurolobus 

barodensis Mask.). Current Sci. 9:134. 

Kapur, A. P. 1942. Bionomics of some Coccinellidae, predaceous on aphids and coccids in North India. Indian 

J. Entomol. 4:49-66.

Kapur, A. P. 1954. A new species of Coccinellidae (Coleoptera) predacious on the citrus whitefly in India. 

Records Indian Museum. 52:189-193. 

Karsh, F. 1888. Kritik von Westhoff's Arbeit. Ent. Nachr. 14: 31. 

Khan, A. G, and A. I. Mohyuddin. 1983. Aleurolobus barodensis (Maskell) (Homoptera: Aleyrodidae) the 

sugarcane whitefly and possibilities of its control by augmentation of its native parasites. Pakistan J. Zool. 

15:191-196. 

Khan, A. G, A. I. Mohyuddin, and A. A. Goraya. 1991. Studies on citrus whiteflies and their natural enemies in 

Pakistan. Pakistan J. Zool. 23:127-132. 

Khan, M. Q, and A. S. Rao. 1948. Annual report of the scheme for research for the pests and diseases of castor 

and other oilseeds in Hyderabad State. 1943-1945. The Nizam's Government, Hyderabad Division, Government 

Press, Hyderabad, India. 

Khan, M.Y. and S.A. Shafee. 1976. Indian species of the genus Azotus (Hymenoptera, Aphelinidae). Geobios, 

Jodhpur. 3(5):172-174. 

Khan, M.Y. and S.A. Shafee. 1980. Species of the genus Eretmocerus (Aphelinidae: Eretmocerinae) from India. 

Oriental Insects. 14(3):363-370. 

Kirejtshuk, A. G, D. G. James, and R. Heffer. 1997. Description and biology of a new species of Cybocephalus 

Erichson (Coleoptera: Nitidulidae), a predator of Australian citrus whitefly. Australian J. Entomol. 36:81-86. 

569

Kiriukhin, G. 1947. Quelques Aleurododea de l'Iran. Entomologie Phytopath. app. 5:8-10. 

Kirk, A, L. Lacey, J. Goolsby, M. Ciomperik, W. Able, C. Pickett, and J. Gould. 1996. Results of foreign 

exploration for parasitoids of SLWF. In T. J. Henneberry, N. C. Toscano, R. M. Faust, and J. R. Coppedge 

[eds.], Silverleaf whitefly: 1996 supplement to the 5-year national research and action plan, USDA/ARS 1996- 

01. pp. 124. 

Kirk, A, and L. Lacey. 1996. Efficacy of Clitostethus arcuatus a predator of Bemisia argentifolii. In T. J. 

Henneberry, N. C. Toscano, R. M. Faust, and J. R. Coppedge [eds.], Silverleaf whitefly: 1996 supplement to the 

5-year national research and action plan, USDA/ARS 1996-01. pp. 122. 

Kirk, A. A, L. A. Lacey, N. Roditakis, and J. K. Brown. 1993. The status of Bemisia tabaci (Hom, 

Aleyrodidae), Trialeurodes vaporariorum (Hom, Aleyrodidae) and their natural enemies in Crete. 

Entomophaga. 38:405-410. 

Kirkaldy, G. W. 1907. A catalogue of the Hemipterous family Aleyrodidae. Bulletin, Board of Commisioners of 

Agriculture and Forestry, Hawaii, Division of Entomology. 2: 1-92. 

Klimaszewski, S. and H. Szelegiewicz. 1962. Materialen zur Kenntnis der Mottenlause (Homoptera: 

Aleyrodidae) Polens. Fragmenta Faunistica. 10: 35-45. 

Ko, C. C, Chou, L.Y. and W. J. Wu. 1998. Aleyrodidae of Taiwan (Homoptera). Part 4. Unrecorded species. 

Entomological Science. 1(1) 77-79. 

Ko, C. C. and A. K. Dubey. 2007. Aleurocyperus humus gen. et sp. n. (Hemiptera: Aleyrodidae) 

from Taiwan, with interesting feeding behaviour. Zootaxa. 1443: 37-54. 

570

571 

Ko, C, Chou, L. and W. Wu. 2001. Agrostaleyrodes arcanus, a new genus and species of whitefly (Homoptera: 

Aleyrodidae) from Taiwan. Zoological Studies. 40(2): 177-186. 

Ko, C. C, T. C. Hsu & W.J. Wu. 1990. The whitefly of Rhododendron from Taiwan (Homoptera: Aleyrodidae). 

Chinese Journal of Entomology. 10: 225-233. 

Ko, C. C, T. C. Hsu & W.J. Wu. 1992. Aleyrodidae of Taiwan. Part 1. Rhachisphora Quaintance et Baker. 

Chinese Journal of Entomology. 60(1): 243-260. 

Ko, C. C, T. C. Hsu & W.J. Wu. 1993. Aleyrodidae of Taiwan. Part II. Asialeyrodes Corbett. Chinese Journal of 

Entomology. 61(3): 613-618. 

Ko, C.C, T.C. Hsu & W.J. Wu. 1995. Aleyrodidae (Homoptera) of Taiwan Part III. Aleuromarginatus Corbett. 

Japanese Journal of Entomology. 63(4): 892-896. 

Ko, C. C. and Z.Y. Luo. 1999. On some whiteflies (Homoptera: Aleyrodidae) from China. Chinese Journal of 

Entomology. 19(1), March, 1999: 19 25, illustr. 

Ko, C. C, Wu, W. J. and L. Y. Chou. 1998. A new species of Rhachisphora (Homoptera: Aleyrodidae) from 

Taiwan. Chinese Journal of Entomology. 18(1), March, 1998: 61 65, illustr. 

Ko, C. C, W.J. Wu and L. Y. Chou. 1998a. Aleurocybotus (Homoptera: Aleyrodidae) of Taiwan. Chinese 

Journal of Entomology. 18: 177-186. 

Ko, C.C, W.J. Wu and L.Y. Chou. 1998b. Aleyrodidae of Taiwan (Homoptera). Part 4. Unrecorded species. 

Entomological Science. 18: 177-186.

Koch, C. L. 1857. Die Pflanzenlause Aphiden. Nurnberg. 330 pp. 

Kodama, G. 1931. Studies on Aleurocanthus spiniferus Quaint. Kagoshima-Ken, Kyushu, Japan. -. 

Korobitsin, V. G. 1967. New and little known species of aleyrodids (Hemiptera: Aleyrodoidea) from Crimea [in 

Russian]. Entomologicheskoe Obozrenie. 46: 857-859. 

Kotinsky, J. 1907. Aleyrodidae of Hawaii and Fifi with descriptions of new species. Bulletin, Board of 

Commissioners of Agriculture and Forestry Hawaii, Division of Entomology. 2: 93-102. 

Kozar, F, M. A. Fowjhan & M. Zarrabi. 1996. Check-list of Coccoidea and Aleyrodoidea (Homoptera) of 

Afghanistan and Iran, with additional data to the scale insects of fruit trees in Iran. Acta Phytopathologica et 

Entomologica Hungarica. 31(1-2): 61-74. 

Krishnamurthy, K. V, A. Raman and B. V. David. 1973. Foliar pit-galls of Morinda tinctoria Roxb. Cecidologia 

Ind. 8: 75-77. 

Krishnamurti, B. and Usman, S. 1954. Some insect parasites of economic importance noted in Mysore State. 

Indian Journal of Entomology. 16: 327-344. 

Krishnan, B. and B. V.David. 1996. Records and descriptions of some aphelinid parasitoids of Aleyrodidae 

(Homoptera: Insecta) from India. Jai Res. Foundt., Valvada, Gujaratii. 65 pp. 

Kuchanwar, D. B, M. G. Hardas, M. N. Borle, and B. K. Sharnagat. 1982. Catana parcesetosa, a potential 

predator of the citrus black fly, Aleurocanthus woglumi Ashby. P. K. V. Res. J. 6:74. 

572

Kuenzel, N. T. -. Population dynamics of protelian parasites attacking a natural population of Trialeurodes 

packardi and new host records fro two species. Proceeding of the Entomological Society of Washington. 79(3): 

400-404. 

Kuhne, S, K. Schiller, and U. Dahl. 1997. Beitrag zur Lebensweise, Morphologie und Entwicklungsdauer der 

rpuberischen Fliege Coenosia attenuata Stein (Diptera: Muscidae). Gesunde Pflanzen. 49:100-106. 

Kumashiro, B. R., Lai, P. Y. and Funasaki, G. Y. 1983. Efficacy of Nephaspis amnicola and Encarsia haitiensis 

in controlling Aleurodicus dispersus in Hawaii. Proceedings of the Hawaiian Entomological Society. 24 (2-3): 

261-269. 

Kunhof, E, C. Sengonca, and E. Montagne. 1992. Laboratory studies on the development and feeding activity of 

Uloborus plumipes Lucas (Araneae, Uloboridae). Mitteilungen der Deutschen Gesellschaft fur Allgemeine und 

Angewandte Entomologie. 8:204-208. 

Kunow, G. 1880. Zweineue Schildiause. Ent. Nachr. 6: 46-47. 

Kurdjumov, N. V. 1916. New genus and species of Aphelininae (Chalcidoidea). Zhurnal Prikladnoi 

Entomologii (Kiev). 1: 80-81. 

Kuwana, I. 1911. The whiteflies of Japan. Pomona College Journal of Entomology. 3: 620-627. 

Kuwana, I. 1922. Bemisia shinanoensis n. sp. A new whitefly from Japan. Journal of Plant Protection. 9:461- 

464. 

573

574 

Kuwana, I. 1927. On the genus Bemisia [Family Aleyrodidae] found in Japan, with description of a new 

species. Annotationes Zoologicae Japonensis. 11: 245-253. 

Kuwana, I. 1928. Aleyrodidae or white flies attacking citrus plants in Japan. Sci. Bull. Min. Agric. Forest. Dept. 

1:41-78. 

Kuwana, I, and T. Ishii. 1927. On Prospaltella smithi Silv. and Cryptognatha sp., the enemies of Aleurocanthus 

spiniferus, imported from Canton, China. J. Okitsu Hort. Soc. 22:77-80. 

Kuwana, I. and K. Muramatsu. 1931. New scale insects and whitefly found upon plants entering Japanese ports 

[In Japanese]. Zool. Mag. Tokyo. 43:647-660. 

Kuznetov, V. N. 1997. Lady Beetles of the Russian Far East. Center for Systematic Entomology Memoir. 1: 248 

pp. 

LaSalle, J. 1999. A new species group and two new species of Euderomphale Girault (Hymenoptera: 

Eulophidae) from North America. Journal of Hymenoptera Research. 8(1): 116-119. 

LaSalle, J. and M. E. Schauff. 1994. Systematics of the tribe Euderomphalini (Hymenoptera: Eulophidae): 

parasitoids of whiteflies (Homoptera: Aleyrodidae). Systematic Entomology. 19:235-258. 

LaSalle, J, A. Polaszek, J. S. Noyes and G. Zolnerowich. 1997. A new whitefly parasitoid (Hymenoptera: 

Pteromalidae: Eunotinae) with comments on its placement, an implications for classification of Chalcidoidea 

with particular reference to the Eriaporinae (Hymenoptera: Aphelinidae). Systematic Entomology. 22(2):131- 

150. 

Laing, F. 1927. Coccidae, Aphidae and Aleyrodidae. Insects Samoa. 1(2): 35-45.

Laing, F. 1928. Description of a new white fly pest of Rhododendron. Entomologist's Monthly Magazine. 

64:228-230. 

Laing, F. 1930. Description of a new species of Aleyrodidae (Rhynch.). Stettin. Ent. Zig. 91: 219-221. 

Lal, K. B. 1938. The first record of the eulophid genus Azotus from India (Chalcidoidea: Hymenoptera). Rec. 

Indian Mus. 40: 1-4. 

Landis, B. J, K. E. Gibson, and R. Schopp. 1958. The Iris whitefly in the Pacific Northwest. Ann. Entomol. Soc. 

Am. 51:486-490. 

Latreille, P. A. 1795. Magazin encyclopedique. 4:304-310. 

Latreille, P. A. 1801-2. Histoire naturelle des crustaces et des insectes. Paris. -. 3: 468 pp. 

Lee, L. H. Y. and R. Winney. 1981. Checklist of Agricultural Insects of Hong Kong 1981. Agricultural and 

Fisheries Department Bulletin No. 2. 

Legaspi, J. C, R. I. Carruthers, D. A. Nordlund, J. A. Correa, A. C. Cohen, and D. Holbrook. 1994. Effect of 

inundative releases of the predator, Chrysoperla rufilabris (Neuroptera) to control sweetpotato whitefly in an 

organic field crop. In T. J. Henneberry, N. C. Toscano, R. M. Faust, and J. R. Coppedge [eds.], Sweetpotato 

whitefly: 1994 Supplement to the five-year national research and action plan. USDA/ARS, Springfield, VA. pp. 

96. 

Legaspi, J. C, J. A. Correa, B. C. Carruthers, J. Legaspi, and D. A. Nordlund. 1996. Effect of short-term releases 

575

576 

of Chrysoperla rufilabris (Neuroptera: Chrysopidae) against silverleaf whitefly (Homoptera: Aleyrodidae) in 

field cages. J. Entomol. Sci. 31:102-111. 

Legaspi, J. C, B. C. Legaspi Jr, R. L. Meagher, and M. A. Ciomperlik. 1996. Evaluation of Serangium 

parcesetosum (Coleoptera: Coccinellidae) as a biological control agent of the silverleaf whitefly (Homoptera: 

Aleyrodidae). Environ. Entomol. 25:1421-1427. 

Legaspi, J. C, D. A. Nordlund, and B. C. Legaspi Jr. 1996. Tri-trophic interaction and predation rates in 

Chrysoperla spp. attacking the silverleaf whitefly. Southwestern Entomol. 21:33-42. 

Leon, G, G. A. Evans and J. C. Campos. 2001. Parasitoides de plagas (Homoptera) de los citricos en el 

departamento de Meta, Colombia. Revista Colombiana de Entomologia. 27(3-4): 143-146. 

Leonardi, G. 1910. Due nuove specie di Aleurodicus Douglas. Bollettino del Laboratorio di Zoologia Generale e 

Agraria della R. Scuola Superiore d'Agricoltura, Portici. 4:316-322. 

Lewis, R. T. 1893. On a new species of Aleurodes. Rep. Ealing microse. nat. Hist. Soc. 1893:1-3. 

Lima, A. F. and R. Racca-Filho. 2005. Peracchius durantae, a new genus and species of whitefly (Hemiptera: 

Aleyrodidae) from Brazil. Zootaxa. 1045: 39-43. 

Lindinger, L. 1932. Randbemerkungen. Entomologische Rundschau. Stuttgart. 1932: 222-223. 

Link, D, and E. C. Costa. 1980. Ocorrpncia de inimigos naturais da mosca blanca, Bemisia tabaci (Gennadius, 

1989), na cultura da soja. Rev. Centro Cipncias Rurais. 10:111-113.

Linnaeus, C. 1758. Systema Naturae. -. 824 pp Uppsula. 

Liotta, G. 1978. Effetti secondari dei fitofarmaci comunemente adoperati contro Dialeurodes citri (Ashm.) 

(Hom. Aleyrodidae) su Clitostethus arcuatus (Rossi) (Col. Coccinellidae). Atti XI Congr. Ital. Ent. Portici- 

Sorrento 10-19 maggio 1976. pp. 437-444. 

Liotta, G. 1983. Osservaziioi bio-etologiche su Clitostethus arcuatus (Rossi) (Coleoptera: Coccinellidae) in 

Siccilia. Redia. 64:173-185. 

Liu, T. X, and P. A. Stansly. 1996. Functional response of Nephaspis oculatus (Coleoptera: Coccinellidae) to 

egg density of Bemisia argentifolii. In T. J. Henneyberry [ed.], Silverleaf Whitefly 1996 Supplement to the 

Five-Year National Research and Action Plan: Progress Review, Technology Transfer, and New Research and 

Action Plan (1997-2001). USDA/ARS, Washington DC. pp. 157. 

Liu, T. X, and P. A. Stansly. 1996a. Morphology of Nephaspis oculatus and Delphastus pusillus (Coleoptera: 

Coccinellidae), predators of Bemisia argentifolii (Homoptera: Aleyrodidae). Proc. Entomol. Soc. Wash. 98:292- 

300. 

Liu, T. X, and P. A. Stansly. 1996c. Toxicological effects of selected insecticides on Nephaspis oculatus (Col, 

Coccinellidae), a predator of Bemisia argentifolii (Hem, Aleyrodidae). J. Appl. Entomol. 120:369-373. 

Liu, T. X, P. A. Stansly, K. A. Hoelmer, and L. S. Osborne. 1997. Life history of Nephaspis oculatus 

(Coleoptera: Coccinellidae), a predator of Bemisia argentifolii (Homoptera: Aleyrodidae). Ann. Entomol. Soc. 

Am. 90:776-782. 

Loi, G. 1979. Osservazioi eco-etologiche sul coleottero Coccinellidae scimnino Clitostethus arcuatus (Rossi), 

predatore di Dialeurodes citri (Ashmead) in Toscana. Frustula Entomol. 1:123-145. 

577

Longo, S, C. Rapisarda, A. Russo, and G. Siscaro. 1990. Preliminary studies on biology of Parabemisia 

myricae (Kuwana) and its entomophagous in Sicily and Calabria (Italy). Boll. Zool. agr. Bachic. 22:161-172. 

Longo, S, C. Rapisarda, and A. Russo. 1985. Risultati del controllo biologico dell' Aleurothrixus floccosus 

(Maskell) in agrumeti della Sicilia Orientale. In [eds.], Atti XIV Congr. naz. ital. Ent, Palermo-Erice-Bagheria. 

Palermo-Erice-Bagheria, Palermo-Erice-Bagheria. pp. 841-848. 

Lopez-Avila, A. 1987. Two new species of Encarsia Foerster (Hymenoptera: Aphelinidae) from Pakistan, 

associated with the cotton whitefly, Bemisia tabaci (Gennadius) (Homoptera: Aleyrodidae). Bulletin of 


Low, F. 1867. Zool. Notizen Zweite Serie. Verhandlungen der Zoologische-Botanishcen Gesellschaft in Wien. 

17: 745-752. 

MacGown, M. W. and T. E. Nebeker. 1978. Taxonomic review of Amitus (Hymenoptera: Proctotrupoidea, 

Platygasteridae) of the Western Hemisphere. Canadian Entomologist. 110: 275-283. 

Mackie, D. B. 1912. A new coconut pest. Philippine Agriculture Rev. 5: 142-143. 

Maki, M. 1915. Studies concerning the more important insects pests of the street trees and ornamental plant [In 

Japanese]. Spec. Rep. Formosa Forest Exp. Station. 1: 30-31. 

Malausa, J. C, E. Franco, and P. Brun. 1988. Establishment of the Azur Coast and in Corsica (France) of 

Serangium parcesetosum (Coleoptera: Coccinellidae), a predator of the citrus whitefly, Dialeurodes citri 

(Homoptera: Aleyrodidae). Entomophaga. 33:517-519. 

578

579 

Malausa, J. C, J. C. Onillon, and F. Franco. 1986. Distribution on the Cpte d'Azur of Encarsia laborensis 

(How.) a parasite introduced against the citrus whitefly, Dialeurodes citri (Ashm.). In Integrated pest control in 

citrus groves, R. Cavalloro and D. Martino [eds.], Proc. Experts' Meeting Acireale 26-29 March 1985. A. A. 

Balkema, Rotterdam. pp. 365-367. 

Mani, M. S. 1939. Descriptions of new and records of some known chalcidoid and other hymenopteous 

parasites from India. Indian Journal of Entomology. 1: 69-99. 

Mani, M. S. 1940. Studies on Indian parasitic Hymenoptera. I. Indian Journal of Entomology. 3: 25-36. 

Mani, M. S. 1941. Studies on Indian parasitic Hymenoptera. I. Indian Journal of Entomology. 3: 25-36. 

Manzari, S and H. Alemansoor. 2005. A new species of Acaudaleyrodes Takahashi (Hemiptera: Aleyrodidae) 

from Iran, with a key to the species of the genus. Zootaxa. 829: 1-6. 

Manzari, S, A. Polaszek, R. Belshaw and D. L. J. Quick. 2002. Morphometric and molecular analysis of the 

Encarsia inaron species-group (Hymenoptera: Aphelinidae), parasitoids of whiteflies (Hemiptera: Aleyrodidae). 

Bulletin of Entomological Research. 92: 165-175. 

Martin, J. H. 1985. The whitefly of New Guinea. Bulletin of the British Museum (Natural History) 

(Entomology). 50(3): 303-351. 

Martin, J. H. 1985a. Aleurochiton acerinus Haupt, a maple-feeding whitefly (Hom, Aleyrodidae) new to Britain. 

Entomologist's Monthly Magazine. 121: 143-144. 

Martin, J. H. 1985b. The whitefly genus Dialeurolonga (Hemiptera: Aleyrodidae) on citrus, with the 

redescription of a Malagasian species. Bulletin of Entomological Research. 75: 459-462.

Martin, J. H. 1987. An identification guide to common whitefly pest species of the world (Homoptera: 

Aleyrodidae). Tropical Pest Management. 33: 298-322. 

Martin, J. H. 1988. Whitefly of northern Sulawesi, including new species from clove and avocado (Homoptera: 

Aleyrodidae). Indo-Malayan Zoology. 5: 57-85. 

Martin, J. H. 1990a. The whitefly pest species Aleurodicus dispersus and its rapid extension of range across the 

Pacific and South East Asia. Newsletter, Malaysian Plant Protection Society. 14: 36. 

Martin, J. H. 1996. Neotropical whiteflies of the subfamily Aleurodicinae established in the western Palearctic 

(Homoptera: Aleyrodidae). Journal of Natural History. 30, 1849-1859. 

Martin, J.H. 1997. The type species of Aleurodicus Douglas, a whitefly genus of economic importance 

(Homoptera: Aleyrodidae). Memoirs Museum Victoria. 56(1): 125-128. 

Martin, J. H. 1999. The whitefly fauna of Australia (Sternorrhyncha: Aleyrodidae). A taxonomic account and 

identification guide. Technical Paper, Division of Entomology, Commonwealth Scientific and Industrial 

Research Organization, Canberra. 38: 1-197. 

Martin, J. H. 2000. The occurrence of Dialeurodes citrifolii and Aleurochiton pseudoplatani (Hemiptera: 

Aleyrodidae) in Lebanon - important new records in the Europe-Mediterranean region. Bulletin of 


Martin, J. H. 2001. Description of an invasive new species of Neotropical aleurodicine whitefly (Hemiptera: 

Aleyrodidae) - a case of complete or partial misidentification?. Bulletin of Entomological Research. 91: 101- 

107. 

580

Martin, J. H. 2004. Whiteflies of Belize (Hemiptera: Aleyrodidae) Part 1 - introduction and account of the 

subfamily Aleurodicinae Quaintance & Baker. Zootaxa. 681: 1-199. 

Martin, J. H. 2005. Aleurodicus talamancensis, a new whitefly species damaging plantation bananas in Costa 

Rica, with discussion of a montane orchid-feeding population (Sternorrhyncha, Aleyrodidae). Zootaxa. 843: 1- 

10. 

Martin, J. H. 2005a. Whiteflies of Belize (Hemiptera: Aleyrodidae), Part 2 - a review of the subfamily 

Aleyrodinae Westwood. Zootaxa. 1098: 1-116. 

Martin, J. H. 2007. Giant whiteflies (Sternorrhyncha: Aleyrodidae): a discussion of their taxonomic and 

evolutionary significance with a description of a new species of Udamoselis Enderlein from Ecuador. 

Tijdschrift voor Entomologie 150: 13-29. 

Martin, J. H, A. M. F. Aguiar and P. Baufeld. 2001. Crenidorsum aroidephagus Martin & Aguiar sp. nov. 

(Sternorrhyncha: Aleyrodidae), a New World whitefly species now colonising cultivated Araceae in Europe, 

Macronesia and the Pacific Region. Zootaxa. 4: 1-8. 

Martin, J. H, A. M. F. Aguiar and M. T. Pita. 1996. Aleyrodidae of Madeira - descriptions of three new species 

and notes on a pan-Mediterranean species of Aleurotrachelus. Journal of Natural History. 30: 113-125. 

Martin, J. H. and J. M. Camus. 2001. Whiteflies (Sternorrhyncha, Aleyrodidae) colonising fern (Pteridophyta: 

Filicopsida), with descriptions of two new Trialeurodes and one new Metabemisia species from south-east Asia. 

Zootaxa. 2: 1-19. 

Martin, J H, Hernandez, S, Suarez, E. and A. Carnero. 1997. An introduced new species of Lecanoideus 

581

582 

(Homoptera: Aleyrodidae) established and causing economic impact on the Canary Islands. Journal of Natural 

History. 31(8), August, 1997: 1261 1272, illustr. 

Martin, J. H. and G. R. Lucas. 1984. Aleurodicus dispersus Russell (Homoptera, Aleyrodidae), a whitefly 

species new to Asia. Philippine Scientist. 21: 168-171. 

Martin, J. and C. P. Malumphy. 2002. A new species of South American whitefly (Sternorrhyncha: 

Aleyrodidae) colonising cultivated bay laurel. Zootaxa. 83: 1-10. 

Martin, J. H, D. Mifsud and C. Rapisarda. 2000. The whiteflies (Hemiptera: Aleyrodidae) of Europe and the 

Mediterranean Basin. Bulletin of Entomological Research. 90: 407-448. 

Martin, J. H. and L. A. Mound. 2007. An annotated check list of the world's whiteflies (Insecta: Hemiptera: 

Aleyrodidae. Zootaxa 1492: 1-84. 

Martin, J. H. and A. Polaszek. 1999. A new genus of Neotropical whitefly, secreting blue iridescent wax 

(Sternorrhyncha, Aleyrodidae, Aleurodicinae), and its parasitoids (Hymenoptera, Aphelinidae). Journal of 

Natural History. 33(10), October, 1999: 1545 1559, illustr. 

Martin, J. H. and J. Streito. 2003. A second species of the enigmatic whitefly genus Stenaleyrodes Takahashi 

(Sternorrhyncha, Aleyrodidae). Zootaxa. 349: 1-15. 

Masi, L. 1909. Contribuzioni alla conoscenza dei Calcididi Italiani (parte 3a). Bollettino del Laboratorio di 

Zoologia Generale e Agraria della R. Scuola Superiore d'Agricoltura, Portici. 4:3-37, 29 figs. 

Masi, L. 1910. Contribuzioni alla conoscenza dei Chalcididi italiana (parte 3a). Bollettino del Laboratorio di 

Zoologia Generale e Agraria, Portici. 4: 1-37.

Masi, L. 1911. Contribuzioni alla conoscenza dei Calcididi Italiani (parte IV). Bollettino del Laboratorio di 

Zoologia Generale e Agraria della R. Scuola Superiore d'Agricoltura, Portici. 5:140-171,. 

Maskell, W. M. 1879. On some Coccidae in New Zealand. Trans. Proc. N.Z. Inst. 11(1878):187-228. 

Maskell, W. M. 1890. On some Aleurodidae from New Zealand and Fiji. Trans. Proc. N.Z. Inst. 22(1889): 170- 

176. 

Maskell, W. M. 1890a. On some species of Psyllidae in New Zealand. Trans. Proc. N.Z. Inst. 22(1889): 157- 

170. 

Maskell, W. M. 1895. Contributions towards a monograph of the Aleurodidae, a family of Hemiptera - 

Homoptera. Transactions of the New Zealand Institute. 28: 411-449. 

Maskell, W. M. 1896. Aleurodes eugeniae, a new species of bug. Indian Museum Notes. 4: 52-53. 

Masner, L. and L. Huggert. 1989. World review and keys to genera of Inostemmatinae. Memoirs of the 

Entomological Society of Canada. 147: 3-224. 

Mason, R. T, H. M. Fales, R. T. Eisner, and H. M. Eisner. 1991. Wax of a whitefly and its utilization by a 

chrysopid larva. Naturwissenschaften. 78:28-30. 

Mayne, R. and J. Ghesquiere. 1934. Hemipteres nuisile aux vegetaux du Congo belge. Famille des Aleurodidae. 

Annls. Gembloux. 40: 29-31. 

583

584 

Mayne, R, and J. Ghesquiere. 1934. Hemipteres nuisibles aux vegetaux du Congo Belge. Ann. Gembloux. 

40:29-31. 

Meade, D. 1995. A dipteran predator of whiteflies, Drapetis (Diptera: Empididae) in Arizona. -. -. 

Medina Gaud, S. 1961. The Thysanoptera of Puerto Rico. Agricultural Experimental Station, Technical Paper 

32 ed. Univ. of Puerto Rico, San Jose. -. 

Medina Gaud, S, A. E. Segarra Carmona, and R. A. Franqui. 1991. La mosca negra de los citircos, 

Aleurocanthus woglumi Ashby (Hyomoptera: Aleyrodidae), en Puerto Rico. J. Agric. Univ. Puerto Rico. 

75:301-305. 

Meganathan, P. and B. V. David. 1994. Aleyrodid fauna (Aleyrodidae: Homoptera) of Silent Valley, a tropical 

evergreen rain forest, in Kerala, India. Fippat Entomology Series 5. 1994: 1 66, illustr. 

Menge, J. A. 1856. Lebenszeichen vorwelticher, im Bernstein eingeschlossener Thiere. In. Programm. Schuler 

der Petrischule, Danzig. 42 pp. 

Mentzelos, I. A. 1967. Contribution to the study of entomophagous insects of Siphonius phillyreae Haliday (= 

inaequalis Guatier) (Aleyrodidae) on pear trees in Central Macedonia. Rep. Pl. Prot. Agric. Res. Station 

Thessaloniki. 3:92-102. 

Mercet, G. M. 1925. El genero Aphycus y sus afines. EOS. 1: 7-31. 

Mercet, R. G. 1912a. Los enemigos de los parasitos de las planas. Los Afelinidos. Trabajos de Museo de 

Ciencias Naturales, Madrid. 10: 1-306 pp.

Mercet, R. G. 1916. Signiphorinos de Espana (Hymenoptera; Chalc.). Boletin de la Real Sociedad Espanola de 

Historia Natural. 16:519-533. 

Mercet, R. G. 1917. Especies espanoles del genero Aphycus Mayr. Boletin de la Real Sociedad Espanola de 

Historia Natural. 17: 128-139. 

Mercet, R. G. 1921. Fauna Iberica. Himenopteros Fam. Encirtidos. Museo Nacional de Ciencas Naturales, 

Madrid. 

Mercet, R. G. 1927. Afelinidos de Chile. Revista Chileana de Historia Natural. 31: 129-131. 

Mercet, R. G. 1928. Afelinidos palearcticos (Hym. Chalc.). 1a nota. Boletin de la Real Sociedad Espanola de 


Mercet, R. G. 1928a. Afelinidos palearcticos (Hym. Chalc.) 2a nota. Boletin de la Real Sociedad Espanola de 


Mercet, R. G. 1930. Los aphelinidos de Espana. Revista de Biologia Forestal y Limnologia (B). 2(2):29-106. 

Mercet, R. G. 1930a. Afelinidos palearticos (Hym. Chalc. ) 4, nota. EOS. 6: 191-199. 

Mercet, R. G. 1931. Afelinidos paleacticos (Hym. Chalc.) 7a nota. Boletin de la Real Sociedad Espanola de 


Mercet, R. G. 1931a. Le genre Encarsia et description d' Encarsia indifferentis nov. sp. d'Egypte (Hymen. 

Chalc.: Aphelinidae). Bulletin de la Societe Entomologique d'Egypte. 1930:220-223. 

585

Mercet, R. G. 1931c. Notas sobre Aphelinidos (Hym. Chalc.) 4a nota. Revista Espanola di Entomologia. 

Madrid. 

Mercet, R.G. 1916. Signiphorinos de Espana (Himenopteros Chalcides). Boletin de la Real Sociedad Espanola 

de Historia Natural. 16: 519-533. 

Meyerdirk, D. E, J. B. Kreasky and W. G. Hart. 1980. Whiteflies (Aleyrodidae) attacking citrus om southern 

Texas with notes on natural enemies. Canadian Entomologist. 112: 1253-1258. 

Meyerdirk, D. E, and D. L. Coudriet. 1985. Predation and developmental studies of Euseius hibisci (Chant) 

(Acarina: Phytoseiidae) feeding on Bemisia tabaci (Gennadius) (Homoptera: Aleyrodidae). Environ. Entomol. 

14:24-27. 

Meyerdirk, D. E, and D. L. Coudriet. 1986. Evaluation of two biotypes of Euseius scutalis (Acari: Phytoseiidae) 

as predators of Bemisia tabaci (Homoptera: Aleyrodidae). J. Econ. Entomol. 79:659-663. 

Mifsud, D. and V. Palmeri. 1996. A new species of Aleurolobus Quaintance et Baker (Homoptera, Aleyrodidae) 

from southern Europe. Bollettino Del Laboratorio Di Entomologia Agraria Filippo Silvestri. 52 1996: 89 95, 

illustr. 

Mills, N. J. 1981. Essential and alternative foods for some British Coccinellidae (Coleoptera). Entomol. Gazette. 

32:197-202. 

Mimeur, J. M. 1844. Neomaskellia bergii Signoret (Hemiptera: Aleurodidae). Bull. Soc. Sci. nat. Maroc. 24: 89. 

586

587 

Mimeur, J. M. 1946. Aleurodidae du Maroc (1re note). Bul. Soc. sci. nat. phy. Maroc. 24:87-89. 

Minelli, A, S. Ruffo and S. La Posta. 1995. Homoptera: Sternorrhyncha- Checklist delle specie della fauna 

italiana. Ministero dell-ambiente e comitato scientifico per la Fauna d'Italia. 55 pp. 

Mineo, G. and G. Viggiani. 2001. Notizie preliminari su una Signiphora (Hymenoptera: Signiphoridae) ottenutu 

da Aleurothrixus floccosus (Maskell) (Homoptera: Aleyrodidae) in Italia. -. -. 

Misra, C. S. 1924. The citrus whitefly, Dialeurodes citri in India and its parasite, together with the life history 

of Aleurodes ricini, n. sp. In Report Proceedings Fifth Entomology Meeting, Pusa, 1923. Indian Agriculture, 

Pusa, Sir-Lanka. pp. 129-135. 

Misra C. S. and K. L. Singh. 1929. The cotton whitefly (Bemisia gossypiperda n. sp.). Bull. agric. Res. Inst. 

Pusa. 196: 1-7. 

Miyatake, M. 1961. A new genus of the Coccinellidae from Japan and the Ryukyu Islands (Coleoptera). Mem. 

Ehime Univ. Sect. VI (Agriculture). 6:37-43. 

Modeer, A. 1778. Om Fastflyet Coccus. Gotteborgs Kungl. Vetenskaps och Vitterhets Samhalles Handlingar. 

Ny Tidsfoljd. 1:11-50. 

Mokrecki, S. A. 1916. A short review of the work of the Salir Pomologial Station at Simferpol for 1913-1915. 

Simferpol. 40 pp. 

Morgan, A. C. F. 1892. A new genus and species of Aleurodidae. Entomologist's Monthly Magazine. 28: 29-33.

588 

Morgan, A. C. F. 1893. Aleyrodes citrifolii. pp. 70-74. In. Stubbs, W.C. & Morgan, H.A. The orange and other 

citrus fruits from seed to market, with insects beneficial and injurious with remedies for the latter. Special 

Bulletin of the Louisanna State Experiment Station. 70-74. 

Morrill, A.W. 1903. Life history and description of the strawberry Aleyrodes. A. packardi n. sp. Canadian 

Entomologist. 35: 25-35. 

Morrill, A.W. 1903a. Notes on some Aleyrodes from Massachusetts, with description of new species. Psyche, 

Cambridge. 10: 80-85. 

Morrill, A. W, and E. A. Back. 1912. Natural control of whiteflies in Florida. Bureau of Entomology, Bull. 102, 

USDA, Washington. 

Motshulsky, V. de. 1859. IV. Entomologie appliquee. Insectes utiles et nuisibles. Etudes Entomologiques. 8: 

169-174. 

Motshulsky, V. de. 1863. Essai d'un catalogue des insectes de 1'ile Ceylan. Bulletin de la Societe Imperial des 

Naturalistes de Moscou. 36(3): 1-153. 

Moulton, D. 1932. The Thysanoptera of South America. Rev. de Entomologia. 2:451-484. 

Mound, L. A. 1956. A revision of the British Aleyrodidae (Hemiptera: Homoptera). Bulletin of the British 

Museum (Natural History) (Entomology). 17: (9): 397-428. 

Mound, L. A. 1961. A new genus and four new species of whitefly from ferns (Homoptera: Aleyrodidae). 

Revue de Zoologie et de Botanique Africaines. 64: 127-132.

Mound, L. A. 1965. An introduction to the Aleyrodidae of Western Africa (Homoptera). Bulletin of the British 


Mound, L. A. 1966. A revision of the British Aleyrodidae (Hemiptera: Homoptera). Bulletin of the British 


Mound, L. A. 1967. A new species of whitefly (Homoptera: Aleyrodidae) from ferns in British greenhouses. 

Proceedings of the Royal Entomological Society of London (B). 36: 30-32. 

Mound, L. A. and S. H. Halsey. 1978. Whitefly of the World. A systematic catalog of the Aleyrodidae 

(Homoptera) with host plant and natural enemy data. British Museum (Natural History)/John Wiley & Sons, 

Chichester. 340 pp. 

Mound, L.A, Martin, J.H. and A. Polaszek. 1994. The insect fauna of Selaginella (Pteridophyta: Lycopsida), 

with descriptions of three new species. Journal of Natural History. 28(6):1403-1415. 

Mulsant, E. 1850. Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et et 

Naturelles d'Agriculture et d'Industrie. 2(2): 1-1104. 

Muma, M. H. 1955. Lady beetles (Coccinellidae: Coleoptera) found on citrus in Florida. Fla. Entomol. 38:117- 

124. 

Muma, M. H. 1957. Effects of larval nutrition on the life cycle, size, coloration, and longevity of Chrysopa 

lateralis Guer. Fla. Entomol. 40:5-9. 

589

Muma, M. H. 1959. Chrysopidae associated with citrus in Florida. Fla. Entomol. 42:21-29. 

Muma, M. H. 1967. Biological notes on Coniopteryx vicina (Neuroptera: Coniopterygidae). Fla. Entomol. 

50:285-293. 

Muma, M. H. 1969. Typhlodromalus peregrinus (Muma)(Acarina: Phytoseiidae) on Florida citrus. In G. O. 

Evans [ed.], Proceedings of the 2nd International Congress of Acarology, Sutton Boninton, UK, 1967, 

Akademiai Kiado, Budapest, Hungary. pp. 135-148. 

Muma, M. H. 1971. Food habits of Phystoseiidae (Acarina: Mesostigmata) including common species on 

Florida citrus. Fla. Entomol. 54:21-34. 

Muma, M. H. 1971a. Biological and behavioral notes on Gasteracantha cancriformis (Arachnidae: Araneidae). 

Fl. Entomol. 54:345-351. 

Muma, M. H. 1975. Spiders in Florida Citrus Groves. Fla. Entomol. 58:83-90. 

Muma, M. H, A. G. Selhime, and H. A. Denmark. 1961. An annotated list of predators and parasites associated 

with insects and mites on Florida citrus. Tech. Bull. 634, Agri. Exp. Sta. Univ. Fla, Gainesville, FL. 

Myartseva, S. V. 2003. New species of the genus Euderomphale Girault, 1916 (Hymenoptera: Eulophidae) - 

parasitoides of whiteflies (Homoptera: Aleyrodidae) from Mexico. Russian Entomol. J. 12(4): 381-384. 

Myartseva, S. V. 2004. New species of the genus Entedononecremnus Girault, 1915 (Hymenoptera: 

Eulophidae) - parasitoids of whiteflies (Homoptera: Aleyrodidae) from Mexico. Russian Entomol. J. 13(1): 77- 

82. 

590

Myartseva, S.N. 1973. Eretmocerus nikolskajae sp.n. (Hym, Chalcidoidea, Aphelinidae) - a new parasite of the 

rose whitefly in Turkmenia. (in Russian). Izvestiya Akademii Nauk Turkmenskoy SSR (Seriya Biologicheskikh 

Nauk). 1973(4):81-85. 

Myartseva, S. N. and J. M. Coronado-Blanco. 2004. A new species of Encarsiella Hayat (Hymenoptera: 

Aphelinidae) with a key to the Mexican species. Folia Entomologica Mexicana. 43(2): 227-232. 

Myartseva, S. N. and C. E. Ruiz. 2002. A new species of Ooencyrtus Ashmead (Hymenoptera: Encyrtidae) 

reared from an aleyrodid (Homoptera) in Mexico. Zoosystematica Rossica. 11: 175-177. 

Myartseva, S. N. & E. Ruiz-Cancino. 2003. Two new species of the genus Metaphycus Mercet (Hymenoptera: 

Encyrtidae), parasitoides of whiteflies (Homoptera: Aleyrodidae) in Mexico. Entomologia Mexicana. 2: 751- 

755. 

Myartseva, S. N. and V. A. Yasnosh. 1994. Parasites of Greenhouse and Cotton Whiteflies (Homoptera: 

Aleyrodidae) in Central Asia. Entomological Review. 73(6): 1-10. 

Nair, R. G, and T. A. Nambiar. 1984. Annual Progress Report 1983. Central Tuber Crop Research Institute, 

Trivandrum, India. 

Nakahara, S. 1989. A new species of Aleurotulus (Homoptera: Aleyrodidae). Proceedings of the Entomological 

Society of Washington. 91 (3): 436-440. 

Nakahara, S. 1995. Taxonomic studies of the genus Tetraleurodes (Homoptera: Aleyrodidae). Insecta Mundi. 9: 

105-150. 

591

592 

Nakao, S. 1962. A list of insects collected in a citrus grove near Fukuoka City (Ecological studies on the insect 

community of citrus groves, IV). Kontyu. 30:50-71. 

Nalepa, C. A. 1996. Release and recovery of the citrus whitefly parasite Encarsia lahorensis (Hymenoptera: 

Aphelinidae) in North Carolina. Florida Entomologist. 79(4): 611-614. 

Naranjo, S, and J. Hagler. 1996. Characterizing and estimating the effect of hereropteran predation. In T. J. 

Henneberry, N. C. Toscano, R. M. Faust, and J. R. Coppedge [eds.], Silverleaf whitefly: 1996 supplement to the 

5-year national research and action plan, USDA/ARS 1996-01. -. 

Nasir, M. M. 1947. Biology of Chrysopa scelestes Banks. Indian J. Entomol. 9:177-192. 

Nasir, M. M. 1947. Chrysopa cymbele Banks and its two new varieties. Indian J. Entomol. 8:119-120. 

Natarajan, K. 1990. Natural enemies of Bemisia tabaci (Gennadius) and effect of insecticide on their activity. J. 

Biol. Control. 4:86-88. 

Natarajan, K, V. T. Sundaramoorthy, T. Surilvelu, A. K. Basu, and A. S. Reddy. 1987. Cotton Whitefly. Central 

Institute for Cotton Research, Coimbatore, Tamilnadu. 

Nawar, M. S, and A. A. El-Sherif. 1993. Neoseiulus cucumeris (Oudemans), a predator of whitefly Bemisia 

tabaci (Gennadius). Bull. Entomol. Soc. Egypt. 71:9-17. 

Nechols, J. R, and D. M. Nafus. 1995. Biological control in the Western United States. Univ. Calif. Publ. 3361, 

Oakland.

593 

Newstead, R. 1908. On the gum-lac insect of Madagascar,a nd other coccids affecting the citrus and tobacco in 

that island. Q. Jl Inst. comml Res. Trop. 3(6): 3-13. 

Newstead, R. 1911. On a collection of Coccidae and Aleurodidae, chiefly African, in the collection of the Berlin 

Zoological Museum. Mitteilungen aus dem Zoologischen Museum in Berlin. 5: 153-174. 

Newstead, R. 1921. A new southern Nigerian Aleurodes (Aleurodidae). Transactions of the Entomological 

Society of London. 54: 528-529. 

Nikol'skaya, M.N and V. A. Yasnosh. 1968. A review of the aphelinid fauna (Hymenoptera, Aphelinidae) of the 

Caucasus. (in Russian). Trudy Vsesoyuznogo Entomologicheskogo Obshchestva. 52:3-42. 

Nikolskaya, M. N. 1950. Representatives of the family Signiphoridae (Hymenoptera: Chalcidoidea) in the fauna 

of the USSR. Doklady Akademii Nauk SSSR. 75: 319-321. 

Noguchi, T. 1943. A list of the natural enemies of the injurious insects of citrus plants. Bull. Shizuoka Citrus 

Exp. Sta. 9:1-30. 

Noshomusho. 1928. Studies on white-flies injurious to the citrus trees in Japan. Byokin-Gaichu-Iho. 16:1-28. 

Nowicki, S. 1929. Bemerkungen zu den europaischen Apheliiden-Gattung (Hym, Chalc). Neue Beitrage zuur 

systematischen Insektenkunde. 4: 153-160. 

Noyes, J. N. 1980. A review of the genera of Neotropical Encyrtidae (Hymenoptera: Chalcidoidea). Bulletin of 

the British Museum (Natural History) (Entomology). 41(3): 107-253.

594 

Noyes, J. N. 2004. Encyrtidae of Costa Rica (Hymenoptera: Chalcidoidea), 2. Metaphycus and related genera, 

parasitoids of scale insects (Coccoidea) and whiteflies (Aleyrodidae). Memoirs of the American Entomological 

Institute. 73: 1-459. 

Noyes, J. S. and P. W. Lozada. 2005. Metaphycus zdeneki sp. nov. (Hymenoptera: Encyrtidae) from Peru, a 

parasitoid of Bakerius sp. (Hemiptera: Aleyrodidae). Acta Soc. Zool. Bohem. 69: 209-214. 

Noyes, J. S. and J. B. Woolley. 1994. North American encyrtid fauna (Hymenoptera: Encytidae): taxonomic 

changes and new taxa. Journal of Natural History. 28: 1327-1401. 

Nunez, E. 1988. Chrysopidae (Neuroptera) del Peru y sus especies mas comunes. Rev. Per. Entomol. 31:69-75. 

Oliveira, M. R. V, E. Amancio, R. Laumann and L. de Gomes. 2003. Natural enemies of Bemisia tabaci 

(Gennadius) B biotype and Trialeurodes vaporariorum (Westwood) (Hemiptera: Aleyrodidae) in Brasilia, 

Brazil. Neotropical Entomology 32(1): 151-154. 

Onillon, J. C, and M. Abbassi. 1973. Notes bio-ecologiques sur l'aleurode floconneux des Aleurothrixus 

floccosus Mask (Homopt, Aleurodidae) et moyens de lutte. Al Awamia. 49:99-117. 

Or, R. 1986. Examination of factors that limit or encourage contamination of the whitefly Bemisia tabaci 

(Gennadius) (Aleyrodidae) on cotton plants in the Bet Shean Valley. M.Sc. Thesis, Tel Aviv Univ. -. 

Or, R, and D. Gerling. 1985. The green lacewing, Chrysoperla carnea, as a predator of Bemisia tabaci. 

Phytoparasitica. 13:75. 

Orian, A. 1972. The Psylloidea of Mauritius with a description of Trioza eastopi sp. nov. Fauna Mauritius. 

Insecta. 1(1): 1-4.

Osborne, L. S, K. Hoelmer, and D. Gerling. 1990. Prospects for biological control of Bemisia tabaci. 

IOBC/WPRS Bull. XIII/. 5:153-160. 

Ossiannilsson, F. 1944. A new whitefly from Sweden, Aleurolobus puripennis n. sp. (Hom.Aleurodidae). K. 

fysiogr. Sallsk. Lund Forh. 14: 186-196. 

Ossiannilsson, F. 1947. Bemisia callunae n. sp. A new Swedish whitefly (Hom. Aleurodidae). Entomologisk 

Tidskrift. 68: 1-3. 

Ossiannilsson, F. 1952. Tetralicia ericae Hesl. Harr. - en for Sverige ny mjollus (Hem. Hom.). Opuscula 

Entomologica, Lund. 17: 80. 

Ossiannilsson, F. 1955. Till kannedomen om de svenska mjollossen (Hem. Hom. Aleyrodina). Opuscula 

Entomologica, Lund. 20: 192-199. 

Ossiannilsson, F. 1966. A new name in the Aleyrodidae (Hem.). Opuscula Entomologica, Lund. 31: 155. 

Pai, K. M, and A. V. Dhuri. 1991. Incidence of insect pest in early variety of cowpea, Vigna unguiculata (L.) 

Walp. Indian J. Entomol. 53:329-331. 

Palmer, J. M, and L. A. Mound. 1990. Predators: Thysanoptera. In D. Rosen [eds.], Armored Scale Insects 

Their Biology Natural Enemies and Control, vol. B. Elseiver, Amsterdam. pp. 67-76. 

Palmer, J. M, L. A. Mound, and G. J. du Heaume. 1989. Thysanoptera. In C. R. Betts [ed.], CIE Guides to 

Insects of Importance to Man 2. C.A.B. International Institute of Entomology, Wallingford, Oxon, UK. 

595

Pappas, S. and G. Viggiani. 1979. Introdotta a Corfe la Prospaltella lahorensis How, (Hym. Aphelinidae), 

parassita del Dialeurodes citri (Ashm.) (Hom. Aleyrodidae). Bollettino del Laboratorio di Entomologia Agraria 

'Filippo Silvestri', Portici. 33:219-231. 

Parrella, M. P, T. S. Bellows Jr, R. J. Gill, J. K. Brown, and K. M. Heinz. 1991. Sweetpotato whitefly: prospects 

for biological control. Calif. Agri. 46:25-26. 

Paulson, G. S, and B. R. Kumashiro. 1985. Hawaiian Aleyrodidae. Proc. Hawaiian Entomol. Soc. 25:103-123. 

Peal, H. W. 1903. Contribution towards a monograph of the oriental Aleurodidae. J. Asiat. Soc. Beng. 72: 61- 

98. 

Peck, O. 1951. Superfamily Chalcidoidea. Pages 410-594. in: Muesebeck, C. F. W., Krombein, K. V. & 

Townes, H. E.,. Hymenoptera of America North of Mexico. Synoptic Catalog. USDA, Agriculture Monograph. 

2: 1-1420. 

Peck, O. 1963. A catalogue of the Nearctic Chalcidoidea (Insecta: Hymenoptera). Canadian Entomologist, 

Supplement. 30: 1092 pp. 

Pedata, P. A. and A. Polaszek. 2003. A revision of the Encarsia longifasciata species group (Hymenoptera: 

Aphelinidae. Systematic Entomology. 28: 361-374. 

Pedata, P.A. and G. Viggiani. 1991. Preliminary morphobiological observations on Azotus perspeciosus 

(Girault) (Hymenoptera: Aphelinidae), hyperparasitoid of Pseudaulacaspis pentagona (Targioni, Tozzetti) 

(Homoptera: Diaspididae). Redia. 74 (3,Appendix):343-350. 

596

597 

Pelov, V, and G. Trenchev. 1973. Siphoninus phillyreae Hal. and its entomophages. Zashchita Rastemii 

(Moskva). 21:26-27. 

Penny, D. D. 1922. A catalog of the California Aleyrodidae and the description of four new species. J. Ent. 

Zool. 14: 21-35. 

Penny, N. D. and J. R. Arias. 1980. Two species of Bakerius from the Amazon basin (Homoptera: Aleyrodidae). 

Acta Amazonica. 10(3): 691-696. 

Penny, N. D. and J. R. Arias. 1980a. A new species of Ceraleurodicus from the Amazon basin (Homoptera: 

Aleyrodidae). Acta Amazonica. 10: 903-905. 

Peracchi. A. L. 1971. Dois aleirodideos pragas de Citrus no Brasil (Homoptera: Aleyrodidae). Archos. Mus. 

Nacional do Rio de Janeiro. 54: 145-151. 

Pettit, F. L. 1992. Biological control in the integrated pest management program at the Land, EPCOT Center. 

Bull. OILB/SROP. 16:2. 

Pickett, C. H, K. A. Hoelmer, and W. J. Roltsch. 1997. Systematics of Delpahstus and biological control of 

whiteflies. In T. J. Hennebery, N. C. Toscano, T. M. Perring, and R. M. Faust [eds.], Silverleaf whitefly: 1997 

supplement to the five-year research and action plan. USDA/ARS 1997-02. pp. 165. 

Pickett, C. H, and D. Kellum. 1996. Release of natural enemies for control of the giant whitefly Aleurodicus 

dugesii, in San Diego County. In L. G. Bezark [ed.], Biological Control Program Annual Summary, 1995. 

California Department of Food and Agriculture, Division of Plant Industry, Sacramento, CA. pp. 58. 

Polaszek, A. 1991. Egg parasitism in Aphelinidae (Hymenoptera: Chalcidoidea) with special reference to

Centrodora and Encarsia species. Bulletin of Entomological Research. 81: 97-106. 

Polaszek, A. 1993. The identity of Neocales Risbec (Hymenoptera: Signiphoridae). Entomologische Berichten, 

Amsterdam. 53(7):99-102. 

Polaszek, A. 1997. Amitus Haldeman (Hymenoptera: Platygasteridae): A genus of whitefly parasitoids new to 

Britain. Entomologist's Monthly Magazine. 1133: 77-79. 

Polaszek, A. unpub. Whiteflies of economic importance and their natural enemies. International Institute of 

Entomology, Natural History Museum. 102 pp. 

Polaszek, A, S. Abd-Rabou and J. Huang. 1999. The Egyptian species of Encarsia (Hymenoptera: Aphelinidae) 

a preliminary review. Zool. Med. Leiden. 73: 131-163. 

Polaszek, A, Evans, G.A. and F. D. Bennett. 1992. Encarsia parasitoids of Bemisia tabaci (Hymenoptera: 

Aphelinidae, Homoptera: Aleyrodidae): a preliminary guide to identification. Bulletin of Entomological 

Research. 82:375-392. 

Polaszek, A. and M. Hayat. 1992. A revision of the genera Dirphys Howard and Encarsiella Hayat 

(Hymenoptera: Aphelinidae). Systematic Entomology. 17:181-197. 

Polaszek, A, S. Manzari & D. Quicke. 2004. Morphological and molecular taxonomic analysis of the Encarsia 

meritoria species-complex (Hymenoptera, Aphelinidae), parasitoids of whiteflies (Hemiptera, Aleyrodidae) of 

economic importance. Zoologica Scripta. 33: 403-421. 

Poorani, J. 2002. An annotated checklist of the Coccinellidae (Coleoptera) (Excluding Epilachninae) of the 

Indian subregion. Oriental Insects. 36: 307-383. 

598

Prasad, V. G. 1955. A second eulophid parasite (Encarsia neomaskelliae n. sp.) on Neomaskellia bergii 

Signoret, on sugar cane in Bihar. Indian Journal of Entomology. 17: 49-51. 

Priesner, H. and M. Hosny. 1932. Contributions to a knowledge of the whiteflies (Aleurodidae) of Egypt (1). 

Bulletin Ministry of Agriculture, Egypt. Technical and Scientific Service. 121: 1-8. 

Priesner, H. and M. Hosny. 1934. Contributions to a knowledge of the whiteflies (Aleurodidae) of Egypt (II). 

Bulletin Ministry of Agriculture, Egypt. Technical and Scientific Service. 139: 1-21. 

Priesner, H. and M. Hosny. 1934a. Contributions to a knowledge of the whiteflies (Aleurodidae) of Egypt (III). 

Bull. Minist. Agric. Egypt tech. scient. Serv. 145: 1-11. 

Priesner, H. and M. Hosny. 1937. A new Trialeurodes on Zizyphus (Hemiptera: Aleurodidae). Bulletin de la 

Societe Entomologique d'Egypte. 21: 45-46. 

Priesner, H, and M. Hosny. 1940. Notes on parasites and predators of Coccidae and Aleurodidae in Egypt. Bull. 

Soc. Fouad 1er d'Ent. 24:58-70. 

Priore, R. 1969. Il Dialeurodes citri (Ashmead) (Homoptera: Aleyrodidae) in Campania (note di morfologia e 

biologia). Boll. Lab. Entomol. Agrar. 'Filippo Silvestri'. 27:286-316. 

Pruszynski, S. 1992. Research on and use of biological methods of pest control in glasshouse crops in Poland. 

Bull. OEPP. 22(3):405-410. 

Pruthi, H. S, and C. K. Samuel. 1942. Entomological investigations on the leaf-curl disease of tobacco in 

599

600 

northern India. V. Biology an population of the white-fly vector (Bemisia tabaci Gen.) in relation to the 

incidence of the disease. Indian J. Agri. Sci. 12:35-57. 

Puttarudriah, M, and G. P. C. Basavanna. 1953. Beneficial Coccinellidae of Mysore India. Indian J. Ent. 15:87- 

96. 

Quaintance, A. L. 1899. New or little known Aleurodidae I. Canadian Entomologist. 31: 1-4. 

Quaintance, A. L. 1899a. New or little known Aleurodidae II. Canadian Entomologist. 31: 89-93. 

Quaintance, A. L. 1900. Contribution towards a monograph of the American Aleurodidae. Technical Series, 

Bureau of Entomology, United States Department of Agriculture. 8: 9-64. 

Quaintance, A. L. 1903. New oriental Aleurodidae. Canadian Entomologist. 35: 61-64. 

Quaintance, A. L. 1907. The more important Aleurodidae infesting economic plants with description of new 

species infesting the orange. Technical Series, Bureau of Entomology, United States Department of Agriculture. 

12: 89-94. 

Quaintance, A. L. 1909. A new genus of Aleyrodidae, with remarks on Aleyrodes nubifera Berger and 

Aleyrodes citri Riley & Howard. Technical Series, Bureau of Entomology, United States Department of 

Agriculture. 12: 169-174. 

Quaintance, A. L. and A. C. Baker. 1913. Classification of the Aleyrodidae Part I. Technical Series, Bureau of 

Entomology, United States Department of Agriculture. 27: 93 pp.

601 

Quaintance, A. L. and A. C. Baker. 1914. Classification of the Aleyrodidae Part II. Technical Series, Bureau of 

Entomology, United States Department of Agriculture. 27: 95-109. 

Quaintance, A. L. and A. C. Baker. 1915. A new genus and species of Aleyrodidae from British Guiana. Annals 

of the Entomological Society of America. 8: 369-371. 

Quaintance, A. L. and A. C. Baker. 1915a. Classification of the Aleyrodidae - Contents and Index. Technical 

Series, Bureau of Entomology, United States Department of Agriculture. 27: 111-114. 

Quaintance, A. L. and A. C. Baker. 1916. Aleurodidae or whiteflies attacking the orange with descriptions of 

three new species of economic importance. Journal of Agricultural Research. 6: 459-472. 

Quaintance, A. L. and A. C. Baker. 1917. A contribution to our knowledge of the whiteflies of the sub-family 

Aleurodicinae (Aleyrodidae). Proceedings of the U. S. National Museum. 51: 335-445. 

Quaintance, A. L. and A. C. Baker. 1937. Notes on some Mexican Aleyrodidae. An. Inst. Biol. Univ. Mex. 8: 

599-629. 

Quaintance, A. L. and A. C. Baker. 1962. A new genus and species of Aleyrodidae from British Guiana. Annals 


Quezada, J. R. 1974. Biological control of Aleurocanthus woglumi (Homoptera: Aleyrodidae) in El Salvador. 

Entomophaga. 19:243-254. 

Quezada, J. R. 1978. El control biologico natural. Un recurso para la agricultura. In Congreso de ingenieros 

agronomos de El Salvador. San Salvador, 1978, San Salvador. pp. 132-138.

Quintela, E. D, S. E. M. Sanchez and M. Yokoyama. 1992. Parasitismo de Encarsia sp. (Hymenoptera: 

Aphelinidae) sobre Bemisia tabaci (Homoptera: Aleyrodidae). An. Soc. Ent. Brasil. 21(3): 471-475. 

Qureshi, J. I. 1970. Dialeurodes dothioensis Dumbleton (Homoptera: Aleyrodidae), a new record from West 

Pakistan. Pakistan Journal of Zoology. 2(1): 114-115. 

Qureshi, J. I. 1979. Genus Singhiella (Aleyrodidae: Homoptera) of Pakistan. Pakistan Entomologist. 1: 13-22. 

Qureshi, J. I. 1980. On a new species of Bemisia (Homoptera: Aleyrodidae) from Pakistan with a key to 

Pakistani species. Oriental Insects. 14(4): 405-407. 

Qureshi, J. I. 1980a. Genus Dialeurodes (Homoptera: Aleyrodidae) of Pakistan. Pakistan Journal of Scientific 

Research. 32(1-2): 62-67. 

Qureshi, J. I. 1982. Aleurotuberculatus (Homoptera: Aleyrodidae) of Pakistan. Pakistan J. Zool. 14(1): 45-57. 

Qureshi, J. I. and M. A. Halimie. 1980. Trialeurodes (Aleyrodidae: Homoptera) of Pakistan. Pakistan Journal of 

Scientific Research. 32(3/4): 232-238. 

Qureshi, J. I. and H. A. Qayyam. 1969. Redescription of the genus Singhiella, and description of a new species, 

Singhiella crenulata (Homoptera: Aleyrodidae) from Lyallpur. Pakistan Journal of Zoology. 1(2): 177-179. 

Rahman, K. A. 1940. Important insect predators of India. Proc. Indian Acad. Sci, Animal Sci. 12:67-74. 

Rao, A. S. 1958. Notes on Indian Aleurodidae (Whiteflies), with special reference to Hyderabad. Proceedings, 

602

X International Congress of Entomology. 1: 331-336. 

Rao, A. S. 1958. Notes on Indian Aleurodidae [sic.] (whiteflies), with special reference to Hyderabad. In 

Proceedings 10th International Congress of Entomology, Montreal, 1956, vol. 1, Montreal, Canada. pp. 331- 

336. 

Rapisarda, C. 1982. Appunti morfologici ed ecologici su alcuni aleirodi (Homoptera: Aleyrodidae) della fauna 

siciliana. Bollettino Del Laboratorio Di Entomologia Agraria Filippo Silvestri. 39: 71-95. 

Rapisarda, C. 1985. Presenza in Italia di Aleurolobus niloticus Priesner & Hosny, nuovo parassita delle piante di 

cappero (Homoptera: Aleyrodidae). Bollettino di Zoologia Agraria e Bachicoltura, Milano. II: 18: 75-86. 

Rapisarda, C. 1986. Trialeurodes (Ericaleyrodes) sardiniae, subgen. n, sp. n. a new heather-feeding whitefly 

(Homoptera: Aleyrodidae). Frustula Entomologica (N.S.). 7-8: 487-499. 

Reddy, A. S, B. Rosaiah, and T. B. Rao. 1985. Control of cotton whitefly in Andhra Pradesh. Indian Farming. 

35:19,21-22. 

Regu, K. and B. V. David. 1990. Rhachisphora elongatus sp. nov (Aleyrodidae: Homoptera) - a new species of 

whitefly from India. Entomon 15(3-4): 277-279. 

Regu, K. and B. V. David. 1991. A new species of Bemisia (Homoptera: Aleyrodidae) from India with a key to 

Indian species. Entomon. 16: 77-81. 

Regu, K. and B. V. David. 1991a. A new species of whitefly Dialeuropora heptapora sp. nov. (Aleyrodidae: 

Homoptera) from India. Journal of the Bombay Natural History Society. 88: 413-414. 

603

Regu, K. and B. V. David. 1992. A new whitefly Aleurotrachelus saklespurensis sp. nov. (Aleyrodidae: 

Homoptera) from India. Entomon. 17(1): 135-136. 

Regu, K. and B. V. David. 1992. On two species of Aleurodicus Douglas (Aleurodicinae: Aleyrodidae: 

Homoptera) from India with a key to the Indian species. Entomon. 17: 99-102. 

Regu, K. and B. V. David. 1993. Taxonomic studies on Indian aleyrodids of the tribe Aleurolobini 

(Aleyrodinae: Aleyrodidae: Homoptera). Frederick Institute of Plant Protection & Technology, Entomology 

Series. 4: 1-17. 

Regu, K. and B. V. David. 1993a. Two new species of whiteflies (Aleyrodidae: Homoptera) from India. 

Hexapoda. 5(1): 53-56. 

Regu, K. and B. V. David. 1993b. Asialeyrodes saklespurensis sp. nov. (Aleyrodidae: Homoptera) from India. 

Entomon 18(1): 91-93. 

Regu, K. and B. V. David. 1993c. On three new species of whiteflies of the tribe Dialeurodine Sampson, 1943 

(Aleyrodidae: Homoptera) from India. Journal of the Bombay Natural History Society. 89: 82-87. 

Reyes, M, A. J. Abud, M. Sencion, E. Estevez, J. Dupuy, and F. A. Barreiro. 1989. Incidencia de Bemisia tabaci 

Gennadius (Homoptera: Aleyrodidae) y sus enemigos naturales en el cultivo del tomate Lycopersicum 

esculentum Mill, asociado con otros cultivos en el Valle de Azua,. In Colloques de L'INRA, No. 58. Caribbean 

meetings on biological control. Guadeloupe, France. Nov. 5-7, 1990, 58th ed. INRA, Guadeloupe, France. pp. 

471-477. 

Ricci, C, and G. Cappelletti. 1988. Relationship between some morphological structures and locomotion of 

604

605 

Clitostethus arcutatus Rossi (Coleoptera Coccinellidae), a whitefly predator. Frustula Entomol. 11:195-202. 

Richardson, H. H. 1948. Present status of the citrus blackfly and its parasite Eretmocerus serius at Nassau, 

Bahamas. J. Econ. Entomol. 41:980. 

Rietschel, P. 1983. Aleurochiton petri n. sp, eine Mottenschildlaus (Homoptera, Aleyrodina) aus dem Pliozan 

von Neu-Isenburg, Hessen. Carolinea. 41: 97-100. 

Riley, C. V. and L. O. Howard. 1893. The orange Aleyrodes (Aleyrodes citri n. sp.). Insect Life. 5: 219-226. 

Ripolles, J. L, and A. Melia. 1982. Primeras observaciones sobre la proliferacion de Conwentzia psoiformis 

(Curt.) (Neuroptera: Coniopterygidae), en los citricos de Castellon de la Planta. Rev. App. Entomol. 70:5361. 

Risbec, J. 1950. La faune entomologique des cultures au Senegal et au Soudan Francaise II. Contribution 

al'etude des Proctotrupidae. Travaux du Laboratoire d'Entomologie Francais d'Afrique Noire. 2: 509-640. 

Risbec, J. 1951. Lec Chalcidoides de l'Afrique occidentale Francaise. Memoires del'Institute Francais d'Afrique 

Noire. 13: 7-409. 

Risbec, J. 1957. Aphelinidae de Madagascar (Hym, Chalcidoidea). Le Naturaliste Malgache. 9(2): 263-272. 

Ritchie, A. H. 1917. Annual report of the entomologist. Jamaica Annual Report of the Department of 

Agriculture. 28-34. 

Riudavets, J, R. Gabarra, and D. Castane. 1993. Frankliniella occidentalis predation by native natural enemies. 

IOBC/WPRS Bull. 16(2):137-140.

Rivnay, T. and D. Gerling. 1987. Aphelinidae parasitoids (Hymenoptera: Chalcidoidea) of whiteflies 

(Hemiptera: Aleyrodidae) in Israel, with description of three new species. Entomophaga. 32: 463-475. 

Roltsch, W. J, and C. H. Pickett. 1994. Areawide establishment of Delphastus pusillus, a predator of silverleaf 

whitefly, in the Imperial Valley. In T. J. Henneberry, N. C. Toscano, R. M. Faust, and J. R. Coppedge [eds.], 

Sweetpotato whitefly: 1994 Supplement to the five-year national research and action plan. USDA/ARS, 

Springfield, VA. pp. 103. 

Rose, M. 2000. Eretmocerus Haldeman (Hymenoptera: Aphelinidae) reared from Aleurothrixus floccosus 

(Maskell) (Homoptera: Aleyrodidae) in the Americas. Vedalia. 

Rose, M, P. DeBach, and J. Woolley. 1981. Potential new citrus pest: Japanese bayberry whitefly. Calif. Agri. 

:22-24. 

Rose, M. and D. Rosen. 1992. Eretmocerus debachi n.sp. (Hymenoptera: Aphelinidae), an effective parasite of 

Parabemisia myricae (Homoptera: Aleyrodidae). Israel. Journal of Entomology XXV-XXVI:199-207. 

Rose, M. and G. Zolnerowich. 1994. Eretmocerus furuhashii sp. n. (Hymenoptera: Aphelinidae) a parasite of 

Parabemisia myricae (Kuwana) (Homoptera: Aleyrodidae) in Japan. Japanese Journal of Entomology. 62(2): 

285-292. 

Rose, M. and G. Zolnerowich. 1997. Eretmocerus Haldeman (Hymenoptera: Aphelinidae) in the United States, 

with descriptions of new species attacking Bemisia (tabaci complex) (Homoptera: Aleyrodidae). Proceedings of 

the Entomological Society of Washington. 99(1), January 1997: 1 27, illustr 

Rose, M. and G. Zolnerowich. 2003. Eretmocerus picketti new species (Hymenoptera: Chalcidoidea: 

606

607 

Aphelinidae) reared from Tetraleurodes acaciae (Quaintance) (Homoptera: Aleyrodidae: Aleyrodinae). Pan- 

Pacific Entomologist. 79(2): 119-127. 

Rose, M. & G. Zolnerowich. 2004. Eretmocerus perseae n. sp. (Hymenoptera: Chalcidoidea: Aphelinidae) 

reared from Tetraleurodes perseae Nakahara (Homoptera: Aleyrodidae: Aleyrodinae) living on avocado in 

Michoacan, Mexico. Vedalia. 11: 45-52. 

Rosen, D. 1966. Notes on the parasites of Acaudeleurodes citri (Priesner & Hosni) (Hem.: Aleurodidae) in 

Israel. Entomologische Berichten, Amsterdam. 26:55-59. 

Rozanov, L. V. 1965. Review of the genera of parasitic Hymenoptera of the family Signiphoridiae 

(Hymenoptera: Chalcidoidea). Entomological Review. 44: 508-515. 

Ruiz, M. E. R. 1993. A New Predator on the Whitefly. Southwestern Entomol. 18:147-148. 

Russell, L. M. 1943. A new genus and four new species of whiteflies from the West Indies (Homoptera: 

Aleyrodidae). Proceedings of the Entomological Society of Washington. 45: 131-141. 

Russell, L. M. 1944. Descriptions of nine species of Aleuroplatus from eastern North America (Homoptera: 

Aleyrodidae). Journal of the Washington Academy of Sciences. 34: 333-341. 

Russell, L. M. 1944a. A taxonomic study of the genus Aleuroglandulus Bondar (Homoptera: Aleyrodidae). 

Proceedings of the Entomological Society of Washington. 46: 1-9. 

Russell, L. M. 1945. A new genus and twelve new species of Neotropical whiteflies (Homoptera: Aleyrodidae). 

Journal of the Washington Academy of Sciences. 35: 55-65.

Russell, L. M. 1947. A classification of the whiteflies of the new tribe Trialeurodini (Homoptera: Aleyrodidae). 

Revista de Entomologia, Rio de Janeiro. 18: 1-44. 

Russell, L. M. 1948. The North American species of whiteflies of the genus Trialeurodes. Miscellaneous 

Publication of the USDA. 635: 1-85. 

Russell, L. M. 1957. Synonyms of Bemisia tabaci (Gennadius) (Homoptera: Aleyrodidae). Bulletin of the 

Brooklyn Entomological Society. 52: 122-123. 

Russell, L. M. 1958. Orchamoplatus. An Australasian genus (Homoptera: Aleyrodidae). Proceedings of the 

Hawaiian Entomological Society. 16: 389-410. 

Russell, L. M. 1959. New name combinations in a list of the species of Dialeuropora Quaintance & Baker 

(Homoptera: Aleyrodidae). Proceedings of the Entomological Society of Washington. 61: 185-186. 

Russell, L. M. 1960. A taxonomic study of the genus Corbettia Dozier (Homoptera: Aleyrodidae). Revue de 

Zoologie et de Botanique Africaines. 62: 120-137. 

Russell, L. M. 1960a. A whitefly living on roses (Homoptera: Aleyrodidae). Proceedings of the Royal 

Entomological Society of London (B). 29: 29-32. 

Russell, L. M. 1962. New name combinations and notes on some African and Asian species of Aleyrodidae 

(Homoptera). Bulletin of the Brooklyn Entomological Society. 57: 63-65. 

Russell, L. M. 1962a The citrus blackfly. Plant Protection Bulletin, F.A.O. 10: 36-38. 

608

Russell, L. M. 1963. Hosts and distribution of five species of Trialeurodes (Homoptera: Aleyrodidae). Annals 


Russell, L. M. 1964. A new species of Aleurocybotus (Homoptera: Aleyrodidae). Proceedings of the 


Russell, L. M. 1964a. Dialeurodes kirkaldyi (Kotinsky), a whitefly new to the United States (Homoptera: 

Aleyrodidae). Florida Entomologist. 47: 1-4. 

Russell, L. M. 1965. A new species of Aleurodicus Douglas and two close relatives (Homoptera: Aleyrodidae). 

Florida Entomologist. 48: 47-55. 

Russell, L. M. 1967. Venezaleurodes pisoniae, a new genus and species of whitefly from Venezuela 

(Homoptera: Aleyrodidae). Florida Entomologist. 50: 235-241. 

Russell, L. M. 1978. The whitefly genus Aleurocerus Bondar (Hemiptera, Homoptera, Aleyrodidae). 

Entomography. 4: 137-183. 

Russell, L. M. 1986. The whitefly genus Aleurocerus (Hemiptera, Homoptera, Aleyrodidae). Entomography. 4: 

137-183. 

Russell, L. M. 1993. A new species of Trialeurodes (Homoptera: Aleyrodidae) from Phlox. Proceedings of the 

Entomological Society of Washington. 95(4): 583-586. 

Russell, L. M. 1993a. Description of Disiphon, new genus, and its type species Asterochiton dubienus Bondar 

609

610 

(Homoptera, Aleyrodidae). Proceeding of the Entomological Society of Washington. 93(3): 463-467. 

Russell, L. M. 1994. Notes on the genus Paraleurolobus: redescription of the genus and its type species and 

description of a new species (Homoptera, Aleyrodidae). Proceedings of the Entomological Society of 

Washington. 96(3): 537-547. 

Russell, L. M. 2000. Notes on the family Aleyrodidae and its subfamilies: redescription of the genus 

Aleurocybotus Quaintance and Baker and description of Vasdavidius, a new genus (Homoptera: Aleyrodidae). 


Russell, L. M, and J. Etienne. 1985. A list of the Aleyrodidae of the Island of Reunion. Proc. Entomol. Soc. 

Wash. 87:202-206. 

Ryberg, O. 1938. Bidrag till kannedomen om de nordiska mjollossen, Aleurodidae (Hem. Hom.) jamte 

provisorisk katalog over de europeiska arternas vardvaxter. Kungl. Fysiografiska Sallskapets I Lund 

Forhandlingar. 8: 10-25. 

Rychewaert, P. and C. Alauzet. 2002. The natural enemies of Bemisia argentifolii in Martinique. Biocontrol. 47: 

115-126. 

Saalas, U. 1942. Eine neue Mottenlaus, Aleurodes campanulae n. sp. (Hem. Aleurodidae) an Camanula. 

Suomen hyont. Aikak. 8: 127-134. 

Sampson, A. C, and V. J. King. 1996. Macrolophus caliginosus, field establishment and pest control effect in 

tomatoes. Bull. OILB/SROP. 19:143-146. 

Sampson, W. W. 1943. A generic synopsis of the Hemipterous Superfamily Aleyrodoidea. Entomologica 

Americana. 23: 173-223.

Sampson, W. W. 1944. Additions to the Aleyrodidae of Mexico (Hem. Hom.). An. Esc. nac. Cienc. biol. Mex. 

3: 437-444. 

Sampson, W. W. 1945. Five new species of Aleyrodidae from California (Homoptera). Pan-Pacific 

Entomologist. 21: 58-62. 

Sampson, W. W. and E. A. Drews. 1940. Gymnaleurodes, a new genus of Aleyrodidae from California 

(Homoptera). Pan-Pacific Entomologist. 16: 29-30. 

Sampson, W. W. and E. A. Drews. 1941. Fauna Mexicana IV. A review of the Aleyrodidae of Mexico (Insecta, 

Homoptera). Anales Nacional Ciencias Biologicas. 2(2-3): 143-189. 

Sampson, W. W. and E. A. Drews. 1956. Keys to the genera of the Aleyrodidae and notes on certain genera. 

(Homoptera: Aleyrodidae). Annals and Magazine of Natural History (12). 9: 689-697. 

Sampson, W. W. and E. A. Drews. 1999. Keys to the genera of the Aleyrodidae and notes on certain genera. 

(Homoptera: Aleyrodidae). Annals and Magazine of Natural History (12). 9: 689-697. 

Sasaji, H. 1971. Fauna Japonica. Coccinellidae (Insecta: Coleoptera). Toyko, 345 pp. -. -. 

Sasaki, C. 1908. Aleurodes kuchinasii. [In Japanese]. Transactions of the Entomological Society of Japan. 2(3): 

55-56. 

Satpute, P. P., Y. M. Taley, S. A. Nimblkar and N. R. Kadu. 1986. Natural enemies of citrus black fly, 

Aleurocanthus woglumi Ashby, in Nagpur region. Bull. Entomol. 2:128-131. 

611

Schaefer, P. W, S. E. Barth, and H. B. White. 1996. Predation by Enallagma civile (Odonata: Coenagrionidae) 

on adult sweetpotato whitefly, Bemisia tabaci (Homoptera: Aleyrodidae). Entomological News. 107:275-276. 

Schauff, M.E., Evans, G.A. and J. M. Heraty. 1996. A pictorial guide to the species of Encarsia (Hymenoptera: 

Aphelinidae) parasitic on whiteflies (Homoptera: Aleyrodidae) in North America. Proceedings of the 

Entomological Society of Washington. 98(1):1-35. 

Schilder, F. A, and M. Schilder. 1928. Die Nahrung der Coccinelliden und ihre beziehung zur Verwandtschaft 

der Arten. Arb. Biol. BundAnst. Land-u. Forstw. 16:213-282. -. 

Schmidt, S., I. D. Naumann and P. J. Barro. 2001. Encarsia species (Hymenoptera: Aphelinidae) of Australia 

and the Pacific Islands attacking Bemisia tabaci and Trialeurodes vaporariorum (Hemiptera: Aleyrodidae) - a 

pictorial key and descriptions of four new species. Bulletin of Entomological Research. 91: 369-387. 

Schmiedeknecht, O. 1909. Hymenoptera f_r Chalcididae. Wytsman Genera Insectorum. 97: 1-550, 8 pls. 

Schrank, F. von P. 1801. Fauan Bioca. -. 2(1): 274 pp. Ingolstadt. 

Seguy, E. 1934. DiptSres (BrachycSres). Faune France. 28:371. 

Selhime, A, M. Muma, and D. W. Clancy. 1963. Biological, chemical and ecological studies on the predatory 

thrips Aleurodothrips faciapennis in Florida citrus groves. Ann. Entomol. Soc. Am. 56:709-712. 

Serra, C. A. 1992. Practical experiences in the use of seed extracts of the neem tree backing bioregulation of 

important pests in Dominican tomato fields. In U. Koepke and D. G. Schulz [eds.], Proceedings 9th IFOAM 

612

conf. IFOAM, Sao Paulo, Brazil. pp. 173-180. 

Serra, C. A. 1992a. Untersuchungen zum Einsatz von Niemsamenextrakten im Rahmen integrieter Ansaetze zur 

Bekaempfung von Tomatenschaedlingen in der Dominikanischen Repiblik. Ph.D. Dissertation, Universidad de 

Giessen, Wissenschaftlicher Fachverlag: Giessen. -. 

Serra, C. A., M. Ortiz, J. B. Nupez, and P. F. Benoit. 1996. Survey of indigenous natural enemies of whiteflies 

in the Dominican Republic. In T. J. Henneberry, N. C. Toscano, R. M. Faust, and J. R. Coppedge [eds.], 

Silverleaf whitefly: 1996 supplement to the 5-year national research and action plan, USDA/ARS 1996-01. pp. 

133. 

Serrano, L. 1978. Identification, multiplication y liberacion de parasitoides himenopteros del picudo el 

algodonero Anthonomus grandis Boheman. Ing. Agr. Thesis, Univ. de El Salvador. -. 

Serrano, L., J. M. Sermeno, and J. F. Larios. 1993. Las moscas blancas en el Salvador. In L. Hilje and O. 

Arboleda [eds.], Serie Tpcnica, Informe Tpcnico No. 205: Las Moscas Blancas (Homoptera: Aleyrodidae) en 

Amprica Central Y El Caribe. CATIE, Turrialba, C. 

Shafee, S. A. 1973. Indian species of the genus Prospaltella Ashm. (Hym.: Aphelinidae). Entomophaga. 

18(3):251-258. 

Shafee, S. A. 1974. Species of the genera Promuscidea Girault and Prospaltella Ashmead (Hymenoptera: 

Aphelinidae) from India. Entomophaga. 19(1):33-39. 

Shafee, S. A. and D. Bela. 1984. A new species of Encarsia (Hym: Aphelinidae) from Budapest, Hungary. 

Indian Journal of Systematic Entomology. 1: 29-30. 

613

614 

Shafee, S.A. and S. Rizvi. 1982. A new species of Encarsia (Hymenoptera: Aphelinidae) from Pakistan. Journal 

of Entomological Research. 6(2):157-158. 

Shafee, S.A. and S. Rizvi. 1984. Taxonomic notes on some Indian Aphelinidae (Hymenoptera: Chalcidoidea). 

Mitteilungen der Schweizerischen Entomologischen Gesellschaft. 57:379-381. 

Shafee, S.A., Rizvi, S. and M. Y. Khan. 1988. A new tribe and three new species of Eulophidae (Hymenoptera: 

Chalcidoidea). Indian Journal of Systematic Entomology. 5(1):1-4. 

Shah, A. H., M. B. Patel, and G. M. Patel. 1986. Record of a coccinellid predator, Serangium parcesetosum Sic. 

of sugarcane whitefly in South Gujarat. Gujarat Agri. Univ. Res. J. 12:63-64. 

Sharkov, A. V. and N. D. Voynovich. 1988. A new genus of encyrtids (Hymenoptera, Encyrtidae) from 

northern Karelia and Finland. Entomologicheskoe Obozrenie. 67: 826-830. 

Shcherbakov, D. E. 2000. The most primitive whiteflies (Hemiptera; Aleyrodidae; Bernaeinae subfam. nov.) 

from the Mesozoic of Asia and Burmese amber, with an overview of Burmese amber hemipterans. Bulletin of 

the Natural History Museum London (Geology). 56: 29-37. 

Shenderovskaya, L. P. 1976. Introduced insect enemies and microorganisms. Zashchita Rastenii (Moskva). 

3:52-53. 

Shimer, H. 1867. Description of a new species of Aleyrodes. Transactions of the American Entomological 

Society. 1: 281. 

Shiraki, T. 1913. Researches concerning insects pests in Formosa. [in Japanese]. Spec. Rep. Formosa Agric. 

Exp. Station. 8: 104-110.

Signoret, V. 1868. Essai monographique sur les aleurodes. Annales de la Societe Entomologique de France. 

(4)8:369-402. 

Signoret, V. 1882. Seance du 14 december 1881, 4o note. Annales de la Societe Entomologique de France. 6(1) 

CLVIII. 

Signoret, V. 1883. Seance du 23 mai 1883. 1o Note. Annales de la Societe Entomologique de France. 6(3): 

LXIII. 

Silvestri, F. 1911. Di una nuova specie di Aleurodes vivente sull'olivo. Bollettino del Laboratorio di Zoologia 

Generale e Agraria della R. Scuola Superiore d'Agricoltura, Portici. 5: 215-225. 

Silvestri, F. 1915a. Contributo all conoscenza degli insetti dell'olivo dell'Eritrea e dell'Africa meridionale. (in 

Italian). Bollettino del Laboratorio di Zoologia Generale e Agraria della R. Scuola Superiore d'Agricoltura, 

Portici. 9:240-334. 

Silvestri, F. 1915b. Descrizione di nueovi Imenotteri Calcididi africani. Bollettino del Laboratorio di Zoologia 

Generale e Agraria, Portici. 9: 337-377. 

Silvestri, F. 1926. Descrizione di tre di Prospaltella e di un di Encarsia parassite di Aleurocanthus 

(Aleyrodidae). Eos. Revista Espanola di Entomologia. Madrid. 2:179-189. 

Silvestri, F. 1926a. Lotta contro alcune cocciniglie degli agrumi. (in Italian). Nuovi Annali dell'Agricoltura, 

Roma. 6:97-101. 

615

616 

Silvestri, F. 1927. Contribuzione all conoscenza degli Aleurodidae (Insecta: Hemiptera) viventi su Citrus in 

Estremo Oriente e dei loro parassiti. Bollettino del Laboratorio di Zoologia Generale e Agraria della R. Scuola 

Superiore d'Agricoltura, Portici. 21: 1-60. 

Silvestri, F. 1928. Contribuzione alla conoscenza degli Aleurodidae (Insecta: Hymenoptera) viventi su Citrus in 

estremo Oriente e die loro parassiti. II Descrizione e notizie biologische dei parassitidi Aleurodidi vivienti si 

Citrus. Bollettino del Laboratorio di Zoologia Generale e Agraria, Portici. 21: 20-60(1927). 

Silvestri, F. 1931. Descrizione di una nouva specie di Prospaltella della Somalia. Bollettino della Societa 

Entomologica Italiana. 63: 20-22. 

Silvestri, F. 1934. Compendio di Entomologia Applicata. -. 1(1): 448-899. 

Silvestri, F. and R. G. Mercet. 1928. Afelinidos palearticos. Boletin de la Real Sociedad Espanola de Historia 

Natural. 28:289-294. 

Simonishvili, R. A. 1976. Trophic relations of Amblyseius largoensis (Muma) (Phytoseiidae, Gamasoidae). 

Sybtropicheskie Kul'tury. 3/4:200-202. 

Singh, K. 1931. A contribution towards our knowledge of the Aleyrodidae (Whiteflies) of India. Memoirs of the 

Department of Agriculture in India. 12: 1-98. 

Singh, K. 1932. On some new Rhynchota of the family Aleyrodidae from Burma. Rec. Indian Mus. 34: 81-88. 

Singh, K. 1933. On four new Rhynchota of the family Aleurodidae from Burma. Rec. Indian Mus. 35: 343-346.

Singh, K. 1938. Notes on Aleurodidae (Rhynchota) from India I. Rec. Indian Mus. 40: 189-192. 

Singh, K. 1940. Notes on Aleurodidae (Rhynchota) from India II. Rec. Indian Mus. 42: 453-456. 

Singh, K. 1945. Notes on Aleurodidae (Rhynchota) from India III. Indian Journal of Entomology. 6: 75-78. 

Smith, H. A., G. A. Evans and R. McSorley. 2000. A survey of parasitoids of Trialeurodes vaporariorum and 

Bemisia tabaci in Eastern Guatemala. Florida Entomologist. 83(1): 492-496. 

Smith, H. and G. Compere. 1928. Black scale Saissetia oleae. University of California Publications in 

Entomology. 4: 314. 

Smith, H. D., H. L. Maltby and J. E. Jimenez. 1964. Biological control of the citrus blackfly in Mexico. 

Technical Bulletin 1311, USDA, Washington, DC. 

Soliman, Z. R., K. K. Shemata and E. A. Gomaa. 1976. On the food range and economic importance of the 

predatory of the predatory mite Agistemus exsertus Gonzalez. (Acari, Prostigmata). Anzeiger fur 

Schadlingskunde, Pflanzenschutz, Umweltschutz. 49:87-90. 

Solomon, M. E. 1935. On a new genus and two new species of Western Australian Aleyrodidae. Proceedings of 

the Royal Society of Western Australia. 21: 75-91. 

Sorauer, P. 1956. Handbuck der Pflanzenkrankheiten. -. 5(2): 399 pp Berlin & Hamburg. 

Southwood, T. R. E, and P. M. Reader. 1988. The impact of predation on the viburnum whitefly, 

617

Aeurotrachelus jelinekii. Oecologia. 74:566-570. 

Southwood, T. R. E, and P. M. Reader. 1988. The impact of predation on the viburnum whitefly, 

Aleurotrachelus jelinekii. Oecologia. 74:566-570. 

Souza, B., L. V. C. Santacecilia, and C. F. Carvalho. 1996. Selectivity of some insecticides and acaricides to 

eggs and larvae of Ceraeochrysa cubana (Hagen, 1861) (Neuroptera: Chrysopidae) in laboratory. Pesquisa 

Agropecuaria Brasileira. 31:775-779. 

Soylu, O. Z. and Urel 1977. 1977. Investigations on parasites and predators of the pests that cause damage on 

citrus plantations in the Anatolia region. Bitki Koruma Bpl. (Plant Protection Bull.). 17:77-122. 

Soylu, O. Z. 1980. Investigations on the biology and control of citrus whitefly Daleurodes citri (Ashmead) 

injurious in citrus orchards in the Mediterranean region of Turkey. Bitki Koruma Bpl. (Plant Protection Bull.). 

20:36-53. 

Soylu, O. Z. and N. Urel. 1977. Investigations on parasites and predators of the pests that cause damage on 

citrus plantations in the Anatolia region. Bitki Koruma Bpl. (Plant Protection Bull.). 17:77-122. 

Stam, P. A. and A. Tune. 1983. Prospects for integrated pest control on cotton in the Cukarova region of 

Turkey. Technical Report, FAO/TUR/83/003, FAO, Rome. -. 

Stephens, J. F. 1829. A Systematic Catalogue of British Insects. Part II. -. 369 pp. London. 

Streito, J. C., J. Ollivier and B. Ollivier. 2004. Deaux ravageurs nouveaux cocotier (Cocos nucifera) pour le 

fune des Comores: Aleurotrachelus atratus Hempel 1922, et Paraleyrodes bondari Perrachi 1971 (Hemiptera: 

Aleyrodidae). Bulletin de la Societe Entomologie de France. 109(1): 67-72. 

618

Subba Rao, B. R. 1984. Descriptions of new species of Oriental Mymaridae and Aphelinidae (Hymenoptera: 

Chalcidoidea). Proceedings of the Indian Academy of Sciences (Animal Sciences). 93: 251-262. 

Subba Rao, B.R. 1974. The genera of Signiphoridae (Hymenoptera). Proceedings of the Indian Academy of 

Sciences (B). 

Sugonjaev, E. S. 1960. On the species of the genera allied to Aphycus Mayr (Hymenoptera: Encyrtidae) from 

the european part of the USSR. Entomologicheskoe Obozrenie. 39: 364-383. 

Suh, S, G. Hodges & M. Lee. 2006. Key to Korean species of whiteflies (Hemiptera: Aleyrodidae). 

Suh, S, G. Hodges, M. Lee and G. A. Evans. 2006. Key to the Korean species of whiteflies (Hemiptera: 

Aleyrodidae). 

Sundararaj, R. 1999. Redescription and a new host record for Aleuroclava jasmini (Takahashi) from India. 

Indian Journal of Entomology. 61(2): 192-194. 

Sundararaj, R. 2001. Description of a New Species of Dialeurolonga Dozier (Hemiptera: Aleyrodidae) 

Breeding on Polyalthia longifolia Hook with Key to Indian species. Entomon, 26(2): 191-194. 

Sundararaj, R. and B. V. David. 1990. A new whitefly Bemisia multituberculata sp. nov. (Aleyrodidae: 

Homoptera) from India. Entomon. 15 (1): 113-115. 

Sundararaj, R. and B. V. David. 1991. On the whiteflies of the genus Rhachisphora Quaintance and Baker 

(Aleyrodidae: Homoptera) from India. Entomon. 16 (4): 311-315. 

Sundararaj, R. and B. V. David. 1991a. Ten new species of Dialeuronomada Quaintance & Baker from Indian 

subcontinent. Hexapoda. 3: 27-47. 

619

620 

Sundararaj, R. and B. V. David. 1992. On the genera Asialeyrodes Corbett and Cockerelliella gen. nov. from 

Indian. Journal, Bombay Natural History Society. 88: 415-424. 

Sundararaj, R. and B. V. David. 1992a. On the genera Fippataleyrodes n. gen. and Taiwanaleyrodes Takahashi 

from India. Reichenbachia. 29(4): 15-18. 

Sundararaj, R. and B. V. David. 1992b. On a new species of Singhius Takahashi (Aleyrodidae: Homoptera) 

with a key to Indian species. Journal of the Bombay Natural History Society. 89: 103-105. 

Sundararaj, R. and B. V. David. 1993. New species of Aleuroclava Singh from India (Homoptera: Aleyrodidae). 


Sundararaj, R. and B. V. David. 1993a. First record of the whitefly genus Martiniella Alexander & David 

(Aleyrodidae: Homoptera) from India. Entomon. 18 (1): 95-99. 

Sundararaj, R. and B. V. David. 1994. On the Indian species of the genus Gigaleurodes Quaintance and Baker 

(Aleyrodidae: Homoptera) from India. Journal of the Bombay Natural History Society. 91: 328-330. 

Sundararaj, R. and B. V. David. 1995. Aleuroclava afriae, a new whitefly species from India (Insecta: 

Homoptera: Sternorrhyncha: Aleyrodidae). Reichenbachia. 31 (4): 17-18. 

Sundararaj, R and B. V. David. 1998. On the genus Rabdostigma Quaintance & Baker (Aleyrodidae: 

Homoptera) from India. Journal of Entomological Research. 22(4), December, 1998. 

Sundararaj, R. and B. V. David. 2003. Massilieurodes homonoiae (Jesudasan and David) comb. nov. 

(Aleyrodidae: Homoptera). Entomon. 28(4): 371-372. 

Sundararaj, R. and A. K. Dubey. 2003. Whiteflies (Hemiptera: Aleyrodidae) associated with sandal (Santalum 

album L.) in southern India with description of a new species. Entomon. 28(4): 293-298. 

Sundararaj, R. and A. K. Dubey. 2004. The whitefly genus Martiniella Jesudasan and David (Aleyrodidae: 

Homoptera), with description of one new species from India. Entomon. 29 (4): 357-360. 

Sundararaj, R and A. K. Dubey. 2005. Parabemisia myricae (Kuwana) (Hemiptera: Aleyrodidae), a new record 

from India. Zoos' Print Journal. 20(7): 1933. 

Sundararaj, R. and A. K. Dubey. 2006. Description of five new species of the whitefly genus Dialeurolonga

Dozier (Hemiptera: Aleyrodidae) from India. Oriental Insects 40: 162. 

Sundararaj, R. and A. K. Dubey 2006a. A review of the whitefly genus Dialeurodes Cockerell (Aleyrodidae: 

Hemiptera) with descriptions of two new species from India. Journal of the Bombay Natural History Society. 

103(1): 62-67. 

Sundararaj, R. and A. K. Dubey 2006b. On the genus Rhachisphora Quaintance & Baker (Hemiptera: 

Aleyrodidae) of India with description of a new species. Journal of the Bombay Natural History Society. 

103(1): 68-70. 

Sussman, I. 1988. The cotton insects of Israel and aspects of the biology of Deraeocoris pallens Reuter 

(Heteroptera: Miridae). M.S. Thesis, Tel Aviv Univ, Tel Aviv. -. 

Suta, A. R, and N. M. Esguerra. 1993. Recent history of biological control in the freely associated states of 

Micronesia. In D. F. Waterhouse and A. Vargo [eds.], Micronesia, Supp. 4: Biological control of exotic pests in 

the Pacific. Proceedings of a Plenary Session and Symposium XIX International Congress of Entomology, 

Beijing, 19. 

Swezey, S. L, and E. Cano Vasquez. 1991. Biological control of citrus blackfly (Homoptera: Aleyrodidae) in 

Nicaragua. Environ. Entomol. 20:1691-1698. 

Swirski, E, S. Amitai, and N. Dorzia. 1967. Laboratory studies on the feeding, development and reproduction of 

the predacious mite Typhlodromus athiasae P. and S. (Acarina, Phytoseiidae) on various kinds of food 

substances. Israel J. Agri. Res. 17:213-218. 

Swirski, E, S. Amitai, and N. Dorzia. 1967. Laboratory studies on the feeding, development and reproduction of 

the predacious mites Amblyseius rubini Swirski and Amitai and Amblyseius swirskii Athisa (Acarina, 

Phytoseiidae) on various kinds of food substances. Israel J. Agri. Res. 17:101-119. 

Swirski, E, S. Amitai, and N. Dorzia. 1970. Laboratory studies on the feeding habits, post embryonic survival 

and oviposition of the predaceous mites Amblyseius chilenensis Dosse and Amblyseius habisci Chant (Acarina: 

Phytoseiidae) on various kinds of food substances. Entomophaga. 15:93-106. 

Swirski, E, D., Blumberg, M. Wysoki, and Y. Izhar. 1985. Data on the phenology and biological control of the 

Japanese bayberry whitefly, Parabemisia myricae, in Israel. Phytoparasitica. 13:73. 

Swirski, E, D., Blumberg, M. Wysoki, and Y. Izhar. 1987. Biological control of the Japanese bayberry whitefly, 

Parabemisia mycricae (Kuwana) (Homoptera: Aleyrodidae), in Israel. Israel J. Entomol. 21:11-18. 

621

Swirski, E. and N. Dorzia. 1968. Studies of the feeding, development and oviposition of the predacious mite 

Amblyseius limonicus Garman and McGregor (Acarina: Phytoseiidae) on various kinds of food substances. 

Israel J. Agri. Res. 18:71-75. 

Swirski, E. and N. Dorzia. 1969. Laboratory studies on the feeding, development, and fecundity of the 

predaceous mite Typhlodromus occidentalis Nesbitt (Acarina: Phytoseiidae) on various kinds of food 

substances. Israel J. Agri. Res. 19:143-145. 

Swirski, E., M. Wysoki, and Y. Izhar. 1988. Integrated pest management in the avocado orchards of Israel. 

Appl. Agric. Res. 3:1-7. 

Szelegiewicz, H. 1972. Notatki faunistyczne of maczlikach (Homoptera: Aleyrodidae) Polski. Fragm. faun. 18: 

25-30. 

Takahashi, R. 1931a. Some Formosan whiteflies. J. Soc. trop. Agric. Taiwan. 3: 218-224. 

Takahashi, R. 1931b. Some white-flies of Formosa (Part I.). Trans. nat. Hist. Soc. Formosa. 21: 203-209. 

Takahashi, R. 1931c. Some white-flies of Formosa (Part II.). Trans. nat. Hist. Soc. Formosa. 21: 261-265. 

Takahashi, R. 1932. Aleyrodidae of Formosa, Part I. Report. Department of Agriculture. Government Research 

Institute. Formosa. 59: 1-57. 

Takahashi, R. 1933a. Aleyrodidae of Formosa, Part I. Report. Department of Agriculture. Government Research 

Institute. Formosa. 59: 1-57. 

Takahashi, R. 1933b. Aleyrodidae of Formosa, Part II. Report. Department of Agriculture. Government 

Research Institute. Formosa. 60: 1-24. 

Takahashi, R. 1934a. A new whitefly from China (Aleyrodidae, Homoptera). Lingnan Science Journal. 13: 137- 

141. 

Takahashi, R. 1934b. Aleyrodidae of Formosa, Part III. Rep. Dept. Agric. Govt. Res. Inst. Formosa. 63: 39-71. 

Takahashi, R. 1934c. Notes on the Aleyrodidae of Japan (Homoptera) I. Kontyu. 8: 223-224. 

622

Takahashi, R. 1934d. Some species of Aleyrodidae (Homoptera) from Madagascar, with a species from 

Mauritius. Memoires de l'institut Scientifique de Madagascar (A). 6: 353-385. 

Takahashi, R. 1935a. Aleyrodidae of Formosa, Part IV. Report. Department of Agriculture. Government 

Research Institute. Formosa. 66: 39-65. 

Takahashi, R. 1935b. Notes on the Aleyrodidae of Japan (Homoptera) II. Kontyu. 9: 25-27. 

Takahashi, R. 1935c. Notes on the Aleyrodidae of Japan (Homoptera) III (with Formosan species). Kontyu. 9: 

279-283. 

Takahashi, R. 1935d. Some foreign Aleyrodidae (Homoptera) VIII. Species from Thailand and French Indo- 

China. Trans. nat. Hist. Soc. Formosa. 32: 300-311. 

Takahashi, R. 1936a. A new Aleuroplatus from Africa (Hemiptera: Aleyrodidae. Arb. morph. taxon. Ent. Berl. 

3: 87-88. 

Takahashi, R. 1936b. An interesting whitefly from Africa (Hemiptera: Aleyrodidae). Arb. morph. taxon. Ent. 

Berl. 3: 52-53. 

Takahashi, R. 1936c. New whiteflies from the Philippines and Formosa (Aleyrodidae, Hemiptera). Philippine 

Journal of Science. 59: 217-221. 

Takahashi, R. 1936d. Notes on the Aleyrodidae of Japan (Homoptera) IV. Kontyu. 10: 150-151. 

Takahashi, R. 1936e. Some Aleyrodidae, Aphididae, Coccidae (Homoptera) and Thysanoptera from 

Micronesia. Tenthredo. 1: 109-120. 

Takahashi, R. 1936f. Three species of Aleyrodidae from China (Homoptera). Lingnan Science Journal. 15: 453- 

455. 

Takahashi, R. 1937a. A new species of the Aleyrodidae from New Zealand (Hemiptera). Trans. nat. Hist. Soc. 

Formosa. 27: 251-253. 

Takahashi, R. 1937b. Notes on the Aleyrodidae of Japan (Homoptera) V. Kontyu. 11: 310-311. 

623

Takahashi, R. 1937c. Three new species of Dialeurodes from China (Homoptera: Aleyrodidae). Lingnan Sci. J. 

16: 21-25. 

Takahashi, R. 1937d. Two new species of Aleyrodidae from Mauritius. Arb. morph. taxon. Ent. Berl. 4: 43-45. 

Takahashi, R. 1938a. A few Aleyrodidae from Mauritius and China (Hemiptera). Trans. nat. Hist. Soc. 

Formosa. 28: 27-29. 

Takahashi, R. 1938b. Notes on the Aleyrodidae of Japan (Homoptera) VI. Kontyu. 12: 70-74. 

Takahashi, R. 1938c. A new genus and species of Aleyrodidae from the Island of Reunion (Homoptera). Trans. 

nat. Hist. Soc. Formosa. 28: 269-271. 

Takahashi, R. 1938d. Three new species of Aleyrodidae from Mauritius (Homoptera). Annotnes zool. jap. 17: 

260-263. 

Takahashi, R. 1939a. Notes on the Aleyrodidae of Japan (Homoptera) VII. Kontyu. 13: 76-81. 

Takahashi, R. 1939b. Some Aleyrodidae from Mauritius (Homoptera). Insecta Matsumurana. 14: 1-5. 

Takahashi, R. 1939c. Some Aleyrodidae, Aphididae, and Coccidae from Micronesia (Homoptera). Tenthredo. 2: 

234-272. 

Takahashi, R. 1940a. A new species of Aleyrodidae from Yugoslavia. Arb. morph. taxon. Ent. Berl. 7: 148-149. 

Takahashi, R. 1940b. Insects of the Sternorrhyncha (Hemiptera) of Daito Jima, the Loochoo Islands. Trans. nat. 

Hist. Soc. Formosa. 30: 327-332. 

Takahashi, R. 1940c. Notes on the Aleyrodidae of Japan (Homptera) VIII. Kontyu. 14: 26-32. 

Takahashi, R. 1940d. Some foreign Aleyrodidae (Hemiptera) I. Trans. nat. Hist. Soc. Formosa. 30: 43-47. 

Takahashi, R. 1940e. Some foreign Aleyrodidae (Hemiptera) II. Trans. nat. Hist. Soc. Formosa. 30: 381-382. 

624

625 

Takahashi, R. 1941a. Some foreign Aleyrodidae (Hemiptera) III. Species from Hong Kong and Mauritius. 

Trans. nat. Hist. Soc. Formosa. 31: 351-357. 

Takahashi, R. 1941b. Some foreign Aleyrodidae (Hemiptera) IV. Species from Hong Kong. Trans. nat. Hist. 

Soc. Formosa. 31: 388-393. 

Takahashi, R. 1941c. Some species of Aleyrodidae, Aphididae and Coccidae from Micronesia (Homoptera). 

Tenthredo. 3: 208-220. 

Takahashi, R. 1942a. Aleurocanthus of Thailand and French Indo-China (Aleyrodidae, Homoptera). Kontyu. 

16: 57-61. 

Takahashi, R. 1942b. Some foreign Aleyrodidae (Homoptera) V. Species from Thailand and Indo-China. Trans. 

nat. Hist. Soc. Formosa. 32: 168-175. 

Takahashi, R. 1942c. Some foreign Aleyrodidae (Homoptera) VI. Species from Thailand and French Indo- 


Takahashi, R. 1942d. Some foreign Aleyrodidae (Homoptera) VII. Species from Thailand and French Indo- 


Takahashi, R. 1942e. Some foreign Aleyrodidae (Homoptera) VIII. Species from Thailand and French Indo- 


Takahashi, R. 1942f. Some foreign Aleyrodidae (Homoptera) VIII. Species from Thailand and French Indo- 


Takahashi, R. 1942g. Two new genera and species of Aleyrodidae from Thailand and French Indo-China 

(Homoptera). Annotnes zool. jap. 21: 102-105. 

Takahashi, R. 1943. Trialeurodes of Thailand (Aleyrodidae, Homoptera). Mushi. 15: 28-32. 

Takahashi, R. 1949. Some Aleyrodidae from Riouw Islands (Homoptera). Mushi. 20: 47-53. 

Takahashi, R. 1950. Four new species of Aleyrodidae (Homoptera) from Australia, India and Borneo. Annotnes 

zool. jap. 23: 85-88.

626 

Takahashi, R. 1951a. Descriptions of six interesting species of Aleyrodidae from Malaya (Homoptera). Kontyu. 

19: 1-8. 

Takahashi, R. 1951b. Some species of Aleyrodidae (Homoptera) from Japan. Misc. Rep. Res. Inst. nat. Resour. 

Tokyo. 19-21: 19-25. 

Takahashi, R. 1951c. Some species of Aleyrodidae (Homoptera) from Madagascar, with a species from 

Mauritius. Memoires de l'institut Scientifique de Madagascar (A). 6: 353-385. 

Takahashi, R. 1952a. Aleurotuberculatus and Parabemisia of Japan (Aleyrodidae, Homoptera). Miscellaneous 

Reports of the Research Institute for Natural Resources. Tokyo. 25: 17-24. 

Takahashi, R. 1952b. Some Malayan species of Aleyrodidae (Homoptera). Mushi. 24: 21-27. 

Takahashi, R. 1954a. Aleurolobus of Japan (Aleyrodidae, Homoptera). Kontyu. 20(3,4): 1-6. 

Takahashi, R. 1954b. Keys to the tribes and genera of Aleyrodidae of Japan, with descriptions of three new 

genera and one new species (Homoptera). Insecta Matsumurana. 18: 47-53. 

Takahashi, R. 1955a. Bemisia and Acanthobemisia of Japan (Aleyrodidae, Homoptera). Kontyu. 23(1): 1-5. 

Takahashi, R. 1955b. Descriptions of some new and little known species of Aleyrodidae from China and 

Malaya (Homoptera). Acta Entomologica Sinica. 5: 221-235. 

Takahashi, R. 1955c. Odontaleyrodes and Pealius of Japan (Aleyrodidae, Homoptera). Mushi. 29: 9-16. 

Takahashi, R. 1955d. Some species of Aleyrodidae from Madagascar (Homoptera) II. Mem. Inst. Scient. 

Madagascar (E). 1: 111-133. 

Takahashi, R. 1955e. Some species of Aleyrodidae from Madagascar III (Homoptera). Mem. Inst. Scient. 

Madagascar (E). 6: 375-441. 

Takahashi, R. 1956. Insects of Micronesia: Homoptera: Aleyrodidae). Insects Micronesia. 6: 1-13. 

Takahashi, R. 1957. Some Aleyrodidae from Japan (Homoptera). Insecta Matsumurana. 21: 12-21.

627 

Takahashi, R. 1958. Aleyrodes, Tuberaleyrodes, and Dialeurodes from Japan. Mushi. 31: 63-68. 

Takahashi, R. 1960a. Some species of Aleyrodidae of Reunion Island (Homoptera). Naturaliste malgache. 12: 

145-153. 

Takahashi, R. 1960b. Three species of Aleyrodidae from Reunion Island (Homoptera). Naturaliste malgache. 

12: 139-143. 

Takahashi, R. 1961a. Some new species of Aleyrodidae from Madagascar IV (Homoptera). Mem. Inst. Scient. 

Madagascar (E). 12: 323-339. 

Takahashi, R. 1961b. Two new genera and species of Aleyrodidae from Thailand and French Indo-China 

(Homoptera). Annotnes zool. jap. 21: 102-105. 

Takahashi, R. 1962. Two new genera and species of Aleyrodidae from Madagascar (Homoptera). Proceedings 

of the Royal Entomological Society of London (B). 31: 100-102. 

Takahashi, R. 1963. Some species of Aleyrodidae from Japan (Homoptera). Kontyu. 31: 49-57. 

Takahashi, R. 1978. Two new genera and species of Aleyrodidae from Madagascar (Homoptera). Proceedings 

of the Royal Entomological Society of London (B). 31: 100-102. 

Takahashi, R. and R. Mamet. 1952. Some species of Aleyrodidae from Madagascar (Homoptera) II. Mem. Inst. 

Scient. Madagascar (E). 1: 111-133. 

Tapia, E. A. 1970. [Preliminary study of Aleyrodidae found in Argentina (Homoptera).]OT: Estudio preliminar 

de los aleirodideos hallados en la Argentina (Homoptera). Actas Del Congreso Latinoamericano De Zoologia. 1, 

1970: 219 234, illustr. 

Tauili'ili, P. and A. M. Vargo. 1993. History of biological control in American Samoa. In D. F. Waterhouse and 

A. Vargo [eds.], Micronesia, Supp. 4: Biological control of exotic pests in the Pacific. Proceedings of a Plenary 

Session and Symposium XIX International Congress of Entomology, Beijing, 1992, Univ. of Guam, Guam. (. 

pp. 57-60. 

Teich, Y. 1966. Mites of the family Phytoseiidae as predators of the tobacco whitefly Bemisia tabaci. Israel J. 

Agri. Res. 16:141-142.

628 

Teran, A. L. 1979. Dos Paraleyrodes (Homoptera: Aleyrodidae) nue vos en los citricos de Tucuman. III. 

Jornada Fitosanitaria Argentina. 161-189. 

Teran, A. L. 1989. Description de la larva de Nephaspis picturata Gordon (Coleoptera: Coccinellidae: 

Scynmini), predator de aleyrodidos (Homoptera) en citricos de Tucuman, Republica Argentina. CIRPON, Rev. 

Invest. 7:75-81. 

Teran, A. L, and E. Frias. 1984. Los estados preimaginales de Nephaspis cocois Gordon (Coleoptera: 

Coccinellidae, Scymnini), predator de Aleurodidos (Homoptera) en citricos. Acta zool. lilloana. 37:201-206. 

Thomas, R. 1932. Periodic failure of the Punjab-American cotton crop. Agric. Live-stock India. 2:243-274. 

Thompson, F. C. 1981. The flower flies of the West Indies (Diptera: Syrphidae). Mem. Entomol. Soc. Wash. 

9:1-200. 

Thompson, W. R. 1950. A catalogue of the parasites and predators of insect pests. Section I. Parasite host 

catalogue. Part 3. Parasites of the Hemiptera. 2nd Edition, Ottawa, Canada. -. 149 pp. 

Thompson, W. R. and F. J. Simmonds. 1964. A catalog of the parasites and predators of insect pests. Section 3. 

Predator host catalog, Farnham Royal, Bucks. -. 204 pp. 

Timberlake, P. H. 1926. New species of Hawaiian chalcid-flies (Hymenoptera). Proceedings of the Hawaiian 

Entomological Society. 6: 305-320. 

Timofeyeva, T. V., and H. D. Nhuan. 1978. Morphology and biology of the Indian ladybird Serangium 

parcesetosum Sicard (Coleoptera: Coccinellidae) predacious on the citrus whitefly in Adzharia. Entomol. Rev. 

57:210-214. 

Traghard, I. 1909. On the biology and development of Clitostethus arcuatus (Rossi), an enemy of the 

Aleyrodidae. Arkiv Zool. 5:13-14. 

Trehan, K. N. 1938. Two new species of Aleurodidae found on ferns in greenhouses in Britain (Hemiptera). 

Proceedings of the Royal Entomological Society of London (B). 7: 182-189. 

Trehan, K. N. 1939. Studies on the British Aleurodidae. Current Science. 8: 266. 

Trehan, K. N. 1940. Studies on the British Whiteflies (Homoptera-Aleyrodidae). Transactions of the Royal

Entomological Society of London. 90: 575-616. 

Tremblay, E. 1969. Il controllo del Siphoninus phillyreae (Haliday) in Campania. Studio del gruppo di lavoro 

C.N.R. per la lotta integrata contro I nemici delle piante: XL. Boll. Lab. Entomol. Agrar. 'Filippo Silvestri'. 

27:161-176. 

Tremblay, E. and F. M. Iaccarino. 1978. Aleurotuba jelinski (Frauen.) per Aleurotrachelus jelineki (Frauen.) 

(Homoptera: Aleyrodidae) su viburno. Bollettino Del Laboratorio Di Entomologia Agraria Filippo Silvestri. 35: 

57-66. 

Trjapitzin, V. A. 1978. Hymenoptera II. Chalcidoidea 7. Encyrtidae. Opred. Faune SSR. pp. 236-328. 

Tullgren, A. 1907. Uber einige Arten der Familie Aleurodidae. Arkiv for Zoologi. 3(26): 1-18. 

USNM Whitefly Collection, website. 

Ulusoy, M. R., N. Uygun, U. Kersting, I. Karaca and S. Satar. 1996. Present status of citrus whiteflies 

(Homoptera: Aleyrodidae) in Turkey and their control. Zeitschrift fur Pflanzenkrankheiten und Pfalanzenschutz. 

103: 397-402. 

Uygun N., B. Ohnesorge and R. Ulusoy. 1990. Two new whiteflies on citrus in Eastern Mediterrean: 

Parabemisia myricae (Kuwana) and Dialeurodes citri (Ashmead). Journal of Applied Entomology. 110: 471- 

482. 

van Ooststroom, S. J. 1944. Sciopus exul Parent (Dipt, Dolchopodid.) in den Leidschen Hortus. Ent. Ber. 

11:196. 

van Schelt, J, J. Klapwijk, M. Letard, and C. Aucouturier. 1995. The use of Macrolophus caliginosus as a 

whitefly predator in protected crops. In D. Gerling and R. T. Mayer [eds.], Bemisia 1995: taxonomy, biology, 

damage control and management. Intercept Ltd, Andover, Hants UK. pp. 515-521. 

Vazquez, L. L. 1999. Whiteflies (Homoptera: Aleyrodidae) of Cuba and their host plants. Bollettino Del 

Laboratorio Di Entomologia Agraria Filippo Silvestri. 55: 139-149. 

Venkataramaiah, G. H. 1971. A note on Dialeurodes vulgaris on coffee. Journal of Coffee Research. 1: 13-14. 

Venugopal Rao, N., A. S. Reddy and D. D. R. Reddy. 1990. Effects of some insecticides on the parasitoids and 

629

predators of the cotton whitefly, Bemisia tabaci. Genn. J. Bio. Cont. 4:4-7. 

Venugopal Rao, N., A. S. Reddy, and K. Tirumala Rao. 1989. Natural enemies of cotton whitefly, Bemisia 

tabaci Gennadius in relation to host populations and weather factors. J. Bio. Control. 3:10-12. 

Vesey-Fitzgerald, D. 1938. Biological notes on some Coccinellidae from East Africa. Bull. Entomol. Res. 

29:185-189. 

Viggiani, G. 1977. Recerche sugli Hymenoptera Chalcidoidea LIV. Descrizione di Euderomphale bemisiae n. 

sp. (Eulophidae) parassita di Bemisia citricola Gom. Men. (Homoptera: Aleyrodidae). Bollettino del 

Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 34: 16-18. 

Viggiani, G. 1981. Two new species of Encarsia Foerster (Hym. Aphelinidae) from known hosts. Bollettino del 

Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 38: 61-65. 

Viggiani, G. 1981a. Afelinidi parassiti di Asterobemisia avellanae (Signoret) (Homoptera, Aleyrodidae), con 

descrizione di una nuova specie. Bollettino del Laboratario di Entomologia Agraria 'Filippo Silvestri' di Portici. 

38: 67-72. 

Viggiani, G. 1982a. New species and host records of African aphelinids (Researches on Hymenoptera 

Chalcidoidea LXX). Journal of the Entomological Society of South Africa. 45: 27-32. 

Viggiani, G. 1982b. Notes on Encarsia olivina (Masi) with descriptions of two new species of Encarsia Foerster 

(Hym, Aphelinidae). Bollettino del Laboratario di Entomologia Agraria 'Filippo Silvestri' di Portici. 39: 11-17. 

Viggiani, G. 1985. A new species of Encarsia (Hym. Aphelinidae) from Papua New Guinea and redescription 

of Coccophagus pulliclavus Girault, 1917. Bollettino del Laboratorio di Entomologia Agraria 'Filippo Silvestri', 

Portici. 42:11-14. 

Viggiani, G. 1985a. Additonal notes and illustrations on some species of aphelinids described by A.A. Girault 

and A.P. Dodd in the genera Coccophagus Westw, Encarsia Foerst. and Prospaltella Ashm. (Hym.: 

Chalcidoidea). Bollettino del Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 42:233-255. 

Viggiani, G. 1985b. Notes on a few Aphelinidae, with descriptions of five new species of Encarsia Foerster 

(Hymenoptera, Chalcidoidea). Bollettino del Laboratario di Entomologia Agraria 'Filippo Silvestri' di Portici. 

42: 81-94. 

Viggiani, G. 1986. Notes on some species of Coccophagus Westwood, Coccophagoides Girault, Encarsia 

630

631 

Foerster and Encarsiella Hayat (Hymenoptera: Aphelinidae), mainly from the Nearctic and Neotropical regions. 

Bollettino del Laboratario di Entomologia Agraria 'Filippo Silvestri' di Portici. 43: 59-78. 

Viggiani, G. 1987. La specie italiane del genre Encarsia Foerster (Hymenoptera: Aphelinidae). Bollettino del 

Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 44:121-179. 

Viggiani, G. 1987a. New species of Encarsia Foerster (Hymenoptera: Aphelinidae), parasitoids of whiteflies. 

Bollettino del Laboratario di Entomologia Agraria 'Filippo Silvestri' di Portici. 44: 33-41. 

Viggiani, G. 1989. Notes on some nearctic and neotropical Encarsia Forster (Hymenoptera: Aphelinidae). 

Bollettino del Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 46:207-213. 

Viggiani, G. 1989a. Two new species of Encarsia Foerster (Hymenoptera: Aphelinidae) from Africa. Redia. 72: 

513-517. 

Viggiani, G. 1991. Ridescrizione di Amitus fuscipennis MacG. & Neb. (Hym.: Platygasteridae), parasitoide 

esotico di Trialeurodes vaporariorum (Westw.) con notizie preliminari sulla sua introduzione in Italia. 

Bollettino del R. Laboratorio di Entomologia Agraria di Portici. 74(1): 177-183. 

Viggiani, G. 1993. New species of Encarsia Forster (Hymenoptera: Aphelinidae), parasitoids of whiteflies, 

from Hawaii and Yemen. Redia. 76(1):121-127. 

Viggiani, G. 1996. Discovery of the male of Amitus fuscipennis MacGown et Nebeker (Hym. Platygasteridae), a 

parasitoid of Trialeurodes vapororarirum (Westwood) (Hom. Aleyrodidae). Bollettino del Laboratorio di 

Entomologia Agraria 'Filippo Silvestri', Portici. 52: 97-99. 

Viggiani, G. 1996a. Fenologia, dinamica delle popolazioni e parassitoidi di Parabemisia myricae (Kuwana) 

(Homoptera: Aleyrodidae) in Campania. Bollettino Del Laboratorio Di Entomologia Agraria Filippo Silvestri. 

51: 137-153. 

Viggiani, G. and D. Battaglia. 1983. Le specie italiane del genere Eretmocerus Hald. (Hymenoptera: 

Aphelinidae). Bollettino del Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 40:97-101. 

Viggiani, G. and M. Carver. 1988. Cales orchamoplata sp.n. (Hymenoptera: Aphelinidae) from Australia. 

Journal of the Australian Entomological Society. 27:43-45. 

Viggiani, G. and G. A. Evans. 1992. Descriptions of three new species of Amitus Haldeman (Hymenoptera: 

Platygasteridae), parasitoids of known whiteflies from the New World. Bollettino del Laboratorio di

Entomologia Agraria 'Filippo Silvestri', Portici. 49:189-194. 

Viggiani, G. and R. Hui. 1993. New species and records of Aphelinidae (Hymenoptera: Chalcidoidea) from 

China. Bollettino del Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 48:219-239. 

Viggiani, G. and S. Laudonia. 1991. On the occurrence in Italy of Encarsia meritoria Gahan (Hymenoptera; 

Aphelinidae), exotic parasitoid of whiteflies (Homoptera; Aleyrodidae). (in Italian with English summary). 

Redia. 74(1):135-140. 

Viggiani, G. and P. Mazzone. 1978. Morphologia, biologia e utilizzazione di Prospaltella lahorensis How. 

(Hym., Aphelinidae), parassita esotico introdotto in Italia per la lotta biologia ad Dialeuordes citri (Ashm.) 

(italian). Bollettino del Laboratario di Entomologia Agraria 'Filippo Silvestri' di Portici. 35: 99-161. 

Viggiani, G. and P. Mazzone. 1979. Contributi alla conoscenza morfo-biologica delle specie del complesso 

Encarsia Foerster - Prospaltella Ashmead (Hym. Aphelinidae). Bollettino del Laboratario di Entomologia 

Agraria 'Filippo Silvestri' di Portici. 36: 42-50. 

Viggiani, G. and P. Mazzone. 1980. Encarsia pseudopartenopea n.sp, parassita di Siphoninus phillyreae 

(Haliday) (Hom. Aleyrodidae). Bollettino del Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 

37:9-12. 

Viggiani, G. and P. Mazzone. 1980a. Le specie paleartiche di Encarsia del gruppo lutea Masi (Hym, 

Aphelinidae), con descrizione di due nuove specie. Bollettino del Laboratorio di Entomologia Agraria 'Filippo 

Silvestri', Portici. 37:51-57. 

Viggiani, G. and P. Mazzone. 1980b. Sull'introduzione in Italia di Encarsia pergandiella Howard 

(Hymenoptera: Aphelinidae), parassita di Trialeurodes vaporariorum (Wetw.) (Homoptera: Aleyrodidae). 

Bollettino del Laboratorio di Entomologia Agraria 'Filippo Silvestri', Portici. 37:39-43. 

Viggiani, G. and P. Mazzone. 1982. Amitus Hald. (Hym. Platygasteridae) of Italy, with descriptions of three 

new species. Bollettino del R. Laboratorio di Entomologia Agraria di Portici. 39: 55-69. 

Viggiani, G. and A. van Harten. 1996. Aphelinidae, Signiphoridae, Mymaridae and Tricogrammatidae of the 

Cape Verde Islands (Hymenoptera: Chalcidoidea). Bollettino Del Laboratorio Di Entomologia Agraria Filippo 

Silvestri. 51: 67-81. 

Visnya, A. 1936. Tovabbi molytetvek koszegrol es videkerol. Weitere Mottenlause aus der Umgebung von 

Koszeg. Folia sabariensis vasi Szle. 3: 116-117. 

632

Visnya, A. 1941a. A gigantic species of Aleurodidae (Homoptera) from greenhouse-Orchideas. Folia Ent. hung. 

6: 4-15. 

Visnya, A. 1941b. Vorarbeiten zur Kenntnis der Aleurodiden-Fauna von Ungarn, nebst systematischen 

Bemerkungen uber die Gattungen Aleurochiton, Pealius und Bemisia (Homoptera). Fragmenta Faunistica 

Hungarica. 4 (Suppl) 1-19. 

von Frauenfeld, G. R. 1866. Zoologische Miscellen IX. Verhand. Kaiser.-Konigl. zool.-bot. Gesell. Wien. 

16:555. 

von Frauenfeld, G. R. 1867. Zoologische Miscellen XIV. Acletoxenus syrphoides. Verhand. Kaiser.-Konigl. 

zool.-bot. Gesell. Wien. 18:150-153. 

von Frauenfeld, G. R. 1868. Zoologische Miscellen XV. Ueber Gattungsund Artrecht von Acletoxenus 

syrphoides. Verhand. Kaiser.-Konigl. zool.-bot. Gesell. Wien. 18:897-899. 

Walker, F. 1852. List of the specimens of Homopterous insects in the collection of the British Museum. 

Supplement. 369 pp. London. 4: 909-1188. 

Walker, F. 1853. Diptera Britannica, vol. 2, London. 

Walker, F. 1858. List of the specimens of Homopterous insects in the collection on the British Museum. 

Supplement. 369 pp. 

Walker, F. 1939. Monographia Chalciditum, I. Hyppolitus Bailliere, London. 333 pp. 

Watson, J. R. 1914. Whitefly control. Agri. Exp. Sta. Bulletin ed, vol. 123, Univ. Florida, Gainesville. -. 

Watson, J. R. 1918. Insects of a citrus grove. Agri. Exp. Sta. Bulletin ed, Univ. Florida, Gainesville. vol. 148. 

Watson, J. R. 1918a. The California Delphastus. Florida Bug. 2:88. 

Watson, J. R. 1923. Delphastus at Bratentown. Fl. Entomol. 6:57. 

Watson, J. R. 1926. Citrus insects and their control. Agricultural Experiment Station Bulletin, Univ. of Florida. 

633

183. 

Watve, C. M and D. F. Clower. 1976. Natural Enemies of the Bandedwing Whitefly in Louisiana. Environ. 

Entomol. 5(6):1075-1078. 

Weems, H. V. 1971. A hover fly, Allograpta obliqua (Say) (Diptera: Syrphidae). Entomology Circular ed, 

Florida Department of Agriculture, Gainesville. vol. 106. 

Weems, H. V. 1974. Orange spiny whitefly. Aleurocanthus spiniferus (Quaintance) (Homoptera: Aleyrodidae). 

Fl. Dept. Agr. & Consumer Serv. DPI Entomology Circular. 151:1-2. 

Westhoff, Fr. 1887. Die Phytophthiren-Gattung Aleurodes und ihre in der Umgegend von Munster 

aufgefundenen Arten. Jber. westf. Proc. Wiss. Kunst. 1886: 55-63. 

Westwood, J. O. 1840. An introduction to the modern classification of insects; founded on the natural habits and 

corresponding organization of different families. 2:587 pp. London. Longman, Orme, Brown & Green, London. 

587 pp. 

Westwood, J. O. 1856. The new Aleyrodes of the greenhouse. Gardener's Chronicle. 1856: 852. 

Whittaker, O. 1930. Some new species and a new genus of parasitic Hymenoptera from British Colombia. 


Wijesekera, G. A. W. and C. Kudagamage. 1990. Life history and control of 'spiralling' whitefly Aleurodicus 

dispersus (Homoptera: Aleyrodidae) fast spreading pest in Sri Lanka. Quart. Newslet. Asia Pacific Plant Prot. 

Comm. 33:22-25. 

Williams, F. X. 1944. A survey of insect pests of New Caledonia. Hawaii Plrs. Rec. 48(2): 93-124. 

Woglum, R. S. 1913. Report of a trip to India and the Orient in search of the natural enemies of the citrus 

whitefly. USDA Bureau of Entomology Bulletin. 120:1-58. 

Woolley, J.B. 1988. Phylogeny and classification of the Signiphoridae (Hymenoptera: Chalcidoidea). 

Systematic Entomology. 13:465-501. 

Wunn, H. 1926. In. Elsass-Lothringen vorkommede Schildlausarten. Zeitschrift fnr Wissenschaftliche 

Insektenbiologie, Berlin. 21: 22-28. 

634

Wysoki, M. and M. Cohen. 1983. Mites of the family Phytoseiidae (Acarina: Mesotigmata) as predators of the 

Japanese bayberry whitefly, Parabemisia myricae Kuwana (Hom.:Aleyrodidae). Agronomie. 3:823-825. 

Yasnosh, V. A. 1973. On the faune of aphelinids (Hymenoptera: Aphelinidae) of middle Asia and Kazakhstan. 

Ent. Obozr. 52: 899-909. 

Yasnosh, V. A. 1973a. The Aphelinidae fauna (Hymenoptera) of central Asia and Kazakh SSR. (in Russian 

with English summary). Entomologicheskoe Obozrenie. 52(4):899-909. 

Yasnosh, V. A. 1974. Novyi vid Eretmocerus Haldeman (Hymenoptera, Aphelinidae) iz Gruzii. Soobshcheniya 

Akademii Nauk Gruzinskoy SSR. 73(3):713-716. 

Yasnosh, V. A. 1989. Species of the genus Encarsia Foerster (Hymenoptera, Aphelinidae) - parasites of 

aleyrodids in the USSR. (in Russian). Trudy Zoologicheskogo Instituta. Akademiya Nauk SSSR. Leningrad. 

191:109-120. 

Yasnosh, V. A. 1991. Whiteflies and their natural enemies. Zash. Rast. (Moskva). 2:24-26. 

Yasnosh, V. A. 1995. Family Aphelinidae. (in Russian). Key to the insects of Russian far east in six volumes. 

4(2):506-551. 

Yasumatsu, K. and C. Watanabe. 1964. A tentative catalogue of insect natural enemies of injurious insects in 

Japan. Kyushu University, Fukuoka, Japan. -. 

Yigit, A. 1991. Serangium parcesetosum Sicard (Col, Coccinellidae): New records as a citrus whitefly 

predatory lady beetle in Turkey. Turk. Entomol. Derg. 16:163-167. 

Yigit, A. 1992. Method for culturing Serangium parcesetosum Sicard (Coleoptera: Coccinellidae) on Bemisia 

tabaci Genn. (Homoptera: Aleyrodidae). J. Plant Dis. Prot. 99:525-527. 

Yigit, A. 1992b. Turkiye'de yeni tesbit edilen Turuncgil beyazsinegi avcisi bir bocek: Serangium parcesetosum 

Sicard (Coleoptera: Coccinellidae). Turk. Entomol. Derg. -. 

Yoshida, H. A. and R. F. L. Mau. 1985. Life history and feeding behavior of Nephaspis amnicola Wingo. Proc. 

Hawaii. Entomol. Soc. 25:155-160. 

Young, B. 1942. White flies attacking citrus in Szechwan. Sinensia, Shanghai. 13: 95-101. 

635

Young, B. 1944. Aleurodidae from Szechwan, I. Sinensia, Shanghai. 15: 129-139. 

Zachrisson, B. and J. Poveda. 1993. Las moscas blancas in Panam . In L. Hilje and O. Arboleda [eds.], Serie 

Tpcnica, Informe Tpcnico No. 205: Las Moscas Blancas (Homoptera: Aleyrodidae) en Amprica Central Y El 

Caribe. CATIE, Turrialba, C. 

Zahradnik, J. 1956. Trois nouvelles especes des aleyrodides pour la faune tchecoslovaque. Sb. faun. Praci ent. 

Odd. nar. Mus. Praze. 1: 43-45. 

Zahradnik, J. 1961. Nouvelles connaissances faunistiques et taxonomiques sur les aleyrodides de la 

Tchecoslovaquie (Homoptera, Aleyrodinea). Acta Faunistica Entomologica Musei Nationalis Pragae. 7: 61-80. 

Zahradnik, J. 1962. Donnees taxonomiques et faunistiques sur Japaneyrodes nov. gen. similis europaeus n. spp. 

(Homoptera, Aleyrodinea). Acta Faunistica Entomologica Musei Nationalis Pragae. 8: 13-19. 

Zahradnik, J. 1962a. Uber einige Aleyrodiden-Arten aus Belin und Umgebung (Homoptera, Aleyrodinea). Acta 

Faunistica Entomologica Musei Nationalis Pragae. 8: 37-41. 

Zahradnik, J. 1963a. Aleyrodina. In. Die Tierwelt Mitteleuropas. 4(3): 1-19. 

Zahradnik, J. 1963b. Notes faunistiques sur les aleurodes en Yougoslavie (Homoptera, Aleyrodinea). Acta 

Faunistica Entomologica Musei Nationalis Pragae. 9: 231-235. 

Zahradnik, J. 1970. Ramsesseus follioti gen. n, sp. n. aleurodes nouveau d'Egypt (R.A.U.) (Homoptera, 

Aleyrodinea). Acta Entomologica Bohemoslovaca. 67: 47-49. 

Zahradnik, J. 1987. Asterobemisia nigrini sp. n, un aleurode nouveau de la Tchecoslovaquie (Sternorrhyncha, 

Aleyrodinea) II. Acta Entomologica Bohemoslovaca. 84: 350-352. 

Zahradnik, J. 1991. Taxonomisches und Faunistisches uber europaische Mottenlause (Aleyrodinea). Acta 

Universitatis Carolinae. Biologica. 35: 111-118. 

Zehntner, L. 1896. De Plantenluizen van het Suikerriet op Java. Meded. Proefstn. Suik. Riet W. Java 'Kagok'. 

Nieuwe Serie. 29: 935-950. 

Zehntner, L. 1897. Mededeelingen uit en voor de Praktijk. Voorloopige Mededeeeling over ees Luizenplaag. 

Meded. v.h. Proefstation Oost. Java en Archief Java Suikerindustrie. 5: 381. 

636

Zehntner, L. 1897a. Overzicht van de Zieken van het Suikerriet op Java. Meded. Proefstn. Suik. Riet W. Java 

'Kagok'. Nieuwe Serie. 37: 525-575. 

Zehntner, L. 1899. De Plantenluizen van het Suikerriet op Java VIII. Meded. Proefstn. Suik. Riet W. Java 

'Kagok'. Nieuwe Serie. 38: 10. 

Zhang, Y., Z. Tao, and Z. Luo. 1991. Citrus whitefly population dynamics and effect of pesticide on it. Chin. J. 

Appl. Ecol. 2:323-328. 

Zimmermann, E. C. 1957. Insects of Hawaii. Univ. of Hawaii, Honolulu. vol. 6. 

Zolnerowich, G. and M. Rose. 1996. A new species of Entedononecremnus (Hymenoptera: Chalcidoidea: 

Eulophidae) parasitic on the giant whitefly, Aleurodicus dugesii Cockerell (Homoptera: Aleyrodidae). 


Zolnerowich, G. and M. Rose. 1998. Eretmocerus Haldeman (Hymenoptera: Aphelinidae) imported and 

released in the United States for control of Bemisia (tabaci complex) (Homoptera: Aleyrodidae). Proceedings of 

the Entomological Society of Washington. 100(2), April, 1998: 310 323, illustr. 

Zolnerowich, G. and M. Rose. 2004. Eretmocerus rui n. sp. (Hymenoptera: Chalcidoidea: Aphelinidae) an 

exotic natural enemy of Bemisia (tabaci group) (Homoptera: Aleyrodidae) released in Florida. Florida 


637

Cited Name Valid Name 

ALEURODICINAE QUAINTANCE & BAKER 1913 ALEURODICINAE QUAINTANCE & BAKER 1913 

ALEYRODIDAE WESTWOOD 1840 ALEYRODIDAE WESTWOOD 1840 

ACANTHALEYODES TAKAHASHI 1931 ACANTHALEYRODES TAKAHASHI 1931 

Acanthaleyrodes callicarpae Takahashi 1931 Acanthaleyrodes callicarpae Takahashi 1931 

Acanthaleyrodes spiniferosa (Corbett) 1933 Tuberaleyrodes spiniferosa (Corbett) 1933 

Acanthaleyrodes styraci Takahashi 1942 Acanthaleyrodes styraci Takahashi 1942 

ACANTHOBEMISIA TAKAHASHI 1935 ACANTHOBEMISIA TAKAHASHI 1935 

Acanthobemisia distylii Takashashi 1935 Acanthobemisia distylii Takahashi 1935 

Acanthobemisia indicus Meganathan & David 1994 Acanthobemisia indicus Meganathan & David 1994 

ACAUDALEYRODES TAKAHASHI 1951 ACAUDALEYRODES TAKAHASHI 1951 

Acaudaleyrodes africanus (Dozier) 1934 Acaudaleyrodes africanus (Dozier) 1934 

Acaudaleyrodes alhagi (Priesner & Hosny) 1934 Acaudaleyrodes rachipora (Singh) 1931 

Acaudaleyrodes citri (Priesner & Hosny) 1934 Acaudaleyrodes rachipora (Singh) 1931 

Acaudaleyrodes ebeni Manzari & Alemansoor 2005 Acaudaleyrodes ebeni Manzari & Alemansoor 2005 

Acaudaleyrodes pauliani Takahashi 1951 Acaudaleyrodes pauliani Takahashi 1951 

Acaudaleyrodes rachipora (Singh) 1931 Acaudaleyrodes rachipora (Singh) 1931 

Acaudaleyrodes tuberculata Bink-Moenen 1983 Acaudaleyrodes tuberculata Bink-Moenen 1983 

ACUTALEYRODES TAKAHASHI 1960 ACUTALEYRODES TAKAHASHI 1960 

Acutaleyrodes palmae Takahashi 1960 Acutaleyrodes palmae Takahashi 1960 

AFRICALEURODES DOZIER 1934 AFRICALEURODES DOZIER 1934 

Africaleurodes adami Cohic 1968 Africaleurodes adami Cohic 1968 

Africaleyrodes ananthakrishnani Dubey & Sundararaj Africaleyrodes ananthakrishnani Dubey & Sundararaj 

Africaleurodes balachowskyi Cohic 1968 Africaleurodes balachowskyi Cohic 1968 

Africaleurodes capgrasi Cohic 1968 Africaleurodes capgrasi Cohic 1968 

Africaleurodes citri (Takahashi) 1932 Africaleurodes citri (Takahashi) 1932 

Africaleurodes coffeacola Dozier 1934 Africaleurodes coffeacola Dozier 1934 

Africaleurodes fulakariensis Cohic 1966 Africaleurodes fulakariensis Cohic 1966 

Africaleurodes hexalobi Bink-Moenen 1983 Africaleurodes hexalobi Bink-Moenen 1983 

Africaleurodes indicus Regu & David 1993 Africaleurodes indicus Regu and David 1993 

Africaleyrodes karwarensis Dubey & Sundararaj Africaleyrodes karwarensis Dubey & Sundararaj 

Africaleurodes lamottei Cohic 1969 Africaleurodes coffeacola Dozier 1934 

Africaleurodes loganiaceae Dozier 1934 Africaleurodes loganiaceae Dozier 1934 

Africaleurodes martini Cohic 1968 Africaleurodes martini Cohic 1968 

Africaleurodes ochnaceae Dozier 1934 Africaleurodes ochnaceae Dozier 1934 

Africaleurodes pauliani Cohic 1968 Africaleurodes pauliani Cohic 1968 

Africaleurodes simula (Peal) 1903 Africaleurodes simula (Peal) 1903 

638

Africaleurodes souliei Ardaillon & Cohic 1970 Africaleurodes souliei Ardaillon & Cohic 1970 

Africaleurodes tetracerae Cohic 1966 Africaleurodes tetracerae Cohic 1966 

Africaleurodes uvariae Cohic 1968 Africaleurodes uvariae Cohic 1968 

Africaleurodes vrijdaghii (Ghesquiere) 1934 Africaleurodes vrijdaghii (Ghesquiere) 1934 

AGROSTALEYRODES KO 2001 AGROSTALEYRODES KO 2001 

Agrostaleyrodes arcanus Ko 2001 Agrostaleyrodes arcanus Ko 2001 

ALEUROCANTHUS QUAINTANCE & BAKER 1914 ALEUROCANTHUS QUAINTANCE & BAKER 1914 

Aleurocanthus aberrans Cohic 1968 Aleurocanthus zizyphi Priesner & Hosny 1934 

Aleurocanthus alternans Cohic 1966 Cohicaleyrodes alternans (Cohic) 1966 

Aleurocanthus angolensis Cohic 1966 Cohicaleyrodes alternans (Cohic) 1966 

Aleurocanthus arecae David & Manjunatha 2003 Aleurocanthus arecae David & Manjunatha 2003 

Aleurocanthus ayyari Regu & David 1993 Aleurocanthus ayyari Regu & David 1993 

Aleurocanthus bambusae (Peal) 1903 Aleurocanthus bambusae (Peal) 1903 

Aleurocanthus bangalorensis Dubey & Sundararaj 2005 Aleurocanthus bangalorensis Dubey & Sundararaj 2005 

Aleurocanthus banksiae (Maskell) 1895 Aleurocanthus banksiae (Maskell) 1895 

Aleurocanthus brevispinosus Dumbleton 1961 Aleurocanthus brevispinosus Dumbleton 1961 

Aleurocanthus calophyllii (Kotinsky) 1907 Aleurocanthus calophyllii (Kotinsky) 1907 

Aleurocanthus cameroni Corbett 1935 Aleurocanthus citriperdus Quaintance & Baker 1916 

Aleurocanthus canangae Corbett 1935 Aleurocanthus cocois Corbett 1927 

Aleurocanthus ceracroceus Martin 1999 Aleurocanthus ceracroceus Martin 1999 

Aleurocanthus cheni Young 1942 Aleurocanthus cheni Young 1942 

Aleurocanthus chiengmaiensis Takahashi 1942 Aleurocanthus chiengmaiensis Takahashi 1942 

Aleurocanthus cinnamomi Takahashi 1913 Aleurocanthus cinnamomi Takahashi 1913 

Aleurocanthus citricolus (Newstead) 1911 Aleurocanthus spiniferus (Quaintance) 1903 

Aleurocanthus citriperdus Quaintance & Baker Aleurocanthus citriperdus Quaintance & Baker 

Aleurocanthus citriperdus Quaintance & Baker 1916 Aleurocanthus citriperdus Quaintance & Baker 1916 

Aleurocanthus clitoriae Jesudasan & David 1991 Aleurocanthus clitoriae Jesudasan & David 1991 

Aleurocanthus cocois Corbett 1927 Aleurocanthus cocois Corbett 1927 

Aleurocanthus corbetti Takahashi 1951 Aleurocanthus corbetti Takahashi 1951 

Aleurocanthus davidi David & Subramaniam 1976 Aleurocanthus davidi David & Subramaniam 1976 

Aleurocanthus delottoi Cohic 1969 Aleurocanthus delottoi Cohic 1969 

Aleurocanthus delottoi nr. Cohic 1969 Aleurocanthus delottoi Cohic 1969 

Aleurocanthus descarpentriesi Cohic 1968 Cohicaleyrodes descarpentriesi (Cohic) 1968 

Aleurocanthus dissimilis Quaintance & Baker 1917 Aleurocanthus dissimilis Quaintance & Baker 1917 

Aleurocanthus esakii Takahashi 1936 Aleurocanthus esakii Martin 1985 

Aleurocanthus eugeniae Takahashi 1933 Aleurocanthus eugeniae Takahashi 1933 

Aleurocanthus euphorbiae Jesudasan & David 1991 Aleurocanthus euphorbiae Jesudasan & David 1991 

Aleurocanthus ficicola David 1993 Aleurocanthus ficicola David 1993 

Aleurocanthus firmianae Dubey & Sundararaj 2005 Aleurocanthus firmianae Dubey & Sundararaj 2005 

Aleurocanthus froggatti Martin 1999 Aleurocanthus froggatti Martin 1999 

Aleurocanthus fumipennis (Hempel) 1899 Crescentaleyrodes fumipennis (Hempel) 1899 

639

Aleurocanthus gateri Corbett 1927 Aleurocanthus gateri Corbett 1927 

Aleurocanthus goaensis Dubey & Sundararaj 2005 Aleurocanthus goaensis Dubey & Sundararaj 2005 

Aleurocanthus gordoniae Takahashi 1941 Aleurocanthus gordoniae Takahashi 1941 

Aleurocanthus gymnosporiae Jesudasan & David 1991 Aleurocanthus gymnosporiae Jesudasan & David 1991 

Aleurocanthus hansfordi Corbett 1935 Aleurocanthus zizyphi Priesner & Hosny 1934 

Aleurocanthus hibisci Corbett 1935 Aleurocanthus hibisci Corbett 1935 

Aleurocanthus hirsutus (Maskell) 1895 Aleurocanthus hirsutus (Maskell) 1895 

Aleurocanthus husaini Corbett 1939 Aleurocanthus husaini Corbett 1939 

Aleurocanthus imperialis Cohic 1968 Aleurocanthus imperialis Cohic 1968 

Aleurocanthus inceratus Silvestri 1927 Aleurocanthus inceratus Silvestri 1927 

Aleurocanthus indicus David & Regu 1989 Aleurocanthus indicus David & Regu 1989 

Aleurocanthus ixorae Jesudasan & David 1991 Aleurocanthus ixorae Jesudasan & David 1991 

Aleurocanthus leptadeniae Cohic 1968 Aleurocanthus leptadeniae Cohic 1968 

Aleurocanthus lobulatus Jesudasan & David 1991 Aleurocanthus lobulatus Jesudasan & David 1991 

Aleurocanthus longispinus Quaintance & Baker 1917 Aleurocanthus longispinus Quaintance & Baker 1917 

Aleurocanthus loyolae David & Subramaniam 1976 Aleurocanthus loyolae David & Subramaniam 1976 

Aleurocanthus lumpurensis Corbett 1935 Aleurocanthus longispinus Quaintance & Baker 1917 

Aleurocanthus luteus Martin 1985 Aleurocanthus luteus Martin 1985 

Aleurocanthus m'voutiensis Cohic 1966 Aleurocanthus mvoutiensis Cohic 1966 

Aleurocanthus mackenziei Cohic 1969 Aleurocanthus mackenziei Cohic 1969 

Aleurocanthus mangiferae Quaintance & Baker 1917 Aleurocanthus mangiferae Quaintance & Baker 1917 

Aleurocanthus martini David 1993 Aleurocanthus martini David 1993 

Aleurocanthus marudamalaiensis David & Subramaniam 1976 Aleurocanthus marudamalaiensis David & Subramaniam 1976 

Aleurocanthus mayumbensis Cohic 1966 Aleurocanthus mayumbensis Cohic 1966 

Aleurocanthus multispinosus Dumbleton 1961 Aleurocanthus multispinosus Dumbleton 1961 

Aleurocanthus musae David & Jesudasan 2002 Aleurocanthus musae David & Jesudasan 2002 

Aleurocanthus nagpurensis Raina et al. Aleurocanthus woglumi Ashby 1915 

Aleurocanthus niger Corbett 1926 Aleurocanthus niger Corbett 1926 

Aleurocanthus nigricans Corbett 1926 Aleurocanthus nigricans Corbett 1926 

Aleurocanthus nubilans (Buckton) 1900 nomen dubium 

Aleurocanthus nudus Dumbleton 1961 Aleurocanthus nudus Dumbleton 1961 

Aleurocanthus obovalis Corbett 1926 Aleurocanthus nigricans Corbett 1926 

Aleurocanthus palauensis Kuwana 1931 Aleurocanthus palauensis Kuwana 1931 

Aleurocanthus palmae Ghesquiere Cerataphis lataniae Boisduval [Psyllidae] 

Aleurocanthus papuanus Martin 1985 Aleurocanthus papuanus Martin 1985 

Aleurocanthus pauliani Cohic 1968 Cohicaleyrodes pauliani (Cohic) 1968 

Aleurocanthus pendleburyi Corbett 1935 Aleurocanthus pendleburyi Corbett 1935 

Aleurocanthus piperis (Maskell) 1895 Aleurocanthus piperis (Maskell) 1895 

Aleurocanthus platysepali Cohic 1966 Aleurocanthus platysepali (Cohic) 1966 

Aleurocanthus punjabensis Corbett 1935 Aleurocanthus woglumi Ashby 1915 

Aleurocanthus recurvispinus Cohic 1966 Cohicaleyrodes recurvispinus (Cohic) 1966 

640

Aleurocanthus regis Mound 1965 Aleurocanthus regis Mound 1965 

Aleurocanthus rosae Singh 1931 Aleurocanthus spiniferus (Quaintance) 1903 

Aleurocanthus rugosa Singh 1931 Aleurocanthus rugosa Singh 1931 

Aleurocanthus russellae Jesudasan & David 1991 Aleurocanthus russellae Jesudasan & David 1991 

Aleurocanthus satyanarayani Dubey & Sundararaj 2005 Aleurocanthus satyanarayana Dubey & Sundararaj 2005 

Aleurocanthus serratus Quaintance & Baker 1917 Aleurocanthus serratus Quaintance & Baker 1917 

Aleurocanthus seshadrii David & Subramaniam 1976 Aleurocanthus seshadrii David & Subramaniam 1976 

Aleurocanthus shillongensis Jesudasan & David 1991 Aleurocanthus shillongensis Jesudasan & David 1991 

Aleurocanthus siamensis Takahashi 1942 Aleurocanthus siasmensis Takahashi 1942 

Aleurocanthus signatus [sic] (Maskell) 1895 Aleurocanthus t-signatus (Maskell) 1895 

Aleurocanthus simplex Singh 1931 Singhiella simplex (Singh) 1931 

Aleurocanthus singhi Jesudasan & David 1991 Aleurocanthus singhi Jesudasan & David 1991 

Aleurocanthus spiniferus (Quaintance) 1903 Aleurocanthus spiniferus (Quaintance) 1903 

Aleurocanthus spiniferus Quaintance Aleurocanthus spiniferus Quaintance 

Aleurocanthus spiniferus var. intermedia Silvestri 1927 Aleurocanthus spiniferus (Quaintance) 1903 

Aleurocanthus spinithorax Dumbleton 1961 Aleurocanthus spinithorax Dumbleton 1961 

Aleurocanthus spinosus (Kuwana) 1911 Aleurocanthus spinosus (Kuwana) 1911 

Aleurocanthus splendens David & Subramaniam 1976 Aleurocanthus splendens David & Subramaniam 1976 

Aleurocanthus strychnosicola Cohic 1966 Aleurocanthus strychnosicola Cohic 1966 

Aleurocanthus t-signatus (Maskell) 1895 Aleurocanthus t-signatus (Maskell) 1895 

Aleurocanthus terminaliae Dubey & Sundararaj 2005 Aleurocanthus terminaliae Dubey & Sundararaj 2005 

Aleurocanthus trispina Mound 1965 Aleurocanthus trispina Mound 1965 

Aleurocanthus uvariae Cohic 1968 Cohicaleyrodes uvariae (Cohic) 1968 

Aleurocanthus valenciae Martin 1999 Aleurocanthus valenciae Martin 1999 

Aleurocanthus valparaiensis David & Subramaniam 1976 Aleurocanthus valparaiensis David & Subramaniam 1976 

Aleurocanthus vindhyachali Dubey & Sundararaj 2005 Aleurocanthus vindhyhachali Dubey & Sundararaj 2005 

Aleurocanthus voeltzkowi (Newstead) 1908 Aleurocanthus voeltzkowi (Newstead) 1908 

Aleurocanthus woglumi Ashby 1915 Aleurocanthus woglumi Ashby 1915 

Aleurocanthus woglumi var. formosana Takahashi 1935 Aleurocanthus woglumi Ashby 1915 

Aleurocanthus yusopei Corbett 1935 Aleurocanthus gateri Corbett 1927 

Aleurocanthus zizyphi Priesner & Hosny 1934 Aleurocanthus zizyphi Priesner & Hosny 1934 

ALEUROCERUS BONDAR 1923 ALEUROCERUS BONDAR 1923 

Aleurocerus ceriferus (Sampson & Drew) 1941 Aleurocerus ceriferus (Sampson & Drew) 1941 

Aleurocerus chiclensis Russell 1986 Aleurocerus chiclensis Russell 1986 

Aleurocerus coccolobae Russell 1986 Aleurocerus coccolobae Russell 1986 

Aleurocerus colombiae Russell 1986 Aleurocerus colombiae Russell 1986 

Aleurocerus flavomarginatus Bondar 1923 Aleurocerus flavomarginatus Bondar 1923 

Aleurocerus luxuriosus Bondar 1923 Aleurocerus luxuriosus Bondar 1923 

Aleurocerus musae Russell 1986 Aleurocerus musae Russell 1986 

Aleurocerus palmae Russell 1986 Aleurocerus palmae Russell 1986 

Aleurocerus petiolicola Russell 1986 Aleurocerus petiolicola Russell 1986 

641

Aleurocerus tumidosus Bondar 1923 Aleurocerus tumidosus Bondar 1923 

ALEUROCHITON TULLGREN 1907 ALEUROCHITON TULLGREN 1907 

Aleurochiton (Nealeurochiton) Sampson 1943 Aleurochiton Tullgren 1907 

Aleurochiton acerina Haupt 1934 Aleurochiton acerinus Haupt 1934 

Aleurochiton acerinus Haupt 1934 Aleurochiton acerinus Haupt 1934 

Aleurochiton aceris (Modeer) 1762 Aleurochiton aceris (Modeer) 1778 

Aleurochiton aceris (Modeer) 1778 Aleurochiton aceris (Modeer) 1778 

Aleurochiton avellanae (Signoret) 1868 Asterobemisia carpini (Koch) 1857 

Aleurochiton complanatus (Baerensprung) 1849 Aleurochiton aceris (Modeer) 1778 

Aleurochiton complanatus Baerensprung 1849 Aleurochiton aceris (Modeer) 1778 

Aleurochiton dubius (Heeger) 1859 Siphoninus phillyreae (Haliday) 1835 

Aleurochiton forbesii (Ashmead) 1893 Aleurochiton forbesii (Ashmead) 1893 

Aleurochiton immaculatus (Heeger) 1856 Siphoninus immaculatus (Heeger) 1856 

Aleurochiton orientalis Danzig 1966 Aleurochiton orientalis Danzig 1966 

Aleurochiton petri Rietschel 1983 Aleurochiton petri Rietschel 1983 

Aleurochiton pseudoplatani Visnya 1936 Aleurochiton pseudoplatani Visnya 1936 

Aleurochiton quercus (Signoret) Pealius quercus (Signoret) 1868 

Aleurochiton quercus (Signoret) 1868 Pealius quercus (Signoret) 1868 

Aleurochiton vaccinii (Kunow); misid. Ryberg 1938 (in part) Aleuroclava similis (Takahashi) 1938 

Aleurochiton vaccinii Kunow 1880 Asterobemisia carpini (Koch) 1857 

ALEUROCLAVA SINGH 1931 ALEUROCLAVA SINGH 1931 

Aleuroclava afriae Sundararaj & David 1995 Aleuroclava afriae Sundararaj & David 1995 

Aleuroclava angkorensis (Takahashi) 1942 Aleuroclava angkorensis (Takahashi) 1942 

Aleuroclava artocarpi (Corbett) 1935 Aleuroclava artocarpi (Corbett) 1935 

Aleuroclava aucubae (Kuwana) 1911 Aleuroclava aucubae (Kuwana) 1911 

Aleuroclava baccaureae (Corbett) 1935 Aleuroclava baccaureae (Corbett) 1935 

Aleuroclava bauhiniae (Corbett) 1935 Aleuroclava bauhiniae (Corbett) 1935 

Aleuroclava bifurcata (Corbett) 1933 Aleuroclava bifurcata (Corbett) 1933 

Aleuroclava bilineata Sundararaj & David 1993 Aleuroclava bilineata Sundararaj & David 1993 

Aleuroclava burmanicus (Singh) 1938 Aleuroclava burmanicus (Singh) 1938 

Aleuroclava calicutensis Dubey & Sundararaj 2005 Aleuroclava calicutensis Dubey & Sundararaj 2005 

Aleuroclava caloncobae (Cohic) 1966 Aleuroclava nigeriae (Mound) 1965 

Aleuroclava calycopteriseae Dubey & Sundararaj 2005 Aleuroclava calycopteriseae Dubey & Sundararaj 2005 

Aleuroclava canangae (Corbett) 1935 Martiniella canangae (Corbett) 1935 

Aleuroclava cardamomi (David & Subramaniam) 1976 Singhiella cardamomi (David & Subramaniam) 1976 

Aleuroclava carpini (Takahashi) 1939 Aleuroclava carpini (Takahashi) 1939 

Aleuroclava carpini var. yushuniae (Takahashi) 1939 Aleuroclava carpini (Takahashi) 1939 

Aleuroclava cherasensis (Corbett) 1935 Minutaleyrodes cherasensis (Corbett) 1935 

Aleuroclava chittendeni (Laing) 1928 Massilieurodes chittendeni (Laing) 1928 

Aleuroclava cinnamomi Jesudasan & David 1991 Aleuroclava cinnamomi Jesudasan & David 1991 

Aleuroclava citri Jesudasan & David 1991 Aleuroclava citri Jesudasan & David 1991 

642

Aleuroclava citrifolii (Corbett) 1935 Aleuroclava citrifolii (Corbett) 1935 

Aleuroclava complex Singh 1931 Aleuroclava complex Singh 1931 

Aleuroclava davidi Dubey & Sundararaj 2005 Aleuroclava tarennae Dubey & Ko 2006 

Aleuroclava dehradunensis Jesudasan & David 1991 Aleuroclava dehradunensis Jesudasan & David 1991 

Aleuroclava doddabettaensis Dubey & Sundararaj 2005 Aleuroclava doddabettaensis Dubey & Sundararaj 2005 

Aleuroclava doveri (Corbett) 1935 Aleuroclava doveri (Corbett) 1935 

Aleuroclava dubius (Bink-Moenen) 1983 Aleuroclava dubius (Bink-Moenen) 1983 

Aleuroclava ehretiae Jesudasan & David 1991 Aleuroclava ehretiae Jesudasan & David 1991 

Aleuroclava elatostemae (Takahashi) 1932 Aleuroclava elatostemae (Takahashi) 1932 

Aleuroclava ellipticae Dumbleton 1956 Dumbletoniella ellipticae (Dumbleton) 1956 

Aleuroclava erythrinae (Corbett) 1935 Aleuroclava erythrinae (Corbett) 1935 

Aleuroclava eucalypti Dumbleton 1957 Dumbletoniella eucalypti (Dumbleton) 1957 

Aleuroclava eugeniae (Corbett) 1935 Aleuroclava eugeniae (Corbett) 1935 

Aleuroclava euphoriae (Takahashi) 1942 Aleuroclava euphoriae (Takahashi) 1942 

Aleuroclava euryae (Kuwana) 1911 Aleuroclava euryae (Kuwana) 1911 

Aleuroclava evanantiae Jesudasan & David 1991 Aleuroclava evanantiae Jesudasan & David 1991 

Aleuroclava fici (Corbett) 1935 Aleuroclava fici (Corbett) 1935 

Aleuroclava ficicola (Takahashi) 1932 Aleuroclava ficicola (Takahashi) 1932 

Aleuroclava filamentosa (Corbett) 1933 Aleuroclava filamentosa (Corbett) 1933 

Aleuroclava flabellus (Takahashi) 1949 Aleuroclava flabellus (Takahashi) 1949 

Aleuroclava fletcheri (Sundararaj & David) 1992 Aleuroclava fletcheri (Sundararaj & David) 1992 

Aleuroclava goaensis Jesudasan & David 1991 Aleuroclava goaensis Jesudasan & David 1991 

Aleuroclava gordoniae (Takahashi) 1932 Aleuroclava gordoniae (Takahashi) 1932 

Aleuroclava grewiae Sundararaj & David 1993 Aleuroclava grewiae Sundararaj & David 1993 

Aleuroclava guyavae (Takahashi) 1932 Aleuroclava guyavae (Takahashi) 1932 

Aleuroclava hexcantha (Singh) 1940 Aleuroclava hexcantha (Singh) 1940 

Aleuroclava hikosanensis (Takahashi) 1938 Aleuroclava hikosanensis (Takahashi) 1938 

Aleuroclava hindustanicus (Meganathan & David) 1994 Aleuroclava hindustanicus (Meganathan & David) 1994 

Aleuroclava indica (Singh) 1931 Aleuroclava indica (Singh) 1931 

Aleuroclava indica Meganathan & David 1994 Aleuroclava kerala Martin & Mound 2007 

Aleuroclava jasmini (Takahashi) 1932 Aleuroclava jasmini (Takahashi) 1932 

Aleuroclava kanyakumariensis Sundararaj & David 1993 Aleuroclava kanyakumariensis Sundararaj & David 1993 

Aleuroclava kavalurensis Jesudasan & David 1991 Aleuroclava kavalurensis Jesudasan & David 1991 

Aleuroclava kerala Martin & Mound 2007 Aleuroclava kerala Martin & Mound 2007 

Aleuroclava kolliensis (David) 1977 Minutaleyrodes kolliensis (David) 1977 

Aleuroclava kudremukekhensis Dubey & Sundararaj 2005 Aleuroclava kudremukhensis Dubey & Sundararaj 2005 

Aleuroclava kusheriki (Mound) 1965 Aleuroclava nigeriae (Mound) 1965 

Aleuroclava kuwanai (Takahashi) 1934 Aleuroclava kuwanai (Takahashi) 1934 

Aleuroclava lagerstroemiae (Takahashi) 1934 Aleuroclava lagerstroemiae (Takahashi) 1934 

Aleuroclava lanceolata (Takahashi) 1949 Aleuroclava lanceolata (Takahashi) 1949 

Aleuroclava latus (Takahashi) 1934 Aleuroclava latus (Takahashi) 1934 

643

Aleuroclava lefroyi Sundararaj & David 1993 Aleuroclava lefroyi Sundararaj & David 1993 

Aleuroclava lithocarpi (Takahashi) 1934 Aleuroclava lithocarpi (Takahashi) 1934 

Aleuroclava longisetosus Jesudasan & David 1991 Aleuroclava longisetosus Jesudasan & David 1991 

Aleuroclava longispinus (Takahashi) 1934 Aleuroclava longispinus (Takahashi) 1934 

Aleuroclava louiseae Sundararaj & David 1993 Aleuroclava louiseae Sundararaj & David 1993 

Aleuroclava macarangae (Corbett) 1935 Martiniella macarangae (Corbett) 1935 

Aleuroclava madhucae Jesudasan & David 1991 Aleuroclava madhucae Jesudasan & David 1991 

Aleuroclava magnoliae (Takahashi) 1952 Aleuroclava magnoliae (Takahashi) 1952 

Aleuroclava malloti (Takahashi) 1932 Aleuroclava malloti (Takahashi) 1932 

Aleuroclava manii (David) 1978 Aleuroclava manii (David) 1978 

Aleuroclava martini Dubey & Sundararaj 2005 Aleuroclava martini Dubey & Sundararaj 2005 

Aleuroclava melastomae (Takahashi) 1934 Aleuroclava melastomae (Takahashi) 1934 

Aleuroclava meliosmae (Takahashi) 1932 Aleuroclava meliosmae (Takahashi) 1932 

Aleuroclava minutus (Singh) 1931 Minutaleyrodes minuta (Singh) 1931 

Aleuroclava montanus (Takahash) 1939 Aleuroclava montanus (Takahashi) 1939 

Aleuroclava multipori (Takahashi) 1935 Aleuroclava multipori (Takahashi) 1935 

Aleuroclava multituberculata Sundararaj & David 1993 Aleuroclava multituberculata Sundararaj & David 1993 

Aleuroclava murrayae (Singh) 1931 Aleuroclava murrayae (Singh) 1931 

Aleuroclava mysorensis Jesudasan & David 1991 Aleuroclava mysorensis Jesudasan & David 1991 

Aleuroclava nachiensis (Takahashi) 1963 Aleuroclava nachiensis (Takahashi) 1963 

Aleuroclava nagercoilensis Sundararaj & David 1993 Aleuroclava nagercoilensis Sundararaj & David 1993 

Aleuroclava nanjangudensis Jesudasan & David 1991 Aleuroclava nanjangudensis Jesudasan & David 1991 

Aleuroclava neolitseae (Takahashi) 1934 Aleuroclava neolitseae (Takahashi) 1934 

Aleuroclava nephelii (Corbett) 1935 Aleuroclava nephelii (Corbett) 1935 

Aleuroclava nigeriae (Mound) 1965 Aleuroclava nigeriae (Mound) 1965 

Aleuroclava nitidus (Singh) 1932 Aleuroclava nitidus (Singh) 1932 

Aleuroclava orientalis Jesudasan & David 1991 Aleuroclava orientalis Jesudasan & David 1991 

Aleuroclava papillata Dubey & Sundararaj 2005 Aleuroclava subindica Martin & Mound 2007 

Aleuroclava parvus (Singh) 1938 Aleuroclava parvus (Singh) 1938 

Aleuroclava pentatuberculata Sundararaj & David 1993 Aleuroclava pentatuberculata Sundararaj & David 1993 

Aleuroclava philomenae Jesudasan & David 1991 Aleuroclava philomenae Jesudasan & David 1991 

Aleuroclava phyllanthi (Corbett) 1935 Aleuroclava phyllanthi (Corbett) 1935 

Aleuroclava piperis (Takahashi) 1935 Aleuroclava piperis (Takahashi) 1935 

Aleuroclava pongamiae Jesudasan & David 1991 Aleuroclava pongamiae Jesudasan & David 1991 

Aleuroclava porosus (Priesner & Hosny) 1937 Aleuroclava porosus (Priesner & Hosny) 1937 

Aleuroclava psidii (Singh) 1931 Aleuroclava psidii (Singh) 1931 

Aleuroclava pulcherrimus (Corbett) 1935 Aleuroclava pulcherrima (Corbett) 1935 

Aleuroclava pyracanthae (Takahashi) 1933 Aleuroclava pyracanthi (Takahashi) 1933 

Aleuroclava regui Sundararaj & David 1993 Aleuroclava regui Sundararaj & David 1993 

Aleuroclava rhododendri (Takahashi) 1935 Aleuroclava rhododendri (Takahashi) 1935 

Aleuroclava russellae (David & Subramaniam) 1976 Aleuroclava russellae (David & Subramaniam) 

644

Aleuroclava sandorici (Corbett) 1935 Aleuroclava nitidus (Singh) 1932 

Aleuroclava saputarensis Sundararaj & David 1993 Aleuroclava saputarensis Sundararaj & David 1993 

Aleuroclava selvakumarani Sundararaj & David 1993 Aleuroclava selvakumarani Sundararaj & David 1993 

Aleuroclava sepangensis Martin & Mound 2007 Aleuroclava sepangensis Martin & Mound 2007 

Aleuroclava siamensis (Takahashi) 1942 Aleuroclava siamensis (Takahashi) 1942 

Aleuroclava similis (Takahashi) 1938 Aleuroclava similis (Takahashi) 1938 

Aleuroclava similis europaeus (Zahradnik) Aleuroclava similis (Takahashi) 1938 

Aleuroclava simplex (Takahashi) 1949 Aleuroclava simplex (Takahashi) 1949 

Aleuroclava singhi Jesudasan & David 1991 Aleuroclava singhi Jesudasan & David 1991 

Aleuroclava sivakasiensis Sundararaj & David 1993 Aleuroclava sivakasiensis Sundararaj & David 1993 

Aleuroclava srilankaensis (David) 1993 Aleuroclava srilankaensis (David) 1993 

Aleuroclava stereospermi (Corbett) 1935 Aleuroclava stereospermi (Corbett) 1935 

Aleuroclava subindica Martin & Mound 2007 Aleuroclava subindica Martin & Mound 2007 

Aleuroclava suishanus (Takahashi) 1934 Minutaleyrodes suishanus (Takahashi) 1934 

Aleuroclava takahashii (David & Subramaniam) 1976 Aleuroclava takahashii (David & Subramaniam) 1976 

Aleuroclava tentactuliformis (Corbett) 1935 Aleuroclava tentactuliformis (Corbett) 1935 

Aleuroclava terminaliae Sundararaj & David 1993 Aleuroclava terminaliae Sundararaj & David 1993 

Aleuroclava thysanospermi (Takahashi) 1934 Aleuroclava thysanospermi (Takahashi) 1934 

Aleuroclava trachelospermi (Takahashi) Aleuroclava trachelospemi (Takahashi) 1938 

Aleuroclava trivandricus Dubey & Sundararaj 2006 Aleuroclava trivandricus Dubey & Sundararaj 2006 

Aleuroclava trilineata Sundararaj & David 1993 Aleuroclava trilineata Sundararaj & David 1993 

Aleuroclava tripori (Dubey & Sundararaj)2006 Aleuroclava tripori (Dubey & Sundararaj)2006 

Aleuroclava trochodendri Takahashi 1957 Aleuroclava trochodendri Takahashi 1957 

Aleuroclava ubonensis (Takahashi) 1942 Aleuroclava ubonensis (Takahashi) 1942 

Aleuroclava uraianus (Takahashi) 1932 Aleuroclava uraianus (Takahashi) 1932 

Aleuroclava vernoniae Meganathan & David 1994 Aleuroclava vernoniae Meganathan & David 1994 

Aleuroclava vitexae Sundararaj & David 1993 Aleuroclava vitexae Sundararaj & David 1993 

Aleuroclava wrightiae Jesudasan & David 1991 Aleuroclava wrightiae Jesudasan & David 1991 

Aleuroclava yambiae (Gameel) 1971 Aleuroclava nigeriae (Mound) 1965 

ALEUROCYBOTUS QUAINTANCE & BAKER 1914 ALEUROCYBOTUS QUAINTANCE & BAKER 1914 

Aleurocybotus cereus Martin 2005 Aleurocybotus cereus Martin 2005 

Aleurocybotus cobarensis Martin 1999 Vasdavidius cobarensis (Martin) 1999 

Aleurocybotus concursus Ko 1998 Vasdavidius concursus (Ko) 1998 

Aleurocybotus graminicolus (Quaintance) 1899 Aleurocybotus graminicolus (Quaintance) 1899 

Aleurocybotus indicus David & Subramaniam 1976 Vasdavidius indicus (David & Subramaniam) 1976 

Aleurocybotus millettiae (Cohic) 1968 Aleurocybotus millettiae (Cohic) 1968 

Aleurocybotus miscanthus Ko 1998 Vasdavidius miscanthus (Ko) 1998 

Aleurocybotus occiduus Russell 1964 Aleurocybotus occiduus Russell 1964 

Aleurocybotus setiferus Quaintance & Baker Vasdavidius setiferus (Quaintance & Baker) 

Aleurocybotus setiferus Quaintance & Baker 1917 Vasdavidius setiferus (Quaintance & Baker) 1917 

ALEUROCYPERUS KO & DUBEY 2007 ALEUROCYPERUS KO & DUBEY 2007 

645

Aleurocyperus humus Ko & Dubey 2007 Aleurocyperus humus Ko & Dubey 2007 

Aleurodes (Ogivaleurodes) Goux 1948 Trialeurodes Cockerell 1902 

Aleurodes abutiloneus Haldeman 1850 Trialeurodes abutiloneus (Haldeman) 1850 

Aleurodes acaciae Quaintance 1900 Tetraleurodes acaciae (Quaintance) 1900 

Aleurodes aceris Bouche 1851 Aleurochiton aceris (Modeer) 1778 

Aleurodes aceris Forbes 1885 Aleurochiton forbesii (Ashmead) 1885 

Aleurodes aepim Goldi 1886 Aleurothrixus aepim (Goldi) 1886 

Aleurodes africana Newstead 1921 Dialeurolonga africana (Newstead) 1921 

Aleurodes albofloccosa Froggatt 1918 Aleurodicus destructor Mackie 1912 

Aleurodes alcocki Peal 1903 Aleuroplatus alcocki (Peal) 1903 

Aleurodes altissima Quaintance 1900 Nealeurodicus altissimus (Quaintance) 1900 

Aleurodes asari Wunn 1926 Aleurolobus wunni (Ryberg) 1938 

Aleurodes asparagi Lewis 1893 Aleurotrachelus asparagi (Lewis) 1893 

Aleurodes asplenii Maskell 1890 Trialeurodes asplenii (Maskell) 1890 

Aleurodes avellanae Signoret 1868 Asterobemisia carpini (Koch) 1857 

Aleurodes bambusae Peal 1903 Aleurocanthus bambusae (Peal) 1903 

Aleurodes banksiae Maskell 1895 Aleurocanthus banksiae (Maskell) 1895 

Aleurodes barodensis Maskell 1895 Aleurolobus barodensis (Maskell) 1895 

Aleurodes bengalensis Peal 1903 Pealius bengalensis (Peal) 1903 

Aleurodes bergii Signoret 1868 Neomaskellia bergii (Signoret) 1868 

Aleurodes bragini Mokrzecki 1916 nomen nudum 

Aleurodes carpini Koch 1857 Asterobemisia carpini (Koch) 1857 

Aleurodes cerata Maskell 1895 Asterochiton cerata (Maskell) 1895 

Aleurodes citri Riley & Howard 1893 Dialeurodes citri (Ashmead) 1885 

Aleurodes citricola Newstead 1911 Aleurocanthus spiniferus (Quaintance) 1903 

Aleurodes citrifolii Foster 1892 Dialeurodes citri (Ashmead) 1885 

Aleurodes cockerelli Ihering 1897 Aleuroplatus cockerelli (Ihering) 1897 

Aleurodes comata Maskell 1895 Neomaskellia bergii (Signoret) 1868 

Aleurodes corni Haldeman 1850 Tetraleurodes corni (Haldeman) 1850 

Aleurodes coronata Quaintance 1900 Aleuroplatus coronatus (Quaintance) 1900 

Aleurodes cotesii Maskell 1895 Bulgarialeurodes cotesii (Maskell) 1895 

Aleurodes crataegi Kiriukhin 1947 Aleyrodes crataegi Kiriukhin 1947 

Aleurodes croceata Maskell 1895 Aleuroduplidens croceata (Maskell) 1895 

Aleurodes croceata Maskell 1896 Aleuroduplidens croceata (Maskell) 1896 

Aleurodes decipiens Maskell 1895 Bemisia decipiens (Maskell) 1895 

Aleurodes decipiens Maskell 1895; misidentifation in part. Dumbletoniella ellipticae Dumbleton 1956 

Aleurodes dubia Heeger 1859 Siphoninus phillyreae (Haliday) 1835 

Aleurodes erigerontis Maskell 1895 Trialeurodes abutiloneus (Haldeman) 1850 

Aleurodes erigerontis Maskell 1896 Trialeurodes abutiloneus (Haldeman) 1850 

Aleurodes eugeniae Maskell 1895 Rusostigma eugeniae (Maskell) 1895 

Aleurodes eugeniae var. aurantii Maskell 1895 Dialeurodes citri (Ashmead) 1885 

646

Aleurodes fagi Maskell 1890 Asterochiton fagi (Maskell) 1890 

Aleurodes filicium Goeldi; misidentification in part Aleurotulus nephrolepidis (Quaintance) 1900 

Aleurodes fitchi Quaintance 1900 Trialeurodes abutiloneus (Haldeman) 1850 

Aleurodes floccosa Maskell 1895 Aleurothrixus floccosus (Maskell) 1895 

Aleurodes floccosus Maskell 1895 Aleurothrixus floccosus (Maskell) 1895 

Aleurodes floridensis Quaintance 1900 Trialeurodes floridensis (Quaintance) 1900 

Aleurodes fodiens Maskell 1895 Aleyrodes fodiens Maskell 1895 

Aleurodes forbesii Ashmead 1893 Aleurochiton forbesii (Ashmead) 1893 

Aleurodes fumipennis Hempel 1899 Aleurotrachelus fumipennis (Hempel) 1899 

Aleurodes gelatinosus Cockerell 1898 Aleuroplatus gelatinosus (Cockerell) 1898 

Aleurodes goyabae Goldi 1886 Paraleyrodes goyabae (Goldi) 1886 

Aleurodes graminicolus Quaintance 1899 Aleurocybotus graminicolus (Quaintance) 1899 

Aleurodes herbericola Cockerell 1896 Bemisia berbericola (Cockerell) 1896 

Aleurodes hirsuta Maskell 1895 Aleurocanthus hirsutus (Maskell) 1895 

Aleurodes holmesii Maskell 1895 Aleurodicus holmesii (Maskell) 1895 

Aleurodes hoyae Peal 1903 Aleuroplatus hoyae (Peal) 1903 

Aleurodes immaculatus Heeger 1856 Siphoninus immaculatus (Heeger) 1856 

Aleurodes inconspicua Quaintance 1900 Bemisia tabaci (Gennadius) 1889 

Aleurodes jelinekii Frauenfeld 1867 Aleurotuba jelinekii (Frauenfeld) 1867 

Aleurodes lactea Zehntner 1897 Aleyrodes lactea Zehntner 1897 

Aleurodes lauri Signoret 1882 Trialeurodes lauri (Signoret) 1882 

647

Aleurodes leakii Peal 1903 Bemisia leakii (Peal) 1903 

Aleurodes limbatus Maskell 1895 Aleurotrachelus limbatus (Maskell) 1895 

Aleurodes longicornis Zehntner 1897 Aleurolobus barodensis (Maskell) 1895 

Aleurodes lonicerae Koch 1857 Aleyrodes lonicerae Walker 1852 

648 

Aleurodes marginata Newstead 1911 Aleurotrachelus marginatus (Newstead) 1911 

Aleurodes marlatti Quaintance 1903 Aleurolobus marlatti (Quaintance) 1903 

Aleurodes melicyti Maskell 1890 Asterochiton aureus Maskell 1879 

Aleurodes mirabilis Cockerell 1898 Lecanoideus mirabilis (Cockerell) 1898 

Aleurodes mori Quaintance 1899 Tetraleurodes mori (Quaintance) 1899 

Aleurodes nephrolepidis Quaintance 1900 Aleurotulus nephrolepidis (Quaintance) 1900 

Aleurodes nicotiana Maskell 1895 Trialeurodes vaporariorum (Westwood) 1856 

Aleurodes niger Maskell 1895 Zaphanera niger (Maskell) 1895 

Aleurodes nubilans Buckton 1900 nomen dubium 

Aleurodes olivinus Silvestri 1911 Aleurolobus olivinus (Silvestri) 1911 

Aleurodes palatina Wunn. Trioza palatina (Wunn.) 

Aleurodes parvus Hempel 1899 Aleurotrachelus parvus (Hempel) 1899 

Aleurodes pergandei Quaintance 1900 Trialeurodes pergandei (Quaintance) 1900 

Aleurodes perseae Quaintance 1900 Paraleyrodes perseae (Quaintance) 1900 

Aleurodes phalaenoides Blanchard 1852 Metaleurodicus phalaenoides (Blanchard) 1852 

Aleurodes phalaroides Lataste 1852 nomen nudum 

Aleurodes phaseoli Schilling nomen nudum 

Aleurodes phillyreae Haliday 1835 Siphoninus phillyreae (Haliday) 1835 

Aleurodes phylliceae Bouche 1851 Siphoninus phillyreae (Haliday) 1835 

Aleurodes pinicola Kirkaldy 1907 nomen nudum 

Aleurodes piperis Maskell 1895 Aleurocanthus piperis (Maskell) 1895 

Aleurodes plumosa Quaintance 1900 Aleuroplatus plumosus (Quaintance) 1900 

Aleurodes pulvinatus Maskell 1895 Aleurodicus pulvinatus (Maskell) 1895 

Aleurodes quaintancei Peal 1903 Aleuroplatus quaintancei (Peal) 1903 

Aleurodes quercus Signoret 1868 Pealius quercus (Signoret) 1868 

Aleurodes quercus Signoret; Gomez-Menor 1945, misid Aleuroviggianus polymorphus Bink-Moenen 1992 

Aleurodes quercus-aquaticae Quaintance 1900 Aleuroplatus quercusaquaticae (Quaintance) 1900 

Aleurodes religiosa Peal 1903 Bemisia religiosa (Peal) 1903 

Aleurodes ribium Douglas 1888 Asterobemisia carpini (Koch) 1857 

Aleurodes rolfsii Quaintance 1899 Trialeurodes abutiloneus (Haldeman) 1850 

Aleurodes rosae Kiriukhin 1947 Bulgarialeurodes cotesii (Maskell) 1895 

Aleurodes rubicola Douglas 1891 Asterobemisia carpini (Koch) 1857 

Aleurodes ruborum Cockerell 1897 Trialeurodes ruborum (Cockerell) 1897 

Aleurodes sacchari Maskell 1890 Neomaskellia bergii (Signoret) 1868 

Aleurodes simplex Maskell 1890 Asterochiton simplex (Maskell) 1890

Aleurodes simula Peal 1903 Africaleurodes simula (Peal) 1903 

Aleurodes spinifera Quaintance 1903 Aleurocanthus spiniferus (Quaintance) 1903 

Aleurodes spinosus Kuwana; misidentification Aleurocanthus cinnamomi Takahashi 1913 

Aleurodes spiraeoides Quaintance 1900 Aleyrodes spiraeoides Quaintance 1900 

Aleurodes stellata Maskell 1895 Tetraleurodes stellata (Maskell) 1895 

Aleurodes stypheliae Maskell 1895 Aleurotrachelus stypheliae (Maskell) 1895 

Aleurodes t-signatus Maskell 1895 Aleurocanthus t-signatus (Maskell) 1895 

Aleurodes tabaci Gennadius 1889 Bemisia tabaci (Gennadius) 1889 

Aleurodes tinaeoides Blanchard 1852 Aleyrodes tinaeoides Blanchard 1852 

649 

Aleurodes tracheifer Quaintance 1900 Aleurotrachelus tracheifer (Quaintance) 1900 

Aleurodes vaporariorum Westwood 1856 Trialeurodes vaporariorum (Westwood) 1856 

Aleurodes variabilis Quaintance 1900 Trialeurodes variabilis (Quaintance) 1900 

Aleurodes vinsonioides Cockerell 1898 Aleuroplatus vinsonioides (Cockerell) 1898 

Aleurodes vittata Quaintance 1900 Trialeurodes vittata (Quaintance) 

Aleurodes voeltzkowi Newstead 1908 Aleurocanthus voeltzkowi (Newstead) 1908 

Aleurodes wunni Ryberg 1938 Aleurolobus wunni (Ryberg) 1938 

Aleurodes zimmermanni Newstead 1911 Pogonaleyrodes zimmermanni (Newstead) 1911 

ALEURODICUS DOUGLAS 1892 ALEURODICUS DOUGLAS 1892 

Aleurodicus aculeatus Menge 1856 Aleurodicus aculeatus Menge 1856 

Aleurodicus altissima Quaintance 1900 Nealeurodicus altissimus (Quaintance) 1900 

Aleurodicus anonae Morgan 1892 Aleurodicus cocois (Curtis) 1846 

Aleurodicus antidesmae Corbett 1926 Aleurodicus antidesmae Corbett 1926 

Aleurodicus antillensis Dozier 1936 Aleurodicus antillensis Dozier 1936 

Aleurodicus araujoi Sampson & Drew 1941 Aleurodicus araujoi Sampson & Drew 1941 

Aleurodicus asarumis (Shimer) 1867 Aleyrodes asarumis Shimer 1867 

Aleurodicus bifasciatus Bondar 1922 Aleurodicus pulvinatus (Maskell) 1895 

Aleurodicus bondari Costa Lima 1928 Aleuronudus bondari (Costa Lima) 1928 

Aleurodicus burmiticus Cockerell 1919 Aleurodicus burmiticus Cockerell 1919 

Aleurodicus capiangae Bondar 1923 Aleurodicus capiangae Bondar 1923 

Aleurodicus cardini Back 1912 Metaleurodicus cardini (Back) 1912 

Aleurodicus cinnamomi Takahashi 1951 Aleurodicus cinnamomi Takahashi 1951 

Aleurodicus coccolobae Quaintance & Baker 1913 Aleurodicus coccolobae Quaintance & Baker 1913 

Aleurodicus cockerellii Quaintance 1900 Dialeurodicus cockerellii (Quaintance) 1900 

Aleurodicus cocois (Curtis) 1846 Aleurodicus cocois (Curtis) 1846 

Aleurodicus conspurcatus Enderlein 1909 Bakerius conspurcatus (Enderlein) 1909 

Aleurodicus destructor Mackie 1912 Aleurodicus destructor Mackie 1912 

Aleurodicus dispersus Russell 1965 Aleurodicus dispersus Russell 1965 

Aleurodicus dugesii Cockerell 1896 Aleurodicus dugesii Cockerell 1896 

Aleurodicus essigi Sampson & Drews 1941 Aleurodicus essigi Sampson & Drews 1941

Aleurodicus flavus Hempel 1922 Aleurodicus flavus Hempel 1922 

Aleurodicus flumineus Hempel 1918 Aleurodicus neglectus Quaintance & Baker 1913 

Aleurodicus formosanus Takahashi 1932 Aleurodicus machili Takahashi 1931 

Aleurodicus fucatus Bondar 1923 Aleurodicus fucatus Bondar 1923 

Aleurodicus griseus Dozier 1936 Metaleurodicus griseus (Dozier) 1936 

Aleurodicus guppyii Quaintance & Baker 1913 Aleurodicus guppyii Quaintance & Baker 1913 

Aleurodicus holmesii (Maskell) 1895 Aleurodicus holmesii (Maskell) 1895 

Aleurodicus indicus Regu & David 1992 Aleurodicus indicus Regu & David 1992 

Aleurodicus inversus Martin 2004 Aleurodicus inversus Martin 2004 

Aleurodicus iridescens Cockerell 1898 Aleurodicus pulvinatus (Maskell) 1896 

Aleurodicus jamaicensis Cockerell 1902 Aleurodicus jamaicensis Cockerell 1902 

Aleurodicus juleikae Bondar 1923 Aleurodicus juleikae Bondar 1923 

Aleurodicus lacerdae (Signoret) 1883 Metaleurodicus lacerdae (Signoret) 1883 

Aleurodicus lahillei Leonardi 1910 Leonardius lahillei (Leonardi) 1910 

Aleurodicus linguosus Bondar 1923 Aleurodicus maritimus Hempel 1923 

Aleurodicus machili Takahashi 1931 Aleurodicus machili Takahashi 1931 

Aleurodicus magnificus Costa Lima 1928 Aleurodicus magnificus Costa Lima 1928 

Aleurodicus malayensis Takahashi 1951 Aleurodicus holmesii (Maskell) 1895 

Aleurodicus maritimus Hempel 1923 Aleurodicus maritimus Hempel 1923 

Aleurodicus marmoratus Hempel 1923 Aleurodicus marmoratus Hempel 1923 

Aleurodicus minima Quaintance 1900 Metaleurodicus minimus (Quaintance) 1900 

Aleurodicus neglectus Quaintance & Baker 1913 Aleurodicus neglectus Quaintance & Baker 1913 

Aleurodicus niveus Martin 2004 Aleurodicus niveus Martin 2004 

Aleurodicus ornatus Cockerell 1893 Aleurodicus ornatus Cockerell 1893 

Aleurodicus pauciporus Martin 2004 Aleurodicus pauciporus Martin 2004 

Aleurodicus perseae (Quaintance) 1900 Paraleyrodes perseae (Quaintance) 1900 

Aleurodicus phalaenoides (Blanchard) Metaleurodicus phalaenoides (Blanchard) 1852 

Aleurodicus philomenae David 1987 Aleurodicus machili Takahashi 1931 

Aleurodicus photiniana Chen Dialeuropora photiniana (Chen) 

Aleurodicus pigeanus Baker & Moles 1923 Australeurodicus pigeanus (Baker & Moles) 1923 

Aleurodicus pimentae Liang 1924, nomen nudum nomen nudum 

Aleurodicus poriferus Sampson & Drews 1941 Aleurodicus dugesii Cockerell 1896 

Aleurodicus pulvinatus (Maskell) 1895 Aleurodicus pulvinatus (Maskell) 1895 

Aleurodicus rugioperculatus Martin 2004 Aleurodicus rugioperculatus Martin 2004 

Aleurodicus silvestrii Leonardi 1910 Dialeurodicus silvestrii (Leonardi) 1910 

Aleurodicus talamancensis Martin 2005 Aleurodicus talamancensis Martin 2005 

Aleurodicus trinidadensis Quaintance & Baker 1913 Aleurodicus trinidadensis Quaintance & Baker 1913 

Aleurodicus vinculus Martin 2004 Aleurodicus vinculus Martin 2004 

Aleurodicus wallaceus Martin 1988 Aleurodicus wallaceus Martin 1988 

650

Aleurodicus (Dialeurodicus) Cockerell 1902 Dialeurodicus Cockerell 1902 

Aleurodicus (Dialeurodicus) cockerellii Quaintance 1900 Dialeurodicus cockerellii (Quaintance) 1900 

Aleurodicus (Lecanoideus) QUAINTANCE & BAKER 1913 Lecanoideus DOUGLAS 1892 

Aleurodicus (Lecanoideus) giganteus Quaintance & Baker 1913 Lecanoideus mirabilis (Cockerell) 1898 

Aleurodicus (Lecanoideus) mirabilis (Cockerell) 1898 Lecanoideus mirabilis (Cockerell) 1898 

Aleurodicus (Mesaleurodicus) altissimus (Quaintance) 1900 Nealeurodicus altissimus (Quaintance) 1900 

Aleurodicus (Metaleurodicus) Quaintance & Baker 1913 Metaleurodicus Quaintance & Baker 1913 

Aleurodicus (Metaleurodicus) altissimus (Quaintance) 1900 Nealeurodicus altissimus (Quaintance) 1900 

Aleurodicus (Metaleurodicus) cardini Back 1912 Metaleurodicus cardini (Back) 1912 

Aleurodicus (Metaleurodicus) manni Baker 1923 Aleuronudus manni (Baker) 1923 

Aleurodicus (Metaleurodicus) minimus Quaintance 1900 Metaleurodicus minimus (Quaintance) 1900 

Aleurodicus (Metaleurodicus) pigeanus Baker & Moles 1923 Australeurodicus pigeanus (Baker & Moles) 1923 

Aleurodicus (Metaleurodicus) variabilis (Quaintance) 1900 Trialeurodes variabilis (Quaintance) 1900 

ALEURODUPLIDENS MARTIN 1999 ALEURODUPLIDENS MARTIN 1999 

Aleuroduplidens carverae Martin 1999 Aleuroduplidens carverae Martin 1999 

Aleuroduplidens croceata (Maskell) 1896 Aleuroduplidens croceata (Maskell) 1896 

Aleuroduplidens eucalyptifolia Martin 1999 Aleuroduplidens eucalyptifolia Martin 1999 

Aleuroduplidens santali Martin 1999 Aleuroduplidens santali Martin 1999 

Aleuroduplidens triangularis Martin 1999 Aleuroduplidens triangularis Martin 1999 

Aleuroduplidens wellsae Martin 1999 Aleuroduplidens wellsae Martin 1999 

ALEUROGLANDULUS BONDAR 1923 ALEUROGLANDULUS BONDAR 1923 

Aleuroglandulus emmae Russell 1944 Aleuroglandulus subtilis Bondar 1923 

Aleuroglandulus inanis Martin 2005 Aleuroglandulus inanis Martin 2005 

Aleuroglandulus magnus Russell 1944 Aleuroglandulus magnus Russell 1944 

Aleuroglandulus malangae Russell 1944 Aleuroglandulus subtilis Bondar 1923 

Aleuroglandulus striatus Sampson & Drews 1941 Aleuroglandulus striatus Sampson & Drews 1941 

Aleuroglandulus striatus Sampson & Drews 1941; misid part Aleuroglandulus subtilis Bondar 1923 

Aleuroglandulus subtilis Bondar 1923 Aleuroglandulus subtilis Bondar 1923 

ALEUROINANIS MARTIN 1999 ALEUROINANIS MARTIN 1999 

Aleuroinanis myrtacei Martin 1999 Aleuroinanis myrtacei Martin 1999 

ALEUROLOBUS QUAINTANCE & BAKER 1914 ALEUROLOBUS QUAINTANCE & BAKER 1914 

Aleurolobus acanthi Takahashi 1936 Aleurolobus acanthi Takahashi 1936 

Aleurolobus antennata Regu & David 1993 Aleurolobus antennata Regu & David 1993 

Aleurolobus asari (Wunn) 1926 Aleurolobus wunni (Ryberg) 1938 

651 

Aleurolobus azadirachtae Regu & David 1993 Aleurolobus azadirachtae Regu & David 1993 

Aleurolobus azimae Jesudasan & David 1991 Aleurolobus azimae Jesudasan & David 1991 

Aleurolobus barleriae Jesudasan & David 1991 Aleurolobus barleriae Jesudasan & David 1991 

Aleurolobus barodensis (Maskell) 1895 Aleurolobus barodensis (Maskell) 1895 

Aleurolobus bidentatus Singh 1940 Aleurolobus bidentatus Singh 1940

Aleurolobus burliarensis Jesudasan & David 1991 Aleurolobus burliarensis Jesudasan & David 1991 

652 

Aleurolobus cassiae Jesudasan & David 1991 Aleurolobus cassiae Jesudasan & David 1991 

Aleurolobus cephalidistinctus Regu & David 1993 Aleurolobus cephalidistinctus Regu & David 1993 

Aleurolobus dalbergiae Dubey & Sundararaj Aleurolobus dalbergiae Dubey & Sundararaj 

Aleurolobus chinensis Takahashi 1936 Aleurolobus taeonabae (Kuwana) 1911 

Aleurolobus cissampelosae Regu & David 1993 Aleurolobus cissampelosae Regu & David 1993 

Aleurolobus citri Takahashi 1932 Africaleurodes citri (Takahashi) 1932 

Aleurolobus citrifolii Corbett 1935 Aleuroclava citrifolii (Corbett) 1935 

Aleurolobus clematidis Goux 1942 Aleurolobus wunni (Ryberg) 1938 

Aleurolobus cohici Regu & David 1993 Aleurolobus cohici Regu & David 1993 

Aleurolobus confusus David & Subramaniam 1976 Aleurolobus confusus David & Subramaniam 1976 

Aleurolobus delamarei Cohic 1969 Aleurolobus delamarei Cohic 1969 

Aleurolobus delhiensis Regu & David 1993 Aleurolobus delhiensis Regu & David 1993 

Aleurolobus diacritica Regu & David 1993 Aleurolobus diacritica Regu & David 1993 

Aleurolobus diastematus Bink-Moenen 1983 Aleurolobus diastematus Bink-Moenen 1983 

Aleurolobus distinctus Regu & David 1993 Aleurolobus distinctus Regu & David 1993 

Aleurolobus exceptionalis Regu & David 1993 Aleurolobus exceptionalis Regu & David 1993 

Aleurolobus flavus Quaintance & Baker 1917 Aleurolobus flavus Quaintance & Baker 1917 

Aleurolobus fouabii Cohic 1969 Aleurolobus fouabii Cohic 1969 

Aleurolobus gmelinae David, Jesudasan & Mathew 1988 Aleurolobus gmelinae David, Jesudasan & Mathew 1988 

Aleurolobus graminicola Bink-Moenen 1983 Aleurolobus graminicola Bink-Moenen 1983 

Aleurolobus greeni Corbett 1926 Aleurolobus greeni Corbett 1926 

Aleurolobus gruveli Cohic 1968 Aleurolobus gruveli Cohic 1968 

Aleurolobus hargreavesi Dozier 1934 Aleurolobus hargreavesi Dozier 1934 

Aleurolobus hederae Takahashi 1935 Aleurolobus hederae Takahashi 1935 

Aleurolobus hosurensis Regu & David 1993 Aleurolobus hosurensis Regu & David 1993 

Aleurolobus indigoferae Regu & David 1993 Aleurolobus indigoferae Regu & David 1993 

Aleurolobus iteae Takahashi 1957 Aleurolobus iteae Takahashi 1957 

Aleurolobus japonicus Takahashi 1954 Aleurolobus japonicus Takahashi 1954 

Aleurolobus juillieni Cohic 1968 Aleurolobus jullieni Cohic 1968 

Aleurolobus karunkuliensis Jesudasan & David 1991 Aleurolobus karunkuliensis Jesudasan & David 1991 

Aleurolobus lagerstroemia Regu & David 1993 Aleurolobus lagerstroemia Regu & David 1993 

Aleurolobus longisetosus Dubey & Sundararaj Aleurolobus longisetosus Dubey & Sundararaj 

Aleurolobus longicornis (Zehntner) 1897 Aleurolobus barodensis (Maskell) 1895 

Aleurolobus luci Cohic 1969 Aleurolobus luci Cohic 1969 

Aleurolobus macarangae Regu & David 1993 Aleurolobus macarangae Regu & David 1993 

Aleurolobus madrasensis Regu & David 1993 Aleurolobus madrasensis Regu & David 1993 

Aleurolobus marlatti (Quaintance) 1903 Aleurolobus marlatti (Quaintance) 1903 

Aleurolobus mauritanicus Cohic 1969 Aleurolobus mauritanicus Cohic 1969

Aleurolobus monodi Cohic 1969 Crescentaleyrodes monodi (Cohic) 1969 

Aleurolobus moundi David & Subramaniam 1976 Aleurolobus moundi David & Subramaniam 1976 

Aleurolobus musae Corbett 1935 Aleurolobus musae Corbett 1935 

Aleurolobus nagercoilensis Regu & David 1993 Aleurolobus nagercoilensis Regu & David 1993 

Aleurolobus niger (Maskell) Zaphanera niger (Maskell) 1895 

Aleurolobus niloticus Priesner & Hosny 1934 Aleurolobus marlatti (Quaintance) 1903 

Aleurolobus olivinus (Silvestri) 1911 Aleurolobus olivinus (Silvestri) 1911 

Aleurolobus onitshae Mound 1965 Aleurolobus onitshae Mound 1965 

Aleurolobus oplismeni Takahashi 1931 Aleurolobus oplismeni Takahashi 1931 

Aleurolobus orientalis David & Jesudasan 1988 Aleurolobus orientalis David & Jesudasan 1988 

Aleurolobus osmanthi Young 1944 Aleurolobus osmanthi Young 1944 

Aleurolobus ovalis Regu & David 1993 Aleurolobus ovalis Regu & David 1993 

Aleurolobus padappaiensis Regu & David 1993 Aleurolobus padappaiensis Regu & David 1993 

Aleurolobus panvelensis Regu & David 1993 Aleurolobus panvelensis Regu & David 1993 

Aleurolobus patchily Regu & David 1993 Aleurolobus patchily Regu & David 1993 

Aleurolobus paulianae Cohic 1969 Crescentaleyrodes paulianae (Cohic) 1969 

Aleurolobus pauliani Cohic 1969 Aleurolobus pauliani Cohic 1969 

Aleurolobus philippinensis Quaintance & Baker 1917 Aleurolobus philippinensis Quaintance & Baker 1917 

Aleurolobus phyllanthi Corbett 1935 Aleuroclava phyllanthi (Corbett) 1935 

Aleurolobus piliostigmatos Bink-Moenen 1983 Aleurolobus piliostigmatos Bink-Moenen 1983 

Aleurolobus psidii Jesudasan & David 1991 Aleurolobus psidii Jesudasan & David 1991 

Aleurolobus pulcherrimus Corbett 1935 Aleuroclava pulcherrima (Corbett) 1935 

Aleurolobus puripennis Ossiannilsson 1944 Aleurolobus wunni (Ryberg) 1938 

Aleurolobus ravisei Cohic 1968 Aleurolobus marlatti (Quaintance) 1903 

Aleurolobus rhachisphora Regu & David 1993 Aleurolobus rhachisphora Regu & David 1993 

Aleurolobus rhododendri Takahashi 1934 Aleurolobus rhododendri Takahashi 1934 

Aleurolobus riveae Regu & David 1993 Aleurolobus riveae Regu & David 1993 

Aleurolobus russellae Regu & David 1993 Aleurolobus russellae Regu & David 1993 

Aleurolobus sairandhryensis Meganathan & David 1993 Aleurolobus sairandhryensis Meganathan & David 1993 

Aleurolobus sairandhryensis Regu & David 1993 Aleurolobus sairandhryensis Regu & David 1993 

Aleurolobus saklespurensis Regu & David 1993 Aleurolobus saklespurensis Regu & David 1993 

Aleurolobus sandorici Corbett 1935 Aleuroclava sandorici (Corbett) 1935 

653 

Aleurolobus saputarensis Regu & David 1993 Aleurolobus saputarensis Regu & David 1993 

Aleurolobus scolopiae Takahashi 1933 Aleurolobus scolopiae Takahashi 1933 

Aleurolobus selangorensis Corbett 1935 Aleurolobus selangorensis Corbett 1935 

Aleurolobus semilunaris (Corbett) 1926 Crescentaleyrodes semilunaris (Corbett) 1926 

Aleurolobus setigerus Quaintance & Baker 1917 Aleurolobus setigerus Quaintance & Baker 1917 

Aleurolobus shiiae Takahashi 1957 Aleurolobus shiiae Takahashi 1957 

Aleurolobus simula (Peal) 1903 Africaleurodes simula (Peal) 1903

Aleurolobus singhi Regu & David 1993 Aleurolobus singhi Regu & David 1993 

Aleurolobus solitarius Quaintance & Baker 1917 Aleurolobus solitarius Quaintance & Baker 1917 

Aleurolobus spinosus Jesudasan & David 1991 Aleurolobus spinosus Jesudasan & David 1991 

Aleurolobus styraci Takahashi 1954 Aleurolobus styraci Takahashi 1954 

Aleurolobus subrotundus Silvestri 1927 Aleurolobus subrotundus Silvestri 1927 

654 

Aleurolobus sundararaji Regu & David 1993 Aleurolobus sundararaji Regu & David 1993 

Aleurolobus szechwanensis Young 1942 Aleurolobus szechwanensis Young 1942 

Aleurolobus taeonabae (Kuwana) 1911 Aleurolobus taeonabae (Kuwana) 1911 

Aleurolobus tassellatus Regu & David 1993 Aleurolobus tassellatus Regu & David 1993 

Aleurolobus tchadiensis Bink-Moenen 1983 Aleurolobus tchadiensis Bink-Moenen 1983 

Aleurolobus teucrii Mifsud & Palmeri 1996 Aleurolobus teucrii Mifsud and Palmeri 1996 

Aleurolobus tuberculatus Regu & David 1993 Aleurolobus tuberculatus Regu & David 1993 

Aleurolobus valparaiensis Jesudasan & David 1991 Aleurolobus valparaiensis Jesudasan & David 1991 

Aleurolobus vitis Danzig 1966 Aleurolobus vitis Danzig 1966 

Aleurolobus vrijdaghi Ghesquiere 1934 Africaleurodes vrijdaghii (Ghesquiere) 1934 

Aleurolobus walayarensis Jesudasan & David 1991 Pealius walayarensis (Jesudasan & David) 1991 

Aleurolobus wunni (Ryberg) 1938 Aleurolobus wunni (Ryberg) 1938 

Aleurolobus zeylanicus Corbett 1926 Orientaleyrodes zeylanicus Corbett 1926 

ALEUROLONGA MOUND 1965 ALEUROLONGA MOUND 1965 

Aleurolonga cassiae Mound 1965 A Aleurolonga cassiae Mound 1965 

ALEUROMARGINATUS CORBETT 1935 ALEUROMARGINATUS CORBETT 1935 

Aleuromarginatus bauhiniae David 1976 Aleuromarginatus tirumurthiensis David 1988 

Aleuromarginatus corbettiaformis Martin 1985 Aleuromarginatus corbettiaformis Martin 1985 

Aleuromarginatus dalbergiae Cohic 1969 Aleuromarginatus dalbergiae Cohic 1969 

Aleuromarginatus erythrinae Selvakumaran & David Dialeurolobus erythrinae (Corbett) 1935 

Aleuromarginatus indica Singh 1940 Aleuromarginatus tephrosiae Corbett 1935 

Aleuromarginatus kallarensis David & Subramaniam 1976 Aleuromarginatus kallarensis David & Subramaniam 1976 

Aleuromarginatus littoralis Martin 1985 Aleuromarginatus littoralis Martin 1985 

Aleuromarginatus marginiquus Martin 1999 Aleuromarginatus marginiquus Martin 1999 

Aleuromarginatus millettiae Cohic 1968 Aleuromarginatus millettiae Cohic 1968 

Aleuromarginatus moundi Martin 1999 Aleuromarginatus moundi Martin 1999 

Aleuromarginatus nemciae Martin 1999 Aleuromarginatus nemciae Martin 1999 

Aleuromarginatus nigrus Martin 1999 Aleuromarginatus nigrus Martin 1999 

Aleuromarginatus serdangensis Takahashi 1955 Aleuromarginatus serdangensis Takahashi 1955 

Aleuromarginatus shihmenensis Ko 1995 Aleuromarginatus shihmenensis Ko 1995 

Aleuromarginatus tephrosiae Corbett 1935 Aleuromarginatus tephrosiae Corbett 1935 

Aleuromarginatus thirumurthiensis David 1976 Aleuromarginatus thirumurthiensis David 1976 

Aleuromigda Singh 1931 Aleuroclava Singh 1931 

Aleuromigda deghai Dozier & Baker nomen nudum

ALEURONUDUS HEMPEL 1922 ALEURONUDUS HEMPEL 1922 

Aleuronudus acapulcensis (Sampson & Drews) 1941 Aleuronudus acapulcensis (Sampson & Drews) 1941 

Aleuronudus bahiensis (Hempel) 1922 Metaleurodicus bahiensis (Hempel) 1922 

Aleuronudus induratus Hempel 1922 Aleuronudus induratus Hempel 1922 

Aleuronudus melzeri (Bondar) 1928 Aleuronudus melzeri (Bondar) 1928 

ALEUROPAPILLATUS REGU & DAVID 1993 ALEUROPAPILLATUS REGU & DAVID 1993 

Aleuropapillatus kumariensis Regu & David 1993 Aleuropapillatus kumariensis Regu & David 1993 

ALEUROPARADOXUS QUAINTANCE & BAKER 1914 ALEUROPARADOXUS QUAINTANCE & BAKER 1914 

Aleuroparadoxus arctostaphyli Russell 1947 Aleuroparadoxus arctostaphyli Russell 1947 

Aleuroparadoxus chomeliae Russell 1947 Aleuroparadoxus chomeliae Russell 1947 

Aleuroparadoxus gardeniae Russell 1947 Aleuroparadoxus gardeniae Russell 1947 

Aleuroparadoxus ilicicola Russell 1947 Aleuroparadoxus ilicicola Russell 1947 

Aleuroparadoxus iridescens (Bemis) 1904 Aleuroparadoxus iridescens (Bemis) 1904 

Aleuroparadoxus lauri (Signoret) 1882 Trialeurodes lauri (Signoret) 1882 

Aleuroparadoxus punctatus Quaintance & Baker 1917 Aleuroparadoxus punctatus Quaintance & Baker 1917 

Aleuroparadoxus rhodae Russell 1947 Aleuroparadoxus rhodae Russell 1947 

Aleuroparadoxus sapotae Russell 1947 Aleuroparadoxus sapotae Russell 1947 

Aleuroparadoxus trinidadensis Russell 1947 Aleuroparadoxus trinidadensis Russell 1947 

Aleuroparadoxus truncatus Russell 1947 Aleuroparadoxus truncatus Russell 1947 

ALEUROPLATUS QUAINTANCE & BAKER 1914 ALEUROPLATUS QUAINTANCE & BAKER 1914 

Aleuroplatus acaciae Bink-Moenen 1983 Aleuroplatus acaciae Bink-Moenen 1983 

Aleuroplatus affinis Takahashi 1961 Aleuroplatus affinis Takahashi 1961 

Aleuroplatus agauriae Takahashi 1955 Aleuroplatus aguariae Takahashi 1955 

Aleuroplatus akeassii Cohic 1969 Aleuroplatus akeassii Cohic 1969 

Aleuroplatus alcocki (Peal) 1903 Aleuroplatus alcocki (Peal) 1903 

Aleuroplatus alpinus Takahashi 1955 Aleuroplatus alpinus Takahashi 1955 

Aleuroplatus anapatsae Takahashi 1951 Aleuroplatus anapatsae Takahashi 1951 

Aleuroplatus andropogoni Dozier 1934 Tetraleurodes andropogoni (Dozier) 1934 

Aleuroplatus berbericolus Quaintance & Baker 1917 Aleuroplatus berbericolus Quaintance & Baker 1917 

Aleuroplatus bignoniae (near) Russell 1944 Aleuroplatus bignoniae (near) Russell 1944 

Aleuroplatus bignoniae Russell 1944 Aleuroplatus bignoniae Russell 1944 

Aleuroplatus biluminiporus Martin & Malumphy 2002 Aleuroplatus biluminiporus Martin & Malumphy 2002 

Aleuroplatus bossi Takahashi 1936 Aleuroplatus bossi Takahashi 1936 

Aleuroplatus buchananiae Jesudasan & David 1991 Aleuroplatus pectiniferus Quaintance & Baker 1917 

655 

Aleuroplatus cadabae Priesner & Hosny 1934 Aleuroplatus cadabae Priesner & Hosny 1934 

Aleuroplatus cinnamomi Jesudasan & David 1991 Aleuroplatus pectiniferus Quaintance & Baker 1917 

Aleuroplatus citri Takahashi 1940 Orchamoplatus citri (Takahashi) 1940 

Aleuroplatus claricephalus Takahashi 1940 Aleuroplatus claricephalus Takahashi 1940 

Aleuroplatus cockerelli (Ihering) 1897 Aleuroplatus cockerelli (Ihering) 1897

Aleuroplatus cococolus Quaintance & Baker 1917 Aleuroplatus cococolus Quaintance & Baker 1917 

Aleuroplatus coronata (Quaintance) 1900 Aleuroplatus coronatus (Quaintance) 1900 

Aleuroplatus crustatus Bondar 1928 Aleuroplatus crustatus Bondar 1928 

Aleuroplatus culcasiae Cohic 1969 Pectinaleyrodes culcasiae (Cohic) 1969 

Aleuroplatus daitoensis Takahashi 1940 Aleuroplatus daitoensis Takahashi 1940 

656 

Aleuroplatus dentatus Sampson & Drew 1941 Aleuroplatus dentatus Sampson & Drew 1941 

Aleuroplatus denticulatus Bondar 1923 Dialeurodes denticulatus (Bondar) 1923 

Aleuroplatus distinctus Jesudasan & David 1991 Aleuroplatus pectiniferus Quaintance & Baker 1917 

Aleuroplatus dorsipallidus Martin 1989 Aleuroplatus dorsipallidus Martin 1989 

Aleuroplatus dubius Takahashi 1955 Aleuroplatus dubius Takahashi 1955 

Aleuroplatus elemarae Mound & Halsey 1978 Aleuroplatus elemarae Mound & Halsey 1978 

Aleuroplatus epigaeae Russell 1944 Aleuroplatus epigaeae Russell 1944 

Aleuroplatus euryae (Kuwana) 1911 Aleuroclava euryae (Kuwana) 1911 

Aleuroplatus evodiae Takahashi 1960 Aleuroplatus evodiae Takahashi 1960 

Aleuroplatus fici Takahashi 1932 Aleuroplatus fici Takahashi 1932 

Aleuroplatus ficifolii Takahashi 1942 Aleuroplatus ficifolii Takahashi 1942 

Aleuroplatus ficifolii var. chiengsenensis Takahashi 1942 Aleuroplatus ficifolii Takahashi 1942 

Aleuroplatus ficusgibbosae Corbett 1926 Aleuroplatus pectiniferus Quaintance & Baker 1917 

Aleuroplatus ficusgibbosae Corbett 1926 Aleuroplatus alcocki (Peal) 1903 acc. David (1993) 

Aleuroplatus ficusrugosae Quaintance & Baker 1917 Aleuroplatus ficusrugosae Quaintance & Baker 1917 

Aleuroplatus fimbriae Mound 1965 Orstomaleyrodes fimbriae (Mound) 1965 

Aleuroplatus gelatinosus (Cockerell) 1898 Aleuroplatus gelatinosus (Cockerell) 1898 

Aleuroplatus graphicus Bondar 1923 Aleuroplatus graphicus Bondar 1923 

Aleuroplatus hiezi Cohic 1968 Aleuroplatus hiezi Cohic 1968 

Aleuroplatus hoyae (Peal) 1903 Aleuroplatus hoyae (Peal) 1903 

Aleuroplatus ilicis Russell 1944 Aleuroplatus ilicis Russell 1944 

Aleuroplatus incisus Quaintance & Baker 1917 Aleuroplatus incisus Quaintance & Baker 1917 

Aleuroplatus incurvatus Takahashi 1961 Aleuroplatus incurvatus Takahashi 1961 

Aleuroplatus insularis Takahashi 1941 Aleuroplatus insularis Takahashi 1941 

Aleuroplatus integellus Bondar 1923 Aleuroplatus integellus Bondar 1923 

Aleuroplatus joholensis Corbett 1935 Aleuroplatus joholensis Corbett 1935 

Aleuroplatus kewensis Trehan 1938 Aleurotulus nephrolepidis (Quaintance) 1900 

Aleuroplatus lateralis Bondar 1923 Aleuroplatus lateralis Bondar 1923 

Aleuroplatus latus Takahashi 1939 Aleuroplatus latus Takahashi 1939 

Aleuroplatus liquidambaris Russell 1944 Aleuroplatus elemarae Mound & Halsey 1976 

Aleuroplatus liquidambaris Takahashi 1941 Aleuroplatus liquidambaris Takahashi 1941 

Aleuroplatus madagascariensis Takahashi 1951 Marginaleyrodes madagascariensis (Takahashi) 1951 

Aleuroplatus magnoliae Russell 1944 Aleuroplatus magnoliae Russell 1944 

Aleuroplatus malayanus Takahashi 1955 Aleuroplatus malayanus Takahashi 1955

Aleuroplatus mameti Takahashi 1937 Aleuroplatus mameti Takahashi 1937 

Aleuroplatus manjakaensis Takahashi 1955 Aleuroplatus manjakaensis Takahashi 1955 

Aleuroplatus monnioti Cohic 1968 Tetraleurodes monnioti (Cohic) 1968 

Aleuroplatus multipori Takahashi 1940 Aleuroplatus multipori Takahashi 1940 

Aleuroplatus myricae Quaintance & Baker 1917 Aleuroplatus myricae Quaintance & Baker 1917 

Aleuroplatus mysorensis David & Subramaniam 1976 Aleuroplatus mysorensis David & Subramaniam 1976 

Aleuroplatus neovatus Takahashi 1961 Aleuroplatus neovatus Takahashi 1961 

Aleuroplatus oculiminutus Quaintance & Baker 1917 Aleuroplatus oculiminutus Quaintance & Baker 1917 

Aleuroplatus oculireniformis Quaintance & Baker 1917 Aleuroplatus oculireniformis Quaintance & Baker 1917 

657 

Aleuroplatus ovatus Quaintance & Baker 1917 Aleuroplatus ovatus Quaintance & Baker 1917 

Aleuroplatus panamensis Sampson & Drews 1941 Aleuroplatus panamensis Sampson & Drews 1941 

Aleuroplatus pauliani Takahashi 1955 Aleuroplatus pauliani Takahashi 1955 

Aleuroplatus pectenserratus Singh 1945 Aleuroplatus pectiniferus Quaintance & Baker 1917 

Aleuroplatus pectiniferus Quaintance & Baker 1917 Aleuroplatus pectiniferus Quaintance & Baker 1917 

Aleuroplatus periplocae (Dozier) 1934 Aleuroplatus periplocae (Dozier) 1934 

Aleuroplatus perseaphagus Martin, Aguiar & Pita 1996 Aleuroplatus perseaphagus Martin, Aguiar & Pita 1996 

Aleuroplatus pileae Takahashi 1939 Aleuroplatus pileae Takahashi 1939 

Aleuroplatus plumosa Quaintance 1900; misid. in part Aleuroplatus magnoliae Russell 1944 

Aleuroplatus plumosus (Quaintance) 1900 Aleuroplatus plumosus (Quaintance) 1900 

Aleuroplatus plumosus (Quaintance); misid. in part Aleuroplatus vaccinii Russell 1944 

Aleuroplatus polystachyae Takahashi 1955 Aleuroplatus polystachyae Takahashi 1955 

Aleuroplatus premnae Corbett 1926 Aleuroplatus premnae Corbett 1926 

Aleuroplatus quaintancei (Peal) 1903 Aleuroplatus quaintancei (Peal) 1903 

Aleuroplatus quercusaquaticae (Quaintance) 1900 Aleuroplatus quercusaquaticae (Quaintance) 1900 

Aleuroplatus robinsoni Takahashi 1955 Aleuroplatus robinsoni Takahashi 1955 

Aleuroplatus sculpturatus Quaintance & Baker 1917 Aleuroplatus sculpturatus Quaintance & Baker 1917 

Aleuroplatus semiplumosus Russell 1944 Aleuroplatus semiplumosus Russell 1944 

Aleuroplatus serratus Takahashi 1955 Aleuroplatus serratus Takahashi 1955 

Aleuroplatus setiger Goux 1939 Massilieurodes setiger (Goux) 1939 

Aleuroplatus silvaticus Cohic 1969 Pectinaleyrodes silvaticus (Cohic) 1969 

Aleuroplatus sinepecten Singh 1945 Aleuroplatus sinepecten Singh 1945 

Aleuroplatus spinus (Singh) 1931 Pealius spinus (Singh) 1931 

Aleuroplatus stellatus (Hempel) 1922 Aleuroplatus stellatus (Hempel) 1922 

Aleuroplatus subrotundus Takahashi 1938 Aleuroplatus subrotundus Takahashi 1938 

Aleuroplatus translucidus Quaintance & Baker 1917 Aleuroplatus translucidus Quaintance & Baker 1917 

Aleuroplatus triclisiae Cohic 1966 Pectinaleyrodes triclisiae (Cohic) 1966 

Aleuroplatus tsibabenae Takahashi 1955 Aleuroplatus tsibabenae Takahashi 1955 

Aleuroplatus tsimananensis Takahashi 1955 Aleuroplatus tsimananensis Takahashi 1955 

Aleuroplatus tuberculatus Takahashi 1951 Aleuroplatus tuberculatus Takahashi 1951

Aleuroplatus vaccinii Russell 1944 Aleuroplatus vaccinii Russell 1944 

Aleuroplatus validus Quaintance & Baker 1917 Aleuroplatus validus Quaintance & Baker 1917 

Aleuroplatus variegatus Quaintance & Baker 1917 Aleuroplatus variegatus Quaintance & Baker 1917 

Aleuroplatus villiersi Cohic 1968 Rugaleyrodes villiersi (Cohic) 1968 

Aleuroplatus vinsonioides (Cockerell) 1898 Aleuroplatus vinsonioides (Cockerell) 1898 

Aleuroplatus vuattouxi Cohic 1969 Rugaleyrodes vuattouxi (Cohic) 1969 

Aleuroplatus walayarensis Jesudasan & David 1991 Aleuroplatus pectiniferus Quaintance & Baker 1917 

Aleuroplatus weinmanniae Takahashi 1951 Aleuroplatus weinmanniae Takahashi 1951 

Aleuroplatus (Massilierodes) Goux 1939 Massilieurodes Goux 1939 

Aleuroplatus (Massilieurodes) setiger Goux 1939 Massilieurodes setiger (Goux) 1939 

Aleuroplatus (Orchamus) Quaintance & Baker 1917 Orchamoplatus Russell 1958 

Aleuroplatus (Orchamus) [sic] samoanus Laing; misid Williams 1944 (in part) Orchamoplatus noumeae Russell 1958 

Aleuroplatus (Orchamus) [sic] samoanus Laing; misid. Orchamoplatus caledonicus (Dumbleton) 1956 

Aleuroplatus (Orchamus) mammaeferus Quaintance & Baker 1917 Orchamoplatus mammaeferus (Quaintance & Baker) 1917 

Aleuroplatus (Orchamus) samoanus Laing 1927 Orchamoplatus mammaeferus (Quaintance & Baker) 1917 

ALEUROPLEUROCELUS DREWS & SAMPSON 1956 ALEUROPLEUROCELUS DREWS & SAMPSON 1956 

Aleuropleurocelus abnormis (Quaintance) 1900 Aleuropleurocelus abnormis (Quaintance) 1900 

Aleuropleurocelus acaudatus Drews & Sampson 1958 Aleuropleurocelus acaudatus Drews & Sampson 1958 

Aleuropleurocelus ceanothi (Sampson) 1945 Aleuropleurocelus ceanothi (Sampson) 1945 

Aleuropleurocelus coachellensis Drews & Sampson 1958 Aleuropleurocelus coachellensis Drews & Sampson 1958 

Aleuropleurocelus granulata (Sampson & Drews) 1941 Aleuropleurocelus granulata (Sampson and Drews) 1941 

Aleuropleurocelus laingi Drews & Sampson 1956 Aleuropleurocelus laingi Drews & Sampson 1956 

Aleuropleurocelus nigrans (Bemis) 1904 Aleuropleurocelus nigrans (Bemis) 1904 

Aleuropleurocelus oblanceolatus Drews & Sampson 1958 Aleuropleurocelus oblanceolatus Drews & Sampson 1958 

Aleuropleurocelus ornatus Drews & Sampson 1958 Aleuropleurocelus ornatus Drews & Sampson 1958 

658 

Aleuropleurocelus rotunda (J. M. Baker) 1937 Aleuropleurocelus rotunda (J. M. Baker) 1937 

Aleuropleurocelus sierrae (Sampson) 1945 Aleuropleurocelus sierrae (Sampson) 1945 

ALEUROPOROSUS CORBETT 1935 ALEUROPOROSUS CORBETT 1935 

Aleuroporosus lumpurensis Corbett 1935 Aleuroporosus lumpurensis Corbett 1935 

ALEUROPTERIDIS MOUND 1961 ALEUROPTERIDIS MOUND 1961 

Aleuropteridis douglasi Mound 1961 Aleuropteridis filicicola (Newstead) 1911 

Aleuropteridis eastopi Mound 1961 Aleuropteridis eastopi Mound 1961 

Aleuropteridis filicicola (Newstead) 1911 Aleuropteridis filicicola (Newstead) 1911 

Aleuropteridis hargreavesi Mound 1961 Aleuropteridis hargreavesi Mound 1961 

Aleuropteridis jamesi Mound 1961 Aleuropteridis jamesi Mound 1961 

ALEUROPUTEUS CORBETT 1935 ALEUROPUTEUS CORBETT 1935 

Aleuroputeus baccaureae Corbett 1935 Aleuroputeus baccaureae Corbett 1935 

Aleuroputeus chinensis Takahashi 1941 Singhiella chinensis (Takahashi) 1941 

Aleuroputeus perseae Corbett 1935 Aleuroputeus perseae Corbett 1935

Aleuroputeus perseae? Corbett 1935 Aleuroputeus perseae? Corbett 1935 

Aleuroputeus publicus Singh 1938 Zaphanera publicus (Singh) 1938 

ALEUROTHRIXUS QUAINTANCE & BAKER 1914 ALEUROTHRIXUS QUAINTANCE & BAKER 1914 

Aleurothrixus (Philodamus) Quaintance & Baker 1917 Aleurothrixus Quaintance & Baker 1914 

Aleurothrixus aepim (Goldi) 1886 Aleurothrixus aepim (Goldi) 1886 

Aleurothrixus aguiari Costa Lima 1942 Aleurothrixus aguiari Costa Lima 1942 

Aleurothrixus antidesmae Takahashi 1933 Aleurothrixus antidesmae Takahashi 1933 

Aleurothrixus bondari Costa Lima 1942 Aleurothrixus bondari Costa Lima 1942 

Aleurothrixus chivelensis Sampson & Drews 1941 Aleurothrixus chivelensis Sampson & Drews 1941 

Aleurothrixus floccosus (Maskell) 1895 Aleurothrixus floccosus (Maskell) 1895 

Aleurothrixus graneli Blanchard 1918 Aleurothrixus aepim (Goldi) 1886 

Aleurothrixus guareae Costa Lima 1942 Aleurothrixus guareae Costa Lima 1942 

Aleurothrixus guimaraesi Costa Lima 1942 Aleurothrixus guimaraesi Costa Lima 1942 

Aleurothrixus horridus (Hempel) 1899 Aleurothrixus floccosus (Maskell) 1895 

Aleurothrixus indica Singh 1931 Aleuroclava indica (Singh) 1931 

Aleurothrixus interrogationis (Bemis) 1904 Aleurothrixus interrogationis (Bemis) 1904 

Aleurothrixus lucumai Costa Lima 1942 Aleurothrixus lucumai Costa Lima 1942 

Aleurothrixus miconiae Hempel 1923 Aleurothrixus miconiae Hempel 1923 

Aleurothrixus myrtacei Bondar 1923 Aleurothrixus myrtacei Bondar 1923 

Aleurothrixus ondinae Bondar 1923 Aleurothrixus ondinae Bondar 1923 

Aleurothrixus porteri Quaintance & Baker 1916 Aleurothrixus porteri Quaintance & Baker 1916 

Aleurothrixus proximans Bondar 1923 Aleurothrixus proximans Bondar 1923 

Aleurothrixus silvestri Corbett 1935 Aleurothrixus silvestri Corbett 1935 

659 

Aleurothrixus similis Sampson & Drews 1941 Aleurothrixus similis Sampson & Drews 1941 

Aleurothrixus smilaceti Takahashi 1934 Aleurothrixus smilaceti Takahashi 1934 

Aleurothrixus solani Bondar 1923 Aleurothrixus solani Bondar 1923 

ALEUROTITHIUS QUAINTANCE & BAKER 1914 ALEUROTITHIUS QUAINTANCE & BAKER 1914 

Aleurotithius mexicanus Russell 1947 Aleurotithius mexicanus Russell 1947 

Aleurotithius timberlakei Quaintance & Baker 1914 Aleurotithius timberlakei Quaintance & Baker 1914 

ALEUROTRACHELUS QUAINTANCE & BAKER 1914 ALEUROTRACHELUS QUAINTANCE & BAKER 1914 

Aleurotrachelus africanus Dozier 1934 Acaudaleyrodes africanus (Dozier) 1934 

Aleurotrachelus alhagi Priesner & Hosny 1934 Acaudaleyrodes rachipora (Singh) 1931 

Aleurotrachelus alpinus Takahashi 1940 Aleurotrachelus alpinus Takahashi 1940 

Aleurotrachelus ambrensis Takahashi & Mamet 1952 Aleurotrachelus ambrensis Takahashi & Mamet 1952 

Aleurotrachelus ampullatus Bink-Moenen 1983 Aleurotrachelus ampullatus Bink-Moenen 1983 

Aleurotrachelus anonae Corbett 1935 Aleurotrachelus anonae Corbett 1935 

Aleurotrachelus asparagi (Lewis) 1893 Aleurotrachelus asparagi (Lewis) 1893 

Aleurotrachelus atratus Hempel 1922 Aleurotrachelus atratus Hempel 1932 

Aleurotrachelus binkae Jesudasan & David 1991 Aleurotrachelus binkae Jesudasan & David 1991

Aleurotrachelus brazzavillense Cohic 1968 Aleurotrachelus brazzavillense Cohic 1968 

Aleurotrachelus cacaorum Bondar 1923 Tetraleurodes cacaorum (Bondar) 1923 

Aleurotrachelus caerulescens Singh 1931 Aleurotrachelus caerulescens Singh 1931 

Aleurotrachelus camamuensis Bondar 1923 Aleurotrachelus camamuensis Bondar 1923 

Aleurotrachelus camelliae (Kuwana) 1911 Aleurotrachelus camelliae (Kuwana) 1911 

Aleurotrachelus cecropiae Bondar 1923 Aleuropleurocelus cecropiae (Bondar) 1923 

Aleurotrachelus chikungensis Mound & Halsey 1978 Aleurotrachelus chikungensis Mound & Halsey 1978 

Aleurotrachelus cinnamomi Jesudasan & David 1991 Aleurotrachelus cinnamomi Jesudasan & David 1991 

Aleurotrachelus citri Priesner & Hosny 1934 Acaudaleyrodes rachipora (Singh) 1931 

Aleurotrachelus coimbatorensis David & Subramaniam 1976 Aleurotrachelus coimbatorensis David & Subramaniam 

1976 

Aleurotrachelus corbetti Takahashi 1941 Aleurotrachelus corbetti Takahashi 1941 

Aleurotrachelus croceatus (Maskell) 1896 Aleuroduplidens croceata (Maskell) 1896 

Aleurotrachelus cruzi Cassino 1999 Tetraleurodes cruzi (Cassino) 1999 

Aleurotrachelus debregeasiae Young 1944 Aleurotrachelus debregeasiae Young 1944 

Aleurotrachelus distinctus Hempel 1923 Aleurotrachelus distinctus Hempel 1923 

Aleurotrachelus dryandrae Solomon 1935 Aleurotrachelus dryandrae Solomon 1935 

Aleurotrachelus duplicatus Bink-Moenen 1983 Aleurotrachelus duplicatus Bink-Moenen 1983 

Aleurotrachelus elatostemae Takahashi 1932 Aleurotrachelus elatostemae Takahashi 1932 

Aleurotrachelus eriosemae (Hempel) 1923 Aleurotrachelus eriosemae (Hempel) 1923 

Aleurotrachelus erythrinae Corbett 1935 Aleurotrachelus erythrinae Corbett 1935 

Aleurotrachelus espunae Gomez-Menor 1945 Aleurotrachelus rhamnicola (Goux) 1940 

Aleurotrachelus euphorifoliae Young 1944 Aleurotrachelus euphorifoliae Young 1944 

Aleurotrachelus fanalae Takahashi 1951 Marginaleyrodes fanalae (Takahashi) 1951 

Aleurotrachelus fenestellae Hempel 1923 Aleurotrachelus fenestellae Hempel 1923 

Aleurotrachelus filamentosus Takahashi 1938 Aleurotrachelus filamentosus Takahashi 1938 

Aleurotrachelus fissistigmae Takahashi 1931 Aleurotrachelus fissistigmae Takahashi 1931 

Aleurotrachelus fumipennis (Hempel) 1899 Crescentaleyrodes fumipennis (Hempel) 1899 

Aleurotrachelus globulariae Goux 1942 Aleurotrachelus globulariae Goux 1942 

Aleurotrachelus goaensis Jesudasan & David 1991 Aleurotrachelus goaensis Jesudasan & David 1991 

Aleurotrachelus granosus Bondar 1923 Aleurotrachelus granosus Bondar 1923 

Aleurotrachelus gratiosus Bondar 1923 Aleurotrachelus gratiosus Bondar 1923 

Aleurotrachelus grewiae Takahashi 1952 Aleurotrachelus grewiae Takahashi 1952 

Aleurotrachelus hazomiavonae Takahashi 1955 Aleurotrachelus hazomiavonae Takahashi 1955 

Aleurotrachelus ingafolii Bondar 1923 Aleurotrachelus ingafolii Bondar 1923 

Aleurotrachelus ishigakiensis (Takahashi) 1933 Aleurotrachelus ishigakiensis (Takahashi) 1923 

Aleurotrachelus jelinekii (Frauenfeld) 1867 Aleurotuba jelinekii (Frauenfeld) 1867 

Aleurotrachelus joholensis Corbett 1935 Aleurotrachelus joholensis Corbett 1935 

Aleurotrachelus juiyunensis Young 1944 Aleurotrachelus juiyunensis Young 1944 

660

Aleurotrachelus limbatus (Maskell) 1895 Aleurotrachelus limbatus (Maskell) 1895 

Aleurotrachelus longispinus Corbett 1926 Aleurotrachelus longispinus Corbett 1926 

Aleurotrachelus lumpurensis Corbett 1935 Aleurotrachelus lumparensis Corbett 1935 

Aleurotrachelus machili Takahashi 1942 Aleurotrachelus machili Takahashi 1942 

Aleurotrachelus madagascariensis Takahashi 1955 Aleurotrachelus madagascariensis Takahashi 1955 

Aleurotrachelus maesae Takahashi 1935 Aleurotrachelus maesae Takahashi 1935 

661 

Aleurotrachelus marginatus (Newstead) 1911 Aleurotrachelus marginatus (Newstead) 1911 

Aleurotrachelus mauritiensis Takahashi 1940 Aleurotrachelus mauritiensis Takahashi 1940 

Aleurotrachelus mesuae Corbett 1935 Aleurotrachelus mesuae Corbett 1935 

Aleurotrachelus micheliae Takahashi 1932 Crenidorsum micheliae (Takahashi) 1932 

Aleurotrachelus minimus Young 1944 Aleurotrachelus minimus Young 1944 

Aleurotrachelus minutus Takahashi 1952 Aleurotrachelus minutus Takahashi 1952 

Aleurotrachelus multipapillus Singh 1932 Aleurotrachelus multipapillus Singh 1932 

Aleurotrachelus murrayae Singh 1931 Aleuroclava murrayae (Singh) 1931 

Aleurotrachelus myrtifolii Bondar 1923 Aleurothrixus myrtifolii (Bondar) 1923 

Aleurotrachelus nivetae Cohic 1969 Aleurotrachelus nivetae Cohic 1969 

Aleurotrachelus obscurus Bink-Moenen 1983 Aleurotrachelus obscurus Bink-Moenen 1983 

Aleurotrachelus orchidicola Takahashi 1939 Aleurotrachelus orchidicola Takahashi 1939 

Aleurotrachelus oriani Mound & Martin 2007 Aleurotrachelus oriani Mound & Martin 2007 

Aleurotrachelus pandani Takahashi 1951 Aleurotrachelus pandani Takahashi 1951 

Aleurotrachelus papilliferus (Sampson & Drews) Aleurotrachelus papilliferus (Sampson & Drews) 1941 

Aleurotrachelus parvus (Hempel) 1899 Aleurotrachelus parvus (Hempel) 1899 

Aleurotrachelus parvus Young 1944 Aleurotrachelus chikungensis Mound & Halsey 1978 

Aleurotrachelus pauliani Takahashi 1960 Aleurotrachelus oriani Mound & Martin 2007 

Aleurotrachelus plectroniae Takahashi 1955 Aleurotrachelus plectroniae Takahashi 1955 

Aleurotrachelus primitus Young 1944 Aleurotrachelus primitus Young 1944 

Aleurotrachelus psidii Singh 1931 Aleuroclava psidii (Singh) 1931 

Aleurotrachelus pykarae Jesudasan & David 1991 Aleurotrachelus pykarae Jesudasan & David 1991 

Aleurotrachelus pyracanthae Takahashi 1935 Aleurotrachelus pyracanthae Takahashi 1935 

Aleurotrachelus rachipora Singh 1931 Acaudaleyrodes rachipora (Singh) 1931 

Aleurotrachelus reunionensis Takahashi 1960 Aleurotrachelus reunionensis Takahashi 1960 

Aleurotrachelus rhamnicola (Goux) 1940 Aleurotrachelus rhamnicola (Goux) 1940 

Aleurotrachelus rosarius Bondar 1923 Aleurotrachelus rosarius Bondar 1923 

Aleurotrachelus rotundus Corbett 1935 Aleurotrachelus rotundus Corbett 1935 

Aleurotrachelus rubi Takahashi 1933 Aleurotrachelus rubi Takahashi 1935 

Aleurotrachelus rubromaculatus Bondar 1923 Aleurotrachelus rubromaculatus Bondar 1923 

Aleurotrachelus saklespurensis Regu & David 1992 Cohicaleyrodes saklespurensis (Regu & David) 1992 

Aleurotrachelus selangorensis Corbett 1935 Aleurotrachelus selangorensis Corbett 1935 

Aleurotrachelus serratus Takahashi 1949 Aleurotrachelus serratus Takahashi 1949

Aleurotrachelus socialis Bondar 1923 Aleurotrachelus socialis Bondar 1923 

Aleurotrachelus souliei Cohic 1969 Aleurotrachelus souliei Cohic 1969 

Aleurotrachelus stellatus Hempel 1922 Aleuroplatus stellatus (Hempel) 1922 

Aleurotrachelus stypheliae (Maskell) Aleurotrachelus stypheliae (Maskell) 1895 

Aleurotrachelus taiwanus Takahashi 1932 Aleurotrachelus taiwanus Takahashi 1932 

662 

Aleurotrachelus tarennae Bink-Moenen 1983 Aleurotrachelus tarennae Bink-Moenen 1983 

Aleurotrachelus theobromae Bondar 1923 Aleurotrachelus theobromae Bondar 1923 

Aleurotrachelus tracheifer (Quaintance) 1900 Aleurotrachelus tracheifer (Quaintance) 1900 

Aleurotrachelus trachoides (Back) 1912 Aleurotrachelus trachoides (Back) 1912 

Aleurotrachelus tuberculatus Corbett 1935 Aleurotrachelus corbetti Takahashi 1941 

Aleurotrachelus tuberculatus Singh 1933 Aleurotrachelus tuberculatus Singh 1933 

Aleurotrachelus turpiniae Takahashi 1932 Crenidorsum turpiniae (Takahashi) 1932 

Aleurotrachelus urticicola Young 1944 Aleurotrachelus urticicola Young 1944 

Aleurotrachelus vitis Corbett 1935 Aleurotrachelus vitis Corbett 1935 

Aleurotrachelus wendlandiae Jesudasan & David 1991 Aleurotrachelus wendlandiae Jesudasan & David 1991 

Aleurotrachelus zonatus Takahashi 1952 Aleurotrachelus zonatus Takahashi 1952 

ALEUROTUBA TREMBLAY AND IACCARINO 1978 ALEUROTUBA TREMBLAY AND IACCARINO 1978 

Aleurotuba jelinekii (Frauenfeld) 1867 Aleurotuba jelinekii (Frauenfeld) 1867 

Aleurotuberculatus TAKAHASHI 1932 Aleuroclava SINGH 1931 

Aleurotuberculatus angkorensis Takahashi 1942 Aleuroclava angkorensis (Takahashi) 1942 

Aleurotuberculatus artocarpi Corbett 1935 Aleuroclava artocarpi (Corbett) 1935 

Aleurotuberculatus aucubae (Kuwana) 1911 Aleuroclava aucubae (Kuwana) 1911 

Aleurotuberculatus bauhiniae Corbett 1935 Aleuroclava bauhiniae (Corbett) 1935 

Aleurotuberculatus bifurcata (Corbett) 1933 Aleuroclava bifurcata (Corbett) 1933 

Aleurotuberculatus bulbiformi Qureshi 1982 Aleuroclava bulbiformi (Qureshi) 1982 

Aleurotuberculatus burmanicus Singh 1938 Aleuroclava burmanicus (Singh) 1938 

Aleurotuberculatus caloncobae Cohic 1966 Aleuroclava nigeriae (Mound) 1965 

Aleurotuberculatus canangae Corbett 1935 Martiniella canangae (Corbett) 1935 

Aleurotuberculatus cardamomi David & Subramaniam 1976 Singhiella cardamomi (David & Subramaniam) 1976 

Aleurotuberculatus cherasensis Corbett 1935 Minutaleyrodes cherasensis (Corbett) 1935 

Aleurotuberculatus citrifolii (Corbett) 1935 Aleuroclava citrifolii (Corbett) 1935 

Aleurotuberculatus cordii Qureshi 1982 Aleuroclava cordii (Qureshi) 1982 

Aleurotuberculatus davidi Qureshi 1982 Aleuroclava davidi (Qureshi) 1982 

Aleurotuberculatus dubius Bink-Moenen 1983 Aleuroclava dubius (Bink-Moenen) 1983 

Aleurotuberculatus elatostemae Takahashi 1932 Aleuroclava elatostemae (Takahashi) 1932 

Aleurotuberculatus erythrinae Corbett 1935 Aleuroclava erythrinae (Corbett) 1935 

Aleurotuberculatus eugeniae Corbett 1935 Aleuroclava eugeniae (Corbett) 1935 

Aleurotuberculatus euphoriae Takahashi 1942 Aleuroclava euphoriae (Takahashi) 1942 

Aleurotuberculatus euryae (Kuwana) 1911 Aleuroclava euryae (Kuwana) 1911

Aleurotuberculatus ficicola Takahashi 1932 Aleuroclava ficicola (Takahashi) 1932 

Aleurotuberculatus filamentosa (Corbett) 1933 Aleuroclava filamentosa (Corbett) 1933 

Aleurotuberculatus flabellus Takahashi 1949 Aleuroclava flabellus (Takahashi) 1949 

Aleurotuberculatus gordoniae Takahashi 1932 Aleuroclava gordoniae (Takahashi) 1932 

Aleurotuberculatus guyavae Takahashi 1932 Aleuroclava guyavae (Takahashi) 1932 

Aleurotuberculatus hexcantha (Singh) 1940 Aleuroclava hexcantha (Singh) 1940 

Aleurotuberculatus hikosanensis Takahashi 1938 Aleuroclava hikosanensis (Takahashi) 1938 

Aleurotuberculatus jasmini Takahashi 1932 Aleuroclava jasmini (Takahashi) 1932 

Aleurotuberculatus kolliensis David 1977 Minutaleyrodes kolliensis (David) 1977 

Aleurotuberculatus kusheriki Mound 1965 Aleuroclava nigeriae (Mound) 1965 

Aleurotuberculatus kuwanai Takahashi 1934 Aleuroclava kuwanai (Takahashi) 1934 

663 

Aleurotuberculatus lagerstroemiae Takahashi 1934 Aleuroclava lagerstroemiae (Takahashi) 1934 

Aleurotuberculatus latus Takahashi 1934 Aleuroclava latus (Takahashi) 1934 

Aleurotuberculatus lithocarpi Takahashi 1934 Aleuroclava lithocarpi (Takahashi) 1934 

Aleurotuberculatus longispinus Takahashi 1934 Aleuroclava longispinus (Takahashi) 1934 

Aleurotuberculatus macarangae Corbett 1935 Martiniella macarangae (Corbett) 1935 

Aleurotuberculatus magnoliae Takahashi 1952 Aleuroclava magnoliae (Takahashi) 1952 

Aleurotuberculatus malloti Takahashi 1932 Aleuroclava malloti (Takahashi) 1932 

Aleurotuberculatus manii David 1978 Aleuroclava manii (David) 1978 

Aleurotuberculatus maximus Qureshi 1982 Aleuroclava maximus (Qureshi) 1982 

Aleurotuberculatus melastomae Takahashi 1934 Aleuroclava melastomae (Takahashi) 1934 

Aleurotuberculatus minutus (Singh) 1931 Minutaleyrodes minutus (Singh) 1931 

Aleurotuberculatus multipori Takahashi 1935 Aleuroclava multipori (Takahashi) 1935 

Aleurotuberculatus multipori var. thaiensis Takahashi 1942 Aleuroclava multipori (Takahashi) 1935 

Aleurotuberculatus murrayae (Singh) 1931 Aleuroclava murrayae (Singh) 1931 

Aleurotuberculatus nachiensis Takahashi 1963 Aleuroclava nachiensis (Takahashi) 1963 

Aleurotuberculatus neolitseae Takahashi 1934 Aleuroclava neolitseae (Takahashi) 1934 

Aleurotuberculatus nephelii Corbett 1935 Aleuroclava nephelii (Corbett) 1935 

Aleurotuberculatus nigeriae Mound 1965 Aleuroclava nigeriae (Mound) 1965 

Aleurotuberculatus nitidus (Singh) 1932 Aleuroclava nitidus (Singh) 1932 

Aleurotuberculatus parvus Singh 1938 Aleuroclava parvus (Singh) 1938 

Aleurotuberculatus phyllanthi (Corbett) 1935 Aleuroclava phyllanthi (Corbett) 1935 

Aleurotuberculatus piperis Takahashi 1935 Aleuroclava piperis (Takahashi) 1935 

Aleurotuberculatus porosus (Priesner & Hosny) 1937 Aleuroclava porosus (Priesner & Hosny) 1937 

Aleurotuberculatus psidii (Singh) 1931 Aleuroclava psidii (Singh) 1931 

Aleurotuberculatus pulcherrimus (Corbett) 1935 Aleuroclava pulcherrima (Corbett) 1935 

Aleurotuberculatus pyracanthae Takahashi 1933 Aleuroclava pyracanthi (Takahashi) 1933 

Aleurotuberculatus rhododendri Takahashi 1935 Aleuroclava rhododendri (Takahashi) 1935 

Aleurotuberculatus russellae David & Subramaniam 1976 Aleuroclava russellae (David & Subramaniam)

Aleurotuberculatus sandorici (Corbett) 1935 Aleuroclava sandorici (Corbett) 1935 

Aleurotuberculatus siamensis Takahashi 1942 Aleuroclava siamensis (Takahashi) 1942 

Aleurotuberculatus similis Takahashi 1938 Aleuroclava similis (Takahashi) 1938 

Aleurotuberculatus similis europaeus (Zahradnik) 1962 Aleuroclava similis (Takahashi) 1938 

Aleurotuberculatus similis suborientalis (Danzig) 1966 Aleuroclava similis (Takahashi) 1938 

Aleurotuberculatus simplex Takahashi 1949 Aleuroclava simplex (Takahashi) 1949 

Aleurotuberculatus stereospermi Corbett 1935 Aleuroclava stereospermi (Corbett) 1935 

Aleurotuberculatus submarginatus Qureshi 1982 Aleuroclava submarginatus (Qureshi) 1982 

Aleurotuberculatus suishanus Takahashi 1934 Minutaleyrodes suishanus (Takahashi) 1934 

Aleurotuberculatus takahashii David & Subramaniam 1976 Aleuroclava takahashii (David & Subramaniam) 1976 

Aleurotuberculatus tentactuliformis Corbett 1935 Aleuroclava tentactuliformis (Corbett) 1935 

664 

Aleurotuberculatus thysanospermi Takahashi 1934 Aleuroclava thysanospermi (Takahashi) 1934 

Aleurotuberculatus trachelospermi Takahashi 1938 Aleuroclava trachelospemi (Takahashi) 1938 

Aleurotuberculatus ubonensis Takahashi 1942 Aleuroclava ubonensis (Takahashi) 1942 

Aleurotuberculatus uraianus Takahashi 1932 Aleuroclava uraianus (Takahashi) 1932 

Aleurotuberculatus yambiae Gameel 1971 Aleuroclava nigeriae (Mound) 1965 

ALEUROTULUS QUAINTANCE & BAKER 1914 ALEUROTULUS QUAINTANCE & BAKER 1914 

Aleurotulus anthuricola Nakahara 1989 Aleurotulus anthuricola Nakahara 1989 

Aleurotulus arundinacea Singh 1931 Aleurotulus arundinacea Singh 1931 

Aleurotulus bodkini Quaintance & Baker 1923 Aleurotrachelus trachoides (Back) 1912 

Aleurotulus filicium (Goldi) 1886 nomen dubia 

Aleurotulus laneus Martin 2005 Aleurotulus laneus Martin 2005 

Aleurotulus maculata Singh 1931 Dialeurolonga maculata (Singh) 1931 

Aleurotulus mundururu Bondar 1923 Aleurotulus mundururu Bondar 1923 

Aleurotulus nephrolepidis (Quaintance) 1900 Aleurotulus nephrolepidis (Quaintance) 1900 

Aleurotulus pteridophytae Martin 1994 Aleurotulus pteridophytae Martin 1994 

ALEUROVIGGIANUS IACCARINO 1982 ALEUROVIGGIANUS IACCARINO 1982 

Aleuroviggianus adanaensis Bink-Moenen 1992 Aleuroviggianus adanaensis Bink-Moenen 1992 

Aleuroviggianus adrianae Iaccarino 1982 Aleuroviggianus adrianae Iaccarino 1982 

Aleuroviggianus graecus Bink-Moenen 1992 Aleuroviggianus graecus Bink-Moenen 1992 

Aleuroviggianus halperini Bink-Moenen 1992 Aleuroviggianus halperini Bink-Moenen 1992 

Aleuroviggianus polymorphus Bink-Moenen 1992 Aleuroviggianus polymorphus Bink-Moenen 1992 

Aleuroviggianus zonalus Bink-Moenen 1992 Aleuroviggianus zonalus Bink-Moenen 1992 

ALEUROVITREUS MARTIN 2005 ALEUROVITREUS MARTIN 2005 

Aleurovitreus insignis (Bondar) 1923 Aleurovitreus insignis (Bondar) 1923 

Aleurovitreus risor Martin 2005 Aleurovitreus risor Martin 2005 

ALEYRODES LATREILLE 1796 ALEYRODES LATREILLE 1796 

Aleyrodes abnormis Quaintance 1900 Aleuropleurocelus abnormis (Quaintance) 1900 

Aleyrodes acerum Kirkaldy 1907 Aleurochiton aceris (Modeer) 1762

Aleyrodes acteae [sic] Britton 1905 Aleyrodes asarumis Shimer 1867 

Aleyrodes akebiae Kuwana 1911 Pealius akebiae (Kuwana) 1911 

Aleyrodes albescens Hempel 1923 Aleyrodes albescens Hempel 1923 

Aleyrodes ambrosiae Cockerell 1910 Trialeurodes abutiloneus (Haldeman) 1850 

Aleyrodes amnicola Bemis 1904 Aleyrodes amnicola Bemis 1904 

Aleyrodes asari (Schrank) 1801 Aleyrodes asari (Schrank) 1801 

Aleyrodes asarumis Shimer 1867 Aleyrodes asarumis Shimer 1867 

Aleyrodes atriplex Froggatt 1911 Lipaleyrodes atriplex (Froggatt) 1911 

Aleyrodes aucubae Kuwana 1911 Aleuroclava aucubae (Kuwana) 1911 

Aleyrodes aureocincta Cockerell 1897 Aleyrodes aureocincta Cockerell 1897 

Aleyrodes avellanae (Signoret), corrected misid. Pealius quercus (Signoret) 1868 

Aleyrodes avellanae (Signoret), misid. Pealius quercus (Signoret) 1868 

Aleyrodes avellanae Signoret 1868 Asterobemisia carpini (Koch) 1857 

Aleyrodes azaleae Baker & Moles 1920 Pealius azaleae (Baker & Moles) 1920 

Aleyrodes azaleae Baker & Moles 1920:81 Pealius azaleae (Baker & Moles) 1920 

Aleyrodes baja Sampson 1943 Aleyrodes baja Sampson 1943 

Aleyrodes bifasciata Stephens 1829 nomen dubium 

Aleyrodes borchsenii Danzig 1966 Aleyrodes lonicerae Walker 1852 

Aleyrodes brassicae Walker 1852 Aleyrodes proletella (Linnaeus) 1758 

Aleyrodes calophyllii Kotinsky 1907 Aleurocanthus calophyllii (Kotinsky) 1907 

Aleyrodes camelliae Kuwana 1911 Aleurotrachelus camelliae (Kuwana) 1911 

Aleyrodes campanulae Saalas 1942 Aleyrodes proletella? (Linnaeus) 1758 

Aleyrodes campanulae Takahashi 1963 Aleyrodes takahashi Ossiannilsson 1966 

Aleyrodes capreae Signoret 1868 nomen dubia 

Aleyrodes chelidonii Latreille 1807 Aleyrodes proletella (Linnaeus) 1758 

Aleyrodes ciliata Takahashi 1955 Aleyrodes ciliata Takahashi 1955 

Aleyrodes cinnamomi Takahashi 1932 Pentaleyrodes cinnamomi (Takahashi) 1932 

Aleyrodes citri Ashmead 1885 Dialeurodes citri (Ashmead) 1885 

Aleyrodes citrifolii Morgan 1893 Singhiella citrifolii (Morgan) 1893 

Aleyrodes cocois Curtis 1846 Aleurodicus cocois (Curtis) 1846 

Aleyrodes coryli Britton 1907 Trialeurodes packardi (Morrill) 1903 

Aleyrodes crataegi Kiriukhin 1947 Aleyrodes crataegi Kiriukhin 1947 

Aleyrodes diasemus Bemis 1904 Aleyrodes diasemus Bemis 1904 

Aleyrodes dorseyi Kotinsky 1907 Tetraleurodes mori (Quaintance) 1899 

Aleyrodes dubia Stephens 1829 Coniopteryx tineiformis Curtis [Neuroptera] 

Aleyrodes elevatus Silvestri 1934 Aleyrodes elevatus Silvestri 1934 

Aleyrodes errans Bemis 1904 Tetraleurodes mori (Quaintance) 1899 

Aleyrodes essigi Penny 1922 Aleyrodes essigi Penny 1922 

Aleyrodes eugeniae var aurantii Maskell 1895 Dialeurodes citri (Ashmead) 1885 

665

Aleyrodes euphorbiae Low 1867 Aleyrodes proletella (Linnaeus) 1758 

Aleyrodes euryae Kuwana 1911 Aleuroclava euryae (Kuwana) 1911 

Aleyrodes extraniens Bemis 1904 Aleurotulus nephrolepidis (Quaintance) 1900 

Aleyrodes fernaldi Morrill 1903 Trialeurodes fernaldi (Morrill) 1903 

Aleyrodes fijensis Kotinsky 1907 Rhachisphora fijiensis (Kotinksy) 1907 

Aleyrodes filicicola Newstead 1911 Aleuropteridis filicicola (Newstead) 1911 

Aleyrodes filicium Goldi 1886 nomen dubia 

Aleyrodes fodiens Maskell 1895 Aleyrodes fodiens Maskell 1895 

Aleyrodes fragariae Walker 1852 Aleyrodes lonicerae Walker 1852 

Aleyrodes fraxini Signoret 1868 nomen dubia 

Aleyrodes gelatinosus Cockerell 1898 Aleuroplatus gelatinosus (Cockerell) 1898 

Aleyrodes giffardi Kotinsky 1907 Bemisia giffardi (Kotinsky) 1907 

Aleyrodes glacialis Bemis (in part) Trialeurodes vittata (Quaintance) 

666 

Aleyrodes glacialis Bemis (misidentification in part) Trialeurodes vaporariorum (Westwood) 1856 

Aleyrodes glacialis Bemis 1904 Trialeurodes glacialis (Bemis) 1904 

Aleyrodes gossypii (Fitch) 1857 Aleyrodes gossypii (Fitch) 1857 

Aleyrodes gracialis Bemis 1904; misidentification in part Trialeurodes drewsi Sampson 1945 

Aleyrodes hibisci Kotinsky 1907 Singhius hibisci (Kotinsky) 1907 

Aleyrodes horridus Hempel 1899 Aleurothrixus floccosus (Maskell) 1895 

Aleyrodes howardi Quaintance 1907 Aleurothrixus floccosus (Maskell) 1895 

Aleyrodes hutchingsi Bemis 1904 Trialeurodes hutchingsi (Bemis) 1904 

Aleyrodes hyperici Corbett 1926 Aleyrodes hyperici Corbett 1926 

Aleyrodes immaculata Stephens 1829 nomen nudum 

Aleyrodes insignis Bondar 1923 Aleurovitreus insignis (Bondar) 1923 

Aleyrodes interrogationis Bemis 1904 Aleurothrixus interrogationis (Bemis) 1904 

Aleyrodes iridescens Bemis 1904 Aleuroparadoxus iridescens (Bemis) 1904 

Aleyrodes iridescens Bemis 1904; misidentification in part Aleuroparadoxus arctostaphyli Russell 1947 

Aleyrodes japonica Takahashi 1963 Aleyrodes japonica Takahashi 1963 

Aleyrodes kelloggi Bemis 1904 Pealius kelloggi (Bemis) 1904 

Aleyrodes kirkaldyi Kotinsky 1907 Dialeurodes kirkaldyi (Kotinsky) 1907 

Aleyrodes kushinasii Sasaki 1908 Dialeurodes citri (Ashmead) 1885 

Aleyrodes lacerdae Signoret 1883 Metaleurodicus lacerdae (Signoret) 1883 

Aleyrodes lactea Zehntner 1897 Aleyrodes lactea Zehntner 1897 

Aleyrodes latus Hempel 1923 Aleyrodes latus Hempel 1923 

Aleyrodes leguminicola (Takahashi) 1942 Aleyrodes leguminicola (Takahashi) 1942 

Aleyrodes liquidambari Takahashi 1932 Pealius liquidambari (Takahashi) 1932 

Aleyrodes lonicerae Walker 1852 Aleyrodes lonicerae Walker 1852 

Aleyrodes madroni Bemis 1904 Trialeurodes madroni (Bemis) 1904 

Aleyrodes maskelli Bemis 1904 Pealius maskelli (Bemis) 1904

Aleyrodes melanops Cockerell 1902 Tetraleurodes melanops (Cockerell) 1902 

Aleyrodes menthae Haupt 1934 Aleyrodes lonicerae Walker 1852 

Aleyrodes merlini Bemis 1904 Trialeurodes merlini (Bemis) 1904 

Aleyrodes millettiae Cohic 1968 Aleurocybotus millettiae (Cohic) 1968 

Aleyrodes mori arizonensis Cockerell 1900 Tetraleurodes mori (Quaintance) 1899 

Aleyrodes mori var. maculata Morrill 1903 Tetraleurodes mori (Quaintance) 1899 

Aleyrodes morrilli Britton 1907 Trialeurodes packardi (Morrill) 1903 

Aleyrodes nigra Maskell [sic] Zaphanera niger (Maskell) 1895 

Aleyrodes nigrans Bemis 1904 Aleuropleurocelus nigrans (Bemis) 1904 

Aleyrodes nubifera Berger 1909 Singhiella citrifolii (Morgan) 1893 

Aleyrodes oceanica Takahashi 1939 Metaleyrodes oceanica (Takahashi) 1939 

Aleyrodes osmaroniae Sampson 1945 Aleyrodes osmaroniae Sampson 1945 

Aleyrodes packardi Morrill 1903 Trialeurodes packardi (Morrill) 1903 

Aleyrodes periplocae Dozier 1934 Aleuroplatus periplocae (Dozier) 1934 

Aleyrodes philadelphi Danzig 1966 Aleyrodes philadelphi Danzig 1966 

Aleyrodes prenanthis (Schrank) 1801 Aleyrodes proletella (Linnaeus) 1758 

Aleyrodes proletella (Linnaeus) 1758 Aleyrodes proletella (Linnaeus) 1758 

Aleyrodes pruinosus Bemis 1904 Aleyrodes pruinosus Bemis 1904 

Aleyrodes pruinosus euphorbium Cockerell 1911 Aleyrodes pruinosus Bemis 1904 

Aleyrodes pryrolae Gillette & Baker 1895 Aleyrodes pyrolae Gillette & Baker 1895 

Aleyrodes quaintancei Bemis 1904 Tetraleurodes mori (Quaintance) 1899 

Aleyrodes rhamnicola Goux 1940 Aleurotrachelus rhamnicola (Goux) 1940 

Aleyrodes ricini Misra 1924 Trialeurodes ricini (Misra) 1924 

Aleyrodes rosae Korobitsin 1967 Neopealius rubi Takahashi 1954 

Aleyrodes rubi Signoret 1868 Aleyrodes lonicerae Walker 1852 

Aleyrodes ruborum Cockerell 1897 Trialeurodes ruborum (Cockerell) 1897 

Aleyrodes shizuokensis Kuwana 1911 Aleyrodes shizuokensis Kuwana 1911 

Aleyrodes singularis Danzig 1964 Aleyrodes singularis Danzig 1964 

667 

Aleyrodes sonchi Kotinsky Trialeurodes vaporariorum (Westwood) 1856 

Aleyrodes sonchi Kotinsky 1907 Trialeurodes vaporariorum (Westwood) 1856 

Aleyrodes sorini Takahashi 1958 Aleyrodes sorini Takahashi 1958 

Aleyrodes spinosus Kuwana 1911 Aleurocanthus spinosus (Kuwana) 1911 

Aleyrodes spiraeae Douglas 1894 Aleyrodes lonicerae Walker 1852 

Aleyrodes spiraeoides Quaintance 1900 Aleyrodes spiraeoides Quaintance 1900 

Aleyrodes splendens Bemis 1904 Tetraleurodes mori (Quaintance) 1899 

Aleyrodes stanfordi Bemis 1904 Tetraleurodes perileuca (Cockerell) 

Aleyrodes taeonabae Kuwana 1911 Aleurolobus taeonabae (Kuwana) 1911 

Aleyrodes taiheisanus Takahashi 1939 Aleyrodes taiheisanus Takahashi 1939 

Aleyrodes takahashii Ossiannilsson 1966 Aleyrodes takahashi Ossiannilsson 1966

Aleyrodes tentaculatus Bemis 1904 Trialeurodes tentaculatus (Bemis) 1904 

Aleyrodes tentaculatus Bemis 1904, in part Trialeurodes bemisiae Russell 1948 

Aleyrodes tinaeoides Blanchard 1852 Aleyrodes tinaeoides Blanchard 1852 

Aleyrodes tokyonis Kuwana 1911 Rusostigma tokyonis (Kuwana) 1911 

Aleyrodes trachoides Back 1912 Aleurotrachelus trachoides (Back) 1912 

Aleyrodes ursorum Cockerell 1910 Tetraleurodes ursorum (Cockerell) 1910 

Aleyrodes vitrinellus Cockerell 1903 Trialeurodes vitrinellus (Cockerell) 1903 

Aleyrodes waldeni Britton Trialeurodes packardi (Morrill) 1903 

Aleyrodes waldeni Britton 1907 Trialeurodes packardi (Morrill) 1903 

Aleyrodes wellmanae Bemis Trialeurodes vittata (Quaintance) 

Aleyrodes wellmanae Bemis 1904 Trialeurodes vittata (Quaintance) 

Aleyrodes winterae Takahashi 1937 Aleyrodes winterae Takahashi 1937 

Aleyrodes youngi Hempel 1901 Aleyrodes proletella (Linnaeus) 1758 

Aleyrodes zygia Danzig 1966 Aleyrodes zygia Danzig 1966 

Aleyrodes (Asterochiton) Maskell; Cockerell 1902 Asterochiton MASKELL 1879 

Aleyrodes (Asterochiton) Maskell; misidentication Trialeurodes Cockerell 1902 

Aleyrodes (Asterochiton) aurea [sic] Maskell Asterochiton aureus Maskell 1879 

Aleyrodes (Dialeurodes) Cockerell 1902 Dialeurodes Cockerell 1902 

Aleyrodes (Dialeurodes) aurantii (Maskell) 1895 Dialeurodes citri (Ashmead) 1885 

Aleyrodes (Dialeurodes) citri Riley & Howard 1892 Dialeurodes citri (Ashmead) 1885 

Aleyrodes (Dialeurodes) eugeniae (Maskell) Rusostigma eugeniae (Maskell) 1895 

Aleyrodes (Dialeurodes) eugeniae (Maskell) 1885 Rusostigma eugeniae (Maskell) 1895 

Aleyrodes (Frauenfeldiella) Gomez-Menor 1943 Aleurotrachelus QUAINTANCE & BAKER 1914 

Aleyrodes (Frauenfeldiella) jelinekii (Frauenfeld) 1867 Aleurotuba jelinekii (Frauenfeld) 1867 

Aleyrodes (Frauenfeldiella) jelinekii form alba 1867 Aleurotuba jelinekii (Frauenfeld) 1867 

Aleyrodes (Ogivaleurodes) lauri (Signoret) 1882 Trialeurodes lauri (Signoret) 1882 

Aleyrodes (Philodamus) interrogationis Bemis Aleurothrixus interrogationis (Bemis) 1904 

Aleyrodes (Tetraleurodes) Cockerell 1902 Tetraleurodes Cockerell 1902 

Aleyrodes (Tetraleurodes) melanops Cockerell 1902 Tetraleurodes melanops (Cockerell) 1902 

Aleyrodes (Tetraleurodes) perileuca Cockerell 1902 Tetraleurodes perileuca (Cockerell) 1902 

Aleyrodes (Trialeurodes) erigerontis Maskell 1895 Trialeurodes abutiloneus (Haldeman) 1850 

Aleyrodes (Trialeurodes) fitchi Quaintance Trialeurodes abutiloneus (Haldeman) 1850 

Aleyrodes (Trialeurodes) pergandei Quaintance Trialeurodes pergandei (Quaintance) 1900 

Aleyrodes (Trialeurodes) ruborum Cockerell Trialeurodes ruborum (Cockerell) 1897 

Aleyrodes (Trialeurodes) variabilis Quaintance 1900 Trialeurodes variabilis (Quaintance) 1900 

Aleyrodes (Trialeurodes) vittata Quaintance Trialeurodes vittata (Quaintance) 

Aleyrodicus (sic) destructor Mackie 1912 Aleurodicus destructor Mackie 1912 

ALEYRODIELLA DANZIG 1966 ALEYRODIELLA DANZIG 1966 

Aleyrodiella lamellifera Danzig 1966 Aleyrodiella lamellifera Danzig 1966 

668

ANOMALEYRODES TAKAHASHI & MAMET 1952 ANOMALEYRODES TAKAHASHI & MAMET 1952 

Anomaleyrodes palmae Takahashi & Mamet 1952 Anomaleyrodes palmae Takahashi & Mamet 1952 

APOBEMISIA TAKAHASHI 1954 APOBEMISIA TAKAHASHI 1954 

Apobemisia celti (Takahashi) 1932 Apobemisia celti (Takahashi) 1932 

Apobemisia kuwanai (Takahashi) 1934 Apobemisia kuwanai (Takahashi) 1934 

ARACHNALEYRODES BINK-MOENEN 1983 ARACHNALEYRODES BINK-MOENEN 1983 

669 

Arachnaleyrodes insignis Bink-Moenen 1983 Arachnaleyrodes insignis Bink-Moenen 1983 

ASIALEYRODES CORBETT 1935 ASIALEYRODES CORBETT 1935 

Asialeyrodes corbetti Takahashi 1949 Asialeyrodes corbetti Takahashi 1949 

Asialeyrodes dubius Martin & Mound 2007 Asialeyrodes dubius Martin & Mound 2007 

Asialeyrodes elegans Meganathan & David 1994 Asialeyrodes elegans Meganathan & David 1994 

Asialeyrodes euphoriae Takahashi 1942 Asialeyrodes euphoriae Takahashi 1942 

Asialeyrodes indica Sundararaj & David 1991 Asialeyrodes indica Sundararaj & David 1991 

Asialeyrodes lumpurensis Corbett 1935 Asialeyrodes lumpurensis Corbett 1935 

Asialeyrodes lushanensis Ko 1993 Asialeyrodes lushanensis Ko 1993 

Asialeyrodes maesae (Takahashi) 1934 Asialeyrodes maesae (Takahashi) 1934 

Asialeyrodes meghalayensis Regu & David 1991 Asialeyrodes meghalayensis Regu & David 1991 

Asialeyrodes menoni Meganathan & David 1994 Asialeyrodes menoni Meganathan &David 1994 

Asialeyrodes multipori Takahashi 1942 Asialeyrodes multipori Takahashi 1942 

Asialeyrodes papillatus Regu & David 1991 Asialeyrodes papillatus Regu & David 1991 

Asialeyrodes saklespurensis Regu & David 1993 Asialeyrodes indicus Sundararaj and David 1993 

Asialeyrodes selangorensis Corbett 1935 Asialeyrodes selangorensis Corbett 1935 

Asialeyrodes spherica (Sundararaj & Dubey) 2006 Asialeyrodes spherica (Sundararaj & Dubey) 2006 

Asialeyrodes splendens Meganathan & David 1994 Asialeyrodes splendens Meganathan & David 1994 

Aspidiotus gossypii Fitch 1857 Aleyrodes gossypii (Fitch) 1857 

ASTEROBEMISIA TREHAN 1940 ASTEROBEMISIA TREHAN 1940 

Asterobemisia atraphaxius (Danzig) 1969 Asterobemisia atraphaxius (Danzig) 1969 

Asterobemisia avellanae (Signoret) 1868 Asterobemisia carpini (Koch) 1857 

Asterobemisia carpini (Koch) 1857 Asterobemisia carpini (Koch) 1857 

Asterobemisia curvata (Qureshi) 1980 Asterobemisia curvata (Qureshi) 1980 

Asterobemisia dentata Danzig 1969 Asterobemisia dentata Danzig 1969 

Asterobemisia helyi Dumbleton 1956 Bemisia giffardi (Kotinsky) 1907 

Asterobemisia lata Danzig 1966 Asterobemisia lata Danzig 1966 

Asterobemisia moringae David & Subramaniam 1976 Bemisia moringae (David & Subramaniam) 1976 

Asterobemisia nigrini Zahradnik 1987a Asterobemisia paveli (Zahradnik) 1961 

Asterobemisia obenbergeri (Zahradnik) 1961 Asterobemisia obenbergeri (Zahradnik) 1961 

Asterobemisia paveli (Zahradnik) 1961 Asterobemisia paveli (Zahradnik) 1961 

Asterobemisia salicaria (Danzig) 1969 Asterobemisia salicaria (Danzig) 1969 

Asterobemisia silvatica (Danzig) 1964 Asterobemisia silvatica (Danzig) 1964

Asterobemisia takahashii Danzig 1966 Asterobemisia takahashii Danzig 1966 

Asterobemisia trifolii (Danzig) 1966 Asterobemisia trifolii (Danzig) 1966 

Asterobemisia yanagicola (Takahashi) 1933 Asterobemisia yanagicola (Takahashi) 1933 

Asterobemisia yanagicola (Takahashi) 1934 Asterobemisia yanagicola (Takahashi) 1934 

ASTEROCHITON MASKELL 1879 ASTEROCHITON MASKELL 1879 

Asterochiton Maskell; misidentification Trialeurodes Cockerell 1902 

Asterochiton abutiloneus (Haldeman) 1850 Trialeurodes abutiloneus (Haldeman) 1850 

Asterochiton ambrosiae (Cockerell) 1910 Trialeurodes abutiloneus (Haldeman) 1850 

Asterochiton asplenii (Maskell) 1890 Trialeurodes asplenii (Maskell) 1890 

Asterochiton aureus Maskell 1879 Asterochiton aureus Maskell 1879 

Asterochiton auricolor (Bondar) 1923 Asterochiton auricolor (Bondar) 1923 

Asterochiton bagnalli Harrison 1920 nomen dubium 

Asterochiton cerata (Maskell) 1895 Asterochiton cerata (Maskell) 1895 

Asterochiton cordiae David & Subramaniam 1976 Asterochiton cordiae David & Subramaniam 1976 

Asterochiton corollis Penny 1922 Trialeurodes corollis (Penny) 1922 

Asterochiton diasemus (Bemis) 1904 Aleyrodes diasemus Bemis 1904 

Asterochiton diminutis Penny 1922 Trialeurodes diminutis (Penny) 1922 

Asterochiton dubienus Bondar 1923 Disiphon dubienus (Bondar) 1923 

Asterochiton erigerontis (Maskell) 1895 Trialeurodes abutiloneus (Haldeman) 1850 

Asterochiton fagi (Maskell) 1890 Asterochiton fagi (Maskell) 1890 

Asterochiton floridensis (Quaintance) 1900 Trialeurodes floridensis (Quaintance) 1900 

Asterochiton glacialis (Bemis) 1904 Trialeurodes glacialis (Bemis) 1904 

Asterochiton hutchingsi (Bemis) 1904 Trialeurodes hutchingsi (Bemis) 1904 

670 

Asterochiton lecanioides Maskell 1879 (in part) Trialeurodes vaporariorum (Westwood) 1856 

Asterochiton madroni (Bemis) 1904 Trialeurodes madroni (Bemis) 1904 

Asterochiton manihoti Bondar 1923 Trialeurodes manihoti (Bondar) 1923 

Asterochiton merlini (Bemis) 1904 Trialeurodes merlini (Bemis) 1904 

Asterochiton packardi (Morrill) 1903 Trialeurodes packardi (Morrill) 1903 

Asterochiton papillifer Maskell 1890 Trialeurodes vaporariorum (Westwood) 1856 

Asterochiton pergandei (Quaintance) 1900 Trialeurodes pergandei (Quaintance) 1900 

Asterochiton phillyreae (Haliday) 1835 Siphoninus phillyreae (Haliday) 1835 

Asterochiton pittospori Dumbleton 1957 Asterochiton pittospori Dumbleton 1957 

Asterochiton rolfsii (Quaintance) 1899 Trialeurodes abutiloneus (Haldeman) 1850 

Asterochiton ruborum (Cockerell) 1850 Trialeurodes ruborum (Cockerell) 1897 

Asterochiton simplex (Maskell) 1890 Asterochiton simplex (Maskell) 1890 

Asterochiton sonchi (Kotinsky) 1907 Trialeurodes vaporariorum (Westwood) 1856 

Asterochiton tentaculatus (Bemis) 1904; in part Trialeurodes tentaculatus (Bemis) 1904 

Asterochiton vaporariorum (Westwood) 1856 Trialeurodes vaporariorum (Westwood) 1856 

Asterochiton variabilis (Quaintance) 1900 Trialeurodes variabilis (Quaintance) 1900

Asterochiton vitrinellus (Cockerell) Trialeurodes vitrinellus (Cockerell) 1903 

Asterochiton vittata (Quaintance) 1900 Trialeurodes vittata (Quaintance) 1900 

Asterochiton wellmanae (Bemis) 1904 Trialeurodes vittata (Quaintance) 1900 

AUSTROALEURODICUS TAPIA 1970 AUSTROALEURODICUS TAPIA 

Austroaleurodicus lomatiae Tapia 1970 Austroaleurodicus lomatiae Tapia 1970 

Austroaleurodicus pigeanus (Baker & Moles) 1921) Austroaleurodicus pigeanus (Baker & Moles) 1921 

AXACALIA DANZIG 1969 AXACALIA DANZIG 1969 

Axacalia spiraeanthi Danzig 1969 Axacalia spiraeanthi Danzig 1969 

AZURALEURODICUS MARTIN 1999 AZURALEURODICUS MARTIN 1999 

Azuraleurodicus pentarthus Martin 1999 Azuraleurodicus pentarthus Martin 1999 

BERNAEINAE SHCHERBAKOV 2000 BERNAEINAE SHCHERBAKOV 2000 

BAKERIUS BONDAR 1923 BAKERIUS BONDAR 1923 

Bakerius amazonicus Penny & Arias 1980 Bakerius amazonicus Penny & Arias 1980 

Bakerius attenuatus Bondar 1923 Bakerius attenuatus Bondar 1923 

Bakerius calmoni Bondar 1928 Bakerius calmoni Bondar 1928 

Bakerius coelhoi [sic] (Bondar) Costa Lima 1936 Dialeurodicus coelhi Bondar 1928 

Bakerius conspurcatus (Enderlein) 1909 Bakerius conspurcatus (Enderlein) 1909 

Bakerius glandulosus Hempel 1938 Bakerius glandulosus Hempel 1938 

Bakerius maculatus Penny & Arias 1980 Bakerius maculatus Penny & Arias 1980 

Bakerius phrygilanthi Bondar 1923 Bakerius phrygilanthi Bondar 1923 

Bakerius sanguineus Bondar 1928 Bakerius sanguineus Bondar 1928 

Bakerius sublatus Bondar 1928 Bakerius sublatus Bondar 1928 

BELLITUDO RUSSELL 1943 BELLITUDO RUSSELL 1943 

Bellitudo campae Russell 1943 Bellitudo campae Russell 1943 

Bellitudo cubae Russell 1943 Bellitudo cubae Russell 1943 

Bellitudo hispaniola Russell 1943 Bellitudo hispaniola Russell 1943 

Bellitudo jamaicae Russell 1943 Bellitudo jamaicae Russell 1943 

Bemisaleyrodes COHIC 1969 Bemisaleyrodes COHIC 1969 

Bemisaleyrodes balachowskyi Cohic 1969 Bemisaleyrodes balachowskyi Cohic 1969 

671 

Bemisaleyrodes brideliae Bink-Moenen 1983 Bemisaleyrodes brideliae Bink-Moenen 1983 

Bemisaleyrodes grjebinei (Cohic) 1968 Bemisaleyrodes grjebinei (Cohic) 1968 

Bemisaleyrodes pauliani Cohic 1969 Bemisaleyrodes pauliani Cohic 1969 

BEMISIA QUAINTANCE & BAKER 1914 BEMISIA QUAINTANCE & BAKER 1914 

Bemisia aceri Takahashi 1931 Parabemisia aceris (Takahashi) 1931 

Bemisia achyranthes Singh 1931 Bemisia tabaci (Gennadius) 1889 

Bemisia afer (Priesner & Hosny) 1934 Bemisia afer (Priesner & Hosny) 1934 

Bemisia alni Takahashi 1957 Bemisia alni Takahashi 1957 

Bemisia antennata Gameel 1968 Bemisia antennata Gameel 1968 

Bemisia argentifolii Bellows & Perring 1994 Bemisia tabaci (Gennadius) 1889

Bemisia artocarpi Corbett 1935 Pealius artocarpi (Corbett) 1935 

Bemisia bahiana Bondar 1928 Bemisia tabaci (Gennadius) 1889 

Bemisia bambusae Takahashi 1942 Bemisia bambusae Takahashi 1942 

Bemisia berbericola (Cockerell) 1896 Bemisia berbericola (Cockerell) 1896 

672 

Bemisia callunae Ossiannilsson 1947 Calluneyrodes callunae (Ossiannilsson) 1947 

Bemisia capitata Regu & David 1991 Bemisia capitata Regu & David 1991 

Bemisia caudasculptura Quaintance & Baker 1937 Bemisia caudasculptura Quaintance & Baker 1937 

Bemisia centroamericana Martin 2005 Bemisia centroamericana Martin 2005 

Bemisia citricola Gomez-Menor 1945 Bemisia afer (Priesner & Hosny) 1934 

Bemisia combreticula Bink-Moenen 1983 Bemisia combreticula Bink-Moenen 1983 

Bemisia confusa Danzig 1964 Bemisia confusa Danzig 1964 

Bemisia cordylinidis Dumbleton 1961 Bemisia cordylinidis Dumbleton 1961 

Bemisia costa-limai Bondar 1928 Bemisia tabaci (Gennadius) 1889 

Bemisia curvata Qureshi 1980 Asterobemisia curvata (Qureshi) 1980 

Bemisia decipiens (Maskell) 1895 Bemisia decipiens (Maskell) 1895 

Bemisia elliptica Takahashi 1960 Bemisia elliptica Takahashi 1960 

Bemisia emiliae Corbett 1926 Bemisia tabaci (Gennadius) 1889 

Bemisia eoa Danzig 1966 Bemisia eoa Danzig 1966 

Bemisia formosana Takahashi 1933 Bemisia formosana Takahashi 1933 

Bemisia giffardi (Kotinsky) 1907 Bemisia giffardi (Kotinsky) 1907 

Bemisia giffardi hispina Young 1942 Bemisia giffardi (Kotinsky) 1907 

Bemisia gigantea Martin 1999 Bemisia gigantea Martin 1999 

Bemisia goldingi Corbett 1935 Bemisia tabaci (Gennadius) 1889 

Bemisia gossypiperda Misra & Lamba 1929 Bemisia tabaci (Gennadius) 1889 

Bemisia gossypiperda var mosaicivectura Ghesquiere 1934 Bemisia tabaci (Gennadius) 1889 

Bemisia graminis David & Augustine 1988 Bemisia formosana Takahashi 1933 

Bemisia grjebinei Cohic 1968 Bemisaleyrodes grjebinei (Cohic) 1968 

Bemisia grossa Singh 1931 Bemisia grossa Singh 1931 

Bemisia guieriae Bink-Moenen 1983 Bemisia guieriae Bink-Moenen 1983 

Bemisia hancocki Corbett 1936 Bemisia afer (Priesner & Hosny) 1934 

Bemisia hibisci Takahashi 1933 Bemisia tabaci (Gennadius) 1889 

Bemisia hirta Bink-Moenen 1983 Bemisia hirta Bink-Moenen 1983 

Bemisia inconspicua (Quaintance) 1900 Bemisia tabaci (Gennadius) 1889 

Bemisia iole Danzig 1966 Bemisia iole Danzig 1966 

Bemisia jasminum David & Subramaniam 1976 Bemisia giffardi (Kotinsky) 1907 

Bemisia kuwanai Takahashi 1934 Apobemisia kuwanai (Takahashi) 1934 

Bemisia lampangensis Takahashi 1942 Bemisia lampangensis Takahashi 1942 

Bemisia lauraceae Martin, Aguiar & Pita 1996 Bemisia lauracea Martin, Aguiar & Pita 1996 

Bemisia leakii (Peal) 1903 Bemisia leakii (Peal) 1903

Bemisia leguminicola Takahashi 1942 Lipaleyrodes leguminicola (Takahashi) 1942 

Bemisia longispina Priesner & Hosny 1934 Bemisia tabaci (Gennadius) 1889 

Bemisia lonicerae Takahashi 1957 Bemisia tabaci (Gennadius) 1889 

Bemisia manihotis Frappa 1938 Bemisia tabaci (Gennadius) 1889 

Bemisia medinae Gomez-Menor 1954 Bemisia medinae Gomez-Menor 1954 

Bemisia mesasiatica Danzig 1969 Bemisia mesasiatica Danzig 1969 

Bemisia minima Danzig 1964 Bemisia tabaci (Gennadius) 1889 

Bemisia miniscula Danzig 1964 Bemisia tabaci (Gennadius) 1889 

Bemisia moringae (David & Subramaniam) 1976 Bemisia moringae (David & Subramaniam) 1976 

Bemisia multituberculata Sundararaj & David 1990 Bemisia multituberculata Sundararaj & David 1990 

Bemisia myricae Kuwana 1927 Parabemisia myricae (Kuwana) 1927 

Bemisia nigeriensis Corbett 1935 Bemisia tabaci (Gennadius) 1889 

Bemisia ovata (Goux) 1940 Bemisia ovata (Goux) 1940 

Bemisia poinsettiae Hempel 1923 Bemisia poinsettiae Hempel 1923 

Bemisia pongamiae Takahashi 1931 Bemisia pongamiae Takahashi 1931 

Bemisia porteri Corbett 1935 Bemisia porteri Corbett 1935 

Bemisia psiadiae Takahashi 1955 Bemisia psiadiae Takahashi 1955 

Bemisia puerariae Takahashi 1955 Bemisia puerariae Takahashi 1955 

Bemisia religiosa (Peal) 1903 Bemisia religiosa (Peal) 1903 

Bemisia rhodesiansis Corbett 1936 Bemisia tabaci (Gennadius) 1889 

Bemisia rosae Danzig 1969 Neopealius rubi Takahashi 1954 

Bemisia salicaria Danzig 1969 Asterobemisia salicaria (Danzig) 1969 

Bemisia shinanoensis Kuwana 1922 Bemisia shinanoensis Kuwana 1922 

Bemisia signata Bondar 1928 Bemisia tabaci (Gennadius) 1889 

Bemisia signus Bondar 1928 Bemisia tabaci (Gennadius) 1889 

Bemisia silvatica Danzig 1964 Asterobemisia silvatica (Danzig) 1964 

Bemisia spiraeae Young 1944 Bemisia spiraeae Young 1944 

Bemisia spiraeoides Mound & Halsey 1978 Bemisia spiraeoides Mound & Halsey 1978 

Bemisia spireae [sic] Gomez-Menor 1954 Bemisia spiraeoides Mound & Halsey 1978 

Bemisia subdecipiens Martin 1999 Bemisia subdecipiens Martin 1999 

Bemisia sugonjaevi Danzig 1969 Bemisia sugonjaevi Danzig 1969 

Bemisia tabaci (Gennadius) 1889 Bemisia tabaci (Gennadius) 1889 

Bemisia tabaci "B"(Gennadius) 1889 Bemisia tabaci B biotype 1889 

Bemisia tabaci Q biotype Bemisia tabaci Q biotype 

Bemisia tuberculata Bondar 1923 Bemisia tuberculata Bondar 1923 

Bemisia vayssierei Frappa 1939 Bemisia tabaci (Gennadius) 1889 

Bemisia yanagicola Takahashi 1933 Asterobemisia yanagicola (Takahashi) 1933 

Bemisia (Neobemisia) VISNYA 1941 Asterobemisia TREHAN 1940 

Bemisia (Neobemisia) afer [sic] (Priesner & Hosny) 1934 Bemisia afer (Priesner & Hosny) 1934 

673

Bemisia (Neobemisia) afra [sic] (Priesner & Hosny) 1934 Bemisia afer (Priesner & Hosny) 1934 

Bemisia (Neobemisia) avellanae (Signoret) 1868 Asterobemisia carpini (Koch) 1857 

Bemisia (Neobemisia) hancocki Corbett 1936 Bemisia afer (Priesner & Hosny) 1934 

Bemisia (Neobemisia) hibisci Takahashi Bemisia tabaci (Gennadius) 1889 

Bemisia (Neobemisia) hibisci Takahashi 1933 Bemisia tabaci (Gennadius) 1889 

Bemisia (Neobemisia) rhodesiaensis Corbett 1936 Bemisia tabaci (Gennadius) 1889 

Bemisia (Neobemisia) ribium (Douglas) 1888 Asterobemisia carpini (Koch) 1857 

Bemisia (Neobemisia) yanagicola Takahashi 1933 Asterobemisia yanagicola (Takahashi) 1933 

Bemisia (Neobemisia) yanagicola Takahashi 1934 Asterobemisia yanagicola (Takahashi) 1934 

Bemisia (Roucasia) medinae Gomez-Menor 1954 Bemisia medinae Gomez-Menor 1954 

BEMISIELLA DANZIG 1966 BEMISIELLA DANZIG 1966 

Bemisiella artemisiae Danzig 1966 Bemisiella artemisiae Danzig 1966 

Bemisiella lespedezae Danzig 1966 Bemisiella lespedezae Danzig 1966 

BERNAEA SCHLEE 1970 BERNAEA SCHLEE 1970 

Bernaea neocomica Schlee 1970 Bernaea neocomica Schlee 1970 

Bondaria SAMPSON & DREWS 1941 Dialeurodicus COCKERELL 1902 

Bondaria radifera Sampson & Drews 1941 Dialeurodicus radifera (Sampson & Drews) 1941 

BRAZZALEYRODES COHIC 1966 BRAZZALEYRODES COHIC 1966 

Brazzaleyrodes bergerardi Cohic 1966 Viennotaleyrodes bergerardi (Cohic) 1966 

Brazzaleyrodes eriococciformis Cohic 1966 Brazzaleyrodes eriococciformis Cohic 1966 

Brazzaleyrodes platysepali Cohic 1966 Viennotaleyrodes platysepali (Cohic) 1966 

BULGARIALEURODES CORBETT 1936 BULGARIALEURODES CORBETT 1936 

Bulgarialeurodes cotesii (Maskell) 1895 Bulgarialeurodes cotesii (Maskell) 1895 

Bulgarialeurodes rosae Corbett 1936 Bulgarialeurodes cotesii (Maskell) 1895 

BURMOSELIS SHCHERBAKOV 2000 BURMOSELIS SHCHERBAKOV 2000 

Burmoselis evelynae Shcherbakov 2000 Burmoselis evelynae Shcherbakov 2000 

CALLUNEYRODES ZAHRADNIK 1961 CALLUNEYRODES ZAHRADNIK 1961 

674 

Calluneyrodes callunae (Ossiannilsson) 1947 Calluneyrodes callunae (Ossiannilsson) 1947 

CERALEURODICUS HEMPEL 1922 CERALEURODICUS HEMPEL 1922 

Ceraleurodicus altissimus (Quaintance) 1900 Nealeurodicus altissimus (Quaintance) 1900 

Ceraleurodicus assymmetricus (Bondar) 1922 Ceraleurodicus assymmetricus (Bondar) 1922 

Ceraleurodicus bakeri (Bondar) 1923 Nealeurodicus bakeri (Bondar) 1923 

Ceraleurodicus duckei Penny & Arias 1980 Ceraleurodicus duckei Penny & Arias 1980 

Ceraleurodicus hempeli Costa Lima 1928 Ceraleurodicus hempeli Costa Lima 1928 

Ceraleurodicus ingae (Baker) 1937 Nealeurodicus ingae (Baker) 1937 

Ceraleurodicus keris Martin 2004 Ceraleurodicus keris Martin 2004 

Ceraleurodicus kesselyaki (Visnya) 1941 Ceraleurodicus varus (Bondar) 1928 

Ceraleurodicus moreirai Costa Lima 1928 Nealeurodicus moreirai (Costa Lima) 1928 

Ceraleurodicus neivai (Bondar) 1928 Ceraleurodicus neivai (Bondar) 1928

Ceraleurodicus nitidus (Hempel) 1922 Octaleurodicus nitidus Hempel 1922 

Ceraleurodicus octifer (Bondar) 1923 Nealeurodicus octifer (Bondar) 1923 

Ceraleurodicus pulcherrimus (Quaintance & Baker) 1913 Octaleurodicus pulcherrimus (Quaintance & Baker) 1913 

Ceraleurodicus splendidus Hempel 1922 Ceraleurodicus spendidus Hempel 1922 

Ceraleurodicus varus (Bondar) 1928 Ceraleurodicus varus (Bondar) 1928 

Chermes aceris Geoffroy 1762 Aleurochiton aceris (Modeer) 1778 

Chermes aceris ovatus Geoffroy 1762 Aleurochiton aceris (Modeer) 1778 

CHITONALEYRODES MARTIN 1999 CHITONALEYRODES MARTIN 1999 

Chitonaleyrodes canberrensis Martin 1999 Chitonaleyrodes canberrensis Martin 1999 

Coccus aceris Modeer 1778 Aleurochiton aceris (Modeer) 1778 

Coccus asari Schrank 1801 Aleyrodes asari (Schrank) 1801 

Coccus preanthis Schrank 1801 Aleyrodes proletella (Linnaeus) 1758 

Coccus prenanthis Schrank 1801 Aleyrodes proletella (Linnaeus) 1758 

COCKERELLIELLA SUNDARARAJ & DAVID 1992 COCKERELLIELLA SUNDARARAJ & DAVID 1992 

Cockerelliella adinandrae (Corbett) 1935 Cockerelliella adinandrae (Corbett) 1935 

Cockerelliella curcumae (Corbett) 1935 Cockerelliella curcumae (Corbett) 1935 

Cockerelliella dehradunensis (Jesudasan & David) 1991 Cockerelliella dehradunensis (Jesudasan & David) 1991 

Cockerelliella dioscoreae Sundararaj & David 1992 Cockerelliella dioscoreae Sundararaj & David 1992 

Cockerelliella indica Sundararaj & David 1992 Cockerelliella indica Sundararaj & David 1992 

Cockerelliella kamardini (Corbett) 1935 Cockerelliella kamardini (Corbett) 1935 

Cockerelliella meghalayensis Sundararaj & David 1992 Cockerelliella meghalayensis Sundararaj & David 1992 

Cockerelliella psidii (Corbett) 1935 Cockerelliella psidii (Corbett) 1935 

Cockerelliella quaintancei Sundararaj & David 1992 Cockerelliella quaintancei Sundararaj & David 1992 

Cockerelliella rhodamniae (Corbett) 1935 Cockerelliella rhodamniae (Corbett) 1935 

Cockerelliella rotunda Regu & David 1993 Cockerelliella rotunda Regu & David 1993 

Cockerelliella sembilanensis (Corbett) 1935 Cockerelliella sembilanensis (Corbett) 1935 

Cockerelliella somnathensis Sundararaj 2000 Cockerelliella somnathensis Sundararaj 2000 

Cockerelliella splendens Meganathan & David 1994 Cockerelliella splendens Meganathan & David 1994 

Cockerelliella zingiberae Sundararaj & David 1992 Cockerelliella zingiberae Sundararaj & David 1992 

COHICALEYRODES BINK-MOENEN 1983 COHICALEYRODES BINK-MOENEN 1983 

Cohicaleyrodes alterans (Cohic) 1966 Cohicaleyrodes alterans (Cohic) 1966 

Cohicaleyrodes blanzyi (Cohic) 1968 Cohicaleyrodes blanzyi (Cohic) 1968 

Cohicaleyrodes crossopterygis Bink-Moenen 1983 Cohicaleyrodes crossopterygis Bink-Moenen 1983 

675 

Cohicaleyrodes descarpentriesi (Cohic) 1968 Cohicaleyrodes descarpentriesi (Cohic) 1968 

Cohicaleyrodes indicus Jesudasan and David 1991 Cohicaleyrodes jesudasani Jesudasan and David 2005 

Cohicaleyrodes jesudasani David 2005 Cohicaleyrodes jesudasani David 2005 

Cohicaleyrodes mappiae Selvaakumaran & David 1996 Cohicaleyrodes mappiae Selvaakumaran & David 1996 

Cohicaleyrodes obscura Bink-Moenen 1983 Cohicaleyrodes obscura Bink-Moenen 1983 

Cohicaleyrodes padminiae Phillips & Jesudasan 2006 Cohicaleyrodes padminiae Phillips & Jesudasan 2006

Cohicaleyrodes pauliani (Cohic) 1968 Cohicaleyrodes pauliani (Cohic) 1968 

Cohicaleyrodes platysepali (Cohic) 1966 Cohicaleyrodes platysepali (Cohic) 1966 

Cohicaleyrodes quadrilongispinae Bink-Moenen 1983 Cohicaleyrodes quadrilongispinae Bink-Moenen 1983 

Cohicaleyrodes recurvispina (Cohic) 1966 Cohicaleyrodes recurvispina (Cohic) 1966 

Cohicaleyrodes saklespurensis (Regu & David) 1992 Cohicaleyrodes saklespurensis (Regu & David) 1992 

Cohicaleyrodes uvariae (Cohic) 1968 Cohicaleyrodes uvariae (Cohic) 1968 

COMBESALEYRODES COHIC 1966 COMBESALEYRODES COHIC 1966 

Combesaleyrodes bouqueti Cohic 1966 Combesaleyrodes bouqueti Cohic 1966 

Combesaleyrodes tauffiebi Cohic 1966 Combesaleyrodes tauffiebi Cohic 1966 

Conantulus GOUX 1987 Aleyrodes LATREILLE 1796 

Conantulus lacombiensis Goux 1988 Aleyrodes lonicerae Walker 1852 

Conantulus lacombiensis Goux 1988:65 Aleyrodes lonicerae Walker 1852 

Corbettella Sampson 1943 Pealius Quaintance & Baker 1914 

CORBETTIA DOZIER 1934 CORBETTIA DOZIER 1934 

Corbettia baphiae Russell 1960 Corbettia millettiacola Dozier 1934 

Corbettia bauhiniae Cohic 1968 Corbettia bauhiniae Cohic 1968 

Corbettia graminis Mound 1965 Corbettia graminis Mound 1965 

Corbettia grandis Russell 1960 Corbettia grandis Russell 1960 

Corbettia indentata Russell 1960 Corbettia millettiacola Dozier 1934 

Corbettia isoberliniae Bink-Moenen 1983 Corbettia insoberliniae Bink-Moenen 1983 

Corbettia lamottei Cohic 1969 Corbettia lamottei Cohic 1969 

Corbettia lonchocarpi Bink-Moenen 1983 Corbettia lonchocarpi Bink-Moenen 1983 

Corbettia millettiacola Dozier 1934 Corbettia millettiacola Dozier 1934 

Corbettia pauliani Cohic 1966 Corbettia pauliani Cohic 1966 

Corbettia tamarindi Takahashi 1951 Corbettia tamarindi Takahashi 1951 

Cortesiana GOUX 1988 Bemisia (Gennadius) 1889 

Cortesiana restonicae Goux 1988 Bemisia tabaci (Gennadius) 1889 

CRENIDORSUM RUSSELL 1945 CRENIDORSUM RUSSELL 1945 

Crenidorsum armatae Russell 1945 Crenidorsum armatae Russell 1945 

Crenidorsum aroidephagus Martin & Aguiar 2001 Crenidorsum aroidephagus Martin and Aguiar 2001 

Crenidorsum bladhiae (Takahashi) 1931 Crenidorsum bladhiae (Takahashi) 1931 

Crenidorsum celebes Martin 1988 Crenidorsum celebes Martin 1988 

Crenidorsum commune Russell 1945 Crenidorsum commune Russell 1945 

Crenidorsum debordae Russell 1945 Crenidorsum debordae Russell 1945 

Crenidorsum diaphanum Russell 1945 Crenidorsum diaphanum Russell 1945 

Crenidorsum differens Russell 1945 Crenidorsum differens Russell 1945 

Crenidorsum lasangensis Martin 1985 Crenidorsum lasangensis Martin 1985 

Crenidorsum leve Russell 1945 Crenidorsum leve Russell 1945 

Crenidorsum magnisetae Russell 1945 Crenidorsum magnisetae Russell 1945 

676

Crenidorsum malpighiae Russell 1945 Crenidorsum malpighiae Russell 1945 

Crenidorsum marginale Russell 1945 Crenidorsum marginale Russell 1945 

Crenidorsum micheliae (Takahashi) 1932 Crenidorsum micheliae (Takahashi) 1932 

Crenidorsum millennium Martin 1999 Crenidorsum millennium Martin 1999 

Crenidorsum morobensis Martin 1985 Crenidorsum morobensis Martin 1985 

Crenidorsum ornatum Russell 1945 Crenidorsum ornatum Russell 1945 

Crenidorsum rubiae (David) 2000 Crenidorsum rubiae (David) 2000 

Crenidorsum russellae David & David 2000 Crenidorsum russellae David & David 2000 

Crenidorsum stigmaphylli Russell 1945 Crenidorsum stigmaphylli Russell 1945 

Crenidorsum tuberculatum Russell 1945 Crenidorsum tuberculatum Russell 1945 

Crenidorsum turpiniae (Takahashi) 1932 Crenidorsum turpiniae (Takahashi) 1932 

CRESCENTALEYRODES DAVID & JESUDASAN 1987 CRESCENTALEYRODES DAVID & JESUDASAN 1987 

677 

Crescentaleyrodes fumipennis (Hempel) 1899 Crescentaleyrodes fumipennis (Hempel) 1899 

Crescentaleyrodes monodi (Cohic) 1969 Crescentaleyrodes monodi (Cohic) 1969 

Crescentaleyrodes paulianae (Cohic) 1969 Crescentaleyrodes paulianae (Cohic) 1969 

Crescentaleyrodes semilunaris (Corbett) 1926 Crescentaleyrodes semilunaris (Corbett) 1926 

Crescentaleyrodes vetiveriae Dubey & Ko 2006 Crescentaleyrodes vetiveriae Dubey & Ko 12006 

CRYPTOLINGULA MARTIN & CARVER 1999 CRYPTOLINGULA MARTIN & CARVER 1999 

Cryptolingula perplexa Martin & Carver 1999 Cryptolingula perplexa Martin & Carver 1999 

Dialeurodes (Dialeurolonga) DOZIER 1928 Dialeurolonga DOZIER 1928 

Dialeurodes (Dialeurolonga) elongata Dozier 1928 Dialeurolonga elongata Dozier 1928 

Dialeurodes (Dialeuronomada) dissimilis Quaintance & Baker 1917 Dialeuronomada dissimilis (Quaintance & Baker) 1917 

Dialeurodes (Dialeuroplata) townsendi Quaintance & Baker 1917 Dialeurodes townsendi Quaintance & Baker 1917 

Dialeurodes (Dialeuropora) Quaintance & Baker 1917 Dialeuropora Quaintance & Baker 1917 

Dialeurodes (Dialeuropora) brideliae Takahashi 1932 Dialeuropora brideliae (Takahashi) 1932 

Dialeurodes (Dialeuropora) decempuncta Quaintance & Baker 1917 Dialeuropora decempuncta (Quaintance & Baker) 1917 

Dialeurodes (Dialeuropora) murrayae Takahashi 1931 Dialeuropora murrayae (Takahashi) 1931 

Dialeurodes (Dialeuropora) setigerus Takahashi 1932 Dialeuropora decempuncta (Quaintance & Baker) 1917 

Dialeurodes (Dialeuropora) viburni Takahashi 1933 Dialeuropora viburni (Takahashi) 1933 

Dialeurodes (Gigaleurodes) Young 1942 Dialeurodes citricola Young 1942 

Dialeurodes (Gigaleurodes) elbaensis Priesner & Hosny 1934 Singhiella elbaensis (Priesner & Hosny) 1934 

Dialeurodes (Gigaleurodes) formosensis Takahashi 1933 Massilieurodes formosensis (Takahashi) 1933 

Dialeurodes (Gigaleurodes) maxima Quaintance & Baker 1917 Dialeurodes maxima Quaintance & Baker 1917 

Dialeurodes (Gigaleurodes) nigeriae Cohic 1966 Dialeurodes nigeriae Cohic 1966 

Dialeurodes (Gigaleurodes) pauliani Cohic 1966 Dialeurodes pauliani Cohic 1966 

Dialeurodes (Gigaleurodes) philippinensis Takahashi 1936 Dialeurodes philippinensis Takahashi 1936 

Dialeurodes (Gigaleurodes) simmondsi Corbett 1927 Dialeurodes simmondsi Corbett 1927 

Dialeurodes (Gigaleurodes) splendens Regu and David 1993 Massilieurodes homonoiae (Jesudasan & David) 1991 

Dialeurodes (Rabdostigma) Quaintance & Baker 1917 Rabdostigma Quaintance & Baker 1917

Dialeurodes (Rabdostigma) minahassai Martin 1988 Rabdostigma minahassai (Martin) 1989 

Dialeurodes (Rabdostigma) radiilinealis Quaintance & Baker 1917 Dialeurodes radiilinealis Quaintance & Baker 1917 

Dialeurodes (Rhabdostigma) shintenensis Takahashi 1933 Dialeurodes shintenensis Takahashi 1933 

Dialeurodes (Rhachisphora) Quaintance & Baker 1917 Rhachisphora Quaintance & Baker 1917 

Dialeurodes (Rhachisphora) capitatis Corbett 1926 Rhachisphora capitatis (Corbett) 1926 

Dialeurodes (Rhachisphora) fici Takahashi 1932 Massilieurodes fici (Takahashi) 1932 

Dialeurodes (Rhachisphora) fijensis (Kotinsky) 1907 Rhachisphora fijiensis (Kotinksy) 1907 

Dialeurodes (Rhachisphora) koshunensis Takahashi Rhachisphora koshunensis (Takahashi) 1933 

Dialeurodes (Rhachisphora) koshunensis Takahashi 1933 Rhachisphora koshunensis (Takahashi) 1933 

Dialeurodes (Rhachisphora) kuraruensis Takahashi 1933 Singhiella kuraruensis (Takahashi) 1933 

Dialeurodes (Rhachisphora) machili Takahashi 1932 Rhachisphora machili (Takahashi) 1932 

Dialeurodes (Rhachisphora) maesae Takahashi 1932 Rhachisphora maesae (Takahashi) 1932 

Dialeurodes (Rhachisphora) reticulata Takahashi 1933 Rhachisphora reticulata (Takahashi) 1933 

Dialeurodes (Rhachisphora) rutherfordi Quaintance & Baker 1917 Rhachisphora rutherfordi (Quaintance & Baker) 1917 

Dialeurodes (Rhachisphora) setulosa Corbett 1926 Rhachisphora setulosa (Corbett) 1926 

Dialeurodes (Rhachisphora) styraci Takahashi 1934 Rhachisphora styraci (Takahashi) 1934 

Dialeurodes (Rhachisphora) trilobitoides Quaintance & Baker 1917 Rhachisphora trilobitoides (Quaintance & Baker) 

1917 

Dialeurodes (Rusostigma) eugeniae (Maskell) 1895 Rusostigma eugeniae (Maskell) 1895 

Dialeurodes (Rusostigma) radiiruigosa Quaintance & Baker 1917 Rusostigma radiiruigosa Quaintance & Baker 1917 

Dialeurodes (Rusostigma) tokyonis (Kuwana) 1911 Rusostigma tokyonis (Kuwana) 1911 

Dialeurodes (Rusostigma) tristylii var. uichancoi (Takahashi) 1936 Rusostigma tristylii (Takahashi) 1935 

Dialeurodes (Singhius) Takahashi 1932 Singhius Takahashi 1932 

Dialeurodes ? erythrinae Corbett Dialeurodes ? erythrinae Corbett 

DIALEURODES COCKERELL 1902 DIALEURODES COCKERELL 1902 

Dialeurodes abbotabadiensis Qureshi 1980 Dialeurodes abbotabadiensis Qureshi 1980 

Dialeurodes adinandrae Corbett 1935 Cockerelliella adinandrae (Corbett) 1935 

Dialeurodes adinobotris Corbett 1935 Dialeurodes adinobotris Corbett 1935 

Dialeurodes agalmae Takahashi 1935 Dialeurodes agalmae Takahashi 1935 

Dialeurodes angulata Corbett 1935 Dialeurodes angulata Corbett 1935 

Dialeurodes anjumi Qureshi 1980 Dialeurodes anjumi Qureshi 1980 

Dialeurodes ara Corbett 1935 Dialeurodes ara Corbett 1935 

Dialeurodes ardisiae Takahashi 1935 Rhachisphora ardisiae (Takahashi) 1935 

Dialeurodes armatus David & Subramaniam 1976 Dialeurodes armatus David & Subramaniam 1976 

Dialeurodes bancoensis Ardaillon & Cohic 1970 Dialeurodes bancoensis Ardaillon & Cohic 1970 

Dialeurodes bankokana Takahashi 1942 Dialeurodes bangokana Takahashi 1942 

Dialeurodes bassiae David & Subramaniam 1976 Singhiella bassiae (David & Subramaniam) 1976 

Dialeurodes bicornicauda Martin 1999 Dialeurodes bicornicauda Martin 1999 

Dialeurodes bifurcata Corbett 1933 Aleuroclava bifurcata (Corbett) 1933 

678

Dialeurodes bipunctata Corbett 1933 Dialeuropora bipunctata (Corbett) 1933 

Dialeurodes bladhiae Takahashi 1931 Cockerelliella bladhiae (Takahashi) 1931 

Dialeurodes brideliae Jesudasan & David 1991 Singhiella brideliae (Jesudasan and David) 1991 

679 

Dialeurodes buscki Quaintance & Baker 1917 Dialeurodes buscki Quaintance & Baker 1917 

Dialeurodes cambodiensis Takahashi 1942 Singhiella cambodiensis (Takahashi) 1942 

Dialeurodes cardamomi David & Subramaniam 1976 Singhiella cardamomi (David & Subramaniam) 1976 

Dialeurodes cardamoni David & Subramaniam 1976 Singhiella cardamoni (David & Subramaniam) 1976 

Dialeurodes celti Takahashi 1942 Dialeurodes celti Takahashi 1942 

Dialeurodes centrosemae Corbett 1935 Dialeuropora centrosemae (Corbett) 1935 

Dialeurodes cephalidistinctus Singh 1932 Dialeurodes cephalidistinctus Singh 1932 

Dialeurodes cerifera Quaintance & Baker 1917 Dialeurodes cerifera Quaintance & Baker 1917 

Dialeurodes chiengsenana Takahashi 1942 Dialeurodes chiengsenana Takahashi 1942 

Dialeurodes chitinosa Takahashi 1937 Singhiella chitinosa (Takahashi) 1937 

Dialeurodes chittendeni Laing 1928 Massilieurodes chittendeni (Laing) 1928 

Dialeurodes cinnamomi Takahashi 1932 Dialeurodes cinnamomi Takahashi 1932 

Dialeurodes cinnamomicola Takahashi 1937 Dialeurodes cinnamomicola Takahashi 1937 

Dialeurodes citri (Ashmead) 1885 Dialeurodes citri (Ashmead) 1885 

Dialeurodes citri var. hederae Takahashi 1936 Dialeurodes citri (Ashmead) 1885 

Dialeurodes citri var. kinyana Takahashi 1935 Dialeurodes citri (Ashmead) 1885 

Dialeurodes citricola Young 1942 Dialeurodes citricola Young 1942 

Dialeurodes citrifolii (Morgan) 1893 Singhiella citrifolii (Morgan) 1893 

Dialeurodes conocephali Corbett 1935 Dialeurodes conocephali Corbett 1935 

Dialeurodes crescentata Corbett 1935 Dialeurodes crescentata Corbett 1935 

Dialeurodes cumiugum Singh 1932 Russellaleyrodes cumiugum (Singh) 1932 

Dialeurodes curcumae Corbett 1935 Cockerelliella curcumae (Corbett) 1935 

Dialeurodes cyathispinifera Corbett 1933 Dialeurodes cyathispinifera Corbett 1933 

Dialeurodes daphniphylli Takahashi 1932 Dialeurodes daphniphylli Takahashi 1932 

Dialeurodes davidi Mound & Halsey 1978 Dialeurodes davidi Mound & Halsey 1978 

Dialeurodes decaspermi Martin 1985 Dialeurodes decaspermi Martin 1985 

Dialeurodes delamarei Cohic 1968 Singhiella delamarei (Cohic) 1968 

Dialeurodes delhiensis David & Sundararaj 1992 Dialeurodes delhiensis David & Sundararaj 1992 

Dialeurodes denticulatus (Bondar) 1923 Dialeurodes denticulatus (Bondar) 1923 

Dialeurodes dicksoni Corbett 1935 Dialeurodes dicksoni Corbett 1935 

Dialeurodes didymocarpi Corbett 1935 Dialeurodes didymocarpi Corbett 1935 

Dialeurodes dioscoreae Takahashi 1934 Singhiella dioscoreae (Takahashi) 1934 

Dialeurodes dipterocarpi Takahashi 1942 Singhiella dipterocarpi (Takahashi) 1942 

Dialeurodes dissimilis Quaintance & Baker 1917 Dialeuronomada dissimilis (Quaintance & Baker) 1917 

Dialeurodes distincta Corbett 1933 Dialeurodes distincta Corbett 1933 

Dialeurodes distinctus David & Subramaniam 1976 Dialeurodes davidi Mound & Halsey 1978

Dialeurodes dorsidemarcata Singh 1932 Asialeyrodes dorsidemarcata (Singh) 1932 

Dialeurodes dothioensis Dumbleton 1961 Dialeuropora decempuncta (Quaintance & Baker) 1917 

Dialeurodes doveri Corbett 1935 Aleuroclava doveri (Corbett) 1935 

Dialeurodes dryandrae Takahashi 1950 Gomenella dryandrae (Takahashi) 1950 

680 

Dialeurodes drypetesi Martin & Mound 2007 Dialeurodes drypetesi Martin & Mound 2007 

Dialeurodes dubia Corbett 1935 Dialeurodes dubia Corbett 1935 

Dialeurodes dumbeaensis Dumbleton 1961 Dialeurodes dumbeaensis Dumbleton 1961 

Dialeurodes egregissima Sampson & Drews 1941 Dialeurodes egregissima Sampson & Drews 1941 

Dialeurodes elaeagni Takahashi 1935 Singhiella elaeagni (Takahashi) 1935 

Dialeurodes endospermi Corbett 1935 Dialeurodes endospermi Corbett 1935 

Dialeurodes erythrinae Corbett 1935 Dialeurolobus erythrinae (Corbett) 1935 

Dialeurodes eugeniae (Maskell) Rusostigma eugeniae (Maskell) 1895 

Dialeurodes eugeniae (Maskell) 1895 Rusostigma eugeniae (Maskell) 1895 

Dialeurodes euryae Takahashi 1940 Massilieurodes euryae (Takahashi) 1940 

Dialeurodes evodiae Corbett 1935 Dialeurodes evodiae Corbett 1935 

Dialeurodes fici Takahashi 1932 Massilieurodes fici (Takahashi) 1932 

Dialeurodes ficicola Takahashi 1935 Massilieurodes ficicola (Takahashi) 1935 

Dialeurodes ficifolii Takahashi 1942 Singhiella ficifolii (Takahashi) 1942 

Dialeurodes fijensis (Kotinsky) 1907 Rhachisphora fijiensis (Kotinksy) 1907 

Dialeurodes filamentosa Corbett 1933 Aleuroclava filamentosa (Corbett) 1933 

Dialeurodes fletcheri Singh 1931 Singhius hibisci (Kotinsky) 1907 

Dialeurodes fodiens (Maskell) 1895 Aleyrodes fodiens Maskell 1895 

Dialeurodes formosensis Takahashi 1933 Massilieurodes formosensis (Takahashi) 1933 

Dialeurodes gardeniae Corbett 1935 Dialeurodes gardeniae Corbett 1935 

Dialeurodes gemurohensis Corbett 1935 Dialeurodes gemurohensis Corbett 1935 

Dialeurodes glomerata Singh 1931 Singhiella simplex (Singh) 1931 

Dialeurodes glutae Corbett 1935 Dialeurodes glutae Corbett 1935 

Dialeurodes greenwoodi Corbett 1936 Dialeurodes greenwoodi Corbett 1936 

Dialeurodes heterocera Bondar 1923 Dialeurodes heterocera Bondar 1923 

Dialeurodes hexpuncta Singh 1932 Dialeurodes hexpuncta Singh 1932 

Dialeurodes hongkongensis Takahashi 1941 Dialeurodes hongkongensis Takahashi 1941 

Dialeurodes icfreae Sundararaj & Dubey 2003 Dialeurodes icfreae Sundararaj & Dubey 2003 

Dialeurodes imperalis Bondar 1923 Dialeurodes imperialis Bondar 1923 

Dialeurodes indicus David & Subramaniam 1976 Dialeurodes indicus David & Subramaniam 1976 

Dialeurodes ixorae Singh 1931 Dialeuronomada ixorae (Singh) 1931 

Dialeurodes jendera Corbett 1935 Dialeuropora jendera (Corbett) 1935 

Dialeurodes joholensis Corbett 1935 Dialeurodes joholensis Corbett 1935 

Dialeurodes kamardini Corbett 1935 Cockerelliella kamardini (Corbett) 1935 

Dialeurodes kepongensis Corbett 1935 Dialeurodes kepongensis Corbett 1935

Dialeurodes kirishimensis Takahashi 1963 Massilieurodes kirishimensis (Takahashi) 1963 

Dialeurodes kirkaldyi (Kotinsky) 1907 Dialeurodes kirkaldyi (Kotinsky) 1907 

Dialeurodes kumargiriensis Sundararaj & Dubey 2006 Dialeurodes kumargiriensis Sundararaj & Dubey 2006 

Dialeurodes lanceolata Takahashi 1949 Aleuroclava lanceolata (Takahashi) 1949 

Dialeurodes langsat Corbett 1935 Dialeuropora langsat (Corbett) 1935 

Dialeurodes laos Takahashi 1942 Indoaleyrodes laos (Takahashi) 1942 

Dialeurodes lithocarpi Takahashi 1931 Dialeurodes lithocarpi Takahashi 1931 

Dialeurodes loranthi Corbett 1926 Dialeurodes loranthi Corbett 1926 

Dialeurodes lumpurensis Corbett 1935 Cockerelliella psidii (Corbett) 1935 

Dialeurodes machilicola Takahashi 1942 Dialeurodes machilicola Takahashi 1942 

Dialeurodes maculatus Bondar 1928 Dialeurodes maculata Bondar 1928 

Dialeurodes maculipennis Bondar 1923 Dialeurodes maculipennis Bondar 1923 

Dialeurodes malabaricus Jesudasan & David 1991 Singhiella malabaricus (Jesudasan and David) 1991 

Dialeurodes mangiferae Corbett 1935 Dialeuropora mangiferae (Corbett) 1935 

681 

Dialeurodes martini Jesudasan & David 1991 Dialeurodes drypetesi Martin & Mound 2007 

Dialeurodes mauritiensis Takahashi 1938 Dialeurolonga mauritiensis (Takahashi) 1938 

Dialeurodes maxima Quaintance & Baker 1917 Dialeurodes maxima Quaintance & Baker 1917 

Dialeurodes malabaricus Jesudasan and David 1991 Dialeurodes malabaricus Jesudasan and David 1991 

Dialeurodes mekonensis Takahashi 1942 Singhiella mekonensis (Takahashi) 1942 

Dialeurodes michoacanensis Sampson & Drews 1941 Dialeurodes denticulatus (Bondar) 1923 

Dialeurodes minuta Singh 1931 Minutaleyrodes minutus (Singh) 1931 

Dialeurodes mirabilis Takahashi 1942 Dialeurodes mirabilis Takahashi 1942 

Dialeurodes monticola Takahashi 1934 Massilieurodes monticola (Takahashi) 1934 

Dialeurodes multipori Takahashi 1932 Massilieurodes multipori (Takahashi) 1932 

Dialeurodes musae Corbett 1935 Dialeurodes musae Corbett 1935 

Dialeurodes natickis Baker & Moles 1923 Dialeurodes natickis Baker & Moles 1923 

Dialeurodes navarroi Bondar 1928 Dialeurodes navarroi Bondar 1928 

Dialeurodes nigeriae Cohic 1966 Dialeurodes nigeriae Cohic 1966 

Dialeurodes nitidus Singh 1932 Aleuroclava nitidus (Singh) 1932 

Dialeurodes nubifera (Berger) 1909 Singhiella citrifolii (Morgan) 1893 

Dialeurodes octoplicata Corbett 1935 Dialeurodes octoplicata Corbett 1935 

Dialeurodes ouchii Takahashi 1937 Dialeurodes ouchii Takahashi 1937 

Dialeurodes oweni Singh 1932 Dialeurodes oweni Singh 1932 

Dialeurodes pallida Singh 1931 Singhiella pallida (Singh) 1931 

Dialeurodes panacis Corbett 1935 Dialeurodes panacis Corbett 1935 

Dialeurodes papulae Singh 1932 Dialeurodes papulae Singh 1932 

Dialeurodes pauliani Cohic 1966 Dialeurodes pauliani Cohic 1966 

Dialeurodes perseae Corbett 1935 Dialeuropora decempuncta (Quaintance & Baker) 1917 

Dialeurodes philippinensis Takahashi 1936 Dialeurodes philippinensis Takahashi 1936

Dialeurodes pilahensis Corbett 1935 Dialeurodes pilahensis Corbett 1935 

Dialeurodes piperis Takahashi 1934 Singhiella piperis (Takahashi) 1934 

Dialeurodes platicus Bondar 1923 Dialeurodes platicus Bondar 1923 

Dialeurodes pseudocitri Takahashi 1942 Dialeurodes pseudocitri Takahashi 1942 

Dialeurodes psidii Corbett 1935 Cockerelliella psidii (Corbett) 1935 

Dialeurodes psychotriae Dumbleton 1961 Dialeurodes psychotriae Dumbleton 1961 

Dialeurodes punctata Corbett 1933 Dialeurodes punctata Corbett 1933 

Dialeurodes radiilinealis Quaintance & Baker 1917 Rabdostigma radiilinealis (Quaintance & Baker) 1917 

Dialeurodes radiipuncta Quaintance & Baker 1917 Dialeurodes radiipuncta Quaintance & Baker 1917 

Dialeurodes rangooni Singh 1932 Dialeurodes rangooni Singh 1932 

Dialeurodes rarasana Takahashi 1934 Massilieurodes rarasana (Takahashi) 1934 

Dialeurodes razalyi Corbett 1935 Dialeurodes razalyi Corbett 1935 

Dialeurodes rempangensis Takahashi 1949 Dialeurodes rempangensis Takahashi 1949 

Dialeurodes rengas Corbett 1935 Dialeurodes rengas Corbett 1935 

Dialeurodes reticulosa Corbett 1935 Dialeurodes reticulosa Corbett 1935 

Dialeurodes rhodamniae Corbett 1935 Cockerelliella rhodamniae (Corbett) 1935 

Dialeurodes rotunda Singh 1931 Dialeurodes rotunda Singh 1931 

Dialeurodes sakaki Takahashi 1958 Massilieurodes sakaki (Takahashi) 1958 

Dialeurodes saklaspurensis David 1976 Dialeurodes saklaspurensis David 1976 

Dialeurodes sandorici Corbett 1935 Dialeurodes sandorici Corbett 1935 

Dialeurodes selangorensis Corbett 1933 Rhachisphora selangorensis (Corbett) 1933 

Dialeurodes sembilanensis Corbett 1935 Cockerelliella sembilanensis (Corbett) 1935 

Dialeurodes sepangensis Corbett 1935 Dialeurodes sepangensis Corbett 1935 

Dialeurodes serdangensis Corbett 1935 Singhiella serdangensis (Corbett) 1935 

Dialeurodes sheryli (David) 2000 Dialeurodes sheryli (David) 2000 

Dialeurodes shintenensis Takahashi 1933 Dialeurodes shintenensis Takahashi 1933 

Dialeurodes shoreae Corbett 1933 Dialeurodes shoreae Corbett 1933 

Dialeurodes siemriepensis Takahashi 1942 Dialeurodes siemriepensis Takahashi 1942 

Dialeurodes simmondsi Corbett 1927 Dialeurodes simmondsi Corbett 1927 

Dialeurodes spina Singh 1931 Pealius spinus (Singh) 1931 

Dialeurodes spiniferosa Corbett 1933 Tuberaleyrodes spiniferosa (Corbett) 1933 

Dialeurodes striata Corbett 1935 Dialeurodes striata Corbett 1935 

Dialeurodes struthanthi Hempel 1901 Dialeurodes struthanthi Hempel 1901 

Dialeurodes subrotunda Takahashi 1935 Singhiella subrotunda (Takahashi) 1935 

Dialeurodes sundararajani Sundararaj & Dubey 2006 Dialeurodes sundararajani Sundararaj & Dubey 2006 

Dialeurodes sutepensis Takahashi 1942 Singhiella sutepensis (Takahashi) 1942 

Dialeurodes tanakai Takahashi 1942 Dialeurodes tanakai Takahashi 1942 

Dialeurodes tetrastigmae Takahashi 1934 Singhiella tetrastigmae (Takahashi) 1934 

Dialeurodes tokyonis (Kuwana) 1911 Rusostigma tokyonis (Kuwana) 1911 

682

Dialeurodes townsendi Quaintance & Baker 1917 Dialeurodes townsendi Quaintance & Baker 1917 

Dialeurodes tricolor Quaintance & Baker 1917 Dialeurodes tricolor Quaintance & Baker 1917 

Dialeurodes tridentifera Corbett 1933 Rhachisphora selangorensis (Corbett) 1933 

Dialeurodes tristylii Takahashi 1935 Rusostigma tristylii (Takahashi) 1935 

Dialeurodes tuberculatus Takahashi 1932 Dialeurodes citri (Ashmead) 1885 

Dialeurodes tuberculosa Corbett 1935 Dialeurodes tuberculosa Corbett 1935 

Dialeurodes turpiniae (Meganathan & David) 1994 Dialeurodes turpiniae (Meganathan & David) 1994 

Dialeurodes vanierieae Takahashi 1935 Singhiella vanieriae (Takahashi) 1935 

Dialeurodes vitis Corbett 1935 Dialeurodes vitis Corbett 1935 

Dialeurodes vulgaris Singh 1931 Dialeurodes vulgaris Singh 1931 

Dialeurodes wendlandiae Meganathan & David 1994 Dialeurodes wendlandiae Meganathan & David 1994 

Dialeurodes yercaudensis Jesudasan & David 1991 Dialeurodes yercaudensis Jesudasan & David 1991 

DIALEURODICUS COCKERELL 1902 DIALEURODICUS COCKERELL 1902 

Dialeurodicus bondariae Martin 2004 Dialeurodicus bondariae Martin 2004 

Dialeurodicus caballeroi Martin 2004 Dialeurodicus caballeroi Martin 2004 

Dialeurodicus cockerellii (Quaintance) 1900 Dialeurodicus cockerellii (Quaintance) 1900 

Dialeurodicus coelhi Bondar 1928 Dialeurodicus coelhi Bondar 1928 

Dialeurodicus cornutus Bondar 1923 Dialeurodicus cornutus Bondar 1923 

Dialeurodicus elongatus Dumbleton 1956 Stenaleyrodes vinsoni Takahashi 1938 

Dialeurodicus frontalis Bondar 1923 Dialeurodicus frontalis Bondar 1923 

Dialeurodicus maculatus Bondar 1928 Dialeurodicus maculatus Bondar 1928 

Dialeurodicus niger Bondar 1923 Dialeurodicus niger Bondar 1923 

Dialeurodicus pulcherrimus Quaintance & Baker 1913 Octaleurodicus pulcherrimus (Quaintance & Baker) 1913 

Dialeurodicus silvestri (Leonardi) 1910 Dialeurodicus silvestri (Leonardi) 1910 

Dialeurodicus silvestrii (Leonardi) 1910 Dialeurodicus silvestrii (Leonardi) 1910 

Dialeurodicus similis Bondar 1923 Dialeurodicus similis Bondar 1923 

Dialeurodicus tessellatus Quaintance & Baker 1913 Dialeurodicus tessellatus Quaintance & Baker 1913 

Dialeurodicus tracheiferus Sampson & Drews 1941 Dialeurodicus silvestrii (Leonardi) 

Dialeurodoides QUAINTANCE & BAKER 1914 Asterochiton QUAINTANCE & BAKER 1914 

Dialeurodoides [sic] auricolor Bondar 1923 Asterochiton auricolor (Bondar) 1923 

Dialeurodoides afer Priesner & Hosny 1934 Bemisia afer (Priesner & Hosny) 1934 

Dialeurodoides fagi (Maskell) 1890 Asterochiton fagi (Maskell) 1890 

Dialeurodoides simplex (Maskell) 1890 Asterochiton simplex (Maskell) 1890 

DIALEUROLOBUS DANZIG 1964 DIALEUROLOBUS DANZIG 1964 

Dialeurolobus erythrinae (Corbett) 1935 Dialeurolobus erythrinae (Corbett) 1935 

Dialeurolobus pulcher Danzig 1964 Dialeurolobus pulcher Danzig 1964 

Dialeurolobus rhamni Bink-Moenen 1992 Dialeurolobus rhamni Bink-Moenen 1992 

DIALEUROLONGA DOZIER 1928 DIALEUROLONGA DOZIER 1928 

Dialeurolonga africana (Newstead) 1921 Dialeurolonga africana (Newstead) 1921 

683

Dialeurolonga agauriae Takahashi 1951 Dialeurolonga aguariae Takahashi 1951 

Dialeurolonga akureensis Mound 1965 Extensaleyrodes akureensis (Mound) 1965 

Dialeurolonga ambilaensis Takahashi 1955 Dialeurolonga ambilaensis Takahashi 1955 

Dialeurolonga angustata Takahashi 1961 Dialeurolonga ambilaensis Takahashi 1961 

Dialeurolonga aphloiae Takahashi 1955 Dialeurolonga aphloiae Takahashi 1955 

Dialeurolonga bambusae Takahashi 1961 Dialeurolonga bambusae Takahashi 1961 

Dialeurolonga bambusicola (Takahashi) Dialeurolonga bambusicola Takahashi 1951 

Dialeurolonga brevispina Takahashi 1951 Dialeurolonga brevispina Takahashi 1951 

Dialeurolonga davidi Dubey & Sundararaj 2006 Dialeurolonga davidi Dubey & Sundararaj 2006 

Dialeurolonga elliptica Takahashi 1955 Dialeurolonga elliptica Takahashi 1955 

Dialeurolonga elongata Dozier 1928 Dialeurolonga elongata Dozier 1928 

Dialeurolonga emarginata Mound 1965 Dialeurodes emarginata (Mound) 1965 

684 

Dialeurolonga erythroxylonis Takahashi 1955 Dialeurolonga erythroxylonis Takahashi 1955 

Dialeurolonga eugeniae Takahashi 1951 Dialeurolonga eugeniae Takahashi 1951 

Dialeurolonga fici David & Subramaniam 1976 Dialeurolonga fici David & Subramaniam 1976 

Dialeurolonga graminis (Takahashi) 1951 Dialeurolonga graminis (Takahashi) 1951 

Dialeurolonga guettardae Martin 2005 Dialeurolonga guettardae Martin 2005 

Dialeurolonga hoyti Mound 1965 Dialeurolonga hoyti Mound 1965 

Dialeurolonga kumargiriensis Dubey & Sundararaj 2006 Dialeurolonga kumargiriensis Dubey & Sundararaj 2006 

Dialeurolonga lagerstroemiae Jesudasan & David 1991 Dialeurolonga lagerstroemiae Jesudasan & David 1991 

Dialeurolonga lamtoensis Cohic 1969 Dialeurolonga lamtoensis Cohic 1969 

Dialeurolonga lata Takahashi 1955 Dialeurolonga lata Takahashi 1955 

Dialeurolonga maculata (Singh) 1931 Dialeurolonga maculata (Singh) 1931 

Dialeurolonga maculata Takahashi 1951 Dialeurolonga takahashii David & Jesudasan 1989 

Dialeurolonga malleshwaramensis Sundararaj 2001 Dialeurolonga malleshwaramensis Sundararaj 2001 

Dialeurolonga mameti Takahashi 1955 Dialeurolonga mameti Takahashi 1955 

Dialeurolonga mauritiensis (Takahashi) 1938 Dialeurolonga mauritiensis (Takahashi) 1938 

Dialeurolonga milloti Takahashi 1951 Dialeurolonga milloti Takahashi 1951 

Dialeurolonga multipapilla Takahashi 1955 Dialeurolonga multipapilla Takahashi 1955 

Dialeurolonga multipori Dubey & Sundararaj 2006 Dialeurolonga multipori Dubey & Sundararaj 2006 

Dialeurolonga multituberculata Dubey & Sundararaj 2006 Dialeurolonga multituberculata Dubey & Sundararaj 2006 

Dialeurolonga nigra Takahashi & Mamet 1952 Dialeurolonga nigra Takahashi & Mamet 1952 

Dialeurolonga operculobata Martin & Carver 1999 Dialeurolonga operculobagta Martin & Carver 1999 

Dialeurolonga paradoxa Takahashi 1955 Dialeurolonga paradoxa Takahashi 1955 

Dialeurolonga paucipapillata Cohic 1969 Dialeurolonga paucipapillata Cohic 1969 

Dialeurolonga pauliani Takahashi 1951 Dialeurolonga pauliani Takahashi 1951 

Dialeurolonga perinetensis Takahashi & Mamet 1952 Dialeurolonga perinetensis Takahashi & Mamet 1952 

Dialeurolonga phyllarthronis Takahashi 1955 Dialeurolonga phyllarthronis Takahashi 1955 

Dialeurolonga pseudocephalidistincta Sundararaj & Dubey 2006 Dialeurolonga pseudocephalidistincta Sundararaj &


Dubey 2006 


Dialeurolonga rhamni Takahashi 1961 Dialeurolonga rhamni Takahashi 1961 

Dialeurolonga robinsoni Takahashi & Mamet 1952 Dialeurolonga robinsoni Takahashi & Mamet 1952 

Dialeurolonga rotunda Takahashi 1961 Dialeurolonga rotunda Takahashi 1961 

Dialeurolonga rusostigmoides Martin 1999 Dialeurolonga rusostigmoides Martin 1999 

Dialeurolonga sarcocephali Cohic 1966 Dialeurolonga hoyti Mound 1965 

Dialeurolonga similis Takahashi 1955 Dialeurolonga similis Takahashi 1955 

Dialeurolonga simplex Takahashi 1955 Dialeurolonga simplex Takahashi 1955 

Dialeurolonga strychnosicola Cohic 1966 Dialeurolonga strychnosicola Cohic 1966 

Dialeurolonga subrotunda Takahashi 1955 Dialeurolonga subrotunda Takahashi 1955 

Dialeurolonga swainei Martin 1999 Dialeurolonga swainei Martin 1999 

Dialeurolonga takahashii David & Jesudasan 1989 Dialeurolonga takahashii David & Jesudasan 1989 

Dialeurolonga tambourissae Takahashi 1955 Dialeurolonga tambourissae Takahashi 1955 

Dialeurolonga trialeuroides Takahashi & Mamet 1952 Dialeurolonga trialeuroides Takahashi & Mamet 1952 

Dialeurolonga vendranae Takahashi 1961 Dialeurolonga vendranae Takahashi 1961 

DIALEURONOMADA QUAINTANCE & BAKER 1917 DIALEURONOMADA QUAINTANCE & BAKER 1917 

Dialeuronomada ayyanarensis Sundararaj & David 1991 Dialeuronomada ayyanarensis Sundararaj & David 1991 

Dialeuronomada binkae Sundararaj & David 1991 Dialeuronomada binkae Sundararaj & David 1991 

Dialeuronomada biventralis Sundararaj & David 1991 Dialeuronomada biventralis Sundararaj & David 1991 

Dialeuronomada canthiae Sundararaj & David 1991 Dialeuronomada canthiae Sundararaj & David 1991 

Dialeuronomada dissimilis (Quaintance & Baker) 1917 Dialeuronomada dissimilis (Quaintance & Baker) 1917 

Dialeuronomada gigantica Sundararaj & David 1991 Dialeuronomada gigantica Sundararaj & David 1991 

Dialeuronomada granulata Sundararaj & David 1991 Dialeuronomada granulata Sundararaj & David 1991 

Dialeuronomada ixorae (Singh) 1931 Dialeuronomada ixorae (Singh) 1931 

Dialeuronomada keralaensis Meganathan & David 1994 Dialeuronomada keralaensis Meganathan & David 1994 

Dialeuronomada martini Sundararaj & David 1991 Dialeuronomada martini Sundararaj & David 1991 

Dialeuronomada nagpurensis Sundararaj & David 1991 Dialeuronomada nagpurensis Sundararaj & David 1991 

Dialeuronomada palmata Sundararaj & David 1991 Dialeuronomada palmata Sundararaj & David 1991 

Dialeuronomada remadeviae Dubey & Sundararaj 2004 Dialeuronomada remadeviae Dubey & Sundararaj 2004 

Dialeuronomada rubiphaga Dubey & Sundararaj 2004 Dialeuronomada ruiphaga Dubey & Sundararaj 2004 

Dialeuronomada russellae Sundararaj & David 1991 Dialeuronomada russellae Sundararaj & David 1991 

Dialeuronomada saklespurensis Regu & David 1993 Dialeuronomada saklespurensis Regu & David 1993 

DIALEUROPORA QUAINTANCE & BAKER 1917 DIALEUROPORA QUAINTANCE & BAKER 1917 

Dialeuropora brideliae (Takahashi) 1932 Dialeuropora brideliae (Takahashi) 1932 

Dialeuropora cogniauxiae Cohic 1966 Dialeuropora papillata Cohic 1966 

Dialeuropora congoensis Cohic 1966 Dialeuropora congoensis Cohic 1966 

Dialeuropora cumiugum (Singh) 1932 Russellaleyrodes cumiugum (Singh) 1932 

Dialeuropora decempuncta Quaintance & Baker 1917 Dialeuropora decempuncta (Quaintance & Baker) 1917 

685

Dialeuropora dothioensis (Dumbleton) 1961 Dialeuropora decempuncta (Quaintance & Baker) 1917 

686 

Dialeuropora hassensanensis Takahashi 1934 Dialeuropora hassensanensis Takahashi 1934 

Dialeuropora heptapora Regu & David 1991 Dialeuropora heptapora Regu & David 1991 

Dialeuropora holboelliae Young 1944 Dialeuropora holboelliae Young 1944 

Dialeuropora indochinensis Takahashi 1942 Dialeuropora indochinensis Takahashi 1942 

Dialeuropora malayensis (Corbett) 1935 Dialeuropora malayensis (Corbett) 1935 

Dialeuropora mangiferae (Corbett) 1935 Dialeuropora mangiferae (Corbett) 1935 

Dialeuropora murrayae (Takahashi) 1931 Dialeuropora murrayae (Takahashi) 1931 

Dialeuropora ndiria Gameel 1971 Dialeuropora papillata Cohic 1966 

Dialeuropora papillata Cohic 1966 Dialeuropora papillata Cohic 1966 

Dialeuropora perseae (Corbett) Dialeuropora decempuncta (Quaintance & Baker) 1917 

Dialeuropora perseae (Corbett) 1935 Dialeuropora decempuncta (Quaintance & Baker) 1917 

Dialeuropora photiniana (Chen) Dialeuropora photiniana (Chen) 

Dialeuropora platysepali Cohic 1966 Dialeuropora papillata Cohic 1966 

Dialeuropora portugaliae Cohic 1966 Dialeuropora portugaliae Cohic 1966 

Dialeuropora pterolobiae David & Subramaniam 1976 Dialeuropora pterolobiae David & Subramaniam 1976 

Dialeuropora setigerus (Takahashi) 1932 Dialeuropora decempuncta (Quaintance & Baker) 1917 

Dialeuropora silvarum (Corbett) 1935 Dialeuropora silvarum (Corbett) 1935 

Dialeuropora urticata Young 1944 Dialeuropora urticata Young 1944 

Dialeuropora viburni (Takahashi) 1933 Dialeuropora viburni (Takahashi) 1933 

DIALEUROTRACHELUS TAKAHASHI 1942 DIALEUROTRACHELUS TAKAHASHI 1942 

Dialeurotrachelus cambodiensis Takahashi 1942 Dialeurotrachelus cambodiensis Takahashi 1942 

DISIPHON RUSSELL 1993 DISIPHON RUSSELL 1993 

Disiphon dubienus (Bondar) 1923 Disiphon dubienus (Bondar) 1923 

Disiphon russellae Martin 2005 Disiphon russellae Martin 2005 

Distinctaleyrodes Dubey and Sundararaj 2006 Distinctaleyrodes Dubey and Sundararaj 2006 

Distinctaleyrodes setosus Dubey and Sundararaj 2006 Distinctaleyrodes setosus Dubey and Sundararaj 2006 

DOTHIOIA DUMBLETON 1961 DOTHIOIA DUMBLETON 1961 

Dothioia bidentatus Dumbleton 1961 Dothioia bidentatus Dumbleton 1961 

DUMBLETONIELLA JESUDASAN & DAVID 1990 DUMBLETONIELLA JESUDASAN & DAVID 1990 

Dumbletoniella callistemoni Martin 1999 Dumbletoniella callistemoni Martin 1999 

Dumbletoniella ellipticae (Dumbleton) 1956 Dumbletoniella ellipticae (Dumbleton) 1956 

Dumbletoniella eucalypti (Dumbleton) 1957 Dumbletoniella eucalypti (Dumbleton) 1957 

Dumbletoniella pittospori Martin & Carver 1999 Dumbletoniella pittospori Martin & Carver 1999 

Dumbletoniella rotunda Martin & Carver 1999 Dumbletoniella rotunda Martin & Carver 1999 

Dumbletoniella xanthorrhoeae Martin 1999 Dumbletoniella xanthorrhoeae Martin 1999 

EDITAALEYODES DAVID 2005 EDITAALEYRODES DAVID 2005 

Editaaleyrodes indicus David 2005 Editaaleyrodes indica David 2005 

EUDIALEURODICUS QUAINTANCE & BAKER 1915 EUDIALEURODICUS QUAINTANCE & BAKER 1915

Eudialeurodicus bodkini Quaintance & Baker 1915 Eudialeurodicus bodkini Quaintance & Baker 1915 

687 

EXTENSALEYRODES BINK-MOENEN 1983 EXTENSALEYRODES BINK-MOENEN 1983 

Extensaleyrodes akureensis (Mound) 1965 Extensaleyrodes akureensis (Mound) 1965 

FASCALEYRODES BINK-MOENEN 1983 FASCALEYRODES BINK-MOENEN 1983 

Fascaleyrodes palmae (Gameel) 1968 Fascaleyrodes palmae (Gameel) 1968 

Fascaleyrodes rara Bink-Moenen 1983 Fascaleyrodes rara Bink-Moenen 1983 

FILICALEYRODES TAKAHASHI 1962 FILICALEYRODES TAKAHASHI 1962 

Filicaleyrodes bosseri Takahashi 1962 Filicaleyrodes bosseri Takahashi 1962 

Filicaleyrodes williamsi (Trehan) 1938 Filicaleyrodes williamsi (Trehan) 1938 

FIPPATALEYRODES SUNDARARAJ & DAVID 1992 FIPPATALEYRODES SUNDARARAJ & DAVID 1992 

Fippataleyrodes cinnamomi Dubey & Sundararaj 2005 Fippataleyrodes cinnamomi Dubey & Sundararaj 2005 

Fippataleyrodes indica Sundararaj & David 1992 Fippataleyrodes indica Sundararaj & David 1992 

Fippataleyrodes litseae Sundararaj & David 1992 Fippataleyrodes litseae Sundararaj & David 1992 

Fippataleyrodes multipori Dubey & Sundararaj 2005 Fippataleyrodes multipori Dubey & Sundararaj 2005 

Fippataleyrodes yellapurensis Dubey & Sundararaj 2005 Fippataleyrodes yellapurensis Dubey & Sundararaj 2005 

Frauenfeldiella Gomez-Menor 1943 Aleurotrachelus Quaintance & Baker 1914 

Frauenfeldiella jelinekii (Frauenfeld) 1867 Aleurotuba jelinekii (Frauenfeld) 1867 

GAGUDJUIA MARTIN 1999 GAGUDJUIA MARTIN 1999 

Gagudjuia allosyncarpiae Martin 1999 Gagudjuia allosyncarpiae Martin 1999 

Gigaleurodes Quaintance & Baker Dialeurodes Cockerell 1902 

Gigaleurodes multipori (Takahashi) 1932 Dialeurodes multipori Takahashi 1932 

Gigaleurodes splendens (Regu & David) 1993 Massilieurodes homonoiae (Jesudasan & David) 1991 

GOMENELLA DUMBLETON 1961 GOMENELLA DUMBLETON 1961 

Gomenella dryandrae (Takahashi) 1950 Gomenella dryandrae (Takahashi) 1950 

Gomenella multipora Dumbleton 1961 Gomenella multipora Dumbleton 1961 

Gomenella reflexa Dumbleton 1961 Gomenella reflexa Dumbleton 1961 

Greeniella David 1993 Rhachisphora Quaintance & Baker 1917 

Gymnaleurodes Sampson & Drews 1940 Trialeurodes Cockerell 1902 

Gymnaleurodes bellissima Sampson & Drews 1940 Trialeurodes bellissima (Sampson & Drews) 1940 

Harpaleyrodes Bink-Moenen 1983 Harpaleyrodes Bink-Moenen 1983 

Harpaleyrodes tuberculata Bink-Moenen 1983 Harpaleyrodes tuberculata Bink-Moenen 1983 

Heidea cretacica Schlee 1970 Heidea cretacica Schlee 1970 

Hempelia Sampson & Drews 1941 Aleurothrixus Quaintance and Baker 1914 

Hempelia chivelensis Sampson & Drews 1941 Aleurothrixus chivelensis (Sampson & Drews) 1941 

HESPERALEYRODES SAMPSON 1943 HESPERALEYRODES SAMPSON 1943 

Hesperaleyrodes michoacanensis Sampson 1943 Hesperaleyrodes michoacanensis Sampson 1943 

HETERALEYRODES TAKAHASHI 1942 HETERALEYRODES TAKAHASHI 1942 

Heteraleyrodes bambusae Takahashi 1942 Heteraleyrodes bambusae Takahashi 1942 

Heteraleyrodes bambusicola Takahashi 1951 Heteraleyrodes bambusicola Takahashi 1951

HETEROBEMISIA TAKAHASHI 1957 HETEROBEMISIA TAKAHASHI 1957 

Heterobemisia alba Takahashi 1957 Heterobemisia alba Takahashi 1957 

Hexaleurodicus HEMPEL 1923 Aleuronudus HEMPEL 1922 

Hexaleurodicus (Drewsia) SAMPSON 1943 Aleuronudus HEMPEL 1922 

Hexaleurodicus (Drewsia) ferrisi Sampson & Drews 1941 Aleuronudus manni (Baker) 1923 

Hexaleurodicus ferrisi Sampson & Drews 1941 Aleuronudus manni (Baker) 1923 

Hexaleurodicus jaciae Bondar 1923 Aleuronudus jaciae (Bondar 1923) 

Hindaleyrodes Meganathan & David 1994 Aleuroclava Singh 

Hindaleyrodes hindustanicus Meganathan & David 1994 Aleuroclava hindustanicus (Meganathan & David) 1994 

ICFREALEYRODES DUBEY & SUNDARARAJ 2007 ASIALEYRODES CORBETT 1935 

Icfrealeyrodes indica Dubey & Sundararaj 2007 Asialeyrodes dubius Martin & Mound 2007 

INDOALEYRODES DAVID & SUBRAMANIAM 1973 INDOALEYRODES DAVID & SUBRMANIAM 1973 

Indoaleyrodes glochidioni Martin & Carver 1999 Indoaleyrodes glochidioni Martin & Carver 1999 

Indoaleyrodes laos (Takahashi) 1942 Indoaleyrodes laos (Takahashi) 1942 

Indoaleyrodes pseudoculatus Martin 1985 Indoaleyrodes pseudoculatus Martin 1985 

Indoaleyrodes pustulatus David & Subramaniam 1973 Indoaleyrodes laos (Takahashi) 1942 

Japaneyrodes Zahradnik 1962 Aleuroclava Singh 1931 

Japaneyrodes hikosanensis (Takahashi) 1938 Aleuroclava hikosanensis (Takahashi) 1938 

Japaneyrodes similis (Takahashi) 1938 Aleuroclava similis (Takahashi) 1938 

Japaneyrodes similis europaeus Zahradnik 1962 Aleuroclava similis (Takahashi) 1938 

Japaneyrodes similis suborientalis Danzig 1966 Aleuroclava similis (Takahashi) 1938 

Japaneyrodes trachelospermi (Takahashi) Aleuroclava trachelospemi (Takahashi) 1938 

688 

Jeannelaleyrodes Cohic 1966 Neoaleurotrachelus Takahashi & Mamet 1952 

Jeannelaleyrodes bertilloni Cohic 1966 Neoaleurotrachelus bertilloni (Cohic) 1966 

Jeannelaleyrodes graberi Cohic 1968 Neoaleurotrachelus graberi (Cohic) 1968 

JUGLASALEYRODES COHIC 1966 JUGLASALEYRODES COHIC 1966 

Juglasaleyrodes orstomensis Cohic 1966 Juglasaleyrodes orstomensis Cohic 1966 

JULEYRODES SHCHERBAKOV 2000 JULEYRODES SHCHERBAKOV 2000 

Juleyrodes gilli Shcherbakov 2000 Juleyrodes gilli Shcherbakov 2000 

Juleyrodes visnyai Shcherbakov 2000 Juleyrodes visnyai Shcherbakov 2000 

Kanakarajiella David & Sundararaj 1993 Dialeurodes Cockerell 

Kanakarajiella bassiae (David and Subramaniam) 1976 Singhiella bassiae (David and Subramaniam) 1976 

Kanakarajiella cardamomi (David and Subramaniam) 1976 Singhiella cardamomi (David and Subramaniam) 1976 

Kanakarajiella pallida (Singh) 1931 Singhiella pallida (Singh) 1931 

Kanakarajiella turpiniae Meganathan & David 1994 Dialeurodes turpiniae (Meganathan & David) 1994 

Kanakarajiella vulgaris (Singh) 1931 Dialeurodes vulgaris Singh 1931 

KERALALEYRODES MEGANATHAN & DAVID 1994 KERALEYRODES MEGANATHAN & DAVID 1994 

Keralaleyrodes indicus Meganathan & David 1994 Keraleyrodes indicus Meganathan & David 1994 

LAINGIELLA CORBETT 1926 LAINGIELLA CORBETT 1926

Laingiella bambusae Corbett 1926 Laingiella bambusae Corbett 1926 

Lankaleurodes David 1993 Dialeurodes Cockerell 

Lankaleurodes radiipuncta (Quaintance & Baker) 1917 Dialeurodes radiipuncta Quaintance & Baker 1917 

LECANOIDEUS QUAINTANCE & BAKER 1913 LECANOIDEUS QUAINTANCE & BAKER 1913 

Lecanoideus floccissimus Martin et al 1997 Lecanoideus floccissimus Martin et al. 1997 

Lecanoideus giganteus (Quaintance & Baker) 1913 Lecanoideus mirabilis (Cockerell) 1898 

Lecanoideus mirabilis (Cockerell) 1898 Lecanoideus mirabilis (Cockerell) 1898 

LEONARDIUS QUAINTANCE & BAKER 1913 LEONARDIUS QUAINTANCE & BAKER 1913 

Leonardius kellyae Martin 2004 Leonardius kellyae Martin 2004 

Leonardius lahillei (Leonardi) 1910 Leonardius lahillei (Leonardi) 1910 

Leonardius loranthi Bondar 1923 Leonardius lahillei (Leonardi) 1910 

LEUCOPOGONELLA DUMBLETON 1961 LEUCOPOGONELLA DUMBLETON 1961 

Leucopogonella apectenata Dumbleton 1961 Leucopogonella apectenata Dumbleton 1961 

Leucopogonella pallida Dumbleton 1961 Leucopogonella pallida Dumbleton 1961 

Leucopogonella simila Dumbleton 1961 Leucopogonella simila Dumbleton 1961 

Leucopogonella sinuata Dumbleton 1961 Leucopogonella sinuata Dumbleton 1961 

LIPALEYRODES TAKAHASHI 1962. LIPALEYRODES TAKAHASHI 1962 

Lipaleyrodes atriplex (Froggatt) 1911 Lipaleyrodes atriplex (Froggatt) 1911 

Lipaleyrodes breyniae (Singh) 1931 Lipaleyrodes breyniae (Singh) 1931 

Lipaleyrodes crossandreae David & Subramaniam 1976 Lipaleyrodes crossandreae David & Subramaniam 1976 

Lipaleyrodes emiliae Chen & Ko 2006 Lipaleyrodes emiliae Chen & Ko 2006 

Lipaleyrodes euphorbiae David & Subramaniuam 1976 Lipaleyrodes euphorbiae David & Subramaniam 1976 

Lipaleyrodes hargreavesi (Corbett) 1935 Lipaleyrodes hargreavesi (Corbett) 1935 

Lipaleyrodes leguminicola (Takahashi) 1942 Lipaleyrodes leguminicola (Takahashi) 1942 

Lipaleyrodes phyllanthi Takahashi 1962 Lipaleyrodes phyllanthi Takahashi 1962 

Lipaleyrodes vernoniae David & Thenmozhi 1996 Lipaleyrodes vernoniae David & Thenmozhi 1996 

Luederwaldtiana Hempel 1923 Aleurotrachelus Quaintance & Baker 1922 

Luederwaldtiana eriosemae Hempel 1923 Aleurotrachelus eriosemae (Hempel) 1923 

MALAYALEYRODES CORBETT 1935 MALAYALEYRODES CORBETT 1935 

Malayaleyrodes lumpurensis Corbett 1935 Malayaleyrodes lumpurensis Corbett 1935 

MARGINALEYRODES TAKAHASHI 1961 MARGINALEYRODES TAKAHASHI 1961 

Marginaleyrodes angolensis Cohic 1966 Rugaleyrodes angolensis (Cohic) 1966 

Marginaleyrodes fanalae (Takahashi) Marginaleyrodes fanalae (Takahashi) 1951 

689 

Marginaleyrodes fenestrata (Takahashi) Marginaleyrodes fenestrata (Takahashi) 1955 

Marginaleyrodes ixorae Takahashi 1961 Marginaleyrodes ixorae Takahashi 1961 

Marginaleyrodes madagascariensis (Takahashi) Marginaleyrodes madagascariensis (Takahashi) 1951 

Marginaleyrodes madagascariensis (Takahashi) 1951 Marginaleyrodes madagascariensis (Takahashi) 1951 

Marginaleyrodes tetracerae Cohic 1966 Rugaleyrodes tetracerae (Cohic) 1966 

Marginaleyrodes tsinjoarivona (Takahashi) Marginaleyrodes tsinjoarivona (Takahashi) 1955

Marginaleyrodes tsinjoarivona (Takahashi) 1955 Marginaleyrodes tsinjoarivona (Takahashi) 1955 

MARTINIELLA JESUDASAN & DAVID 1990 MARTINIELLA JESUDASAN & DAVID 1990 

Martiniella ayyari Sundararaj & David 1993 Martiniella ayyari Sundararaj & David 1993 

Martiniella canangae (Corbett) 1935 Martiniella canangae (Corbett) 1935 

Martiniella lefroyi Sundararaj & David 1993 Martiniella lafroyi Sundararaj & David 1993 

Martiniella macarangae (Corbett) 1935 Martiniella macarangae (Corbett) 1935 

Martiniella papillata Sundararaj & Dubey 2004 Martiniella papillata Sundararaj & David 2004 

MASSILIEURODES GOUX 1939 MASSILIEURODES GOUX 1939 

Massilieurodes alabamensis Jensen 2001 Massilieurodes alabamensis Jensen 2001 

Massilieurodes americanus Jensen 2001 Massilieurodes americanus Jensen 2001 

Massilieurodes chittendeni (Laing) 1928 Massilieurodes chittendeni (Laing) 1928 

Massilieurodes curiosa Jensen 2001 Massilieurodes curiosa Jensen 2001 

Massilieurodes euryae (Takahashi) 1940 Massilieurodes euryae (Takahashi) 1940 

Massilieurodes fici (Takahashi) 1932 Massilieurodes fici (Takahashi) 1932 

Massilieurodes ficicola (Takahashi) 1935 Massilieurodes ficicola (Takahashi) 1935 

Massilieurodes ficifolii (Takahashi) 1942 Singhiella ficifolii (Takahashi) 1942 

690 

Massilieurodes formosensis (Takahashi) 1933 Massilieurodes formosensis (Takahashi) 1933 

Massilieurodes homonoiae (Jesudasan & David) 1991 Massilieurodes homonoiae (Takahashi) 1933 

Massilieurodes kirishimensis (Takahashi) Massilieurodes kirishimensis (Takahashi) 1963 

Massilieurodes monticola (Takahashi) 1934 Massilieurodes monticola (Takahashi) 1934 

Massilieurodes multipori (Takahashi) 1932 Massilieurodes multipori (Takahashi) 1932 

Massilieurodes myricae Jensen 2001 Massilieurodes myricae Jensen 2001 

Massilieurodes rarasana (Takahashi) 1934 Massilieurodes rarasana (Takahashi) 1934 

Massilieurodes sakaki (Takahashi) 1958 Massilieurodes sakaki (Takahashi) 1958 

Massilieurodes setiger Goux 1939 Massilieurodes setiger Goux 1939 

Massilieurodes splendens (Regu and David) 1993 Massilieurodes homonoiae (Jesudasan & David) 1991 

METABEMISIA TAKAHASHI 1963 METABEMISIA TAKAHASHI 1963 

Metabemisia distylii Takahashi 1963 Metabemisia distylii Takahashi 1963 

Metabemisia filicis Mound 1967 Metabemisia filicis Mound 1967 

Metabemisia palawana Martin 2001 Metabemisia palawana Martin 2001 

METALEURODICUS QUAINTANCE & BAKER 1913 METALEURODICUS QUAINTANCE & BAKER 1913 

Metaleurodicus altissimus (Quaintance) 1900 Nealeurodicus altissimus (Quaintance) 1900 

Metaleurodicus arcanus Martin 2004 Metaleurodicus arcanus Martin 2004 

Metaleurodicus cardini (Back) 1912 Metaleurodicus cardini (Back) 1912 

Metaleurodicus griseus (Dozier) 1936 Metaleurodicus griseus (Dozier) 1936 

Metaleurodicus jequiensis Bondar 1928 Aleuronudus jequiensis (Bondar) 1928 

Metaleurodicus lacerdae (Signoret) 1833 Metaleurodicus lacerdae (Signoret) 1883 

Metaleurodicus manni (Baker) 1923 Aleuronudus manni (Baker) 1923 

Metaleurodicus melzeri Bondar 1928 Aleuronudus melzeri (Bondar) 1928

Metaleurodicus minimus (Quaintance) 1900 Metaleurodicus minimus (Quaintance) 1900 

Metaleurodicus phalaenoides (Blanchard) Metaleurodicus phalaenoides (Blanchard) 1852 

Metaleurodicus pigeanus (Baker & Moles) 1923 Australeurodicus pigeanus (Baker & Moles) 1923 

Metaleurodicus stelliferus Bondar 1923 Metaleurodicus stelliferus Bondar 1923 

Metaleurodicus tenuis Martin 2004 Metaleurodicus tenuis Martin 2004 

Metaleurodicus variporus Martin 2004 Metaleurodicus variporus Martin 2004 

METALEYRODES SAMPSON 1943 METALEYRODES SAMPSON 1943 

Metaleyrodes oceanica (Takahashi) 1939 Metaleyrodes oceanica (Takahashi) 1939 

Mexicaleyrodes Sampson & Drews 1941 Tetraleurodes Cockerell 1902 

Mexicaleyrodes contigua Sampson & Drews 1941 Tetraleurodes contigua (Sampson & Drews) 1941 

MINUTALEYRODES JESUDASAN & DAVID 1990 MINUTALEYRODES JESUDASAN & DAVID 1990 

Minutaleyrodes cherasensis (Corbett) 1935 Minutaleyrodes cherasensis (Corbett) 1935 

Minutaleyrodes indicus Meganathan & David 1994 Minutaleyrodes indicus Meganathan & David 1994 

Minutaleyrodes kolliensis (David) 1977 Minutaleyrodes kolliensis (David) 1977 

Minutaleyrodes minutus (Singh) 1931 Minutaleyrodes minuta (Singh) 1931 

MIXALEYRODES TAKAHASHI 1936 MIXALEYRODES TAKAHASHI 1936 

Mixaleyrodes elongatus Meganathan & David 1994 Cohicaleyrodes elongatus Meganathan & David 1994 

Mixaleyrodes indicus David & Selvakumaran 1987 Cohicaleyrodes indicus (David and Selvakumaran) 1987 

Mixaleyrodes polypodicola Takahashi 1963 Mixaleyrodes polypodicola Takahashi 1963 

Mixaleyrodes polystichi Takahashi 1936 Mixaleyrodes polystichi Takahashi 1936 

Mohanasundaramiella David 2000 Crenidorsum Russell 

Mohanasundaramiella rubiae David 2000 Crenidorsum rubiae (David) 2000 

Moundiella David 1974 Viennotaleyrodes Cohic 1968 

691 

Moundiella indicus Jesudasan & David 1991 Viennotaleyrodes megapapillae (Singh) 1932 

Moundiella megapapillae (Singh) 1932 Viennotaleyrodes megapapillae (Singh) 1932 

Nealeurochiton Sampson 1943 Aleurochiton TULLGREN 1907 

Nealeurochiton forbesii (Ashmead) 1893 Aleurochiton forbesii (Ashmead) 1893 

Nealeurochiton pseudoplatani (Visnya) 1936 Aleurochiton pseudoplatani Visnya 1936 

NEALEURODICUS HEMPEL 1923 NEALEURODICUS HEMPEL 1923 

Nealeurodicus altissimus (Quaintance) 1900 Nealeurodicus altissimus (Quaintance) 1900 

Nealeurodicus bakeri (Bondar) 1923 Nealeurodicus bakeri (Bondar) 1923 

Nealeurodicus fallax Martin 2004 Nealeurodicus fallax Martin 2004 

Nealeurodicus paulistus Hempel 1923 Nealeurodicus paulistus Hempel 1923 

Nealeurodicus petiolaris Martin 2004 Nealeurodicus petiolaris Martin 2004 

NEALEYRODES HEMPEL 1923 NEALEYRODES HEMPEL 1923 

Nealeyrodes bonariensis Hempel 1923 Nealeyrodes bonariensis Hempel 1923 

NEOALEURODES BONDAR 1923 NEOALEURODES BONDAR 1923 

Neoaleurodes clandestinus Bondar 1923 Neoaleurodes clandestinus Bondar 1923 

Neoaleurolobus Takahashi 1951 Aleurolobus Quaintance & Baker 1914

Neoaleurolobus musae (Corbett) 1935 Aleurolobus musae Corbett 1935 

NEOALEUROTRACHELUS TAKAHASHI & MAMET 1952 NEOALEUROTRACHELUS TAKAHASHI & MAMET 1952 

Neoaleurotrachelus aphloiae Takahashi & Mamet 1952 Neoaleurotrachelus aphloiae Takahashi & Mamet 1952 

Neoaleurotrachelus bertilloni (Cohic) 1966 Neoaleurotrachelus bertilloni (Cohic) 1966 

Neoaleurotrachelus graberi (Cohic) 1968 Neoaleurotrachelus graberi (Cohic) 1968 

Neoaleurotrachelus sudanensis (Gameel) 1968 Neoaleurotrachelus sudanensis (Gameel) 1968 

Neobemisia VISNYA 1941 Asterobemisia TREHAN 1940 

Neobemisia atraphaxius Danzig 1969 Asterobemisia atraphaxius (Danzig) 1969 

Neobemisia obenbergeri Zahradnik 1961 Asterobemisia obenbergeri (Zahradnik) 1961 

Neobemisia paveli Zahradnik 1961 Asterobemisia paveli (Zahradnik) 1961 

Neobemisia trifolii Danzig 1966 Asterobemisia trifolii (Danzig) 1966 

Neobemisia yanagicola (Takahashi) 1933 Asterobemisia yanagicola (Takahashi) 1933 

NEOMASKELLIA QUAINTANCE & BAKER 1913 NEOMASKELLIA QUAINTANCE & BAKER 1913 

Neomaskellia andropogonis Corbett 1926 Neomaskellia andropogonis Corbett 1926 

Neomaskellia bergii (Signoret) 1868 Neomaskellia bergii (Signoret) 1868 

Neomaskellia comata (Maskell) 1896 Neomaskellia bergii (Signoret) 1868 

Neomaskellia eucalypti Dumbleton 1956 Xenaleyrodes eucalypti (Dumbleton) 1956 

Neomaskellia hainanensis Chou & Yan 1988 Neomaskellia andropogonis Corbett 1926 

NEOPEALIUS TAKAHASHI 1954 NEOPEALIUS TAKAHASHI 1954 

Neopealius nilgiriensis David & Subramaniam 1976 Pealius azaleae (Baker & Moles) 1920 

Neopealius rubi Takahashi 1954 Neopealius rubi Takahashi 1954 

NIGRASIALEYRODES MARTIN & CARVER 1999 NIGRASIALEYRODES MARTIN & CARVER 1999 

Nigrasialeyrodes convexus Martin & Carver 1999 Nigrasialeyrodes convexus Martin & Carver 1999 

NIPALEYRODES TAKAHASHI 1951 NIPALEYRODES TAKAHASHI 1951 

Nipaleyrodes elongata Takahashi 1951 Nipaleyrodes elongata Takahashi 1951 

OCTALEURODICUS HEMPEL 1922 OCTALEURODICUS HEMPEL 1922 

Octaleurodicus nitidus Hempel 1922 Octaleurodicus nitidus Hempel 1922 

Octaleurodicus pulcherrimus (Quaintance & Baker) 1913 Octaleurodicus pulcherrimus (Quaintance & Baker) 1913 

Odontaleyrodes Takahashi 1954 Pealius Quaintance & Baker 1914 

Odontaleyrodes akebiae (Kuwana) 1911 Pealius akebiae (Kuwana) 1911 

Odontaleyrodes artocarpi David, Sundararaj & Regu 1991 Pealius nagerkoilensis Jesudasan & David 1991 

Odontaleyrodes damnacanthi (Takahashi) 1935 Pealius damnacanthi Takahashi 1935 

Odontaleyrodes elongatus David, Sundararaj & Regu 1991 Pealius elongatus (David, Sundararaj & Regu) 1991 

Odontaleyrodes euryae Takahashi 1955 Pealius euryae (Takahashi) 1955 

Odontaleyrodes indicus David 1972 Pealius indicus (David) 1972 

Odontaleyrodes kongosana Takahashi 1955 Pealius kongosana (Takahashi) 1955 

Odontaleyrodes mitakensis Takahashi 1955 Pealius mitakensis (Takahashi) 1955 

Odontaleyrodes nilgiriensis David 1972 Pealius nilgiriensis (David) 1972 

Odontaleyrodes rhododendri (Takahashi) 1935 Pealius rhododendri Takahashi 1935 

692

Odontaleyrodes sairandhryensis Meganathan & David 1994 Pealius sairandhryensis (Meganathan & David) 1994 

Odontaleyrodes selvakumarani David, Sundararaj & Regu 1991 Pealius nagerkoilensis Jesudasan & David 1991 

Odontaleyrodes splendens David, Sundararaj & Regu 1991 Pealius splendens (David, Sundararaj & Regu) 1991 

Ogivaleurodes Goux 1948 Trialeurodes Cockerell 1902 

Ogivaleurodes lauri (Signoret) 1882 Trialeurodes lauri (Signoret) 1882 

Orchamoplatus Russell 1958 Orchamoplatus Russell 1958 

Orchamoplatus caledonicus (Dumbleton) 1956 Orchamoplatus caledonicus (Dumbleton) 1956 

Orchamoplatus calophylli Russell 1958 Orchamoplatus calophylli Russell 1958 

Orchamoplatus citri (Takahashi) 1940 Orchamoplatus citri (Takahashi) 1940 

Orchamoplatus dentatus (Dumbleton) 1956 Orchamoplatus dentatus (Dumbleton) 1956 

Orchamoplatus dumbletoni (Cohic) 1959 Orchamoplatus dumbletoni (Cohic) 1959 

693 

Orchamoplatus incognitus (Dumbleton) 1956 Orchamoplatus incognitus (Dumbleton) 1956 

Orchamoplatus louiserussellae Martin 1999 Orchamoplatus louiserussellae Martin 1999 

Orchamoplatus mammaeferus (Quaintance & Baker) 1917 Orchamoplatus mammaeferus (Quaintance & Baker) 1917 

Orchamoplatus montanus (Dumbleton) 1956 Orchamoplatus montanus (Dumbleton) 1956 

Orchamoplatus niuginii Martin 1985 Orchamoplatus niuginii Martin 1985 

Orchamoplatus noumeae Russell 1958 Orchamoplatus noumeae Russell 1958 

Orchamoplatus perdentatus Dumbleton 1961 Orchamoplatus perdentatus Dumbleton 1961 

Orchamoplatus plumensis (Dumbleton) 1956 Orchamoplatus plumensis (Dumbleton) 1956 

Orchamoplatus porosus (Dumbleton) 1956 Orchamoplatus porosus (Dumbleton) 1956 

Orchamoplatus sudanensis Gameel 1968 Neoaleurotrachelus sudanensis (Gameel) 1968 

Orchamus Quaintance & Baker 1917 Orchamoplatus Russell 1958 

Orchamus caledonicus Dumbleton 1956 Orchamoplatus caledonicus (Dumbleton) 1956 

Orchamus dentatus Dumbleton 1956 Orchamoplatus dentatus (Dumbleton) 1956 

Orchamus dumbletoni Cohic 1959 Orchamoplatus dumbletoni (Cohic) 1959 

Orchamus incognitus Dumbleton 1956 Orchamoplatus incognitus (Dumbleton) 1956 

Orchamus montanus Dumbleton 1956 Orchamoplatus montanus (Dumbleton) 1956 

Orchamus plumensis Dumbleton 1956 Orchamoplatus plumensis (Dumbleton) 1956 

Orchamus porosus Dumbleton 1956 Orchamoplatus porosus (Dumbleton) 1956 

ORIENTALEYRODES DAVID 1993 ORIENTALEYRODES DAVID 1993 

Orientaleyrodes indicus David 1993 Orientaleyrodes indicus David 1993 

ORSTOMALEYRODES COHIC 1966 ORSTOMALEYRODES COHIC 1966 

Orstomaleyrodes fimbriae (Mound) Orstomaleyrodes fimbriae (Mound) 1965 

Orstomaleyrodes fimbriae (Mound) 1965 Orstomaleyrodes fimbriae (Mound) 1965 

PAPILLIPES BINK-MOENEN 1983 PAPILLIPES BINK-MOENEN 1983 

Papillipes spinifer Bink-Moenen 1983 Papillipes spinifer Bink-Moenen 1983 

PARABEMISIA TAKAHASHI 1952 PARABEMISIA TAKAHASHI 1952 

Parabemisia aceris (Takahashi) 1931 Parabemisia aceris (Takahashi) 1931 

Parabemisia indica Meganathan & David 1994 Parabemisia indica Meganathan & David 1994

Parabemisia jawani Martin 1985 Parabemisia jawani Martin 1985 

Parabemisia lushanensis Ko & Luo 1999 Parabemisia lushanensis Ko & Luo 1999 

Parabemisia maculata Takahashi 1952 Parabemisia maculata Takahashi 1952 

Parabemisia myricae (Kuwana) 1927 Parabemisia myricae (Kuwana) 1927 

Parabemisia myrmecophila Martin 1985 Parabemisia myrmecophila Martin 1985 

Parabemisia reticulata Dumbleton 1961 Parabemisia reticulata Dumbleton 1961 

Parabemisia sairandhryensis (Meganathan & David) 1994 Pealius sairandhryensis (Meganathan & David) 1994 

Paraleurolobus Sampson & Drews 1941 Paraleurolobus Sampson & Drews 1941 

Paraleurolobus chamaedoreae Russell 1994 Paraleurolobus chamaedoreae Russell 1994 

Paraleurolobus imbricatus Sampson & Drews 1941 Paraleurolobus imbricatus Sampson & Drews 1941 

Paraleyodes pseudonaranjae Martin 2001 Paraleyrodes pseudonaranjae Martin 2001 

PARALEYRODES QUAINTANCE 1909 PARALEYRODES QUAINTANCE 1909 

Paraleyrodes ancora Martin 2004 Paraleyrodes ancora Martin 2004 

Paraleyrodes bondari Peracchi 1971 Paraleyrodes bondari Peracchi 1971 

Paraleyrodes cervus Martin 2004 Paraleyrodes cervus Martin 2004 

Paraleyrodes citri Bondar 1931 Paraleyrodes citri Bondar 1931 

Paraleyrodes citricolus Costa Lima 1928 Paraleyrodes citricolus Costa Lima 1928 

Paraleyrodes crateraformans Bondar 1922 Paraleyrodes crateraformans Bondar 1922 

Paraleyrodes goyabae (Goldi) 1886 Paraleyrodes goyabae (Goldi) 1886 

Paraleyrodes minei Iaccarino 1990 Paraleyrodes minei Iaccarino 1990 

Paraleyrodes naranjae Dozier 1927 Paraleyrodes naranjae Dozier 1927 

Paraleyrodes perplexus Martin 2004 Paraleyrodes perplexus Martin 2004 

Paraleyrodes perseae (Quaintance) 1900 Paraleyrodes perseae (Quaintance) 1900 

Paraleyrodes proximus Teran 1979 Paraleyrodes proximus Teran 1979 

Paraleyrodes pseudonaranjae Martin 2001 Paraleyrodes pseudonaranjae Martin 2001 

Paraleyrodes pulverans Bondar 1923 Paraleyrodes pulverans Bondar 1923 

Paraleyrodes singularis Bondar 1923 Paraleyrodes singularis Bondar 1923 

Paraleyrodes triungulae Martin 2004 Paraleyrodes triungulae Martin 2004 

Paraleyrodes urichii Quaintance & Baker 1913 Paraleyrodes urichii Quaintance & Baker 1913 

Parudamoselis Visnya 1941 Ceraleurodicus Hempel 1922 

Parudamoselis kesselyaki Visnya 1941 Ceraleurodicus varus (Bondar) 1928 

PAULIANALEYRODES COHIC 1966 PAULIANALEYRODES COHIC 1966 

Paulianaleyrodes pauliani Cohic 1966 Paulianaleyrodes pauliani Cohic 1966 

Paulianaleyrodes splendens Cohic 1966 Paulianaleyrodes splendens Cohic 1966 

Paulianaleyrodes tetracerae Cohic 1966 Paulianaleyrodes tetracerae Cohic 1966 

PEALIUS QUAINTANCE & BAKER 1914 PEALIUS QUAINTANCE & BAKER 1914 

Pealius akebiae (Kuwana) 1911 Pealius akebiae (Kuwana) 1911 

Pealius amamianus Takahashi 1963 Pealius amamianus Takahashi 1963 

Pealius artocarpi (Corbett) 1935 Pealius artocarpi (Corbett) 1935 

694

Pealius azaleae (Baker & Moles) 1920 Pealius azaleae (Baker & Moles) 1920 

Pealius bangkokensis Takahashi 1942 Pealius bangkokensis Takahashi 1942 

Pealius bengalensis (Peal) 1903 Pealius bengalensis (Peal) 1903 

Pealius cambodiensis Takahashi 1942 Pealius cambodiensis Takahashi 1942 

Pealius celti Takahashi 1932 Apobemisia celti (Takahashi) 1932 

Pealius chinensis Takahashi 1941 Singhiella chinensis (Takahashi) 1941 

Pealius cryptus Martin 1999 Pealius cryptus Martin 1999 

Pealius damnacanthi Takahashi 1935 Pealius damnacanthi Takahashi 1935 

Pealius elatostemae (Takahashi) 1932 Pealius elatostemae (Takahashi) 1932 

Pealius elongatus (David, Sundararaj & Regu) 1991 Pealius elongatus (David, Sundararaj & Regu) 1991 

Pealius euryae (Takahashi) 1955 Pealius euryae (Takahashi) 1955 

Pealius ezeigwi Mound 1965 Pealius ezeigwi Mound 1965 

Pealius fici Mound 1965 Pealius fici Mound 1965 

Pealius hibisci (Kotinsky) 1907 Singhius hibisci (Kotinsky) 1907 

Pealius homonoiae Jesudasan & David 1991 Massilieurodes homonoiae (Jesudasan & David) 1991 

Pealius indicus (David) 1972 Pealius indicus (David) 1972 

Pealius indicus David & Subramaniam 1976 Singhiella simplex (Singh) 1931 

Pealius kalawi Singh 1933 Pealius kalawi Singh 1933 

Pealius kankoensis (Takahashi) Pealius kankoensis (Takahashi) 1933 

Pealius kankoensis (Takahashi) 1933 Pealius kankoensis (Takahashi) 1933 

Pealius kelloggi (Bemis) 1904 Pealius kelloggi (Bemis) 1904 

Pealius liquidambari (Takahashi) 1932 Pealius liquidambari (Takahashi) 1932 

Pealius longispinus Takahashi 1932 Pealius longispinus Takahashi 1932 

Pealius machili Takahashi 1935 Pealius machili Takahashi 1935 

Pealius maculatus Takahashi 1942 Pealius maculatus Takahashi 1942 

Pealius madeirensis Martin, Aguiar & Pita 1996 Pealius madeirensis Martin, Aguiar & Pita 1996 

Pealius maskelli (Bemis) 1904 Pealius maskelli (Bemis) 1904 

Pealius misrae Singh 1931 Pealius misrae Singh 1931 

Pealius mori (Takahashi) 1932 Pealius mori (Takahashi) 1932 

Pealius nagerkoilensis Jesudasan & David 1991 Fippataleyrodes nagerkoilensis Jesudasan & David 1991 

Pealius polygoni Takahashi 1934 Pealius polygoni Takahashi 1934 

Pealius psychotriae Takahashi 1935 Pealius psychotriae Takahashi 1935 

Pealius quercus (Signoret) 1868 Pealius quercus (Signoret) 1868 

Pealius rhododendri Takahashi 1935 Pealius rhododendri Takahashi 1935 

Pealius rubi Takahashi 1936 Pealius rubi Takahashi 1936 

Pealius sairandhryensis (Meganathan & David) 1994 Pealius sairandhryensis (Meganathan & David) 1994 

Pealius schimae Takahashi 1950 Pealius schimae Takahashi 1950 

Pealius setosus Danzig 1964 Pealius setosus Danzig 1964 

Pealius simplex (Singh) 1931 Singhiella simplex (Singh) 1931 

695

Pealius spinosus Jesudasan & David 1991 Pealius spinosus David & Jesudasan 1991 

Pealius spinus (Singh) 1931 Pealius spinus (Singh) 1931 

Pealius splendens (David, Sundararaj & Regu) 1991 Pealius splendens (David, Sundararaj & Regu) 1991 

Pealius sutepensis Takahashi 1942 Pealius sutepensis Takahashi 1942 

Pealius tuberculatus Takahashi 1942 Pealius tuberculatus Takahashi 1942 

Pealius walayarensis (Jesudasan & David) 1991 Pealius walayarensis (Jesudasan & David) 1991 

696 

PECTINALEYRODES BINK-MOENEN 1983 PECTINALEYRODES BINK-MOENEN 1983 

Pectinaleyrodes culcasiae (Cohic) 1969 Pectinaleyrodes culcasiae (Cohic) 1969 

Pectinaleyrodes silvaticus (Cohic) 1969 Pectinaleyrodes silvaticus (Cohic) 1969 

Pectinaleyrodes triclisiae (Cohic) 1966 Pectinaleyrodes triclisiae (Cohic) 1969 

Pentaleurodicus Bondar 1923 Aleuronudus Hempel 1923 

Pentaleurodicus bahiensis (Hempel) 1922 Metaleurodicus bahiensis (Hempel) 1922 

Pentaleurodicus induratus (Hempel) 1922 Aleuronudus induratus Hempel 1922 

PENTALEYRODES TAKAHASHI 1937 PENTALEYRODES TAKAHASHI 1937 

Pentaleyrodes cinnamomi (Takahashi) Pentaleyrodes cinnamomi (Takahashi) 1932 

Pentaleyrodes cinnamomi var. raishana Takahashi 1937 Pentaleyrodes cinnamomi (Takahashi) 1932 

Pentaleyrodes hongkongensis Takahashi 1941 Pentaleyrodes hongkongensis Takahashi 1941 

Pentaleyrodes linderae Chou & Yan 1988 Pentaleyrodes linderae Chou & Yan 1988 

Pentaleyrodes yasumatsui Takahashi 1939 Pentaleyrodes yasumatsui Takahashi 1939 

PERACCHIUS LIMA & RACCA-FILHO 2005 PERACCHIUS LIMA & RACCA-FILHO 2005 

Peracchius durantae Lima & Racca Filho 2005 Peracchius durantae Lima & Racca Filho 2005 

Permaleurodes rotundatum Becker Migdisova 1959 Permaleurodes? rotundatum? Becker Migdisova 1959 

Phalaena (Tinea) proletella Linnaeus 1758 Aleyrodes proletella (Linnaeus) 1758 

Phalaena culiciformis Geoffroy 1795 Aleyrodes proletella (Linnaeus) 1758 

PLATALEYRODES TAKAHASHI & MAMET 1952 PLATALEYRODES TAKAHASHI & MAMET 1952 

Plataleyrodes anthocleistae Takahashi & Mamet 1952 Plataleyrodes anthocleistae Takahashi & Mamet 1952 

POGONALEYRODES TAKAHASHI 1955 POGONALEYRODES TAKAHASHI 1955 

Pogonaleyrodes fastuosa Takahashi 1955 Pogonaleyrodes fastuosa Takahashi 1955 

Pogonaleyrodes zimmermanni (Newstead) 1911 Pogonaleyrodes zimmermanni (Newstead) 1911 

Powellia Maskell 1879 Trioza pellucida Maskell [Psyllidae] 

Pseudaleurodicus Hempel 1922 Aleuronudus Hempel 1922 

Pseudaleurodicus acapulcensis Sampson & Drews 1941 Aleuronudus acapulcensis (Sampson & Drews) 1941 

Pseudaleurodicus bahiensis Hempel 1922 Metaleurodicus bahiensis (Hempel) 1922 

PSEUDALEUROLOBUS HEMPEL 1923 PSEUDALEUROLOBUS HEMPEL 1923 

Pseudaleurolobus dorsidemarcata (Singh) 1932 Asialeyrodes dorsidemarcata (Singh) 1932 

Pseudaleurolobus jaboticabae Hempel 1923 Pseudaleurolobus jaboticabae Hempel 1923 

Pseudaleurolobus maesae Takahashi 1934 Asialeyrodes maesae (Takahashi) 1934 

PSEUDALEUROPLATUS MARTIN 1999 PSEUDALEUROPLATUS MARTIN 1999 

Pseudaleuroplatus kiensis Martin 1999 Pseudaleuroplatus kiensis Martin 1999

Pseudaleuroplatus litseae (Dumbleton) 1956 Pseudaleuroplatus litseae (Dumbleton) 1956 

PSEUDALEYRODES HEMPEL 1923 PSEUDALEYRODES HEMPEL 1923 

Pseudaleyrodes depressus Hempel 1923 Pseudaleyrodes depressus Hempel 1923 

PSEUDOSYNALEURODICUS GILLESPIE 2006 PSEUDOSYNALEURODICUS GILLESPIE 2006 

Pseudosynaleurodicus mayoi Gillespie 2006 Pseudosynaleurodicus mayoi Gillespie 2006 

Pseudosynaleurodicus nigrimarginatus Gillespie 2006 Pseudosynaleurodicus nigrimarginatus Gillespie 2006 

PSEUDOZAPHANERA MANZARI 2006 PSEUDOZAPHANERA MANZARI 2006 

Pseudozaphanera niger (Maskell) 1895 Pseudozaphanera niger (Maskell) 1895 

Pseudozaphanera papyrocarpae (Martin) 2001 Pseudozaphanera papyrocarpae (Martin) 2001 

Pseudozaphanera rhachisreticulata (Martin) 1999 Pseudozaphanera rhachisreticulata (Martin) 1999 

Pseudozaphanera splendida (Martin) 1999 Pseudozaphanera splendida (Martin) 1999 

Pseudozaphanera wariensis (Martin) 1999 Pseudozaphanera wariensis (Martin) 1999 

Quaintancius BONDAR 1922 Octaleurodicus HEMPEL 1922 

Quaintancius pulcherrimus (Quaintance & Baker) 1913 Octaleurodicus pulcherrimus (Quaintance & Baker) 1913 

Quaintancius rubrus Bondar 1922 Octaleurodicus nitidus Hempel 1922 

RABDOSTIGMA QUAINTANCE & BAKER 1917 RABDOSTIGMA QUAINTANCE & BAKER 1917 

Rabdostigma mahableshwarensis David & Sundararaj 1998 Rabdostigma mahableshwarensis David & Sundararaj 1998 

Rabdostigma mahableshwarensis Sundararaj and David 1998 Rabdostigma mahableshwarensis Sundararaj and David 1998 

Rabdostigma minahassai (Martin) 1988 Rabdostigma minahassai (Martin) 1989 

Rabdostigma radiilinealis (Quaintance & Baker) 1917 Rabdostigma radiilinealis (Quaintance & Baker) 1917 

Radialeurodicus Bondar 1922 Ceraleurodicus Hempel 1922 

Radialeurodicus altissimus (Quaintance) 1900 Nealeurodicus altissimus (Quaintance) 1900 

Radialeurodicus altissimus ingae Baker 1937 Nealeurodicus ingae (Baker) 1937 

Radialeurodicus assymmetricus Bondar 1922 Ceraleurodicus assymmetricus (Bondar) 1922 

Radialeurodicus bakeri Bondar 1923 Nealeurodicus bakeri (Bondar) 1923 

Radialeurodicus cinereus Bondar 1922 Ceraleurodicus spendidus Hempel 1922 

Radialeurodicus melzeri Laing 1930 Nealeurodicus paulistus Hempel 1922 

Radialeurodicus neivai Bondar 1928 Ceraleurodicus neivai (Bondar) 1928 

Radialeurodicus octifer Bondar 1923 Nealeurodicus octifer (Bondar) 1923 

Radialeurodicus varus Bondar 1928 Ceraleurodicus varus (Bondar) 1928 

RAMSESSEUS ZAHRADNIK 1970 RAMSESSEUS ZAHRADNIK 1970 

Ramsesseus follioti Zahradnik 1970 Ramsesseus follioti Zahradnik 1970 

RHACHISPHORA QUAINTANCE & BAKER 1917 RHACHISPHORA QUAINTANCE & BAKER 1917 

Rhachisphora alishanensis Ko 1992 Rhachisphora alishanensis Ko 1992 

Rhachisphora ardisiae (Takahashi) 1935 Rhachisphora ardisiae (Takahashi) 1935 

Rhachisphora capitatis (Corbett) 1926 Rhachisphora capitatis (Corbett) 1926 

Rhachisphora elongatus Regu & David 1990 Rhachisphora elongatus Regu & David 1990 

Rhachisphora fici (Takahashi) 1932 Massilieurodes fici (Takahashi) 1932 

Rhachisphora franksae Martin 1999 Rhachisphora franksae Martin 1999 

Rhachisphora indica Sundararaj & David 1991 Rhachisphora indica Sundararaj & David 1991 

697

Rhachisphora ixorae Sundararaj & David 1991 Rhachisphora ixorae Sundararaj & David 1991 

Rhachisphora kallarensis Jesudasan & David 1991 Rhachisphora kallarensis Jesudasan & David 1991 

Rhachisphora koshunensis (Takahashi) 1933 Rhachisphora koshunensis (Takahashi) 1933 

Rhachisphora kuraruensis (Takahashi) 1933 Singhiella kuraruensis (Takahashi) 1933 

Rhachisphora machili (Takahashi) 1932 Rhachisphora machili (Takahashi) 1932 

Rhachisphora madhucae Jesudasan & David 1991 Rhachisphora trilobitoides (Quaintance & Baker) 1917 

Rhachisphora maesae (Takahashi) 1932 Rhachisphora maesae (Takahashi) 1932 

Rhachisphora malayensis Takahashi 1952 Rhachisphora malayensis (Takahashi) 1952 

Rhachisphora oblongata Ko 1988 Rhachisphora oblongata Ko 1988 

Rhachisphora queenslandica Martin 1999 Rhachisphora queenslandica Martin 1999 

Rhachisphora reticulata (Takahashi) 1933 Rhachisphora reticulata (Takahashi) 1933 

Rhachisphora rutherfordi (Quaintance & Baker) 1917 Rhachisphora rutherfordi (Quaintance & Baker) 1917 

Rhachisphora sanhsianensis Ko 1992 Rhachisphora sanhsianensis Ko 1992 

Rhachisphora selangorensis (Corbett) 1933 Rhachisphora selangorensis (Corbett) 1933 

Rhachisphora setulosa (Corbett) 1926 Rhachisphora setulosa (Corbett) 1926 

Rhachisphora spherica Sundararaj & Dubey 2006 Asialeyrodes spherica (Sundararaj & Dubey) 2006 

Rhachisphora styraci (Takahashi) 1934 Rhachisphora styraci (Takahashi) 1934 

Rhachisphora taiwana Ko 1992 Rhachisphora taiwana Ko 1992 

Rhachisphora trilobitoides (Quaintance & Baker) 1917 Rhachisphora trilobitoides (Quaintance & Baker) 1917 

ROSANOVIA DANZIG 1969 ROSANOVIA DANZIG 1969 

Rosanovia hulthemiae Danzig 1969 Rosanovia hulthemiae Danzig 1969 

Rositaleyrodes Meganathan & David 1994 Aleurobus Quaintance & Baker 

Rositaleyrodes oplismeni (Takahashi) 1931 Aleurolobus oplismeni Takahashi 1931 

Roucasia Goux 1940 Bemisia Quaintance & Baker 1914 

Roucasia ovata Goux 1940 Bemisia ovata (Goux) 1940 

RUGALEYRODES BINK-MOENEN 1983 RUGALEYRODES BINK-MOENEN 1983 

Rugaleyrodes angolensis (Cohic) 1966 Rugaleyrodes angolensis (Cohic) 1966 

Rugaleyrodes bidentata Bink-Moenen 1983 Rugaleyrodes bidentata Bink-Moenen 1983 

Rugaleyrodes tetracerae (Cohic) 1966 Rugaleyrodes tetracerae (Cohic) 1966 

Rugaleyrodes villiersi (Cohic) 1968 Rugaleyrodes villiersi (Cohic) 1968 

Rugaleyrodes vuattouxi (Cohic) 1969 Rugaleyrodes vuattouxi (Cohic) 1969 

RUSOSTIGMA QUAINTANCE & BAKER 1917 RUSOSTIGMA QUAINTANCE & BAKER 1917 

Rusostigma eugeniae (Maskell) Rusostigma eugeniae (Maskell) 1895 

Rusostigma eugeniae (Maskell) 1895 Rusostigma eugeniae (Maskell) 1895 

Rusostigma radiiruigosa (Quaintance & Baker) 1917 Rusostigma radiiruigosa Quaintance & Baker 1917 

Rusostigma tokyonis (Kuwana) 1911 Rusostigma tokyonis (Kuwana) 1911 

Rusostigma tristylii (Takahashi) 1935 Rusostigma tristylii (Takahashi) 1935 

RUSSELLALEYRODES DAVID 1973 RUSSELLALEYRODES DAVID 1973 

Russellaleyrodes cumiugum (Singh) 1932 Russellaleyrodes cumiugum (Singh) 1932 

698

Ryoichiella Jesudasan & David 1990 Dumbletoniella Jesudasan & David 1990 

Ryoichiella ellipticae (Dumbleton) 1956 Dumbletoniella ellipticae (Dumbleton) 1956 

SEPTALEURODICUS SAMPSON 1943 SEPTALEURODICUS SAMPSON 1943 

Septaleurodicus mexicanus Sampson 1943 Septaleurodicus mexicanus Sampson 1943 

SETALEYRODES TAKAHASHI 1931 SETALEYRODES TAKAHASHI 1931 

699 

Setaleyrodes litseae David & Sundararaj 1991 Setaleyrodes litseae David & Sundararaj 1991 

Setaleyrodes mirabilis Takahashi 1931 Setaleyrodes mirabilis Takahashi 1931 

Setaleyrodes quercicola Takahashi 1934 Setaleyrodes quercicola Takahashi 1934 

Setaleyrodes takahashia Singh 1933 Setaleyrodes takahashia Singh 1933 

Setaleyrodes thretaonai David 1981 Setaleyrodes thretaonai David 1981 

Setaleyrodes vigintiseta Martin 1999 Setaleyrodes vigintiseta Martin 1999 

Shanthiniae David 2000 Dialeurodes Cockerell 

Shanthiniae sheryli David 2000 Dialeurodes sheryli (David) 2000 

SIMPLALEURODES GOUX 1945 SIMPLALEURODES GOUX 1945 

Simplaleurodes hemisphaerica Goux 1945 Simplaleurodes hemisphaerica Goux 1945 

SINGHIELLA SAMPSON 1943 SINGHIELLA SAMPSON 1943 

Singhiella bassiae (David and Subramaniam) 1976 Singhiella bassiae (David and Subramaniam) 1976 

Singhiella bauhiniae Dubey & Sundararaj 2006 Singhiella bauhiniae Dubey & Sundararaj 2006 

Singhiella bicolor (Singh) 1931 Singhiella bicolor (Singh) 1931 

Singhiella brideliae (Jesudasan & David) 1991 Singhiella brideliae (Jesudasan and David) 1991 

Singhiella cambodiensis (Takahashi) 1942 Singhiella cambodiensis (Takahashi) 1942 

Singhiella cardamomi (David and Subramaniam) 1976 Singhiella cardamomi (David and Subramaniam) 1976 

Singhiella chinensis (Takahashi) 1941 Singhiella chinensis (Takahashi) 1941 

Singhiella chitinosa (Takahashi) 1937 Singhiella chitinosa (Takahashi) 1937 

Singhiella citrifolii (Morgan) 1893 Singhiella citrifolii (Morgan) 1893 

Singhiella crenulata Qureshi & Qayyum 1969 Singhiella crenulata Qureshi & Qayyum 1969 

Singhiella delamarei (Cohic) 1968 Singhiella delamarei (Cohic) 1968 

Singhiella denticulata Qureshi 1979 Singhiella denticulata Qureshi 1979 

Singhiella dioscoreae (Takahashi) 1934 Singhiella dioscoreae (Takahashi) 1934 

Singhiella dipterocarpi (Takahashi) 1942 Singhiella dipterocarpi (Takahashi) 1942 

Singhiella elaeagni (Takahashi) 1935 Singhiella elaeagni (Takahashi) 1935 

Singhiella elbaensis (Priesner and Hosny) 1934 Singhiella elbaensis (Priesner and Hosny) 1934 

Singhiella ficifolii (Takahashi) 1942 Singhiella ficifolii (Takahashi) 1942 

Singhiella kuraruensis (Takahashi) 1933 Singhiella kuraruensis (Takahashi) 1933 

Singhiella longisetae Chou & Yan 1988 Singhiella longisetae Chou & Yan 1988 

Singhiella malabaricus (Jesudasan & David) 1991 Singhiella malabaricus (Jesudasan and David) 1991 

Singhiella mekonensis (Takahashi) 1942 Singhiella mekonensis (Takahashi) 1942 

Singhiella melanolepis Young & Ko 2007 Singhiella melanolepis Young & Ko 2007 

Singhiella pallida (Singh) 1931 Singhiella pallida (Singh) 1931

Singhiella piperis (Takahashi) 1934 Singhiella piperis (Takahashi) 1934 

Singhiella premnae Martin 1999 Singhiella premnae Martin 1999 

Singhiella serdangensis (Corbett) 1935 Singhiella serdangensis (Corbett) 1935 

Singhiella simplex (Singh) 1931 Singhiella simplex (Singh) 1931 

Singhiella subrotunda (Takahashi) 1935 Singhiella subrotunda (Takahashi) 1935 

Singhiella sutepensis (Takahashi) 1942 Singhiella sutepensis (Takahashi) 1942 

Singhiella tetrastigmae (Takahashi) Singhiella tetrastigmae (Takahashi) 1934 

Singhiella vanieriae (Takahashi) Singhiella vanieriae (Takahashi) 1935 

SINGHIUS TAKAHASHI 1932 SINGHIUS TAKAHASHI 1932 

Singhius hibisci (Kotinsky) 1907 Singhius hibisci (Kotinsky) 1907 

Singhius russellae (David & Subramaniam) 1976 Aleuroclava russellae (David & Subramaniam) 1976 

Siphonaleyrodes formosanus Takahashi 1932 Trioza cinnamomi Boselli 1930 [Psyllidae] 

SIPHONINUS SILVESTRI 1915 SIPHONINUS SILVESTRI 1915 

Siphoninus blanzyi Cohic 1968 Cohicaleyrodes blanzyi (Cohic) 1968 

Siphoninus dubiosa Haupt 1935 Siphoninus phillyreae (Haliday) 1835 

Siphoninus finitimus Silvestri 1915 Siphoninus phillyreae (Haliday) 1835 

Siphoninus granati Priesner & Hosny 1932 Siphoninus phillyreae (Haliday) 1835 

Siphoninus gruveli Cohic 1968 Siphoninus gruveli Cohic 1968 

Siphoninus heegeri Haupt 1935 Siphoninus immaculatus (Heeger) 1856 

Siphoninus immaculatus (Heeger) 1856 Siphoninus immaculatus (Heeger) 1856 

Siphoninus phillyreae (Haliday) 1835 Siphoninus phillyreae (Haliday) 1835 

Siphoninus phillyreae inequalis Goux 1949 Siphoninus phillyreae (Haliday) 1835 

Siphoninus phillyreae mulititubulatus Goux 1949 Siphoninus phillyreae (Haliday) 1835 

SPHERICALEYRODES SELVAKUMARAN AND DAVID SPHERICALEYRODES SELVAKUMARAN AND DAVID 

Sphericaleyrodes bambusae Selvakumaran and David Sphericaleyrodes bambusae Selvakumaran and David 

Sphericaleyrodes regui Dubey & Sundararaj Sphericaleyrodes regui Dubey & Sundararaj 

STENALEYRODES TAKAHASHI 1938 STENALEYRODES TAKAHASHI 1938 

Stenaleyrodes elongatus (Dumbleton) 1956 Stenaleyrodes vinsoni Takahashi 1938 

700 

Stenaleyrodes papillote Martin & Streito 2003 Stenaleyrodes papillote Martin & Streito 2003 

Stenaleyrodes vinsoni Takahashi 1938 Stenaleyrodes vinsoni Takahashi 1938 

SYNALEURODICUS SOLOMON 1935 SYNALEURODICUS SOLOMON 1935 

Synaleurodicus hakeae Solomon 1935 Synaleurodicus hakeae Solomon 1935 

Synaleurodicus serratus Martin 1999 Synaleurodicus serratus Martin 1999 

TAIWANALEYRODES TAKAHASHI 1932 ALEUROCLAVA SINGH 1931 

Taiwanaleyrodes baccaureae Corbett 1935 Aleuroclava baccaureae (Corbett) 1935 

Taiwanaleyrodes carpini Takahashi 1939 Aleuroclava carpini (Takahashi) 1939 

Taiwanaleyrodes carpini var. yushuniae Takahashi 1939 Aleuroclava carpini (Takahashi) 1939 

Taiwanaleyrodes fici Corbett 1935 Aleuroclava fici (Corbett)1935 

Taiwanaleyrodes fletcheri Sundararaj & David 1992 Aleuroclava fletcheri (Sundararaj & David) 1992

Taiwanaleyrodes hexcantha Singh 1940 Aleuroclava hexcantha (Singh) 1940 

Taiwanaleyrodes indica (Singh) 1931 Aleuroclava indica (Singh) 1931 

Taiwanaleyrodes macarangae Corbett 1935 Aleuroclava sepangensis Martin & Mound 2007 

Taiwanaleyrodes meliosmae Takahashi 1932 Aleuroclava meliosmae (Takahashi) 1932 

Taiwanaleyrodes montanus Takahashi 1939 Aleuroclava montanus (Takahashi)1939 

Taiwanaleyrodes nitidus (Singh) 1932 Aleuroclava nitidus (Singh) 1932 

Taiwanaleyrodes srilankaensis David 1993 Aleuroclava srilankaensis (David) 1993 

Taiwanaleyrodes tripori Dubey & Sundararaj 2006 Aleuroclava tripori (Dubey & Sundararaj) 2006 

TEGMALEURODES MARTIN 2005 TEGMALEURODES MARTIN 2005 

Tegmaleurodes lentus Martin 2005 Tegmaleurodes lentus Martin 2005 

Tetralecia granulata Sampson & Drews 1941 Aleuropleurocelus granulata (Drews & Sampson) 1941 

TETRALEURODES COCKERELL 1902 TETRALEURODES COCKERELL 1902 

Tetraleurodes abnormis (Quaintance) 1900 Aleuropleurocelus abnormis (Quaintance) 1900 

Tetraleurodes acaciae (Quaintance) 1900 Tetraleurodes acaciae (Quaintance) 1900 

Tetraleurodes adabicola Takahashi 1955 Tetraleurodes adabicola Takahashi 1955 

Tetraleurodes andropogoni (Dozier) 1934 Tetraleurodes andropogoni (Dozier) 1934 

Tetraleurodes asparagi (Lewis) 1893 Aleurotrachelus asparagi (Lewis) 1893 

Tetraleurodes aucubae (Kuwana) 1911 Aleuroclava aucubae (Kuwana) 1911 

Tetraleurodes bambusae Jesudasan & David 1991 Tetraleurodes bambusae Jesudasan & David 1991 

Tetraleurodes banksi Martin 1999 Tetraleurodes banksi Martin 1999 

Tetraleurodes bararakae Takahashi 1955 Tetraleurodes bararakae Takahashi 1955 

Tetraleurodes bicolor Bink-Moenen 1992 Tetraleurodes bicolor Bink-Moenen 1992 

Tetraleurodes bidentatus Sampson & Drews 1941 Tetraleurodes bidentatus Sampson & Drews 1941 

Tetraleurodes bireflexa Nakahara 1995 Tetraleurodes bireflexa Nakahara 1995 

Tetraleurodes burliarensis Jesudasan & David 1991 Tetraleurodes burliarensis Jesudasan & David 1991 

Tetraleurodes cacaorum (Bondar) 1923 Tetraleurodes cacaorum (Bondar) 1923 

Tetraleurodes caulicola Nakahara 1995 Tetraleurodes caulicola Nakahara 1995 

Tetraleurodes chivela Nakahara 1995 Tetraleurodes chivela Nakahara 1995 

Tetraleurodes citriculus Dozier Chopra 1928 nomen nudum 

Tetraleurodes confusa Nakahara 1995 Tetraleurodes confusa Nakahara 1995 

Tetraleurodes contigua (Sampson & Drews) 1941 Tetraleurodes contigua (Sampson & Drews) 1941 

Tetraleurodes corni (Haldeman) 1850 Tetraleurodes corni (Haldeman) 1850 

Tetraleurodes croceata (Maskell) 1895 Aleuroduplidens croceata (Maskell) 1895 

Tetraleurodes cruzi (Cassino) 1999 Tetraleurodes cruzi (Cassino) 1999 

Tetraleurodes dendrocalamae Dubey & Sundararaj 2005 Tetraleurodes dendrocalamae Dubey & Sundararaj 2005 

Tetraleurodes dorseyi (Kotinsky) 1907 Tetraleurodes mori (Quaintance) 1899 

Tetraleurodes dorsirugosa Nakahara 1995 Tetraleurodes dorsirugosa Nakahara 1995 

Tetraleurodes elaeocarpi Takahashi 1950 Tetraleurodes elaeocarpi Takahashi 1950 

Tetraleurodes errans (Bemis) 1904 Tetraleurodes mori (Quaintance) 1899 

701

702 

Tetraleurodes fenestrata Takahashi 1955 Marginaleyrodes fenestrata (Takahashi) 1955 

Tetraleurodes fici Quaintance & Baker 1937 Tetraleurodes fici Quaintance & Baker 1937 

Tetraleurodes fici Takahashi 1955 Tetraleurodes malayensis Mound & Halsey 1978 

Tetraleurodes ghesquierei Dozier 1934 Tetraleurodes ghesquierei Dozier 1934 

Tetraleurodes graminis Takahashi 1934 Tetraleurodes graminis Takahashi 1934 

Tetraleurodes granulata Bink-Moenen 1983 Tetraleurodes granulata Bink-Moenen 1983 

Tetraleurodes hederae Goux 1939 Tetraleurodes hederae Goux 1939 

Tetraleurodes herberti (Penny) 1922 Tetraleurodes acaciae (Quaintance) 1900 

Tetraleurodes herberti Penny 1922 Tetraleurodes acaciae (Quaintance) 1900 

Tetraleurodes hirsuta Takahashi 1955 Tetraleurodes hirsuta Takahashi 1955 

Tetraleurodes kunnathoorensis Regu & David 1993 Tetraleurodes kunnathoorensis Regu & David 1993 

Tetraleurodes leguminicola Bink-Moenen 1983 Tetraleurodes leguminicola Bink-Moenen 1983 

Tetraleurodes litseae Dumbleton 1956 Pseudaleuroplatus litseae (Dumbleton) 1956 

Tetraleurodes madagascariensis Takahashi 1951 Tetraleurodes madagascariensis Takahashi 1951 

Tetraleurodes malayensis Mound & Halsey 1978 Tetraleurodes malayensis Mound & Halsey 1978 

Tetraleurodes mameti Takahashi 1938 Tetraleurodes mameti Takahashi 1938 

Tetraleurodes marginata (Newstead) 1911 Aleurotrachelus marginatus (Newstead) 1911 

Tetraleurodes marshalli Bondar 1928 Tetraleurodes marshalli Bondar 1928 

Tetraleurodes melanops (Cockerell) 1902 Tetraleurodes melanops (Cockerell) 1902 

Tetraleurodes mexicana Nakahara 1995 Tetraleurodes mexicana Nakahara 1995 

Tetraleurodes mirabilis Takahashi 1961 Tetraleurodes mirabilis Takahashi 1961 

Tetraleurodes mori (Quaintance) 1899 Tetraleurodes mori (Quaintance) 1899 

Tetraleurodes mori arizonensis (Cockerell) 1900 Tetraleurodes mori (Quaintance) 1899 

Tetraleurodes mori var. maculata (Morrill) 1903 Tetraleurodes mori (Quaintance) 1899 

Tetraleurodes moundi Cohic 1968 Tetraleurodes moundi Cohic 1968 

Tetraleurodes neemani Bink-Moenen 1992 Tetraleurodes neemani Bink-Moenen 1992 

Tetraleurodes niger (Maskell) Zaphanera niger (Maskell) 1895 

Tetraleurodes niger (Maskell) 1895 Zaphanera niger (Maskell) 1895 

Tetraleurodes nigrans (Bemis) 1904 Aleuropleurocelus nigrans (Bemis) 1904 

Tetraleurodes nudus Sampson & Drews 1941 Tetraleurodes fici Quaintance & Baker 1937 

Tetraleurodes opilsmeni Takahashi 1934 Tetraleurodes opilsmeni Takahashi 1934 

Tetraleurodes palmae Gameel 1968 Fascaleyrodes palmae (Gameel) 1968 

Tetraleurodes papilliferus Sampson & Drews 1941 Aleurotrachelus papilliferus (Sampson & Drews) 1941 

Tetraleurodes pauliani Takahashi 1955 Tetraleurodes pauliani Takahashi 1955 

Tetraleurodes perileuca (Cockerell) 1902 Tetraleurodes perileuca (Cockerell) 1902 

Tetraleurodes perseae Nakahara 1995 Tetraleurodes perseae Nakahara 1995 

Tetraleurodes plumosa (Quaintance) 1900 Aleuroplatus plumosus (Quaintance) 1900 

Tetraleurodes pluto Dumbleton 1956 Tetraleurodes pluto Dumbleton 1956 

Tetraleurodes pringlei Quaintance & Baker 1937 Tetraleurodes pringlei Quaintance & Baker 1937

Tetraleurodes pseudacaciae Nakahara 1995 Tetraleurodes pseudacaciae Nakahara 1995 

Tetraleurodes psidii David 1993 Tetraleurodes psidii David 1993 

Tetraleurodes pusana Takahashi 1950 Tetraleurodes pusana Takahashi 1950 

Tetraleurodes quadratus Sampson & Drews 1941 Tetraleurodes quadratus Sampson & Drews 1941 

Tetraleurodes quercicola Nakahara 1995 Tetraleurodes quercicola Nakahara 1995 

Tetraleurodes rugosus Corbett 1926 Tetraleurodes rugosus Corbett 1926 

Tetraleurodes russellae Cohic 1968 Tetraleurodes russellae Cohic 1968 

Tetraleurodes selachidentata Bink-Moenen 1983 Tetraleurodes selachidentata Bink-Moenen 1983 

Tetraleurodes semibarbata Takahashi 1955 Tetraleurodes semibarbata Takahashi 1955 

703 

Tetraleurodes semilunaris Corbett 1926 Crescentaleyrodes semilunaris (Corbett) 1926 

Tetraleurodes simplicior Bink-Moenen 1983 Tetraleurodes simplicior Bink-Moenen 1983 

Tetraleurodes splendens (Bemis) 1904 Tetraleurodes mori (Quaintance) 1899 

Tetraleurodes stanfordi (Bemis) 1904 Tetraleurodes perileuca (Cockerell) 1902 

Tetraleurodes stellata (Maskell) 1895 Tetraleurodes stellata (Maskell) 1895 

Tetraleurodes stirlingiae Martin 1999 Tetraleurodes stirlingiae Martin 1999 

Tetraleurodes stypheliae (Maskell) Aleurotrachelus stypheliae (Maskell) 1895 

Tetraleurodes submarginata Dumbleton 1961 Tetraleurodes submarginata Dumbleton 1961 

Tetraleurodes subrotunda Takahashi 1937 Tetraleurodes subrotunda Takahashi 1937 

Tetraleurodes sulcistriatus Martin 1999 Tetraleurodes sulcistriatus Martin 1999 

Tetraleurodes supraxialis Martin 1999 Tetraleurodes supraxialis Martin 1999 

Tetraleurodes thenmozhiae Jesudasan & David 1991 Tetraleurodes thenmozhiae Jesudasan & David 1991 

Tetraleurodes truncatus Sampson & Drews 1941 Tetraleurodes truncatus Sampson & Drews 1941 

Tetraleurodes tuberculata Bink-Moenen 1983 Tetraleurodes tuberculata Bink-Moenen 1983 

Tetraleurodes tsinjoarivona Takahashi 1955 Marginaleyrodes tsinjoarivona (Takahashi) 1955 

Tetraleurodes tuberculosa Nakahara 1995 Tetraleurodes tuberculosa Nakahara 1995 

Tetraleurodes ursorum (Cockerell) 1910 Tetraleurodes ursorum (Cockerell) 1910 

TETRALICIA HARRISON 1917 TETRALICIA HARRISON 1917 

Tetralicia ceanothi Sampson 1945 Aleuropleurocelus ceanothi (Sampson) 1945 

Tetralicia debarroi Martin & Carver 1999 Tetralicia debarroi Martin & Carver 1999 

Tetralicia erianthi Danzig 1969 Tetralicia erianthi Danzig 1969 

Tetralicia ericae Harrison 1917 Tetralicia ericae Harrison 1917 

Tetralicia granulata Sampson & Drews 1941 Aleuropleurocelus granulata (Sampson and Drews) 1941 

Tetralicia graminicola Bink-Moenen 1983 Tetralicia graminicola Bink-Moenen 1983 

Tetralicia iberiaca Bink-Moenen 1989 Tetralicia iberiaca Bink-Moenen 1989 

Tetralicia nigrans (Bemis) 1904 Aleuropleurocelus nigrans (Bemis) 1904 

Tetralicia rotunda J. M. Baker 1937 Aleuropleurocelus rotunda (J. M. Baker) 1937 

Tetralicia sierrae Sampson 1945 Aleuropleurocelus sierrae (Sampson) 1945 

Tetralicia tuberculata Bink-Moenen 1983 Tetralicia tuberculata Bink-Moenen 1983 

Trialeurodes (Ericaleyrodes) ericae Bink-Moenen 1976 Trialeurodes (Ericaleyrodes) ericae Bink-Moenen 1976

Trialeurodes (Ericaleyrodes) sardiniae Rapisarda 1986 Trialeurodes (Ericaleyrodes) sardiniae Rapisarda 1986 

TRIALEURODES COCKERELL 1902 TRIALEURODES COCKERELL 1902 

Trialeurodes [sic] cotesi (Maskell) 1895 Bulgarialeurodes cotesii (Maskell) 1895 

Trialeurodes abdita Martin 2005 Trialeurodes abdita Martin 2005 

Trialeurodes abutiloneus (Haldeman) 1850 Trialeurodes abutiloneus (Haldeman) 1850 

Trialeurodes ambrosiae (Cockerell) 1910 Trialeurodes abutiloneus (Haldeman) 1850 

Trialeurodes amealcensis Carapia 2003 Trialeurodes amealcensis Cararapia 2003 

Trialeurodes asplenii (Maskell) 1890 Trialeurodes asplenii (Maskell) 1890 

Trialeurodes bambusae Takahashi 1943 Trialeurodes bambusae Takahashi 1943 

Trialeurodes bambusicola Takahashi 1951 Dialeurolonga bambusicola Takahashi 1951 

Trialeurodes bauhiniae Corbett 1935 Aleuromarginatus bauhiniae (Corbett) 1935 

Trialeurodes bellissima (Sampson & Drews) 1940 Trialeurodes bellissima (Sampson & Drews) 1940 

Trialeurodes bemisae Russell 1948 Trialeurodes bemisiae Russell 1948 

Trialeurodes bicolor Singh 1931 Singhiella bicolor (Singh) 1931 

Trialeurodes breyniae Singh 1931 Lipaleyrodes breyniae (Singh) 1931 

Trialeurodes bruneiensis Martin 2001 Trialeurodes bruneiensis Martin 2001 

Trialeurodes californicus Sampson 1945 Trialeurodes madroni (Bemis) 1904 

Trialeurodes caricae Corbett 1935 Trialeurodes variabilis (Quaintance) 1900 

Trialeurodes celti Takahashi 1943 Trialeurodes celti Takahashi 1943 

Trialeurodes chinensis Takahashi 1955 Trialeurodes chinensis Takahashi 1955 

Trialeurodes coccolobae Russell 1948 Trialeurodes coccolobae Russell 1948 

Trialeurodes colcordae Russell 1948 Trialeurodes colcordae Russell 1948 

Trialeurodes corollis (Penny) 1922 Trialeurodes corollis (Penny) 1922 

Trialeurodes cryptus Martin 2005 Trialeurodes cryptus Martin 2005 

Trialeurodes darwiniensis Martin 1999 Trialeurodes darwiniensis Martin 1999 

Trialeurodes desmodii Corbett 1935 Trialeurodes ricini (Misra) 1924 

Trialeurodes diasemus (Bemis) Aleyrodes diasemus Bemis 1904 

Trialeurodes dicksoniae Martin 1999 Trialeurodes dicksoniae Martin 1999 

Trialeurodes drewsi Sampson 1945 Trialeurodes drewsi Sampson 1945 

Trialeurodes dubienus (Bondar) 1923 Disiphon dubienus (Bondar) 1923 

Trialeurodes elaphoglossi Takahashi 1960 Trialeurodes elaphoglossi Takahashi 1960 

Trialeurodes elatostemae Takahashi 1932 Pealius elatostemae (Takahashi) 1932 

Trialeurodes ericae Bink-Moenen 1976 Trialeurodes ericae Bink-Moenen 1976 

Trialeurodes erigerontis (Maskell) 1895 Trialeurodes abutiloneus (Haldeman) 1850 

Trialeurodes eriodictyonis Russell 1948 Trialeurodes eriodictyonis Russell 1948 

Trialeurodes euphorbiae Russell 1948 Trialeurodes euphorbiae Russell 1948 

Trialeurodes fernaldi (Morrill) 1903 Trialeurodes fernaldi (Morrill) 1903 

Trialeurodes floridensis (Quaintance) 1900 Trialeurodes floridensis (Quaintance) 1900 

Trialeurodes glacialis (Bemis) 1904 Trialeurodes glacialis (Bemis) 1904 

704

Trialeurodes graminis Takahashi 1951 Dialeurolonga graminis (Takahashi) 1951 

Trialeurodes hargreavesi Corbett 1935 Lipaleyrodes hargreavesi (Corbett) 1935 

Trialeurodes heucherae Russell 1948 Trialeurodes heucherae Russell 1948 

Trialeurodes hutchingsi (Bemis) 1904 Trialeurodes hutchingsi (Bemis) 1904 

Trialeurodes immaculatus (Heeger) 1856 Siphoninus immaculatus (Heeger) 1856 

Trialeurodes inaequalis Gautier 1923 Siphoninus phillyreae (Haliday) 1835 

Trialeurodes intermedia Russell 1948 Trialeurodes intermedia Russell 1948 

Trialeurodes ipomoeae Carapia 2003 Trialeurodes ipomoeae Carapia 2003 

Trialeurodes ishigakiensis Takahashi 1933 Aleurotrachelus ishigakiensis (Takahashi) 1923 

Trialeurodes kankoensis Takahashi 1933 Pealius kankoensis (Takahashi) 1933 

Trialeurodes klemmi Takahashi 1940 Trialeurodes lauri (Signoret) 1882 

Trialeurodes lauri (Signoret) 1882 Trialeurodes lauri (Signoret) 1882 

Trialeurodes longispina Takahashi 1943 Trialeurodes longispina Takahashi 1943 

Trialeurodes lubia El Khidir & Khalifa 1962 Trialeurodes ricini (Misra) 1924 

Trialeurodes madroni (Bemis) 1904 Trialeurodes madroni (Bemis) 1904 

Trialeurodes magnoliae Russell 1948 Trialeurodes magnoliae Russell 1948 

Trialeurodes malayensis Corbett 1935 Dialeuropora malayensis (Corbett) 1935 

Trialeurodes mameti Takahashi 1951 Trialeurodes mameti Takahashi 1951 

Trialeurodes manihoti (Bondar) Trialeurodes manihoti (Bondar) 1923 

705 

Trialeurodes megapapillae Singh 1932 Viennotaleyrodes megapapillae (Singh) 1932 

Trialeurodes meggitti Singh 1933 Trialeurodes meggitti Singh 1933 

Trialeurodes merlini (Bemis) 1904 Trialeurodes merlini (Bemis) 1904 

Trialeurodes mirissimus Sampson & Drews 1941 Trialeurodes mirissimus Sampson & Drews 1941 

Trialeurodes mori Takahashi 1932 Pealius mori (Takahashi) 1932 

Trialeurodes mossopi Corbett 1935 Trialeurodes vaporariorum (Westwood) 1856 

Trialeurodes multipori Russell 1948 Trialeurodes multipori Russell 1948 

Trialeurodes notata Russell 1948 Trialeurodes notata Russell 1948 

Trialeurodes oblongifoliae Russell 1948 Trialeurodes oblongifoliae Russell 1948 

Trialeurodes packardi (Morrill) 1903 Trialeurodes packardi (Morrill) 1903 

Trialeurodes palaquifolia Corbett 1935 Trialeurodes palaquifolia Corbett 1935 

Trialeurodes paucipapilla Martin 2005 Trialeurodes paucipapilla Martin 2005 

Trialeurodes perakensis Corbett 1935 Trialeurodes perakensis Corbett 1935 

Trialeurodes pergandei (Quaintance) 1900 Trialeurodes pergandei (Quaintance) 1900 

Trialeurodes phillyreae (Haliday) 1835 Siphoninus phillyreae (Haliday) 1835 

Trialeurodes phlogis Russell 1993 Trialeurodes phlogis Russell 1993 

Trialeurodes porosus Priesner & Hosny 1937 Aleuroclava porosus (Priesner & Hosny) 1937 

Trialeurodes rara Singh 1931 Trialeurodes ricini (Misra) 1924 

Trialeurodes rex Martin 2001 Trialeurodes rex Martin 2001 

Trialeurodes ricini (Misra) 1924 Trialeurodes ricini (Misra) 1924

Trialeurodes rolfsii (Quaintance) 1899 Trialeurodes abutiloneus (Haldeman) 1850 

Trialeurodes ruborum (Cockerell) 1897 Trialeurodes ruborum (Cockerell) 1897 

Trialeurodes sardiniae Rapisarda 1986 Trialeurodes sardiniae Minelli et al. 1995 

706 

Trialeurodes sesbania Corbett 1936 Trialeurodes vaporariorum (Westwood) 1856 

Trialeurodes shawundus Baker & Moles 1923 Trialeurodes shawundus Baker & Moles 1923 

Trialeurodes silvarum Corbett 1935 Dialeuropora silvarum (Corbett) 1935 

Trialeurodes similis Russell 1948 Trialeurodes similis Russell 1948 

Trialeurodes sonchi (Kotinsky) 1907 Trialeurodes vaporariorum (Westwood) 1856 

Trialeurodes tabaci Bondar 1928 Trialeurodes tabaci Bondar 1928 

Trialeurodes tentaculatus (Bemis) Trialeurodes tentaculatus (Bemis) 1904 

Trialeurodes tephrosiae Russell 1948 Trialeurodes tephrosiae Russell 1948 

Trialeurodes thaiensis Takahashi 1943 Trialeurodes thaiensis Takahashi 1943 

Trialeurodes unadutus Baker & Moles 1923 Trialeurodes unadutus Baker & Moles 1923 

Trialeurodes vaporariorum (Westwood) 1856 Trialeurodes vaporariorum (Westwood) 1856 

Trialeurodes varia Quaintance & Baker 1937 Trialeurodes varia Quaintance & Baker 1937 

Trialeurodes variabilis (Quaintance) 1900 Trialeurodes variabilis (Quaintance) 1900 

Trialeurodes vitrinellus (Cockerell) 1903 Trialeurodes vitrinellus (Cockerell) 1903 

Trialeurodes vitrinellus (Cockerell); misidentification in part Trialeurodes intermedia Russell 1948 

Trialeurodes vittata (Quaintance) 1900 Trialeurodes vittata (Quaintance) 1900 

Trialeurodes wellmanae (Bemis) Trialeurodes vittata (Quaintance) 

Trialeurodes williamsi Trehan 1938 Filicaleyrodes williamsi (Trehan) 1938 

TRIALEUROLONGA MARTIN 2005 TRIALEUROLONGA MARTIN 2005 

Trialeurolonga trifida Martin 2005 Trialeurolonga trifida Martin 2005 

TRICHOALEYRODES TAKAHASHI & MAMET 1952 TRICHOALEYRODES TAKAHASHI & MAMET 1952 

Trichoaleyrodes carinata Takahashi & Mamet 1952 Trichoaleyrodes carinata Takahashi & Mamet 1952 

TUBERALEYRODES TAKAHASHI 1932 TUBERALEYRODES TAKAHASHI 1932 

Tuberaleyrodes bobuae Takahashi 1934 Tuberaleyrodes bobuae Takahashi 1934 

Tuberaleyrodes machili Takahashi 1932 Tuberaleyrodes machili Takahashi 1932 

Tuberaleyrodes neolitseae Young 1944 Tuberaleyrodes neolitseae Young 1944 

Tuberaleyrodes rambutana Takahashi 1955 Tuberaleyrodes rambutana Takahashi 1955 

Tuberaleyrodes spiniferosa (Corbett) 1933 Tuberaleyrodes spiniferosa (Corbett) 1933 

URALEYRODINAE SAMPSON & DREWS) 1941 URALEYRODINAE SAMPSON & DREWS) 1941 

UDAMOSELINAE ENDERLEIN 1909 UDAMOSELINAE ENDERLEIN 1909 

UDAMOSELIS ENDERLEIN 1909 UDAMOSELIS ENDERLEIN 1909 

Udamoselis estrellamarinae Martin 2007 Udamoselis estrellamarinae Martin 2007 

Udamoselis pigmentaria Enderlein 1909 Udamoselis pigmentaria Enderlein 1909 

Uraleyodes Sampson & Drews 1941 Aleurocerus Bondar 1923 

Uraleyrodes ceriferus Sampson & Drew 1941 Aleurocerus ceriferus (Sampson & Drew) 1941 

Uraleyrodes ceriferus Sampson & Drews 1941 Aleurocerus ceriferus (Sampson & Drews) 1941

VASANTHARAJIELLA DAVID 2000 VASANTHARAJIELLA DAVID 2000 

Vasantharajiella kalakadensis David 2000 Vasantharajiella kalakadensis David 2000 

VASDAVIDIUS RUSSELL 2000 VASDAVIDIUS RUSSELL 2000 

Vasdavidius cobarensis (Martin) 1999 Vasdavidius cobarensis (Martin) 1999 

Vasdavidius concurus (Ko) 1998 Vasdavidius concursus (Ko) 1998 

Vasdavidius indicus (David & Subramaniam) 1976 Vasdavidius indicus (David & Subramaniam) 1976 

Vasdavidius miscanthus (Ko) 1998 Vasdavidius miscanthus (Ko) 1998 

Vasdavidius setiferus (Quaintance & Baker) 1917 Vasdavidius setiferus (Quaintance & Baker) 1917 

VENEZALEURODES RUSSELL 1967 VENEZALEURODES RUSSELL 1967 

Venezaleurodes pisoniae Russell 1967 Venezaleurodes pisoniae Russell 1967 

VIENNOTALEYRODES COHIC 1968 VIENNOTALEYRODES COHIC 1968 

Viennotaleyrodes bergerardi (Cohic) Viennotaleyrodes bergerardi (Cohic) 1966 

Viennotaleyrodes bicolorata Martin 1999 Viennotaleyrodes bicolorata Martin 1999 

707 

Viennotaleyrodes bosciae Bink-Moenen 1983 Viennotaleyrodes bosciae Bink-Moenen 1983 

Viennotaleyrodes bourgini Cohic 1968 Viennotaleyrodes bourgini Cohic 1968 

Viennotaleyrodes curvisetosus Martin 1999 Viennotaleyrodes curvisetosus Martin 1999 

Viennotaleyrodes dichrostachi Bink-Moenen 1983 Viennotaleyrodes dichrostachi Bink-Moenen 1983 

Viennotaleyrodes fallax Bink-Moenen 1983 Viennotaleyrodes fallax Bink-Moenen 1983 

Viennotaleyrodes incomptus Martin 1999 Viennotaleyrodes incomptus Martin 1999 

Viennotaleyrodes indicus (Jesudasan & David) 1991 Viennotaleyrodes megapapillae (Singh) 1932 

Viennotaleyrodes lacunae Martin 1999 Viennotaleyrodes lacunae Martin 1999 

Viennotaleyrodes lafonti Cohic 1968 Viennotaleyrodes megapapillae (Singh) 1932 

Viennotaleyrodes megapapillae (Singh) 1932 Viennotaleyrodes megapapillae (Singh) 1932 

Viennotaleyrodes nilagiriensis David & Krishnan 1994 Viennotaleyrodes nilagiriensis David & Krishnan 1994 

Viennotaleyrodes platysepali (Cohic) Viennotaleyrodes platysepali (Cohic) 1966 

XENALEYRODES TAKAHASHI 1936 XENALEYRODES TAKAHASHI 1936 

Xenaleyrodes artocarpi Takahashi 1936 Xenaleyrodes artocarpi Takahashi 1936 

Xenaleyrodes broughae Martin 1985 Xenaleyrodes broughae Martin 1985 

Xenaleyrodes eucalypti (Dumbleton) 1956 Xenaleyrodes eucalypti (Dumbleton) 1956 

Xenaleyrodes fauceregius Martin 1999 Xenaleyrodes fauceregius Martin 1999 

Xenaleyrodes irianicus Martin 1985 Xenaleyrodes irianicus Martin 1985 

Xenaleyrodes timonii Martin 1985 Xenaleyrodes timonii Martin 1985 

XENOBEMISIA TAKAHASHI 1951 XENOBEMISIA TAKAHASHI 1951 

Xenobemisia coleae Takahashi 1951 Xenobemisia coleae Takahashi 1951 

YLEYRODES BINK-MOENEN 1983 YLEYRODES BINK-MOENEN 1983 

Yleyrodes isoberliniae Bink-Moenen 1983 Yleyrodes isoberliniae Bink-Moenen 1983 

ZAPHANERA CORBETT 1926 ZAPHANERA CORBETT 1926 

Zaphanera cyanotis Corbett 1926 Zaphanera cyanotis Corbett 1926 

Zaphanera indicus Jesudasan & David 1991 Zaphanera indicus Jesudasan & David 1991

Zaphanera nigra (Maskell) 1895 Pseudozaphanera niger (Maskell) 1895 

Zaphanera papyrocarpae Martin 2001 Pseudozaphanera papyrocarpae (Martin) 2001 

Zaphanera publicus (Singh) 1938 Zaphanera publicus (Singh) 1938 

Zaphanera rhachisreticulata Martin 1999 Pseudozaphanera rhachisreticulata (Martin) 1999 

Zaphanera splendida Martin 1999 Pseudozaphanera splendida (Martin) 1999 

Zaphanera wariensis Martin 1999 Pseudozaphanera wariensis (Martin) 1999 

708

The Whiteflies of the World and Their Host - Keys - Lucid Key Server ...

Create successful ePaper yourself

Delete template?

Save as template?