Attributes of British and Irish Plants - Natural Environment Research ...

t 

Cent re for 

Ecology & Hydrology 

N AT U RAL ENV I RO N M ENT RES EA RC H C O U N C I L

PLANTATT 

Attributes of British and Irish Plants: 

Status, Size, Life History, 

Geography and Habitats 

for use in connection with the 

New atlas of the British and Irish fl ora 

M. 0 . Hill, C. D. Preston & D. B. Roy 

Biological Records Centre 

NERC Centre for Ecology and Hydrology 

Centre tor Et ctotry entl Hydrology 

Bush Es te:e 

Penicuik 

Midlottna t 

6 326 OAS 

2004 

10 0 0 (T 4 - 2 0 0 1

@ NERC Copyright 2004 

Print ed in 2004 by Raven Marketing Group, Cambridgeshire 

ISBN I 870393 74 0 

The Cent re for Ecology and Hydrology (CEH) is one of t he Cent res and Surveys of the 

Nat ural Environment Research Council (NERC). Established in 1994, CEH is a mult idisciplinary, 

environmental research organisat ion wit h some 600 staf and well-equipped 

laboratories and fi eld facilities at nine sites t hroughout t he Unit ed Kingdom. CEH's 

mission is to "advance t he science of ecology, environmental microbiology and hydrology 

t hrough high quality and internationally recognised research leading to bett er 

understanding and quant ifi cat ion of the physical, chemical and biological processes relating 

to land and freshwat er and living organisms wit hin t hese environments". 

The Biological Records Cent re is operated by CEH, based at CEH Monks Wood. BRC is 

joint ly funded by CEH and t he Joint Nature Conservation Committ ee (www.jncc.gov.uk), 

t he latter act ing on behalf of the statutory nat ure conservation agencies in England, 

Scotland,Wales and Nort hern Ireland. CEH and JNCC support BRC as an important 

component of t he developing National Biodiversity Network. BRC seeks to help 

nat uralists and research biologists to co-ordinate t heir eff ort s in studying the occurrence 

of plant s and animals in Britain and Ireland, and to make t he results of these st udies 

available to others. 

For furt her informat ion, visit www.ceh.ac.uk. 

Cover photograph shows the endemic plant Coincya wright ii on the cliffs of Lundy (courtesy of 

Roger Key, English Nature). 

Published by 

Cent re for Ecology & Hydrology 

Monks Wood 

Abbot s Ripton 

Hunt ingdon 

Cambridgeshire 

PE28 2LS 

Copies can be ordered by writing to the above address or by telephoning 

0 1487 772400, or faxing 0 1487 772558.

CON TEN TS 

Int roduction 

Species, fam ilies, stat us and change 3 

Species and families . 3 

Native/afi en status . 3 

Conservation status . . . 3 

Rarity status . 3 

Change Index . 4 

Size and life history at tributes . . . 

Height and length . 

Perennation . 5 

Life form . . . . . 5 

Woodiness 

Clonality 

Geographic attributes 

European and wider distributions, and continentality . . 9 

Native distribution of species alien in Britain and Ireland . . . 10 

Counts of 10-km squares in the New Atlas . 10 

Climatic means 11 

H abitat att ributes . . . . . 12 

Coastal species . 12 

Broad Habitats . 12 

Ellenberg's indicator values . 14 

Light values (L) . 14 

Moisture values (F) . 15 

Reaction (R) . 16 

Nitrogen (N) . . . 16 

Sak tolerance (S) . . 17 

Listing of species with their attributes 18 

A ppendix 6 1 

Sources for plant height . . 6 1 

Defi nition of life forms . 6 1 

Examples of how fi fe-form criteria are applied . . 62 

Links between life forms and plant height . 63 

Clonality . 64 

Broad Habitats . 64 

Initial calculation of species' preferences for Broad Habitats . 68 

Extension and validation of species' preferences . 69 

Acknowledgem ent s 70 

Bibliography 7 1

IN TR OD UCTI ON 

In the course of our research on geographical distributions and climate change, we have 

frequent ly wanted to characterize the plants t hat const itute the wild fl ora of Britain and Ireland. 

Several sources of information were available, including t he Electronic Comparative Plant Ecology 

(Hodgson et al., 1994), t he Czech clonal plant database Clop la I (http.//www.butbn.cas.ce kfi mes/ 

clopla I .htm), t he BSBI database (http://www.bsbi.org.uk/html/database.html), t he Ecoflora database 

(Fitter & Peat, 1994), and the indicator values of Ellenberg (Ellenberg et S., 1991). Inevitably, these 

sources were not sufi cient for our purposes, and we have gradually assembled our own set of 

att ribute data, linked to t he database of the Biological Records Centre (BRC). 

The establishment of the National Biodiversity Network (NBN) provided an additional st imulus 

to the assembling of att ribute data, of ering a suitable platform for online searches for species 

att ributes. Some of the information published here is also available from the NBN gateway 

(www.searchnbn.net). In 2002, the publicat ion of the New Atlas of the British and Irish Flora (Preston, 

Pearman & Dines, 2002), toget her with results of a companion project to analyse change 

(Preston,Telfer et aL, 2002), allowed us to complete the dataset that is presented here. 

The dataset is partly old and partly new (Table 1). Biogeographic elements and Ellenberg values 

are taken with few modifi cat ions from earlier publications (Preston & Hill, 1997, 1999; Hill et 

1999). Information that depends directly on the New Atlas is either new or (Change Index) is 

taken from the text of the New Atlas. Data on t he Broad Habitats of plants are mostly drawn 

from an unpublished report (Preston et al., 2003). Information on plant height, perennat ion, life 

form and clonal growt h was assembled from a variety of fl oras and monographs, with some 

personal observation; it is a new compilat ion. Furt her details of t hese att ributes are provided in 

t he following pages. The attribute categories are tabulated (Tables 2- 13), with the numbe of taxa 

in each category listed in the column 'N='.

Table I . Attributes, codes and names listed as columns in PLA NTATT 

Colum n nam e 

(a) Status and taxonomic 

Taxon name 

Family 

Nat ive status 

Conservation status 

Rarity status 

Change Index 

(b) Size and life form 

Height (terrest rial) 

Length (aquatic) 

Perennation 

Life form 

Woodiness 

Clonal spread 

(c) Geography & climate 

Major Biome (European 

dist ribut ion) 

Eastern limit code 

Continentality in Europe 

O rigin of alien taxa 

Number of 10-km squares in 

Britain (including Isle of Man) 

Number. of 10-km squares in 

Ireland 

Number of 10-km squares in 

Channel Islands 

January mean terhperature 

July mean temperature 

Annual precipitat ion 

(d) Habitat 

Coastal 

Broad Habitats 

Ellenberg indicator value 

A bbreviat 

ion 

Fam 

NS 

CS 

RS 

Chg 

Hght 

Len 

P I , P2 

LF1, LF2 

Clonel , 

Clone2 

E I 

E2 

O rigin 

GB 

IR 

CI 

Tjan 

Tjul 

Prec 

Co 

Br Habitats 

Source or ot her comment 

Name in current flora (Stace, 1997) 

Stace (1997) 

New Atlas (Preston et al., 2002) 

Chefi ngs (2004) 

Calculated from New Atlas data 

New Atias (Preston et al., 2002) 

Several sources, listed below 

Several sources, listed below 

Mainly Clapham,Tutin & Warburg (1962) 

Clapham et al. ( 1962), heavily revised 

Mainly Clapham et al. ( 1962) 

Sources are listed below; clone2 is mainly blank 

Part of biogeographic element 

accor ding to Preston & Hill ( 1997) 

Part of biogeographic element 

according to Preston & Hill ( 1997) 

Preston & Hill ( 1997) plus additions 





Calculated from New Atlas plus climat ic data 

Calculat ed from New Atlas plus climat ic data 

Calculat ed from New Atlas plus climat ic data 

Calculated from New Atlas plus Ellenberg salt value 

Preston et al. (2003), plus previously unreported data 

for aliens 

L, F, R, N, S Hill et aL ( 1999)

SPE CIES, FAMILIES, S TA TUS A ND CHAN GE 

Species and families 

The list of 1885 taxa includes all nat ive British and Irish species, as well as natives of the Channel 

Islands not known from Britain or Ireland. There are no species restricted to the Isle of Man. All 

archaeophyt e species are included, plus a number of well-established neophytes.A small number 

of subspecies, hybrids and aggregate species are also included. The number of taxa has been 

slightly increased from the 1791 t hat were listed for Ellenberg indicator values (Hill et al., 1999). 

Most of t he addit ions are of neophytes, which, following the New Atlas (Preston et al., 2002), we 

now t hink t o be suffi cient ly common or potent ially interest ing to be included. 

The names of families have been abbreviated to four letters and follow Stace ( 1997). 

Native/alien status 

Species of uncertain stat us are classified as 'nat ive or alien'. Int roduced species have been 

classifi ed as archaeophytes, neophytes and casuals (Preston et al., 2002). Bot h archaeophytes and 

neophytes are introduced species which are present in t he wild as naturalized populations, that is 

t hey are spreading vegetat ively or reproducing ef ectively by seed. An archaeophyte is a plant 

t hat became naturalized before AD 1500. A neophyte is one that was first int roduced after 1500, 

or was only present as a casual before 1500 and is naturalized now only because it was 

reint roduced subsequent ly. In contrast to archaeophytes and neophytes, a casual is a plant that is 

present only as populat ions which fail to persist in the wild for periods of more t han 

approximately fi ve years. Such a species is dependent on constant reint roduction (Macpherson et 

at , 1996). 

Hybrids between two alien parents which were introduced to the wild as hybrids (e.g. Crocosmia 

x crocosmiifl ora, Euphorbia x pseudovirgata) are classified as neophytes. Only hybrids between 

which have been formed spontaneously in the wild between two alien parents are classified as 

'AX'. 

Conservation status 

The conservat ion status of plant s is coded according to the system proposed by Hodgetts, 

Palmer & W igginton ( 1996); see also Palmer et al. ( 1997). This status is an assessment of threat 

rather t han rarity, alt hough closely related to rarity in that only species known at the time of the 

assessment in less t han 10 1 10-km squares have been allocated t o one of the categories listed. 

The stat us list ed is t hat given by Chefi ngs (2004). This replaces t he stat uses provided in the Red 

Data Book (W igginton, 1999) and Scarce plants in Britain (Stewart , Pearman & Preston, 1994). 

Cheffi ngs assigns a conservat ion status to nat ive species and archaeophytes, with only a few 

exceptions. PLA NTATT cannot be used to provide a complete list of threatened taxa as many 

subspecies listed by Cheffi ngs are not t reated separately by us. 

Rarity status 

Rarity status is based on counts of the number of 10-km squares in Britain and t he Isle of Man in 

which the plant was recorded as a nat ive in t he period 1987- 1999. Rare plants are those 

recorded in 1- 15 10-km squares during t his period; scarce plants are recorded in 16- 100.The 

t hresholds were those that were used in defining rare and scarce species for the Red Data Book 

and Scarce plants. Rarity status, however, is solely a measure of rarity rat her than threat, and is 

based on data from the New Atlas. Cheffi ngs assigns a conservat ion status to nat ive species and 

archaeophyt es, with only a few except ions. PLANTATT cannot be used to provide a complete 

list of t hreat ened taxa as many subspecies listed by Cheffi ngs are not t reated separately by us. 

Alien species have by defi nition no nat ive records, and therefore are not given a rarity status.

Table 2. Nat ive stat us, Conservation stat us and Rarity stat us 

At tribute and codes N= Explanation 

(a) Native status 

(b) Conservation status 

(c) Rarity status 

(d) Change Index 

AC 12 Alien casual; many are crop plants 

A N 259 Neophyte, alien introduced after 1500 

A R 15 1 A rchaeophyte, alien introduced before 1500 

AX 1 Spontaneous hybrid between two alien parents 

1362 Native, not endemic 

NA 46 Native or alien (nat ive status doubtful) 

N E 47 Native endemic 

N H 7 Spontaneous hybrid between two native parents 

CR 20 Critically endangered 

D D I Data defi cient 

EN 34 Endangered 

EW 4 Ext inct in the wild 

EX 10 Ext inct 

VU 119 Vulnerable 

(values) 

933 Present, not rare or scarce 

234 Rare ( 1- 15 10-km squares in Britain, 1987- 1999) 

254 Scarce (16- 100 10-km squares in Britain, 1987- 1999) 

27 

(blank) 423 Alien taxa 

Absent from Britain and Isle of Man as a native, but native in 

Ireland or t he Channel Islands 

12 Apparently ext inct (not recorded since 1986) 

2 Insufi cient data available to assess rarity 

Change between 1930- 1960 and 1987- 1999 

Change Index 

The Change Index (Telfer et al., 2002) measures the relat ive magnitude of change, comparing the 

period 1930- 1960 with 1987- 1999. It is explained in more detail in t he Appendix. It ranges from 

-4.78 for the critically endangered Gahum tricornutum to 4.70 for t he much more frequent ly 

recorded Prunus laurocerasus. The Change Index has not been calculated for species not included 

in the fi rst Atlas of the British Flora (Perring & Walters, 1962). Consequent ly, several aliens such as 

Amsinckia micrantha and Crassula helmsii, which have increased spectacularly in the intervening 

period, lack a value for the Change Index.

SIZE A ND LI FE HIS TOR Y A TTRIB UTES 

H eight and length 

Heights, measured in cent imet res, are given for terrestrial (or emergent) plants and lengths (cm) 

for submerged aquatic plants. A few amphibious or emergent plants such as Persicaria amphibia 

are scored for both height and length. 

These values are int ended as a general indication of the size of the plant. For procumbent plants, 

heights are an indicat ion of the height of the less procumbent shoots. For scapose plants and 

ferns, heights are not stem lengths but leaf lengths. Likewise, lengths for isoetids (i.e. linear-leaved 

rosette-forming root ed aquat ics) are in fact leaf lengths. For example Lobelia dortmanna has 

length 4 cm, which is certainly not the length of t he scape. 

Height and length values were drawn from a variety of sources, which are detailed in the 

Appendix. 

Perennat ion 

The categories are annual, biennial and perennial. Biennial is used as a short hand to denote also 

monocarpic perennials. For example Car /Ma vulgaris and Heracleum mantegazzianum normally 

take several years to reach maturity but are categorized as biennials. The sources of data for 

perennation are most ly t he same as t hose used for height and length. In a few cases (e.g. Linum 

catharticum), we have relied on personal communication and scientifi c papers rat her than fl oras. 

A relat ively small number of species fall into more t han one of these categories. The secondary 

category is intended to be either less frequent or equally frequent. Thus Poe annua has primary 

perennation category P I =annual and secondary category P2 = perennial. Several fl oras suggest 

t hat Cardamine flexuosa and Montia fontana can be annual, biennial or perennial. These have been 

coded as PI = p, P2 = a and PI = a, P2 = p respectively. 

Life form 

The Raunkiaer system of life forms was set out by Clapham,Tutin & Warburg (1962), who listed 

life forms for most species. Their assignments provide an excellent starting point but contain 

numerous inconsistencies. Life forms have been fully revised for PLANTATT. Following German 

and Swiss aut hors (e.g. Lindacher, 1995), the category of helophytes or marsh plants has been 

omitt ed. The majority of t hese can grow in places that are not permanently fl ooded in winter. 

The position of the overwintering buds in such situations is taken as a guide to t heir life form. 

An additional life form of annual water plants has been added, so t hat t he category t herophytes 

are all terrest rial. 

As with perennat ion, a secondary life form is given for some species. Details of how life forms 

were assigned and the relation of life form to plant height are given in t he Appendix. 

Woodiness 

Woodiness is an attribut e that applies to some chamaephytes, nanophanerophytes and 

phanerophytes. All ot her life forms are t reated as herbaceous. Three categories are recognized, 

woody, semi-woody and herbaceous. Although we have been guided in part by Clapham,Tutin & 

W arburg ( 1962), we have not hesitated to express our own opinions where these diff er.

Table 3. Size and life history attributes other t han clonality; counts of primary 

att ributes are in the column N 1=, count s of secondary attributes are in N 2= 

Attribute or code N 1= N2= Explanat io n 

(a) Height (cm) Hght Height (leaf length for scapose plants and ferns) 

(b) Length (cm) Len Length (aquatic plants; leaf length for isoetids) 

(c) Perennation PI P2 Primary and secondary type of perennation 

a 455 14 Annual 

b 96 14 Biennial, including monocarpic perennials 

p 1334 42 Perennial 

(d) Life form L Fl LF2 Primary and secondary fi fe form (see Table Al ) 

(e) Woodiness 

Ch 148 24 Chamaephyte 

Gb 32 Bulbous geophyte 

Gn 134 11 Non-bulbous geophyte (rhizome, corm or tuber) 

hc 8 11 50 Hemicryptophyte 

Hy 124 34 Perennial hydrophyt e (perennial water plant) 

H z 25 1 Annual hydrophyte (aquatic therophyte) 

Ph 108 15 Mega-, meso- and microphanerophyte 

Pn 73 2 1 Nanophanerophyte 

T h 430 14 Therophyte (annual land plant) 

1651 Herbaceous 

sw 27 Semi-woody 

Vt 207 Woody 

Clonality 

Clonal growt h is defi ned here as vegetative reproduct ion combined with lateral spread. Like 

perennat ion, clonality may have more t han one value for a given species, refl ecting the various 

ways in which clonal growth may occur. In some species t here are varieties with clonal growt h 

(Arrhenatherum elatius van bulbosum, Caltha palustris var. radicans) while the normal form is not 

markedly clonal. In these two species, the variety with clonal growth is less frequent than the 

type variety, so that both plants are given Clone] = 0 (not spreading clonally) as the primary 

state, whereas Clone2 = D Rg and Sto12 respectively. Two species with proliferous 

infl orescences really ought to have been assigned to three categories, namely Juncus bulbosus 

(N ode2, Irreg and omitted DRi) and Butomus umbellatus (Rhiz2, DRg and omitted D Ri). It did 

not seem wort h having a third column just for these two, so the inflorescence character has 

been omitt ed.

Table 4. Cat egor ies of clonality; counts of primary attributes are in the column 

N 1=, count s of secondary attributes are in N 2= 

Attribute or code N 1 = N 2= Explanat io n 

(I) Clonafi ty Gone Clone2 

0 1228 Little or no vegetative spread 

Ogr 25 Tussock-forming graminoid, may slowly spread 

Ot b 26 Tuberous or bulbous, slowing cloning by of sets 

D Ra 20 10 Detaching ramets above ground (oft en axillary) 

D Rg 4 16 Detaching ramets at or below ground 

D Ri 11 Detaching ramets on infl orescence 

D R1 1 Detaching rarnets on leaves (Hammarbya) 

D Rp 1 Detaching ramets on prot hallus (Trichomanes) 

Frag 7 Fragmenting as part of normal growth 

Ir reg 38 17 Irregularly fragmenting (mainly water plants) 

Leaf 1 Plantlets formed on leaves (Cardamine pratensis) 

N ode l 19 20 Short ly creeping and rooting at nodes 

N ode2 76 9 Ext ensively creeping and rooting at nodes 

Rhiz l 152 8 Rhizome shortly creeping 

Rhiz2 209 15 Rhizome far-creeping 

Root 37 3 Clones formed by suckering from roots 

St ol l 14 Short ly creeping, stolons in illuminated medium 

St o12 26 5 Far-creeping by stolons in illuminated medium 

T ip 5 Tip rooting (the stems often turn downwards) 

The following comments may be helpful in distinguishing the clonal categories. 

I. The category Ogr is rather poorly defi ned. If a perennial graminoid (grass, sedge or rush) is 

repeatedly mown or heavily grazed, it may gradually form clones, even though this is not the 

normal condition. 

2. The category Ot b assumes that the ground is litt le disturbed. If there is regular disturbance, 

t he of sets will be dispersed to new sites and t he plant would then be indicated as DRg 

(detaching ramets at or below ground). 

3. The small category Frag applies to duckweeds and Azola 

4. The categories N ode (creeping and root ing at nodes), Rhiz (rhizomatous or with 

subterranean stolons) and Stol (stoloniferous in the illuminated medium, eit her air or 

water) are divided into two, according to whether they spread a short distance in a season 

or are far-creeping. For rhizomatous plants, a rule of t humb is that a species whose new 

shoots arise at a distance less t han a quarter of the height of the plant is short ly creeping,

while those that spread further are deemed to be far-creeping. Analogous distinct ions are 

made for N ode and Stol. 

5. The distinction between Node and Stol is that in N ode the horizontal or decumbent 

stem extends with indeterminate growth, rooting at the leafy nodes and not forming a new 

stem axis at each rooted position. In St ol, a new stem axis normally arises at the point of 

root ing, or the stolon is determinate, bending upwards and not progressing with 

indeterminat e growt h. 

6. In the tip-rooting category, T ip, leafy stems that are not obviously stolons turn down at the 

apex and root there.

GE OGRAPHI C A TTRIB UTES 

Euro ean and wider distribut ions and cont inent alit 

The categorizat ion of taxa to biogeographic elements and their designat ion as 'continental' or 

ot herwise (Table 5) follow Preston & Hill ( 1997), with minor revisions and t he addit ion of the 

hyperoceanic category from a subsequent paper (Preston & Hill, 1999). Element s are provided 

for nat ive species and some archaeophytes; for archaeophyt es the classifi cation describes the 

archaeophyt ic range. In addition, some but not all alien species originat ing in Europe have also 

been assigned to biogeographic elements for PLANTATT and designated as continental. 

Geographic att ribut es are not provided for some hybrids. 

Table 5. European and wider dist ribution, and continentality 

Att ribut e 

and codes 

N = Ex planat io n

Native distribution o s ecies alien in Britain and Ireland 

The native dist ribution of aliens (Table 6) is taken from the New Atlas. This information is 

provided for neophytes and some archaeophytes. In Table 6, taxa present in two areas are 

included in bot h totals but taxa are excluded from the totals if they are only doubtfully native in 

the relevant area. 

Table 6. Nat ive dist ribution of species alien in Britain and Ireland 

Code N = Explanat ion 

Am 24 North America 

Am4 22 West ern North America 

Am6 5 Eastern North America 

As 10 Asia east of 60°E 

As I 24 Asia between 60°E and I 20°E 

As2 9 Asia E of I 20°E 

Aus 3 Aust ralia 

Crop 15 Crop plant, does not have a native range 

Eur 149 Europe 

Gard 13 Garden or igin, does not have a native range 

NHem 6 N Hemisphere (Europe,Asia and Nort h America) 

NZ 4 New Zealand 

SAf 5 Southern Africa 

SAm 20 South America and/or Cent ral America 

Unk 10 Unknown 

Counts o I O-km s uares in the New Atlas 

For each taxon, the number of 10-km squares in Britain, Ireland and the Channel Islands is. 

enumerated (Table 7). For most natives, only t he native distribution has been counted (blue dots 

in the New Atlas). For the small number of nat ive species in which it proved impossible even to 

attempt to delimit the nat ive range in the New Atlas, the count is for all squares. For alien taxa, 

all squares with records have been counted. Squares have been counted without regard to date, 

so that Otanthus maritimus, ext inct in Britain, is recorded from 23 squares t here. 

10

Table 7. Counts of squares and climat ic means 

Attribute 

(a) Counts 

GB 

Min Max Explanation 

IR 0 985 Ireland 

Counts of 10-km squares 

0 2805 Great Britain and Isle of Man 

CI 0 14 Channel Islands 

(b) Climatic means Mean values for 10-km squares 

Tian - 1.3 7.0 January mean temperat ure (°C) 

Tjul 10.4 17.0 July mean temperature (°C) 

Prec 553 32 18 Annual precipitation (mm) 

Climatic means 

Climat ic values for plants were calculated as t he mean climate of the 10-km squares where they 

occur in Britain, Ireland and the Channel Islands, averaging over the squares enumerated for the 

counts. Climate data for 10-km squares were taken from baseline climate summaries of t he UK 

Climate Impacts Programme (Hu Ime & Jenkins, 1998). These baseline summaries were 

const ructed by interpolat ion of daily weat her measurements from individual met stat ions, 

averaged over the 30-year period 1961- 1990 (Barrow et al., 1993).

HA BI TA T A TTRIB UTES 

Coastal species 

Species are deemed to be coastal if 80% of occupied squares are on the coast and if they depend 

on coastal habitat. Most coastal species have Ellenberg salt values (S values, defi ned below) 

greater than 0. Those with Ellenberg S = 0 were scrutinized carefully to ascertain whether their 

habitat was indeed coastal. 

Broad Habitats 

The preferences of species for Broad Habitats of the UK BAP are listed in the column labelled Br 

Habitats. Species' main habitat (s) are_list ed. No species is deemed to have a preference for 

more t han four habitats. Minor habitats are ignored. For more informat ion about Broad 

Habitats and species' preferences, refer to the Appendix.

Table 8. Coastal species and preferences for Broad H abitats 

A ttribut e 

and codes 

N = Explanat io n 

(a) Co Coastal species 

Co 145 At least 80% of occupied squares contain sea at high tide 

(b) Br Nabs Broad Habitats 

3 10 Broad leaved, mixed and yew woodland 

2 32 Coniferous woodland 

3 543 Boundary and linear features (eg hedges, roadsides, walls) 

4 198 A rable and horticultural (includes orchards, excludes domest ic gardens) 

5 22 Improved grassland 

6 163 Neut ral grassland (includes coarse Arrhenatherum grassland) 

7 2 18 Calcareous grassland (includes lowland and montane types) 

8 89 Acid grassland (includes non-calcareous sandy grassland) 

9 10 Bracken 

10 83 Dwarf shrub heat h (cover of dwarf shrubs at least 25%) 

I I 254 

12 4 1 

Fen, marsh and swamp (not wooded; includes flushes, rush-pastures, springs and 

mud communit ies) 

Bog (on deep peat; includes bog pools as well as acid lowland valley mires on 

slight ly shallower peat) 

13 174 Standing wat er and canals 

14 149 Rivers and st reams 

I S 103 Montane habitats (acid grassland and heat h with montane species) 

16 292 

Inland rock (heterogeneous - includes quarries, limestone pavement, clif s, screes 

and skeletal soils over rock) 

17 231 Built-up areas and gardens 

18 74 Supra littoral rock (does not include maritime grassland) 

19 135 Supra littoral sediment (st randlines, shingle, coastal dunes) 

2 1 65 Littoral sediment (includes saltmarsh and saltmarsh pools) 

23 1 Inshore sublittoral sediment (only Zostera marina)

Ellenbe s indicator values 

Ellenberg defi ned seven major scales, of which fi ve are presented here. The two that are 

omitted, T (temperat ure) and K (cont inentality), correspond quite closely to the major biome 

and east ern limit categories defi ned for European dist ribut ions by Preston & Hill (1997). Neither 

T nor K values are satisfactory in an oceanic climate such as that of Britain; t hose for K are 

particularly unreliable, especially as Ellenberg's defi nition was geographical rather than climat ic. 

The fi ve remaining scales have values defi ned in the tables that follow. The values are based on 

those of Ellenberg et at (1991). They are mostly reproduced from a previous publication (Hill et 

al., 1999), with some additions because of the ext ra species included here. A few example 

species are given for each value, by way of explanation. 

Light values (L) 

The full range of Ellenberg values for light (Table 9) is not represent ed in the British fl ora. For 

canopy t rees, light values refer to the tolerance of t he sapling stage of the life cycle. 

Table 9. Ellenberg values for light (L) 

Code N = Explanat io n 

0 Plant in deep shade (no examples for Britain or Ireland) 

2 4 Between 1 and 3 (Epipogium aphyllum, Neottia nidus-avis,Trichomenes speciosum) 

Shade plant, most ly less than 5% relative illuminat ion, seldom more than 30% 

3 20 illumination when t rees are in full leaf (Ga hurn odoraturn, Listera cordate, Mercurialis 

perennis) 

4 70 Between 3 and 5 (Circaea lutetiono, Lamiastrum galeobdolum, Poa nérnoralis) 

Semi-shade plant, rarely in full light, but generally with more than 10% relat ive 

5 120 illumination when t rees are in leaf (Carex pendula, Hyacinthoides non-scripta, Primula 

vulgaris) 

6 2 13 Between 5 and 7 (Anthriscus sylvestris, Digitalis purpurea, Teucriurn scorodonia) 

7 680 

8 576 

9 202 

Plant generally in well lit places, but also occurring in part ial shade (Arrhenatherurn 

elatius, Carex flacco, Poo trivialis,Vicia cracca) 

Light-loving plant rarely found where relat ive illumination in summer is less than 40% 

(Cardarnine hirsute, Orchis morio,Thymus polytrichus,Vacciniurn oxycoccus) 

Plant in full light, found mostly in full sun (Aster tripolium, Mel/ lotus albus, Poa compressa, 

Primula farinosa) 

14

M oisture values (F) 

Unlike the ot her Ellenberg values, moisture is on a scale of I to 12 (Table 10). We use the 

abbreviation F from the German Feuchtigkeit. 

Table 10. Ellenberg values for moisture (F) 

Code N = Explanat ion 

1 3 

Indicator of ext reme dryness, restricted to soils that oft en dry out for some time 

(Corynephorus canescens, Helianthemum apenninum, Koeleria vallesiana) 

2 28 Between 1 and 3 (Clinopodium acinos, Saxifraga tridactyfi tes, Sedum acre) 

3 168 

4 378 

5 492 

Dry-site indicator, more oft en found on dry ground than in moist places (Asplenium 

trichomanes, Centaurea scabiosa, Spergularia rubra) 

Between 3 and 5 (Arctium minus, Hefi ctotrichon pratense, Iris foetidissima,Thymus 

polytrichus) 

Moist-site indicator, mainly on fresh soils of average dampness (Anthriscus sylvestris, 

Euphorbia amygdaloides, Hyacinthoides non-scripta, Solarium nigrum) 

6 226 Between 5 and 6 (Agrostis stolonifera, Empetrum nigrum, Rumex crispus) 

7 14 1 

8 170 

9 126 

10 64 

I I 34 

12 

55 

Dampness indicator, mainly on constant ly moist or damp, but not on wet soils (Carex 

Dactylorhiza maculata, Pulicaria dysenterica, Ranunculus repens) 

Between 7 and 9 (Cardamine pratensis, Equisetum telmateia, Phalaris arundinacea, Schoenus 

nigricans) 

Wet-site indicator, oft en on water-saturated, badly aerated soils (Drosera rotundifofi a, 

Myosotis scorpioides,Vaccinium oxK occus, Viola palustris) 

Indicator of shallow-water sites that may lack standing water for ext ensive periods 

(Afi sma plantago-aquatica, Carex limosa, Ranunculus fi ngua,Typha latifofi a) 

Plant root ing under water, but at least for a time exposed above, or plant fl oat ing on 

t he surface (Lemna minor, Nuphar lutea, Sagit aria sagittifolia, Schoenoplectus lacustris) 

Submerged plant, permanently or almost constantly under water (Isoetes lacustris, 

Potamogeton crispus, Ranunculus circinatus, Zostera marina) 

15

Reaction (R) 

Reaction (Table I I) refers to environmental acidity, which would ordinarily be measured by pH. 

Except for water plants, R values refl ect preferences for soil acidity. 

Table I I . Ellenberg values for reaction (R) 

Code N = Ex planat io n 

Indicator of ext reme acidity, never found on weakly acid or basic soils (Andromeda 

polifolia, Lycopodium clavatum, Rubus chamaemorus, Ulex minor) 

Between I and 3 (Agrostis curtisii, Calluna vulgaris, Drosera rotundifofi a, Po lygala serpyllifolia) 

Acidity indicator, mainly on acid soils, but exceptionally also on nearly neutral ones 

(Agrostis vinealis, Dactylorhiza M a de ra, Ga lium saxatile, Pteridium aquilinum) 

Between 3 and 5 (Agrostis capillaris, Carex panicea,Juncus eff usus,Teucrium scorodonia) 

Indicator of moderately acid soils, only occasionally found on very acid or on neutral to 

basic soils (Cardamine pratensis, Cirsium palustre, Rubus idaeus, Ulex eumpaeus) 

Between 5 and 7 (Ammophila arenaria, Carex sylvatica, Lofi um perenne, Ranunculus ficaria) 

Indicator of weakly acid to weakly basic conditions; never found on very acid soils 

(Agrimonia eupatoria,Atriplex prostrata, Nuphar lutea, Phleum pretense) 

Between 7 and 9 (Artemisia vulgaris, Carduus nutans, Iris foetidissima, Viola hirsuta) 

Indicator of basic reaction, always found on calcareous or other high-pH soils (Bunium 

bulbocastanum, Clinopodium calamintha, Dryopteris submontana, Primula farMosa) 

Nitrogen (N) 

Nitrogen values (Table 12) are in fact a general indication of preference for soil fert ility. They are 

closely correlated with t he st ress values of Grime ( 1979, 200 1), low N values corresponding to 

plants with high st ress tolerance (Grime et al., 1997) and vice-versa. 

Table 12. Ellenberg values for nitrogen (N) 

Code N= Explanat io n 

1 

84 

Indicator of ext remely infertile sites (Agrostis curtisii, Cli nopodium acinos, Drosera 

rotundifolia, Rubus chamaemorus) 

2 323 Between 1 and 3 (Aira praecox, Carex panicea, Linum catharticum, Scabiosa columbaria) 

3 286 

Indicator of more or less infert ile sites (Centaured scabiosa, Go lium saxatile, Pimpinella 

saxifraga,Teucrium scorodonia) 

4 245 Between 3 and 5 (Agrostis capillaris, Cirsium palustre, Plantago lanceolata, Primula vulgaris) 

5 

342 

Indicator of sites of intermediate fert ility (Angelica sylvestris, Digitalis purpurea, Iris 

foetidissima,Trifolium pratense) 

6 3 19 Between 5 and 7 (Cirsium arvense, Glyceria fluitans, Poa trivialis, Rumex crisp s) 

7 223 

Plant often found in richly fertile places (Atriplex prostrate, Epilobium hirsutum, Stellaria 

media,Typha latifolia) 

8 55 Between 7 and 9 (Beta vulgaris, Galium aparine, Lamium album, Urtica dioica) 

9 8 

Indicator of ext remely rich situat ions, such as cattle resting places or near pollut ed 

rivers (Arctium laPpa,Artemisia absinthium, Hyoscyamus niger, Rumex obtusifolius)

Salt tolerance (5) 

Values for salt tolerance (Table 13) start at zero, corresponding to no tolerance of salt. 

Table 13. Ellenberg values for salt tolerance (S) 

Explanat ion 

Absent from saline sites; if in coastal situations, only accidental and non-persistent if 

subjected to saline spray or water (85% of the fl ora) 

Slightly salt-tolerant species, rare to occasional on saline soils but capable of persisting 

in the present of salt (includes dune and dune-slack species where the ground water is 

fresh but where some inputs of salt spray are likely) (Calystegia sepium, Chenopodium 

album, Oenanthe crocata, Sedum angfi cum) 

Species occurring in both saline and non-saline situat ions, for which saline habitats are 

not stro ngly predominant (Atriplex prostrata, Elytrigia repens, Phragmites australis, Rumex 

crispus) 

Species most common in coastal sites but regularly present in freshwater or on nonsaline 

soils inland (includes strictly coastal species occurring in sites such as cliff crevices 

and sand dunes that are not obviously salt-af ected) (Cakile maritima, Cochlearia officinalis, 

Juncus gerardii, Spergularia rupicola) 

Species of salt meadows and upper saltmarsh, subject to at most only very occasional 

tidal inundation (includes species of brackish conditions, i.e. of consistent but low 

salinity) (Atriplex littoralis, Elytrigia atherica, Glaux maritima,Triglochin maritimum) 

Species of the upper edge of saltmarsh, where not inundated by all tides (includes 

obligate halophytes of clif s receiving regular salt spray) (Aster tripofi um, Crithmum 

maritimum, Puccinellia maritima, Suaeda vera) 

Species of mid-level saltmarsh (Atriplex portulacoides, Cochlearia anglica, Limonium vulgare) 

Species of lower saltmarsh (Spartina anglica, Suaeda maritima) 

Species more or less permanently inundated in sea water (Zostera spp.) 

Species of extremely saline conditions, in sites where sea water evaporates, 

precipitating salt (Safi cornia europaea agg.; these could equally well be t reated as 

species of t he lower marsh)

LIS TIN G OF SPE CIES WI TH THEIR A TTRIB UTES 

The listing of species with their attributes has 32 columns. Codes used in these columns are set 

out in the foregoing tables. The short and full names of the columns are given here (Table 14), as 

are the pages on which the tables of codes appear 

Table 14. Plant attributes cross-referenced to tables and page numbers where t hese 

are described more fully 

Short name Attribute Table N o Page 

Taxon name Name in New At las 1 2 

Fam Family (4-letter abbreviation) 1 2 

NS Native status 2 4 

CS Conservation status 2 4 

RS Rarity status 2 4 

Chg Change Index 2 

Hght Height in cm (terrest rial) 3 6 

Len Length in cm (aquatic) 3 6 

PI Perennat ion - primary 3 6 

P2 Perennation - secondary 3 6 

LF 1 Life form - primary 3 6 

LF2 Life form - secondary 3 6 

Woodiness 3 6 

Clone 1 Clonal spread .- primary 4 7 

Clone 2 Clonal spread - secondary 4 7 

E I Major Biome (European di stribution) 5 9 

E2 Eastern limit code 5 9 

C' Continentality in Europe 5 9 

O rigin Origin of alien taxa 6 10 

GB Number of 10-km squares in Britain (including Isle of Man) 7 1 I 

IR Number of I 0-km squares in Ireland 7 I I 

CI Number of 10-km squares in Channel Islands 

7 I I 

Tjan January mean temperature 7 I I 

Tjul July mean temperature 7 I I 

Prec Annual precipitation 7 

Co Coastal 8 13 

Br Habitats Broad Habitats 8 13 

Ellenberg indicator value - light 9 14 

Ellenberg indicator value - moisture 10 15 

ft Ellenli erg indicator value - reaction I I 16 

Ellenberg indicator value - nit rogen 12 16 

Ellenberg indicator value - salt tolerance 13 17 

18

Taxo n name Fa m NS CS RS C hg Hght Len 1 P2 LF 1 LF2 W lo ne l C lo ne 2 E l E2 C Ori in G B IR C I r an r ul Prec Co Br Habitats L F R N S 

Acae na no vae-se landiae Rosa A N 11 C h ode2 A us , NZ 82 9 3 .8 15 .2 83 1 3 , 10 , 18 , 19 8 3 6 3 

Acer campestre Acer N n 0 .35 15 00 Ph 7 3 1389 0 3 .6 15 .7 839 1, 3 5 5 7 6 

A cer plata noides Acer A N 3000 Ph 7 3 c Eur 14 19 43 3 .4 15 .2 903 1, 3, 17 4 5 7 7 ' 

Ace r pseu dop latanus A cer A N -0 .40 3000 Ph 7 3 c Eur 2 59 9 942 1 3 .6 14 .6 1083 1, 3, 17 4 5 6 6 

Ace ras anthm pop hor um O rch N s -0 .76 40 G n 9 1 109 0 3 .6 16 .3 690 7 7 1 4 8 3 

Achillea millefolium A ste N n 0.29 45 C h h hiz2 5 5 277 4 968 1 3 .5 14 .5 110 3 6 7 5 6 4 

Ach illea ptarmica A ste N n -0.6 5 60 hc h hiz l 5 5 2 38 2 492 3 .3 14 .3 1146 11 7 7 5 3 

Acon itu m nap ell us Ranu NA 5 100 G n h hiz t 7 3 104 0 4 .0 15 .8 964 1, 3 . 14 , 17 5 7 7 6 

Acon iturn nap ellus se ns . lat. Ranu NA s 1.4 2 100 G n h hiz l 7 3 104 0 4 .0 15 .8 964 1, 3 , 14 , 17 5 7 7 6 

Acor us calam us A ra c A N 0.69 112 Hy h hiz2 Unk 5 15 17 3 .6 15 .8 770 13 , 14 8 10 7 71 

Actaea spica ta Ranu N s -0.44 60 G n h hiz l 4 6 33 0 2 .5 14 .3 10 54 16 3 5 8 6 

Adiantum cap illus- ven eris A dia N 5 0.54 30 hc 9 1 38 29 5 .2 15 .2 110 1 16 4 7 8 3 

Adonis annua Ranu A R V U -2.19 40 Th 9 1 234 3 3 .8 16 .0 745 4 7 4 7 4 

Adox a m osehatellin a Adox N n -0 .05 12 Gn h hiz l 5 6 1720 2 3 .3 14 .9 1005 1 4 5 6 5 

Aegopodium dagraria A pia A R -0 .45 100 hc h hiz2 7 4 Eur, A s 1 2533 8 19 1 3 .6 14 .6 1064 3, 17 6 5 6 71 

Aes culus hippoca stanum Hip p A N 1.0 8 32 00 Ph Eur 2 186 557 I 3.6 14 .8 10 14 3 , 17 5 5 7 7 

Aethusa cynap ium A pia NA n -0 .4 1 100 T h 7 3 164 0 0 3.6 15 .5 863 3, 4 , 17 6 4 7 6 

Agrimo nia eup atoria Ro sa N n -0 .89 60 hc 8 4 1859 532 3.8 15 .1 953 6 7 4 7 4 

Agrimo nia proc ere Ro sa N n -0 .38 100 hc 7 3 8 19 16 1 3.9 15 .1 10 26 1, 3 , 6 5 6 7 5 

Agros tem ma githago Cary A R EW -0 .75 10 0 T h Unk 8 15 96 3 .9 15 .6 852 4 7 5 6 5 

Agros tis can ine Po ac N n 60 hc to12 5 6 1609 36 1 3 .5 14 .5 1128 11, 13 7 7 3 3 

Agros tis can ine sen s.lat. Poac N n 1.3 2 60 hc iz2 St o12 5 6 19 37 44 3 3 .4 14 .3 114 5 8 , 11 7 6 3 3 

Agros tis cap illaris Poet N n 1.2 8 62 hc iz2 5 4 2758 922 1 3 .5 14 .5 1104 8 6 5 4 4 

Agro stis curtisii Poac N n -0 .26 60 he 8 1 207 0 4 .9 15 .8 1082 8 , 10 7 6 2 1 

Agros tis gigantea Poac A R 1.39 80 hc iz2 8 5 16 13 109 3 .6 15 .3 889 3, 4 7 6 6 71 

Agros tis stolonifera Poac N n 3 .66 4 5 hc to12 6 6 2 77 6 977 1 3 .6 14 .5 110 1 4, 6 7 6 7 6 

Agrostis vinea lis Poac N n 60 he iz2 7 3 1126 150 3 .3 14 .0 1200 8, 10 7 6 3 2 

Aire ca ryop hy llea Poa c N n -0 .52 25 Th 8 3 2046 6 56 1 3 .7 14 .6 1087 10 , 16 8 2 5 2 

Aire praeco x Po ac N n -0.19 10 Th 8 2 2450 6 59 1 3 .5 14 .3 1149 8 , 16 8 2 4 2 

Aj uga cha maep itys La mi NA V U r -0 .62 20 hc 8 3 43 0 3 .8 16 .5 7 13 4 , 7 71 4 8 2 

Aj uga py ramid alis La rni N 5 -0 .34 30 hc 4 3 106 8 3 .3 12 .6 1425 10 , 16 7 ' 5 5 2 

Aj uga reptans La m i N n -0 .56 30 hc to 12 7 3 2439 7 17 3 .4 14 .6 1095 1 5 7 5 5 

Alchem illa ac utiloba Ro sa N r 60 hc 5 3 c 15 0 1.6 13 .2 969 6 7 4 6 5 

Alch emilla alpina Ro sa N n -0 .6 1 15 hc 1 3 384 4 1.6 12 .1 1843 7 , 15 , 16 7 5 4 3 

Alch emilla filica ulis Ro sa N n 30 hc 4 3 1407 308 2 .9 14 .1 1185 7, 15 . 16 8 6 6 3 

Alch emilla glabra Ro sa N n 40 hc 5 3 127 1 223 2.6 13 .4 1338 6 , 15 . 16 7 6 6 4 

Alch emilla gla uce scens Ro sa N s 15 hc 5 3 c 20 3 2.0 13 .3 14 08 7 T 5 7 5 

Alch emilla glomerulans Ro sa N s 30 hc 2 3 57 0 0 .4 11.5 19 78 8 , 15 7 5 5 4 

Alchem illa micans . Ro sa N V U r 40 hc 5 3 c 4 0 2 .0 13 .5 10 56 7 7 5 7 5 

Alchem illa min ima Ro sa N E V U r 5 hc 4 1 3 0 1.7 13 .5 166 7 7 71 6 8 3 

Alchem illa mollis Rosa A N 60 hc Eur 805 17 3.5 15 .0 10 09 3 , 17 6 5 7 6 

Alchem illa monticola Rosa N r 35 hc 5 3 c 9 0 1.3 13 .0 10 84 6 7 4 6 4 

Alchem illa sub crena ta Rosa N EN r 35 hc 5 3 c 2 0 0 .9 12 .6 1125 6 7 4 6 5 

Alchem illa vulgaris agg . Rosa N n -0 .0 1 35 hc 5 3 19 35 496 3.1 14 .1 1179 6 , 7 , 15 , 16 T 5 6 4 

Alchemilla wichu rae Rosa N s 20 hc 4 3 6 5 0 0 .9 12 .0 1992 7 71 5 5 3 

Alchemilla xa nthoc hlora Rosa N n 40 . hc 7 3 1060 24 0 2 .8 13 .9 1174 6 , 7 6 5 6 4

Tax on name Fam NS CS RS Chg. Hght Len P1 P2 LF 1 LF2 W Clona l Clo ne2 El E2

S 

Taxon name Fam NS CS RS Chg Hght Len 1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tja n Tjul Prec Co Br Habitats L F R N 

Anthoxanthum aristatum Poac AN -2.65 40 Th 0 Eur 93 0 I 3.9 16.0 775 4, 17 7 4 4 5 

Anthoxanthum odoratum Poac N n 0.90 50 hc 0 6 4 2782 965 14 3.5 14.4 1106 6 7 6 4 3 

Anthriscus caucalis Apia N n -0.16 70 Th 0 7 3 659 471 12 3.9 15.71 747 3, 4, 8 71 5 6 5 

Anthriscus sylvestris Apia N n -0.19 100 hc 0 5 5 2480 805 5 3.6 14.6 1039 3 6 5 7 T 

Anthylris vulneraria Faba N n 0.45 60 he 0 5 3 1798 465 9 3.7 14.6 1030 7 8 4 7 2 

Anfi rrhinum maj us Scro AN 2.84 30 Ch 0 Eur 1043 84 8 3.9 15.7' 837 3, 17 8 3 7' 5 

Apera interrupta Poac AN 0.80 40 Th 0 8 4 Eur, As l 104 0 0 3.4 15.9 671 3, 4, 16 9 5 8 6 

Apera spica-venti Poac AR -0.21 100 Th 0 5 4 326 1 3 3.7 16.0 726 3, 4 7i 4 5 5 

Aphanes arvensis Rosa N n 10 Th 0 7 3 1608 432 4 3.7 15.0 956 4, 16 8 4 6 4 

Aphanes arvensis agg. Rosa N n -0.32 10 Th 0 7 3 2302 659 14 3.6 14.7 1038 4, 8, 16 7 4 6 4 

Aphanes australis Rosa N n 10 Th 0 7 3 1549 230 14 3.5 14.6 1070 8, 16 7 4 5 4 

Apium grayeolens Apia N n -0.63 80 hc 0 8 4 519 82 8 4.3 15.8 849 13 8 8 7 7 

Apium inundatum Apia N n -0.54 30 so Hy Irreg 7 2 9271 351 6 3.8 14.8 1023 11, 13 71 10 6 4 

Apium nodiflorum Apia N n -0.31 60 100 Hy Irreg 8 4 1661 855 12 4.0 15.2 961 14 7 10 7 7' 

Apium repens Apia N CR r 15 hc Node2 7 3 3 0 0 3.6 16.4 651 13 9 9 7 7 

Aquilegia vulgaris Ranu N n 1.70 100 hc 0 7 3 1504 197 5 3.6 15.0 986 1, 3, 16 6 4 6 5 

Arabidopsis thaliana Bras N n 1.21 30 Th 0 7 4 2218 509 13 3.5 14.8 1034 16, 17 8 3 6 2 

Arabis alpina Bras N EN r 15 Ch 0 1 4 1 0 0 2.5 11.8 3218 16 7 5 71 3 

Arabis glabra Bras N VU s -1.16 100 hc 0 7 4 151 0 1 3.5 16.0 686 8 7 3 8 5 

Arabis hirsuta Bras N n -1.02 40 p he 0 5 6 1042 143 4 3.3 14.6 1104 7, 16 7 5 8 3 

Arabis petraea Bras N s -0.64 25 Ch 0 1 5 78 2 0 1.7 11.9 1966 15, 16 9 3 8 1 

Arabis scabra Bras N VU r 20 Ch 0 9 3 1 0 0 4.4 16.5 844 16 7 3 8 2 

Arbutus unedo Eric N o 1.18 500 Ph w O 9 1 0 8 0 4.7 14.4 1335 1, 16 6 5 71 2 

Arctium lappa Aste AR 0.51 150 hc h 0 7 4 971 4 1 3.8 16.0 769 3 9 5 7 9 

Arctium minus Aste N n -0.41 150 hc h 0 7 5 2424 846 14 3.7 14.7 1051 3 6 4 7 5 

Arctostaphybos alpinus Eric N s -0.22 20 Ch w Node2 1 6 134 0 0 1.6 11.6 1750 15 7 6 2 2 

Arctostaphylos uva-ursi Eric N n -0.75 20 Ch w Node2 4 6 473 37 0 2.1 12.3 1615 10, 15 7 5 2 2 

Arenaria ciliata Cary N o 6 Ch 0 1 3 0 1 0 2.6 12.9 1454 15 9 5 8 2 

Arenaria norvegica Cary N r 0.21 6 a Ch Th 0 1 3 16 1 0 2.5 12.3 1904 16 9 3 8 2 

Arenaria serpyllifolia Cary N n -0.76 30 Th 0 8 4 2147 522 13 3.6 14.9 986 16 8 3 T 5 

Armeria arenaria Plum N o 20 Ch 0 8 2 0 0 5 6.1 16.9 843 Co 18, 19 8 3 6 2 

Armada maritima Plum N n -0. 14 15 Ch 0 3 6 1137 319 14 3.9 14.1 1265 Co 18, 21 8 7 5 5 

Armoracia rusticana Bras AR 0.05 150 hc Rhiz2 Crop 1532 111 10 3.8 15.6 850 3 8 5 7 Tj 

Arnoseris minima Aste AR EX -3.72 30 Th 0 7 3 83 0 0 3.6 16.1 685 4 7 4 3 3 

Arrhenatherurn elatius Poac N n 0.37 150 hc 0 DRg 7 3 2678 958 13 3.6 14.5 1089 3, 6 7 5 7 7 

Artemisia absinthium Aste AR -0.46 90 Ch sw 0 7 4 1005 41 6 3.8 15.6 861 3, 16, 17 7 4 7 9 

Art emisia campestris Aste N EN r -0,42 60 Ch sw Rhizl 7 4 c 9 0 0 3.3 16. 1 620 3, 8 8 3 6 5 

Artemisia norvegica Aste N VU r 8 hc h 0 1 3 3 0 0 0.9 10.9 1982 15 9 4 4 1 

Artemisia vulgaris Aste AR -0.20 150 hc h 0 7 4 2109 489 12 3.7 14.9 984 3, 17 7 4 8 7 

Arum italicum Arac N s 2.09 60 Gn h Rhizl 9 1 39 0 8 5.5 16.2 962 1, 3 4 5 6 6 

Arum maculatum Arae N n -0.28 50 Gn h Rhizl 7 3 1604 650 11 3.9 15.3 931 1 4 5 7 7' 

Asparagus of icinalis Lili N n 1.78 150 Gn h 0 7 1 17 7 7 5.4 15.9 953 3, 18, 19 7 5 6 5 

Asparagus of icinalis 

subsp.officinalis 

Lili AR 150 Gn h 0 7 4 5871 7 4 3.8 16.2 713 3 7j 5 6 5, 

Asparagus ofi cinalis 

I 

subsp.prostratus Uh N VU r 30 Gn 0 7 1 171 7 7 5.9 15.9 950 18, 19 8 4 6 3

S 

Taxon name Fam NS CS RS Chg li ght Len 1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N 

Asperula cynanchica Rubi N n -0.47 50 hc Rhiz2 7 3 329 47 3 4.0 15.9 842 7 7 8 2 

Asp leniurn adianturn-nigrum Asp N n 45 hc 0 7 3 2146 708 14 3.7 14.6 1110 3, 16 6 5 5 

Asp lenium adianturn-nigrum 

sens.lat Asp N n 0.35 45 hc 0 8 3 2146 708 14 3.7 14.6 1110 3 6 5 5 

Asp lenium marinum Asp N n 0.02 35 hc 0 8 2 530 172 12 4.4 14.0 1261 Co 18 9 5 5 

Asp leniurn obovatum Asp N s -0.18 30 hc 0 9 1 135 19 11 5.3 15.4 1197 16 5 4 3 

Asp lenium onopteris Asp N o 45 hc 0 9 1 0 34 0 4.6 14.5 1200 1, 3, 16 5 7 3 

Asp leniurn ruta-muraria Asp N n 0.15 12 hc 0 7 6 2184 898 11 3.7 14.7 1074 3, 16, 17 7 7 2 

Asp leniurn septentrionale Asp N s -0.08 15 hc 0 7 3 55 2 0 2.5 13.2 1825 16 8 2 2 

Aspleniurn trichomanes Asp N n 0.07 35 hc 0 8 6 2294 911 12 3.6 14.5 1121 3, 16, 17 5 8 2 

Asplenium viride Asp N n -0.21 20 hc 0 4 6 435 43 0 2.0 12.8 1701 15, 16 4 8 3 

Aster (alien N American taxa) Aste AN 150 hc Rhizl Am 1211 53 5 3.7 15.3 934 3, 17, 21 7 7 6 

Aster lanceolatus Aste AN 120 hc Rhizl Am6 211 5 0 3.9 15.8 844 3, 17 7 7 6 

Aster lanceolatus x novi-belgii 

(A. x salignus) Aste AN 150 hc Rhizl Gard 373 9 0 3.8 15.6 890 3, 17 7 7 6 

Aster linosyris Aste N r -0.10 50 hc 0 7 3 c 9 0 0 4.9 15.8 1006 Co 18 8 8 1 

Aster novae-angliae Aste AN 200 hc Rhizl Am6 83 0 1 3.9 16.1 806 3 7 7' 6 

Aster novi-belgii Aste AN 150 hc Rhir l Am6 557 12 3 3.7 15.2 936 3, 17 7 7 6 

Aster tripolium Aste N n -0.44 100 hc 0 7 5 703 267' 3 4.3 14,8 1109 Co 21 9 7 6 

Astragalus alpinus Faba N VU r 30 hc 0 1 6 4 0 0 -0.9 11.0 1275 7, 15 9 6 2 

Astragalus danicus Faba N n -0.88 30 hc 0 7 6 c 242 3 0 3.3 14.8 745 7 8 8 2 

Astragalus glycyphyllos Faba N n -0.36 100 hc 0 7 3 357 0 0 3.4 15.7 711 1, 3, / 6 6 7 3 

NJ 

1‘ ) Athyrium distentifolium Wood N s 0.38 30 hc 0 1 6 98 0 0 0.3 11.2 2098 15 6 3 4 

Athyrium filix-femina Wood N n 0.25 90 hc 0 5 6 2574 849 12 3.5 14.4 1132 1, 16 5 5 6 

Athyriurn flexile Wood NE VU r 20 hc 0 1 1 16 0 0 -0.9 10.7 1753 15, 16 7 3 4 

Atriplex glabriuscula Chen N n -0.93 33 Th 0 5 2 780 159 10 4.2 14.4 1163 Co 19 9 7 8 

Atriplex laciniata Chen N n 0.38 30 Th 0 7 1 418 92 13 4.5 14.8 979 Co 19 9 7 7 

Atriplex littoralis Chen N n 1.59 100 Th 0 7 6 362 36 3 4.2 15.4 822 Co 19 9 7 6 

Atriplex longipes Chen N s 90 Th 0 4 3 39 0 0 4.2 15.6 982 Co 21 9 7 8 

Atrip/ex patula Chen N n -0.34 87 Th 0 6 4 2318 787 12 3.7 14.8 1013 3, 4 7 7 7 

Atriplex pedunculata Chen N CR r 30 Th 0 7 4 20 0 0 3.9 16.4 603 Co 21 91 7 8 

Atriplex portulacoides Chen N n 0.06 80 Pn 0 9 1 322 68 10 4.6 15.7 869 Co 21 9 8 6 

Atriplex praecox Chen N s 10 Th 0 4 3 62 0 0 3.6 12.8 1503 Co 19 9 7 6 

Atriplex prostrata Chen N n 1.10 100 Th 0 6 4 1847 349 5 3.9 15.1 955 3, 4, 19, 21 8 7 7 

Atropa belladonna Sola N n -0.33 150 hc 0 7 3 414 0 0 3.6 16.1 717 1, 3 5 6 6 

Avena fatua Poac AR 1.17 150 Th 0 Eur 1556 170 5 3.71 15A 849 3, 4 71 7 7 

Avena sativa Poac AC 150 Th 0 Eur 728 131 4 3.9 15A 903 4 7 7 7 

Avena strigosa Poac AC -3.01 120 Th 0 Crop 270 72 6 4.0 14.6 1042 4 7 5 5 

Azol a filicuboides . Azol AN 2.76 1 Hy Frag Am4, SAm 660 21 9 4.0 16.0 774 13, 14 7 11 8 8 

Baldellia ranunculoides Alis N n -1.08 20 20 Hy 0 Sto12 8 2 536 344 6 4. 1 15.1 974 13, 14 8 10 6 2 

Bal ota nigra Lami AR -0.37 100 hc Rhiz2 8 3 1365 70 12 3.8 15.7 797 3 71 4 8 6 

Barbarea intermedia Bras AN 1.92 60 hc 0 Eur 735 207 7 3.7 15.0 975 4 8 5 6 7 

Barbarea stricta Bras AN 0.50 100 p hc 0 5 4 c Eur, Asl 104 1 0 3.7 15.9 705 11, 13, 14 8 7 7 8 

Barbarea verna Bras AN 1.34 90 b Th hc 0 Eur 514 28 7 4, 1 15.6 943 3 8 5 6 6 

Barbarea vulgaris Bras N n -0.02 90 p hc 0 7 4 1869 615 3 3.7 15.0 943 3, 14 7 6 7 8 

Bartsia alpina Scro N r -0.10 22 hc Rhizl 1 3 18 .".0 0 0.4 11.9 2069 7, 11, 15 5 8 71 2

S 

Taxon name Fam NS CS RS Chg Hght Len 1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tja n Tjul Prec Co Br Habitats L F R N 

Bell is pe rennis Aste N n 0.89 8 hc h Rhizl 7 3 2797 984 14 3,5 14.5 1105 6 8 5 6 4 

Berberis vulgaris Serb NA n -0 61 250 Pn w Rhizl 7 3 974 26 1 3.4 15.2 892 1, 3 7 4 8 3 

Baru la erecta Apia N n -0.02 100 Hy h Irreg 7 3 1111 265 1 3.8 15.5 843 11 7 10 7 7 

Beta vulgaris Chen N n 1.23 150 b he h 0 9 1 607 205 14 4.6 15.4 952 Co 18, 19 9 5 7 8 

Betula nana Betu N s -0.09 100 Pn w Rhizl 2 6 125 0 0 0.71 11.5 1585 10, 12, 15 7 8 1 1 

Betula pendula Betu N n -0.23 2500 Ph w o 5 4 2293 390' 0 3.3 14.6 1073 1 7 5 4 4 

Betula pubescens Betu N n 0.40 2000 Ph w 0 5 4 2399 779 0 3.4 14.4 1123 1 7 7 4 4 

Bidens cernua Aste N n -0.54 67 Th 0 7 6 874 281 2 3.9 15.5 883 13, 14 8 9 7 7 

Bide ns tripartita Aste N n -0.43 67 Th o 7 5 1055 222 5 3.9 15.5 884 11, 13 8 8 7 7 

Blacks tonia perfoliata Gent N n 0.12 45 Th 0 9 2 7871 198 2 4.0 15.8 812 7 8 5 8 2 

Blechnum spicant Blec N n -0.39 50 hc o 7 3 2159 831 9 3.5 14. 1 1198 1, 2, 10, 16 5 6 3 3 

Blysmus compressus Cype N n -1.28 37 hc Rhiz2 7 3 c 384 0 3.2 15.2 840 11 8 8 8 3 

Blysmus rufus Cype N n -0.53 37' hc Rhiz2 4 3 367 7 0 3.8 13.6 1371 Co 21 8 8 7 4 

Bo lboschoe nus maritimus Cype N n 0.00 100 Hy Rhizl DRg 8 4 766 21 7 4.3 15.1 1020 Co 21 8 10 8 7' 

Bo trychium lunaria Ophi N n -0.43 15 Gn 0 5 6 1109 13 1 3.0 13.8 1259 7, 16 8 4 6 2 

Brachy podium pinnatum Poac N n 0.15 90 he Rhizl 7 4 612 2 0 3.6 15.9 745 7 7 3 8 3 

Brachy podium sylvaticum Poac N n -0.17 95 he 0 7 3 2310 78 12 3.7 14.7 1062 1 6 5 6 5 

Brassica napus Bras AN 2.88 130 b Th hc 0 Gard 1758 14 7 3.6 15.1 914 3, 4, 17 7 4 7 7 

Brassica nigra Bras NA n -0.02 150 Th 0 7 3 1080 5 11 4.0 15.8 822 3, 4 8 5 7 6 

Brassica oleracea Bras NA s 0.90 130 Pn sw 0 8 2 98 1 4.9 15.6 922 17, 18 8 4 71 8 

Brassica rapa Bras AR 0.74 100 b Th hc 0 7 4 1407 63 7 3.9 15.0 993 4 7 5 7 6 

Briza maxima Poac AN 62 Th 0 0 3 Eur 239 11 4.6 15.8 932 3, 17, 19 7 3 4 2 

Briza media Poac N n 0.75 62 he Rhizl 7 3 1853 58 2 3.5 14.9 971 7 8 5 7 3 

Briza minor Poac AR 0.28 55 Th 0 9 1 92 8 5.2 16. 1 919 4 7 4 5 5 

Bromopsis benekenii Poac N s 0.25 105 hc 0 7 3 c 64 0 3.0 15.3 896 1 5 5 7 5 

Bromopsis erecta Poac N n -0.01 110 hc 0 7 3 798 4 3 3.7 15.9 764 7 7 4 5 3 

Bromopsis inermis Poac AN 1.71 150 hc Rhiz2 7 4 Eur, Asl 263 0 3.6 15.9 757 3 8 4 8 5 

Bromopsis ramosa Poac N n -0.18 170 hc 0 7 3 1887 45 0 3.5 14.9 995 1 4 6 7 7 

Bromus commutatus Poac N n 1.07 95 Th 0 7 3 675 3 0 3.9 16. 1 772 3 7 4 8 6 

Bromus hordeaceus Poac N n -0.37 80 Th o 8 3 2406 83 14 3.71 14.7 1051 4, 6 8 4 71 4 

Bromus racemosus Poac N n 0.74 95 Th 0 7 3 516 7 0 4.0 15.8 852 3, 6 6 6 7 8 

Bromus secalinus Poac AR -1.15 90 Th 0 Unk 403 1 4 3.9 15.8 816 4 6 4 5 4 

Bryonia dioica Cucu N n -0.50 400 Gn 0 9 2 1004 4 3.6 16.0 726 3 7 5 7 7 

Bu ddleja davidii Budd AN 3.73 500 Ph w 0 As 1434 26 14 3.9 15.4 930 3, 17 7 5 7 5 

Bu nium bulbo cas tanum Apia N r 0. 14 50 Gn 0 8 2 13 0 3.3 16.3 633 4 71 4 9 4 

Bupleururn baldense Apia N EN r 10 Th 0 9 1 3 8 6.1 16.5 808 Co 18, 19 9 3 8 2 

Bupleurum falcatum Apia AN 100 hc 0 8 5 Eur, As 8 0 3.4 16.0 776 3 6 3 9 3 

Bupleurum rotun difolium Apia AR EW -4.58 30 Th 0 Eur? 287 1 3.8 16.0 753 4 8 3 9 4 

Bupleurum tenuissimum Apia N s -0.97 50 Th 0 8 3 161 0 4.2 16.4 684 3, 6 9 7 8 4 

Butomus umbellatus Buto N n -0.04 150 150 Hy Rhiz2 DRg 7 4 685 0 3.71 16.0 715 13, 14 7i 11 71 71 

Buxus sempervirens Buxa N r 2.54 500 Ph w O 9 2 2 0 3.7 16.6 716 1, 3 4 4 8 5 

Cakile maritima Bras N n -0.38 30 Th h 0 6 3 576 131 13 4.4 14.6 1011 Co 19 9 6 7 7 

Calamagrostis canescens Poac N n -0.33 120 hc h Rhiz2 5 4 c 293 0 0 3.4 157 723 11 7 9 7 5 

Calamagrostis epigej os Poac N n 0.47 200 hc h Rhiz2 5 5 936 9 4 3.7 15.7 812 3, 11 7 7 7 6 

Calamagrostis purpurea Poac N r 150 hc h Rhiz2 4 4 10 0 0 1.6 13.1 1346 1, 11 71 8 6 3 

Calamagrostis scotica Poac NE VU r 100 hc h Rhizl 4 1 1 0 0 3.6 12.8 888 11 8 8 6 4

Taxon name Fam NS CS RS Chg_ Hght Len P1 P2 LF1 LF2 W Clonel Clone2 E1 E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N S 

Calamagrostis stricta Poac N r -0.74 100 hc Rhiz1 2 6 22 6 0 2.9 14.2 928 11 9 9 4 2 

Calendula officinalis Aste AN 50 a Th 0 Unk 764 24 10 4.0 15.7 822 3, 17 8 5 7 7 

Callitriche brutia Cal N 25 a Hz Node2 9 1 213 63 3 4.1 14.9 1087 13 8 10 5 5 

Ca Ilitriche hamulata Cal N 80 a p Hz Hy Node2 Irreg 5 2 1758 282 6 3.3 14.2 1162 13, 14 7 11 6 5 

Callitriche hamulata sens.lat Cal N 1.12 25 80 a p Hz Hy Node2 lrreg 6 3 1758 282 6 3.3 14.2 1162 11, 13, 14 7 10 5 

Callitriche hermap hm ditica Cal N 0.21 50 p Hy Irreg 4 6 392 115 0 3.1 13.7 1120 13 7 12 7 5 

Callitriche obtusangula Cal N 1.35 60 Hy Irreg 8 2 757 157 4 4.1 15.7 840 13, 14 7 11 7 6 1 

Ca llitriche platycarpa Cal N 15 100 p Hz Hy Node2 lrreg 7 3 1006 173 5 3.7 15.1 947 11, 13 6 10 7 7 0 

Callitriche stagnalis Cal N 15 60 p Hz Hy Node2 Irreg 7 3 1488 346 5 3.71 14.6 1083 13 7 10 6 6 1 

Ca & riche stagnalis sens.lat Cal N n 1.51 15 80 p Hz Hy Node2 Irreg 7 3 2647 769 11 3.5 14.4 1105 11, 13, 14 6 10 6 6 

Callitriche truncata Cal N s 0.47 20 Hz 0 Irreg 9 1 52 1 2 4. 1 16.1 719 13, 14 7 12 7 7 

Eric N n -0.64 60 Ch Pn 0 Nodel 5 3 2434 905 11 3.5 14.3 1157 10, 12 7 6 2 2

Taxon name Fan/ NS CS RS Chg Hght Len 1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N S 

Carex binervis Cype N n -0.17 90 hc 0 7 / 1927 647 3 3.3 14.0 1243 8, 10, 16 7 6 3 2 

Carex buxbaumii Cype N VU r 70 hc Rhizl 4 4 c 3 1 0 3.3 13.5 1410 11 8 8 7 2 

Carex capillaris Cype N s -0.35 40 hc Ogr 2 6 120 0 0 1.2 11.9 1636 7 9 6 8 2 

Carex caryophyllea Cype N n -0.20 15 hc Rhizl 7 4 1866 437 11 3.4 14.7 1079 7 7 4 7 2 

Carex chordorrhiza Cype N VU r 40 hc Rhiz2 2 6 c 4 0 0 1.4 11.9 1176 11 9 9 4 3 

Carex curia Cype N n 0.17 50 hc Rhizl 4 6 1190 219 0 2.7 13.6 1303 11 8 9 3 2 

Carex davalliana Cype N EX x 25 hc 0 7 3 c 1 0 0 4.0 16.3 833 11 9 9 8 2 

Carex depauperata Cype N CR r 60 hc Rhizl 9 2 9 1 0 4. 1 15.8 823 3 5 4 71 4 

Carex diandra Cype N n 0.22 60' hc Rhizl 5 6 378 294 0 3.6 14.4 1042 11 8 9 5 3 

Carex digitate Cype N s 0.04 15 hc 0 5 3 c 39 0 0 3.3 15.5 859 1, 16 5 5 8 4 

Carex dioica Cype N n -0.35 30 hc Rhizl 2 6 1100 204 0 2.7 13.2 1404 11 8 9 6 2 

Carex distans Cype N n -0.47 72 hc 0 8 3 814 213 9 4.3 15.0 1039 Co 21 8 6 7 5 

Carex disticha Cype N n -0.03 90 hc Rhiz2 7 4 12271 48 1 3 3.6 15.0 910 11 7 8 6 4 

Carex divisa Cype N s -0.35 70 hc Rhizl 9 2 162 1 4.4 16.4 727 6 8 7 7 6 

Carex divulsa Cype N n 82 hc 0 8 4 780 24 4.2 15.8 844 3, 6, 7 7' 4 7 6 

Carex echinata Cype N n -0.75 40 hc 0 5 3 2100 78 3.4 14.1 1207 11, 12, 14 8 8 3 2 

Carex elate Cype N n -0.32 95 Hy hc 0 7 5 294 22 3.8 15.2 903 11 7 10 7 5 

Carex elongate Cype N s 0.06 80 hc 0 5 4 c 72 1 3.6 15.5 899 1 5 8 6 6 

Carex erk etorum Cype N s -0.46 17 hc Rhizl 5 4 c 33 2.9 15.3 801 7 6 4 7 1 

Carex extensa Cype N n -0.23 40 hc Ogr 8 3 440 19 4.5 14.5 1228 Co 21 8 7 7 5 

Carex filiformis Cype N r 0.23 50 hc Rhiz2 7 4 c 13 3.5 16.3 708 6, 7 7 7 8 5 

NJ Carex /lam a Cype N n 0.53 50 hc Rhiz2 8 3 2706 95 3.5 14.5 1104 7, 11 7 5 6 2 

in Carex fi ava Cype N VU r 70 hc 0 5 3 1 3.2 14.7i 1585 1, 11 7i 9 8 2 

Carex hirta Cype N n 0.17 70 hc Rhiz2 7 3 1958 57 3.7 15.0 963 6 T 7 7 6 

Carex hostiana Cype N n -0.05 57 hc 0 7 3 1578 42 3.2 13.8 1290 11 8 9 6 2 

Carex humifi s Cype N s -0.01 10 hc Ogr 7 5 c 30 3.9 16. 1 845 7 8 3 8 2 

Carex lachenalii Cype N r -0.22 20 hc Rhizl 1 6 9 -1.1 10.4 1948 11, 15 8 7 4 1 

Carex laevigata Cype N n -0.01 120 hc 0 7 1 1012 29 3.6 14.4 1240 1, 16 5 8 5 4 

Carex lasiocarpa Cype N n 0.73 120 Hy hc Rhiz2 4 6 461 16 3.1 13.6 1383 11 8 10 6 3 

Carex fi mosa Cype N n 0.14 40 Hy hc Rhiz2 4 6 423 19 3.3 13.4 1431 11, 12 8 10 4 1 

Carex magellanica Cype N s -0.02 40 hc Rhizl 4 6 131 1.7 12.8 1808 12 9 9 2 1 

Carex marifi ma Cype N s -1.34 18 hc Rhiz2 1 6 83 3.6 12.9 964 Co 19 9 8 7 2 

Carex microglochin Cype N VU r 12 hc Rhiz2 1 3 1 -0.7 11.0 1855 11 9 9 8 2 

Carex montane Cype N s 0.68 35 hc Rhizl 7 3 c 48 4.0 15.5 1193 6, 8 7 6 4 1 

Carex muricata Cype N n 85 he 0 8 4 912 6 1 3.8 15.4 939 3, 7, 8 T 4 6 4 

Carex nigra Cype N n -0.01 70 hc Rhiz2 5 4 2582 87 3.5 14.3 1131 11 7 8 4 2 

Carex norvegica Cype N VU r 30 hc 0 1 6 6 -0.8 10.8 1901 15 8 7 7 2 

Carex ornithopoda Cype N r 0.28 15 hc 0 4 3 15 0 2.1 13.8 1272 7, 16 8 3 9 3 

Carex otrubae Cype N n -0. 14 100 hc 0 8 4 1636 40 9 4.0 15.3 939 11 6 8 7 7 

Carex Ova rls Cype N n -0.21 90 hc 0 5 4 2422 74 7 3.4 14.3 1138 3, 10 7 7 5 4 

Carex pallescens Cype N n -0.51 60 hc 0 5 4 1596 18 1 3.0 14.2 1236 1 6 6 5 4 

Carex panicea Cype N n -0.31 50 hc Rhiz2 5 3 2526 89 6 3.5 14.3 1139 11 8 8 4 2 

Carex paniculata 

Cype N n -0.11 150 

hc Hy 0 7 3 

1516 50 5 3.7 14.9 1036 1, 11 6 9 6 6, 

Carex pauciflora 

Cype N n -0.59 25' 

hc 

Rhizl 4 6 

377 0 1.7 12.1 1789 12 

8 9 1 1 

Carex pendula Cype N n 1.30 150 hc 0 8 3 1409 30 10 3.9 15.3 928 1, 14 5 8 7 6 

Carex pilulifera Cype N n -0.04 35 hc 0 7 3 2111 44 8 3.3 14.1 1197 8 7 5 3 2

R N 

6 6 

5 2 

7 3 

3 2 

7 5 

6 6 

7 7 

4 2 

7 2 

7 3 

6 4 

7 6 

6 5 

6 3 

5 4 

6 2 

onel Clone2 El E2 C Origin GB IR CI Tjan Tjul Pree Co Br Habitats L F 

7 

8, 

9 

8 

8 

4 

71 

8 1 

8 

8 

7 

3 

4 

7' 

8 1 

8 

754 11 

1239 11, 16 

1125 Co 18 

1541 15 

936 11 

1063 1, 14 

814 11, 14 

1193 11 

1699 15, 16 

2262 15 

823 3, 6, 7 

841 1, 14 

1050 1 

1904 15 

1072 11 

1169 11, 14, 19 

16.0 

13.9 

15.4 

10.6 

13.1 

14.8 

15.7 

14.0 

11.5 

11.3 

15.7 

15.7 

14.8 

11.3 

14.6 

14.2 

3.8 

3.3 

5.3 

-1.2 

3.1 

3.7 

3.9 

3.3 

0.7 

0.3 

3.6 

3.8 

3.6 

-0.1 

3.3 

3.5 

3 

4 

7 

0 

0 

5 

41 

0 

0 

0 

1 

0 

2 

0 

1 

9 

72 

656 

42 

0 

0 

675 

160 

731 

0 

0 

48 

102 

608 

0 

230 

855 

672 

1874 

56 

17 

4 

1879 

1186 

1924 

31 

71 

1043 

417 

1899 

83 

913 

2337 

7 4 

7 2 

8 2 

1 6 

4 1 

7 3 

7 4 

5 6 

1 6 

1 6 

7 3 

7 2 

7 5 

2 6 

5 6 

5 6 

hizl 

hizz21 h 

hiz2 

hiz2 

hIz21 h 

hiz2 

hizl 

a, 

84 2 1 

1200 11,14 

13.9 

3.1 

2 

434 

1168 

7 3 

2 0 

7 3 1 

8 6 0 

7 2 0 

s 6 0 

4 5 3 

7 6 1 

4 2 0 

s 5 0 

7 71 1 

7 3 

7 2 

7 3 

6 5 

6 5 

8 3 

6 3 

a 3 

8 3 

5 2 0 

8 

8 

7 

8 

4 

9' 

8 

7 

5 

6 

9 

a 

7 

7 

71 

8 

8 

9 

8 

9 

8 

8 

4 

s 

4 

9 

7 

1194 14 

1278 11, 19 

731 13 

947 7 

895 1 

962 Co 18, 19 

868 3, 6, 17 

1578 11, 14 

950 1 

936 13, 14 

1027 Co 18 

878 7, 17 

744 3 

836 3, 4, 17 

1093 6, 7 

632 6, 7 

980 7, 19 

1063 Co 19, 21 

841 Co 21 

1170 10 

825 Co 18 

896 3, 16, 17, 

763 1 

1166 1 

797 1 

2003 15, 16 

2178 . 15, 16 

14.2 

13.9 

16.4 

15.3 

15.4 

15.9 

15.0 

13.9 

15.2 

15.0 

15.1 

15.5 

16.1 

15.4 

14.5 

15.6 

15.1 

14.3 

16.1 

15.4 

16.5 

15.5 

16.2 

14.7 

16.2 

11.4 

11.3 

3.4 

3.8 

3.9 

3.9 

3.6 

5.7 

3.6 

3.4 

3.6 

3.8 

4.8 

4.1 

4.0 

3.8 

3.6 

3.8 

4.0 

3.8 

4.4 

5.8 

5.5 

4.0 

3.7 

3.7 

3.8 

0.4 

0.8 

6 

5 

0 

10 

9 

10 

2 

1 

12 

3 

14 

10 

3 

5 

11 

2 

14 

0 

8 

0 

2 

14 

0 

0 

0 

0 

0 

761 

179 

0 

243 

52 

8 

30 

42 

140 

304 

172 

494 

1 

56 

975 

125 

710 

3 

17 

0 

0 

357 

0 

31 

0 

0 

0 

2203 

477 

24 

1131 

1488 

79 

303 

296 

1693 

955 

463 

1216 

153 

884 

2658 

1239 

1811 

111 

457 

3 

3 

1361 

233 

131 

10 

77 

46 

c 

5 2 

5 3 

7 4 

7 4 

7 3 

SAf 

Eur?, Asl ? 

ode2 

8 1 

Eur 

5 3 

9 1 

9 2 

ode2 

Eur 

7 3 

7 2 

7 4 

8 3 

7 3 

a 4 

8 0 

9 1 

18 

Eur 

7 3 

7 3 

7 3 

1 3 

1 3 

odel 

hizl

Taxon name Fam NS CS RS Chg Hght Len P1 P2 LF1 LF2 W Clonal Clone2 El E2 C Origin GB IR CI Tja n Tjul Prec Co Br Habitats L F R N S 

Cerastium arvense Cary N n -1.05 30 p Ch Rhiz2 5 6 814 39 0 3.3 15.3 773 8 8 4 5. 3 0 

Co a st/UM brachypetalum Cary AN 30 a Th 0 8 3 Eur 2 0 0 3,4 16.3 657 3, 7 9 3 8 2 0 

Cerastiurn cerastoides Cary N s -0.05 12 p Ch Nodal 1 3 29 0 0 -0.4 10.8 1994 15 8 8 5 4 0 

Cerastiurn diffusum Cary N n 0.38 23 a Th 0 7 3 1174 247 12 4.0 14.6 1062 Co 18, 19 9 4 61 3 1 

Cerastium fontanum Cary N n 1.40 30 p Ch Rhizl Nodal 5 4 2805 985 14 3.5 14.4 1106 6 7 5 5 4 0 

Cerastium glomeratum Cary N n 1.44 30 a Th 0 8 3 2631 823 14 3.6 14.5 1079 3, 5 7 5 6 5 0 

Cerastium nigrescens Cary NE VU r 5 p Ch 0 4 1 2 0 0 3.0 11.5 1256 16 9 3 5 1 0 

Cerastiurn pumilum Cary N s -0.171 11 a Th 0 7 3 87 0 0 4,1 16.2 806 7 8 2 8 1 0 

Cerastium sernidecandrum Cary N n 0.50 14 a Th 0 7 3 1117 61 12 3.7 15.3 846 8, 18, 19 8 3 6 3 0 

Cerastium tomentosurn Cary AN 2.97 30 p Ch Rhiz2 Node2 Eur 1303 140 8 3.7 15.2 898 3, 17, 19 8 3 7 5 1 

Ceratocapnos claviculata Furna N n 0.57 75 a Th ' 0 7 1 1122 20 2 3.3 14.6 1111 1, 9 5 5 4 5 0 

Ceratochloa carinata Poac AN 2.09 80 p hc 0 Am4 183 1 2 3,8 16. 1 744 3, 4 8 5 6 6 0 

Ceratoch/oa cathartica Poac AN 0.63 100 p hc 0 Am, SAm 184 0 9 4.2 16.1 796 3 , 4 7i 4 5, 5 0 

Ceratophyllum demersum Gera N n 0.87 100 p Hy Irreg 6 6 927 62 1 3,7 15.8 770 13 7 12 7 7 

Ceratophyllum submersum Cera N n 0.39 100 p Hy Irreg 7 4 208 3 5 3,9 16.3 705 13 7 12 8 8 

Ceterach of icinarum AspI N n -0.30 20 p he 0 9 2 1093 787 9 4.0 15.0 1061 3, 16 7 3 8 1 

Chaenorhinum minus Scro AR -0.63 25 a Th 0 7 3 1468 170 0 3.6 15.3 899 3, 4, 17 8 4 7 4 

Chaerophyllum temulum Apia N n -0.64 100 b hc 0 7 3 1786 0 5 3.6 15.4 890 3 6 5 71 7i 

Chamaecyparis lawsoniana Cupr AN 4100 p Ph w 0 Arn4 827 46 4 3.6 15.3 965 17 5 5 6 4 

Chamaemelum nobile Aste N n -0.92 30 p hc h Node2 8 2 308 115 13 4.6 15.7 990 8 8 7 5 5 

Charnerion angustifolium Onag N n -0.01 150 p Gn hc h Root 5 6 2603 650 10 3.4 14.5 1075 3, 17 6 5 6 5 

Chelidonium rnaj us Papa AR -0.72 90 p he h 0 7 5 1662 220 9 3.7 15.3 905 3 6 5 8 7 

Chenopodium album Chen N n 100 a Th h 0 6 5 2340 782 14 3.7 14.8 1018 4 7 5 71 71 1 

Chenopodium album agg. Chen N n -0.73 100 a Th h 0 6 5 2340 782 14 3.7 14.8/ 1018 4 7 5 7 7 1 

Chenopodiurn bonus-henricus Chen AR -1.79 50 p hc h 0 Eur 1363 81 1 3,4 15.2 883 3 8 5 7 8 0 

Chenopodium 

chenopodioides Chen N s -0.17 30 a Th h 0 7 4 39 0 2 4.2 16.7 612 Co 6, 21 8 7 7 8 4 

Chenopodiurn ficifoliurn Chen AR 1.90 90 a Th h 0 7 4 745 11 7 3.9 16. 1 745 4 7 6 6 7 0 

Chenopodiurn glaucurn Chen AR -1.32 50 a Th h 0 7 6 157 2 2 4.0 16. 1 724 17 8 6, 7i 9 3 

Chenopodiurn hybridum Chen AR -0.32 100 a Th h 0 7 6 285 0 1 3.7 16.2 695 4 7 4 7 7 0 

Chenopodiurn murale Chen AR -1.63 100 a Th h 0 8 4 412 8 13 4.2 16.0 797 4 8 6 6 7 0 

Chenopodiurn polyspermum Chen AR 0.62 50 a Th h 0 7 4 998 8 12 3.9 16.0 794 4 7 6 7 8 0 

Chenopodiurn rubrum Chen N n' 1.00 70 a Th h 0 7 4 1267 142 8 3.8 15.7 800 4, 11 7 7 7 8 1 

Chenopodiurn urbicurn Chen AR 1 -4.57 100 a Th h 0 7 4 239 1 0 4.0 16.1 762 4 71 5 71 7t 

Chenopodiurn Chen AR VU -2.60 35 a Th h 0 8 4 180 0 6 4. 1 16.1 752 Co 15, 19 7 4 7 9 

Chrysanthemum segetum Aste AR -1.80 60 a Th h 0 8 3 1682 471 10 3.8 14.8 1022 4 7 5 6 5 

Chrysospleniurn alternifolium Saxi N n 0.62 20 p he h Sto12 3 6 790 0 0 2.8 14.4 1042 11, 14 5 8 6 6 

Chrysosplenium 

oppositifolium Saxi N n -0.36 15 p Ch hc h Node2 7 2 2067 688 6 3.4 14.3 1168 1, 14 5 9 5 5 

Cicendia filiformis Gent N s -0.70 10 a Th h 0 9 2 66 371 5 5.4 15.6 1063 10 9, 8 3 2 

Cicerbita alpina Aste N VU r 130 p hc h Rhiz2 4 3 4 0 0 -1.3 10.5 1375 15, 16 7 6 6 6 

Cicerbita rnacrophylla Aste AN 175 p hc h Rhiz2 Eur 724 42 0 3.3 14.9 920 3, 17 7 5 6 6 

Cichorium intybus Aste AR . -1.27' 110 p hc h 0 8 4 1312 60 10 3.8 15.5 840 3 5 4 7 5 

Cicuta virosa Apia N s 0.55 150 p Hy h 0 5 5 c 139 139 0 3.5 14.8 926 1/ , 13, 14 7 10 7 5 

Circaea alpina Onag N 5 . 30 p Gn h Rhiz2 4 6 40 0 0 2.2 13.0 1836 1 4 71 5; 5 

tJ

Taxon name Fam NS CS RS Ch Hght Len P P2 LF1 LF2 W Clonal Clone2 El E2 C Origin GB IR CI T an T'ul Prec Co Br Habitats L F R N S 

Circaea alpina x lutetiana (C 

x intermedia) Onag NH n 0.48 45 Gn Rhiz2 7 3 570 49 0 2.7 13.4 1477 1 6 6 

Circaea lutetiana Onag N n -0.38 60 Gn Rhiz2 7 3 2053 748 8 3.7 14,9 1041 1 6 6 

Cirsium acaule Aste N n -0.52 15 hc Rhial 7 3 734 0 3 3.7 16.1 742 7 4 3 

Cirsium arvense Aste N n 0.47 120 Gn Root 7 5 2736 968 14 3.6 14.5 1092 3, 4, 6 6 6 

Cirsium dissectum Aste N n -0. 14 60 hc Rhizl 7 1 539 553 2 4.0 15.0 1043 11 8 2 

Cirsium eriophorum Aste N n -0.08 150 hc 0 7 3 447 0 0 3.6 16.0 745 7 4 5 

Cirsium heterophyllum Aste N n -0.44 120 he Rhizl 4 4 761 1 0 2.0 12.9 1476 6, 16 6 5 

Cirsium palustre Aste N n 0.15 175 hc 0 5 4 2697 95 12 3.5 14.5 1105 11, 14 8 4 

Cirsium tuberosurn Aste N VU r 0.41 80 hc o 8 2 17 0 3.9 16.1 882 7 6 3 

Cirsium vulgare Aste N n 0.80 150 hc 0 7 4 2789 98 14 3.5 14.5 1103 3, 5, 6, 7 5 6 

Cladium mariscus Cype N n 0.11 200 Hy Gn Rhir l 8 4 232 25 1 4.1 14.7 1085 11 10 4 

Claytonia perfoliata Port AN 0.50 30 a Th 0 Am4 704 10 3.6 15.6 768 4, 17, / 9 6 5 

Claytonia sibirica Port AN 1.28 40 a p Th hc 0 As2, Am4 1166 2 6 3.2 14.4 1092 1, 3 7' 6 

Clematis vitalba Ranu N n 0.00 3000 p Ph 0 7 3 954 0 4.0 16.0 815 3 4 5 

Clinopodium acinos Lami N n -1.59 22 a Th 0 7 3 551 0 3.6 15.8 778 16 2 1 

Clinopodium ascendens Lami N n 0.04 60 P hc Rhizl 7 3 657 6 2 4.2 15.9 854 3, 16 5 6 

Clinopodium calamintha Lami N s -0.31 60 P hc Rhiz2 9 2 129 4 3.7 16.4 641 3 3 3 

Clinopodium menthifolium Lami N EN r 60 p hc Rhizl 7 3 1 0 4.8 16.3 844 1 5 5 

Clinopodium vulgare Lami N n -0.67 77 P hc Rhizl 7 6 1317 2 3.5 15.4 880 7 4 4 

Cochlearra anglica Bras N n 0.02 40 b hc 0 7 1 297 9 0 4.5 15.6 917 Co 21 8 6 

Cochlearia atlantica Bras NE DD r 20 

t.. ) 

P he 0 4 1 2 0 4.4 12.8 1830 Co 18 6 5 

CO Cochlearia danica Bras N n 3.31 25 a Th 0 7 1 588 13 14 4.5 14.9 1056 Co 3, 18 6 5 

Cochlearia micacea Bras NE s 10 b p hc 0 1 1 32 0 0.2 11.4 2221 15 2 

Cochlearia officinalis Bras N n 30 P hc 0 3 6 1051 27 2 3.9 14.0 1246 Co 18, 21 6 5 

Cochlear la officinalis sens.lat Bras N n -0.18 30 b p hc 0 3 6 1245 30 3 3.7 13.9 1287 18 7 i 4 

Cochleada pyrenaica Bras N s 30 b p hc 0 1 3 124 0 1.5 12.9 1567 11, 16 7 3 

Coeloglossum viride Orch N n -1.34 22 P Gn 0 4 6 964 21 0 3.2 14.1 1148 7 4 2 

Coincya monensis Bras N s 0.43 50 b hc 0 8 2 61 3 4.2 14.8 1037 3, 17, 19 4 3 

Coincya wrightii Bras NE VU r 90 P Ch hc 0 7 1 1 0 5.8 15.7 986 Co 18 4 3 

Colchicum autumnale Li li N n -0. 14 35 p Gn Dtb 7 3 301 1 0 3.6 15.8 809 6 6 4 

Colutea arborescens Faba AN 400 p Ph 0 Eur 166 0 3.7 16.3 671 3, 17 4 3 

Conium maculatum Apia AR -0.02 250 b hc 0 8 4 1847 47 13 3.9 15.2 915 3 8 

Conopodium maj us Apia N n -0.19 40 a Gn 0 7 1 2520 75 7 3.5 14.5 1102 1, 6 5 5 

Consolida aj acis Ranu AN 60 a Th 0 Eur 360 7 3.8 16.2 703 3, 4, 17 4 4 

Convallaria maj alis Li li N n 0,25 25 P Gn Rhiz2 5 3 439 0 3.3 15.5 831 1, 7 5 5 

Convolvulus arvensis Cony N n -0.70 100 P Gn Rhiz2 8 4 1841 43 14 3.8 15.2 929 3, 4 4 6 

Conyza canadensis Aste AN 1.12 100 a Th 0 Am 1048 11 10 3.8 16.0 766 3, 4, 17, 19 4 6 

Corallorhiza trifida Orch N s 0.61 22 p Gn 0 4 6 c 102 0 0 2.5 13.5 956 1, 2 5 4 

Coriandrum sativum Apia AN 50 a Th 0 Eur? 201 2 3 3.9 16.0 763 3, 17 4 5 

Cornus sanguinea Corn N n -0.06 400 P Ph Root 7 3 1179 52 1 3.7 15.8 810 1, 3 5 6 

Cornus sericea Corn AN 300 p Pn Ph 0 Nodal Am 421 111 0 3.5 15.0 936 1, 3, 17 7 6 

Cornus suecica Corn N n -0.42 20 p hc Rhiz2 2 3 218 0 0 0.9 11.6 1853 10, 15 6 1 2 

Coronopus didymus Bras AN 1.77 18 a Th 0 Unk 1284 315 14 4.1 15.5 921 4, 17 5 6 7 

Coronopus squamatus Bras AR 0.33 25 a Th 0 8 3 1290 149 9 4.0 15.7 821 3, 4 5 7 7 

Corrigiola litoralis Cary N CR r -0.96 25 a Th 0 8 2 2 0 0 6.2 16.0 1052 Co 19 7 5 5

S 

, 

Taxon name Fam NS CS RS Chg Hght Len P1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N 

Corylus avellana Beth N n -0.54 600 p Ph w 0 7 3 2470 870 5 3.5 14.6 1094 1, 3 4 5 6 6 

Cory nephorus canes cens Poac N r 0.01 32 P hc h 0 8 3 22 0 3 4. 1 16.2 649 Co 19 9 1 3 1 

Cotoneaster bullatus Rosa AN 400 p Ph w 0 As l 237 10 0 3.7 15.3 965 1, 3, 16, 17 7 4 7 4 

Cotoneaster cambricus Rosa AN EN 150 p Pn vv 0 7 1 Eur 1 0 0 5,4 15.5 796 7, 16 8 3 7 2 

Cotoneaster horiz ontalis Rosa AN 100 p Pn w 0 As 1 855 63 2 3.8 15.5 933 3, 16, 17 8 3 8 4 

Cotoneaster integrifolius Rosa AN 100 p Pn w 0 As l 537 103 2 3.9 14.7 1194 3, 17 7 3 7 4 

Cotoneaster microphyllus 

agg. Rosa AN 1.54 100 Pn w 0 Asl 576 135 3 3.9 14.7 1174 3, 16, 17 7 3 7 4 

Cotoneaster simonsii Rosa AN 3.55 300 Pn Ph w 0 Asl 973 122 5 3.7 14.9 1086 3, 10, 16, 17 6 5 6 4 

Crambe maritima Bras N n 0.29 62 hc h 0 7 3 25 1 46 11 4.8 15.5 938 Co 19 9 5 8 7 

Crassula aquatica Cras NA VU 8 a Hz h 0 Nodel 7 6 c 2 0 0 3.2 14.0 1459 14 8 9 5 5 

Crassula helmsii Cras AN 10 30 p Hy Ch h Irreg Node2 Aus, NZ 604 8 8 3.8 15.8 826 13, 14 7 10 6 7 

Crassula tillaea Cras N $ 0.86 5 a Th h 0 9 2 104 0 11 4.1 16.2 734 3 8 7 4 2 

Crataegus laevigata Rosa N n 0.32 1000 Ph w 0 7 3 597 0 1 3.5 16.2 674 1 5 5 7 5 

Crataegus m onogyna Rosa N n -0.76 1000 Ph w 0 7 3 2496 946 13 3.6 14.7 1073 1, 3 6 5 7 6 

Crepis biennis Aste N n -0.02 120 hc 0 7 3 288 th 0 3.7 16.3 711 3, 6 8 5 7 6 

Crepis capillaris Aste N n -0.17 75 a b Th hc 0 7 3 2525 907 14 3.6 14.6 1074 7 7 4 7 4 

Crepis foetida Aste AR EN 60 a hc Th 0 8 4 33 0 2 4.0 16.4 707 Co 19 9 4 6 3 

Crepis morns Aste N r -1.20 60 hc 0 7 3 c 75 0 0 1.7 13.2 1079 7 8 5 7 5 

Crepis paludosa Aste N n -0.27 80 hc 0 5 3 1162 357 0 2.7' 13.4 1334 11, 16 6 7 6 4 

Crepis praemorsa Aste N EN 60 hc 0 7 4 c 1 0 0 1.71 13.3 1333 7 8 3 9 3 

Crepis vesicaria Aste AN 0.60 80 hc 0 Eur 1227 302 14 4.1 15.7 858 3, 5, 6, 17 8 5 7 7 

Crithmurn maritimum Apia N n 0.23 45 hc 0 9 1 301 138 14 5.1 15.4 1042 Co 18 9 6 7 5 

Crocosmia aurea x poli sh (C. 

x crocosmiiflora) Irid AN 3.11 60 Gn Rhizl ORg Gard 1446 731 12 4.0 14.6 1129 1, 3, 17 7 6 4 4 

Cruciata laevipes Rubi N n -0.77 60 hc Rhiz2 7 4 1475 0 1 3.2 15.2 892 6 6 5 7 5 

Cryptogramma crispa Adia N n -0.63 15 hc 0 4 3 466 23 1.8 12.9 1678 10, 15, 16 7 5 2 3 

Cuscuta epithyrnum Cuse N n -1.28 1 a Th Sto12 8 4 494 27 1 4.3 15.9 846 10 7 6 2 2 

Cuscuta europaea Cusc N s 0.04 2 a Th Sto12 7 4 126 0 3.7 16.4 679 14 6 7 6 7 

Cymbalaria muralis Scro AN 

-0.10 8, 

Ch 

Node2 

Eur 2059 619 1 3.7 15.0 990 3, 16, 17, 19 7 5 7 6 

Cynodon dactylon 

Poac NA VU r -0.10 30 

hc 

Rh1z2 Node2 8 5 

2 0 6.4 15.6 1156 3, 5, 17 8 4 7 5 

Cynoglossum ge rmanicurn Bora N VU r -0.52 75 hc 0 7 3 c 50 0 3.71 16.5 676 1 6 5 8 71 

Cynoglossum ofi cinale Bora N n -1.09 75 hc 0 7 4 718 42 3.9 15.8 780 7, 19 8 4 8 6 

Cynosurus cristatus Poac N n 0.02 75 hc 0 7 3 2745 962 1 3.5 14.5 1103 6 7 5 6 4 

Cyperus fuscus Cype N VU r -0.32 20 a Th 0 8 4 11 0 4.4 16.6 758 13 9 8 5 4 

Cyperus longus Cype N s 2.22 100 he Rhizl 8 3 33 0 1 5.7 16.1 943 11, 13 8 9 7 5 

Cypripedium calceolus Orch N CR r 30 Gn Rhizl 5 6 c 22 0 2.2 13.9 1270 7 5 4 8 4 

Cystopteris dickie ana Wood N VU r 20 hc 0 ? ? 5 0 0.9 12.3 1379 16 5 7 8 2 

Cystopteris fragilis Wood N n -0.69 20 hc 0 3 6 1118 191 2.6 13.5 1361 16 6 7 8 4 

Cystopteris montana Wood N r -0.25 15 hc Rhiz2 4 6 22 0 -0.2 11.3 2151 15, 16 5 7 9 2 

Cytisus seopar/us Faba N n 0.00 200 Pn w 0 7 3 2288 553 1 3.4 14.6 1085 3 8 5 4 4 

Daboe cia cantabrica Eric N o 0.12 50 Ch Pn w 0 8 1 0 33 4.5 14.4 1265 10 8 5 3 2 

Dactylis glomerata Poac N n -0.06 120 he h 0 8 4 2707 981 1 3.6 14.5 1091 6 7 5 7 6 

Dacty/orhiza fuchsii Orch N n 0.33 50 Gn h 0 7 4 2214 774 3.5 14.7 1063 11 7 8 7 3 

Dactylorhiza incarnata Orch N n -0.33 45 Gn h 0 5 4 1189 331 3.6 14.4 1127 11 8 9 6 2 

Dactylorhiza lapponica Orch N s 21 Gn h 0 4 3 15 0 3.2 12.5 1910 11 5 8 6 2

F R N S 

77 53 23 0 0 

6 7 3 

8 7 2 0 

6 7 2 1 

10 5 3 0 

Br Habitats 

12 

11 

11 

11 

11 

13 

7, 8 

1 

1 

17 

6, 7 

17 

1, 6 

8, 10 

12, 13 

4 

3, 16 

7 

16 

15 

8, 9 

3, 4, 17 

3, 4, 17 

15 

10 

3, 4, 16, 17 

3, 16, 17 

3, 6, 17 

3, 6, 17 

1, 3, 16 

3, 18, 19 

1, 3 

16 

7, 16 

3, 16 

15, 16 

11, 12 

12, 14 

12 

7, 16 

1, 16 

1, 16 

1 

11 . 

1, 2 

1, 15, 16 

El E2 C Origin GB IR CI Tjan Tjul Prec Co 

1221 

1237 

849 

1244 

978 

707 

1160 

760 

890 

796 

989 

913 

1099 

1177 

1211 

738 

805 

797 

979 

2845 

1120 

777 

783 

1686 

1465 

810 

790 

863 

863 

729 

898 Co 

921 

1191 

1584 

1151 

2311 

1422 

1337 

1269 

1734 

1373 

1160 

1075 

659 

1114 

1772 

W Clonel Clone2 

h 0 

h 0 

h 0 

h 0 

h 0 

h 0 

h 0 

w 0 

w 0 

h 0 

h 0 

h 0 

h 0 DRi 

h Ogr 

h 0 

h 0 

LF2 

65 74 2 5 0 

5 7 6 0 

4 7 8 0 

4 71 3 1 

65 75 8 4 

5 2 3 

9 2 1 

4 7i 6 

5 3 5 3 2 

2 7 1 

3 4 1 

6 4 5 

4 5 5 

4 5 5 

5 2 2 

4 1 2 

4 71 6 

h 0 

0 

0 

sw 

sw 

w 0 

h 0 

h 0 

h 0 

Taxon name Fam NS CS RS Chg Hght Len P1 P2 LF1 

40 p Gn 

30 p Gn 

50 p Gn 

25 p Gn 

30 p Gn 

30 a Hz 

40 p hc 

100 p Pn 

100 p Pn 

100 a Th 

100 b hc 

120 b hc 

150 p hc 

60 p hc 

70 p hc 

100 a Th 

60 a Th 

37 p Ch 

20 p Ch 

6 p Ch 

150 b hc 

35 a Th 

50 a Th 

10 p Ch 

8 p Ch 

60 a Th 

80 p Ch 

200 b hc 

200 b hc 

150 b hc 

9 p Ch 

80 p hc 

10 p Ch 

35 b hc 

50 a Th 

5 

Ch 

13 

hc 

5 

hc 

5 

hc 

10 

Ch 

60 

hc 

80 

he 

80 

hc 

60 

hc 

150 

hc 

80 

hc 

f.a 

c) 

57 77 76 

sw Rhiz2 

sw Rhiz2 

h 0 

h 0 

h 0 

h 0 

hc 

76 78 7 7 

3 4 5 

5 6 5 

4 9 3 

5 7 2 

6 7 6 

5 71 3 

99 22 1 1 

9 2 1 

4 7 2 

6 2 3 

6 5 5 

8 5 4 

9 4 4 

6 4 5 

6 3 2 

h 0 

sw Node2 

Rhizl 

0 

0 

0 

0 

0 

0 

0 

w Node2 

0 

0 

0 Rhizl 

0 

0 

0

N 

IR 

Origin 

LF2 

1 P2 

Chg 

RS 

n 

n 

n 

x 

s 

CS 

5 

5 

2 

4 

3 

8 

5 

4 

4 

5 

5 

5 

1 

4 

5 

2 

4 

2 

R 

6 5 

6 5 

5 2 

6 4 

5 9, 

5 7 

3 5 

4 7 

5 

5 

7 

9 5 

NS 

N 

N 

N 

N 

N 

AN 

AR 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N 

AN 

AN 

N 

N 

N 

N 

N 

N 

N 

AN 

AN 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N 

NE 

N 

N 

Fam 

Dry o 

Taxon name 

Dryopteris filix-mas 

0.03 

0.24 

NHem 

LF1 

hc 

hc 

he 

hc 

he 

Th 

hc 

hc 

Hz 

Hz 

Hy 

hc 

hc 

Hy 

hc 

hc 

hc 

Hy 

Hy 

Hy 

he 

he 

he 

hc 

Ch 

hc 

he 

he 

hc 

hc 

hc 

hc 

hc 

hc 

he 

hc 

hc 

Gn 

Gn 

Gn 

Gn 

Gn 

Gn 

Gn 

Gn 

Gn 

Hght Len 

120 

120 

50 

80 

60 

120 

67 

90 

5 10 

5 10 

10 

60 

35 

60 

8 

30 

60 

45 

300 

300 

110 

105 

55 

125 

30 

20 

10 

4 

75 

150 

60 

75 

75 

60 

75 

75 

75 

30 

80 

60 

45 

42 

65 

60 

22 

90 

0.10 

0.75 

0.36 

-0.24 

1.07 

0.66 

-0.11 

, 

n 

n 

s 

n 

r 

n 

n 

r 

n 

n 

n 

F 

EN 

Hy 

p 

he 

9 4 

6 

9 71 

9 7 

9 7 

4 

L F 

5 

5 

7 

6 

8 

6 

9 

8 

7 10 

71 10 

7 10 

8 

8 

8 10 

6 

9 

8 

8 11 

W Clonel Clone2 El E2 C 

0 7 6 

0 5 6 

0 4 2 

0 4 3 

0 9 3 

0 

0 9 1 

0 7 4 

0 Nodel 7 3 

0 Nodel 5 4 c 

Rhiz2 5 6 

Rhiz2 4 3 

0 7 2 

Rhiz2 6 5 

Rhiz2 7 3 

Rhizl 5 3 

Rhiz2 7 6 

Irreg 8 1 

Irreg 

Irreg 

0 5 4 

Rhiz2 8 3 

Rhiz2 8 3 

Rhiz2 6 4 

w Node2 2 6 

Rhizl 1 3 

Stoll 1 6 

Node2 

0 

Rhiz2 8 5 

0 9 2 

0 7 3 

0 7 3 

Rhizl 5 6 

0 7 3 

0 7 4 

0 7 4 

0 5 4 

0 7 5 

hc 

0.47 

0.91 

0.00 

0.02 

0.60 

0.37 

0.37 

VU 

67 

8 

6 

6 

7 

1 

4 

3 

3 

6 

71 

5 

6 

5 

3 

5 

7 

5 

1 

4 

4 

3 

4 

4 

3 

4 

6 

771 

7 

7 

7 

7 

2 

6 

6 

4 

6 

71 

6 

6 

5 

5 

71 

7 

5 

8 

7 

9 

7 

7 

8 

5 

7i 

6 

7 6 11 

7 

9 

9 

7 

7 

8 

8 

7 

7 

71 

7 

6 

6 

7 

71 

6 

6 

7 

4 

3 

8 

3 

2 

3 

2 

7 

Co Br Habitats 

1, 2 

1, 2 

15, 16 

1, 2 

16 

17 

4 

7 

13 

13 

13, 14 

13, 14 

11, 12, 13 

11 

Co 21 

11 

11 

11 

13, 14 

13, 14 

1, 3 

Co 19, 21 

Co 19 

3, 4, 19 

10, 12, 15 

11, 16 

11, 15 

14, 15, 16 

3, 4, 17 

11, 14 

3, 16, 17 

3, 16, 17 

11, 14 

11, 14 

11 

1, 3, 14, 17 

3, 17 

16 

1, 7 

1 

11 

1 

1 

1 

1 

3, 4 

Prec 

1096 

1096 

1810 

1998 

1527 

785 

848 

822 

1365 

1072 

924 

1348 

1348 

1098 

1130 

1323 

1182 

1259 

936 

836 

981 

863 

1050 

1046 

1381 

1832 

1880 

1357 

972 

971 

937 

1091 

1104 

1146 

996 

893 

834 

1328 

946 

808 

915 

774 

728 

820 

721 

1087 

Tjul 

14.5 

14.5 

12.6 

13.4 

13.7 

16.0 

15.9 

15.6 

14.2 

14.71 

15.2 

13.2 

14.0 

14.5 

15.3 

13.5 

14.4 

14.2 

15.1 

15.8 

14.9 

15.9 

14.5 

14.6 

13.2 

11.9 

11.6 

13.6 

15.0 

15.0 

16.0 

14.5 

14.5 

14.3 

14.9 

15.5 

15.9 

13.8 

15.3 

15.7 

15.3 

16.0 

16.3 

14.3 

16.1 

14.5 

CI Tjan 

13 3.5 

13 3.5 

0 1.9 

0 2.7 

0 2.1 

10 4.0 

7 4.7 

9 3.8 

3 3.7 

0 3.6 

1 3.6 

0 1.4 

7 3.7 

10 3.5 

0 4.6 

4 3.1 

2 4.0 

3 3.7 

3 3.6 

3 3.7 

0 3.3 

8 4.6 

9 4.4 

13 3.6 

0 2.7 

0 1.0 

0 0.9 

2 3.0 

9 3.7 

12 3.8 

12 4.4 

10 3.5 

8 3.5 

3 3.4 

11 3.8 

0 3.6 

7 3.9 

0 3.0 

0 3.6 

0 3.6 

3 3.9 

0 3.8 

0 3.6 

0 2.8 

0 3.6 

12 3.5 

928 

928 

5 

1 

0 

4 

0 

41 

79 

19 

89 

0 

330 

756 

5 

267 

95 

286 

424 

15 

122 

41 

180 

837 

236 

1 

0 

423 

400 

795 

0 

894 

758 

812 

824 

24 

0 

13 

161 

0 

160 

9 

0 

0 

0 

921 

GB 

2650 

2650 

308 

1 

33 

370 

79 

1066 

212 

33 

366 

14 

962 

2577 

15 

1236 

574 

889 

1696 

808 

1669 

370 

572 

2530 

1359 

218 

236 

1226 

2005 

2036 

371 

2630 

2347 

2417 

2070 

937 

1171 

60 

1218 

86 

447 

134 

235 

10 

8 

2666 

Am 

Am 

0.27 

0.32 

-0.28 

-0.01 

-0.29 

-0.41 

-0.76 

1.42 

3.88 

0.12 

0.07 

-0.39 

0.38 

-0.18 

-0.41 

-0.25 

1.66 

0.16 

0.08 

0.26 

-0.39 

0.19 

-0.08 

n 

n 

n 

n 

n 

n 

n 

NZ 

Am 

Ch 

0Rhiz2 7 7 43 

0 7 3 

0 7 3 

0 7 1 

Rhiz2 4 5 

Rhiz2 3 6 

n 

n 

n 

n 

n 

n 

n 

n 

s 

n 

s 

n 

s 

n 

r 

x 

n 

EN 

CR 

0.39

F R N S 

10 6 4 

78 76 6 3 

7 5 4 

4 8 5 

8 5 5 

8 7 6 

8 8 3 

5, 7 6 

1 P2 LF1 LF2 W 

Gn Hy 

Gn 

Gn 

Gn 

Gn 

Gn 

Gn 

Gn 

Chg 

0.42 

0.30 

0.18 

0.11 

RS 

n 

n 

n 

s 

-0.35 

0.41 

-0. 12 

1.59 

-0.11 

-0.94 

n 

n 

s 

Gn 

7 1 1 

56 22 2 2 

8 2 1 

8 2 1 

6 4 1 

5 7 3 

5 7 2 

3 7 2 

3 8 2 

11 4 1 

CI 

8 

0 

8 

0 

0 

0 

3 

0 

0 

0 

13 

0 

0 

2 

0 

5 

0 

11 

1 

E2 C Origin 

6 

6 

6 

6 

5 Eur, As 

6 

3 

6 

Eur 

1 

1 

1 

0 

2 

1 

6 

4 

SAm 

Eur 

0 

6 

6 c 

3 

6 

4 

4 

2 

2 

1 

Ch Pn 

Ch Pn 

Pn 

Ch Pn 

Ch Pn 

Pn 

b Th hc 

Ch 

Ch 

Ch 

Hy 

hc 

hc 

hc 

hc 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

hc 

hc 

Th 

Ch Pn sw 

Ph 

hc 

Ch 

hc 

Fight Len 

100 

100 

60 

60 

120 

90 

180 

60 

15 

60 

60 

120 

60 

60 

80 

50 

20 

25 

20 

20 

60 

60 

60 

50 

40 

40 

15 

20 

40 

9 

9 

10 

10 

60 

75 

60 

90 

60 

600 

150 

70 

40 

r 

n 

o 

o 

n 

r 

n 

r 

-0.91 

-0.07 

0.33 

-0. 11 

2.37 

1.52 

0.18 

-0.79 

-0.20 

0.36 

-0.36 

99 44 21 

9 7 2 

8 2 1 

4 6 4 

4 6 4 

4 7 2 

4 6 6 

4 6 5 

3 7 3 

3 7 3 

3 6 3 

3 7 3 

4 7 7 

3 6 3 

4 6 5 

5 7 7 

4 8 5 

5 8 5 

8 6 7 

5 6 6 

3 7 3 

4 8 5 

Br Habitats L 

11 

1, 11, 14 

11 

11 

6 

1, 16 

11 

11 

1, 3, 17 

10, 12 

10 

10, 13, 14 

10, 12 

10, 12 

10 

3, 16 

15, 16 

3, 16, / 7 

3, 16 

13 

12 

11 

11 

12 

19 

19 

19 

18, 19 

3, 4 

3, 16, 17 

3, 16, 17 

3, 16, 17 

3, 16, 17 

3, 16 

3, 5 

19 

3, 4 

3, 16, 17 

1, 3 

11 

1 

3, 4, 7, 19 

Prec Co 

1116 

1108 

1091 

1406 

740 

1269 

950 

1224 

716 

1011 

1226 

1395 

1593 

1230 

1047 

774 

1653 

904 

1121 

1435 

1194 

1006 

1394 

1307 

902 

902 

959 Co 

996 Co 

880 

997 

844 

918 

1011 

809 

856 

1010 Co 

804 

848 

904 

974 

827 

838 

Tjul 

14.4 

14.0 

14.5 

12.7 

16.3 

13.7 

15.2 

14.0 

15.6 

15.9 

14.0 

14.0 

13.7 

14.0 

15,5 

15.9 

10.6 

15.9 

14.5 

14.1 

14.1 

15.4 

13.6 

13.6 

15.2 

15.2 

15.5 

15.6 

15.8 

14.9 

15.5 

15.2 

14.7 

15.9 

16.1 

152 

15.7 

15.6 

15.4 

15.2 

16. 1 

15.6 

Tjan 

3.5 

3.0 

3.4 

1.8 

4.1 

2.9 

3.8 

3.4 

3.4 

5.7 

3.5 

4.3 

4.0 

3.4 

6.0 

3.8 

-1.2 

4.6 

3.3 

4.4 

3.4 

4.5 

2.7 

3.1 

3.8 

3.8 

5.0 

5.3 

4.6 

3.5 

3.7 

3.6 

3,4 

IR 

847 

112 

704 

35 

0 

334 

426 

129 

0 

0 

712 

24 

10 

781 

1 

80 

0 

34 

55 

70 

831 

14 

100 

560 

167 

167 

13 

33 

80 

40 

11 

177 

423 

25 

1 

106 

80 

97 

483 

410 

0 

2 

GB 

2494 

238 

2534 

170 

2 

1561 

1248 

170 

614 

19 

1999 

0 

0 

1962 

6 

974 

10 

292 

341 

8 

2134 

17 

575 

1516 

1666 

1666 

74 

195 

338 

359 

123 

1074 

2180 

141 

46 

291 

929 

907 

1254 

1715 

704 

369 

Clonel Clone2 El 

Rhiz2 5 

Rhiz2 5 

Rhiz2 5 

Rhiz2 4 

Rhiz2 8 

Rhiz2 5 

Rhiz2 8 

Rhiz2 2 

0 

w 0 8 

w 7 

w 8 

w 0 7 

w Nodel 7 

w O 8 

0 5 

0 1 

0 

0 

Rhizl DRa 

Rhiz2 

Rhiz2 

Rhiz2 

Ogr 

0 

0 

0 

0 

0 

0 

0 

0 

0 

0 

0 

0 

0 

r 

n 

r 

n 

n 

n 

n 

s 

n 

n 

s 

n 

n 

NS CS 

N 

N 

N 

N 

AN 

N 

N 

N 

AN 

N 

N 

N 

N 

N 

N 

N 

N VU 

AN 

AN 

N 

N 

N VU 

N 

N 

N 

N 

N 

N 

AR 

N 

N 

N 

N 

AN 

AR VU 

N 

AR 

AR 

N 

N 

N 

AN 

Taxon name Fam 

Equi 

Equi 

Equi 

Equi 

Equi 

Equi 

Equi 

Equi 

Ranu 

Eric 

Eric 

Eric 

Eric 

Eric 

Eric 

Aste 

Aste 

Aste 

Orob 

Eno 

Cype 

Cype 

Cype 

Cype 

Gera 

Gem 

Gem 

Gera 

Gem 

Bras 

Bras 

Bras 

Bras 

Bras 

Apia 

Apia 

Bras 

Bras 

Cela 

Aste 

Euph 

Euph 

1 

1 

-0.11 

1 

1 

0.38 

0.47 

11 

8 4 

0 3,9 

2 4.6 

4.7 

2 3.7 

8 4.0 

0 3.9 

3.9 

9 4.1 

2 3.6 

Eur 

11 

8 3 

8 3 

0.52 

-0.02 

-0.41 

-0.80 

-0.65 

1.05 

0.15 

-0.15 

-0.22 

0.98 

n 

Unk 

Gard 

7 3 

7 3 

7 3 

7 3 c Eur 

11 

w Root 

h Rhizl 

h 0 

h Root 

n 

n 

n 

4 71 5 

3 

4 

4, 17 

715 

783 

973 

16.1 

15.8 

14.9 

1 3.7 

3 3.9 

12 3.8 

0 

125 

642 

169 

1039 

2114 

Eur 

5 6 6 

8 3 

8 5 

h Root 

h 0 

h 0 

hc 

Th 

Th 

80 

20 

40 

-1.18 

-0.77 

AN 

AR 

AR 

Euph 

Euph 

Euph

F R N S 

5 57 45 0 

7 5 3 

7 5 3 

0 

0 

3 

0 

0 

0 

0 

0 

0 

1 

0 

3 

, 

Taxon name Fam NS CS RS Chg_ Hght Len P1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L 

Euphorbra hyberna Euph N VU r 52 p hc 0 8 2 

2 137 0 5.2 14.9 1203 1, 3 5 

Euphorbia lathyris Euph AR 2.16 120 b Ch 0 8 3 

956 21 13 3.8 15.8 804 3, 16, 17 6 

Euphorbia paralias Euph N n -0.35 52 Ch 0 9 1 

182 65 8 5./ 15.6 987 Co 19 9 

Euphorbia peplis Euph N EW x -1.49 5 Th 0 9 1 

26 1 8 5.8 16. 1 969 Co 19 9 

Euphorbia peplus Euph AR -0.17 30 Th 0 8 3 

1911 52 14 3.8 15.1 950 4, 17 7 

Euphorbia platyphyllos Euph AR -0.24 70 Th 0 8 3 

248 0 3.9 16.3 733 4 7 

Euphorbia portlandica Euph N n -0.09 35 Ch 0 8 / 

148 8 14 5.3 15.4 1060 Co 18, 19 8 

Euphorbia serrulata Euph NA VU r 1.20 65 Th 0 8 3 

13 0 4. 1 16.3 812 1, 3 5 

Euphrasia anglica Scro NE n 20 Th 0 7 1 

375 5 0 4. 1 15.3 1081 10, 16 71 

Euphrasia arctica Scro N n 30 Th 0 5 1 

1095 47 0 3.2 13.6 1315 6, 11 7 

Euphrasia cambrica Scro NE VU r 8 Th 0 1 1 

5 0 2.8 13.2 2458 15 8 

Euphrasia campbelliae Scro NE r 10 Th 0 4 1 

11 0 4.0 12.5 1638 10, 11 8 

Euphrasia confusa Scro N n 20 Th 0 5 1 

970 3 3 3.2 13.8 1267 7, 8, 16 8 

Euphrasia foulaensis Scro N s 6 Th 0 4 1 

143 0 3.6 12.4 1263 Co 18, 21 8 

Euphrasia frigida Scro N s 20 Th 0 1 4 

118 0 1.5 11.8 1981 15, 16 81 

Euphrasia heslop-hanisonii Scro NE r 15 Th 0 4 1 

20 0 3.7 12.6 1695 Co 21 8 

Euphrasia marshallii Scro NE r 12 Th 0 4 1 

39 0 3.8 12.3 1334 Co 18 8 

Euphrasia micrantha Scro N n 25 Th 0 7 3 

941 19 0 3.0 13.2 1400 8, 10 7 

Euphrasia nemorosa Scro N n 35 Th 0 7 3 

1498 20 71 3.6 14.8 1037 7, 10 

Euphrasia ofi cinalis agg. Scro N n -1.61 30 Th 0 6 3 

2600 87 12 3.5 14.4 1131 6, 7, 8, 10 8 

Euphrasia ostenfeldii Scro N r 12 Th 0 2 1 

85 1 2.8 12.2 1694 16 9 

Euphrasia pseudokerneri Scro NE s 20 Th 0 7 1 

167 0 3.7 16.1 753 7 7' 

Euphrasia rivularis Scro NE r 15 Th 0 4 1 

14 0 2.3 13.2 2450 15, 16 7 

Euphrasia rostko viana Scro N n 35 Th 0 5 3 

29 1 12 1 3.7 14.3 1336 6 71 

Euphrasia rotundifolia Scro NE EN r 10 Th 0 4 1 

4 

3.5 12.4 1004 Co 18 8 

Euphrasia salisburgensis Scro N o 12 Th 0 2 3 

0 3 4.6 14.7 1134 16 7 

Euphrasia scottica Scro N n 25 Th 0 4 3 

601 7 2.5 12.8 1575 10, 15 8 

Euphrasia tetraquetra Scro N n 15 Th 0 7 1 

313 10 1 4.7 14.9 1105 6, 10 8 

Euphrasia vigursii Scro NE VU r 20 Th 0 7 1 

33 

5.4 15.3 1263 10 

Exact, /um pusillum Gent N o 8 Th 0 8 2 

0 

6.8 16.7 712 19 9 

Fagopyrum esculentum Poly AN -0.53 60 

Th 0 

Asl ? 501 

3.8 15.8 788 4 8 

Fagus sylvatica Faga N n -0.62 3000 

Ph sAf 

7 3 2397 79 3.5 14.7 1061 1 3 

Fallopia convolvulus Poly AR -1.31 100 

Th 0 

6 4 2139 51 3.7 14.9 971 3, 4 7 

Fallopia dumetorum Poly N s -0.33 200 

Th 0 

7 5 69 

3.9 16.4 784 1, 3 6 

Fallopia j aponica Poly AN 1.83 200 

Gn Rhiz2 

As2 2060 68 1 3.7 14.8 1043 3, 14, 17 6 

Fallopia sachalinensis Poly AN 1.05 300 

Gn Rhiz2 

As2 507 6 3.7 15.3 976 3 6 

Festuca altissima Poac N n 0.83 120 

hc 0 

7 3 241 8 2.6 13.6 1410 1, 16 3 

Festuca arenaria Poac N S 75 

hc Rhiz2 

7 1 108 

4.2 15.4 828 19 8 

Festuca armoricana Poac N o 40 

hc 0 

7 1 0 

6.2 16.9 796 19 8 

Festuca arundinacea Poac N n 1.71 120 

hc 0 

8 4 2082 55 7 3.6 14.9 980 6, 7 8 

Festuca filiformis Poac N n 35 

hc Ogr 

7 2 821 6 4 3.0 14.3 1100 8 8 

Festuca gigantea Poac N n 0.46 125 

hc 0 

7 3 1885 35 0 3.6 15.1 981 1 5 

Festuca huonii Poac N a 25 

hc 0 

7 0 0 7 6.4 16.5 780 16, 18 8 

Festuca lemanii Poac NA r 66 

hc 0 

7 1 71 1 3.4 15.3 1079 7, 8, / 6 8 

Festuca longifolia Poac N VU r 40 

hc 0 

7 1 15 6 4.8 16.2 751 8 8 

Festuca ovina Poac N n 43 

hc Ogr 

5 5 1788 5 4 3.5 14.5 1109 7, 8 7 

3 

0

NS CS RS Chg_ Hght Len P1 P2 LFI LF2 W Clonal Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N S 

n -0.15 45 p hc h Ogr 3 6 2718 726 13 3.5 14.4 1118 7, 8 7 5 4 2 0 

n -0. 16 80 p hc h O 5 4 1976 569 4 3.6 14.9 985 6 7 6 6 6 0 

Taxon name Fam 

Festuca ovina agg. Poac 

Festuca pratensis Poac 

Festuca pratensis x Lolium 

perenne (x Festulolium 

6 71 6 

5 6 5 

62 35 2 2 

3 4 2 

3 4 2 

4 7 3 

a 

a 

a 

a 

a 

48 6 4 5 

4 8 2 

5 8 5 

5 6 4 

5 6 7 

8 5 5 

8 8 5 

6 7 6 

8 7 4 

6 5 5 

4 6 6 

4 6 71 

790 80 3.5 15.5 851 6 

iz2 3 6 2799 974 1 3.5 14.5 1104 3, 5, 6, 2/ 

iz2 3 6 2799 974 1 3.5 14.5 1104 6 

gr DRi 2 6 801 182 2.7 12.7 1592 15, 16 

Eur 21 0 3.8 16.4 647 3, 4 

7 3 85 0 3.71 16.4 660 3, 4 

7 3 788 105 1 3.6 15.1 904 16 

9 2 132 0 3.8 16.3 687 4 

8 3 980 78 3.8 15.5 832 3 

iz2 5 5 2692 967 3.5 14.5 1105 11 

izl DRg 7 4 578 8 3.6 15.8 776 7 

0 3 Eur 961 60 1 4.1 15.9 823 3, 16, 17 

oI2 7 4 2380 819 3.5 14.6 1091 1, 7 

ol2 Gard 740 20 3.7 15.5 863 3, 17 

7 4 712 56 3.8 15.8 881 1 

8 2 58 0 4.4 16.5 680 Co 19, 21 

7 3 2459 930 1 3.6 14.7 1069 1 

7 3 98 0 3.6 16.3 688 6 

SAm 367 542 4.3 14.3 1223 3, 17 

9 1 423 330 1 4.5 14.8 1086 4 

9 2 482 180 11 4.2 14.9 997 3, 16 

8 3 307 16 0 3.6 15.5 763 4 

8 1 1200 368 13 4.0 14.9 1004 4 

7 1 31 0 0 6.5 15.8 1073 3, 4 

8 3 1978 304 12 3.6 15.0 912 3, 4 

8 3 128 0 0 3.6 16.1 703 4 

7 1 191 37 2 4.1 14.8 978 3, 4 

8 1 Eur 13 0 1 5.4 16. 1 958 ' 3, 4, 17 

7 4 116 0 0 3.5 16.2 702 4 

b DRa 8 3 c 1 0 0 2.9 14.7 1008 16 

Otb DRg 7 3 c 212 0 0 3.0 15.1 861 1, 3, 14, 16 

Otb DRg 8 3 Eur 1763 36 5 3.5 15.2 919 1, 3, 17 

0 7 3 c Eur 352 0 1 3.7 16.2 725 3, 17 

0 7 3 616 43 0 3.8 15.8 791 4, 16 

0 5 5 1135 138 1 3.6 14.9 1011 4 

0 7 2 32 01 0 3.7 15.6 789 4 

0 5 4 999 84 0 3.1 14.5 1043 4 

0 5 3 1725 409 1 3.6 14.8 1025 3 

0 5 5 2508 623 4 3.5 14.5 1073 3 

0 SAm 429 2 9 3.8 16. 1 721 4, 17 

0 SAm 524 5 6 3.8 16.0 778 3, 4, 17 

0 7 3 2672 960 14 3.6 14.6 1080 3, 4, 17 

hc 

hc 

hc 

hc 

Th 

Th 

Th 

Th 

Th 

hc 

hc Gn 

hc 

hc 

hc 

Ph 

Ch 

Ph 

Gb 

Pn 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Gb 

Gb 

Gb 

hc 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

loliaceum) Poac NH n 65 

Festuca rubra Poac N n 72 

Festuca rubra agg. Poac N n 2.96 72 

Festuca vivipara Poac N n 0.13 44 

Filago gallica Aste AR CR 0.01 20 

Filago lutescens Aste NA VU s -0.34 25 

Filago minima . Aste N n -0.91 20 

Filago pyramidata Aste AR EN -1.14 30 

Filago vulgaris Aste N n -1.20 35 

Filipendula ulmaria Rosa N n -0.10 120 

Filipendula vulgaris Rosa N n -0.07 50 

Foeniculum vulgate Apia AR 1.17 250 

Fragaria vesca Rosa N n -1.09 30 

Fragaria x ananassa Rosa AN 0.42 40 

Frangula alnus Rham N n -0.16 500 

Frankenia laevis Fran N s 0.03 8 

Fraxinus excelsior Olea N n -0.73 2500 

Fritillaria meleagris Lili NA s 0.86 30 

Fuchsia magellanica Onag AN 1.85 150 

Fumaria bastardii Fuma N n 0.39 75 

Fumaria capreolata Fuma N n 0.31 100 

Fumaria densillora Fuma AR -0.37 52 

Fumaria muralis Fuma N n 1.75 100 

Fumaria occidentalis Fuma NE s 0.04 100 

Fumaria ofi cinalis Fuma AR -0.34 52 

Fumaria parvillora Fuma AR -0.55 471 

Fumaria purpurea Fuma N s 0.25 60 

Fumaria reuteri Fuma AN -0.62 100 

Fumaria vaillantii Fuma AR -0.51 42 

Gagea bohemica Lili N VU r 9 

Gagea lutea Lili N n 0.16 25 

Galanthus nivalis Lili AN 3.01 22 

Galega dfi cinalis Faba AN 150 

Galeopsis angustifi lia Lami AR -3.31 50 

Galeopsis bifida Lami N n 100 

Galeopsis segetum Lami AR EX 50 

Galeopsis speciosa Lami AR -1.82 100 

Galeopsis tetrahit Lami N n 100 

Galeopsis tetrahit agg. Lami N n -0.61 100 

Galinsoga parVillora Aste AN 0.63 80 

Galinsoga quadriradiata Aste AN 1.071 80 

Garium aparine Rubi N n -0.09 150 

La) 

35 86 65 

4 6 5 

5 7 6 

4 8 5 

4 6 5 

4 6 5 

3 8 5 

2 5 2 

6 7 7 

6 7 7 

5 7 8 

2 8 4 

5 6 6 

4 3 3 

5 7 7 

5 6 6 

5 6 6 

4 6 7 

5 6 6 

6 7 8 

a 

a 

a 

a 

a 

a 

a 

a 

a 

a 

a 

a 

a 

a 

a 

a 

a 

a

S 

Taxon name Fan/ NS CS RS Chg li ght Len 1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N 

Galiurn boreale Rubi N n -0.52 45 hc h Rhiz2 5 6 561 101 0 2.2 12.8 1580 7, 16 7 5 7 3 

Galium conshictum Rubi N r 40 hc h Rhizl 9 1 12 0 5 5.0 16.5 799 13 8 9 3 2 

Galium mollugo Rubi N n 0.04 112 hc h Rhiz2 5 3 1665 0 14 3.5 15.3 905 7 7 4 7 4 

Galiurn odoratum Rubi N n -0.62 45 hc h Rhiz2 7 3 1836 350 1 3.4 14.7 1086 1 3 5 71 6 

Gahm palustre Rubi N n 0.07 75 hc h Rhizl 5 4 2652 932 9 3.5 14.5 1105 11 7 9 5 4 

Galium parisiense Rubi NA $ -0.57 30 Th h O 9 2 63 0 0 3.7 16.3 693 3 8 3 7 2 

Galium pumilum Rubi N r -1.32 35 hc h Nodel 7 3 65 0 0 3.7 16.2 754 7 7 4 8 3 

Galium saxatile Rubi N n -0. 15 30 hc h Node2 7 2 2489 793 9 3.5 14.3 1148 8, 9 6 6 3 3 

Galturn spurium Rubi AN -1.871 100 Th h O 7 6 c NHem 55 0 0 3.9 16.1 755 4 71 5 8 5 

Galiurn sterneri Rubi N n 0.69 30 hc h Nodel 4 2 273 35 0 2.0 13.0 1457 7, 16 9 4 7 1 

Galium tricomutum Rubi AR CR -4.78 50 Th h 0 8 4 386 1 0 3.8 16.1 736 4 7 4 7 4 

Galium uliginosum Rubi N n -0.14 60 hc h Rhizl 5 5 1435 102 0 3.2 14.9 96 1 11 7 9 6 4 

Galium verurn . Rubi N n -0.85 75 hc h Rhiz2 5 5 2516 730 14 3.5 14.6 1045 7 7 4 6 2 

Gastridium ventricosum Poac NA $ -0.48 50 Th h o 9 1 159 0 2 4.6 16.3 839 4, 7 9 3 8 2 

Gaudinia fragilis Poac NA s 45 Th h 9 2 47 0 2 4.5 16.2 865 3, 6 8 5 6 6 

Gaultheria shallon Eric AN 150 Pn w Rhiz2 Am4 179 14 1 3.6 14.9 1077 1, 10 3 4 4 3 

Genista anglica Faba N n -1.09 50 Ch Pn wi 0 7 1 858 0 0 3.0 14.6 1019 10 8 5 3 2 

Genista pilosa Faba N r -0.26 40 Ch Pn wo 0 7 3 23 0 0 5.0 15.6 1011 10 8 5 4 

Genista tind oria Faba N n -0.77 60 Ch Pn w 0 7 3 932 0 1 3.6 15.6 880 6 8 6 71 2 

Gentiana nivalis Gent N VU r 15 Th 0 1 3 4 0 -0.5 11.2 1853 15, 16 9 5 7 3 

Gentiana pneurnonanthe Gent N s -0.31 40 hc 0 7 4 130 0 3.7 15.7 806 8, 10 8 7 4 1

Taxo n name Fam NS CS RS Chg._ Hght Len P1 P2 LF 1 LR2 W Clona l Clo ne2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N S 

Hordelymus europaeus Poac N s 0. 12 120 p hc 0 7 3 185 1 0 3.2 15.5 788 1 6 4 7 7 0 

Hordeum distichon Poac AC 75 a Th 0 Cro p 714 76 3 3.8 15.4 898 3, 4 8 4 7 7 0 

Hordeum distichon sensi at Poac AC 87 a Th 0 Crop 714 76 3 3.8 15.4 898 3, 4 8 4 71 7 0 

Hordeum j ubatum Poac AN 60 p hc 0 As2, Am 349 6 1 3.5 15.4 800 3, 5 9' 6 7 6 2 

Hordeum marinum Poac N s -0.85 37 a Th 0 9 1 146 0 1 4.3 16.4 720 6 9 6 8 6 4 

Hordeum murinum Poac AR I -0.04 60 a Th 0 8 4 1497 48 14 3.8 15.6 817 3, 17 8 4 7 6 0 

Hordeum secalinum Poac N n -0. 19 70 p he 0 7 3 926 30 0 3.8 16.0 742 5 8 6 7 6 

Hordeum vulgare Poac AC 100 a Th 0 Cro p 196 8 1 3.8 15.6 833 3, 4 9 4 71 7 

Hornungia petraea Bras N s 0.31 10 a Th 0 7 3 56 0 3 3.5 14.8 116 1 16 9 2 8 1 

Hottonia palustris Prim N n -0 .63 120 p Hy brag 7 3 463 2 0 3.6 16.0 697 13 7 11 7 5 

Humulus lupu/us Cann N n -0 .09 450 p hc Rhiz2 7 4 1273 0 8 3.8 15.8 833 3 6 7 7 8 

Huperzia selago Lyco N n -0 .41 10 p Ch sw 0 2 6 989 234 0 2.7 13.2 1472 15, 16 7 6 2 2 

Hyacinthoides hispanica Lili AN L 60 p Gb OM Eur 847 76 12 3.9 15.5 866 1, 3 , 17 5 4 6 6 

Hyacinthoides non-scripta Lili N n -0.4 1 50 p Gb Ot b 7 1 243 9 760 13 3.6 14.6 1092 1, 9 5 5 5 6 

Hydrilla vert icillata Hydr N r 100 p Hy DRa DRg 8 5 c 2 1 0 3.5 14.2 1555 13 6 12 9 3 

Hydrocharis morsus-ranae Hyd r N n -0 .89 50 p Hy Stoll 7 4 323 84 2 3.9 15.8 772 13 7 11 7 7 

Hydrocotyle ranunculoides Apia AN 20 40 p Hy lrreg Am 43 0 0 4.0 16.6 680 13, 14 7 10 7 7 

Hydrocotyle vulgaris Apia N n -0 .53 20 p hc Node2 8 2 2091 842 10 3.7 14.4 1126 11 8 8 6 3 

Hymenophyllum tunbrigense Hym e N n -0 .54 8 p hc Rhiz l 7 0 197 112 0 3.8 14.0 1567 16 4 6 2 3 

Hymenophyllum wilsonii Hyme N n -0 .87 10 p hc Rhiz l 5 0 577 176 0 3.1 13.1 1685 1, 16 5 5 3 3 

Hyoscyamus niger Sob AR , - 1.38 80 b hc 

0 

8 4 

796 87 5 4.0 15.7 799 4 

8 4 7 9 

Hypericum androsaemum Clus N n 0.78 80 p Pn w 0 

9 2 

1139 744 8 4.0 14.8 1184 1, 3 

S 6 6 , 5 

Hypericum calycinum Clus A N 

0.74 60 p Ch Pn w Rhiz2 

Eur 

702 63 2 4.0 15.6 884 3 

5 71 5, 5 

Hypericum canadense Clus AN 

20 a Th 

0 

Am 

0 3 0 4.6 14.4 1252 11, 14 8 9 2 2 

Hypericum elodes Clus N n -0 .46 20 40 hc Hy Node2 7 1 583 260 7 4.2 14.8 1194 11 8 10 3 2 

Hypericum hirsutum Clus N n -0 .18 100 hc 0 7 4 1276 10 0 3.3 15.3 832 6, 7 6 5 7 5 

Hypericum humilusum Clus N n -0 .40 10 Ch 0 7 3 1732 428 12 3.6 14.8 1085 3 7 6 4 3 

Hypericum Iinadilolium Clus N r 0.09 40 hc Ch 0 8 1 14 0 9 5.5 15.8 1069 16 71 3 3 2 

Hypericum maculatum Clus N n 2.11 60 hc Rhiz2 5 3 1180 300 0 3.6 15.0 1016 1, 3 , 16 6 6 5 5 

Hypericum montanum Clus N n -0 .49 80 hc 0 7 3 269 0 0 3.8 15.6 867 1, 3 7 4 8 2 

Hypericum co l oratura Clus N n 80 hc Rhiz2 Root 8 4 1906 385 6 3.6 15. 1 95 1 7 7 4 7 5 

Hypericum pulchrum Clus N n -0.32 60 hc 0 7 2 2427 887 9 3.5 14.3 1148 10, 16 6 5 4 3 

Hypericum tetrapterum Clus N n -0 .41 60 hc Stoll 7 3 2 101 842 11 3.71 14.9 1025 11 71 8 6 4 

Hypericum undulatum Clus N s -0 .12 60 hc Stol l 8 1 81 0 0 5.6 15.5 1152 11 8 8 4 2 

Hypochaeris glabra Ast e N n - 1.01 20 a Th 0 8 3 270 5 14 4.0 15.9 754 8 8 4 4 2 

Hypochaeris maculata Aste N VU r -0 .10 60 p hc 0 7 4 c 17 0 1 3.9 16.1 710 7 8 4 8 3 

Hypochaeds radicata Aste N n 0.6 1 60 p hc 0 8 3 272 5 977 14 3.5 14.5 1104 6 8 4 5 3 

/beds amara Bras N s - 1.21 35 a Th 0 8 2 471 0 0 3.5 16.3 710 7 71 4 8 3 

Ilex aquifolium Aqui N n -0. 16 1500 p Ph w 0 8 2 235 3 86 1 14 3.6 14.7 1079 1 5 5 5 5 

Illecebrum vedicillatum Cary N r -0 .60 20 a Th h 0 Nodal 8 2 37 ' 0 0 5.6 15.7 1062 3, 13 8 7 3 2 

Impatiens capensis Bals AN 0.71 60 a Th h 0 Am6 323 1 0 3.7 16.2 723 13, 14 7 9 7 6 

Impatiens glandulifera Bals AN 1.85 200 a Th h 0 As l 1599 286 6 3.7 15.1 957 14 6 8 7 7 

Impatiens noli-tangere Bals N s -0 .77 60 a Th h 0 7 5 2 1 0 0 2.6 13.9 1833 1 4 71 71 6 

Impatiens pandfi ora Bals AN 

0.10 100 a Th h 0 

AO 470 1 0 3.5 15.7 82 1 1 

4 5, 7 8 

/nu/a conyzae 

Ast e N n -0 .15 125 p hc h 0 7 3 

860 0 7 3.9 15.9 82 1 7, 16 

7 3 8 3 

Mula crithmoides Ast e N e 0.09 95 p hc Ch h 0 9 1 121 19 8 5. 1 16.1 869 Co 18 9 6 7 5 

l a 

V

Taxon name Fam NS CS RS Chg_ Hght Len P1 P2 LF1 LF2 W Clonal Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats F R N 

Mu la helenium Aste AR -0.80 150 p hc 0 Eur, As l 631 100 3 4.0 15.1 973 3, 17 6 5 

Mu la salicina Aste N o 60 p hc Rhiz2 7 5 c 0 3 0 4.7 14.9 1017 16 9 3 

Iris foetidissima Ind N n 1.47 80 p hc Rhizl 8 2 728 0 14 4.2 16.1 821 1 8 5 

Iris germanica Ind AN 95 p hc Rhizl Gard 211 1 3 3.9 16.0 734 3, 17 6 4 

Iris pseudacorus Irid N n 0.16 150 p Gn Hy Rhiz2 8 3 2563 959 10 3.7 14.6 1081 11 6 6 

(sail s tinctoria Bras AR 1.08 150 b p hc 0 Eur, As 102 2 1 3.8 16.0 733 3, 16 8 3 

Isoetes echinospora Isoe N n 0.65 15 p Hy 0 4 6 178 32 0 3.2 13.0 1613 13 1 5 2 

Isoetes histrix lsoe N r 4 p hc 0 9 1 3 0 5 6.6 16.2 829 16 5 1 

Isoetes lacustris Isoe N n 0.95 25 p Hy 0 4 4 522 147 0 2.8 12.9 1660 13 1 4 1 

Iso lapis cernua Cype N n 0.23 15 a p Th hc 0 9 1 242 215 10 4.8 14.8 1181 11 5 3 

(so lapis setacea Cype N n 0.53 15 a p Th he 0 7 4 2038 638 10 3.5 14.4 1149 11, 14 5 3 

Jasione montana Camp N n -1.08 50 hc 0 7 3 1076 420 14 4.0 14.8 1152 8, 10 4 2 

Jug bans regia Jugl AN 2400 Ph w 0 Eur?, Asl ? 803 11 3 3.7 15.9 779 1, 3 8 7 

Juncus acutiflorus Junc N n 1.16 100 he Rhiz2 7 3 2498 845 11 3.5 14.5 1115 11 4 2 

Juncus acutus Junc N s 0.01 150 hc Ogr 9 1 41 28 12 5.4 15.7 1031 Co 19 7 3 

Juncus alpinoarticulatus June N $ -0. 12 30 hc Rhizl 4 6 53 0 0 0.7 12.1 1477 11 7 2 

Juncus ambiguus Junc N n 17 Th 0 8 3 175 50 1 4.4 14.9 999 Co 19, 21 71 5 

Juncus articulatus Junc N n 1.26 60 hc IRhiz2 8 4 2740 956 11 3.5 144 1108 11 6 3 

Juncus balticus Junc N s -0.34 45 Gn Rhiz2 2 6 92 0 0 3.3 13.1 1012 Co 19 5 2 

Juncus bigIumis June N s -0.17 12 hc Ogr 1 6 37 0 0 0.5 11.4 2194 11, 15 8 2 

Juncus bufonius June N n 25 Th 0 6 6 1986 724 3 3.6 14.6 1102 3, 11, 13, 14 6 5 

Juncus bufonius sensi at. Junc N n 1.13 25 Th 0 6 6 2736 937 14 3.5 14.5 1105 3, / , 13, 14 6 5 

La 

co Juncus bulbosus June N n 0.34 30 90 hc Hy Node2 Irreg 5 3 2250 827 8 3.5 14.2 1180 14 1 4 2 

Juncus capitatus Junc N 5 Th 0 8 3 12 0 10 6.3 16.1 889 10 5 1 

Juncus castaneus Junc N s -0.40 30 he Rhiz2 1 6 44 0 0 0.1 11.2 2291 15 7 3 

Juncus compressus Junc N n -1.09 30 Gn Rhizl 7 4 430 4 1 3.6 15.9 746 6, 11 7 5 

Juncus conglomeratus Junc N n 0.84 100 hc Ogr 7 3 2622 798 4 3.5 14.4 1117 11 4 3 

Juncus effusus Junc N n 1.06 120 hc Ogr 8 3 2753 974 13 3.5 14.4 1108 8, 11 4 4 

Juncus filiformis Junc N s 0.79 30 hc Ogr 4 6 32 0 0 2.3 13.6 1530 13 6 4 

Juncus foliosus Junc N n 25 Th 0 8 2 218 88 2 4.3 14.7 1241 11, 13 6 6 

Juncus gerardii June N n -0. 13 30 hc Rhiz2 6 6 919 271 11 4.2 14.4 1174 Co 21 7 6 

Juncus infi exus Junc N n 0.04 90 hc Ogr 8 4 1758 631 9 3.8 15.2 925 6, 11 71 5 

Juncus maritimus June N n -0.26 100 Gn Ogr 8 3 390 196 11 4.7 15.1 1081 Co 21 8 5 

Juncus pygmaeus Junc N EN r 8 Th 0 9 1 4 0 0 6.7 15.9 965 3 4 2 

Juncus squarrosus Junc N n 30 p hc Ogr 7 2 1849 484 0 3.1 13.8 1254 8, 12 2 2 

Juncus subnodulosus Junc N n 0.15 120 p hc Rhizl 8 3 680 224 2 3.9 15.5 838 11 8 4 

JUnCUS tenuis Junc AN 0.83 40 p hc Ogr Am, SAm 1053 145 3 3.6 14.6 1267 1, 3, 13 5,1 4 

Juncus trifidus Ju nc N n -0.38 30 p hc Ogr 1 4 177 0 0 1.0 11.5 2162 15 2 2 

Juncus triglumis Junc N n -0.38 20 p hc Ogr 1 6 200 0 0 0.9 11.8 2085 11, 16 6 2 

Jun/penis communis Cupr N n -0.42 500 p Ph Ch w O 5 6 1020 145 0 2.8 13.4 1380 7, 10, 15, 16 5 3 

Kickxia elatine Scro AR -0.18 25 a Th 0 8 3 911 41 11 4.1 16.0 813 4 6 5 

Kickxia spuria Scro AR -0.071 25 a Th 0 8 3 622 0 2 3.9 16.2 737 4 7 5 

Knautia arvensis Dips N n -0.88 100 p hc 0 7 4 1707 478 4 3.8 15.2 919 6, 7 8 4 

Kobresia sitnpliciuscula Cype N r 0.58 20 p he 0 1 6 18 0 0 -0.1 11.5 2044 11, 15 6 1 

Koeleria macrantha Poac N n -0.29 50 p hc 0 7 6 1250 266 7 3.7 14.7 989 7 7 2 

Koeleria vallesiana Poac N r 40 p hc 0 8 2 4 0 0 4.5 16.4 868 7 8 1

S 

L F R N 

8 9 6 1 

6 5 7 7 

8 4 7 6 

8 5 7 6 

8 4 7 7 

6 12 7 6 

4 5 7 6 

7 5 7 8 

7 4 7 6 

7 5 7 7 

71 5 7 6 

5 6 7 8 

6 5 7 7 

6 4 7 7 

7 4 6 3 

Br Habitats 

11, 16 

3, 17 

19 

3, 17 

3, 16 

13 

1 

3, 17 

3, 4 

4 

4 

3, 17 

3, 4, 17 

3, 17 

1, 2, 17 

LF2 W Clonal Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co 

2175 

884 

601 Co 

741 

693 

839 

855 

911 

842 

1097 

877 

869 

1027 

1059 

1049 

LF1 

Th 

Ph 

Th 

hc 

hc 

Hy 

Ch 

hc 

Th 

Th 

Th 

hc 

Th 

Th 

Ph 

1 P2 

Taxon name Fam NS CS RS Chg Hght Len 

6 

700 

75 

200 

200 

300 

60 

60 

30 

25 

30 

35 

30 

95 

4600 

7j 6, 5 3 

7 6 5 3 

3 6 7 6 

7 3 8 4 

9 5 7 6 

71 4 8 3 

6 5 4 3 

8 6 7 6 

7 9 7 4 

7 6 6 5 

71 4 8 2 

6 5 7 6 

6 5 7 6 

9 6 7 8 

9 4 5 5 

8 9 8 71 

7 4 7 4 

7 11 7 8 

7 11 7 6 

7 11 7' 7 

71 12 71 5 

8 6 6 4 

8 4 7' 3 

8 5 6 3 

7 4 7 6 

8 4 8 6 

7 4 5 4 

8 5 7 8 

9 4 7 7' 

2, 17 

2, 17 

1, 3 

4, 7 

19 

3, 18 

8, 15 

3, 6, 7 

11 

6 

3 

3 

3, 17, 18, 19 

18 

3, 4, 16 

13 

4 

13 

11, 13 

13, 14 

11, 13 

6 

7 

7 

3, 4, 17 

3, 19, 21 

3 

6 

3, 17 

1134 

1133 

955 

718 

886 Co 

782 

1172 

742 

823 

1079 

835 

776 

942 

1023 Co 

875 

766 

717 

739 

986 

787 

832 

1103 

918 

960 

801 

786 

1074 

629 

757 

Ph 

Ph 

Gn 

Th 

Gn 

hc 

Gn 

Th 

hc 

hc 

hc 

Gn 

Ph 

Pn 

Th 

hc 

Th 

Hy 

Hy 

Hy 

Fly 

hc 

hc 

hc 

Th 

hc 

hc 

hc 

Th 

3000 

3700 

30 

60 

20 

300 

40 

75 

120 

80 

200 

120 

600 

300 b 

100 a 

90 

30 

0.5 

0.4 

0.3 

1 

60 

35 

20 

40 

60 

50 

120 

40 

LO 

0 

p 

3

Taxon name Fam NS CS RS Chg Hght Len P1 P2 LF1 LF2 W Clonal Clone2 E/ E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N 

Lobelia dortm anna Camp N n -0.05 - 4 p Hy h 0 

4 3 

570, 178 0 3.0 12.9 1635 13 

8 12 5 1 

Lobelia urens Camp N VU r -0. 19 70 p hc h 0 

8 1 

12 0 0 4.7 16.1 988 8, 10 

8 8 4 2 

Lobularia maritima Bras AN ' 2.34 30 a p Th hc h 0 0 3 Eur 738 33 13 4.0 15.71 854 18, 19 9 3 71 4 

Loiseleuria procum bens Eric N n -0.58 25 p Ch w Node2 1 6 182 0 0 0.9 11.4 1994 15 9 5 2 2 

Lolium multifl orum Poac AN : -1.06 100 a b Th hc h 0 Eur 2069: 340 10 3.6 14.9 968 5 7 5 7 7 

Lolium perenne Poac N n -0.29 50 p hc h 0 8 3 2743 964 14 3.6 14.5 1096 3, 6 8 5 6 6 

Lolium te m ulentum Poac AR -4.05 90 a Th h 0 Eur 341 37' 7 4.0 15.6 855 4 7 4 8 7 

Lonice ra p ericlym e num Capr N n -0.11 600 p Ph w 0 Node2 8 2 2622 943 14 3.6 14.5 1102 1 5 6 5 5 

Lonicera xylosteum Capr AN 0.58 200 p Pn w 0 7 4 Eur, As1 242 16 0 3.4 15.2 874 1, 3 5 5 7 6 

Lotus angustissimus Faba N $ -0.23 30 a Th h 0 8 3 55 0 12 5.7 16.2 952 8 8 3 4 3 

Lotus corniculatus Faba N n 1.09 40 p hc h 0 8 5 2801 975 14 3.5 14.5 1104 6, 7 7 4 6 2 

Lotus glab er Faba N n -0.55 90 p hc h 0 8 3 509 0 4 3.9 16. 1 732 3, 6, 7 7 7 7 5 

Lotus pe dunculatus Faba N n -0.06 60 p he h 0 Rhizl 7 3 2380 729; 13 3.6 14.7i 1072 11 7 8 6 4 

Lotus s ubbillorus Faba N s 0.22 30 a Th h 0 8 2 861 8 14 5.8 16.0 987 8 7 5 6 5 

Ludwigia palustris Onag N r 0. 19 15 p Th Hy h Node2 8 3 11 0 3 4.6 16.6 812 13 8 9 4 4 

Lupinus arboreus Faba AN 1 1.84 200 Pn w 0 Am4 341 21 11 4.2 15.7 825 3. 19 9 4 7 3 

Lupinus polyphyl/us Faba AN : 150 hc h 0 Am4 215 3 1 3.2 15.2 828 3, 14, 17 7 5 5 5 

Luron ium natans Alis N s 0.24 50 Hy h Sto12 7 2 91 3, 0 3.3 14.71 1229 13, 14 8 11 5 3 

Luzula arcuata Junc N r -0.43 10 he h Rhizl 1 3 22 0 0 0.0 10.8 2044 15, 16 9 5 2 2 

Luzula campestris Junc N n -0.18 15 hc h Rhiz2 7 3 2725 835 14 3.5 14.5 1100 6 7 4 5 2 

Luzula forsteri Junc N n 0.25 35 hc h 0 9 2 309 0 5 4.2 16.2 859 1 4 4 5 2 

Luzula multillora Junc N n 0.28 50 hc h 0 3 6 2451 834 7 3.5 14.3 1148 8 7 6 3 3 

Luzula pallidula Junc N VU r 30 hc h 0 5 5 c 2 0, 0 3.4 16.3 553 1, 11 7f 7 5 2 

Luzula pilosa June N n -0.35 32 hc h 0 5 4 2132 266 1 3.2 14.4 1127 1, 2 5 5 5 3 

Luzula spicata June N n -0.72 25 hc h Rhizl 1 3 189 0 0 1.0 11.5 2085 7, 15, 16 8 5 3 2 

Luzula sylvatica Jun c N n -0.02 80 hc h Rhizl 7 3 2058 621 4 3.3 14.1 1208 1, 16 5 5 4 4 

Ly chnis alpina Cary N VU r 20 Ch h 0 2 3 2 0 0 0.5 12.0 1640 15 8 3 4 2 

Lychnis flos-cuculi Cary N n -0.79 75 he h 0 7 4 2569 765 9 3.5 14.5 1095 11 71 9 6 4 

Lychnis viscaria Cary N VU r 0.01 45 Ch h 0 7 4 c 28 0 0 2.2 13.8 926 16 8 3 4 2 

Lycium Sola AN 0.13 250 Pn w Root As 1104 68 9 3.8 15.7 784 3, 19 8 5 7 4 

Lyciurn barbarum Sola AN 250 Pn w Root As 3, 19 8 5 7 4 

Lycium chinense Sola AN 250 Pn Ph w Root As 3, 19 8 5 7 4 

Lycop e rsicon e sculentum Sola AN 150 a Th h 0 SAm 523 41 4.0 15.6 876 17 71 5 71 8 

Lyc opodie lla inundata Lyco N s -0.65 5 Ch h Nodal 5 3 233' 18 3.5 14.9 1122 10, 12 9 9 2 1 

Lycopodium annotinum Lyco N s -0.38 10 Ch sw Node2 2 6 171 0 0.7 11.7 1760 10, 15 6 6 3 3 

Lycopodium clavatum Lyco N n -0.52 15 Ch sw Node2 5 6 948 81 2.4 13.3 1368 10 7 5 1 2 

Lycopus europaeus Lami N n -0.01 100 hc h Rhizl 7 4 1689 347 3.8 15.2 995 11 7 8 7 6 

Lysichiton am ericanus Arac AN 110 hc h Rhizl Am4 174 24 3.9 15.1 1073 13, 14 4 9 6 8 

Lysimachia nem orum Prim N n -0.46 20 Ch hc h Node2 7 2 2217 740 3.4 14.4 1150 1 5 7 4 5 

Lysimachia nummularia Prim N n -0.02 5 hc Ch h Node2 7 3 1266 227 3.7 15.5 895 6, 11, 14 5 7 5 5 

Lysimachia punctata Prim AN , 4.62 120 hc h Rhiz1 Eur 1127 48 3.5 15.1 999 1, 3, 17 6 6 7 5 

Lys ima chia thyrsillora Ri m N s 0.38 70 hc Hy h Rhiz2 4 6 51 0 2.7 14.2 1200 11, 13 8 10 4 3 

Lysimachia vulgaris 

Prim N n 0.22 105 

hc h Rhiz2 

7 5 

1227f 288 3.71 15.2 943 11 

7i 9 71 5, 

Lythrum hyssopifolium Lyth AR VU -1.12 15 

Th h 0 

8 4 

112 3 3.9 16.0 777 4 

8 6 6 4 

Lythrum portula Lyth N n 0.32 8 Th Hz h 0 Nodel 7 3 1262 321 3.8 14.8 1084 11, 13 8 9 5 3 

Lythrum salicaria Lyth N n -0.08 120 hc h 0 7 5 1692 827 4.0 15.0 1024 11 7 9 7 5

Taxon name Fam NS CS RS Chg_ Hght Len 1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N S 

Mahonia aquifoliurn 

Berb AN 1.61 150 Pn w Rhiz2 Am4 991 4 2 3.4 15.6 786 1, 3 4 6 5 

Maianthemurn bifolium Li li NA VU r 0.32 20 Gn h Rhiz2 5 5 0 4 0 0 2.9 15.0 753 1, 2 5 3 3 

Malus domestica Rosa AR 1000 Ph w 0 Gard 1532 327 8 3.7 15.0 975 3 5 6 7' 

Malus sylvestris sens.lat. Rosa N n 0.57 1000 Ph w 0 7 3 2023 598 11 3.7 15.0 975 1, 3 5 6 6 

Malus sylvestris sens.str. Rosa N n 1000 Ph w 0 7 3 1335 218 0 3.7 15.1 964 3 5 6 6 

Ma Iva moschata Maly N n -0.04 80 hc h 0 7 3 1423 0 10 3.7 15.6 871 6 3 7 4 

Ma Iva neglecta 

Maly AR -0.22 60 Th h 0 7 3 1196 70 14 3.8 15.7 781 3 4 8 71 

Melva pusilla Maly AN 50 a Th h 0 7 5 c Eur, As 107 6 1 4.1 16.0 788 3, 17, 19 5 5 5 

Ma Iva sylvestris Maly AR -0.30 150 p he h 0 8 4 1788 354 14 3.8 15.3 899 3, 17 4 8 7 

Marrubium vulgere Lami N s -2.02 60 p hc h Rhizl 8 4 46 0 0 4.6 16.1 861 7 5 7 8 

Matricaria discoidea Aste AN -0.49 35 a Th h 0 As2?, Am? 2677 945 14 3.6 14.5 1086 3, 4 5 7 7 

Matricaria recutita Aste AR 0.92 60 a Th h O 8 3 1588 51 11 3.7 15.5 868 3, 4 5 7 7 

Matteuccia struthiopteris Wood AN 60 p hc h O 4 6 c NHem 74 4 3.2 14.7 1108 1 8 7 7 

Matthiola incana Bras AN 0.75 80 p Pn sw O 9 1 Eur 107 2 5.0 15.9 874 Co 18, 19 3 5 2 

Matthiola sinuata Bras NA VU r 60 b hc h O 9 1 19 8 5.4 15.6 1089 Co 18, 19 3 7 2 

Meconopsis cambrica Papa N s 2.36 60 p hc h O 5 1 54 77 3.5 14.0 1321 1 5 7 5 

Medicago arabica Faba N n 0.69 60 a Th h O 9 2 744 0 1 4.2 16.2 781 3, 17 5 6 5 

Medicago lupulina Faba N n -0.43 50 a p Th hc h o 7 4 2064 686 1 3.8 15.0 963 7, 17 4 8 4 

Medicago minima Faba N s -1.97 20 a Th h O 8 4 54 0 3.9 16.5 623 8 3 7 2 

Medicago polymorphe Faba N s -1.34 60 a Th h o 9 2 118 0 1 5.0 16.4 801 8 4 5 5 

Medicago sativa Faba N s -0.56 90 p hc h 0 Rhir l 8 4 c 54 0 3.5 16.2 604 3, 6 4 6 5 

Medicago sativa 

subsp falcata Faba N s 60 hc h Rhizl 8 4 c 541 01 3.51 16.2 604 3 3 7 3 

Medicago sativa subsp.sativa Faba AN 90 hc h 0 Crop 1065 17 3.8 15.8 768 3, 6 4 6 5 

Melampyrum arvense Scro AN -0.49 60 a Th h 0 7 3 c Eur 50 0 3.6 16.2 692 3, 4, 16 4 8 3 

Melampyrum cristatum Scro N r -0.88 50 a Th h 0 7 4 c 62 0 3.4 16.3 606 3 3 8 2 

Melampyrum pratense Scro N n -0.88 60 a Th h 0 5 4 1696 323 3.2 14.3 1220 1, 2 5 2 3 

Melampyrurn sylvaticum Scro N r -0.58 35 a Th h 0 4 3 75 20 1.5 12.5 1532 1, 16 5 2 2 

Merica nutans Poac N n -0.17 60 hc h Rhiz1 5 5 c 408 0 1.9 13.0 1483 1, 7, 16 5 7 3 

Merica uniflora Poac N n -0.04 60 hc h Rhir l 7 3 1511 246 3,5 15.0 1015 1 5 7 5 

Mehlotus elbus Faba AN -0.20 150 a hc Th h 0 Eur?, Asl ? 913 15 3.7 15.8 778 3 3 7 4 

Mefi lotus altissimus Faba AR 0.73 150 hc h 0 7 3 1122 16 3.8 15.8 785 3, 17 6 7 7 

Melilotus indicus Faba AN -1.59 40 Th h 0 Eur, As 1 427 12 1 3.8 15.7 790 3, 17 5 71 71 

Melilotus ofi cinalis Faba AN 0.02 150 hc h 0 Eur?, Asl ? 1142 26 3.7 15.7 790 3, 17 5 7 5 

Melissa officinalis Lami AN 1.73 60 hc h Rhizl Eur 667 36 4.1 15.9 854 3, 17 5 7 6 

Melittis melissophyllum Lami N s -0.47 60 hc h Rhizl 7 3 119 0 5.1 15.7 1091 1, 3 4 7 5 

Menthe aquatica Lami N n -0.11 90 hc h Rhiz2 7 3 2475 922 3.7 14.7 1065 11 8 7 5 

Mentha arvensis Lami N n -1.30 60 hc h Rhiz2 5 6 1965 428 3.6 14.9 1017 4, 11 7i 71 6 

Mentha pulegium Lami N VU r -0.70 30 hc h Rhiz2 8 3 242 45 4.4 15.9 869 6, 13 7 5 7 

Mentha spicata Lami AR 1.69 90 hc h Rhiz2 Crop 1563 74 3.5 15.0 942 3, 17 5 7 7 

Menthe suaveolens Lami N $ -0.32 100 hc h Rhiz2 9 2 118 0 5.4 15.5 1151 3 8 6 6 

Menyanthes trifoliata Meny N n -0.04 30 150 Gn Hy h Rhiz2 5 6 1905 756 3.4 14.1 1201 11 10 4 3 

Mercurialis annua Euph AR 0.28 50 Th h 0 9 2 793 55 1 4.0 16.0 779 3, 4, 17 5 7 7 

Mercurialis perennis Euph N n -0.65 40 hc h Rhiz2 7 3 2214 4 3.2 14.8 1045 1 6 7 7 

Mertensia maritima Bora N s -0.53 60 hc h 0 2 3 222 29 3.8 13.4 1083 Co 19 5 T 7 

Mespilus germanica Rosa AR 900 Ph w o Eur 98 1 4.4 16.1 861 1, 3, 17 4 6 6

Taxon name Fam NS CS RS Chg. Hght Len P1 P2 LF1 LF2 W Clonal Clone2 El E2 C Origin GB IR CI Tja n Tjul Prec Co Br Habitats L F R N S 

Meum athamanticum Apia N s -0.40 60 p hc 0 4 3 164 0 0 1.6 12.9 1459 3, 6 8 5 4 3 

Mibora minirna 

Poac N r -0.01 8 a Th 

0 

8 2 

7, 0 10 5.8 16.1 861 Co 19 

9 3 71 1 

Milium effusum 

Poac N n 0.31 150 p hc 

0 

5 6 

1391 91 0 3.6 15.3 928 1 

4 5 6 5 

Milium vernale Poac N 0 5 a Th 0 9 1 0 0 2 6.8 16.6 718 19 9 3 6 2 

Mimulus Scro AN -0.47 50 p hc Node2 Am4, SAm 1767 237 5 3.2 14.3 1106 13, 14 7 9 6 5 

Mimulus guttatus Scro AN 50 p hc Node2 Am4 996 44 4 3.2 14.5 1020 13, 14 7 9 6 6 

Mimulus guttatus x luteus (M 

x robert sin 

Scro AN 

50 p hc 

Node2 

Gard 

473, 125 0 3.1 14.0 1162 13, 14 7 8 7i 5 

Mimulus luteus 

Scro AN 

50 p hc 

Node2 

SAm 

186 5 0 2.8 13.6 1144 13, 14 7 9 5 5 

Mimulus moschatus Scro AN 40 p hc Node2 Am4 361 13 2 3.3 14.6 1099 13, 14 7 8 5 5 

Minuartia hybrida Cary N s -1.70 20 a Th 0 9 2 294 0 1 3.5 16.0 716 7 9 3 8 3 

Minuartia recurva Cary N o 5 p Ch 0 1 3 0 1 0 4.8 14.2 1414 15 8 4 3 1 

Minuartia rubella Cary N r 0.01 6 p Ch 0 1 6 71 0 0 0.8 11.2 1679 15 8 4 7 1 

Minuartia sedoides Cary N s -0.75 8 p Ch 0 1 3 76' 0 0 1.2 11.4 2095 15 8 5 4 2 

Minuart ia stricta Cary N EN r 10 p Ch 0 1 6 1 0 0 0.4 12.1 1443 11 9 9 5 2 

Minuartia verna Cary N s -0.42 15 p Ch 0 4 5 139 30 0 2.7 13.9 1215 7, 16 8 4 7 1 

Misopates orontium Scro AR -0.89 50 a Th 0 8 4 488 24 11 4.3 15.9 864 4 7 5 6 6 

Moehringia trinervia 

Cary N n -0.40 40 a Th 

0 

7 3 

1990 259, 4 3.5 15.0 982 1 

4 5 7 6 

Moenchia erecta 

Cary N n -0.65 12 a Th 

0 

8 2 

418 0 14 4.2 16.0 823 8 

9 4 4 3 

Molinia caerulea Poac N n -0.34 130 p hc Ogr 5 4 2244 897 6 3.5 14.2 1179 12 7 8 3 2 

Moneses uniflora Pyro N VU r 0.14 4 p hc Rhizl 4 6 27 0 0 2.0 13.0 985 2 4 5 4 1 

Monotropa hypopitys Mono N 

-h. 

n -1.09 30 p Gn Rhizl 7 6 288 24 0 3.7 15.8 817 1 4 5 6 2 

to Montia fontana 

Port N n 0.14 20 a p Th Hy Nodal 

5 3 

21971 547, 14 3.4 14. 1 1198 11 

71 9 5 3 

Muscari neglectum 

Lili NA VU r 1.55 30 p Gb 

Gib DRg 8 4 

13 0 0 3.3 16.3 590 3, 8 

7 3 7 5 

Mycelis muralis Aste N n 0.01 100 p hc 0 7 3 1302 0 0 3.3 15.2 944 1, 16 4 5 7 5 

Myosotis alpestris Bora N r -0.22 25 p hc 0 1 6 7 0 0 -0.1 11.6 1623 7, 15, 16 8 4 8 2 

Myosotis arvensis Bora AR -0.34 40 a Th 0 5 4 2577 748 8 3.5 14.6 1056 3, 4 7 5 6 6 

Myosotis discolor Bora N n 0.14 25 a Th 0 7 3 23171 521 14 3.5 14.4 1088 6 7 5 5 3 

Myosotis taxa Bora N n 0.65 40 b hc 0 5 6 2409 757 8 3.5 14.5 1085 11 7 9 6 5 

Myosotis ramosissima Bora N n 0. 11 25 a Th 0 8 3 1174 52 14 3.8 15.5 823 8, 16 8 3 6 3 

Myosotis scorpioides Bora N n -0.77 57 p hc Hy Stoll 7 4 2291 662 3 3.5 14.7 1037 11, 14 7 9 6 6 

Myosotis secunda Bora N n 0.52 55 p hc Stoll 7 1 1736 498 6 3.3 14.0 1237 11 6 9 5 4 

Myosotis sicula Bora N o 15 a Th 0 9 1 0 0 2 6.2 17.0 794 19 6 7 6 3 

Myosotis stolonifera Bora N s 0.77 20 p hc Stoll 4 1 115 0 0 1.6 13.2 1362 11, 14 8 9 5 4 

Myosotis sylvatica Bora N n 2. 18 47 p hc 0 7 5 1690 24 4 3.4 15.1 931 1 6 5 7 5 

Myosoton aquaticum Cary N n 0.00 100 p hc Nodal 7 4 927 0 0 3.6 16.0 747 11, 13, 14 7 8 7 8 

Myosurus nnnimus Ranu NA n -0.66 8 a Th 0 7 3 339 0 2 3.8 16.3 691 4 8 7 6 5 

Myrica gale 

Myri N n -0.75 150 p Pn 

Rhiz2 

5 2 

976, 553 0 3.4 13.8 1353 12 

8 9 3 2 

Myriophyllum alternifl orum Halo N n 1.00 120 p Hy 

Irreg 

5 2 

1390 327 3 3.2 13.8 1299 13, 14 7 12 5 3 

Myriophyllum aquaticum Halo AN , 

200 p Hy 

Irreg SAm 268 2 7 4.3 16.1 838 13 

7 12 5 3 

Myriophyllum spicatum Halo N n 0.63 250 p Hy 

Irreg 7 5 

1409 373 7 3.7 15.1 907 13, 14 7 12 7 7 

Myriophyllum verticillatum Halo N n -0.89 300 p Hy Irreg DRa 7 6 360 130 0 3.8 15.7 765 13, 14 7 12 7 7 

Mynt is odorata 

Apia AN 

-0.25 180 p hc 

0 

Eur 1152 147, 0 3.0 14. 1 1084 3, 17 

7 6 7 71 

Naj as flexilis 

Naja N s 0.48 30 a Hz 

0 4 6 

28 28 0 4.1 13.8 1338 13 

6 12 7 4 1 

Naj as marina Naja N VU r 97 a Hz 0 8 6 c 4 0 0 3.9 16.1 600 13 5 12 9 6 0 

Narcissus pseudonarcissus Lili N n 0.87 35 p Gb Otb 8 2 646 0 3 3.7 15.6 920 3 7' 5 6 5 0

Fam NS CS RS Chg_ Hght Len 1 P2 LF1 LF2 W Clonal Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N S 

Lili Poac N n -0.32 -0.68 45 40 he hc 0 Rhizl 5 3 1 2051 1628 560 716 3 0 33.3 2 14.0 13.8 1226 1296 8 12 79 32 21 0 

Orch N EX n x -0.91 47 30 Gn 0 7 9 4 1 742 1 99 24 0 3.5 4.7 15.2 14.8 1107 938 7, 1 16 4 87 25 0 

Lami AR -1.23 100 he Rhizl 7 4 478 6 1 3.7 15.9 761 3, 7 4 7 6 0 

Nymp N n -0.13 150 Hy Rhizt 5 4 1140 452 0 3.6 15.1 975 13, 14 11 7 6 

Nymp N s 0.87 150 Hy Rhizl 4 6 68 0 0 1 7 12.5 1680 13 11 6 4 

Nymp N n 1.02 150 Hy Rhizl 7 3 1511 340 5 3.6 14.8 1090 13 11 6 4 

Meny N s 2.81 200 Hy Irreg Rhiz2 7 5 45 0 0 3.5 16.4 603 13, 14 11 7 6 

Apia Scro N n -0.35 -0.46 150 50 Th 0 7 5 2320 865 7 3.7 14.7 1047 6 5 6 5 0 

p Hz Hy 0 7 4 505 183 0 3.8 15.6 792 11 10 7 6 

Apia N n -0.04 150 he Hy 0 8 2 1599 633 12 4.0 14.8 1124 11, 14 ' 9 6 7 

Apia N n -1.18 0.19 100 80 Hy hc Hy 0 Node2 7 3 1 804 235 101 45 6 0 3.9 3.7 15.8 16.0 771 718 11 14 10 9 78 6 0 

Apia N n -0.36 0.48 100 hc 0 9 8 2 1 241 551 118 0 0 3 4.3 4.4 15,4 16.2 957 849 6 11 87 86 53 3 0 

Apia N s 0.37 100 hc 0 8 3 76 0 0 3.7 16.3 661 6 9 7 5 0 

Onag AN 1.02 100 hc 0 Am, SAm 1185 38 11 3.9 15.8 839 3, 16, 17, 19 4 6 4 0 

Onag AN 100 hc 0 Am? 614 9 2 3.9 15.9 777 3, 16, 17, 19 4 6 4 0 

Onag AX 100 hc 0 125 2 1 3.7 15.9 798 3, 16, 19 3 6 5 0 

Onag AN 80 he 0 Am? 217 0 4.2 15.9 921 3, 16, 17, 19 4 6 3 0 

Onag AN 100 he 0 Am 966 24 1 4.0 15.9 832 3, 16, 19 4 6 5 0 

Faba N NA VU n r -0.76 0.27 60 15 a Th he 0 9 7 4 1 265 10 0 5,6 3,6 15.8 16.2 999 710 Co 7 1 8 42 8 32 0 1 

Faba N n -0.45 60 Ch he sw Rhiz2 7 3 1664 178 1 3.8 15.3 878 7 4 6 3 0 

Aste Faba AR N n -0.82 0.66 200 70 Ch hc hc sw 0 87 4 724 778 0 5 3.6 3.7 16.0 15.9 718 733 6, 3, 7 17 4 86 37 0 

Ophi N s 10 Gn Root 5 2 72 12 4.3 13.3 1252 8 6 5 2 1 

Ophi N VU r 4 Gn Root 9 1 1 0 6,7 16.2 848 8 6 5 2 

Ophi N n 30 Gn Root 7 6 1474 209 3.6 15.1 964 6, 7 7 7 3 

Ophi N n 0.72 30 Gn Root 7 6 1474 209 3.6 15.1 964 6 7 7 3 

Orch N VU n r 0.83 45 35 Gn 0 9 2 936 6 182 0 3,9 4.0 15.7 16.4 807 766 7 4 89 23 

Orch N n -1.34 60 Gn 0 7 3 264 31 3.7 15.8 803 1. 7, 11 5 9 2 

Taxon name 

Nardus stdcta 

Narthecium ossifragum 

Neotinea maculata 

Neottia nidus-avis 

Nepeta cataria 

Nuphar lutea 

Nuphar pumila 

Nymphaea alba 

Nymphoides pe/tata 

Odontites vernus 

Oenanthe aquatica 

Oenanthe crocata 

Oenanthe fistulosa 

Oenanthe fluviatilis 

Oenanthe lachenalii 

Oenanthe pimpinelloides 

Oenanthe silaifolia 

Oenothera 

Oenothera biennis 

Oenothera biennis x 

glazoviana (0 . x fallax) 

Oenothera cambrica 

Oenothera glazioviana 

Onobrychis 

Orionis reclinata 

Orionis repens 

Ononis spinosa 

Onopordum acanthium 

Ophioglossum azoricum 

Ophioglossum lusitanicum 

Ophioglossum vulgatum 

Ophioglossum vulgatum 

sens.lat 

Ophrys apifera 

Ophrys fuciflora 

Ophrys insectifera 

Ophrys sphegodes 

Orchis laxiflora 

Orchis mascula 

Orchis militaris 

Omhis morio 

Orchis purpurea 

Orchis simia 

Orchis ustulata 

Oreopteris limbosperma 

Origanum vulgare 

Orch N s o -Oi l 20 50 Gn 0 98 2 4 c 62 0 0 8 1 4.0 6.4 16.3 16.7 729 800 6, 7 11 49 98 3 2 

Orch N n -0.72 40 Gn 0 7 3 1962 475 9 3.5 14.7 1073 1, 7, 16 5 7 4 

Orch N VU n r -0.98 45 20 Gn 0 7 4 3 c 931 19 124 0 0 8 3,4 3.9 16.3 15.8 694 801 6, 7 7 43 7 9 3 2 

Orch N s -0.56 50 Gn 0 7 3 36 0 1 4.0 16.5 748 1 4 8 3 

Orch N VU 

Orch N s r -1.77 30 15 Gn 0 87 3 265 10 0 0 3.8 3.5 16.4 15.7 697 777 7 34 8 2 

Thel N n -0.18 90 hc 0 7 3 1585 138 0 2.8 13.7 1323 1, 16 6 4 3 

Lami N n -0.10 65 Ch hc 0 Nodal 8 5 1148 179 0 3.7 15.3 898 7, 16 4 7 4

S 

P2 LF1 LF2 W Clonel Clone2 E1 E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N 

Gb Mb DRg 8 3 Eur 1139 8 12 3.7 15.5 848 1, 3, 7, 8 8 3 6 4 

Th Gb Otb 0 9 7 2 1002 33 20 0 12 0 3,6 3.9 15.5 16.2 773 935 8 1, 3 5 71 5 4 7 5 

4 3 

p Th Gn 0 8 7 2 3 92 3 42 0 7 2 4.0 7.0 16.4 13.7 1493 832 8, 16 10 8 3 3 7 2 

3 6 0 0 4.5 16.5 753 Co 18 8 4 8 5 

34 0 2671 5 0 0 4.1 3.6 16.4 16.2 782 713 3, 18 7 8 3 98 2 3 

2 160 122 11 4.9 15.5 1004 1, 3, 16 4 5 6 5 

3 800 0 14 4.0 16.1 774 4, 5 7 4 8 6 

3 25 0 12 4.9 16.3 776 3, 6 7 4 7 2 

2 422 30 4 4.1 15.7 890 10 7 5 3 2 

3 c 71 0 0 3.1 15.3 666 3, 7 71 6 71 6 

6 228 7 0 1.4 12.2 1568 2, 10, 16 5 51 5 3 

2 735 576 4 4.2 14.5 1225 1, 11 6 9 5 4 

5 1 2393 23 791 6 2 3 3.4 5.1 14.4 16.0 1138 874 Co 1, 19 2, 16 4 9 26 4 5 2 4 

8 

7 

7 

9 

8 

7 

8 

7 

4 

8 

9 

5 

SAm 541 65 14 4.4 15.8 861 3, 17, 19 7 3 4 2 

Unk 998 32 11 3.9 15.8 843 3, 17 7 4 6' 5 

SAm 137 10 12 4.3 16.1 784 3, 17 7 4 6 8 

Aus, NZ 323 11 8 3.8 15.7 782 3, 17 7 4 6 4 

SAT 146 5 11 4.6 15.8 927 3, 17 6 5 6 5 

SAm 67 1 11 5.1 16.1 892 3, 17 6 5, 6 5 

SAl 10 0 10 5.9 16.1 861 3, 4 7 4 6 5 

As2, Am 396 27 4 3.8 15.7 895 3, 17 6 5 6 5 

6 311 24 0 1.7 12.1 1940 15, 16 7 6 5 3 

5 c 3 0 0 0.4 11.5 1392 7 9 4 8 2 

3 16 0 0 2.5 12.9 1141 7, 18, 19 9 3 8 2 

Crop 359 6 6 4.0 16.0 787 1, 4, 17 9 3 7 6 

3 874 48 6 3.7 15.7 759 4 7 4 6 5 

4 1875 402 13 3.7 15.0 910 3, 4 7 5 6 5 

2 357' 33 6 4. 1 15.9 776 4 7 4 8 4 

3 1712 370 14 3.8 15.3 877 3, 4 71 5 71 6 

3 Eur? 1600 190 12 3.7 15.3 868 3, 4, 17 7 4 7 8 

1 109 / 5 4.4 16.4 709 Co 18, 19, 21 9 6 7 4 

2 1 347 165 101 63 11 5 5.1 4.5 15.4 15.7 1129 876 Co 3, 21 5 7 8 67 7 6 5 

2 1336 3171 14 4.1 15.6 865 16, 17 71 4, 8 5 

4 714 0 0 3.2 15.3 877 1 3 61 7 6 

6 1091 302 0 3.0 13.7 1236 11 8 8 7 3 

4 1011 0 7 3.8 16.0 774 3, 6, 7 7 4 7 5 

3 1744 583 1 3.3 14.0 1226 11 8 8 5 2 

3 2118 744 9 3.4 14.1 1202 10, 12, 14 8 8 3 2 

Eur 1753 88 11 3.6 15.2 915 1, 3, 17 6 5 6 7 

6 2013 644 11 3.7 14.9 986 11, 13 7 10 6 6 

0 

0 

0 

0 

0 

0 

0 

0 

Rhiz1 

0 

Rhiz2 

Rhiz1 

Rhiz1 

Node2 

DRg 

Node2 

DRa 

DRg 

DRg 

Rhiz1 

0 

0 

0 

0 

0 

0 

0 

0 

0 

0 

0 

0 

0 

Rhiz2 

0 

0 

0 

0 

0 

Rhiz2 klieg 

p Th Gn 

a Gn Th 

Gn 

a Gn Th 

p Th Gn 

a Gn Th 

Gn 

a Gn Th 

Ch 

hc 

hc 

hc 

hc 

a hc Th 

Gb 

a hc Th 

Gb 

Gb 

Gb 

Th 

hc 

hc 

hc 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

hc 

Gn 

hc 

hc 

hc 

hc 

hc 

Hy hc 


Ornithogalum angustifolium Li li AN : 1.05 30 

Ornithogalum pyrenaicum Li li N s 0.14 75 

Ornithopus perpusillus Faba N n -0.18 10 

Ornithopus pinnatus Faba N r 8 

Orobanche alba Orob N s -0.38 25 

Orobanche artemisiaecampestris 

Orob N EN r 60 

Orobanche caryophyllacea Orob N VU r 0.01 40 

Orobanche elatior Orob N n -0.33 75 

Orobanche he derae Orob N n 0.20 60 

Orobanche minor Orob N n -0.20 60 

Orobanche purpurea Orob N VU r 0.50 45 

Orobanche rapum-genistae Orob N s -0.35 85 

Orobanche reticulata Orob N r 60 

Odin ha s ecunda Pyro N n -0.40 5 

Os m unda regalis Osmu N n 0.56 160 

Otanthus maritimus Aste N EX x -1.49 30 

Oxalis acetosella Oxa N n -0.74 10 

Oxalis art iculata Oxa AN 25 

Oxalis corniculata Oxa AN 1.62 15 

Oxalis de bilis Oxa AN 20 

Oxalis exilis Oxa AN 5 

Oxalis incarnata Oxa AN 20 

Oxalis latifolia Oxa AN 20 

Oxalis pes-caprae Oxa AN 30 

Oxalis stricta . Oxa AN -0.09 40 

Oxyria digy na Poly N n -0.71 30 

Oxytropis campestris Faba N VU r. 20 

Oxytropis hail ed Faba N r 0.16 13 

Panicum mil aceum Poac AC 100 

Papaver argemone Papa AR -1.79 45 

Papaver dubium Papa AR 0.23 60 

Papaver hybridum Papa AR -0.35 50 

Papaver rhoeas Papa AR -0.41 60 

Papaver s om nife rum Papa AR . 2.54 100 

Parapholis incurva Poac N s, 0.09 10 

Parapholis strigosa Poac N n 0.14 25 

Parentucellia viscosa Scro N n 0.64 50 

Parietaria j udaica Urti N n1 0.08 52 

Paris guadrifolia Lili N n1 -0.68 40 

Parnassia palustris Saxi N n; -0.84 20 

Pastinaca sali va Apia N ni -0.39 180 b 

Pedicularis palustris Scro N n -0.88 60 b 

Pedicularis sylvatica Scro N n -1.28 25 P 

Pentaglottis sempervirens Bora AN , 1.81 100 p 

Persicaria amphibia Poly N n 0.27 60 200 p

F R N S 

77 6 6 

6 71 7 

CI Tja n 

0 3.3 

9 3.7 

10 3.7 

12 3.6 

1 3.6 

1 3.8 

0 1.6 

0 3.8 

0 2.8 

12 4. 1 

2 3.6 

5 5.3 

0 3.8 

5 5.3 

10 4.2 

3 4.1 

IR 

43 

740 

434 

934 

93 

31 

5 

145 

10 

582 

569 

0 

0 

0 

42 

0 

GB 

1355 

2041 

1881 

2578 

296 

203 

446 

390 

335 

1328 

1825 

6 

5 

6 

401 

482 

10 

184 

47 

2449 

1102 

80 

10171 

33 

219 

1927 

26 

2294 

2429 

2182 

2115 

3 

14 

55 

8 

1460 

1144 

1191 

885 

1343 

3 

LF2 W Clonal Clone2 E1 E2 C Origin 

Rhiz1 5 5 

0 7 6 

0 8 6 

0 7 5 

0 7 5 

0 7 3 

Rhizl 2 6 

Rhiz2 As 1 

Rhiz2 Eur 

Rhiz2 Eur 

Rhiz2 7 3 

0 8 2 

0 7 3 Eur 

0 8 3 

0 Crop 

0 8 2 

0 8 3 

0 Eur 

0 5 4 c 

Gn Rhiz2 5 6 

0 Eur? 

0 9 1 Eur 

Rhiz2 5 6 

Rhir / 2 6 

0 8 3 

0 8 3 

0 7 4 c 

0 7 4 

0 8 4 

Gn Rhiz2 6 6 

0 7 3 

w 0 1 3 

h 0 7 3 

h 0 5 3 

h 0 7 3 

w 0 4 5 c Eur, As 

w 0 Am4 

he h 0 8 3 

h 0 7 5 

h Rhiz2 Sto12 4 3 Eur 

h Sto12 5 3 c 

LF1 

hc 

Th 

Th 

Th 

Th 

Th 

he 

Gn 

Gn 

Gn 

Gn 

Th 

Th 

Th 

hc 

hc 

hc 

he 

he 

Hy 

Th 

Th 

Gn 

hc 

Th 

hc 

hc 

hc 

hc 

Hy 

he 

Ch 

hc 

hc 

hc 

Ph 

Ph 

Th 

hc 

hc 

hc 

1 P2 

6 8 65 7 8 

8 6 9 

6 6 2 

4 5 6 

5 5 7 

5 6 6 

7 7 7 

2 6 1 

3 5 2 

3 6 2 

4 7 5 

5 8 6 

5 8 4 

5 71 7 

ChQ Hght Len 

-0.44 80 

-0.41 75 

-0.04 100 

-0.95 80 

-0.06 40 

-0.90 75 

-0.58 30 

0.59 150 

0.01 70 

0.80 30 

-0.15 120 

50 

50 

-0.68 50 

-0.34 75 

0.12 100 

0.29 200 

0.03 100 

-0.07 150 

0.23 200 

-0.32 120 

45 

-0.22 40 

-0.30 50 

-0.56 17 

50 

-0.10 60 

150 

-0.33 100 

0.43 270 

0.45 60 

20 

0.07 120 

-0. 16 50 

-0.73 80 

4600 

5500 

0.771 80 

-0.06 95 

20 

13 

RS 

n 

n 

n 

n 

n 

s 

n 

n 

r 

n 

0 0 4.1 

21 0 2.6 

0 3.6 

6 3.6 

10 3.7 

6 4.1 

8 0 2.4 

0 -0.7 

6 9 4.5 

8 4 3.5 

0 3.3 

3 3.5 

10 3.5 

11 3.7 

14 3.8 

0 -1.1 

0 5.4 

0 3.9 

0 4.1 

2 3.2 

1 3.2 

10 3.9 

8 3.8 

6 4 3.4 

0 3.5 

n 

r 

9 7 5 7 

4 7 6 

5 6 5 

6 4 4 

5 6 4 

3 5 3 

s 

n 

75 

6 

n 

s 

n 

n 

r 

n 

n 

n 

n 

r 

r 

s 

r 

Taxon name Fam NS CS 

N 

N 

N 

N 

N 

N 

N 

AN 

AN 

AN 

N 

N EN 

AN CR 

N 

AR 

N 

N 

AR 

N 

N 

AN 

AN 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N VU 

N VU 

N 

NA VU 

AN 

AN 

AR 

N 

AN 

N 

-C. 

G.% 

43 78 4 2 

5 71 6 

Br Habitats 

6 

11, 13, 14 

4, 11 

3, 4 

13, 14 

13, 14 

7, 15 

3 

1, 3 

3 

14 

19 

8 

8, 19 

3, 17 

3, 4 

19 

5 

11 

11, 14 

3, 17 

4, 17 

1, 16 

11, 15 

19 

3, 6 

8 

3, 6 

3, 6 

11 

1, 16 

10, 15 

1, 10 

7 

1, 3 

2 

2 

3, 4 

7 

3, 17 

3, 16 

Prec Co 

1021 

1093 

958 

1077 

1003 

780 

1588 

1149 

1000 

941 

1002 

774 Co 

613 

774 

819 

751 

599 Co 

1127 

631 

1044 

856 

760 

1515 

1770 

918 Co 

942 

621 

1023 

1047 

1055 

1048 

1829 

1279 

806 

826 

1054 

1223 

795 

781 

985 

1153 

Tjul 

14.8 

14.8 

15.1 

14.6 

15.2 

15.9 

12.2 

14.8 

13.9 

15.3 

14.8 

16.7 

16.5 

16.7 

15.8 

16.2 

16.7 

13.5 

16. 1 

14.6 

15.5 

15.9 

13.1 

10.8 

15.3 

15.1 

16.2 

14.8 

14.7 

14.7 

14.8 

10.6 

15.5 

16.3 

16.4 

14.7 

14.1 

15.9 

16.0 

14.9 

11.5 

10 5 7 6 

67 

71 

78 

93 

54 73 5 2 

5 4 4 

4 8 3 

5 6 5 

6 3 4 

7 2 2 

5 71 6 

12 

21 

2 

b 

4 86 3 2 

4 7 4 

5 6 3 

n 

n 

4 7 4 

16 

1153 

11.5 

0 3.5 

3 

h Sto12 4 1 

hc 

13 

r 

NE VU 

4 7 2 

3 

7 

786 

1094 

14.9 

14.5 

0 3.0 

12 3.5 

89 

72 

2629 

h Sto12 Eur 

h Sto12 7 3 

hc 

hc 

13 

-0.59 12 

n 

AN 

N

CI Tja n 

12 5.2 

3 3.6 

0 3.6 

0 3.6 

0 3.0 

0 5.1 

0 3.9 

0 3.0 

2 3.3 

6 3.7 

0 0.8 

2 4.0 

14 4.1 

14 3.5 

14 3.5 

8 3.8 

0 3.5 

1 3.5 

0 3.4 

0 0.6 

6 3.7 

14 3.5 

6 4.6 

1 3.1 

2 3.6 

0 -0.6 

0 1.1 

5 3.4 

13 5.9 

5 3.2 

2 3.7 

10 3.5 

2 3.6 

11 3.6 

0 1.9 

14 6.3 

1 2.8 

0 3.7 

11 3.4 

3.5 

3.6 

3.1 

1.2 

1 3.7f 

3.7 

1 3.6 

IR 

0 

24 

109 

391 

0 

74 

309 

484 

116 

25 

0 

10 

303 

985 

975 

405 

0 

308 

251 

2 

0 

985 

0 

3 

0 

0 

0 

297 

0 

0 

12 

928 

423 

903 

0 

0 

0 

0 

717 

665 

0 

0 

0 

480 

819 

906 

GB 

11 

312 

584 

1938 

1 

0 

500 

1523 

444 

1009 

65 

137 

1445 

2804 

2766 

1295 

1243 

949 

1163 

72 

83 1 

2792 

72 

169 

1063 

8 

62 

1865 

54 

1960 

132 

2766 

1637 

2721 

16 

16 

18 

153 

2150 

2176 

268 

48 

10 

19371 

2030 

2605 

Origin 

Clone2 El E2 C 

5 2 

7 2 

7 3 

' 7 4 

1 4 

7 1 

7 1 

4 6 

W Clonel 

h Sto12 

h Rhiz2 

h 0 

h 0 

h O 

h O 

h o 

h 0 

w 0 

w 0 

w 0 

0 

0 

Rhiz1 

0 

0 

Rhiz1 

0 

0 

0 

Rhiz2 

0 

0 

0 

Rhiz2 

0 

0 

Rhiz2 

0 

0 

0 

Rhiz2 

Rhiz2 

0 

0 

0 

0 

0 

0 

0 

Rhiz2 

Rhiz2 

Rhiz2 

0 

0 

0 

LF2 

LF1 

hc 

Hy 

hc 

hc 

hc 

hc 

hc 

hc 

Ph 

Ph 

Ph 

Th 

hc 

hc 

he 

hc 

hc 

Gn 

Gn 

hc 

hc 

Th 

hc 

hc 

hc 

hc 

hc 

he 

Th 

he 

he 

hc 

hc 

hc 

hc 

Th 

Ch 

Ch 

Ch 

Ch 

Gn 

Gn 

Gn 

Th 

Th 

Th 

1 P2 

Hght Len 

9 

10 10 

100 

70 

5 

8 

3 

8 

2500 

4200 

3000 

200 

6 

15 

15 

15 

6 

40 

50 

40 

70 

20 

35 

100 

50 

22 

40 

30 

10 

75 

100 

58 

75 

70 

90 

25 

10 

10 

15 

25 

80 

40 

80 

20 

30 

30 

Chg 

0 

0 

-0.03 

-0.16 

-0.31 

0 

0 

0 

0 

0 

0 

0 

0 

0 

2 

0 

0 

3 

0 

0 

0 

0 

0 

1 

R N 

8 2 

4 2 

7 6 

7 3 

8 2 

4 2 

4 2 

6 2 

5 2 

5 2 

2 2 

7 7 

6 4 

6 4 

6 7 

6 4 

7 3 

6 2 

71 4 

7 3 

7 5 

6 7 

5 2 

6 5 

7 4 

3 2 

6 3 

6 4 

5 5 

6 5 

7 6 

6 5 

6 5 

6 6 

7 6 

6 4 

9 1 

8 2 

2 2 

6 3 

7 6 

7 3 

5 5 

71 6 

6 7 

7 6 

F 

3 

10 

5 

4 

8 

8 

8 

8 

5 

3 

6 

4 

6 

5, 

5 

7 

4 

6 

5 

5 

5 

5 

3 

5 

4 

5 

5 

6 

4 

5 

9 

5 

5 

6 

5 

4 

6 

3 

71 

5 

5 

3 

5 

5 

51 

5 

L 

8 

8 

7 

71 

9 

7 

8 

8 

7f 

7 

7 

7 

8 

71 

7 

8 

8 

6 

5 

7 

7 

7 

8 

5 

9 

6 

7 

8 

8 

4 

7 

7 

7 

7 

5 

9 

9 

7 

8 

8 

4 

5 

4 

71 

7 

7 

Br Habitats 

7, 16 

13 

6 

7 

11 

10, 12 

10, 11 

11, 12 

2 

1, 2, 17, 19 

2 

3, 4 

3, 6, 18 

6, 7 

3, 5 

15, 21 

7 

1, 10 

1, 6 

15 

7, 8 

3, 4, 5, 6 

19 

1 

3 

15, 16 

15, 16 

6, 19 

3 

1 

11, 13, 14 

3, 5, 6, 7 

3, 5, 6, 7 

1, 3, 6 

16 

3, 4, 17 

7 

7 

8, 10, 12 

7 

1 

1, 16 

1 

3, 4 

4, 17 

3, 4, 17 

Prec Co 

871 

1040 

819 

958 

752 

1270 

1475 

1305 

1302 

889 

1930 

896 

1065 

1105 

1102 

1248 Co 

804 

1235 

1147 

2142 

790 

1105 

741 Co 

959 

850 

1907 

2252 

1134 

935 

1015 

948 

1102 

1017 

1095 

1246 

923 

1037 

783 

1200 

1114 

829 

1091 

1061 

1008 

1039 

1073 

Tjul 

16,1 

14.8 

15.5 

15.0 

13.6 

14.7' 

13.7 

13.6 

14.0 

15.5 

11.7' 

15.6 

14.8 

14.5 

14.5 

14.0 

15.6 

14.3 

14.7 

11.5 

15.8 

14.5 

16.4 

14.5 

15.6 

10.5 

11.6 

14.3 

16.3 

14.8 

15.2 

14.5 

14.8 

14.5 

13.9 

16.3 

15.0 

16.2 

14. 1 

14.4 

16.0 

15.1 

13.1 

14.9 

14.8 

14.6 

-0.83 

-0.76 

RS 

e 

S. 

n 

n 

x 

o 

n 

n 

Am4 

Eur 

0.40 

' 

s. 

Eur 

4 5 

0 3 

8 4 

8 4 

6 5 

3 4 

7 5 

5 5 

7 3 

DRi 1 6 

8 6 

6 4 

DRi 8 4 

0.16 

1.35 

0.09 

-0.28 

-0.79 

-1.67 

-0.88 

-0.31 

hc 

p 

n 

n 

n 

n 

n 

n 

n1 

s 

ni 

e 

s: 

0.83 

0.63 

-0.05 

0.21 

7 3 

1 3 

2 6 

7 ? 

9 1 

5 6 

5 6 

6 6 

6 6 

Nodel 6 4 

5 4 c 

9 1 

5 3 c 

8 1 

7 2 

7 3 

7 3 

7 5 

4 3 

6 5 

6 6 

6 6 

n. 

r : 

s 

n. 

s' 

n 

n. 

n 

n 

r 

r ' 

r , 

n. 

n 

n 

n 

s 

r 

Taxon name Fam NS CS 

N VU 

N 

N 

N 

NA EX 

N 

N 

N 

AN 

AN 

N 

AC 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N 

N 

AN 

N 

N VU 

N 

N 

N 

N 

AN 

N 

N 

N 

N 

NA 

N VU 

N 

N 

N 

N 

N 

N VU 

AR 

N 

N 

Eur 

-0.48 

NHem 

1.33 

0.27 

-1.55 

0.60 

1.10 

1.17 

-0.04 

-0.10 

-0.37 

-0.50 

-1.14 

0.27 

0.34 

-0.70 

n: 

n1 

I

Taxon name Fam NS CS RS Chg_ Hght Len P1 P2 LF1 LF2 W Clonal Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N S 

Polygonum boreale Poly N $ 20 a Th 0 4 2 109 0 0 3.6 12.4 1084 4 7 5 6 0 

Polygonum maritimum Poly N EN r 0.21 20 p Ch 0 9 1 16 1 4 6.0 16.2 950 Co 19 9 3 4 3 

Polygonum oxyspermum Poly N n 0.01 20 a Th 0 6 3 305 75 9 4.6 14.8 1102 Co 19 9 6 8 3 

Polygonum runvagum Poly AR 20 a Th 0 7 3 274 0 4 3,9 16. 1 738 4 8 4 5 0 

Polypodiurn cambricum Poly N n 40 P hc Gn Rhizl 9 1 196 157 1 4.4 15.1 1102 3, 16 6 5 3 0 

Polypodiurn inte rje ctum Poly N n 40 p hc Gn Rhizl 7 2 1188 487 12 3.9 14.9 1074 1, 3, 16 5 5 3 0 

Polypodium vulgare Poly N n 40 P hc Gn Rhizl 5 3 1795 523 1 3.5 14.3 1158 1, 3, 16 5 5 3 0 

Polypodiurn vulgare sens.lat Poly N n -0.03 40 p hc Gn Rhir l 6 3 2496 916 12 3,5 14.4 1133 1 5 5 3 0 

Polypogon rnonspeliensis Poac N $ 0.60 80 a Th 0 9 1 45 0 0 4.3 16.6 661 6 8 8 6 3 

Polystichum aculeaturn Dryo N n 0.54 60 p hc 0 7 5 1618 314 3 3.2 14.4 1145 1, 16 5 5 5 0 

Polystichurn fonchitis Dryo N n -0.76 30 p hc 0 4 6 181 20 0 1.4 12.1 1857 7, 15, 16 6 5 3 0 

Polystichurn setiferurn Dryo N n 1.47 120 p hc 0 9 2 1249 748 10 4,1 15.0 1032 1 4 5 6 0 

Populus alba Sail AN 2000 p Ph w Root Eur, As 1 1531 116 10 3.6 15.3 867 3, 17, 19 6 6 6 0 

Populus alba x tremula (P. x I 

cane s ce ns) Sali AN 0.97 3000 p Ph w Root Eur 1145 110 7 3.8 15.6 819 1, 3 6 6 5 0 

Populus nigra sens.lat Sali N n 0.65 3000 p Ph w Root 7 4 692 66 2 3.6 15.7 800 3, 14 6 8 7 0 

Populus tremula Sall N n 0.88 2000 p Ph w Root 5 5 2248 414 7 3.3 14.5 1102 1, 16 6 5 6 0 

Potarnogeton acutifolius Pota N VU r 0.05 100 p Hy DRa 7 3 35 0 0 4.0 16.4 702 13 7 12 6 0 

Potamogeton alpinus Pota N n 0.30 280 p Hy lrreg DRa 4 6 5371 142 1 3.1 14.1 1188 13 71 12 5 1 

Potamogeton be rchto ldii Pota N n 1.66 60 p Hy DRa 5 6 1461 317 3.6 14.7 1027 13, 14 7 12 5 0 

Potarnogeton co/oratus Pota N s 0.03 70 p Hy Meg Rhiz2 8 3 142 140 4.1 15.1 893 13 7 11 5 0 

Potamogeton compressus Pota N s -1.68 90 p Hy DRa 5 5 134 0 3.4 15.9 697 13 7 12 4 - 

Potamogeton crispus Pota N n 150 p Hy Rhiz2 DRa 8 5 1541 358 3.7 15.2 893 13, 14 7 12 6 / 

Potamogeton epihydrus Pota N VU r 0.11 190 p Hy lrreg DRa 5 0 2 0 4.5 13.0 1288 13 8 12 / 0 

Potamogeton filiformis Pota N s 0.63 30 p Hy Rhiz2 DRg 4 6 161 63 3.6 13.3 1109 13 7 12 5 1 

Potamogeton friesii Pota N $ -1.06 150 p Hy DRa 5 6 270 43 3.6 15.4 782 13 7 12 5 0 

Potamogeton gramineus Pota N n 0.67 80 p Hy lrreg Rhiz2 5 6 473 163 3.3 13.8 1170 13, 14 7 12 3 0 

Potamogeton grarnineus x 

lucens (P. x zizii) Pota NH S 120 p Hy Meg Rhiz2 100 92 3.5 14.3 1138 13, 14 7 12 4 0 

Potamogeton grarnineus x 

perfoliatus (P. x nite ns) Pota NH n 250 p Hy Irreg Rhiz2 216 113 3.5 13.6 1257 14 7 12 5 1 

Potarnogeton lucens Pota N n 0.25 250 p Hy Irreg Rhiz2 7 4 456 171 I 3.7 15.5 832 13, 14 7 12 6 0 

Potarnogeton natans Pota N n 100 p Hy Irreg Rhiz2 5 6 2340 648 6 3.5 14.5 1091 11, 13, 14 7 11 4 0 

Potamogeton nodosus Pota N r -0.18 250 p Hy Me g Rhiz2 8 6 15 0 0 3.9 16.4 761 14 6 12 5 0 

Potamogeton obtusifolius Pota N n 0.96 190 p Hy DRa 5 6 601 155 0 3.4 14.7 994 13 7 12 5 0 

Potarnogeton pectinatus Pota N n 230 p Hy Rhiz2 DRg 6 6 1165 242 4 3.8 15.3 844 13, 14 6 12 7 2 

Potarnogeton perfoliatus Pota N n 300 p Hy Meg Rhiz2 5 6 1056 275 0 3.5 14.6 1026 13, 14 7 12 5 1 

Potam ogeton polygonifolius Pota N n 10 70 p Hy hc Irreg Rhiz2 7 2 1841 582 5 3.3 13.9 1254 11, 12, 13 8 10 2 0 

Potamogeton praelongus Pota N n -0.26 300 p Hy lrreg Rhiz2 4 6 252 90 0 3.2 14.1 1178 13 7 12 5 1 

Potamogeton pusillus Pota N n 0.77 70 p Hy DRa 8 6 806 146 1 3.6 15.1 877 13 7 12 6 1 

Potamogeton rutilus Pota N r 0. 18 45 p Hy DRa 4 3 13 0 0 3.8 12.9 1115 13, 14 7 12 5 0 

Potarnogeton trichoide s Pota N n 0.57 100 p Hy DRa 8 4 185 0 1 3.8 16.2 739 13, 14 6 12 6 0 

Potentil la anglica Rosa N n 0.11 25 p hc Sto12 7 3 1248 733 4 4.0 15.0 1045 6 7 5 5 0 

Potent) Ila ans erina Rosa N n -0.23 25 p hc Sto12 5 6 2662 973 12 3.6 14.6 1083 6 8 7 6 2 

Potentilla argentea Rosa N n -0.78 30 p hc 0 7 4 323 0 5 3.6 16.0 704 3, 8 8 3 2 0 

Potent" Ila crantzir Rosa N s -0.21 20 p hc 0 2 4 98 0 0 0.7 12.0 1855 7, 16 8 5 2 0 

Co

, 

Taxon name Fam NS CS RS Chg Fight Len 1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N S 

Potent/ ha erecta Rosa N n -0.50 25 hc h 0 5 4 2700 947 12 3.5 14.4 1118 8 7 7 3 2 

Potentilla fruticosa Rosa N r 1.44 100 Pn w 4 6 8 7 0 2.8 13.7 1366 16 8 6 8 2 

Potentilla ne um anniana Rosa N s -0. 17 10 hc h 0 7 3 128 0 0 3.1 14.7 994 7, 16 7 3 8 1 

Potentilla palustris Rosa N n -0.21 50 Gn Hy h Rhiz2 5 6 1672 721 'I 3.4 14.0 1186 11 8 9 5 3 

Potentilla reptans Rosa N n -0.62 30 hc h Sto12 8 4 1882 769 11 3.8 15.1 959 6 71 5 7i 5 

Potent/ Ila rupestris Rosa N VU r 60 hc h 0 9 3 4 01 0 3.0 13.8 934 16 7 4 6 2 

Potentilla sterilis Rosa N n -0.30 15 hc h Stoll 7 2 2167 742 8 3.6 14.7 1071 1 5 5 5 5 

Primula elatior Prim N s 0.01 20 hc h 0 7 3 c 38 0 0 3.3 16.3 591 1 4 5 7 6 

Primula farinosa Prim N s -0.46 5 hc h 0 4 5 103 0 0 2.2 13.8 1194 7, 11 9 8 9 2 

Primula scotica Prim NE e -0.18 5 hc h 0 4 1 42 0 0 3.6 12.4 1026 Co 6, 19 9 4 71 2 

Primula veris Prim N n -0.32 15 hc h 0 7 4 1632 369 6 3 7 15.2 900 6, 7 7 4 7 3 

Primula vulgaris Prim N n 0.16 15 hc h 0 7 3 2651 934 11 3.5 14.5 1108 1, 16 5 5 6 4 

Prune lla vulgaris Lami N n 0.60 30 hc h Nodel 6 6 2783 970 14 3.5 14.4 1106 6, 7 7 5 8 4 

Prunus avium Rosa N n 1.29 2500 Ph w Root 7 3 2136 431 7 3.4 14.8 1024 1 4 5 6 6 

Prunus cerasifera Rosa AN 3.43 800 Ph w Eur, As 1 900 11 4 3.7; 15.8 803 1, 3, 17 6 5 7 6 

Prunus cerasus Rosa AR 0.90 800 Ph w Root Eur 706 339 7 3.9 15.2 961 1, 3 6 5 6 5 

Prunus domestica Rosa AR 2.19 800 Ph w Root Eur 1656 406 8 3.7 15.2 928 3, 17 7 5 7 6 

Prunus laurocerasus Rosa AN 4.70 600 Ph w Eur 1253 345 4 3.8 15.3 953 1, 17 4 6 5 6 

Prunus lusitanica Rosa AN , 800 Ph w Eur 468 31 1 3.8 15.5 876 1, 17 6 5 7 6 

Prunus padus Rosa N n 0.58 1500 Ph w 5 5 1089; 189 2.71 13.8 1240 1 5 6 6 71 

Prunus spinosa Rosa N n 0.40 400 Ph w Root 7 3 23081 917 3.7 14.8 1032 3 6 5 7 6 

Pseudofumaria lutea Furna AN 0.59 30 hc h 0 Eur 1425 39 3.6 15.4 876 3, 17 6 6 8 5 

-le 

C . Ps e udorchis albida Orch N n -0.88 20 Gn h 0 4 3 385 110 2.6 13.0 1534 7, 8 8 5 6 2 

Pseudotsuga menziesii Pina AN ! 5800 Ph w Am4 900 13 3.4 15.0 1035 1, 2, 17 6 6 4 4 

Pteridium aquilinum Denn N n -0.71 150 Gn Rhiz2 7 6 2685 964 / 3.5 14.5 1109 1, 9 6 5 3 3 

Puccinellia dis tans Poac N n 3.02 60 hc 0 5 4 405 39 4.1 14.9 859 3, 21 8 8 7 7 

Puccinellia fasciculata Poac N s -0.51 50 hc 0 8 2 97 11 4.5 16.4 722 6 8 7 7 7 

Puccinellia maritima Poac N n -0.27 80 hc Node2 5 1 788 201 4.1 14.4 1162 Co 21 9 8 7 6 

Puccinellia rupestris Poac N s -0.40 40 b Th hc 0 8 1 148 0 4.5 16.2 756 Co 6, 19 9 7 7 5 

Pulicaria dysenterica Aste N n -0.08 80 hc Rhiz2 8 4 14971 391 1 4.0 15.5 885 6, 11 71 71 7 4 

Pulicaria vulgaris Aste N VU r -0.551 45 Th 0 7 4 121 0 4.0 16.5 696 8 9 8 6 7 

Pulmonaria longifolia Bora N s -0.01 40 hc 0 7 1 21 0 4.7 16.5 810 1, 3 6 4 6 5 

Pulmonaria obscura Bora N VU r, 

30 

he 

0 7 3 c 

1 0 3.2 16. 1 577 1 

4 6 8 7 

Pulmonaria ofi cinalis Bora AN 

1.77 30 

hc 

0 7 3 c Eur 

682 8 3.4 15.3 922 1, 3, 17 5 5 8 6 

Pulsatilla vulgaris Ranu N s -0.50 30 hc 0 7 3 c 691 0 3.3 16.0 664 7 71 3 8 3 

Pyrola media Pyro N 5 -1.09 10 hc Rhizl 4 4 258 47 2.2 13.0 1197 2, 10 5 4 5 2 

Pyrola rninor Pyro N n -0.55 7 hc Ch Rhizl 4 6 558 50 2.6 13.9 1089 1, 16 5 5 4 2 

Pyrola rotun difolia Pyro N s' -0.08 12 hc Rhizl 5 4 155 13 2.9 14.2 1072 2, 19 6 7 7 3 

Pyrus communis sens.lat Rosa AR 1.49 1500 Ph w Root Eur?, As l ? 781 6 3.8 15.9 801 1, 3, 17 7 5 6 7 

Pyrus cornm unis sens.str. Rosa AR 1500 Ph w Root Gard 3, 17 7l 5 6 71 

Pyrus cordata Rosa NA EN r 400 Ph w Root 7 1 9 0 5.4 15.9 1129 3 6 5 5 4 

Quercus cerris Faga AN 2.32 3500 Ph w Eur 1247 42 1 3.8 15.6 862 1, 3, 17 6 4 6 6 

Quercus ilex Faga AN ' 2.37 2500 Ph w 9 1 Eur 802 22 1 4.1 15.9 824 1, 17, 19 6 3 7 4 

Quercus petraea Faga N e 0.14 3000 Ph w 7 3 1832 549 3.5 14.6 1136 1 6 6 3 4 0 

Quercus robur Faga N n -0.60 3000 Ph w 7 3 2310 663 1 3.5 14.71 1049 1, 3 71 5 5 4 0 

Radio/a linoides Lina N n -0.87 6 a Th h 0 7 3 548 115 I I 4.2 14.7 1061 10 8 7 4 2 1 

1 

I

S 

Taxon name Fam NS CS RS Chg Hght Len P1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N 

Ranunculus acris Ranu N n 0.30 - 75 p hc 0 3 5 2780 968 13 3.5 14.4 1106 6 6 4 

Ranunculus aquatilis Ranu N n 90 a Hz 0 Irreg 7 3 1123 157 4 3.6 15.2 860 13, 14 1 7 5 

Ranunculus aquatilis sens.lat Ranu N n -1.37 90 a Hz 0 Irreg 6 3 1830 430 6 3.6 15.3 860 13, 14 1 7 5 

Ranunculus arvensis Ranu AR -3.771 60 a Th 0 8 4 824 2 3 3.7 15.9 742 4 7 6 

Ranunculus auncomus Ranu N n -0.33 40 p hc 0 5 3 1379 226 1 3.3 15.1 922 1 6 5 

Ranunculus baudotii Ranu N n -0.04 60 a Hz 0 Node2 8 3 361 61 5 4.4 15.3 905 13 1 7 6 

Ranunculus bulbosus Ranu N n -0.48 40 p hc 0 6 3 2259 616 14 3.6 14.8 1004 6, 7 7 4 

Ranunculus circinatus Ranu N n -0.34 75 p Hy Irreg 7 5 575 76 0 3.7 15.8 753 13, 14 1 7 7 

Ranunculus ficaria Ranu N n 0.16 25 p Gn Otb DRa 8 3 2648 820 14 3.6 14.5 1084 1, 3 6 6 

Ranunculus flammula Ranu N n -0.60 50 p hc Hy Nadel 7 3 2651 938 11 3.5 14.4 1122 11 5 3 

Ranunculus fluitans Ranu N n 1.96 300 p Hy 1rreg Node2 7 3 373 3 0 3.3 15.4 798 14 1 7 6 

Ranunculus he derace us Ranu N n 0.10 9 23 a p Hz Hy 0 Node2 8 2 1703 551 8 3.6 14.5 1093 11, 13 1 5 5 

Ranunculus lingua Ranu N n 1.70 120 p hc Hy Rhiz2 7 4 537 210 0 3.7 15.2 868 11 1 6 7 

Ranunculus muricatus Ranu AN 40 a Th 0 0 3 Eur 18 1 0 5.0 15.6 864 4 5 5 

Ranunculus omiophyllus Ranu N n 0.52 9 25 a p Hz Hy 0 Node2 8 2 813 151 1 3.7 14.7 1180 11 1 5 4 

Ranunculus ophioglossifolius Ranu N EN r 40 a Th 0 8 3 4 0 2 4.9 16.5 824 13 7 5 

Ranunculus paludosus Ranu N o 40 p he Rhizl 9 1 0 0 4 6.1 16.9 843 6 6 3 

Ranunculus parviflorus Ranu N n -0.08 40 a Th 0 6 2 497 0 11 4.3 16.0 834 6 6 5 

Ranunculus peltatus Ranu N n 90 a Hz 0 Irreg 6 3 990 151 5 3.5 15.2 908 11, 13, 14 I l 5 6 

Ranunculus penicillatus Ranu N n 180 p Hy Irreg 7 3 706 284 0 3.8 15.2 942 14 1 8 5 

Ranunculus repens Ranu N n 0.55 60 p hc Sto12 5 5 2784 983 14 3.5 14.5 1105 3, 6 6 7 

Ranunculus reptans Ranu N EN r 10 p hc Hy Sto12 4 6 9 0 0 2.6 13.7i 1467 13 6 2 

Ranunculus sardous Ranu NA n 0.24 45 a Th 0 7 3 544 0 12 4.1 15.9 800 3, 6, 13 6 7 

Ranunculus sceleratus Ranu N n -0.05 60 a Th 0 5 6 1492 353 6 3.8 15.4 861 11, 13, 14 8 8 

Ranunculus trichophy llus Ranu N n -0.07 60 a Hz 0 Wag 3 6 1121 259 6 3.7 15.2 861 11, 13 1 6 6 

Ranunculus tripartitus Ranu N VU s -1.09 9 25 a p Hz 0 8 1 79 1 0 5.2 15.8 1024 13 1 6 3 

Raphanus raphanistrum Bras N n -1.39 70 a p Th hc 0 8 3 1862 259 13 4.6 15.0 1079 4, 19 6 6 

Raphanus raphanistrum 

subsp.maritimus Bras N 80 b p hc 0 9 1 325 108 13 4.7 15.1 1093 19 7 5 

Raphanus raphanistrum 

s ubsp.raphanistrum Bras AR 60 p he 0 8 3 1797 237 8 3.6 15.0 941 4 6 6 

Reseda lutea Rese NA n 0.39 75 hc 0 8 3 1276 0 7 3.6 15.7 808 3 7 5 

Reseda luteola Rese AR 0.69 150 hc 0 8 4 1660 428 12 3.8 15.3 879 3, 17 8 6 

Rhamnus cathartica Rham N n -0.04 600 Ph 0 7 4 857 88 0 3.6 15.8 783 1, 3 7 6 

Rhinanthus angustifolius Scro AN -0.10 60 a Th 0 5 4 Eur, As 1 90 0 0 3.1 14.6 809 3, 4, 7 7 2 

Rhinanthus minor Scro N n -0.49 50 a Th 0 5 3 2629 850 6 3.5 14.4 1115 6 6 4 

Rhodode ndron ponticum Eric AN 1.83 500 Ph 0 Nadel Eur 1966 507 6 3.5 14.6 1111 1, 10, 16 3 3 

R hynchospora alba Cype N n -0.43 30 hc Rhizl 5 6 624 407 0 3.6 14.0 1400 12 2 1 

R hy nch osp ora fusca Cype N s 0.02 30 hc Rhiz2 5 2 46 94 0 4.4 14.7 1205 12 3 1 

Ribes alpinum Gros N s 0.45 200 Pn Ph 0 4 3 50 0 0 2.3 14.2 1084 1, 3, 16 8 6 

R ibe s nigrum Gros AN 1.76 200 Pn 0 5 4 Eur, As1 1749 213 2 3.4 14.9 995 1, 3, 14 6 6 

Ribes rubrum Gros NA n 1.79 200 Pn 0 7 2 1874 0 2 3.3 15.1 944 1 7 6 

Ribes spicatum Gros N 5 -0.12 200 Pn 0 5 3 c 118 0 0 2.0 13.2 1095 1, 16 7 6 

Ribes uva-crispa Gros AN 0.72 100 Pn 0 7 3 Eur 2130 312 4 3.4 14.8 1006 1, 3 7 6 

Robinia pseudoacacia Faba AN 2700 Ph Root Am6 566 1 4 3.7 16.0 748 3, 17 6 6 

Romulea columnae hi d N VU r 6 Gn Otb 9 1 2 0 14 6.2 16.5 842 Co 18, 19 5 2

S 

1

N 

I 7 

5 

8 

4 

3 

4 

1 

3 

5 

2 

4 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

5 

8 

2 

3 

7 

5 

7 

2 

3 

3 

4 

2 

4 

4 

5 

3 

3 

5 

6 

8 

4 

2 

7 

7 

7 

R 

Clonel Clone2 E1 E2 C Origin GB IR CI Tja n Tjul Prec Co Br Habitats L F 

7 

12 

11 

5 

4 

7 

7 

7 

6 

7 

6 

11 

LF2 W 

h 

h 

h 

w 

hc 

1 P2 LF1 

hc 

Hy 

Hy 

Pn 

Th 

Th 

Ch 

Ch 

hc 

Ch 

Ch 

Hy 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Th 

Ph 

Pn 

Pn 

Ph 

Ph 

Ph 

Ch 

Pn 

Pn 

Pn 

Ch 

Ph 

Ph 

Pn 

Pn 

Ch 

Ph 

Ph 

Th 

hc 

hc 

hc 

Ph 

Ph 


60 

60 

40 

77 

15 

15 

3 

15 

20 

10 

10 

95 

36 

45 

35 

38 

40 

25 

40 

35 

25 

40 

2500 

70 

250 

1000 

800 

1500 

6 

100 

100 

300 

40 

700 

400 

300 

150 

15 

1000 

600 

50 

90 

80 

150 

1000 

400 

v i 

t-} 

I 

Ph 

Pn 

Pn 

Ph

S 

Taxon name Fam NS CS RS Chg Hght Len 1 P2 LFI LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan T'ul Prec Co Br Habitats L F R N 

Samolus valerandi Prim N n -0.42 45 hc 0 8 6 845 411 12 4.3 15.1 1013 11 8 8 8 5 

Sanguisorba minor Rosa N n -0.16 50 hc 0 8 4 1216 84 6 3.7 15.6 844 7 7 4 8 3 

Sanguisorba offi cinalis Rosa N n -0.23 120 hc 0 5 6 946 12 0 3 2 15.1 938 6 7 7 6 5 

Sanicula europaea Apia N n -0.98 40 hc o Rhizl 7 3 2025 634 0 3.5 14.7 1090 1 4 5, 7 5 

Saponaria officina(is Cary AR 0.29 90 hc Rhiz2 7 3 1246 256 9 3.9 15.5 890 3, 17 8 5 6 6 

Sarcocornia perennis Chen N s -0.22 30 Ch 0 9 1 80 4 1 4.4 16.4 702 Co 21 9 8 8 6 

Saussurea alpina Aste N n -0.51 45 hc Stoll 1 5 237 21 0 1.6 11.9 20 10 7, 15, 16 8 6 6 3 

Saxifraga aizoides Saxi N n -0.61 20 Ch Node2 1 3 422 17 0 1.7 12.3 1799 7, 11, 15, 16 8 9 6 2 

Saxifraga cernua Saxi N VU r 15 hc DRa DRi 1 6 5 0 0 0.5 11.7 2265 15, 16 6 6 7 1 

Saxifraga cespitosa Sax/ N VU r -0. 10 10 Ch 0 1 6 13 0 0 -0.4 10.7 2003 15, 16 71 5 71 1 

Saxifraga granulate Saxi N n -0.26 36 hc DRa 7 3 975 6 0 3 1 15.1 853 6, 7 8 5 6' 4 

Saxifraga hirculus Saxi N VU r -0.30 20 hc Rhiz/ 2 6 20 16 0 2.4 13.2 1253 11 8 9 6 2 

Saxifraga hirsuta Saxi N o 1.39 30 hc Stoll 7 1 0 36 0 5.0 14.6 1334 1, 15, 16 6 7 5 2 

Saxifraga hypnoides Saxi N n -0.54 20 Ch Node2 4 1 406 61 0 2.0 12.8 1697 7, 15, 16 7 5 6 3 

Saxifraga nivalis Saxi N s -0.50 15 hc 0 1 6 72 1 0 0 6 11.5 2099 15, 16 6 6, 71 3 

Saxifraga oppositifolia Saxi N n -0.45 3 Ch Node2 1 6 276 22 0 1.8 12.1 1980 7, 15, 16 8 6 8 2 

Saxifraga rivularis Saxi N r 0.19 10 hc Rhizl DRa 1 6 21 0 0 -0.3 10.8 1941 11, 15 6 9 5 2 

Saxifraga rosacea Saxi N EW x 20 Ch 0 4 2 2 31 0 4.4 14.3 1312 16 7 4 8 4 

Saxifraga spathularis Saxi N o 40 hc Ch Stoll 7 0 0 137 0 4.7 14.4 1328 16 6 8 3 2 

Saxifraga stellaris Saxi N n -0.58 20 hc Ch o 1 3 440 61 0 1.8 12.3 1811 11, 15, 16 8 8 5: 3 

Saxifraga tridactylites Saxi N n -0.12 10 Th 0 8 3 985 316 11 3.9 15.3 921 16 7 2 7 2 

Scabiosa columbaria Dips N n -0.71 70 hc 0 7 3 804 0 0 3,5 15.6 831 7 8 3 8 2 

Scandix pecten-veneris Apia AR -3.65 50 Th 0 8 4 780 94 6 3.8 15 7 792 4 7 4 7 4 

S che uchz e ria palustris Sche N VU r 22 hc Rhiz2 4 6 c 14 1 0 2.2 13.9 1380 12 9 9 3 1 

Schoenoplectus lacustris Cype N n 0 .47 2 10 Hy Rhizl 6 4 1202 535 0 3.7 14.9 1013 13. 14 8 11 71 6 

Schoenoplectus pungens Cype NA r 60 Hy Rhizl 7 3 0 0 1 6.1 16.7 869 Co / 9 8 10 7 7' 

Schoenoplectus 

tabemaemontani Cype N n 0.67 150 Hy Rhizl 8 5 702 192 4.2 15.3 917 11, 13 9 10 8 7 

Schoenoplectus ttigueter Cype N CR r 125 Hy Rhizl 7 5 8 3 4.7 16.5 799 14 8 10 7 7 

Schoenus ferw gineus Cype N VU r 40 hc o 4 3 3 0 0 4 12.4 1113 11 8 9 7 2 

Schoenus nigricans Cype N n -0.53 75 hc 0 8 4 763 484 3.9 13.9 1261 11 8 8 71 2 

Scilla autumnatis Lili N s -0.37 20 Gb Otb 9 1 49 0 1 5.9 16.2 939 Co 10, 18 9 3 6/ 1 

Scilla Verna Lili N n 0.12 10 Gb Otb DRg 7 1 310 29 4.6 13.9 1103 6, 10 8 5 5 3 

Scirpoides holos choenus Cype N VU r 0.21 100 hc 0 8 4 3 0 5.2 16.3 909 Co 19 8 8 7 6 

Scirpus sylvaticus Cype N n 0.02 120 hc Rhizl 7 4 871 102 3.4 15.2 921 1, 11 6 8 6 6 

S cleranthus annuus Cary N n -2.68 20 Th 0 7 3 983 69 3.6 15.2 883 4, 8, 10, 16 71 4 4 4 

S cleranthus perennis Cary N EN r -0.11 20 Ch o 7 3 11 0 3 3 16.0 656 8, 16 81 3 4 2 

Scorzonera humilis Aste NA VU r 50 hc 0 7 3 4 0 4 8 16. 1 1097 6 8 7 5 2 

Scrophularia auriculata Scro N n -0.21 120 hc 0 8 2 1490 504 3.9 15.4 918 11, 14 7 8 7 7 

Scrophularia nodos a Scro N 6 -0.37 100 hc 0 7 4 2327 850 3.6 14.7 1069 1, 3 5 6 7 6 

Scrophularia s corod onia Scro AN , 0.75 100 

hc 

0 8 1 Eur 68 0 1 5.8 16.0 1033 3, 16, 18 71 4 6 6 

Scrophularia umbrosa Scro N n 0.72 100 

hc 

0 7 4 c 

196 14 3.1 14.9 868 1, 14 

7 9 7 7 

Scrophularia vernalis Scro AN . 0.54 65 p hc 0 Eur 192 0 3.3 15.3 769 1, 3 5 5 7 7 

Scutellaria galericulate Lami N n -0.39 50 hc Rhizl 5 4 1751 212 3.6 14.9 1055 11 7 8 6 5 

Scutellaria minor Lami N 6 0.03 20 hc Rhizl 8 2 803 136 4.1 14.9 1191 11 7 9 4 2 

Secale cereale Poac AC , 150 Th 0 Eur 134 71 4.0 15.8 852 3, 4 8 5 71 7 

/

Taxon name Fam NS CS RS Chg Hght Len 1 P2 LF1 LF2 W Clone') Clone2 El E2 C Origin GB IR CI T an T ul Prec Co Br Habitats L F R N S 

Sedum acre Cras N n -0.24 10 Ch Node2 Irreg 7 3 2108 504 12 3.7 14.9 995 16 8 2 7 2 1 

Sedum album Cras AR 2.41 20 Ch Node2 Irreg 9 2 1603 517 13 3.8 15.1 948 3, 16, 17 8 3 6 2 0 

Sedum anglicum Cras N n -0.21 5 Ch Node2 Irreg 7 1 988 393 14 4. 1 14.3 1321 16, 18 8 3 4 2 

Sedum forsterianum Gras N s 1.54 20 Ch Node2 Irreg 8 1 122 0 0 3.6 14.8 1239 3, 16 7 3 5 1 

Sedum rosea Cras N n -0.41 35 hc 0 1 6 524 82 0 21 12.6 1728 15, 16 7 6 6 3 

Sedum rupestre Gras AN 2.20 30 Ch Node2 Irreg Eur 1212 70 4 3.7 15.5 905 3, 16 7 2 5 4 

Sedum telephium Cras N n -0.34 60 hc 0 7 5 1197 0 0 3.5 15.0 1032 1, 3, 16 7 5 7 5 

Sedum villosum Cras N s -0.76 10 Ch 0 2 3 211 0 0 1.6 12.9 1273 11 8 9 6 2 

Selaginella selaginoides Se la N n -0.47 9 Ch Nodal 4 6 988 288 0 2.8 13.0 1443 7, 11 8 7 6 2 

Selinum carvifolia Apia N VU r 100 hc 0 7 3 3 0 0 3.4 16.1 595 11 7 7 8 4 

Sempervivum tectorum Cras AN 40 Ch Stoll Eur 500 42 2 3.5 15.3 871 3, 17 8 2 4 1 

Senecio aquaticus Aste N n -0.92 80 p hc 0 7 3 2304 919 1 3.6 14.5 1109 11, 14 7 8 6 5 

Senecio cambrensis Aste NE r 30 hc 0 7 1 14 0 0 3.6 15.0 804 3, 17 8 5 7 7 

Senecio cineraria Aste AN 

2.73 60 

Ch Pn sw 0 

0 3 Eur 256 8 5 4.4 15.9 837 3, 17, 18, 19 9 3 7 3 

Senecio erucifolius Aste N n -0.03 120 

hc 

Rhizl 

7 4 

1276 30 0 3.8 15.8 783 3, 6 

7 5 7 5 

Senecio fluviatilis Aste AN 

0.03 150 

hc 

Rhiz2 

7 4 Eur, As l 184 25 0 3.3 14.6 1018 1, 11, 13, 14 7 8 6 7 

Senecio j acobaea Aste N n 0.11 125 

b hc 

0 Root 7 4 

2725 982 14 3.6 14.5 1102 3, 6, 7, 8 7 4 6 4 

Senecio paludosus Aste N CR r 175 hc 0 7 4 c 7 0 0 3.4 16.2 573 11, 13 7 9 7 6 

Senecio smithii Aste AN 100 hc 0 SAm 62 0 0 3.4 12.3 1072 3, 6 8 7 6 7 

Senecio squalidus Aste AN 0.771 50 a p Th Ch 0 Gard 1494 75 4 3.7 15,5 850 3, 17 8 4 7 7 

Senecio sylvaticus Aste N n 0,09 70 a Th 0 7 3 1853 238 12 3.6 14.9 987 2, 8, 9 7 5 5 6 

Senecio viscosus Aste AN 0.63 60 a Th 0 7 3 Eur 1747 69 1 3.4 15.1 920 3, 17, 19 8 5 7 6 

Senecio vulgaris Aste N n -1.08 37 a Th 0 8 3 2630 942 14 3.6 14.6 1075 3, 4, 17 7 5 7 7 

Serapias parviflora Orch NA r 30 p Gn 0 9 1 1 0 0 6.0 16.2 1048 6 8 4 7 3 

Seriphidium maritimum Aste N n -0.42 50 p Ch hc Rhiz2 7 2 222 20 1 4.3 15.8 783 Co 21 9 7 8 6 

Serratula tinctoria Aste N n -0.21 70 p hc 0 7 3 945 0 2 3.8 15.5 965 7 7 6 6 2 

Seseli libanotis Apia N VU r 60 b hc 0 7 5 c 4 0 0 3.7 16.4 645 7 7 4 8 3 

Sesleria caerulea Poac N s -0.09 45 p he 0 5 3 76 89 0 3.1 13.9 1262 7, 16 7 6 8 2 

Setaria viridis Poac AN 0.80 70 a Th 0 Unk 431 27 5 3.9 15.9 798 3, 4, 17 7 4 7 7 

Sherardia arvensis Rubi N n -0.94 40 a Th 0 8 3 1635 369, 12 3.9 15.2 916 7, 16 7 4 6 4 

Sibbaldia procumbens Rosa N s -0.75 5 p hc 0 1 6 133 0 0 0.4 11.3 2109 15 8 5 4 3 

Sibthorpia europaea Scro N s -0.14 5 p Ch Node2 7 1 107 9 8 5.3 15.5 1189 3, 14 5 7 5 5 

Silaum silaus Apia N n -0.42 100 p hc 0 7 4 963 0 0 3.6 15.8 743 6 8 5 7 4 

Silene acaulis Cary N n -0.47 10 p Ch 0 1 3 236 9 0 1.9 11.8 1937 7, 15, 16 8 5 6 1 

Silene conica Cary N $ -1.05 35 a Th 0 5 4 59j 0 10 4.0 16.0 702 8, 19 8 3 4 2 

Silene dioica Cary N n -0.44 90 hc Ch 0 5 3 2514 263 14 3 4 14.6 1065 1 5 6 6 7 

Silene gallica Cary AR -2.78 45 a Th 0 9 2 455 31 11 4,2 15.7 877 3, 4 7 4 5 5 

Silene latifolia Cary AR -0.88 100 p hc 0 8 4 1954 211 13 3.6 15,1 911 3 7 4 7 6 

Silene noctiflora Cary AR -2.04 50 a Th 0 7 3 686 22 1 3.7 15.8 736 4 7 4 7 6 

Silene nutans Cary N $ -0.39 80 p hc 0 7 4 52 0 10 4.3 15.7 883 16 5 3 8 4 

Sfi ene ofi tes Cary N r -0.36 80 p hc 0 7 3 c 18 0 0 3.2 16.1 625 3, 8 8 3 7 2 

Silene uniflora Cary N n -0.39 28 p hc 0 5 2 943 236 13 4.0 14.1 1263 Co 18 8 6 6 4 

Silene vulgaris Cary N n -1.26 80 p hc 0 8 5 1710 280 8 3.7 15.3 897 3, 6 7 4 8 5 

Silybum marianum Aste AR -0.07 100 b a hc Th 0 0 3 Eur 552 64 6 4.0 15.8 781 3 8 4 8 6 

Simethis planifolia Lili N o 42 p Gn 0 5 2 0 4 0 5.7 15.1 1324 10 71 5 4 3 

Sinapis alba Bras AR -0.90 90 a Th 0 8 3 1082 162 10 3.8 15.5 832 3, 4 7 4 7 6

Taxon name Fam NS CS RS Chg Hght Len P1 P2 LF1 LF2 W Cb nel Clone2 El E2 C Origin GB IR CI Tian Tjul Prec Co Br Habitats L F R N S 

Sinapis aneensis Bras AR , -1 76 90 a Th h 0 7 4 2373 772 11 3.7 14.7 1012 3, 4, 17 8 5 7 7' 

Sison amomum Apia N n -0.19 100 b hc h 0 9 2 790 0 0 3.9 16.2 756 3 7 4 7 5 

Sisymbrium altissimum Bras AN -0,84 100 a Th h 0 Eur 614 29 3 3.7 15.6 771 3, 17 8 5 6 4 

Sisymbrium ofi cinale Bras AR i -0.21 95 a b Th hc h 0 8 3 21171 775 13 3.8 15.0 973 3, 4, 17 71 4 7 7 

Sisymbrium orientale Bras AN -0.24 80 Th h 0 Eur 864 75 10 3.8 15.6 815 3, 17 7' 4 7 5 

Sisyrinchium be rm udiana Irid N o -1.80 23 hc h 0 6 0 0 43 0 4.5 14.4 1290 6, 13 8 8 6 3 

Sium latifolium Apia N s -1.83 200 Hy h 0 7 4 259 34 0 3.7 15.9 702 11, 14 7 10 7 7 

Smyrnium olusatrum Apia AR 0.66 150 he h 0 Eur 803 327 13 4.4 15.5 907 3 T 5 7 7 

Solanum dulcamara 

So la N n -0.11 225 

Pn Ch sw Node2 

8 5 

1918, 533 14 3.8 15.2 945 3, 11, 14 71 8 71 7 

Solanum nigrum 

So la NA n 0.44 60 

Th 

0 

8 5 

1258E 0 14 3.9 15.9 809 4 

7 5 7 8 

Solanum sarachoides So la AN 60 a Th 0 SAm 88 0 4 3.8 16.1 696 4, 17 7 4 7 7 

Solanum tubemsum So la AN 100 Gn Rhiz1 SAm 625 56 5 3.9 15.4 883 4 7 4 6 7 

So leiro lia soleirolii Urti AN 2.36 10 Ch Node2 0 2 Eur 773 130 11 4.3 15.7 906 3, 17 4 8 7 6 

Solidago canadensis Aste AN 200 hc Rhiz1 Am 981 24 3.6 15.6 833 3, 17 8 5 6 6 

Solidago gigantea Aste AN 200 hc Rhiz1 Am 645 1 3.5 15.5 854 3, 17 8 5 5 6 

Solidago virgaurea Aste N n -0.89 70 he 0 5 5 2043 492 3.4 14.2 1217 10, 16 5 5 4 3 

Sonchus arvensis Aste N n -0.12 150 hc Rhiz2 7 4 2320 806 3.8 14.8 1013 4 8 6 7' 6 

Sonchus asper Aste N n 0.78 150 a Th 0 8 3 2584 961 1 3.7 14.6 1070 3, 4 T 5 7 6 

Sonchus oleraceus Aste N n -0.42 150 Th 0 8 3 2358 924 1 3.71 14.8 1031 3, 4 71 5 71 71 

Sonchus palustris Aste N s, 0.18 250 he 0 7 4 51 0 3.8 16.4 615 11 7 8 7 7' 

Sorbus anglica Rosa NE VU r 200 Pn w 0 7 1 14 1 3.8 15.2 1082 1, 16 6 4 8 4 

Sorbus aria Rosa N n 1500 Ph w 0 7 3 225 2 3.8 16.3 774 1 6 4 7 4 

u-i 

u, Sorbus aria agg, Rosa N n' 0.82 1000 

Ph w 0 

7 3 

341 75 3.8 16.3 780 17 

6 5 7 4 

Sorbus arrane nsis Rosa NE VU r 

750 

Ph w 0 

4 1 

1 0 2.5 12.1 1530 16 

71 4 4 3 

Sorbus aucuparia Rosa N n 0.86 1500 Ph w 0 5 5 2472 749 3.4 14.4 1128 1, 2, 16 6 6 3 4 

Sorbus bristoliensis Rosa NE EN r , 1000 

Ph w 0 

7 1 

1 0 4.4 16.5 844 1, 16 

6 4 8 4 

Sorbus de von iensis Rosa NE s , 1500 

Ph w 0 

7 1 

32 11 5.0 15.4 1171 1, 3 

6 5 6 5 

Sorbus domestica Rosa N CR r 1 500 

Ph w Root 

3 

4 0 4.5 16.5 857 16 

6 4 8 3 

Sorbus e minens 

Rosa NE VU r 600 

Ph w 0 

7 

8, 0 4.4 16.2 878 1 

6 5 71 5 

Sorbus hibernica 

Rosa NE o 600 

Ph w 0 

7 

0' 63 4.5 14.6 1035 1, 3, 16 6 5 7 5 

Sorbus intermedia Rosa AN 1000 Ph w 0 Eur 686 17 3.5 15.2 901 1, 3, 17 6 6 8 7 

Sorbus lanc as triensis Rosa NE r 500 Ph w 0 7 9 0 3.3 14.8 1285 16 8 4 7 3 

Sorbus leptophylla Rosa NE CR r 300 Pn Ph w 0 7 3 0 2.9 14.5 1501 16 5 5 7 5 

Sorbus leyana 

Rosa NE CR r 

200 

Pn w 0 

7 

2 0, 2.5 14.0 1687 16 

7 5 8 4 

Sorbus minima 

Rosa NE VU r ' 300 

Pn Ph w 0 

7 

1 0' 2.2 14.0 1523 16 

6 4 8 3 

Sorbus porrigentiformis Rosa NE 5 , 500 

Ph w 0 

7 

30 0 3.8 15.2 1273 16 

7' 5 7' 5 

Sorbus ps e udotennica Rosa NE VU r 

700 

Ph w 0 

4 

1 0 2.5 12.1 1530 16 

7 4 4 3 

Sorbus rupicola Rosa N s 200 Pn w 0 4 2 98 13 2.9 13.6 1405 16 8 4 7 3 

Sorbus subcuneata 

Rosa NE VU r 1000 

Ph w 0 

7 1 

4, 0 4.0 14.9 1380 1 

6 5 4 4 

Sorbus torminalis 

Rosa N n 0.22 1900 

Ph w Root 

7 3 

573 0 3.9 16.0 804 1 

4 5 6 5 

Sorbus vexans Rosa NE VU r 600 Ph w 0 7 1 4 0 4.2 14.9 1409 1, 16 6 5 4 4 

Sorbus wilmottiana Rosa NE CR r 600 Ph w 0 7 1 1 0 4.4 16.5 844 1, 16 6 4 8 3 

Sparganium angus tifolium Spar N n 1.66 100 Hy h Rhiz2 4 3 652 207' 3.0 12.9 1571 13 8 11 4 2 

Sparganium emersum Spar N n 0.61 60 Hy h Rhiz2 5 6 1426 393 3.6 15.0 982 13, 14 7 11 7 6 

Sparganium erectum Spar N n 0.48 150 Hy h Rhiz2 7 6 2129 766 3.7' 14.9 1000 11, 13 7' 10 7 7 

Sparganium natans Spar N n : -0.13 50 Hy h Rhiz2 5 6 508 267 3.3 13.8 1303 13 7 11 6 3

S 

Taxon name Fam NS CS RS Chg_ Hght Len P1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tian Tjul Prec Co Br Habitats L F R N 

Spartina alternitiora Poac AN -0.22 110 p hc Rhiz2 Am 12 0 0 4.1 15.7 760 Co 21 9 9 8 7 

Spartina anglica Poac NE n 0.11 130 p hc Rhiz2 7 1 251 71 1 4.6 15.7 904 Co 21 9 9 8 6 

Spartina maritima Poac N s -0.55 50 p hc Rhiz2 8 2 571 0 0 4.1 16.6 625 Co 21 9 9 8 5 

Spergula arvensis Cary N o -2.30 30 a Th 0 6 4 0 0 9 6.3 16.7 811 4 7 4 5 5 

Spergularia bocco ne i Cary AN -0.22 10 a Th 0 9 1 Eur 18 0 9 5.9 16.3 901 16, 17, 19 9 4 6 7 

Spergularia marina Cary N n 1.83 10 a Th 0 8 6 680 185 8 4.3 14.6 1094 Co 3, 21 8 8 8 6 

Spergularia media Cary N n -0.24 15 p Ch 0 8 4 659 186 0 4.2 14.6 1125 Co 21 8 8 8 5 

Spergularia rubra Cary N n 0.05 15 a Th 0 8 3 1363 66 13 3.5 14.9 983 8, 17 8 3 4 2 

Spergulatia rupicola Cary N n 0.30 10 Ch 0 7 0 239 169 13 5.2 15.1 1105 Co 18 9 6 6 5 

Spiranthes aestivalis Orch N EX x 40 hc 0 7 3 1 0 2 5.6 16.5 822 11 9 8 8 2 

Spiranthes romanz offi ana Orch N s 0.45 25 hc 0 4 0 21 44 0 4.4 14. 1 1248 11 8 8 6 4 

Spiranthes spiralis Orch N n -0.95 15 hc 0 8 3 655 117 13 4.4 15.9 884 7 8 4 6 3 

Spirodela polyrhiza Lemn N n -0.18 0.8 Hy Frag 8 6 535 991 2 3.9 15.9 768 13 71 11 7 7 

Stachys alpina Lami AN 100 hc 0 7 3 Eur 3 0 0 3.1 14.7 1007 1, 3 7 5 8 7 

Stachys arvensis Lami AR -1.17 25 Th 0 8 2 1418 206 13 4.0 15.3 964 3, 4 8 5 5 5 

Stachys germanica Lami N EN r -0.27 80 hc 0 7 3 10 0 0 3.4 16. 1 66 1 3 7 3 8 5 

Stachys officinalis Lami N n -0.62 60 he 0 7 3 1571 38 4 3.6 15.4 940 6, 7 7 5 5 3 

Stachys palustris Larni N n 0.01 100 Gn Rhiz2 5 6 2346 919 9 3.71 14.6 1079 11, 14 7 8 7 7 

Stachys palustris x sylvatica 

(S. x ambigua) Lami NH n 100 hc Rhiz2 1020 1171 4 3.5 14.7 1086 3, 14 7 6 6 6 

Stachys sylvatica Lami N n -0.49 100 hc Rhiz2 7 4 2462 806 12 3.6 14.7 1064 3 6 6 7 8 

Stellaria graminea Cary N n -0.02 80 hc Rhizl 5 4 2446 852 8 3.5 14.6 1065 6 7 6 5 4 

i n 

cn Stellaria holostea Cary N n -0.56 60 Ch Rhiz1 Nodel 7 4 2372 671 7 3.5 14.7 1058 1 5 5 6 6 

Stellaria media Cary N n 50 Th 0 Nodel 6 5 2749 962 14 3.6 14.5 1095 3, 4 7 5 6 7 

Stellaria nemorum Cary N n 0.21 60 hc Rhizl 5 3 432 0 0 2.7 14.2 1027 1, 14 4 6 6 7 

Stellaria media egg. Cary N n 0.03 50 Th 0 Nodal 6 5 2749 962 14 3.6 14.5 1095 3, 4 7 5 5 6 

Stellaria neglecta Cary N n 0.42 80 Th 0 Nodel 7 3 775 4 0 3.9 15.6 911 1, 3 6 7 6 7 

Stellaria pallida Cary N n 1.17 50 Th 0 Nodel 8 4 593 27 13 4.0 15.8 766 3, 19 7 4 4 4 

Stellaria palustris Cary N n -0.89 60 hc Rhizl Nodel 5 5 389 73 1 3.6 15.5 793 11 7 8 6 4 

Stellaria uliginosa Cary N n -0.10 40 he Rhizl 7 3 2570 798 9 3.5 14.4 1128 11, 14 7 8 5 5 

Stratiotes ab ides Hydr NA r 1.65 50 Hy Stoll DRa 5 4 c 65 0 0 3.5 16.1 596 13 7' 11 7 6 

Suaeda maritima Chen N n -0.47 30 Th 0 8 5 525 141 6 4.3 14.9 1094 Co 21 9 8 8 6 

Suaeda vera Chen N s -0.11 120 Pn w 0 9 1 47 0 1 4. 1 16.4 620 Co 19, 21 9 7 8 5 

Subularia aquatica Bras N n 0.73 6 Hz h 0 4 6 324 33 0 2.4 12.6 1789 13 7 11 5 2 

Succisa pratensis Dips N n -0.57 100 he h 0 Rhizl 7 4 2633 934 2 3.5 14.4 1124 6 7 7 5 2 

Symphoricarpos albus Capr AN 1.74 200 Pn w Rhiz2 Am 2067 744 4 3.6 14.9 992 1, 3, 17 5 5 6 7 

Symphytum asperum x 

of icinale (S. x upland icum) Bora AN 135 hc h 0 Eur 1924 399 8 3.6 15.0 952 3 6 5 7 7 

Symphy tum officinale Bora N n 0.34 135 hc h 0 7 3 1263 189 10 3.7 15.4 871 11 7 7 7 8 

- - 

Symphytum orientale Bora AN 1.83 70 hc h 0 Eur 442 2 3 3.71 16.0 719 3 6 4 7 6 

Symphytum tuberosum Bora N n 0.11 55 hc h 0 7 3 407 0 0 2.5 13.5 1047 1, 3 6 6 6 6 

Sybnga vulgaris Olea AN 4.48 700 Ph w Rhir l Eur 1296 163 4 3.6 15.3 881 3, 17 6 5 6 5 

Tamus communis Dios N n -0.41 400 Gn h 0 9 2 1421 0 4 3.8 15.7 850 1, 3 6 5 7 6 

Tanacetum parthenium Aste AR 0.23 54 Ch hc h 0 Eur 2126 413 11 3.5 14.9 980 3, 17 7 4 7 6 

Tanacetum vulgare Aste N n -0.23 120 hc h Rhizl 5 5 2004 0 10 3.5 15.0 958 3 7 6 7 7 

Taraxacum Aste N n 0.43 30 hc h 0 6 6 2778 974 14 3.5 14.5 1104 3, 5, 17 7 5 7 6

W Clonel Clone2 El E2 C Origin GB IR CI Tja n Tjul Prec Co Br Habitats L F R N S 

w 0 

7 3 1881 260 7 3 5 15.1 976 1 4 4 7 5 0 

h 0 

7 3 509 11 12 3 4 15.0 949 8 8 3 2 2 0 

h 0 

Am4 308 20 2 3.6 15.2 974 1, 3, 17 3 8 6 4 0 

h 0 

3 6 c 99, 0, 0 3 71 16. 1 774 7 71 3 8 3 0 

h 0 

3 6 26 0 0 3.6 16. 1 617 13 7 9 7 6 0 

h 0 

7 3 c Eur 12 0 0 3.6 16.3 767 3, 4, 7, 16 9 2 8 2 0 

sw Rhizl 

8 3 c Eur 72 2 0 4.2 15.7 856 3, 7, 16 8 2 8 1 0 

h Rhiz2 

8 4 24 12 I 4.0 15.8 740 11, 13, 14 7 8 8 4 1 

h Rhiz2 

8 2 2322 588 1 3.5 14.5 1150 1, 9, 16 6 4 4 3 

h Rhiz2 

1 6 409' 23 2.0 12.1 1845 7, 11, 15, 16 8 7 6 3 

h 0 

5 4 669 91 3.7 15.8 762 11 7 8 7 5 

h Rhiz2 

5 5 504 89 3.4 14.0 1185 16, 19 7 4 6 3 

h Rhiz2 

7 6 171 54 3.9 15.5 869 1, 11 6 8 7' 5 

h 0 

7 1 145 0 3 6 16.2 762 7 8 3 8 3 

h 0 

7 5 1600 143 1 3 7 15.3 858 3, 4 7 4 7 6 

h 0 

4 3 70 0 2 0 13.5 1314 16 8 4 6 1 

h 0 

8 4 9 0 3,3 16.0 732 3, 16 8 4 8 2 

w 0 

Am4 574 7 1 3.6 15.4 977 2, 17 4 5 5 4 

sw Node2 

5 3 2246 486 1 3.4 14.2 1175 7, 16 8 4 6 2 

sw Node2 

7 3 455 3 3.7 16. 1 752 7 8 4 8 2 

sw Node2 

5 3 c 9 0 3,3 16.2 606 8 8 2 5 2 

w 0 

7 4 896 5 20 0 2.4 3 5 15.5 14.4 1064 873 1 5 5 6 5 6 

R 

B 

B 

B 

C 

L 

T 

7 3 84 0 3.3 15.6 816 1 4 5 7 6 

1 6 156 0 0 6 11.6 1964 7, 11 8 9 7 2 

Am4 267 11 3.2 14.3 1131 1 3 6 7 7 

Eur 11 1 4.0 16.3 739 6 7 3 6 5 

8 4 389 0 3.7 16.2 707 4 8 4 8 4 

7 5 2178, 824 3.71 14.9 1004 3 7 5 7 71 

9 1 708 77 4. 1 15.9 792 3, 6 8 5 7 6 1 

7 4 1749 124 1 3.6 15.3 87 1 6 8 4 7 5 0 

DRp 7 0 162 62 3.8 14.2 1390 16 2 7 7 3 0 

4 6 c 1 0 2.8 14.0 858 12 8 9 2 2 0 

4 6 1553, 629 0 3.2 13.71 1312 10, 12 8 8 2 1 0 

DRg 4 6 514 0 0 1,8 12.6 1277 1, 2, 10 5 6 3 3 0 

8 4 1205 56 14 3.8 15.5 84 1 8 9 3 5 2 1 

9 1 3 0 2 6 5 16.2 867 16 9 4 5 2 0 

8 4 1996 381 14 3.7 15.1 939 3, 16, 19 8 4 6 4 0 

7 3 2548 942 14 3 6 14.6 1072 6 71 4 6 5, 0 

8 4 7531 42 6 4.0 16.0 764 6 8 7 7 6 2 

9 1 148 5 13 4.6 16.3 767 8 9 3 5 2 0 

Eur 1940 214 11 3.5 15.0 943 3, 5 7 5 7 6 0 

9 1 3 0 4 6.4 16.4 883 3, 18 8 2 5 2 1 

w 0 

w 0 

h Rhizl 

h 0 

h 0 

h 0 

h 0 

h 0 

h 0 

h Rhizl 

h Rhizl 

h Ogr 

h Rhiz2 

h 0 

h 0 

h 0 

h 0 

h Node2 

h 0 

h 0 

h 0 

h 

T 

T 

Crop 196 1 5 4.0 16. 1 780 3 7 4 6 5 0 

h O

Taxon name Fam NS CS RS Chg Hght Len P1 P2 LF1 LF2 W Clonal Clone2 El E2 C Od in GB IR CI T an T uI Prec Co Br Habitats I. F R N S 

Trifolium incamatum 

subsp.molinerti Faba N VU r 20 Th 0 9 1 3 0 4 6 4 16.4 883 Co 18 6 2 51 2 

Trifolium medium Faba N n -0.53 45 hc 0 5 4 2050 208 2 3.4 14.8 1008 6 7 4 6 4 

Trifolium micranthum Faba N n 0.62 7 Th 0 9 2 909 52 13 4.1 15.9 848 8, 17 8 5 5 5 

Trifolium occidentale Faba N a 10 hc Ch Node2 7 0 21 9 12 6.3 15.9 940 Co 18 9 4 6 2 

Trifolium ochroleucon Faba N s -0.84 45 he 0 7 3 1271 0 0 3.4 16.3 595 3. 6 7 5 8 2 

Trifolium om ithop odioide s Faba N n 0.42 12 Th 0 8 2 281 18 14 4.8 16. 1 874 8 9 6 5 3 

Trifolium pratense Faba N n -0.18 45 hc 0 7 4 2745 976 14 3.6 14.5 1100 6. 7 7 5 7 5 

Trifolium repens Faba N n 1.31 20 hc Ch Node2 5 4 2798 981 14 3.5 14.5 1105 6 7 5 6 6 

Trifolium scabrum Faba N n -0.39 20 Th 0 9 2 390 12 11 4.5 15.9 831 8 9 3 7 2 

Trifoliurn sguamosum Faba N s -0.32 40 Th 0 9 1 116 0 6 4.5 16.5 757 Co 6, 19 9 6 7 6 

Trifolium striatum Faba N n -0.11 20 Th 0 8 3 848 31 12 4.0 15,7 805 8 8 3 5 2 

Trifolium strictum Faba N VU r 15 Th 0 9 2 4 0 2 5.8 15.9 874 16 9 2 5 2 

Trifolium subterraneum Faba N n -0. 10 10 Th 0 9 2 337 1 13 4.5 16.2 805 8, 18 8 3 4 2 

Trifolium suffocatum Faba N s 0.14 5 Th 0 9 1 96 0 13 4.8 16.4 774 8 8 4 4 2 

Triglochin maritimum Junc N n -0.44 55 hc Rhizl 5 6 8371 251 5 4.2 14.3 1181 Co 21 8 7 7 5 

Triglochin palustre Junc N n -0,22 55 hc Rhiz2 5 6 2196 71 5 3.4 14.2 1145 11 8 9 6 2 

Trinia glauca Apia N r 0.12 20 hc 0 8 3 6 0 4.9 16.4 889 7 9 2 8 1 

Tripleurospermum inodorum Aste AR 60 Th 0 7 4 2119 42 12 3.6 14.9 967 4. 17 8 5 6 6 

Tripleurospermum maritimum Aste N n 60 Ch hc 0 3 6 757 27 3 4.4 14.5 1132 Co 18, 19 8 5 6 6 

Tnpleurosp ennum maritimum 

sens.lat. Aste N n 0.3 1 60 p Th hc 0 3 6 757i 27 3 4.4 14,5 1132 4, 17 18, 19 8 5 6 6 

tri 

co Trisetum flavescens Poac N n -0.13 80 he 0 7 3 1734 36 6 3.6 15.2 900 6, 7 7 4 7 4 

Triticum aestivum Poac AC 150 Th 0 Crop 741 5 4 3.8 15.6 861 3, 4 8 5 7 7 

Trollius europaeus Ranu N n -0.73 60 hc 0 4 3 856 0 2.2 13.0 1517 11, 16 7 7 6 4 

Tsuga he terophylla Pina AN 4600 Ph w 0 Am4 598 1 0 3.4 14.9 1102 1, 2, 17 6 6 3 3 

Tuberaria guttata Cist N VU r 15 Th h 0 9 1 5 1 4 5.71 15.5 1069 10 9 2 5 1 

Tussilago farfara Aste N n -0.65 30 Gn h Rhiz2 5 4 2618 90 7 3.5 14.5 1085 16 7 6 6 6 

Typha angustifolia Typh N n 0.35 300 Hy h Rhiz2 7 4 776 6 6 3.7 15.8 775 11 8 10 7 7 

Typha latifolia Typh N n 1.01 275 Hy h Rhiz2 8 6 1860 68 9 3.8 15.1 958 11 8 10 7 7 

Ulex europaeus Faba N n -0.34 200 Pn w 0 7 1 2518 95 14 3.6 14.6 1090 10 7 5 5 3 

(fl ex gallii Faba N n 0.20 150 Pn w 0 7 1 918 38 11 4.1 15.0 1099 10 7 6 3 2 

Ulex minor Faba N n 0.20 100 Pn w 0 8 1 197 4 3.9 16.4 745 10 8 6 1 2 

Ulmus glabra Ulrna N n -0.28 3000 Ph w 0 7 3 2338 60 0 3.4 14.7 1056 1 4 5 7 6 

Ulmus minor Ulrna N n 0.75 3100 Ph w Root 7 3 641 9 3.9 16. 1 764 1, 3 5 5 7 7 

Ulmus plotii Ulma NE s 2000 Ph w Root 7 1 128 0 3.4 16.0 667 3 5 5 7 7 

Litmus procera Ulma NA n -0.48 3300 Ph w Root 7 3 13171 0 3.7 15.8 819 3 5 5 8 6 

Umbilicus rupestris Cras N n -0. 12 38 hc h 0 9 1 784 65 i 14 4.3 14.9 1125 3, 16 6 4 5 4 

Urtica dioica Urti N n 0.28 150 hc h Rhiz2 Sto12 5 4 2773 98 13 3.6 14.5 1102 3, 14, 17 6 6 7 8 

Urtica urens Urti AR -0.70 60 Th h 0 8 4 1924 28 14 3.7 15.0 914 4, 17 8 5 6 8 

Utricularia australis Lent N s 60 Hy h DRa 5 5 162 5 0 4.0 15.0 1120 13 7 12 5 3 

Utricularia intermedia 

sens.lat. Lent N n 0.40 20 Hy h DRa 4 6 412 15 0 3.1 13.2 1545 11, 12, 13 8 12 4 2 

Utricularia intermedia 

sens.str. Lent N n 20 Hy h DRa ? ? 13 8 12 4 2 

Utricularia minor Lent N n 0.20 40 Hy h DRa 5 6 628 377 1 3.6 13.9 1321 11, 12 8 12 4 2

S 

Taxon name Fam NS CS RS Chg_ Hght Len P2 LF1 LF2 W Clonel Clone2 E1 E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N 

Utricularia ochroleuca Lent N i 20 Hy h DRa ? ? 11, 13 8 12 3 1 

Utricularia stygia Lent N i 20 Hy h DRa ? ? 11, 13 8 12 5 2 

Utricularia vulgaris seno lat. Lent N n 0.47 100 Hy h DRa 5 5 744 301 0 3.6 14.4 1148 13 7j 12 6 4 

Utricularia vulgaris sens.str. Lent N s 100 Hy h DRa 7 4 159 77 0 3.8 15.3 874 11 7 12 7 4 

Vaccinium microcarpum Eric N s 0.81 30 Ch h Node2 4 6 100 0 0 0.9 11.9 1339 12 7 8 1 1 

Vaccinium myrtillus Eric N g -0.61 50 Ch Pn w Rhiz2 4 4 1886 701 1 3.3 13.9 1243 10, 16 6 6 2 2 

Vaccinium oxycoccos Eric N n 0.28 30 Ch h Node2 4 6 725 246 0 3.0 14.0 1229 12 8 9 2 1 

Vaccinium uliginosum Eric N n -0.39 50 Ch Pn w Rhiz2 2 6 252 0 0 1.1 11.7i 1897 12, 15, 16 71 6 2 2 

Vaccinium vitis-idaea Eric N n -0. 18 30 Ch w Rhiz2 2 6 938 89 0 2.2 12.9 1474 2, 10, 15 6 5 2 2 

Valeriana dioica Vale N n -0.67 35 hc h Sto12 7 3 1171 0 0 3.2 15.3 896 11 8 8 6 3 

Valeriana ofi cinalis Vale N n -0.64 175 hc h 0 Rhizl 5 5 2408 808 0 3.4 145 1115 11 6 8 6 5 

Valeriana pyrenaica Vale AN -0.35 110 hc h 0 Eur 195 8 0 2.7 13.8 1151 1, 3 5 7 5 5 

Va lerianella carinata Vale AR 2.15 15 Th h 0 8 3 547t 66 13 4.3 15.8 93 1 3, 17 8 4 8 4 

Valerianella dentata Vale AR -1.86 15 Th h 0 7 3 600 45 4 3.9 15.7 800 4 8 4 7 4 

Valerianella eriocarpa Vale AN -0.69 15 Th h 0 9 2 Eur 59 0 6 4.5 16.0 826 3, 16 8 3 8 3 

Valerianella locusta Vale N n -0.11 15 Th h 0 7 3 1358 238 9 3.9 15.3 925 3, 16, 19 8 4 6 4 

Valerianella rimosa Vale AR CR -2.55 15 Th h 0 7 3 181 27 0 4.3 15.9 860 4 8 4 8 3 

Verbascum lychnids Scro N s -0 .23 150 hc h 13 7 3 42 0 0 4.1 16.5 790 3, 7 71 3 71 3 

Verbascum nigrum Scro N n -0.12 120 

hc h 0 

7 4 

479, 0 8 3.9 16.2 742 3 

7 4 7 6 

Verbascum pulverulentum Scro NA s 0.94 135 

hc h 0 

9 2 

51 0 0 3.4 16.1 624 3, 16 

8 3 7 5 

Verbascum thapsus Scro N n 0.27 200 hc h 0 7 4 1874 375 12 3.7 15.2 918 3 7 4 7 5 

Verbascum virgatum Scro AN 0.35 100 hc h 0 8 2 Eur 339 13 

in 

3 4.2 15.8 870 3, 5, 17, 19 8 4 5 5 

so Verbena ofi cinalis Verb AR -0.43 67 hc h 0 8 5 8571 90 10 4 1 15.8 859 3, 16 8 5 71 6 

Veronica agrestis Scro AR -0.38 10 a Th h 0 7 3 1715 177 9 3.5 15.0 963 3, 4, 17 7 6 6 7 

Veronica alpina Scro N s -0 .29 12 Ch h 0 1 4 35 0 0 -0.7 10.9 1783 15 8 6 5 2 

Veronica anagallts -aquabca Scro N n 0.05 50 a p Hz Hy h Nodel 8 5 1225 459 3 3.7 15.1 899 13, 14 7 10 7 7 

Veronica arvensis Scro N n 0.48 25 a Th h 0 8 3 2614 793 14 3.5 14.6 1077 3, 4, / 6 8 4 6 5 

Veronica be cca bunga Scro N n -0.31 45 Hy Ch h Nodel 7 4 2333 912 9 3.6 14.7 1034 11, 14 7f 10 6 6 

Veronica catenata Scro N n 0.37 50 a p Hz Hy h Nodel 7 6 956 249 4 3.8 15.6 816 13, 14 8 10 7 8 

Veronica chamaedrys Scro N n -0.50 30 Ch hc h Node2 5 4 2609 945 14 3.5 14.5 1095 1, 3, 6 6 5 6 5 

Veronica Milormis Scro AN 2.69 5 hc Ch h Node2 Eur 2013 397 9 3.6 14.9 990 3, 17 7 6 7 7 

Veronica froticans Scro N s 0.11 14 Ch sW 0 1 3 26 0 0 -0 .5 11.2 182 1 15, 16 8 5 7 2 

Veronica hecieriforia Scro AR 0.57 30 a Th h 0 8 3 1944 346 14 3.71 15.1 918 1, 3, 4, 17 6 5 7j 6 

Veronica m ontana Scro N n 0.48 15 Ch h Node2 7 3 1808 506 0 3.6 14.9 1029 1 4 6 6 6 

Veronica of icinalis Scro N n -0.84 23 Ch h Nodel 5 3 2507 754 8 3.4 14.4 1125 7, 8 6 5 4 4 

Veronica persica Scro AN -0.37 30 a Th h 0 Nodal Eur 2232 722 13 3.7 14.9 993 3, 4, 17 6 5 7 7 

Veronica polka Scro AN 0.07 10 a Th h 0 8 4 Eur, Asl 1237 107 8 3.8 15.5 867 4, 17 7 4 7 5 

Veronica praecox Scro AN 15 a Th h 0 8 3 c Eur 6 0 0 3.3 16.3 618 3, 4, 8 8 2 8 1 

Veronica scutellata Scro N n -0.06 25 hc h Nodel 5 4 1877 576 3 3.4 14.3 1129 11, 13 8 9 5 3 

Veronica serpyllifolia Scro N n 0.80 10 hc h Node2 5 6 2652 884 11 3.5 14.5 1104 3, 5, 11 7 5 6 5 

Veronica spicata Scro N s 0.13 45 hc Ch h Nodel 7 4 c 28 0 0 3.7 15.3 949 7, 16 8 3 7 2 

Veronica triphyllos Scro AR EN -0.82 15 a Th h 0 7 3 c 33 0 0 3.5 16.1 690 3, 4 7 4 7 3 

Veronica vema Scro N VU r -0.64 15 a Th h 0 7 4 c 71 0 0 3.3 16.2 618 8 8 2 5 1 

Viburn um lantana Capr N n 0.37 600 Ph w 0 7 3 537 0 0 3.8 16.2 763 1, 3 7 5 7 5 

Viburnu m opulus Capr N n -0.15 400 Ph w 0 7 6 1854 565 1 3.6 15.0 1019 1 6 7 6 6 

Vicia bithynica Faba N s -0.52 60 hc h 0 9 1 74 0 4 4.4 16. 1 827 3 7 4 6 4

Taxon name Fam NS CS RS Chg. Hght Len P1 P2 LF1 LF2 W Clonel Clone2 El E2 C Origin GB IR CI Tjan Tjul Prec Co Br Habitats L F R N $ 

Vicia cracca Faba N n -0.37 120 p hc Rhir l 5 5 2647 938 10 3.6 14.5 1083 3, 11 7 6 5 0 

Vicia laba Faba AN 100 a Th 0 Cro 377, 5 2 3.71 16.1 734 4, 17 8 4 71 0 

Vicia hirsuta Faba N n 0.05 80 a Th 0 7 3 1925' 285 12 3.7 15.2 926 3, 6 7 5 6 0 

Vicia lathyroides Faba N n -0.36 10 a Th 0 7 3 501 27 11 3.8 15.2 808 8, 19 8 3 3 0 

Vicia lutea Faba N s -0.85 50 a Th 0 9 2 58 0 7 4.8 15.9 844 Co 18, 19 7 4 5 1 

Vicia orobus Faba N s -0.34 60 p hc 0 7 2 214 15 0 3.0 13.8 1410 16 7 5 4 

Vicia parvillora Faba N s -1.05 60 a Th 0 9 2 136 0 0 3.9 16.3 695 3, 4 7 5 5 

Vicia sativa Faba N n 0.19 90 a Th 0 8 3 1930 472 14 3.7 14.9 991 6 7 4 4 

Vicia sepium Faba N n -0.43 60 p hc Rhizl 5 4 2615 945 4 3.5 14.5 1095 3, 6 6 5 6 

Vicia sylvatica Faba N n -0.71 150 p hc Rhizl 5 4 623 57 0 3.1 14.3 1119 1, 3, 16 7 5 5 

Vicia tetrasperma Faba N n 0.45 60 a Th 0 7 3 1159 0 10 3.9 15.9 804 3, 4, 6 7 5 6 

Vicia villosa Faba AN 150 a Th 0 8 3 c Eur 147, 2 3 3.8 16.1 757 3, 4 7 4 5 

Vinca maj or Apoc AN 1.49 35 p Ch Tip Eur 1336 231 12 4.0 15.6 873 1, 3, 17 5 6 6 

Vinca minor Apoc AR 0.48 15 p Ch Node2 Eur 1359 12 8 3.7 15.3 893 1, 3, 17 4 6 7 

Viola arvensis \do AR -0.29 40 a Th 0 7 4 2065 33 11 3.6 15.0 940 4 8 4 6 

Viola canina Vio N n -0.87 18 p hc 0 5 4 1030. 18 3.5 14.7 1008 8, 10 8 4 2 

Viola hilt s 

Vio N n -0.46 15 P hc 

0 Rhir l 7 4 

964, 1 3.7 15.71 795 7 

7 4 2 

Viola kitaibeliana 

\do N VU r 

10 a Th 

0 8 3 

2 

6.7 16.6 814 19 

9 3 2 

Viola lactea Vio N s -1.08 15 hc 0 7 1 183 2 5.0 15.8 1025 10 7 6 2 

Viola lutea Vio N n -0.69 20 hc Rhiz2 4 3 536 1 2.0 13.3 1281 7, 8, 16 8 5 2 

Viola odorata Vio N n -0.19 15 hc Sto12 7 3 1340 9 3.8 15.7 833 7 5 5 7 

Viola palustris Vio N n -0.30 15 hc Rhiz2 5 3 1880 56 3.3 13.9 1262 11, 14 7 9 2 

a, 

0 Viola persicifolia Vio N EN r -0.62 25 hc 0 7 4 c 18 1 3.9 15.4 826 11 7 8 3 

Viola reichenbachiana Vio N n 0.20 13 hc 0 7 3 1128 28 3.8 15.5 869 1 4 6 5 

Viola riviniana Vio N n 1.07 18 hc 0 Root 7 3 2741 96 1 3.5 14.4 1112 1, 7, 16 6 5 4 

Viola rupestris Vio N r 5 hc 0 7 5 c 7 1,6 13.3 1393 7 8 3 2 

Vio/a tricolor Vio N n -1.52 20 p Th hc 0 7 3 1691 24 3.4 14.6 992 4, 8, 19 8 4 4 

Viscum album Vise N n 0.97 100 Ch Pn w 0 7 3 923 3.8 15.9 788 3, 17 7 5 5 

Vulpia bromoides Poac N n 0.18 37 Th 0 9 2 1777 51 1 3.8 15.0 1006 3 8 4 3 

Vulpia ciliata Poac N s 0.78 37 a Th 0 9 1 109 4.0 16.3 716 8 9 2 2 

Vulpia fasciculata Poac N s 0.37 55 a Th 0 9 1 109 1 5.0 15.8 911 Co 19 9 3 2 

Vulpia myuros Poac AR 1.55 62 a Th 0 8 4 1134 11 i 3.9 15.6 866 3, 17 8 3 3 

Vulpia unilateralis Poac AN -0.56 35 a Th 0 9 2 Eur 39 3.6 16.2 705 3, 7 9 3 2 

Wahlenbergia hederacea Camp N n -0.30 5 p hc Node2 8 1 314 4 4.1 14.9 1280 11, 14 6 8 3 

Wolfi a arrhiza Lemn N s -0.03 0.1 p Fly Frag 8 4 48 4.4 16.6 752 13 7 11 7 

Woodsia alpina Wood N r 0.11 8 p hc 0 2 6 22 0.4 11.6 2371 15, 16 7 4 2 

Woodsia ilvensis Wood N EN r -0.10 10 p hc 0 2 6 14 1.3 12.4 2108 16 7 3 2 

Zannichellia palustris Zann N n 0.17 50 p Hy Irreg Rhiz2 8 6 1196 20 3.8 15.5 820 13, 14 7 12 7 

Zea mays Poac AC 300 a Th 0 Crop 92 1 1 4.0 15.9 820 4, 17 8 3 7 

Zostera angustifolia Zost N s -0.68 30 p Hy Rhiz2 ? ? 131 27 6 4.3 15.0 926 Co 21 7 12 5 

Zostera marina Zost N n -0.86 50 p Hy Rhiz2 6 6 296 68 13 4.5 14.6 1156 Co 21, 23 6 12 6 

Zostera noltei Zost N s -0.51 12 p Hy Rhiz2 8 5 159, 36 4 4.2 14.9 1020 Co 21 8 11 5

A PPE NDIX 

Sources or Mot hei ht 

Plant heights have been calculated from a range of sources and have been subject to personal 

adjust ment . The main sources were Clapham,Tut in & Warburg ( 1962), Stace ( 1991, 1997) and 

Sell & Murrell ( 1996). Values that were thought to be too big, especially the height of 

procumbent species, were modifi ed using informat ion in Butcher (1961). 

A nearly complet e set of additional height values was taken from sources including Flora lberica 

(Cast roviejo et al., I 986-), a Czech fl ora (Dostal, 1958) and BSBI handbooks such as Crucifers of 

Great Britain and Ireland (Rich, 1991). Other such sources included Aeschimann & Burdet ( 1994), 

Hutchinson & Thomas ( 1966), Jermy, Chater & David (1982), Mei kle ( 1984), Page ( 1997) and 

Preston ( 1995). The new values were checked against t hose derived from the main sources. For 

t hose species where t here was a dif erence of more than a factor of 1.5 between the two 

sources, the discrepancy was resolved by making a judgement as to which seemed most suitable 

for British conditions. In addition, all heights of pteridophytes and of trees and shrubs exceeding 

5 m have been critically examined. 

A difi culty with creeping or procumbent plants is that the fl oras give shoot lengt h but not plant 

height. For several such plants, the canopy height was estimated by Hill, using descriptions and 

personal experience. Likewise Preston estimated t he heights of Aizoaceae (cf Preston & Sell, 

1989). 

Many diffi culties in assigning height to a plant remain. The height of a fern is taken to be the 

length of its fronds. Scapose plants such as Pinguicula lusitanica may be very small except for their 

scape. In several scapose genera (e.g. Bellis, Pilosella, Pinguicula, Plantago, Primula, Pyrola), t he length 

of the leaves is used as a measure of size. On the other hand, rosette plants with substantial 

infl orescences (e.g. Sempervivum tectorum) have been accorded t he height of their infl orescence 

rather t han t he lengt h of t he leaves. It was impossible to be completely consistent. Maximum 

height is a general indication of the size of the plant rather than a precise measure. 

The result ing table is inevitably a complex result of cited values and personal judgement. It was 

clear that aut hors repeat each ot her and that their informat ion is oft en wrong. Overestimates 

abound because botanists take pleasure in ext reme values. For example, British authors and Flora 

Europaea (Tutin et al., 1964- 1980) give t he height of Ulmus glabra as 40 m, higher than any ot her 

elm. This is clearly unrealistic, but authors' copying of height data has meant that the value 40 m 

is constantly repeated. Fortunately Bean ( 1950- 1951) indicates a more realist ic value of 30 m, 

which has been used here. We have in many ot her cases t ried to drive height s down from 

ext reme to typical values, by selecting the publication with the lowest values. 

De tuition o Ii e orms 

Life forms (Raunkiaer, 1934) are an example of what are now called 'plant funct ional types' (cf. 

Box, 1996). A modifi ed version of Raunkiaer's system was set out by Clapham,Tutin & Warburg 

( 1962), who specifi ed a life form for almost all species. Their assignments are a valuable starting 

point but contain many inconsistencies. For PLANTATT, they have been comprehensively 

revised. The categories were partially redefi ned (Table A l). 

6 1

Table A I . Life form categor ies from Clapham ,Tutin & W arburg ( 1962) and revised 

categories 

Category 

(CTW ) 

Phanerophyte, 

mega-, meso- 

Phanerophyte, 

micro- 

Phanerophyte, 

nano- 

Defi nition (CTW ) PLA NTAT T Defi nit io n (database) 

>8 m Ph Mega-, meso- and microphanerophyte 

2-8 m Ph (combined with taller phanerophytes) 

25 cm-2 m Pn Nanophanerophyte 

Chamaephytes 0-25 cm Ch Chamaephyte 

Hemicryptophytes 

W intering buds at soil 

surface 

hc Hemicryptophyte 

Herbs with wintering 

Geophyt es buds below the soil Gb, Gn Geophytes 

surface 

Helophyt es Marsh plants (not recognized) 

Hydrophytes Water plants Hy 

Therophytes 

Passing unfavourable 

season as seeds 

Hz 

T h 

62 

Perennat ing buds submerged during 

winter 

Annual water plant 

Annual land plant, including those living 

on summer mud 

The redefi ned categories are generally self-explanatory. As the database contains plant heights, 

the dist inctions between phanerophytes and chamaephytes, and distinct ions within 

phanerophytes, are not ent irely necessary. If required, new categories can be derived using 

combinations of character's. For example German authors such as Mueller-Dombois & Ellenberg 

( 1974) generally allow chamaephytes to be up to 50 cm tall and distinguish a category of dwarf 

shrubs (denot ed by Z - Zwergstraucher). These are woody or semi-woody chamaephytes. It 

would be possible to reconstruct t his category from our database, allowing for t he fact that 

ordinary maximum heights of many of these bushes might be up to 100 cm. 

Ot her categories included by Mueller-Dombois & Ellenberg ( 1974) are lianes, epiphytes and 

errant hydrophytes (unattached water plants). They defi ne Hemicryptophytes as plants that die 

back to near the ground in the unfavourable season. Although worked out in some detail, their 

scheme remains somewhat tentative and we have not adopted it. 

Exam les o how li e- orm criteria are a lied 

The criteria given above result in many discrepancies between our defi nit ions of life for m and 

those of Clapham,Tutin & Warburg ( 1962). For example, t he distinction between geophytes and

hemicryptophytes is often hard to apply with rhizomatous perennials. In principle, plants whose 

new shoots penet rate the surface before the advent of winter are hemicryptophytes, while those 

whose shoots remain well buried are geophytes. Thus Phragmites australis, with a relatively deep 

rhizome (Preston & Croft , 1997), has the life forms perennial Hy and Gn. In Carex acutiformis, 

however, many new shoots have already emerged by the autumn (Preston & Croft, 1997). It is 

also less strongly aquatic. It is t herefore given the life forms hc and Hy. 

In general, biennials have been classifi ed as hemicryptophytes, but a few such as Euphorbia lathyris, 

which are elongated from the beginning, are chamaephytes. 

It is inst ructive to consider some ot her cases (Table A2), which will not be discussed in detail. 

Table A 2. Som e cases where life-form ( LF) is difficult to apply or where our concept 

dif ers from t hat of C lapham,Tut in & W arburg ( 1962) 

Links between Ii e orms and lent hei ht 

Because chamaephytes and our two categories of phanerophytes are defi ned on the basis of 

height, an eff ort was made to defi ne life forms consistently by reference to plant height (Table 

A3). Sometimes, as in Artemisia campestris, t he overwintering buds are at a substantially lower 

height t han the height of the mature plant. Therefore, this scheme has not been followed exact ly 

but has been used as a guide. 

63

These defi nitions are necessarily imprecise. However, knowing that Ca !tuna vulgans has a typical 

maximum height of 60 cm suggests that it will often grow as a chamaephyte and oft en as a 

nanophanerophyte. This is indeed the case; C. vulgaris is the classic dwarf shrub. 

Table A 3. Chamaephyte and phanerophyte categories in relation to plant height 

Typical maximum 

plant height (cm ) 

3-39 Ch 

40-69 Ch 

70-299 Pn 

300-399 Pn 

400- Ph 

Primary life form Secondary life form 

Cionality 

Clonality can be defined in various ways. In an enumeration for Cent ral Europe, Klime§ et al. 

( 1997) defi ned clonal growt h as synonymous with vegetat ive multiplicat ion, which results in t he 

production of new, genetically identical descendants (ramets) with the potential to become 

independent of t he mot her organism. According to this definition, plants that produce a new 

ramet each year but do not spread laterally (for example many orchids) are clonal. Klime§ et aL 

recognize 21 diff erent clonal types. These are described in Klimet 's excellent website Clopla 

(http://wwri butInn.cas.cz/klimes/clopla .htm); all species of Central Europe, including the majority of 

British and Irish plants, are assigned to one or more clonal types. 

Here we adopt a more familiar defi nit ion of clonal growth, namely vegetative reproduction 

combined with lateral spread — i.e. spreading through the formation of clonal patches or clumps, 

or through fragmentation and subsequent dispersal of plant parts. 

Broad Habit ats 

The Broad Habitat Classifi cation, developed as part of the UK Biodiversity Action Plan (Jackson, 

2000; UK Biodiversity Group, 1998; UK Biodiversity Steering Group, 1995), provides a 

comprehensive framework for surveillance of the UK countryside.We categorized the habitats of 

plants by t heir preferences for Broad Habitats. 

In the UK Biodiversity Action Plan (UK Biodiversity Steering Group, 1995), 37 Broad Habitat 

categories were defi ned. These included some habitats, such as limestone pavements, that are 

quite narrow but were specified because t hey are priority habitats for conservat ion. In a later 

publication (UK Biodiversity Group, 1998), the number of terrest rial and freshwater Broad 

Habitats was reduced to 17. Limestone pavements were included in the Broad Habitat inland 

rock. Ten coastal and marine Broad Habitats were added shortly aft erwards (UK Biodiversity 

Group, 1999). Out of the 27 Broad Habitats, 2 1 support vascular plants (see Table 8, above). 

64 

Pn 

Ph

Broad Habitats are intended to be comprehensive and exclusive. In other words, every place in 

which a vascular plant is found ought to belong to one and only one Broad Habitat. Although 

the defi nit ions of Broad Habitats are in many cases obvious, t here was a need for more precise 

circumscript ion to defi ne t he boundaries between them. For terrest rial and freshwater habitats 

this guidance is now available (Jackson, 2000). The guidance not only gives writt en defi nit ions, but 

relates Broad Habitat s to the units of t he National Vegetation Classifi cation (NVC; Rodwell, 

1991-2000). 

Jackson's (2000) notes for guidance are the basis of the cross-reference and defi nitions used here 

(Table A4). In a small number of cases, we have diverged from her interpretations. The basis of 

our disagreement was oft en the scale at which we wished to view the vegetation. Thus we have 

t reated bog pools as st rictly bog features (not as open water) and we treat rock-free maritime 

grassland as neut ral grassland (not as supralitt oral rock). Ot her interpretat ions are possible but 

result in fl orist ically less homogeneous Broad Habitats. 

Table A 4. Assignment of N VC com munities to Broad Habitats for t he purpose of 

ascribing habitat preferences. Sequential runs of numbers are indicated by hyphens, e.g. 2 1- 

23, which signifi es t he sequence 2 1,22,23. Some NVC communit ies were assigned to two Broad 

Habitats; where this is t he case, the community is marked by an asterisk, to show that it is also 

listed under another Broad Habitat. 

N o Nam e N VC com munities 

Broad leaved, mixed and yew W I- 17,19 

woodland 

2 Coniferous woodland W I8 

3 Boundary and linear feat ures W 2 I-23,24*; OV I 8*, I 9*,21*,22*,24*,25*,27 

4 A rable and horticult ural O V I- 11, 13, 14- 17, 18* , 19*,2 1* ,22*,36 

5 Improved grassland MG6*,7; OV 12,23*,25* 

6 Neutral grassland W 24; MC9- 1 2; MG I-5,6*; MG8- I 3; 0 V28* 

7 Calcareous grassland CG I- 14 

8 Acid grassland U I - 6,19 

9 Bracken U 20; W 25 

10 Dwarf shrub heath H I- 10,12,16,18,2 1; M I5*,16; 0 V 34 

11 Fen, marsh and swamp M4- 14,22-24,25*,26-38; S - I 9,20*,22-28; OV 26,28*,29- 

12 Bog M 1-3,15*, 17-21,25* 

13 Standing water and canals A I-7,8*,9*,10,(11- 16)* , 19*,20*, 21-24 

14 Rivers and st reams A 8*,9*,( 1 I - I 6)*,17,18,19*,20* 

15 Montane habitats H I 3- 15,17,19,20,22; U7- I5,18; W 20 

16 Inland rock U 16, 17,2 1; 0 V37-40,4 I* 

17 Built-up areas and gardens 0 V 20,23*,24*,41* ,42 

18 Supralittor al rock MC 1-8 

19 Supralittor al sediment H I I; SD I- 19 

2 1 Littor al sediment 520,21; SM I-28 

65

It is pertinent here to consider the defi nitions, drawing attention to less obvious features and to 

where we have dif ered from Jackson (2000). 

BH I Broad leaved, mixed and yew woodland 

Broad leaved woodland is defi ned as woodland with broadleaved trees having at least 20% canopy 

cover. In practice, most woodland has closed canopy. Juniper scrub is included in this type if it is 

large enough to result in a woodland understorey (W 19). Other scrub types are treated as edge 

features (BH3), because hedges are often where they occur. The underscrub community W 24 is 

shared between woodland (BH1) and neutral grassland (BH6). 

8 H 2 Coniferous woodland 

This category is clearly defi ned and corresponds to nat ive pine woodland (W 18) in the NVC. 

Stands of planted conifers normally have a subset of the normal broadleaved woodland fl ora 

growing under them. There is no corresponding NVC type. Planted conifers have therefore 

been under-represented, and the fern Dryopteris dilatata, which is t he commonest vascular plant 

in upland conifer plantations, did not appear as a conifer specialist in our fl oristic database, but its 

preference was added by us later 

BH 3 Boundary and linear feat ures 

Apart from scrub that forms hedges, boundary and linear features are not directly crossreferenced 

to the NVC. They include hedges, t ree-lines, walls, earth banks, grass strips and dry 

ditches. Note that roads, t racks and railways in urban areas belong to the urban BH I7. 

BH 4 A rable and horticultural 

Commercial orchards are included here, as well as fi eld crops. Note that hort iculture here is 

defi ned as commercial hort iculture and does not include domestic gardens and allotments, which 

are included in BH 17. The NVC communit ies classifi ed as OV 1- 11,13-19,2 1,22,36 are included 

here. 

BH 5 Im proved grassland 

This has few characteristic species, and is typically dominated by sown ryegrass Lolium perenne or 

clover Trifolium repens. OV I2, the Poo annua-Myosotis arvensis community, is indicated in British 

Plant Communities (Rodwell, 1991-2000) as being mainly a disturbed element in improved pasture 

and has been included here. 

8 H 6 N eutral grassland 

The mesot rophic grassland types MG I-6,8- 13 are the core of t he neutral grassland category. 

Coastal grazing marsh is included here. Four of the coastal clif grasslands, MC9- I2, are not 

necessarily or normally rocky and have been included here in BH6. OV28, an inundation 

grassland, is shared between BH6 and BH 1 I. 

BH 7 Calcareous grassland 

By convention, this includes all NVC types included in the CG class. As a result, montane 

calcareous vegetation is in BH7, along with chalk grassland. 

BH 8 Acid grassland 

Only lowland and subalpine grassland are included here; vegetation types that would normally be 

found above the timberline are in BH I5. Communities U I-6 form the core of this type. The 

fern community U 19 is treated in BH8. It is usually a mosaic element in grassland, though it also 

occurs on steep banks by rivers and could therefore be included also in the broad defi nit ion (not 

used here) of BH I4. 

66

BH 9 Bracken 

This is small type, defi ned as unwooded land dominated by bracken Pteridium aquilinum. 8H9 has 

few characterist ic species, because it is usually a derivative of grassland or forest. Just two NVC 

types, U20 and W 25, are included here. 

BH 10 Dwarf shrub heat h 

The heath type BH 10 includes vegetation with at least 25% cover of Ericaceae or dwarf gorse 

Ulex minor. Coastal heath is included in this category, except for dune heath, which is assigned to 

BHI9. Dwarf shrub communit ies on blanket bogs are st rictly included in BH I2, but this is a 

relat ively minor habitat for them and has not been recognized in t he cross-reference. The 

standard heathland communities H 1- 10,12,16,18,21, toget her with the wet heath communities 

M 15,16 form the core of this community. The bizarre 0 V34, a chive M um schoenoprasum 

community forming a mosaic element on shallow soils in coastal heath, is included here. 

B14 1 I Fen, marsh and swamp 

This is a very wide category and includes reedbeds, swamps, tall-herb fens, fl ushes, springs, 

marshes, rush-past ures and wet grassland. Mud communities of dried-up ponds and riverbeds 

(e.g. 0 V30) have been included here, although they belong more st rictly to BH I3 and BH I4 

unless they are very ext ensive (>0.25 ha). Types M4- 14,22-38, all swamp communities except 

Scirpus maritimus S2 1, and OV 26,28-33,35 are included. We have excluded the bog-like 

communities (Erico-Sphagnion) that are found in very acid lowland valley-mires. 

BH I 2 Bog 

Bogs are defi ned strictly only as ombrot rophic bogs. However, we have lumped all Erico- 

Sphagnion vegetation, including M21, in BH 12 even if it the vegetation occurs in valley mires, 

toget her with bog pool vegetation M 1-3. This is to prevent the characteristic species of BH 11 

being too heterogeneous. The NVC types included are M1-3,15,17-21,25. 

BH I 3 Standing water and canals 

Only aquatic vegetat ion has been cross-referenced to this type, i.e. communit ies A l - 16,19-24. 

Mud communities and marginal vegetation are referred to BH11. 

BH I 4 Rivers and st reams 

This Broad Habitat is t reated in much the same way as BH I3. According to t he strict defi nition, 

all vegetation between the banks of a river counts as belonging to BH I4. For the purposes of 

defi ning characteristic species, river-bank vegetat ion has been t reated as belonging to other types 

such as inland rock. Thus t he communities included in BHI4 are A8,9,11-20. Several of these 

communities are also characteristic of B1-113. 

BH 1 S Mont ane habit at s 

This type is defined by having distinctive arctic-alpine species, provided that they are not 

calcicolous. Calcareous montane vegetation is included in BH7. BH 15 includes montane heaths 

H I3- 15,17,19,20,22, grassier types U7- I5,18, and a willow scrub type W 20. 

BH I 6 Inland rock 

This heterogeneous Broad Habitat includes limestone pavement, cliff s, caves, scree, quarries and 

vegetation on skeletal soils over rock. Communities U 16,17,2 1 and 0 V37-4 1 are assigned to 

BH 16. 

BH I 7 Built-up areas and gardens 

Urban habitats were largely ignored by the NVC surveyors. Most of the characteristic species 

are neophytes, while t he commonest species are widespread nat ives and archaeophytes (Hill, Roy 

67

& Thompson, 2002). Very few NVC communities are really characteristic of the built 

environment. Pavement and wall communities 0 V20,42 are assigned wholly to BH I7; 

0 V23,24,4 I are assigned to BH 17 but shared with other Broad Habitats. 

BH 18 Supra littoral rock 

It is clear from the Habitat Statement for supralitt oral rock (UK Biodiversity Group, 1999) that 

maritime grassland is not included. For that reason, MC9- 11,12 have been treated as neutral 

grassland, BH6. Only MCI-8 are included in BH I8. 

BH 19 Supralittoral sedim ent 

This includes strandlines, shingle, machair and coastal dunes. It includes all the NVC types SD I- 

19, toget her with the dune heath type H I I . 

BH 20 Littoral Rock 

Intert idal rock normally lacks vascular plants and no vascular-plant species or vegetat ion type is 

characterist ic of it. 

BH 2 I Littoral sediment 

This includes saltmarshes and saltmarsh pools. 

BH 23 Inshore sublittoral sediment 

One vascular plant. Zostero marina, occurs in this habitat, which is not considered furt her 

Initial calculation o s ecies' re erences or Broad Habitats 

For the majority of species, habitat preferences were based on quadrat data. Two major datasets 

were available, the original data used to derive the association tables in British Plant Communities 

(NVC; Rodwell, 1991-2000) and quadrat samples collected for Count ryside Survey 2000, 

CS2000 (Haines-Young et aL, 2000). 

The quadrat data used to create the NVC categories were supplied by JNCC as an MS Access 

database. This had been created from fi les in the format of the VESPAN package (Malloch, 1985). 

There was no information on the NVC community to which each quadrat had been assigned. 

Therefore, the computer program TABLEFIT (Hill, 1996) was used to assign 312 16 quadrats to 

NVC communities using species' Domin scores. The median goodness-of-fi t was 67%. Only four 

samples had no fi t; t hese were assigned to a community by personal judgement . NVC plant 

communit ies were then attributed to Broad Habitats using the cross-reference outlined above 

(Table A4). All CS2000 samples had been allocated to BAP Broad Habitat types in the fi eld. 

Aft er quadrats had been assigned to Broad Habitats, species' frequencies in each Broad Habitat 

were calculated. Frequencies were used to calculate preferences for Broad Habitats. For each 

species s and BH h, we calculated an odds-ratio index: 

Qsh 

= observed odds for species s in habitat h / expected odds 

nsh 

/ (ns+ "sh + 0.5) 

esh/ Os+ esh) 

where 

nsh = observed number of occurrences of species s in BH h 

ns+ = 1h nsh = total number of occurrences of speCies s 

esh = expected number of occurrences of species s in BH h. 

68

The expected number of occurrences 

where 

esh 

= ns N h / N + 

Nh = number of quadrat samples in BH h 

N 4. = E h N h = total number of quadrat samples. 

Species s was deemed to have a preference for BH h if 

Qsh 4. 

Extension and validation o s ecies' re erences 

Many rare, scarce or uncommon native species are not represented either in the NVC quadrats 

or in CS2000 samples. Furt hermore, t here was the possibility that uneven sampling of Broad 

Habitats could have resulted in some wrong indications of preference. 

First, we ext ended the list of BH preferences to species not adequately sampled by the NVC or 

CS2000, using the text prepared for the New Atlas, together with other literature sources (e.g. 

Stewart, Pearman & Preston, 1994; W igginton, 1999). Much of the initial work was done by Bill 

Meek (CEH); t he resulting table of preferential species was checked by D.A. Pearman (BSB1), as 

well as by ourselves and ot her CEH staf . In addition, preferences were checked against ot her 

data in PLANTATT For example, species attributed to BH I3 (Standing water and canals) and 

BH 14 (Rivers and streams) were checked for their Ellenberg moisture (F) values. Those for 

which F

A CKN O WLED GEMEN TS 

Much of the work presented here was support ed by funding from the W ildlife and Countryside 

Division of Defra and its predecessor DETR. The immediate stimulus was a project to interpret 

the New Atlas of the British and Irish Flora (Preston et al., 2003). A requirement for non-spatial 

attributes of plants was written into the 2002 programme of work for the National Biodiversity 

Network (NBN). Informat ion on native status was prepared for the New Atlas and presented 

there (Preston, Pearman & Dines, 2002). The Ellenberg indicator values were developed under 

the DETR-funded ECO FACT programme and published separately (Hill et al., 1999). 

The other main ot her source of funding was from our parent body the Natural Environment 

Research Council. Not only have they generously support ed the Biological Records Centre 

(BRC) and NBN over the years, but they have allowed us to fi nd additional t ime to work our 

data up into publishable form. The geographical classifi cat ion of European distributions (Preston 

& Hill, 1997) was developed with support from NERC's special programme on global 

environmental change,TIGER. 

Some elements of t his work have been funded by the Joint Nat ure Conservation Committee 

through t heir joint sponsorship of BRC. Other aspects were supported by t he Countryside 

Council for Wales, English Nat ure, Scottish Natural Heritage, Environment and Heritage Service 

(Northern Ireland) and Plantlife as part of the analysis of New Atlas data. 

Publication of this booklet has been jointly funded by CEH and Defra. 

We are grateful to the following for particular help: 

De fra 

Andrew Stott (overall guidance and long-term support) 

Debbie Boobyer (Broad Habitats and analysis of New Atlas data) 

BSBI 

David Pearman (Broad Habitats) 

Cambridge University 

Peter Sell (unpublished fl ora draft s) 

English Nature 

Roger Key (Cover photograph) 

JNCC 

Ian St rachan (NVC quadrat data) 

CEH 

Bob Bunce (Ellenberg values) 

Jon Cooper (liaison with NBN) 

Julie Gaunt (graphic design) 

Bill Meek (Broad Habitats) 

Owen Mountford (Ellenberg values, European fl oras, species attributes) 

Peter Rothery (Change index) 

Mark Telfer (Change index) 

70

B IB LI OGRA PH Y 

Aeschimann, D. & Burdet, H.M. ( 1994) Fiore de la Suisse. Editions du Grif on, Neuchatel. 

Barrow, E.M., Hu Ime, M. and Jiang,T. (1993) A 1961-90 baseline climatology and future climate 

change scenarios for Great Britain and Europe. Part 1: 1961-90 Great Britain baseline climatology. 

Climatic Research Unit, Norwich, 43pp. (plus maps). 

Bean,W.J. ( 1950- 1951) Trees and shrubs hardy in the British Isles. 7 edn. John Murray, London. 

Box, E.O. ( 1996) Plant functional types and climate at the global scalej ournal of Vegetation Science, 

7, 309-320. 

Butcher, R.W. ( 1961) A new illustrated British fl ora. Leonard Hill, London. 

Castrovielo, S. et al., eds. ( 1986-) Flora iberica: plantas vasculares de la Peninsula iberica e Islas 

Baleares. Vol. 1 -. Real Jardin Botanico, Madrid. 

Cheffi ngs, C. (2004) New plant status lists for Great Britain. 8581 News, 95, 36-43. 

Clapham, A.R.,Tut in,T.G. & Warburg, E.F. ( 1962) Flora of the British Isles. 2 edn. Cambridge 

University Press, Cambridge. 

Dostald . ( 1958) Klic k Opine kvetene CSR. 2 edn. Ceskoslovenske Akademie Ved, Prague. 

Ellenberg, H.,Weber, H. E., Dull, R.,W irt h,V,Werner,W. & Paulissen, D. ( 1991) Zeigerwerte von 

Pfl anzen in Mitt eleuropa. Scripta Geobotanica, 18, 1-248. 

Fitt er, A.H. & Peat, H.J. ( 1994) The Ecological Flora Database. Journal of Ecology, 82, 4 15-425. 

Grime, J. R ( 1979) Plant strategies and vegetation processes. John W iley, Chichest er. 

Grime, J. P. (2001) Plant strategies, vegetation processes, and ecosystem properties.W iley, Chichester. 

Gr imel .R,Thompson, K., Hunt, R., Hodgson, J.G., Cornelissen, J.H.C., Rorison, 1.H., Hendry, G.A.F, 

Ashenden,T.W.,A skew,A.R, Band, S.R., Booth, R.E., Bossard, C.C., Campbell, B.D., Cooper, 

J.E.L., Davison, A.W., Gupta, PL., Hall,W , Hand, D.W., Hannah, M.A., Hillier, S.H., Hodkinson, 

D.J., Jahli,A ., Liu, Z., Mackey, J.M.L., Matthews, N., Mowforth, M.A., Neal,A.M., Reader, R.J., 

Reiling, K., Ross-Fraser,W , Spencer, R.E., Sutton, F.,Tasker, D.E., Thorpe, P.C. & W hitehouse, J. 

( 1997) Integrated screening validates primary axes of specialisation in plants. Oikos, 79, 259- 

281. 

Haines-Young, R.H., Barr, C.J., Black, H.1.J., Briggs, D.J., Bunce, R.G.H., Clarke, R.T , Cooper, A., 

Dawson, F.H., Firbank, L.G., Fuller, R.M., Furse, MI , Gillespie, M.K., Hill, R., Hornung, M., 

Howard, D.C., McCann,T , Morecroft, M.D., Petit, S., Sier,A.R.J., Smart , S.M., Smith, G.M., Stott, 

A.R, St uart , R.C. & Watkins, J.W. (2000) Accounting for nature: assessing habitats in the UK 

countryside. Depart ment of the Environment,Transport and the Regions, London. 

Hill, M.O. ( 1996) Tablefit version 1.0, for identification of vegetation types. Institute of Terrestrial 

Ecology, Huntingdon. 

Hill, M.O., Mountford, J.O., Roy, D.B. & Bunce, R.G.H. (1999) Ellenberg's indicator values for British 

plants. ECO FACT Volume 2 technical annex. Institute of Terrestrial Ecology, Huntingdon. 

Hill, M.O., Roy, D.B. & Thompson, K. (2002) Hemeroby, urbanity and ruderality: bioindicators of 

disturbance and human impact. Journal of Applied Ecology, 39, 708-720. 

Hodgson, J.G., Grime, J.R, Hunt, K. & Thompson, K. ( 1994) The electronic comparative plant ecology. 

Chapman & Hall, London. 

Hodgetts, N., Palmer, M. & W igginton, M. ( 1996) The Pink Book of Plants: lists of vascular and nonvascular 

plant species which are nationally threatened, localised or protected in Great Britain. 

JNCC, Peterborough.

Hu Ime, M. & Jenkins, G. ( 1998) Climate change scenarios for the United Kingdom: summary report 

Climate Research Unit, Norwich. 

Hutchinson, G. & Thomas, B.A. ( 1996) Welsh ferns: clubmosses, quillworts and horsetails. 7 edn. 

National Museums and Galleries of Wales, Cardif . 

Jackson, D.L. (2000) Guidance on the interpretation of the Biodiversity Broad Habitat Classifi cation 

(terrestrial and freshwater types): Defi nitions and the relationship with other habitat classifi cations. 

JNCC Repor t No. 307. JNCC, Peterborough. 

Jermy, A.C., Chater, A.O. & David, R.W. ( 1982) Sedges of the British Isles. Botanical Society of the 

British Isles, London. 

Klim8 , L., Klimegova J Hendriks, R. & van Groenendael, J. ( 1997) Clonal plant architecture: a 

comparative analysis of form and funct ion. In The ecology and evolution of clonal plants (eds H. 

de Kroon & J. van Groenendael), pp. 1-29. Backhuys, Leiden. 

Lindacher, R. ( 1995) PHANART: Datenbank der Gefasspfl anzen Mitteleuropas. Vertiffentlichungen 

des Geobotanischen Institutes der ETH, Stiftung RUbel, Zurich, I 25, 1-436. 

Macpherson, P., Dickson, J.H., Ellis, R.G., Kent, D.H. & Stace, C.A. ( 1996) Plant status 

nomenclature. M I News, 72, 13-16. 

Malloch,A.J.C. ( 1985) VESPAN: FORTRAN programs for handling and analysis of vegetation data and 

species distributions. Depart ment of Biological Sciences, University of Lancaster. 

Meikle, R.D. ( 1984) Willows and poplars of Great Britain and Ireland. Botanical Society of t he British 

Isles, London. 

Mueller-Dombois, D. & Ellenberg, H. ( 1974) Aims and methods of vegetation ecology. John W iley & 

Sons, New York. 

Page, C.N. ( 1997) The ferns of Britain and Ireland. Cambridge University Press, Cambridge. 

Palmer, M.A., Hodgetts, N.G.,W igginton, Mj ., Ing, B. & Stewart , N.F. ( 1997) The application to t he 

Brit ish fl ora of the World Conservation Union's revised red list criteria and the signifi cance 

of red lists for species conservation. Biological Conservation, 82, 2 19-226. 

Perring, EH. & Walters, S.M. ( 1962) Atlas of the British Flora.Thomas Nelson & Sons, London. 

Preston, C.D. ( 1995) Pondweeds of Great Britain and Ireland. Botanical Society of the British Isles, 

London. 

Preston, C.D. & Croft, J.M. ( 1997) Aquatic Plants in Britain and Ireland. Harley Books, Colchester. 

Preston, C.D. & Hill, M.O. ( 1997) The geographical relationships of Brit ish and Irish vascular 

plants. Botanical Journal of the Linnean Society, 124, I - 120. 

Preston, C.D. & Hill, M.O. ( 1999) The geographical relationships of the British and Irish fl ora: a 

comparison of pteridophytes, fl owering plants, liverworts and mosses. Journal of Biogeography, 

26, 629-642. 

Preston, C.D., Pearman, D.A. & Dines,T.D., eds. (2002) New atlas of the British and Irish Flora. 

Oxford University Press, Oxford. 

Preston, C.D. & Sell, RD. ( 1989) The Aizoaceae naturalized in the British Isles. Watsonia, 17, 217- 

245. 

Preston, C.D.,Telfer, M.G.,Arnold, H.R., Carey, RD., Cooper, J.M., Dines,T.D., Hill, M.O., Pearman, 

D.A., Roy, D.B. & Smart , S.M. (2002) The changing fl ora of the UK. DEFRA, London. 

Preston, C.D.,Telfer, M.G., Roy, D.B., Carey, RD., Hill, M.O., Meek,W.R., Rot hery, P. Smart, 

Smith, G.M.,Walker, K.J. & Pearman, D.A. (2003) The changing distribution of the fl ora of the 

United Kingdom: technical report Report to DEFRA on CEH project C0 1093. CEH Monks 

Wood, Huntingdon. 

72

Raunkiaer, C. ( 1934) The life forms of plants and statistical plant geography. Oxford University Press, 

Oxford. 

Rich,T.C.G. ( 1991) Crucifers of Great Britain and Ireland. Botanical Society of the Brit ish Isles, 

London. 

Rodwell, J.S. ( 1991-2000) British plant communities. Cambridge University Press, Cambridge. 

Sell, P. & Murrell, G. ( 1996) Flora of Great Britain and Ireland. Volume 5 Butomaceae-Orchidaceae. 

Cambridge University Press, Cambridge. 

Stace, C.A. ( 1991) New fl ora of the British Isles. Cambridge University Press, Cambridge. 

Stace, C.A. ( 1997) New flora of the British Isles. 2 edn. Cambridge University Press, Cambridge. 

Stewart, A., Pearman, D.A. & Preston, C.D., eds. ( 1994) Scarce plants in Britain. JNCC, 

Peterborough. 

Telfer, M.G., Preston, C.D. & Rothery, P. (2002) A general method for the calculation of relat ive 

change in range size from biological atlas data. Biological Conservation, 107, 99- 109. 

Tutin,T.G., Heywood,V.H., Burges, NA ., Moore, D.M.,Valent ine, D.H., Walters, S.M. & Webb, D.A. 

( 1968- 1980) Flora Europaea. Volumes 2-5. Cambridge University Press, Cambridge. 

Tutin,I G., Heywood,V.H., Burges, N.A.,Valent ine, D.H.,Walters, S.M. & Webb, D.A. ( 1964) Flora 

Europaea. Volume I. Cambridge University Press, Cambridge. 

UK Biodiversity Group ( 1998) Action Plan Volume 2:Terrestrial and freshwater habitats. English 

Nat ure, Peterborough. 

UK Biodiversity Group ( 1999) Tranche 2 Action Plans.Volume S. Maritime species and habitats. 

English Nat ure, Peterborough. 

UK Biodiversity Steering Group ( 1995) Biodiversity: the UK Steering Group Report. Volume 1, 

Meeting the Rio challenge; Volume 2: Action plans. HMSO, London. 

W igginton, M.J., ed. ( 1999) British Red Data Books. I Vascular plants, 3 edn. Joint Nature 

Conservat ion Committee, Peterborough. 

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