001.0000
Family
Lycopodiaceae P.Beauv. ex Mirb.; Hist. Nat. Veg. (Lam. & Mirb.) 4: 293 (1802)
17 genera, ca. 470 spp. Phylogeny: Chen et al. (2021).
Species listed in database: 476; hybrids: 20
= Huperziaceae Rothm.; Feddes Repert. 66: 236 (1962), nom. nov.
= Urostachyaceae Rothm.; Feddes Repert. 54: 58 (1944) (nom. illeg.)
= Phylloglossaceae Kunze; Bot. Zeitung (Berlin) 1: 722 (1843)
= Lycopodiellaceae Val.N.Tikhom.; Novosti Sist. Vyssh. Rast. 49: 149 (2018)
001.4002
Genus
Phlegmariurus (Herter) Holub; Preslia 36(1): 179 (1964)
Distribution: Worldwide tropical. There is a clear distinction between a neotropical clade and an Old World clade but this is not sufficient to split two genera (Chen et al. 2021).
Species listed in database: 307; hybrids: 1
species :
Phlegmariurus acerosus
(Sw.) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Guatemala; Costa Rica; Panama; Cuba; Hispaniola (Haiti, Dominican Republic); Guadeloupe; Trinidad; Colombia (Cesar, Valle); Venezuela (Aragua, Falcon, Tachira); Ecuador; Bolivia (La Paz); Brazil (Amazonas, Bahia, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Argentina (Misiones)
80 COS GUA PAN 81 CUB GDL HIS TRT 82 CLM VEN 83 BOL ECU 84 BRA 85 ARG
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia acerosa (Sw.) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Lycopodium acerosum Sw.; Fl. Ind. occ. 3: 1575 (1806)
= Urostachys acerosus (Sw.) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1(4): 399, t. 18 (1927)
species :
Phlegmariurus acifolius
(Rolleri) B.Øllg.; Phytotaxa 57: 11 (2012)
Distribution: Colombia (Huila, Norte de Santander, Putumayo, Quindío, Cauca); Venezuela (Tachira); Ecuador; Peru
82 CLM VEN 83 ECU PER
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia acifolia (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21(100): 155 (1988)
= Lycopodium acifolium Rolleri; Notas Mus. La Plata, Bot. 14(87): 2, t. 1-2 (1985)
species :
Phlegmariurus acutus
(Rolleri) B.Øllg.; Phytotaxa 57: 11 (2012)
Distribution: Bolivia (La Paz)
83 BOL
Remarks: saururus agg. Known only from type collection
Synonyms: hidden
= Lycopodium acutum Rolleri; Notas Mus. La Plata, Bot. 13(71): 61-114 (1981)
= Huperzia acuta (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21(100): 155 (1988)
species :
Phlegmariurus affinis
(Trevis.) B.Øllg.; Phytotaxa 57: 11 (2012)
Distribution: Colombia (Boyacá, Huila, Nariño); Ecuador; Peru
82 CLM 83 ECU PER
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia affinis Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874), nom. nov. for Lycopodium affine Hook. & Grev.
= Huperzia blepharodes (Maxon) Holub; Folia Geobot. Phytotax. 20: 71 (1985) (nom. superfl. for H. affinis Trevis.)
= Lycopodium affine Hook. & Grev.; Bot. Misc. 2: 364 (1831) (nom. illeg., non Lycopodium affine Bory (1804))
= Lycopodium blepharodes Maxon; Contr. U.S. Nat. Herb. 17(4): 423 (1914), nom. nov. for Lycopodium affine Hook. & Grev.
= Urostachys affinis (Hook. & Grev.) Nessel; Repert. Spec. Nov. Regni Veg. 39: 69 (1935)
= Urostachys blepharodes (Maxon) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 53 (1949)
= Urostachys involutus Herter ex Nessel; Revista Sudamer. Bot. 6(5-6): 162, t. 10, f. 39 (1940)
species :
Phlegmariurus afromontanus
(Pic.Serm.) A.R.Field & Bostock; PhytoKeys 20: 36 (2013)
Distribution: E-D.R.Congo [Zaire]; Rwanda; Burundi; Tanzania (Mt. Kilimanjaro)
23 BUR RWA ZAI 25 TAN
Remarks: ophioglossoides agg. Possibly to be included in H. ophioglossoides
Synonyms: hidden
= Huperzia afromontana Pic.Serm.; Webbia 27: 394, fig. 2 (1972 publ. 1973)
= Lycopodium afromontanum (Pic.Serm.) Kornas; Prace Bot. (Krakow) 25: 33 (1993)
species :
Phlegmariurus ambrensis
(Rakotondr.) A.R.Field & Bauret; Molec. Phylogen. Evol. 127: 499 (2018)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Huperzia ambrensis Rakotondr.; Candollea 70(1): 50 (2015)
species :
Phlegmariurus amentaceus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Guatemala; Colombia (Antioquia, Cundinamarca, Quindío); Venezuela (Merida, Trujillo)
80 GUA 82 CLM VEN
Remarks: myrsinites agg.
Synonyms: hidden
= Huperzia amentacea (B.Øllg.) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Lycopodium amentaceum B.Øllg.; Amer. Fern J. 72: 53 (1982)
species :
Phlegmariurus andinus
(Rosenst.) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Colombia (Chocó, Magdalena); ?Ecuador; Peru; Bolivia (Cochabamba, La Paz); Argentina (Buenos Aires, Catamarca, Cordoba, Jujuy, Salta, San Luis, Tucuman)
82 CLM 83 BOL PER 85 ARG
Remarks: saururus agg. Contains obviously records of P. saururus from South America
Synonyms: hidden
= Huperzia andina (Rosenst.) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Lycopodium andinum Rosenst.; Repert. Spec. Nov. Regni Veg. 5: 239 (1908)
= Lycopodium breve Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909)
= Urostachys andinus (Rosenst.) Nessel; Bärlappgew. 76 (1939)
= Urostachys brevis (Herter) Herter ex Nessel; Bärlappgew. 82 (1939)
species :
Phlegmariurus aqualupianus
(Spring) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Cuba; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Trinidad; Colombia (Antioquia, Cundinamarca, Huila, Meta, Putumayo, Valle, Magdalena, Santander); Venezuela (Aragua, Bolívar, Venezuela Distrito Federal, Falcon, Guarico, Lara, Miranda, Monagas, Portuguesa, Tachira, Yaracuy, Zulia); Guyana; Brazil (Minas Gerais)
81 CUB DOM GDL HIS MAR PUE STL STV TRT 82 CLM GUY VEN 84 BRA
Remarks: aqualupiana agg.
Synonyms: hidden
= Huperzia aqualupiana (Spring) Rothm.; Feddes Repert. 54: 62 (1944)
= Lycopodium aqualupianum Spring; Bull. Acad. Roy. Soc. Bruxelles 8: 518 (1841)
= Lycopodium guadalupianum Spring ex Fée; Mém. Foug. 11 (Hist. Foug. Antil.) 1310, t. 33p, f. 1 (1866) (nom. superfl.)
= Urostachys aqualupianus (Spring) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 166 (1923)
species :
Phlegmariurus arcuatus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: S-Colombia (?Cauca, Narino, Risaralda); Ecuador; Peru
82 CLM 83 ECU PER
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia arcuata B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 15, f. 1D (1988)
species :
Phlegmariurus aristei
(Nessel) B.Øllg.; Phytotaxa 630(3): 192 (2023)
Distribution: Colombia (Cundinamarca); Peru; Bolivia (Cochabamba, Chuquisaca, La Paz, Tarija); Argentina (Jujuy, Salta, Tucuman)
82 CLM 83 BOL PER 85 ARG
Remarks: taxifolia agg. Synonymy fide Øllgaard in Kessler et al. (2023)
Synonyms: hidden
= Huperzia aristei (Nessel) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 155 (1988)
= Huperzia buesii (Herter) B.Øllg.; Fieldiana, Bot. n. s. 34: 42 (1994)
= Huperzia sotae (Rolleri) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium aristei (Nessel) Rolleri; Rev. Mus. La Plata (N. S.) Bot. 13: 94 (1981)
= Lycopodium buesii (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium sotae Rolleri; Darwiniana 16: 130, f. 1 (A-C), t. 2, t. 3(A) (1970)
= Phlegmariurus buesii (Herter) B.Øllg.; Phytotaxa 57: 12 (2012)
= Phlegmariurus sotae (Rolleri) B.Øllg.; Phytotaxa 57: 19 (2012)
= Urostachys aristei Nessel; Repert. Spec. Nov. Regni Veg. 36: 181, t. 172 (1934)
= Urostachys buesii Herter; Revista Sudamer. Bot. 10: 126 (1953 publ. 1954)
species :
Phlegmariurus ascendens
(Nessel) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Colombia (Boyacá, ?Narino); Ecuador
82 CLM 83 ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia ascendens (Nessel) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Urostachys ascendens Herter ex Nessel; (Bärlappgew. 74 (1939) (nom. inval.) Revista Sudamer. Bot. 6: 161, t. 9, f. 30 (1940)
species :
Phlegmariurus attenuatus
(Spring) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Costa Rica; Colombia (Boyacá, Cundinamarca, Nariño, Risaralda); Ecuador; Peru; Bolivia
80 COS 82 CLM 83 BOL ECU PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia attenuata (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1875)
= Lycopodium attenuatum Spring; Mém. Acad. Roy. Sci. Belg. 24: 8 (1849)
= Lycopodium attenuatum var. crassum Nessel; Bärlappgew. 74 (1939)
= Lycopodium tobarii Sodiro; Crypt. Vasc. Quit. 565 (1893) (Anales Univ. Quito 12: 320 (1895))
= Urostachys attenuatus (Spring) Herter ex Nessel; Bärlappgew. 101 (1939)
= Urostachys tobarii (Sodiro) Herter ex Nessel; Bärlappgew. 101 (1939)
species :
Phlegmariurus australis
(Willd.) A.R.Field; PhytoKeys 109: 63 (2018)
Distribution: Vanuatu; Tonga; Fiji (Viti Levu, Vanua Levu); Marquesas Isl. (Nuku Hiva, Ua Pou, Hiva Oa, Fatu Hiva); Society Isl.
60 FIJ TON VAN 61 MRQ SCI
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia australis (Willd.) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Huperzia subtrifoliata (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium australe Willd.; Sp. Pl. ed. 4. 5: 11 (1810)
= Lycopodium phlegmaria var. australe (Willd.) Domin; Biblioth. Bot. 20(85): 228 (1915)
= Lycopodium phlegmaria var. marchionicum E.D.Br.; Bernice P. Bishop Mus. Bull. 89: 108, pl. 20 (1931)
= Lycopodium subtrifoliatum Brownlie; Beih. Nova Hedwigia 55: 24 (1977)
= Phlegmariurus subtrifoliatus (Brownlie) A.R.Field & Bostock; PhytoKeys 20: 48 (2013)
= Urostachys australis (Willd.) Herter ex Nessel; Bärlappgew. 224 (1939)
species :
Phlegmariurus austroecuadoricus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Ecuador
83 ECU
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia austroecuadorica B.Øllg.; Fl. Ecuador 33: 31 (1988)
species :
Phlegmariurus austrosinicus
(Ching) Li Bing Zhang; Fl. Reipubl. Popularis Sin. 6(3): 42 (2004)
Distribution: China (Chongqing, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Hongkong, Yunnan)
36 CHI
Remarks: selago agg.
Synonyms: hidden
= Huperzia austrosinica Ching; Acta Bot. Yunnan. 3(3): 298-299 (1981)
species :
Phlegmariurus axillaris
(Roxb.) comb.ined.
Distribution: St. Helena
28 STH
Remarks: saururus agg. Frequently included in H. saururus s. lat. or in H. insularis but kept separate by Roux (2009)
Conservation Status: IUCN V (St. Helena)
Synonyms: hidden
= Huperzia axillaris (Roxb.) Rothm.; Feddes Repert. 54: 60 (1944)
= Lycopodium axillare Roxb.; Beats., St. Helena 312 (1816)
= Urostachys axillaris (Roxb.) Herter ex Nessel; Bärlappgew. 82 (1939)
species :
Phlegmariurus badinianus
(B.Øllg. & P.G.Windisch) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Minas Gerais, Rio de Janeiro, Santa Catarina)
84 BRA
Remarks: saururus agg.
Synonyms: hidden
= Huperzia badiniana B.Øllg. & P.G.Windisch; Bradea 5: 7, fig. 1A (1987)
species :
Phlegmariurus balansae
(Herter) A.R.Field & Bostock; PhytoKeys 20: 36 (2013)
Distribution: New Caledonia
60 NWC
Remarks: myrtifolia agg.
Synonyms: hidden
= Huperzia balansae (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium balansae Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 51 (1909)
= Lycopodium balansae var. pomperyanum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 52 (1909)
= Urostachys balansae (Herter) Herter ex Nessel; Bärlappgew. 208 (1939)
= Urostachys pomperyanus (Herter) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 76 (1949)
species :
Phlegmariurus bampsianus
(Pic.Serm.) A.R.Field & Bostock; PhytoKeys 20: 36 (2013)
Distribution: Rwanda; Tanzania (Kilimanjaro)
23 RWA 25 TAN
Remarks: ophioglossoides agg.
Synonyms: hidden
= Huperzia bampsiana Pic.Serm.; Bull. Jard. Bot. Natl. Belg. 55: 193 (1985)
= Lycopodium bampsianum (Pic.Serm.) Kornas; Práce Bot. Univ. Wroclawsk. 25: 33 (1993)
species :
Phlegmariurus banayanicus
(Herter) A.R.Field & Bostock; PhytoKeys 20: 36 (2013)
Distribution: Philippines; New Guinea (Irian Jaya, Papua New Guinea)
42 NWG PHI
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia banayanica (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Urostachys banayanicus Herter; Bot. Arch. 3: 17 (1923)
= Lycopodium banayanicum (Herter) Copel.; Univ. Calif. Publ. Bot. 12: 384 (1931)
species :
Phlegmariurus beitelii
(B.Øllg.) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Venezuela (Amazonas, Bolívar); Guyana (Mt. Roraima)
82 GUY VEN
Synonyms: hidden
= Huperzia beitelii B.Øllg.; Amer. Fern J. 79: 151 (1989)
species :
Phlegmariurus biformis
(Hook.) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Bahia, Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
84 BRA
Remarks: myrsinites agg.
Synonyms: hidden
= Huperzia biformis (Hook.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium biforme Hook.; Hooker´s Icon. Pl. 3: t. 228 (1839)
= Urostachys biformis (Hook.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 52 (1949)
species :
Phlegmariurus billardierei
(Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
Distribution: New Zealand (North Isl., South Isl., Three Kings Isl., Stewart Isl.); Kermadec Isl.; Chatham Isl.
51 CTM KER NZL
Synonyms: hidden
= Huperzia billardierei (Spring) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 61 (1944)
= Lycopodium billardierei Spring; Mon. Lycop. 1, 56 (1843)
= Lycopodium varium var. billardierei (Spring) Kirk; Trans. & Proc. New Zealand Inst. 16, 377 (1884)
= Phlegmariurus billardierei (Spring) Brownsey & Perrie; New Zealand J. Bot. 58(2): 126 (2019), isonym
= Urostachys billardierei (Spring) Herter ex Nessel; Bärlappgew. 194 (1939)
species :
Phlegmariurus binervius
(Herter) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Peru
83 PER
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia binervia (Herter) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium binervium Spring ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 48 (1909)
= Urostachys binervius (Spring ex Herter) Herter ex Nessel; Bärlappgew. 110 (1939)
species :
Phlegmariurus bolanicus
(Rosenst.) A.R.Field & Bostock; PhytoKeys 20: 36 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia bolanica (Rosenst.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Huperzia kaysseri (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium bolanicum Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 181 (1913)
= Urostachys bolanicus (Rosenst.) Herter ex Nessel; Bärlappgew. 234 (1939)
= Urostachys kaysseri Nessel; Bärlappgew. 31, t. 4, f. 10 (1939) (nom. inval.), et Revista Sudamer. Bot. 6: 156 (1940)
species :
Phlegmariurus brachiatus
(Maxon) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Isla del Coco
80 COC
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia brachiata (Maxon) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium brachiatum Maxon; Contr. U.S. Natl. Herb. 17: 176 (1913)
= Urostachys brachiatus (Maxon) Herter ex Nessel; Bärlappgew. 127 (1939)
species :
Phlegmariurus brachystachys
(Baker) A.R.Field & Bostock; PhytoKeys 20: 36 (2013)
Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]
22 GUI IVO LBR NGA SIE 23 BIO CMN
Remarks: gnidioides agg.
Synonyms: hidden
= Huperzia brachystachya (Baker) Pic.Serm.; Webbia 23: 162 (1968)
= Lycopodium brachystachys (Baker) Alston; Bol. Soc. Broter. Ser. 2, 30: 19 (1956) (non Hook. & Grev. (1832))
= Lycopodium dacrydioides var. brachystachys Baker; Handb. Fern-Allies 18 (1887)
= Urostachys brachystachys (Baker) Herter ex Nessel; Bärlappgew. 189 (1939)
species :
Phlegmariurus bradeorum
(Christ) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Mexico (Chiapas); Guatemala; Nicaragua; Costa Rica; Panama; Isla Margarita
79 MEX 80 COS GUA NIC PAN 81 VEN
Remarks: wilsonii agg.
Synonyms: hidden
= Huperzia bradeorum (Christ) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium bradeorum Christ & Nutt.; Repert. Spec. Nov. Regni Veg. 8: 20 (1910)
= Urostachys bradeorum (Christ) Herter ex Nessel; Bärlappgew. 154 (1939)
species :
Phlegmariurus brevifolius
(Hook. & Grev.) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Costa Rica; Colombia (Valle, Boyacá, Cundinamarca, Meta, Nariño, Putumayo, Risaralda); Ecuador; Peru
80 COS 82 CLM 83 ECU PER
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia brevifolia (Hook. & Grev.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium brevifolium Hook. & Grev.; Bot. Misc. 3: 104 (1832)
= Urostachys brevifolius (Hook. & Grev.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 54 (1949)
= Urostachys rufescens var. brevifolius (Hook. & Grev.) Herter ex Nessel; Bärlappgew. 103 (1939)
species :
Phlegmariurus brongniartii
(Spring) B.Øllg.; Phytotaxa 57: 12 (2012)
Distribution: Colombia (Antioquia, Cauca, Putumayo); Ecuador; Peru; Bolivia (La Paz, Santa Cruz)
82 CLM 83 BOL ECU PER
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia brongniartii (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium brongniartii Spring; Bull. Acad. Roy. Soc. Bruxelles 8(2): 515 (1841)
= Lycopodium taxifolium var. brongniartii (Spring) Baker; Handb. Fern-Allies 16 (1887)
= Urostachys brongniartii (Spring) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 388 (1927)
species :
Phlegmariurus callitrichifolius
(Mett.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cundinamarca, Risaralda, Tolima); Ecuador
82 CLM 83 ECU
Remarks: myrsinites agg.
Synonyms: hidden
= Huperzia callitrichifolia (Mett.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium callitrichifolium Mett.; Ann. Sci. Nat., Bot., sér. V, 3: 309 (1865)
= Urostachys callitrichifolius (Mett.) Herter ex Nessel; Bärlappgew. 253 (1939)
species :
Phlegmariurus campianus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Colombia (Cundinamarca, Norte de Santander); Ecuador; Peru
82 CLM 83 ECU PER
Remarks: myrsinites agg.
Synonyms: hidden
= Huperzia campiana B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 109, f. 22B (1988)
species :
Phlegmariurus cancellatus
(Spring) Ching; Acta Bot. Yunnan. 4(2): 122 (1982)
Distribution: China (Yunnan); Tibet; Japan; Ryukyu Isl.; Bhutan; India (Arunachal Pradesh); Myanmar; Vietnam
36 CHI TIB 38 JAP NNS 40 BHU IND 41 MYA VIE
Remarks: cancellatus agg.
Conservation Status: IUCN VU (Fraser-Jenkins 2012)
Synonyms: hidden
= Huperzia cancellata (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
= Lycopodium cancellatum Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 27 (1849)
= Phlegmariurus cancellatus var. minor Ching; Acta Bot. Yunnan. 4(2): 122 (1982)
= Urostachys cancellata (Spring) Herter ex Nessel; Bärlappgew. 140 (1939)
species :
Phlegmariurus capellae
(Herter) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Colombia (Antioquia, Caldas, Cauca, Cundinamarca, Tolima); Venezuela (Merida, Tachira); Ecuador; Peru
82 CLM VEN 83 ECU PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia capellae (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium capellae (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys capellae Herter; Revista Sudamer. Bot. 10: 114-115 (1953 publ. 1954)
species :
Phlegmariurus capillaris
(Sodiro) B.Øllg.; Rodriguesia 63(2): 480 (2012)
Distribution: Mexico (Chiapas); Guatemala; Nicaragua; Costa Rica; Panama; Colombia (Antioquia, Cauca, Huila, Norte de Santander); Venezuela (Amazonas, Bolívar, Yaracuy); Ecuador; Bolivia (La Paz); Brazil (Roraima)
79 MEX 80 COS GUA NIC PAN 82 CLM VEN 83 BOL ECU 84 BRA
Remarks: linifolia agg.
Synonyms: hidden
= Huperzia capillaris (Sodiro) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Huperzia underwoodiana (Maxon) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium capillare Sodiro; Crypt. Vasc. Quit. 90 (1883)
= Lycopodium guatemalense Maxon; Contr. U.S. Nat. Herb. 17(2): 177, t. 9a (1913)
= Lycopodium sarmentosum var. capillare (Sodiro) Sodiro; Crypt. Vasc. Quit. 573 (1893)
= Lycopodium sarmentosum var. rubescens Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 13 (1849)
= Lycopodium underwoodianum Maxon; Contr. U.S. Natl. Herb. 13: 41 (1909)
= Urostachys capillaris (Sodiro) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 54 (1949)
= Urostachys guatemalensis (Maxon) Herter; Repert. Spec. Nov. Regni Veg. 19: 165 (1923)
= Urostachys rubescens (Spring) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 79 (1949)
= Urostachys sarmentosus var. rubescens (Spring) Herter ex Nessel; Bärlappgew. 158 (1939)
= Urostachys underwoodianus (Maxon) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 87 (1949)
species :
Phlegmariurus carinatus
(Desv. ex Poir.) Ching; Acta Bot. Yunnan. 4: 120 (1982)
Distribution: China (Guangdong, Guangxi, Hebei, Yunnan); Japan; Taiwan; Nicobar Isl.; peninsular Thailand; ?Cambodia; ?Laos; Vietnam; peninsular Malaysia (widespread); +Singapore; Sumatra; Borneo (Sabah, Sawarak); Sulawesi; Philippines; New Guinea (Irian Jaya, Papua New Guinea, Louisiade Arch.: Misima Isl., Rossel Isl., Woodlark Isl.); Bismarck Arch.; Solomon Isl. (Guadalcanal, Santa Ysabel, Kolombangara, Makira, Malaita, New Georgia, Nggela, Rendova, Rennell, Tulagi); Australia (NE-Queensland); Santa Cruz Isl. (Vanikoro); New Caledonia; ?Vanuatu; Rotuma Isl.; Fiji (Viti Levu, Vanua Levu, Ovalau); Wallis & Futuna (Alofi, Futuna, ´Uvea, Wallis); American Samoa (Tutuila, Olosega, Ta´u); [Western] Samoa (Aleipata Isl., Savai´i, ´Upolu); Cook Isl. (Rarotonga)
36 CHI 38 JAP TAI 41 NCB THA VIE 42 BOR MLY NWG PHI SNG SUL SUM 43 BIS SOL 50 AUS 60 ASA FIJ NWC ROT SAM SCZ WAL 61 COO
Remarks: carinata agg.
Conservation Status: IUCN VU (Fraser-Jenkins 2012)
Synonyms: hidden
= Huperzia albescens (F.M.Bailey) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Huperzia carinata (Desv. ex Poir.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
= Huperzia laxa (C.Presl) T.Sen & U.Sen; Fern Gaz. 11(6): 420, f. 3, a-j (1978) (non (Desv.) Khokr.)
= Huperzia laxa (C.Presl) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
= Huperzia struthioloides (C.Presl) Rothm.; Feddes Repert. 54: 61 (1944)
= Huperzia vanikorensis (Copel.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium acrostachyum Hook. & Grev.; Icon. Filic. 2: t. 181 (1831)
= Lycopodium albescens F.M.Bailey; Queensl. Dept. Agric. Brisbane Bot. Bull. 14: 16 (1896)
= Lycopodium carinatum Desv. ex Poir.; Encycl. suppl. 3: 555 (1813 publ. 1814)
= Lycopodium carinatum var. minus Tagawa; Acta Phytotax. Geobot. 14: 91 (1951)
= Lycopodium laxum C.Presl; Reliq. Haenk. 1(1): 83 (1825)
= Lycopodium pendulum Roxb.; Calcutta J. Nat. Hist. 4: 472 (1844)
= Lycopodium struthioloides C.Presl; Reliq. Haenk. 1: 82 (1825)
= Lycopodium struthioloides var. margaritae Nessel; Bärlappgew. 171 (1939)
= Lycopodium urostachyum Wall.; Numer. List no. 117 (1829), nomen
= Lycopodium urostachyum Wall. ex Ridl.; J. Straits Branch Roy. Asiat. Soc. 80: 142 (1919)
= Lycopodium vanikorense Copel.; J. Arnold Arbor. 12: 49 (1931)
= Phlegmariurus laxus (C.Presl) Satou; Hikobia 12(3): 268 (1997)
= Selaginella struthioloides (C.Presl) Underw.; Bull. Torrey Bot. Club 25: 132 (1898)
= Urostachys albescens (F.M.Bailey) Herter; Revista Sudamer. Bot. 8: 86 (1949)
= Urostachys carinata (Desv. ex Poir.) Herter ex Nessel; Bärlappgew. 179 (1939), isonym
= Urostachys carinatus (Desv. ex Poir.) Herter; Bot. Arch. 3: 15 (1923)
= Urostachys struthioloides (C.Presl) Herter ex Nessel; Bärlappgew. 170 (1939)
= Urostachys vanikorensis (Copel.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 87 (1949)
species :
Phlegmariurus catacachiensis
(Nessel) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Colombia (Antioquia, Boyacas, Caldas, Cauca, Cundinamarca, Meta, Risaralda, Valle); Venezuela
82 CLM VEN
Remarks: saururus agg.
Synonyms: hidden
= Huperzia catacachiensis (Nessel) B.Øllg.; Opera Bot. 92: 169 (1987)
= Urostachys catacachiensis Nessel; Revista Sudamer. Bot. 6: 162, t. 10, f. 40 (1940)
= Urostachys catachiensis Herter ex Nessel; Bärlappgew. 99 (1939), no Latin descr.
species :
Phlegmariurus cavifolius
(C.Chr.) A.R.Field & Bostock; PhytoKeys 20: 37 (2013)
Distribution: Madagascar; Mauritius
29 MAU MDG
Remarks: gnidioides agg.
Synonyms: hidden
= Huperzia cavifolia (C.Chr.) Tardieu; Adansonia 10: 18 (1970)
= Huperzia smithiana (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium cavifolium C.Chr.; Dansk Bot. Ark. 7: 190, t. 76, f. 6-10 (1932)
= Urostachys cavifolius (C.Chr.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 55 (1949)
= Urostachys smithianus Nessel; Repert. Spec. Nov. Regni Veg. 36: 185, t. 174 (1934)
species :
Phlegmariurus chamaeleon
(Herter) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Costa Rica
80 COS
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia chamaeleon (Herter) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium chamaeleon (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys chamaeleon Herter; Amer. Fern J. 48: 82 (1958)
species :
Phlegmariurus changii
T.Y.Hsieh; Amer. Fern J. 102(4): 284 (2012)
Distribution: Taiwan
38 TAI
Conservation Status: very rare or extinct in the wild
Synonyms: hidden
= Huperzia changii (T.Y.Hsieh) Ralf Knapp; Index Ferns Fern Allies Taiwan 164 (2014)
species :
Phlegmariurus chiricanus
(Maxon) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Costa Rica (Volcan Turrialba); Panama (Volcan Chiriqui)
80 COS PAN
Remarks: saururus agg.
Synonyms: hidden
= Huperzia chiricana (Maxon) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium chiricanum Maxon; Contr. U.S. Natl. Herb. 17: 176 (1913)
= Urostachys chiricanus (Maxon) Herter ex Nessel; Bärlappgew. 45 (1939)
species :
Phlegmariurus christii
(Silveira) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia christii (Silveira) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium christianum Kümmerle; Mag. Bot. Lapok 26: 99 (1927 publ. 1928) (nom. confus. & superfl.)
= Lycopodium christii Silveira; Bol. Comm. Geogr. Geol. Est. Minas Gerais 2, 5: 117, t. 1 (1898)
= Urostachys christii (Silveira) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 379 (1927)
= Urostachys christii var. orgaosensis (Herter ex Nessel & Hoehne) Nessel; Fl. Bras. (Hoehne) 2, pt. 2: 26 (1955) [according to Ollgaard & Windisch 2019 ´orgaosanus´]
= Urostachys orgaosensis Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 380 (1927) [according to Ollgaard & Windisch 2019 ´orgaosanus´]
species :
Phlegmariurus ciliatospiculatus
B.Øllg.; Nordic J. Bot. 33(2): 192 (2014)
Distribution: Peru
83 PER
species :
Phlegmariurus ciliolatus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Venezuela
82 VEN
Synonyms: hidden
= Huperzia ciliolata B.Øllg.; Nordic J. Bot. 23(1): 31 (2004)
species :
Phlegmariurus cleefianus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Colombia (Huila, Cauca)
82 CLM
Synonyms: hidden
= Huperzia cleefiana B.Øllg.; Nordic J. Bot. 23(1): 34 (2004)
species :
Phlegmariurus cocuyensis
B.Øllg.; Phytotaxa 426(1): 64 (2019)
Distribution: Colombia (Arauca, Boyaca, Santander)
82 CLM
species :
Phlegmariurus colanensis
(B.Øllg.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Peru
83 PER
Synonyms: hidden
= Huperzia colanensis B.Øllg.; Fieldiana, Bot. n. s. 34: 31 (1994)
species :
Phlegmariurus columnaris
(B.Øllg.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Ecuador
83 ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia columnaris B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 45, f. 5A (1988)
species :
Phlegmariurus comans
(Herter ex Nessel) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia comans (Herter ex Nessel & Hoehne) B.Øllg. & P.G.Windisch; Bradea 5: 8 (1987)
= Lycopodium comans Christ; Schwacke, Pl. Nov. Mineiras 2: 40 (1900) (non Hook.f. (1844))
= Lycopodium verticillatum var. apertum Fée; Crypt. Vasc. Brésil 222 (1869)
= Urostachys comans Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 400 (1927) (nom. nov. for Lycopodium comans Christ (non Hook.f. (1844))
= Urostachys pollucis Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 76 (1949 (nom. superfl. for Urostachys comans Nessel))
species :
Phlegmariurus compactus
(Hook.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Ecuador
83 ECU
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia compacta (Hook.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1875)
= Huperzia jamesonii (Baker) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium compactum Hook.; Hooker´s Icon. Pl. 3: t. 244 (1839)
= Lycopodium jamesonii Baker; Handb. Fern-Allies 9 (1887)
= Urostachys compactus (Hook.) Herter ex Nessel; Bärlappgew. 102 (1939)
= Urostachys jamesonii (Baker) Herter ex Nessel; Bärlappgew. 100 (1939)
species :
Phlegmariurus copelandianus
(R.C.Y.Chou & Bartlett) A.R.Field; PhytoKeys 109: 64 (2018)
Distribution: Sumatra
42 SUM
Remarks: macgregorii agg.
Synonyms: hidden
= Huperzia copelandiana (R.C.Y.Chou & Bartlett) Holub; Folia Geobot. Phytotax. 26: 92 (1991)
= Lycopodium copelandianum R.C.Y.Chou & Bartlett; Bull. Torrey Bot. Club 74: 369 (1947)
= Lycopodium petiolatum Copel.; Univ. Calif. Publ. Bot. 14: 377 (1929) (non Herter (1909))
species :
Phlegmariurus coralium
(Spring) A.R.Field & Bostock; PhytoKeys 20: 37 (2013)
Distribution: peninsular Malaysia (Perak); Java
42 JAW MLY
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia coralium (Spring) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium coralium Spring; Miq., Pl. Jungh. 3: 273 (1854)
= Urostachys coralium (Spring) Herter ex Nessel; Bärlappgew. 234 (1939)
species :
Phlegmariurus crassus
(Humb. & Bonpl. ex Willd.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Mexico (Oaxaca); Guatemala; Costa Rica; Panama; Hispaniola (Dominican Republic); Colombia (Arauca, Boyacá, Caldas, Cauca, Chocó, Magdalena, Meta, Nariño, Putumayo, Risaralda, Santander, Tolima); Venezuela (Merida); Ecuador; Peru; Bolivia (La Paz)
79 MEX 80 COS GUA PAN 81 HIS 82 CLM VEN 83 BOL ECU PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia crassa (Humb. & Bonpl. ex Willd.) Rothm.; Feddes Repert. 54: 60 (1944)
= Huperzia erythraea (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1875)
= Huperzia pilgeriana (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia springii (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium bonae-voluntatis (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium crassum Humb. & Bonpl. ex Willd.; Sp. Pl. 5: 50 (1810)
= Lycopodium crassum var. springii (Nessel) Nessel; Bärlappgew. 76 (1939)
= Lycopodium erythraeum Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 7 (1849)
= Lycopodium nesselianum Duek & Lellinger; Amer. Fern J. 68(4): 120 (1978)
= Lycopodium saururus var. crassum (Humb. & Bonpl. ex Willd.) Sodiro; Crypt. Vasc. Quit. 90 (1883)
= Urostachys bonae-voluntatis Herter; Revista Sudamer. Bot. 10: 112. (1953 publ. 1954)
= Urostachys bonae-voluntatis var. minor Herter; Revista Sudamer. Bot. 10: 113 (1953)
= Urostachys crassus (Humb. & Bonpl. ex Willd.) Herter ex Nessel; Bärlappgew. 75 (1939)
= Urostachys crassus var. herteri Nessel; Repert. Spec. Nov. Regni Veg. 36(12-15): 180, t. 171 (1934)
= Urostachys erythraeus (Spring) Herter ex Nessel; Bärlappgew. 74 (1939)
= Urostachys kupperi Nessel; Repert. Spec. Nov. Regni Veg. 36: 180, pl. 176 (1934)
= Urostachys orionis Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 73 (1949) (nom. superfl.)
= Urostachys pilgerianus Herter ex Nessel; Revista Sudamer. Bot. 6(5-6): 161, t. 9, f. 32 (1940)
= Urostachys springii (Klotzsch & H.Karst.) Herter ex Nessel; Revista Sudamer. Bot. 6(5-6): 161, t. 9, f. 31 (1940)
variant :
var. gelidus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Huperzia crassa var. gelida B.Øllg.; Fl. Ecuador 33: 56 (1988)
variant :
var. manus-diaboli
(B.Øllg.) B.Øllg.; Phytotaxa 57: 13 (2012)
Distribution: Ecuador; Bolivia (La Paz)
83 BOL ECU
Synonyms: hidden
= Huperzia crassa var. manus-diaboli B.Øllg.; Fl. Ecuador 33: 56 (1988)
species :
Phlegmariurus creber
(Alderw.) A.R.Field & Bostock; PhytoKeys 20: 37 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: macgregorii agg.
Synonyms: hidden
= Huperzia crebra (Alderw.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium crebrum Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 4 (1917 (crebre))
= Urostachys creber (Alderw.) Herter ex Nessel; Bärlappgew. 181 (1939)
species :
Phlegmariurus crucis-australis
(Herter) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Colombia
82 CLM
Remarks: saururus agg.
Synonyms: hidden
= Lycopodium crucis-australis (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys crucis-australis Herter; Revista Sudamer. Bot. 10: 119 (1953 publ. 1954)
species :
Phlegmariurus cruentus
(Spring) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Colombia (Antioquia, Arauca, Boyacá, Caldas, Cauca, Cundinamarca, Meta, Nariño, Norte de Santander, Quindío, Risaralda, Santander); Venezuela (Tachira)
82 CLM VEN
Remarks: saururus agg.
Synonyms: hidden
= Huperzia cruenta (Spring) Rothm.; Feddes Repert. 54: 59 (1944)
= Lycopodium cruentum Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 34 (1849)
= Urostachys cruentus (Spring) Herter ex Nessel; Bärlappgew. 27 (1939)
= Urostachys erikssonii Nessel; Revista Sudamer. Bot. 6: 163 (1940)
variant :
var. exilis
B.Øllg.; Phytotaxa 426(1): 70 (2019)
Distribution: Colombia
82 CLM
variant :
var. illimitatus
B.Øllg.; Phytotaxa 426(1): 71 (2019)
Distribution: Colombia
82 CLM
variant :
var. innocentium
(Herter) B.Øllg.; Phytotaxa 426(1): 73 (2019)
Distribution: Colombia
82 CLM
Synonyms: hidden
= Lycopodium innocentium (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys innocentium Herter; Revista Sudamer. Bot. 10: 127 (1953 publ. 1954)
species :
Phlegmariurus cryptomerinus
(Maxim.) Satou; Hikobia 12(3): 268 (1997)
Distribution: Russian Far East (Kuril Isl.); China (Zhejiang); South Korea (incl. Cheju Isl.); S-Japan (incl. Yakushima Isl.); Taiwan; ?Philippines
31 RFE 36 CHI 38 JAP KOR NNS TAI
Remarks: hamiltonii agg. Reports from NE-India belong to H. bucahwangensis (Fraser-Jenkins et al. 2017)
Synonyms: hidden
= Huperzia cryptomerina (Maxim.) R.D.Dixit; J. Bombay Nat. Hist. Soc. 77(3): 541 (1981)
= Huperzia formosana Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Huperzia integrifolia (Matsuda) B.Øllg. ex Satou; Hikobia 12(3): 268 (1997)
= Huperzia taiwanensis (C.M.Kuo) C.M.Kuo; Taiwania 47(2): 170 (2002)
= Lycopodium cryptomerinum Maxim.; Bull. Acad. Imp. Sci. Saint-Petersbourg 15: 231 (1870)
= Lycopodium integrifolium (Matsuda) Matsuda & Nakai ex Nakai; Bot. Mag. (Tokyo) 28: 102 (1914) (non Goldie (1822))
= Lycopodium serratum var. integrifolium Matsuda; Bot. Mag. (Tokyo) 25: 124 (1911)
= Lycopodium taiwanense C.M.Kuo; Taiwania 30: 51 (1985)
= Phlegmariurus cryptomerinus (Maxim.) Ching; Acta Phytotax. Sin. 37(1): 51 (1999), isonym
= Phlegmariurus taiwanensis (C.M.Kuo) Li Bing Zhang; Fl. Reipubl. Popularis Sin. 6(3): 38 (2004) (fide Knapp 2011 ff.)
= Urostachys coreanus (Herter ex Nessel) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 57 (1949), non Hayata
= Urostachys cryptomerinus (Maxim.) Herter ex Nessel; Bärlappgew. 70 (1939)
= Urostachys cryptomerinus var. coreanus Herter ex Nessel; Bärlappgew. 72, t. 11, f. 8. (1939), Revista Sudamer. Bot. 6: 160 (1940) (nom. inval., non Hayata)
species :
Phlegmariurus cuernavacensis
(Underw. & F.E.Lloyd) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Mexico (Chiapas, Colima, Durango, Guerrero, Jalisco, Mexico State, Michoacan, Morelos, Oaxaca, Puebla, Sinaloa, Veracruz); Guatemala; El Salvador; Honduras
79 MEX 80 ELS GUA HON
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia cuernavacensis (Underw. & F.E.Lloyd) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium cuernavacense Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 110 (1906)
= Urostachys cuernavacensis (Underw. & F.E.Lloyd) Herter ex Nessel; Bärlappgew. 92 (1939)
species :
Phlegmariurus cumingii
(Nessel) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Colombia; Ecuador
82 CLM 83 ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia cumingii (Nessel) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium cumingii (Nessel) Rolleri; Rev. Mus. La Plata, n. s., Bot. 13(71): 92 (1980)
= Urostachys cumingii Nessel; Repert. Spec. Nov. Regni Veg. 36(12-15): 178, t. 170 (1934)
species :
Phlegmariurus cuneifolius
(Hieron.) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Costa Rica; Panama; Colombia (Antioquia, Arauca, Boyaca, Guajira, Norte de Santander, Santander); Venezuela (Merida, Tachira, Trujillo); N-Peru
80 COS PAN 82 CLM VEN 83 PER
Remarks: myrsinites agg.
Synonyms: hidden
= Huperzia cuneifolia (Hieron.) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium aqualupianum var. obtusifolium Spring; Mém. Acad. Roy. Soc. Bruxelles 24: 31 (1849)
= Lycopodium cuneifolium Hieron.; Bot. Jahrb. Syst. 34: 572 (1905)
= Urostachys cuneifolius (Hieron.) Herter ex Nessel; Bärlappgew. 252, t. 62, f. 14 (1939)
species :
Phlegmariurus cunninghamioides
(Hayata) Ching; Acta Bot. Yunnan. 4(2): 125 (1982)
Distribution: Taiwan; +Ryukyu Isl. (+Amami Oshima)
38 NNS TAI
Conservation Status: CR in Taiwan
Synonyms: hidden
= Huperzia cunninghamioides (Hayata) Holub; Folia Geobot. Phytotax. 20(1): 72 (1985)
= Lycopodium cunninghamioides Hayata; Icon. Pl. Formosan. 4: 131 (1914)
= Urostachys cunninghamioides (Hayata) Herter ex Nessel; Bärlappgew. 206 (1939)
species :
Phlegmariurus curiosus
(Herter) A.R.Field & Bostock; PhytoKeys 20: 38 (2013)
Distribution: Madagascar
29 MDG
Remarks: squarrosa agg.
Synonyms: hidden
= Huperzia curiosa (Herter) Tardieu; Adansonia, n.s., 10: 18 (1970)
= Urostachys curiosus Herter; Notul. Syst. (Paris) 15: 355 (1958)
species :
Phlegmariurus curvifolius
(Kunze) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Honduras; Nicaragua; Costa Rica; Panama; Colombia (Cauca, Choco, Antioquia); Ecuador; Peru
80 COS HON NIC PAN 82 CLM 83 ECU PER
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia curvifolia (Kunze) Holub; Folia Geobot. Phytotax. 20(1): 72 (1985)
= Huperzia curvifolia var. parvifolia (Wercklé ex Nessel) B.Øllg.; Novon 3(1): 71 (1993)
= Lycopodium cernuum var. tenerrimum Herter ex Nessel; Bärlappgew. 366 (1939)
= Lycopodium curvifolium Kunze; Linnaea 9: 5 (1834)
= Lycopodium tenue var. tenuissimum Spring; Mém. Acad. Roy. Soc. Bruxelles 24: 21 (1849)
= Lycopodium verticillatum var. parvifolium (Wercklé ex Nessel) Lellinger; Proc. Biol. Soc. Washington 89(61): 719 (1977)
= Urostachys curvifolius (Kunze) Herter ex Nessel; Bärlappgew. 129 (1939)
= Urostachys verticillatus var. parvifolius Wercklé ex Nessel; Bärlappgew. 123 (1939), et Revista Sudamer. Bot. 6: 163 (1940)
species :
Phlegmariurus dacrydioides
(Baker) A.R.Field & Bostock; PhytoKeys 20: 38 (2013)
Distribution: Guinea; Sierra Leone; Liberia; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Principe Isl.; São Tomé; D.R.Congo [Zaire]; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Comores
22 GUI LBR NGA SIE 23 BIO BUR CMN PRI SAO ZAI 24 ETH SUD 25 KEN TAN UGA 26 MLW MOZ ZIM 27 SAF 29 COM
Remarks: gnidioides agg.
Synonyms: hidden
= Huperzia dacrydioides (Baker) Pic.Serm.; Webbia 23: 162 (1968)
= Lycopodium dacrydioides Baker; Handb. Fern-Allies: 17 (1887)
= Urostachys dacrydioides (Baker) Herter ex Nessel; Bärlappgew. 188 (1939)
= Urostachys dacrydioides var. minor Herter ex Nessel; Bärlappgew. 189 (1939), nom. nud.
species :
Phlegmariurus dalhousieanus
(Spring) A.R.Field & Bostock; PhytoKeys 20: 38 (2013)
Distribution: peninsular Malaysia (Penang, Pahang, Melaka); Borneo (Sabah, Sarawak); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Kolombangara); Australia (NE-Queensland); Fiji (Viti Levu, Vanua Levu, Taveuni)
42 BOR MLY MOL NWG 43 SOL 50 AUS 60 FIJ
Remarks: dalhousiana agg.
Synonyms: hidden
= Huperzia caudifolia (Alderw.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Huperzia clarae (F.M.Bailey) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Huperzia dalhousieana (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
= Huperzia magnifica (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium caudifolium Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 1: 14 (1911)
= Lycopodium clarae F.M.Bailey; Queensl. Dept. Agric. Brisbane Bot. Bull. 21 (Bot. Bull. 7): 69 (1893)
= Lycopodium dalhousieanum Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 25 (1849))
= Lycopodium glaucum Ces.; Atti Reale Accad. Sci. Napoli 7: 35 (1876)
= Lycopodium ingens A.Gepp ex Nessel; Bärlappgew. 185 (1939), nom. nud.
= Lycopodium magnificum Brownlie; Beih. Nova Hedwigia 55: 22-23, t. 2e, f. 3-4 (1977)
= Urostachys caudifolius (Alderw.) Herter ex Nessel; Bärlappgew. 185 (1939)
= Urostachys clarae (F.M.Bailey) Herter ex Nessel; Bärlappgew. 185 (1939)
= Urostachys dalhousieanus (Spring) Herter ex Nessel; Bärlappgew. 185 (1939)
= Urostachys glaucus (Ces.) Herter; Revista Sudamer. Bot. 8: 86 (1949)
species :
Phlegmariurus darwinianus
(Herter ex Nessel) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Peru; Bolivia (Cochabamba, La Paz)
83 BOL PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia darwiniana (Herter ex Nessel) B.Øllg.; Fieldiana, Bot. n. s. 34: 33 (1994)
= Urostachys darwinianus Herter ex Nessel; Revista Sudamer. Bot. 6(5-6): 161 (1940)
species :
Phlegmariurus delbrueckii
(Herter) A.R.Field & Bostock; PhytoKeys 20: 39 (2013)
Distribution: Philippines (Mindanao); ?New Caledonia
42 PHI
Remarks: nummulariifolia agg.
Synonyms: hidden
= Huperzia delbrueckii (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium delbrueckii (Herter) Copel.; Univ. Calif. Publ. Bot. 12: 384 (1931)
= Urostachys delbrueckii Herter; Bot. Arch. 3: 19 (1923)
species :
Phlegmariurus deminuens
(Herter) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Minas Gerais, Rio de Janeiro)
84 BRA
Remarks: brongniartii agg.
Conservation Status: IUCN R
Synonyms: hidden
= Huperzia deminuens (Herter) B.Øllg.; Opera Bot. 92: 169 (1987)
= Huperzia luederwaldtii (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium deminuens Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 44 (1909)
= Urostachys deminuens (Herter) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 384 (1927)
= Urostachys luederwaldtii Nessel; Repert. Spec. Nov. Regni Veg. 36: 178, t. 170 (1934)
species :
Phlegmariurus dentatus
(Herter) Arana; Phytotaxa 252(4): 298 (2016)
Distribution: Azores (Sao Miguel Isl., Terceira, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl.)
21 AZO MDR
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia dentata (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Huperzia selago subsp. dentata (Herter) Valentine; Feddes Repert. 69: 44 (1964)
= Lycopodium dentatum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 41 (1909)
= Urostachys dentatus (Herter) Herter ex Nessel; Bärlappgew. 41 (1939)
species :
Phlegmariurus dianae
(Herter) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Colombia (Antioquia)
82 CLM
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia dianae (Herter) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium dianae (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys dianae Herter; Revista Sudamer. Bot. 10: 116 (1953 publ. 1954)
species :
Phlegmariurus dichaeoides
(Maxon) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Guatemala; Belize; Honduras; Nicaragua; Costa Rica; Panama; Colombia (Antioquia, Cauca, Chocó, Nariño, Santander, Valle); Ecuador
80 BLZ COS GUA HON NIC PAN 82 CLM 83 ECU
Remarks: aqualupiana agg.
Synonyms: hidden
= Huperzia dichaeoides (Maxon) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium dichaeoides Maxon; Proc. Biol. Soc. Washington 18: 231 (1905)
= Urostachys dichaeoides (Maxon) Herter ex Nessel; Bärlappgew. 252 (1939)
species :
Phlegmariurus dichotomus
(Jacq.) W.H.Wagner; Novon 3: 305 (1993)
Name: Hanging Club-Moss
Distribution: USA (Florida: Collier Co., Fakahatchee Strand); Mexico (Chiapas, Oaxaca, Puebla, San Luis Potosi, Tamaulipas, Veracruz, Mexico Distrito Federal, Guerrero); Revillagigedos Isl.; Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Virgin Isl. (Virgin Gorda); St. Eustatius; St. Kitts; Guadeloupe; Dominica; Martinique; St. Vincent; Grenada; Tobago; Trinidad; Colombia (Antioquia, Chocó, Cauca, Magdalena, Valle); Venezuela (Aragua, Barinas, Bolívar, Delta Amacuro, Venezuela Distrito Federal, Falcon, Merida, Miranda, Portuguesa, Sucre, Tachira, Yaracuy, Zulia); Guyana; Surinam; French Guiana; Ecuador; Galapagos; Bolivia (La Paz, Santa Cruz); Brazil (Acre, Amapá, Ceará, Mato Grosso, Pará, Rondônia, Roraima)
78 USA 79 MEX RGI 80 BLZ COS ELS GUA HON NIC PAN 81 CUB DOM GDL GRE HIS JAM MAR PUE STE STK STV TRT VIR 82 CLM FRG GUY SUR VEN 83 BOL ECU GAL 84 BRA
Remarks: dichotoma agg.
Synonyms: hidden
= Huperzia dichotoma (Jacq.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia graminea (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia lindeneri (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Huperzia mortonii (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia schlechtendalii (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium barbatum Christ; Bull. Herb. Boissier, sér. 2 5(3): 254 (1905)
= Lycopodium chamaepeuce Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 50 (1909)
= Lycopodium dichotomum Jacq.; Enum. stirp. Vindob. 3 (1762)
= Lycopodium dichotomum var. maximum Fée ex Nessel; Bärlappgew. 151 (1939)
= Lycopodium dichotomum var. pusillum Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 41 (1842)
= Lycopodium gramineum Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 19 (1849)
= Plananthus dichotomus (Jacq.) P.Beauv.; Prodr. Aethéogam. 111 (1805)
= Urostachys barbatus (Christ) Herter ex Nessel; Bärlappgew. 30 (1939)
= Urostachys chamaepeuce (Herter) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 164 (1923)
= Urostachys chamaepeuce var. maior Herter ex Nessel; Bärlappgew. 151 (1939)
= Urostachys chamaepeuce var. maximus (Fée) Herter ex Nessel; Bärlappgew. 151 (1939)
= Urostachys chamaepeuce var. urbanianus Herter; Repert. Spec. Nov. Regni Veg. 19: 164 (1923)
= Urostachys dichotomus (Jacq.) Herter; Beih. Bot. Centralbl. 39(2): 249 (1922)
= Urostachys dichotomus var. pusillus (Spring) Herter ex Nessel; Bärlappgew. 153 (1939)
= Urostachys gramineus (Spring) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 63 (1949)
= Urostachys lindeneri Herter ex Nessel; Revista Sudamer. Bot. 6(5-6): 164, t. 11, f. 50 (1940)
= Urostachys mortonii Herter ex F.Morton; Repert. Spec. Nov. Regni Veg. 28: 108 (1930)
= Urostachys schlechtendalii Nessel; Revista Sudamer. Bot. 6(5-6): 164, t. 11, f. 51 (1940)
= Urostachys urbanianus Herter ex Nessel; Bärlappgew. 151 (1939)
species :
Phlegmariurus dielsii
(Herter) A.R.Field & Bostock; PhytoKeys 20: 39 (2013)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Remarks: phyllantha agg.
Synonyms: hidden
= Huperzia dielsii (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium dielsii Herter; Bot. Jahrb. Syst. 54: 229 (1916)
= Urostachys dielsii (Herter) Herter ex Nessel; Bärlappgew. 213 (1939)
species :
Phlegmariurus divergens
(Alderw.) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Java
42 JAW
Synonyms: hidden
= Lycopodium divergens Alderw.; Mal. Fern-Allies 45 (1915)
species :
Phlegmariurus durus
(Pic.Serm.) A.R.Field & Bostock; PhytoKeys 20: 39 (2013)
Distribution: Kenya; Tanzania
25 KEN TAN
Synonyms: hidden
= Huperzia dacrydioides subsp. dura (Pic.Serm.) Verdc.; Fl. Trop. E. Africa, Lycopod. 6 (2005)
= Huperzia dura Pic.Serm.; Webbia 27: 392, fig. 1 (1972 publ. 1973)
species :
Phlegmariurus echinatus
(Spring) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Colombia (Cundinamarca, Cauca, Narino, Tolima, Antioquia, Valle); Venezuela (Aragua, Merida, Trujillo); Ecuador; Peru
82 CLM VEN 83 ECU PER
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia echinata (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
= Huperzia molongensis (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium echinatum Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 24 (1849)
= Lycopodium molongense Herter; Bot. Jahrb. Syst. 43: Beibl. 98.- 51 (1909)
= Phlegmariurus molongensis (Herter) B.Øllg.; Phytotaxa 57: 17 (2012)
= Urostachys echinatus (Spring) Herter ex Nessel; Bärlappgew. 178 (1939)
= Urostachys molongensis (Herter) Herter ex Nessel; Bärlappgew. 240 (1939)
species :
Phlegmariurus ellenbeckii
(Nessel) A.R.Field & Bostock; PhytoKeys 20: 39 (2013)
Distribution: Ethiopia
24 ETH
Remarks: ophioglossoides agg.
Synonyms: hidden
= Huperzia ellenbeckii (Nessel) Pic.Serm.; Webbia 23: 162 (1968)
= Urostachys ellenbeckii Nessel; Repert. Spec. Nov. Regni Veg. 36: 188, t. 175 (1934)
species :
Phlegmariurus elmeri
(Herter) A.R.Field & Bostock; PhytoKeys 20: 39 (2013)
Distribution: Palawan; Philippines (Benguet, Tayabas, Mindoro, Cotabato, Davao)
42 PHI PLW
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia elmeri (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Urostachys elmeri Herter; Bot. Arch. 3: 16 (1923)
species :
Phlegmariurus elongatus
(Lag ex Sw.) comb.ined.
Distribution: Ascension Isl.; St. Helena; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz, Tarija); Argentina (Jujuy, Buenos Aires, Catamarca, Cordoba, Salta, San Luis, Tucuman); Marion Isl.; Prince Edward Isl.; Ile Amsterdam; Iles Crozet; Iles Kerguélen; Gough Isl.; Tristan da Cunha (incl. Nightingale Isl., Inaccessible Isl.)
28 ASC STH 83 BOL PER 85 ARG 91 ASP CRZ KEG MPE TDC
Remarks: saururus agg. Frequently included in H. saururus s. lat. but kept separate by Roux (2009). South American and insular material may not be conspecific.
Conservation Status: IUCN I, Ex/E (Ascension), V (St. Helena), I (Tristan da Cunha)
Synonyms: hidden
= Huperzia insularis (Carmich.) Rothm.; Feddes Repert. 54: 60 (1944)
= Huperzia sanctae-barbarae (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Lycopodium elongatum Lag. ex Sw.; Syn. Fil. 175 (1806)
= Lycopodium insulare Carmich.; Trans. Linn. Soc. 12. 509 (1819)
= Lycopodium sanctae-barbarae Rolleri; Darwiniana 16: 129, f. 1(D-E), t. 11, t. 3(B) (1970)
= Lycopodium selago var. hessei Herter; Botanische Jahrbücher 43,1, 2, Beiblatt 98: 25 (1909), nom. nud.
= Urostachys elongatus (Lag. ex Sw.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 60 (1949)
= Urostachys insularis (Carmich.) Herter ex Nessel; Bärlappgew. 81 (1939)
= Urostachys saururus var. elongatus (Lag. ex Sw.) Herter ex Nessel; Bärlappgew. 80 (1939)
= Urostachys selago var. hessei Nessel; Bärlappgew. 40 (1939)
species :
Phlegmariurus engleri
(Hieron. & Herter) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Peru
83 PER
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia engleri (Hieron. & Herter) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium engleri Hieron. & Herter ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909)
= Urostachys engleri (Hieron. & Herter) Nessel; Bärlappgew. 94 (1939)
species :
Phlegmariurus eremorum
(Rolleri) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Colombia (Valle)
82 CLM
Remarks: brevifolia agg.
Conservation Status: IUCN R
Synonyms: hidden
= Huperzia eremorum (Rolleri) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium eremorum Rolleri; Hickenia 1(26): 141, f. 1 (1978)
species :
Phlegmariurus ericifolius
(C.Presl) B.Øllg.; Phytotaxa 57: 14 (2012)
Distribution: Ecuador; Peru; Bolivia (La Paz)
83 BOL ECU PER
Remarks: aqualupiana agg.
Synonyms: hidden
= Huperzia ericifolia (C.Presl) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium ericifolium C.Presl; Reliq. Haenk. 1: 77 (1825)
= Urostachys phlegmaria var. ericifolius (C.Presl) Nessel; Bärlappgew. 217 (1939)
species :
Phlegmariurus erythrocaulos
(Fée) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Bahia, Minas Gerais, Rio de Janeiro)
84 BRA
Remarks: myrsinites agg.
Synonyms: hidden
= Huperzia erythrocaulos (Fée) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium erythrocaulon Fée; Crypt. Vasc. Brésil 2: 95, t. 106, f. 2 (1872 publ. 1873)
= Urostachys erythrocaulos (Fée) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 421 (1927)
species :
Phlegmariurus espinosanus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Ecuador
83 ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia espinosana B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 47, f. 7A (1988) (esp)
species :
Phlegmariurus eversus
(Poir.) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Costa Rica; Panama; Colombia (Antioquia, Bolivar, Boyacá, Caldas, Cauca, Cundinamarca, Huila, Magdalena, Meta, Nariño, Norte de Santander, Putumayo, Quindío, Risaralda, Santander, Tolima, Valle); Venezuela (Merida, Trujillo); Ecuador; Peru; Bolivia (Cochabamba, La Paz)
80 COS PAN 82 CLM VEN 83 BOL ECU PER
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia ecuadorica (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Huperzia eversa (Poir.) B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 28 (1988)
= Lycopodium ecuadoricum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 48 (1909)
= Lycopodium eversum Poir.; Lam., Encycl. suppl. 3: 556 (1814)
= Lycopodium polycarpum (Sodiro) Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 105 (1906) (nom. illeg., non Lycopodium polycarpos Kunze (1834))
= Lycopodium reflexum var. polycarpum Sodiro; Crypt. Vasc. Quit. 90 (1883)
= Lycopodium reflexum Willd.; Sp. Pl. ed. IV, ed. 4 5: 52 (1810) (nom. illeg., non Lam. (1789) and (1795))
= Lycopodium rolandii-principis Herter ex Bonap.; Notes Pteridol. 1, 179 (1915)
= Urostachys dingesianus Nessel; Repert. Spec. Nov. Regni Veg. 39(8): 69, t. 193b (1935)
= Urostachys ecuadoricus (Herter) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 59 (1949)
= Urostachys reflexus var. polycarpus (Sodiro) Herter; Bärlappgew. 116 (1939)
= Urostachys ringshausenii Nessel; Repert. Spec. Nov. Regni Veg. 39: 68, tab. 193a (1935)
= Urostachys rolandii-principis Herter ex Nessel; Bärlappgew. 11 7 (1939)
species :
Phlegmariurus fargesii
(Herter) Ching; Acta Bot. Yunnan. 4(2): 120-121 (1982)
Distribution: China (Chongqing, Guangxi, Sichuan, Yunnan); Japan; Taiwan
36 CHI 38 JAP TAI
Remarks: cancellata agg.
Synonyms: hidden
= Huperzia fargesii (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Huperzia sieboldii var. christenseniana (Christ & Herter) Nakaike; New Fl. Jap. Pterid. 841 (1992)
= Lycopodium christensenianum Christ & Herter ex Herter; Hedwigia 49: 89 (1909)
= Lycopodium fargesii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 48 (1909)
= Lycopodium fargesii var. gracile Tagawa; Acta Phytotax. Geobot. 14(3): 92 (1951)
= Lycopodium fauriei Rosenst.; Hedwigia 56(5): 348 (1915)
= Lycopodium quasiprimaevum Koidz.; Acta Phytotax. Geobot. 1: 32 (1932)
= Lycopodium tereticaule Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 411 (1911) (non Poir. (1814))
= Phlegmariurus christensenianus (Christ & Herter) Satou; Hikobia 12(3): 268 (1997)
= Urostachys christensenianus (Christ & Herter) Herter ex Nessel; Bärlappgew. 139 (1939)
= Urostachys fargesii (Herter) Herter ex Nessel; Bärlappgew. 137 (1939)
= Urostachys fauriei (Rosenst.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 61 (1949)
= Urostachys tereticaulis (Hayata) Herter ex Nessel; Bärlappgew. 137, f. 32 (1939)
species :
Phlegmariurus filicaulos
(Copel.) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015), as ´filicaulon´
Distribution: New Guinea (Papua New Guinea); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, Makira, Rendova)
42 NWG 43 SOL
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia copelandii (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium filicaulon Copel.; Philipp. J. Sci. 60: 100, t. 2 (1936) (non Lycopodium filicaule Hook.f. (1844))
= Urostachys copelandii Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 57 (1949)
species :
Phlegmariurus filiformis
(Sw.) W.H.Wagner; Contr. Univ. Michigan Herb. 20: 242 (1995)
Distribution: peninsular Malaysia (Pahang, Selangor); Borneo (Sabah, Sarawak); Sulawesi; Philippines; Java; Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Australia (NE-Queensland); New Caledonia; American Samoa (Olosega, Ta´u); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
42 BOR JAW MLY MOL NWG PHI SUL 50 AUS 60 ASA NWC 63 HAW
Remarks: verticillata agg. Contains Malesian and Pacific populations of "P. verticillatus" (true verticillatus is restricted to the Western Indian Ocean). Since P. filiformis type is from Hawaii and it predates H. ignambiensis, filiformis should take priority.
Synonyms: hidden
= Huperzia filiformis (Sw.) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Huperzia ignambiensis (Compton) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Huperzia polytrichoides (Kaulf.) Trevis.; Bärlappgew. 129 (1939)
= Huperzia verticillata var. filiformis (Sw.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium acerosum var. filiforme (Sw.) Spring; Mart., Fl. Bras. 1(2): 111 (1840)
= Lycopodium filiforme Sw.; J. Bot. (Schrader) 1800(2): 114 (1801) (Syn. Fil. 174, 398, t. 4, f. 3 (1806))
= Lycopodium ignambiense Compton; J. Linn. Soc., Bot. 45: 458 (1922)
= Lycopodium polytrichoides Kaulf.; Enum. Filic. 6 (1824)
= Lycopodium verticillatum var. filiforme (Sw.) Spring; Mém. Acad. Roy. Sci. Bruxelles 12: 111 (1840)
= Phlegmariurus ignambiensis (Compton) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
= Urostachys filiformis (Sw.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 61 (1949)
= Urostachys polytrichoides (Kaulf.) Herter ex Nessel; Bärlappgew. 129 (1939)
= Urostachys verticillatus var. filiformis (Sw.) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 404 (1927)
species :
Phlegmariurus firmus
(Mett.) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Colombia (Antioquia, Caldas, Cauca, Chocó, Cundinamarca, Huila, Meta, Nariño, Putumayo, Quindío, Risaralda, Tolima, Valle); Venezuela (Tachira); Ecuador
82 CLM VEN 83 ECU
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia coriacea (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Huperzia firma (Mett.) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium coriaceum Rolleri; Rev. Mus. La Plata (N. S.) Bot. 13(71): 85, t. 5 (B, D) (1981)
= Lycopodium firmum Mett.; Ann. Sci. Nat., Bot., sér. V, 3: 307 (1865)
= Urostachys firmus (Mett.) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 391 (1927)
species :
Phlegmariurus flagellaceus
(Kuhn) A.R.Field & Bostock; PhytoKeys 20: 40 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: varia agg.
Synonyms: hidden
= Huperzia flagellacea (Kuhn) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium flagellaceum Kuhn; Forschungsr. Gazelle 4: 15 (1889)
= Urostachys flagellaceus (Kuhn) Herter ex Nessel; Bärlappgew. 237 (1939)
species :
Phlegmariurus flexibilis
(Fée) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Bahia, Ceará, Espírito Santo, Minas Gerais, Pará, Pernambuco, Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Huperzia flexibilis (Fée) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium flexibile Fée; Crypt. Vasc. Brésil 2: 94p, t. 105, f. 3 (1872 publ. 1873)
= Lycopodium linifolium var. sanguineum Spring; Bull. Acad. Roy. Soc. Bruxelles 15: 31 (1842)
= Lycopodium linifolium var. subaristatum Christ; Bull. Soc. bot. Genève 1(5): 236 (1909)
= Urostachys flexibilis (Fée) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 61 (1949)
= Urostachys linifolius var. flexibilis (Fée) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 416 (1927)
= Urostachys linifolius var. sanguineus (Spring) Nessel; Arch. Bot. São Paulo 1: 316 (1927)
= Urostachys schwendeneri var. subaristatus (Christ) Nessel; Bärlappgew. 167 (1939)
species :
Phlegmariurus foliaceus
(Maxon) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Costa Rica; Panama
80 COS PAN
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia foliacea (Maxon) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium foliaceum Maxon; Smithsonian Misc. Collect. 56(29): 1 (1912)
= Urostachys foliaceus (Maxon) Herter ex Nessel; Bärlappgew. 68 (1939)
species :
Phlegmariurus foliosus
(Copel.) A.R.Field & Bostock; PhytoKeys 20: 40 (2013)
Distribution: New Guinea (Papua New Guinea); Fiji (Viti Levu, Vanua Levu, Ovalau, Taveuni)
42 NWG 60 FIJ
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia foliosa (Copel.) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium foliosum Copel.; Bull. Bernice P. Bishop Mus. 59: 7, t. 1 (1929)
= Urostachys foliosus (Copel.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 61 (1949)
species :
Phlegmariurus fontinaloides
(Spring) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Remarks: quadrifariata agg.
Synonyms: hidden
= Huperzia fontinaloides (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia gehrtii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium fontinaloides Spring; Mart., Fl. Bras. 1(2): 112, pl. 5, f. 2 (1840)
= Lycopodium fontinaloides var. brasiliense Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 49 (1842)
= Lycopodium serpyllifolium Fée; Crypt. Vasc. Brésil 1: 222, t. 73, f. 3 (1869)
= Urostachys fontinaloides (Spring) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 401 (1927)
= Urostachys fontinaloides var. brasiliensis (Spring) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 402 (1927)
= Urostachys fontinaloides var. filiformis Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 404 (1927)
= Urostachys fontinaloides var. serpyllifolius (Fée) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 404 (1927)
= Urostachys gehrtii Nessel; Repert. Spec. Nov. Regni Veg. 36: 189, t. 177 (1934)
= Urostachys serpyllifolius (Fée) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 81 (1949)
species :
Phlegmariurus fordii
(Baker) Ching; Acta Bot. Yunnan. 4: 126 (1982) [Acta Phytotax. Sin. 20(4): 447 (1982])
Distribution: China (Guangdong, Guangxi, Guizhou, Fujian, Hainan, S-Hunan, S-Jiangxi, Yunnan, Zhejiang); Japan; Ryukyu Isl.; Taiwan; Myanmar (Chin); Vietnam; Philippines
36 CHI 38 JAP NNS TAI 41 MYA VIE 42 PHI
Remarks: hamiltonii agg. Inclusion of P. juniperistachyus uncertain. Not in India.
Synonyms: hidden
= Huperzia fordii (Baker) Holub; Folia Geobot. Phytotax. 20(1): 72 (1985)
= Huperzia fordii (Baker) R.D.Dixit; Census Indian Pteridophytes: 7 (1984)
= Huperzia juniperistachya (Hayata) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium fordii Baker; Handb. Fern-Allies 17 (1887)
= Lycopodium juniperistachyum Hayata; Icon. Pl. Formosan. 4: 132 (1914)
= Lycopodium poissonii Herter; Bot. Jahrb. Syst. 43: Beibl. 9851 (1909)
= Lycopodium subdistichum Makino; Bot. Mag. (Tokyo) 12: 37 (1898)
= Phlegmariurus juniperistachyus (Hayata) A.R.Field & Bostock; PhytoKeys 20: 42 (2013)
= Phlegmariurus longyangensis C.Y.Ma; Bull. Bot. Res., Harbin 10(3): 58 (1990)
= Phlegmariurus nanus C.Y.Ma; Bull. Bot. Res., Harbin 10(3): 58 (1990)
= Phlegmariurus yandongensis Ching & C.F.Zhang; Bull. Bot. Res. North-East. Forest. Inst. 3(3): 2 (1983)
= Urostachys fordii (Baker) Herter ex Nessel; Bärlappgew. 69 (1939)
= Urostachys juniperistachyus (Hayata) Herter ex Nessel; Bärlappgew. 70 (1939)
= Urostachys poissonii (Herter) Herter ex Nessel; Bärlappgew. 196 (1939)
= Urostachys subdistichus (Makino) Herter ex Nessel; Bärlappgew. 196 (1939)
species :
Phlegmariurus friburgensis
(Herter ex Nessel & Hoehne) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro); Paraguay (Paraguari)
84 BRA 85 PAR
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia friburgensis (Herter ex Nessel & Hoehne) B.Øllg. & P.G.Windisch; Bradea 5: 10 (1987)
= Lycopodium friburgense (Herter ex Nessel & Hoehne) Rolleri; Revista Mus. La Plata, Secc. Bot. 13: 191 (1984)
= Urostachys friburgensis Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 391 (1927)
= Urostachys reflexus var. friburgensis (Herter ex Nessel & Hoehne) Nessel; Bärlappgew. 114 (1939)
species :
Phlegmariurus funiculus
(Herter) comb.ined.
Distribution: D.R.Congo [Zaire]
23 ZAI
Synonyms: hidden
= Huperzia funicula (Herter) J.P.Roux; Strelitzia 23: 13 (2009)
= Urostachys funiculus Herter; Revista Sudamer. Bot. 8: 22 (1949)
species :
Phlegmariurus funiformis
(Cham. ex Spring) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Trinidad; Colombia (Antioquia, Cauca, Narino, Valle); Venezuela (Amazonas, Bolívar, Venezuela Distrito Federal, Nueva Esparta); Guyana; Ecuador; Peru
80 COS PAN 81 CUB DOM GDL HIS JAM MAR PUE STL STV TRT 82 CLM GUY VEN 83 ECU PER
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia funiformis (Cham. ex Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium funiforme Cham. ex Spring; Bull. Acad. Roy. Soc. Bruxelles 8: 516 (1841) (non Brongniart, Hist. Vég. fossil. 2: 10 (1837)) (nom. inval.)
= Urostachys funiformis (Cham. ex Spring) Herter; Symb. Ant. 9: 387 (1925)
species :
Phlegmariurus gagnepainianus
(Herter) A.R.Field & Bostock; PhytoKeys 20: 40 (2013)
Distribution: Madagascar
29 MDG
Remarks: pecten agg.
Synonyms: hidden
= Huperzia gagnepainiana (Herter) Tardieu; Adansonia 10: 18 (1970)
= Urostachys gagnepainianus Herter; Revista Sudamer. Bot. 8: 23 (1949)
species :
Phlegmariurus galapagensis
(O.J.Hamann) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Galapagos
83 GAL
Remarks: verticillata agg.
Conservation Status: IUCN R
Synonyms: hidden
= Lycopodium setaceum subsp. galapagense O.J.Hamann; Bot. Notis. 127(2): 253 (1974)
= Huperzia galapagensis (O.J.Hamann) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
species :
Phlegmariurus giganteus
(Herter) A.R.Field & Bostock; PhytoKeys 20: 40 (2013)
Distribution: Philippines (Davao, Cotabato, Misamis Occidental); New Guinea (Papua New Guinea)
42 NWG PHI
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia gigantea (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Urostachys verticillatus var. giganteus Herter; Bot. Arch. 3: 13 (1923)
= Urostachys giganteus (Herter) Herter ex Nessel; Bärlappgew. 124 (1939)
species :
Phlegmariurus gnidioides
(L.f.) A.R.Field & Bostock; PhytoKeys 20: 40 (2013)
Distribution: D.R.Congo [Zaire]; Rwanda; Burundi; Uganda; Kenya; Tanzania; Malawi; E-Zimbabwe; Mozambique; South Africa (Mpulamanga, Limpopo, E-Free State, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Lesotho; Eswatini [Swaziland]; Madagascar; Comores; Mauritius; La Réunion; Seychelles
23 BUR RWA ZAI 25 KEN TAN UGA 26 MLW MOZ ZIM 27 LES SAF SWZ 29 COM MAU MDG REU SEY
Remarks: gnidioides agg.
Synonyms: hidden
= Huperzia gnidioides (L.f.) Trevis.; Atti Soc. Critt. Ital. 17: 247 (1875)
= Huperzia helmii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lepidotis funiculosa (Lam.) P.Beauv.; Prodr. Aethéogam. 108 (1805)
= Lycopodium ambiguum Schrad.; Gött. Gel. Anz. 1818: 920 (1818)
= Lycopodium flagelliforme Schrad.; Gött. Gel. Anz. 1818: 920 (1818)
= Lycopodium funiculosum Lam.; Encycl. 3: 689 (1789)
= Lycopodium gnidioides L.fil.; Suppl. Pl. 448 (1782)
= Lycopodium gnidioides var. pinifolium (Kaulf.) Pappe & Rawson; Synopsis filicum Africae australis: 49 (1858)
= Lycopodium gnidioides var. strictum (Baker) C.Chr.; Dansk Bot. Ark. 7: 190 (1932)
= Lycopodium pinifolium Kaulf.; Enum. Filic. 7 (1824) (non Blume (1828))
= Plananthus gnidioides (L.f.) P.Beauv.; Prodr. Aethéogam. 110 (1805)
= Urostachys gnidioides (L.f.) Herter ex Nessel; Bärlappgew. 187 (1939)
= Urostachys helmii Nessel; Repert. Spec. Nov. Regni Veg. 36: 186p, t. 174 (1934)
species :
Phlegmariurus goebelii
(Nessel) A.R.Field & Bostock; PhytoKeys 20: 40 (2013)
Distribution: Sumatra; Borneo (Sabah)
42 BOR SUM
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia goebelii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Huperzia heroldii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Urostachys goebelii Nessel; Repert. Spec. Nov. Regni Veg. 36: 188, t. 175 (1934)
= Urostachys heroldii Nessel; Revista Sudamer. Bot. 6: 167, t. 14, f. 69 (1940)
species :
Phlegmariurus gracilis
A.Rojas & R.Calderón; Acta Botanica Malacitana 42: 67-70 (2017)
Distribution: Costa Rica
80 COS
Remarks: reflexus agg.
species :
Phlegmariurus guangdongensis
Ching; Acta Bot. Yunnan. 4(2): 123 (1982)
Distribution: China (Guangdong, Guangxi, Hainan)
36 CHI
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia guangdongensis (Ching) Holub; Folia Geobot. Phytotax. 20(1): 73 (1985)
species :
Phlegmariurus gunturensis
(Alderw.) A.R.Field & Bostock; PhytoKeys 20: 41 (2013)
Distribution: Java
42 JAW
Remarks: carinata agg.
Synonyms: hidden
= Huperzia gunturensis (Alderw.) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium gunturense Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 1: 14 (1911)
= Urostachys gunturensis (Alderw.) Herter ex Nessel; Bärlappgew. 183 (1939)
species :
Phlegmariurus haeckelii
(Herter) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Society Isl. (Tahiti, Raiatea)
61 SCI
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia haeckelii (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium haeckelii Herter; Repert. Spec. Nov. Regni Veg. 5: 22 (1908)
= Urostachys haeckelii (Herter) Herter ex Nessel; Bärlappgew. 138 (1939)
species :
Phlegmariurus haitensis
(Herter) comb.ined.
Distribution: Hispaniola (Haiti)
81 HIS
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia haitensis (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Urostachys haitensis Herter; Repert. Spec. Nov. Regni Veg. 19: 161 (1923)
species :
Phlegmariurus hamiltonii
(Spreng.) Á.Löve & D.Löve; Taxon 26(2-3): 324 (1977)
Distribution: China (Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Sichuan, Yunnan); Tibet; Volcano Isl. (Kita-Iwojima, Minami-Iwojima); Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Chattisgarh, Goa, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Odisha, Sikkim, Tamil Nadu, Uttarakhand, West Bengal, Mizoram, Nagaland); Sri Lanka; Myanmar (Kachin, Chin, etc.); Thailand (widespread); Vietnam
36 CHI TIB 38 KZN 40 BHU IND NEP SRL 41 MYA THA VIE
Remarks: hamiltonii agg. Inclusion of H. aloifolia and H. petiolata following Fraser-Jenkins et al. (2015, 2017)
Synonyms: hidden
= Huperzia aloifolia (Wall. ex Grev. & Hook.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia hamiltonii (Spreng.) T.Sen & U.Sen; Fern Gaz. 11(6): 419-420, f. 2, bb-hh (1978), isonym
= Huperzia hamiltonii (Spreng.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia petiolata (C.B.Clarke) R.D.Dixit; J. Bombay Nat. Hist. Soc. 77(3): 541 (1981)
= Lycopodium aloifolium Wall. ex Hook. & Grev.; Icon. Filic. 2: t. 233 (1831 publ. 1832)
= Lycopodium empetrifolium Dalzell; Hooker´s J. Bot. Kew Gard. Misc. 4: 113 (1852)
= Lycopodium hamiltonii Spreng.; Syst. Veg. 5: 429 (index) (1828)
= Lycopodium hamiltonii var. petiolatum C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 590 (1880)
= Lycopodium obtusifolium Buch.-Ham. ex D.Don; Prodr. Fl. Nepal 18-19 (1825) (nom. illeg., non Lycopodium obtusifolium Sw. (1806))
= Lycopodium petiolatum (C.B.Clarke) Christ; Bull. Acad. Int. Geogr. Bot. 11: 271 (1902)
= Lycopodium petiolatum (C.B.Clarke) Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 36 (1909), isonym
= Phlegmariurus aloifolius (Wall. ex Hook. & Grev.) A.R.Field & Bostock; PhytoKeys 20: 36 (2013)
= Phlegmariurus hamiltonii var. petiolatus (C.B.Clarke) Ching; Acta Bot. Yunnan. 4(2): 126-127 (1982)
= Phlegmariurus petiolatus (C.B.Clarke) H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 37(1): 45-47 (1999)
= Urostachys aloifolius (Wall. ex Grev. & Hook.) Nessel; Bärlappgew. 70 (1939)
= Urostachys hamiltonii (Spreng.) Herter ex Nessel; Bärlappgew. 68 (1939)
= Urostachys hamiltonii var. petiolatus (C.B.Clarke) Herter ex Nessel; Bärlappgew. 68 (1939)
= Urostachys petiolatus (C.B.Clarke) Herter ex Nessel; Bärlappgew. 196 (1939)
species :
Phlegmariurus harmsii
(Nessel) A.R.Field & Bostock; PhytoKeys 20: 41 (2013)
Distribution: Solomon Isl. (Bougainville, Guadalcanal, Santa Ysabel, Kolombangara, Makira, Malaita, Mono, Nggatokae, Vella Lavella, New Georgia, Rendova, Rennell, Shortland, Vangunu); Palau (Babeldaob, Koror, Malakal)
43 SOL 62 PLU
Remarks: some more of the distribution records of P. phlegmaria for the Pacific might belong here
Synonyms: hidden
= Huperzia harmsii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Urostachys harmsii Herter ex Nessel; Bärlappgew. 223 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 166 (1940)
species :
Phlegmariurus hartwegianus
(Spring) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Mexico (Chiapas); Guatemala; Honduras; Colombia (Antioquia, Boyacá, Cauca, Cesar, Cundinamarca, Magdalena, Nariño, Quindío, Risaralda, Guajira, Santander); Venezuela (Aragua, Venezuela Distrito Federal, Lara, Merida, Miranda, Sucre); Ecuador; N-Peru
79 MEX 80 GUA HON 82 CLM VEN 83 ECU PER
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia caracasica (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Huperzia hartwegiana (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium caracasicum Herter; Hedwigia 49(2-3): 88, t. 3A (1909)
= Lycopodium funckii Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 44 (1909)
= Lycopodium hartwegianum Spring; Mém. Acad. Roy. Sci. Belg. 24: 14 (1850)
= Lycopodium hartwegianum var. viridissimum Sodiro; Crypt. Vasc. Quit. 575 (1893)
= Urostachys caracasicus (Herter) Herter ex Nessel; Bärlappgew. 90 (1939)
= Urostachys funckii (Herter) Herter ex Nessel; Bärlappgew. 170, t. 41 (1939)
= Urostachys hartwegianus (Spring) Herter ex Nessel; Bärlappgew. 90 (1939)
= Urostachys hartwegianus var. viridissimus (Sodiro) Herter ex Nessel; Bärlappgew. 90 (1939)
= Urostachys maxonii Herter ex Nessel; Revista Sudamer. Bot. 6(5-6): 165 (1940)
species :
Phlegmariurus hastatus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Ecuador
83 ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia hastata B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 39, f. 6A (1988)
species :
Phlegmariurus hellwigii
(Warb.) A.R.Field & Bostock; PhytoKeys 20: 41 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia hellwigii (Warb.) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium hellwigii Warb.; Monsunia 1: 97 (1900)
= Urostachys hellwigii (Warb.) Herter ex Nessel; Bärlappgew. 230 (1939)
species :
Phlegmariurus hemleri
(Nessel) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Minas Gerais, Rio de Janeiro)
84 BRA
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia hemleri (Nessel) B.Øllg.; Opera Bot. 92: 169 (1987)
= Urostachys hemleri Nessel; Arch. Bot. São Paulo 1: 389, t. 9 (1927)
species :
Phlegmariurus henryi
(Baker) Ching; Acta Bot. Yunnan. 4(2): 125-126 (1982)
Distribution: China (Guangxi, Yunnan); Vietnam
36 CHI 41 VIE
Remarks: hamiltonii agg.
Synonyms: hidden
= Huperzia henryi (Baker) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium henryi Baker; Bull. Misc. Inf. Kew 1906(1): 15 (1906)
= Urostachys henryi (Baker) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 64 (1949)
species :
Phlegmariurus heterocarpos
(Fée) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Argentina (Misiones)
84 BRA 85 ARG
Remarks: heterocarpon agg.
Synonyms: hidden
= Huperzia heterocarpon (Fée) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Huperzia longearistata (Christ) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium dichotomum var. spegazzinii Nessel & Hoehne; Arch. Bot. São Paulo 1: 412 (1927)
= Lycopodium heterocarpon Fée; Crypt. Vasc. Brésil 2: 93 (1872 publ. 1873)
= Lycopodium longearistatum Christ; Schwacke, Pl. Nov. Mineiras 2: 40 (1900)
= Urostachys dichotomus var. spegazzinii (Herter ex Nessel & Hoehne) Nessel; Fl. Bras. (Hoehne) 2(2): 74 (1955)
= Urostachys heterocarpos (Fée) Herter ex Nessel & Hoehne; Repert. Spec. Nov. Regni Veg. 19: 164 (1923)
= Urostachys heterocarpos f. brevibracteatus (Rosenst.) Nessel; Bärlappgew. 156 (1939)
= Urostachys heterocarpos var. longearistatus (Christ) Herter ex Nessel; Bärlappgew. 156 (1939)
= Urostachys longearistatus (Christ) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 414 (1927)
= Urostachys major (Nessel) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 69 (1949)
= Urostachys spegazzinii Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 412 (1927)
species :
Phlegmariurus heteroclitus
(Desv. ex Poir.) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Ecuador; Peru
83 ECU PER
Remarks: myrsinites agg.
Synonyms: hidden
= Lycopodium heteroclitum Desv. ex Poir.; Encycl. suppl. 3: 554 (1813 (1814))
= Huperzia heteroclita (Desv. ex Poir.) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
species :
Phlegmariurus hexastichus
(B.Øllg. & P.G.Windisch) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Remarks: quadrifariata agg.
Synonyms: hidden
= Huperzia hexasticha B.Øllg. & P.G.Windisch; Bradea 5: 11, fig. 2 (1987)
= Lycopodium fontinaloides var. adpressa Fée; Crypt. Vasc. Brésil 2: 295 (1872 publ. 1873)
= Urostachys fontinaloides var. adpressa (Fée) Nessel & Hoehne; Arch. Bot. São Paulo 1: 404 (1927)
species :
Phlegmariurus hildebrandtii
(Herter) A.R.Field & Bauret; Molec. Phylogen. Evol. 127: 499 (2018)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Lycopodium hildebrandtii Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 40 (1909)
= Huperzia hildebrandtii (Herter) Tardieu; Adansonia ser. 2 10: 18 (1970)
= Urostachys hildebrandtii (Herter) Herter ex Nessel; Bärlappgew. 28, t. 4, f. 3 (1939)
species :
Phlegmariurus hippurideus
(Christ) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Mexico (Puebla, Oaxaca, Chiapas); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Colombia (Antioquia, Arauca, Boyacá, Caldas, Cauca, Cesar, Chocó, Cundinamarca, Huila, Magdalena, Meta, Nariño, Norte de Santander, Putumayo, Risaralda, Santander, Tolima, Valle); Venezuela (Amazonas, Bolívar, Lara, Merida, Tachira, Trujillo); Ecuador; Peru; Bolivia (Cochabamba, La Paz); Brazil (Roraima, ?Acre)
79 MEX 80 COS ELS GUA HON NIC PAN 81 CUB HIS JAM 82 CLM VEN 83 BOL ECU PER 84 BRA
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia boliviana (Rosenst.) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21(100): 155 (1988)
= Huperzia hippuridea (Christ) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Huperzia lechleri (Hieron.) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Huperzia montana (Underw. & F.E.Lloyd) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium bolivianum Rosenst.; Repert. Spec. Nov. Regni Veg. 21: 95 (1924)
= Lycopodium bolivianum var. teretiuscula Rosenst.; Repert. Spec. Nov. Regni Veg. 11: 60 (1912)
= Lycopodium hippurideum Christ; Primit. Fl. Costar. 3(1): 56 (1901)
= Lycopodium hippurideum var. montanum (Underw. & F.E.Lloyd) Proctor; Amer. Fern J. 72: 107 (1982)
= Lycopodium lechleri Hieron.; Bot. Jahrb. Syst. 34: 571 (1905)
= Lycopodium montanum Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 107 (1906)
= Lycopodium poseidonis (Herter) C.V.Morton; Amer. Fern J. 54: 42 (1964)
= Urostachys bolivianus (Rosenst.) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 396 (1927)
= Urostachys bolivianus var. ringshausenii (Nessel) Nessel; Bärlappgew. 88 (1939)
= Urostachys bolivianus var. teretiuscula (Rosenst.) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 396 (1927)
= Urostachys hippurideus (Christ ex Pittier) Herter ex Nessel; Bärlappgew. 88 (1939)
= Urostachys hippurideus var. montanus (Underw. & F.E.Lloyd) Herter ex Nessel; Bärlappgew. 88 (1939)
= Urostachys lechleri (Hieron.) Herter ex Nessel; Bärlappgew. 85 (1939)
= Urostachys lechleri var. lehmannii Bogemann ex Nessel; Bärlappgew. 86 (1939) (nom. inval.), et Revista Sudamer. Bot. 6: 161, t. 8, f. 35 (1940)
= Urostachys lehmannii (Nessel) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 67 (1949)
= Urostachys montanus (Underw. & F.E.Lloyd) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 70 (1949)
= Urostachys poseidonis Herter; Revista Sudamer. Bot. 10: 122-123 (1953)
species :
Phlegmariurus hippuris
(Desv. ex Poir.) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Borneo (Sabah); ?Philippines; Java; Moluccas (Seram); New Guinea (Papua New Guinea); ?[Western] Samoa
42 BOR JAW MOL NWG
Remarks: hippuris agg.
Synonyms: hidden
= Huperzia hippuris (Desv. ex Poir.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium blumeanum de Vriese; Tijdschr. Nat. Gesch. 3: 338 (1844)
= Lycopodium hippuris Desv. ex Poir.; Encycl. suppl. 3: 559 (1813 (1814))
= Urostachys hippuris (Poir.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 65 (1949)
species :
Phlegmariurus hoffmannii
(Maxon) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Costa Rica; Panama
80 COS PAN
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia hoffmannii (Maxon) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Lycopodium hoffmannii Maxon; Contr. U.S. Natl. Herb. 17(4): 423 (1914)
= Urostachys hoffmannii (Maxon) Herter ex Nessel; Bärlappgew. 99 (1939)
species :
Phlegmariurus hohenackeri
(Herter) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Colombia (Antioquia, Caldas, Cundinamarca, Meta, Nariño, Risaralda, Tolima, Valle); Ecuador; Peru
82 CLM 83 ECU PER
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia hohenackeri (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium hohenackeri Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 46 (1909)
= Urostachys hohenackeri (Herter) Herter ex Nessel; Bärlappgew. 105 (1939)
species :
Phlegmariurus holstii
(Hieron.) A.R.Field & Bostock; PhytoKeys 20: 41 (2013)
Distribution: Kenya; Tanzania
25 KEN TAN
Remarks: gnidioides agg.
Synonyms: hidden
= Huperzia holstii (Hieron.) Pic.Serm.; Webbia 23: 163 (1968)
= Lycopodium holstii Hieron.; Engl., Pflanzenw. Ost-Afrikas C: 90 (1895)
= Urostachys holstii (Hieron.) Herter ex Nessel; Bärlappgew. 202 (1939)
species :
Phlegmariurus homocarpus
(Herter) B.Øllg.; Phytotaxa 57: 15 (2012)
Distribution: Costa Rica; Panama; Cuba; Hispaniola (Haiti, Dominican Republic); Colombia (Cundinamarca, Narino); Venezuela (Merida); Ecuador
80 COS PAN 81 CUB HIS 82 CLM VEN 83 ECU
Remarks: dichotoma agg.
Synonyms: hidden
= Huperzia homocarpa (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium homocarpum (Herter) C.Chr.; Kongl. Vetensk. Acad. Handl. 3. Ser. 16(2): 92 (1937)
= Urostachys homocarpus Herter; Repert. Spec. Nov. Regni Veg. 19: 163 (1923)
species :
Phlegmariurus horizontalis
(Nessel) A.R.Field & Bostock; PhytoKeys 20: 41 (2013)
Distribution: Borneo (Sabah); New Guinea (Papua New Guinea, d´Entrecasteaux Arch.: Fergusson Isl., Louisiade Arch.: Misima Isl., Sudest Isl.); Bismarck Arch. (Admiralty Isl.)
42 BOR NWG 43 BIS
Remarks: status doubtful, according to Checklist of New Guinea a Selaginellaceae
Synonyms: hidden
= Huperzia horizontalis (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium aellenii (Herter) M.Tagawa; Acta Phytotax. Geobot. 26: 108 (1974)
= Lycopodium horizontale Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 30 (1912) (non C. Presl (1825))
= Urostachys horizontalis Herter ex Nessel; Bärlappgew. 225 (1939)
species :
Phlegmariurus huberi
(B.Øllg.) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Venezuela (Amazonas, Bolívar); Brazil (Roraima)
82 VEN 84 BRA
Synonyms: hidden
= Huperzia huberi B.Øllg.; Amer. Fern J. 79: 153 (1989)
species :
Phlegmariurus hugoi
B.Øllg.; Phytotaxa 246(2): 97 (2016)
Distribution: Ecuador
83 ECU
species :
Phlegmariurus humbertii
(Nessel) A.R.Field & Bostock; PhytoKeys 20: 41 (2013)
Distribution: Madagascar
29 MDG
Remarks: obtusifolia agg.
Synonyms: hidden
= Urostachys humbertii Nessel; Repert. Spec. Nov. Regni Veg. 48: 170, t. 317 (307), f. 3 (1940)
= Huperzia teretirigida Rakotondr.; Candollea 70(1): 53 (2015)
species :
Phlegmariurus humbertii-henrici
(Herter) A.R.Field & Bostock; PhytoKeys 20: 41 (2013)
Distribution: Madagascar
29 MDG
Remarks: gnidioides agg.
Synonyms: hidden
= Huperzia humbertii-henrici (Herter) Tardieu; Adansonia ser. 2 10: 18 (1970)
= Urostachys humbertii-henrici Herter; Revista Sudamer. Bot. 8: 23 (1949)
species :
Phlegmariurus hypogaeus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Colombia (Boyacá); Ecuador; Peru; Bolivia (La Paz)
82 CLM 83 BOL ECU PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia hypogaea B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 58, f. 8E (1988)
species :
Phlegmariurus hystrix
(Herter) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Colombia (Cauca, Chocó, Huila, Nariño, Risaralda); Ecuador
82 CLM 83 ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia hystrix (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium hystrix (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys hystrix Herter; Revista Sudamer. Bot. 10: 120 (1953 publ. 1954)
species :
Phlegmariurus idroboi
B.Øllg.; Phytotaxa 426(1): 80 (2019)
Distribution: Colombia (Cauca, Huila, Narino)
82 CLM
species :
Phlegmariurus iminii
Kiew; PhytoKeys 96: 100 (2018)
Distribution: peninsular Malaysia
42 MLY
species :
Phlegmariurus intermedius
(Trevis.) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Guadeloupe; Venezuela (Bolívar, Monagas); Guyana (Mt. Roraima); Brazil (Bahia, Espírito Santo, Minas Gerais)
81 GDL 82 GUY VEN 84 BRA
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia intermedia Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875), nom. nov. for Lycopodium intermedium Spring, non Blume
= Lycopodium commutatum Spring ex Klotzsch; Linnaea 18: 517 (1844) (nom. inval.)
= Lycopodium intermedium Spring; Mart., Fl. Bras. 1, 2: 111 (1840) (nom. illeg., non Lycopodium intermedium Blume (1828))
= Urostachys commutatus (Spring ex Klotzsch) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 57 (1949)
= Urostachys intermedius (Trevis.) Herter; Repert. Spec. Nov. Regni Veg. 19: 163 (1923)
= Urostachys intermedius var. cipoanus Nessel; Arch. Bot. São Paulo 1: 397 (1927)
species :
Phlegmariurus itambensis
(B.Øllg. & P.G.Windisch) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Minas Gerais)
84 BRA
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia itambensis B.Øllg. & P.G.Windisch; Bradea 5: 12 (1987)
species :
Phlegmariurus jaegeri
(Herter) A.R.Field & Bostock; PhytoKeys 20: 42 (2013)
Distribution: Guinea; Sierra Leone; Liberia; Cameroon
22 GUI LBR SIE 23 CMN
Remarks: hamiltonii agg.
Synonyms: hidden
= Huperzia jaegeri (Herter) Pic.Serm.; Webbia 23: 163 (1968)
= Urostachys jaegeri Herter; Revista Sudamer. Bot. 8: 24 (1949)
= Lycopodium jaegeri (Herter) Alston; Mém. Inst. Fr. Afr. Noire 50: 21 (1957)
species :
Phlegmariurus josesantae
B.Øllg.; Phytotaxa 426(1): 82 (2019)
Distribution: Colombia (Antioquia)
82 CLM
species :
Phlegmariurus kajewskii
(Copel.) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Santa Cruz Isl. (Vanikoro)
60 SCZ
Synonyms: hidden
= Lycopodium kajewskii Copel.; J. Arnold Arbor. 12: 49 (1931)
= Urostachys kajewskii (Copel.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 66 (1949)
species :
Phlegmariurus killipii
(Herter) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Hispaniola (Dominican Republic); Colombia (Cauca, Cundinamarca, Huila, Norte de Santander, Valle); Venezuela (Amazonas, Bolívar, Falcon, Portuguesa); French Guiana; Ecuador
81 HIS 82 CLM FRG VEN 83 ECU
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia killipii (Herter) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium killipii (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys killipii Herter; Revista Sudamer. Bot. 10: 128 (1953 publ. 1954)
species :
Phlegmariurus kuesteri
(Nessel) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Ecuador; Peru
83 ECU PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia kuesteri (Nessel) B.Øllg.; Opera Bot. 92: 169 (1987)
= Urostachys kuesteri Nessel; Repert. Spec. Nov. Regni Veg. 35: 182p, t. 172 (1934)
species :
Phlegmariurus lancifolius
(Maxon) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Costa Rica; Panama
80 COS PAN
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia lancifolia (Maxon) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium lancifolium Maxon; Contr. U.S. Natl. Herb. 17(2): 177 (1913)
= Urostachys lancifolius (Maxon) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 66 (1949)
= Urostachys linifolius var. lancifolius (Maxon) Herter ex Nessel; Bärlappgew. 163 (1939)
species :
Phlegmariurus lauterbachii
(E.Pritz. ex K.Schum. & Lauterb.) A.R.Field & Bostock; PhytoKeys 20: 42 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: squarrosa agg.
Synonyms: hidden
= Huperzia lauterbachii (E.Pritz. ex K.Schum. & Lauterb.) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium lauterbachii E.Pritz. ex K.Schum. & Lauterb.; Fl. Schutzgeb. Südsee 149 (1900)
= Urostachys lauterbachii (E.Pritz. ex K.Schum. & Lauterb.) Herter ex Nessel; Bärlappgew. 203 (1939)
species :
Phlegmariurus lecomteanus
(Nessel) A.R.Field & Bostock; PhytoKeys 20: 42 (2013)
Distribution: Madagascar
29 MDG
Remarks: included in H. squarrosa by Roux (2009)
Synonyms: hidden
= Huperzia lecomteana (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Urostachys lecomteanus Nessel; Repert. Spec. Nov. Regni Veg. 36: 1871, t. 174 (1934)
species :
Phlegmariurus ledermannii
(Herter) A.R.Field & Bostock; PhytoKeys 20: 42 (2013)
Distribution: Sulawesi; New Guinea (Irian Jaya, Papua New Guinea, d´Entrecasteaux Arch.: Normanby Isl., Louisiade Arch.: Misima Isl., Rossel Isl., Woodlark Isl.); Bismarck Arch. (New Hanover); Solomon Isl. (Guadalcanal, Kolombangara, Malaita, New Georgia, Ranongga, Rendova, Santa Ysabel, Vangunu, Vella Lavella)
42 NWG SUL 43 BIS SOL
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia ledermannii (Herter) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium ledermannii Herter; Bot. Jahrb. Syst. 54: 232 (1916)
= Lycopodium phlegmaria var. brachystachyum Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 530 (1913)
= Urostachys ledermannii (Herter) Herter ex Nessel; Bärlappgew. 236 (1939)
species :
Phlegmariurus lehnertii
Testo; Phytotaxa 297(3): 289 (2017), nom. nov.
Distribution: Vietnam
41 VIE
Synonyms: hidden
= Phlegmariurus lancifolius V.T.Tran & Duy; Adansonia 38(2): 152 (2016), nom. illeg., non (Maxon) B. Øllg.
species :
Phlegmariurus lignosus
(Herter) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Colombia (Boyacá, Cesar, Huila, Norte de Santander, Santander)
82 CLM
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia lignosa (Herter) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium lignosum Herter; Bot. Jahrb. Syst. 43: Beibl.- 98: 46 (1909)
= Urostachys lignosus (Herter) Herter ex Nessel; Bärlappgew. 104 (1939)
species :
Phlegmariurus lindenii
(Spring) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Colombia (Cundinamarca, Nariño, Santander, Tolima); Ecuador
82 CLM 83 ECU
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia lindenii (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
= Huperzia sodiroana (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Huperzia stuebelii (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Huperzia wohlberedtii (Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Lycopodium lindenii Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 27 (1849)
= Lycopodium sodiroanum Herter; Hedwigia 49: 91 (1909)
= Lycopodium stuebelii Hieron. & Herter ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 53 (1909)
= Lycopodium wohlberedtii (Nessel) C.V.Morton; Amer. Fern J. 54: 73 (1964)
= Urostachys lindenii (Spring) Herter ex Nessel; Bärlappgew. 177 (1939)
= Urostachys sodiroanus (Herter) Herter ex Nessel; Bärlappgew. 240 (1939)
= Urostachys stuebelii (Hieron. & Herter) Herter ex Nessel; Bärlappgew. 241 (1939)
= Urostachys wohlberedtii Nessel; Repert. Spec. Nov. Regni Veg. 39: 69, t. 193c (1935)
species :
Phlegmariurus linifolius
(L.) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Mexico (Guerrero, Tabasco, Chiapas, Veracruz, Puebla, Oaxaca); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Isla del Coco; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; St. Kitts; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Grenada; Tobago; Trinidad; Colombia (Amazonas, Antioquia, Boyacá, Caquetá, Cauca, Chocó, Cundinamarca, La Guajira, Magdalena, Nariño, Quindío, Risaralda, Santander, Valle); Venezuela (Amazonas, Bolívar, Carabobo, Delta Amacuro, Falcon, Sucre, Yaracuy); French Guiana; Ecuador
79 MEX 80 BLZ COC COS ELS GUA HON NIC PAN 81 CUB DOM GDL GRE HIS JAM MAR PUE STK STL STV TRT 82 CLM FRG VEN 83 ECU
Remarks: linifolia agg.
Synonyms: hidden
= Huperzia linifolia (L.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium linifolium L.; Sp. Pl. 2: 1100 (1753)
= Lycopodium linifolium var. attenuatum Jenman ex Nessel; Bärlappgew. 158 (1939)
= Lycopodium linifolium var. flexuosum S.Sm ex Nessel; Bärlappgew. 163 (1939)
= Lycopodium linifolium var. gracile L´Herm. ex Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 129 (1866)
= Lycopodium linifolium var. laxum Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 129 (1866)
= Lycopodium linifolium var. quitense Christ ex Nessel; Bärlappgew. 173 (1939)
= Plananthus linifolius (L.) P.Beauv.; Prodr. Aethéogam. 111 (1805)
= Urostachys linifolia (L.) Herter; Repert. Spec. Nov. Regni Veg. 19: 165 (1923)
= Urostachys linifolius var. gracilis (L´Herm. ex Fée) Herter ex Nessel; Bärlappgew. 161 (1939)
= Urostachys linifolius var. laxus (Fée) Herter ex Nessel; Bärlappgew. 162 (1939)
= Urostachys linifolius var. mexicanus Herter ex Nessel; Revista Sudamer. Bot. 6: 164 (1940)
= Urostachys linifolius var. minor Herter; Repert. Spec. Nov. Regni Veg. 19: 165 (1923)
variant :
var. jenmanii
(Underw. & F.E.Lloyd) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Colombia (Amazonas, Norte de Santander, Vaupes, Vichada); Venezuela (Amazonas, Bolívar, Delta Amacuro); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (La Paz, Santa Cruz); Brazil (Amazonas, Amapá, Bahia, Mato Grosso, Pará, Rondônia)
82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Huperzia jenmanii (Underw. & F.E.Lloyd) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Huperzia linifolia var. jenmanii (Underw. & F.E.Lloyd) B.Øllg. & P.G.Windisch; Bradea 5: 13 (1987)
= Lycopodium jenmanii Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 112 (1906)
= Urostachys jenmanii (Underw. & F.E.Lloyd) Herter ex Nessel; Bärlappgew. 158 (1939)
variant :
var. planifolius
(B.Øllg.) B.Øllg.; Phytotaxa 57: 16 (2012), nom. inval.
Distribution: Colombia (Cesar, Magdalena, Norte de Santander); Venezuela; Ecuador
82 CLM VEN 83 ECU
Synonyms: hidden
= Huperzia linifolia var. planifolia B.Øllg.; Nordic J. Bot. 23(1): 35 (2004)
variant :
var. tenuifolius
(Schomb. ex Nessel) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Colombia (Cauca, Santander, Valle); Venezuela (Merida, Trujillo); Ecuador; Peru; Bolivia (Beni, Cochabamba, La Paz, Santa Cruz)
82 CLM VEN 83 BOL ECU PER
Synonyms: hidden
= Urostachys linifolius var. tenuifolius Schomb. ex Nessel; Bärlappgew. 162, tab. 39, fig. 14 (1939), et Revista Sudamer. Bot. 6(5-6): 164 (1940)
= Huperzia linifolia var. tenuifolia (Schomb. ex Nessel) B.Øllg.; Fl. Ecuador 33: 90 (1988)
species :
Phlegmariurus llanganatensis
(B.Øllg.) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: ?Colombia (?Narino); Ecuador
83 ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia llanganatensis B.Øllg.; Fl. Ecuador 33: 35 (1988)
species :
Phlegmariurus lockyeri
(D.L.Jones & B.Gray) A.R.Field & Bostock; PhytoKeys 20: 42 (2013)
Distribution: Australia (NE-Queensland)
50 AUS
Remarks: carinata agg.
Conservation Status: IUCN V
Synonyms: hidden
= Huperzia lockyeri (D.L.Jones & B.Gray) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium lockyeri D.L.Jones & B.Gray; Austrobaileya 2: 126, f. 1 (1985)
species :
Phlegmariurus loefgrenianus
(Silveira) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Huperzia leitzii (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Huperzia loefgreniana (Silveira) B.Øllg. & P.G.Windisch; Bradea 5: 13 (1987)
= Lycopodium loefgrenianum Silveira; Fl. serr. Min. 78, t. 30 (1908)
= Urostachys leitzii Nessel; Repert. Spec. Nov. Regni Veg. 36: 184, pl. 173 (1934)
= Urostachys loefgrenianus (Silveira) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 68 (1949)
species :
Phlegmariurus longus
(Copel.) A.R.Field & Bostock; PhytoKeys 20: 42 (2013)
Distribution: Solomon Isl. (Makira, Malaita, Vella Lavella)
43 SOL
Remarks: squarrosa agg.
Synonyms: hidden
= Lycopodium longum Copel.; Philipp. J. Sci. 60: 100, t. 3 (1936)
= Urostachys longus (Copel.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 68 (1949)
species :
Phlegmariurus loxensis
(B.Øllg.) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Ecuador
83 ECU
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia loxensis B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 17, f. 3B (1988)
species :
Phlegmariurus luteynii
B.Øllg.; Nordic J. Bot. 33(2): 194 (2014)
Distribution: Colombia (Choco, Valle)
82 CLM
species :
Phlegmariurus macbridei
(Herter) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Ecuador; Peru
83 ECU PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia macbridei (Herter) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium macbridei (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys macbridei Maxon ex Herter; Revista Sudamer. Bot. 10: 115 (1953 publ. 1954)
species :
Phlegmariurus macgregorii
(Baker) A.R.Field & Bostock; PhytoKeys 20: 42 (2013)
Distribution: New Guinea (Papua New Guinea, Irian Jaya)
42 NWG
Remarks: macgregorii agg.
Synonyms: hidden
= Huperzia keysseri (Herter & Schltr. ex Diels) P.J.Edwards; Guide Alpine Subalpine Fl. Mount Jaya (R. J. Johns et al.) 31 (2006)
= Huperzia macgregorii (Baker) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Huperzia versteegii (Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Huperzia whartoniensis (Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Lycopodium keysseri Herter & Schltr. ex Diels; Bot. Jahrb. Syst. 62: 456 (1929)
= Lycopodium macgregorii Baker; J. Bot. 28: 109 (1890)
= Lycopodium versteegii Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 4 (1917)
= Urostachys macgregorii (Baker) Herter ex Nessel; Bärlappgew. 184 (1939)
= Urostachys versteegii (Alderw.) Herter ex Nessel; Bärlappgew. 182 (1939)
= Urostachys whartoniensis Nessel; Repert. Spec. Nov. Regni Veg. 39: 62, t. 190b (1935)
species :
Phlegmariurus macrostachys
(Hook. ex Spring) N.C.Nair & S.R.Ghosh; J. Econ. Taxon. Bot. 12(1): 194 (1988)
Distribution: India (Karnataka, Kerala, Tamil Nadu); Sri Lanka; Andaman Isl.; Nicobar Isl.; peninsular Malaysia (Penang, Kelantan, Perak, Negeri Sembilan); +Singapore; Borneo (Mt. Kinabalu, Sabah); Philippines; New Caledonia; Fiji (Viti Levu); [Western] Samoa (Savai´i); Fed. States of Micronesia (Pohnpei)
40 IND SRL 41 AND NCB 42 BOR MLY PHI SNG 60 FIJ NWC SAM 62 MIC
Remarks: phyllanthus agg. Validity of name uncertain. Was formerly united with H. phyllanthus s. str., which is an Hawaiian endemic. What is listed here is "everything else".
Synonyms: hidden
= Huperzia macrostachys (Spring) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium macrostachys Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 30 (1849)
= Lycopodium pachystachyon Spring; Bull. Acad. Roy. Soc. Bruxelles 8: 518 (1841) (non Desv. ex Poir. (1814))
= Urostachys macrostachys (Spring) Herter ex Nessel; Bärlappgew. 211 (1939)
= Urostachys pachystachyos (Spring) Herter ex Nessel; Bärlappgew. 211 (1939)
= Urostachys pachystachyos (Spring) Rothm.; Feddes Repert. 54: 62 (1944), isonym
species :
Phlegmariurus magnusianus
(Herter) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Philippines (Luzon, Mindanao)
42 PHI
Remarks: squarrosa agg.
Synonyms: hidden
= Huperzia magnusiana (Herter) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Huperzia whitfordii (Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Lycopodium magnusianum Herter; Hedwigia 49(2-3): 91 (1909)
= Lycopodium whitfordii (Herter) Copel.; Univ. Calif. Publ. Bot. 12: 384 (1931)
= Phlegmariurus whitfordii (Herter) A.R.Field & Bostock; PhytoKeys 20: 49 (2013)
= Urostachys magnusianus (Herter) Herter; Bot. Arch. 3: 14 (1923)
= Urostachys whitfordii Herter; Bot. Arch. 3: 14 (1923)
species :
Phlegmariurus mandiocanus
(Raddi) B.Øllg.; Rodriguésia 63(2): 480 (2012)
Distribution: Brazil (Alagoas, Bahia, Ceará, Espírito Santo, Minas Gerais, Mato Grosso do Sul, Pernambuco, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Paraguay (Canindeyu); Argentina (Misiones, Salta, Tucuman, Jujuy)
84 BRA 85 ARG PAR
Remarks: wilsonii agg.
Synonyms: hidden
= Huperzia mandiocana (Raddi) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium dichotomum Grev. & Hook.; nom. illeg., non Jacq. 1762
= Lycopodium dichotomum var. mandiocana (Raddi) Rosenst.; Hedwigia 46: 165 (1906)
= Lycopodium mandiocanum Raddi; Pl. Bras. 1: 77-78, t. 4 (1825)
= Lycopodium mandiocanum var. brasiliense Spring; Bull. Acad. Roy. Soc. Bruxelles 15: 45 (1842)
= Lycopodium pseudomandiocanum Herter; Beibl. Bot. Jahrb. Syst. 98: 35, 49 (1909)
= Urostachys mandiocanus (Raddi) Herter; Repert. Spec. Nov. Regni Veg. 19: 164 (1923)
= Urostachys mandiocanus var. brasiliensis (Spring) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 410 (1927)
= Urostachys pseudomandiocanus (Herter) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 411 (1927)
species :
Phlegmariurus mannii
(Hillebr.) W.H.Wagner; Contr. Univ. Michigan Herb. 20: 241 (1995)
Distribution: Hawaii (Kauai, Maui, Hawaii Isl.)
63 HAW
Remarks: myrsinites agg.
Conservation Status: IUCN V
Synonyms: hidden
= Huperzia mannii (Hillebr.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium mannii (Hillebr.) Skottsb.; Acta Hort. Gothob. 15: 131-132 (1942)
= Lycopodium phlegmaria var. mannii Hillebr.; Fl. Hawaiian Isl. 645 (1888)
species :
Phlegmariurus marsupiiformis
(D.L.Jones & B.Gray) A.R.Field & Bostock; PhytoKeys 20: 43 (2013)
Distribution: Australia (NE-Queensland)
50 AUS
Remarks: phlegmaria agg.
Conservation Status: IUCN V
Synonyms: hidden
= Huperzia marsupiiformis (D.L.Jones & B.Gray) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium marsupiiforme D.L.Jones & B.Gray; Austrobaileya 2: 128, f. 2 (1985)
species :
Phlegmariurus martii
(Wawra) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Bahia, Espírito Santo, Minas Gerais, Pernambuco)
84 BRA
Synonyms: hidden
= Huperzia martii (Wawra) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium cipoense Damazio; Bull. Soc. bot. Genève, sér. 2, 7: 119, f. 1-8 (1915)
= Lycopodium martii Wawra; Oesterr. Bot. Zeitung (Berlin) 13: 218 (1863)
= Urostachys cipoensis (Damazio) Nessel; Bärlappgew. 169 (1939)
= Urostachys martii (Wawra) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 385 (1927)
= Urostachys passerinoides var. cipoensis (Damazio) Nessel; Fl. Bras. (Hoehne) 2(2): 85 (1955)
species :
Phlegmariurus megastachyus
(Baker) A.R.Field & Bostock; PhytoKeys 20: 43 (2013)
Distribution: Madagascar
29 MDG
Remarks: squarrosa agg.
Synonyms: hidden
= Huperzia megastachya (Baker) Tardieu; Adansonia 10: 18 (1970)
= Lycopodium megastachyum Baker; J. Linn. Soc., Bot. 21: 454 (1885)
= Urostachys megastachyus (Baker) Herter ex Nessel; Bärlappgew. 210 (1939)
species :
Phlegmariurus melanesicus
(Brownlie) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Vanuatu; Fiji (Viti Levu)
60 FIJ VAN
Remarks: varia agg.
Synonyms: hidden
= Huperzia melanesica (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium melanesicum Brownlie; Beih. Nova Hedwigia 55: 20i, t. 2, f. 1-2 (1977)
species :
Phlegmariurus merrillii
(Herter) A.R.Field & Bostock; PhytoKeys 20: 43 (2013)
Distribution: Philippines (Luzon)
42 PHI
Remarks: hamiltonii agg.
Synonyms: hidden
= Huperzia merrillii (Herter) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Urostachys merrillii Herter; Bot. Arch. 3: 15 (1923)
= Lycopodium merrillii (Herter) Copel.; Univ. Calif. Publ. Bot. 12: 384 (1931)
species :
Phlegmariurus mesoamericanus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Nicaragua; Costa Rica; Panama
80 COS NIC PAN
Synonyms: hidden
= Huperzia mesoamericana B.Øllg.; Novon 3(1): 67, f. 1 (1993)
species :
Phlegmariurus mexicanus
(Herter) B.Øllg.; Phytotaxa 57: 16 (2012)
Distribution: Mexico (Chiapas, Oaxaca); Guatemala; Honduras; Nicaragua
79 MEX 80 GUA HON NIC
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia mexicana (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Lycopodium mexicanum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 49 (1909)
= Urostachys mexicanus (Herter) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 164 (1923)
species :
Phlegmariurus mildbraedii
(Herter) A.R.Field & Bostock; PhytoKeys 20: 43 (2013) (milbraedii)
Distribution: Guinea; Sierra Leone; Liberia; Bioko Isl. [Fernando Poo]; Principe Isl.; São Tomé; D.R.Congo [Zaire]; Rwanda; Burundi; ?Uganda; Tanzania
22 GUI LBR SIE 23 BIO BUR PRI RWA SAO ZAI 25 TAN
Remarks: gnidioides agg. Included in P. dacrydioides by Roux (2009)
Synonyms: hidden
= Huperzia mildbraedii (Herter) Pic.Serm.; Webbia 23: 163 (1968)
= Lycopodium mildbraedii Herter; Hedwigia 49(2-3): 90 (1909)
= Urostachys mildbraedii (Herter) Herter ex Nessel; Bärlappgew. 188 (1939)
species :
Phlegmariurus mingcheensis
Ching; Acta Bot. Yunnan. 4: 125 (1982)
Distribution: China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Hainan, Hunan, Jiangxi, Sichuan, Zhejiang)
36 CHI
Remarks: hippuris agg.
Synonyms: hidden
= Huperzia mingcheensis (Ching) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium mingcheense (Ching) Ching; Fl. Fujianica (ed. 2) 1: 619, t. 6 (1991))
= Phlegmariurus mingcheensis (Ching) Li Bing Zhang; Fl. Reipubl. Popularis Sin. 6(3): 45 (2004), non Ching (1982)
= Phlegmariurus mingcheensis var. angustifolius C.Y.Ma; Bull. Bot. Res., Harbin 10(3): 57-58, t. 1 (1990)
= Phlegmariurus mingjoui X.C.Zhang; Higher Pl. China 2: 20 (2008), nom. nov. illeg. superfl.
species :
Phlegmariurus minutifolius
(Alderw.) A.R.Field & Bostock; PhytoKeys 20: 43 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia minutifolia (Alderw.) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium coralium var. minutifolium Alderw.; Mal. Fern-Allies 44 (1915)
= Lycopodium minutifolium (Alderw.) Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 44 (1918)
= Urostachys minutifolius (Alderw.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 70 (1949)
species :
Phlegmariurus mirabilis
(Willd.) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: peninsular Malaysia (Pahang, Melaka, Kedah); New Guinea (Papua New Guinea); New Caledonia; Austral Isl. (Tubuai); Palau (Babeldaob, Ulebsechel); Northern Marianas (Rota Isl.); Guam; Fed. States of Micronesia (Nukuoro, Kosrae)
42 MLY NWG 60 NWC 61 AUI 62 GUM MIC MRN PLU
Conservation Status: IUCN E
Synonyms: hidden
= Huperzia mirabilis (Willd.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Huperzia phlegmaria var. longifolium Spring
= Lycopodium mirabile Willd.; Sp. Pl. ed. 4. 5: 11 (1810), nom. cons. prop.
= Lycopodium phlegmaria var. mirabile (Willd.) R.Fosberg
= Urostachys mirabilis (Willd.) Herter ex Nessel; Bärlappgew. 225 (1939)
species :
Phlegmariurus mollicomus
(Spring) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Mexico (Oaxaca); Costa Rica; Isla del Coco; Panama; Hispaniola (Haiti, Dominican Republic); Colombia (Antioquia, ?Cauca, Choco, Santander); Venezuela (Aragua, Venezuela Distrito Federal, Falcon, Lara, Portuguesa, Tachira, Trujillo, Yaracuy); Ecuador; ?Peru; Bolivia (La Paz); Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
79 MEX 80 COC COS PAN 81 HIS 82 CLM VEN 83 BOL ECU 84 BRA
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia flaccida (Fée) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Huperzia mollicoma (Spring) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia setacea var. mollicoma (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium dichotomum subsp. mollicomum Spring; Flora 21(1): 162 (1838)
= Lycopodium flaccidum Fée; Crypt. Vasc. Brésil 2: 92, t. 106, f. 1 (1872 publ. 1873), nom. illeg., non Lycopodium flaccidum Bory (1833)
= Lycopodium mollicomum (Spring) Spring; Mart., Fl. Bras. 1(2): 113 (1840)
= Lycopodium setaceum var. brasiliense Spring; Mém. Acad. Roy. Sci. Bruxelles 15: 43 (1842)
= Lycopodium williamsii Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33(2): 112 (1906)
= Urostachys flaccidus (Fée) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 164 (1923)
= Urostachys mollicomus (Spring) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1(4): 407, t. 25 (1927)
= Urostachys mollicomus var. majus Nessel; Repert. Spec. Nov. Regni Veg. 36: 183, t. 172 (1934)
= Urostachys neptuni Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 72 (1949) (nom. illeg. superfl.)
= Urostachys setacea var. brasiliensis (Spring) Nessel; Arch. Bot. São Paulo 1: 404 (1927)
= Urostachys williamsii (Underw. & F.E.Lloyd) Herter ex Nessel; Bärlappgew. 164 (1939)
species :
Phlegmariurus monticola
Kiew; PhytoKeys 96: 103 (2018)
Distribution: peninsular Malaysia
42 MLY
species :
Phlegmariurus mooreanus
(Sander ex Baker) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Bahia, Minas Gerais)
84 BRA
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia mooreana (Baker) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia sydowiorum (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21(100): 157 (1988)
= Lycopodium luetzelburgii Rosenst.; Repert. Spec. Nov. Regni Veg. 20: 95 (1924)
= Lycopodium mooreanum Sander ex Baker; Gard. Chron., ser. 3, 12: 582 (1892)
= Lycopodium sydowiorum Herter; Repert. Spec. Nov. Regni Veg. 13: 296 (1914)
= Urostachys mooreanus (Sander ex Baker) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 70 (1949)
= Urostachys sydowiorum (Herter) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 390 (1927)
species :
Phlegmariurus multifarius
(Alderw.) A.R.Field & Bostock; PhytoKeys 20: 43 (2013)
Distribution: Moluccas (Ternate); New Guinea (Irian Jaya, Papua New Guinea)
42 MOL NWG
Remarks: hippuris agg.
Synonyms: hidden
= Huperzia multifaria (Alderw.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium multifarium Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 226 (1922)
= Urostachys multifarius (Alderw.) Herter ex Nessel; Bärlappgew. 207 (1939)
species :
Phlegmariurus myrsinites
(Lam.) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Mexico (Guerrero, Veracruz, Chiapas, Oaxaca); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Hispaniola (Haiti, Dominican Republic); Trinidad; Colombia (Cauca, Magdalena); Venezuela (Amazonas, Aragua, Bolívar, Venezuela Distrito Federal, Falcon, Lara, Miranda, Sucre); Guyana; Ecuador; Peru; Bolivia; Brazil (Amazonas, Roraima)
79 MEX 80 COS ELS GUA HON NIC 81 HIS TRT 82 CLM GUY VEN 83 BOL ECU PER 84 BRA
Remarks: myrsinites agg.
Synonyms: hidden
= Huperzia myrsinites (Lam.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1875)
= Lycopodium myrsinites Lam.; Encycl. 3: 654. (1789 publ. 1792)
= Lycopodium myrsinites var. mettenianum Hieron. ex Nessel; Bärlappgew. 244 (1939)
= Lycopodium patens Willd. ex Spreng.; Syst. Veg. ed. 14 4: 12 (1827)
= Lycopodium roraimense Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 115 (1906)
= Lycopodium skutchii Maxon; Proc. Biol. Soc. Washington 46: 159 (1933)
= Plananthus myrsinites (Lam.) P.Beauv.; Prodr. Aethéogam. 111 (1805)
= Urostachys myrsinites (Lam.) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 166 (1923)
= Urostachys patens (Willd. ex Spreng.) Herter ex Nessel; Bärlappgew. 245, t. 59, f. 25 (1939)
= Urostachys roraimensis (Underw. & F.E.Lloyd) Herter ex Nessel; Bärlappgew. 246 (1939)
species :
Phlegmariurus myrtifolius
(G.Forst.) A.R.Field & Bostock; PhytoKeys 20: 43 (2013)
Distribution: Australia (Queensland, Tasmania); Vanuatu
50 AUS 60 VAN
Remarks: myrtifolia agg.
Synonyms: hidden
= Huperzia myrtifolia (G.Forst.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium myrtifolium G.Forst.; Fl. ins. austr. 87 (1786)
= Stachygynandrum myrtifolium (G.Forst.) P.Beauv.; Prodr. Aethéogam. 113 (1805)
= Urostachys myrtifolius (G.Forst.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 71 (1949)
species :
Phlegmariurus myrtuosus
(Spring) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Colombia (Caldas, Magdalena); Venezuela (Tachira)
82 CLM VEN
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia lellingeri (Rolleri) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Huperzia myrtuosa (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium lellingeri Rolleri; Hickenia 1(26): 143 (1978)
= Lycopodium myrtuosum Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 9 (1849)
= Urostachys myrtuosus (Spring) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 387 (1927)
species :
Phlegmariurus neocaledonicus
(Nessel) A.R.Field & Bostock; PhytoKeys 20: 44 (2013)
Distribution: New Caledonia
60 NWC
Remarks: hippuris agg.
Synonyms: hidden
= Huperzia neocaledonica (Nessel) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Urostachys neocaledonicus Nessel; Repert. Spec. Nov. Regni Veg. 36: 187, t. 175 (1934)
species :
Phlegmariurus nesselii
(Brause ex Nessel) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Peru
83 PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia nesselii (Brause ex Nessel) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21(100): 156 (1988)
= Lycopodium nesselii (Brause ex Nessel) Rolleri; Revista Mus. La Plata, Secc. Bot. 71: 77 (1980)
= Urostachys nesselii Brause ex Nessel; Bärlappgew. 78, f. 12 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 161 (1940)
species :
Phlegmariurus niligaricus
(Spring) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: India (Tamil Nadu, Kerala, Karnataka); Sri Lanka
40 IND SRL
Remarks: hamiltonii agg.
Conservation Status: IUCN VU (Fraser-Jenkins 2012)
Synonyms: hidden
= Huperzia hilliana (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985) (fide Fraser-Jenkins 2008)
= Huperzia niligarica (Spring) R.D.Dixit; J. Bombay Nat. Hist. Soc. 77(3): 541 (1981)
= Lycopodium indicum D.D.Pant & P.S.Pandey; Phyta Monogr. 3: 24 (1985)
= Lycopodium niligaricum Spring; Bull. Acad. Roy. Sci. Bruxelles 8: 517 (1841)
= Phlegmariurus hillianus (Nessel) A.R.Field & Bostock; PhytoKeys 20: 41 (2013)
= Phlegmariurus nilagiricus (Spring) A.R.Field & Bostock; PhytoKeys 20: 44 (2013), sphalm.
= Urostachys hillianus Nessel; Repert. Spec. Nov. Regni Veg. 36: 186, t. 174 (1934)
species :
Phlegmariurus nudus
(Christ ex Nessel & Hoehne) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Espírito Santo, Minas Gerais, Rio de Janeiro, São Paulo)
84 BRA
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia nuda (Christ ex Nessel & Hoehne) B.Øllg. & P.G.Windisch; Bradea 5: 14 (1987)
= Urostachys nudiusculus Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 72 (1949) (nom. superfl. for Urostachys nudus Christ ex Nessel 1927)
= Urostachys nudus Christ ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 396, tab. 14 (1927)
species :
Phlegmariurus nummulariifolius
(Blume) Ching; Acta Bot. Yunnan. 3: 298 (1981)
Distribution: Nicobar Isl.; Thailand; peninsular Malaysia (Kedah, Penang, Kelantan, Pahang, Johor); +Singapore; Sumatra; Borneo (Sabah, Sarawak); Sulawesi; Philippines; Java; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, New Georgia, Nggatokae, Santa Ysabel, Vella Lavella, Makira, Malaita, Mono); Santa Cruz Isl. (Vanikoro); Vanuatu; Fiji (Viti Levu, Ovalau, Taveuni)
41 NCB THA 42 BOR JAW MLY MOL NWG PHI SNG SUL SUM 43 BIS SOL 60 FIJ SCZ VAN
Remarks: nummulariifolia agg.
Conservation Status: IUCN EN (Fraser-Jenkins 2012)
Synonyms: hidden
= Huperzia nummulariifolia (Blume) Jermy; Brit. Fern Gaz. 10: 176 (1971)
= Lycopodium nummulariifolium Blume; Enum. Pl. Javae 2: 263 (1828)
= Lycopodium rotundifolium Roxb.; Calcutta J. Nat. Hist. 4: 473 (1844)
= Urostachys nummulariifolius (Blume) Herter; Philipp. J. Sci.22: 182 (1923)
species :
Phlegmariurus nutans
(Brack.) W.H.Wagner; Contr. Univ. Michigan Herb. 20: 241 (1995)
Distribution: ?New Caledonia; ?Vanuatu; Cook Isl. (Rarotonga); Hawaii (Kauai, Oahu)
61 COO 63 HAW
Remarks: squarrosa agg.
Conservation Status: IUCN E
Synonyms: hidden
= Huperzia nutans (Brack.) Rothm.; Feddes Repert. 54: 62 (1954)
= Lycopodium nutans Brack.; U.S. Expl. Exped., Filic. 16: 327, t. 46 (1854)
= Urostachys nutans (Brack.) Herter ex Nessel; Bärlappgew. 208 (1939)
species :
Phlegmariurus nylamensis
(Ching & S.K.Wu) H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 37(1): 52 (1999)
Distribution: Tibet
36 TIB
Synonyms: hidden
= Huperzia nylamensis Ching & S.K.Wu; Acta Bot. Yunnan. 3(3): 305 (1981)
species :
Phlegmariurus obovalifolius
(Bonap.) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Vietnam
41 VIE
Remarks: obtusifolia agg.
Synonyms: hidden
= Phlegmariurus obovalifolius (Bonap.) V.T.Tran & Duy; Adansonia 38(2): 152 (2016), isonym
= Lycopodium obovalifolium Bonap.; Notes Ptéridol. 14: 190 (1923)
= Urostachys obovalifolius (Bonap.) Nessel; Bärlappgew. 249 (1939)
species :
Phlegmariurus obtusifolius
(P.Beauv.) A.R.Field & Bostock; PhytoKeys 20: 44 (2013)
Distribution: ?Madagascar; La Réunion
29 REU
Remarks: obtusifolia agg.
Synonyms: hidden
= Huperzia obtusifolia (P.Beauv.) Rothm.; Feddes Repert. Regni Veg. 54(1): 61 (1944)
= Lepidotis obtusifolia P.Beauv.; Prodr. Aethéogam. 109 (1805)
= Lycopodium obtusifolium (P.Beauv.) Sw.; Syn. Fil. 177 (1806)
= Urostachys obtusifolius (P.Beauv.) Herter ex Nessel; Bärlappgew. 190 (1939)
species :
Phlegmariurus ocananus
(Herter) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Colombia (Antioquia, Boyacá, Cesar, Chocó, Cundinamarca, Magdalena, Norte de Santander, Valle); Venezuela (Merida, Tachira, Trujillo)
82 CLM VEN
Remarks: saururus agg.
Synonyms: hidden
= Huperzia arthurii (Herter) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Huperzia moritzii (Nessel) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Huperzia ocanana (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia trachyloma (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 157 (1988)
= Lycopodium arcturi (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium arthurii (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium ocananum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909)
= Lycopodium trachyloma (Herter) Maxon ex C.V.Morton; Amer. Fern J. 54: 73 (1964)
= Phlegmariurus arcturi (Herter) B.Øllg.; Phytotaxa 57: 12 (2012) (fide Ollgaard 2019)
= Urostachys arcturi Herter; Revista Sudamer. Bot. 10: 118 (1953)
= Urostachys arthurii Maxon ex Herter; Revista Sudamer. Bot. 10: 114 (1953 publ. 1954)
= Urostachys moritzii Herter ex Nessel; Bärlappgew. 70 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 1601, t. 9, f. 29 (1940)
= Urostachys ocananus (Herter) Herter ex Nessel; Bärlappgew. 101 (1939)
= Urostachys trachyloma Maxon ex Herter; Revista Sudamer. Bot. 10: 113 (1953 publ. 1954)
species :
Phlegmariurus oceanianus
(Herter) A.R.Field & Bostock; PhytoKeys 20: 44 (2013)
Distribution: Vanuatu
60 VAN
Remarks: myrtifolia agg.
Synonyms: hidden
= Huperzia oceanica (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985), sphalm.
= Lycopodium oceanianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 52 (1909)
= Phlegmariurus oceanianus (Herter) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015), isonym
= Urostachys oceanianus (Herter) Herter ex Nessel; Bärlappgew. 237 (1939)
species :
Phlegmariurus oellgaardii
(A.Rojas) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Costa Rica
80 COS
Synonyms: hidden
= Huperzia oellgaardii A.Rojas; Lankesteriana 5(2): 110 (2005)
species :
Phlegmariurus oltmannsii
(Herter ex Nessel) A.R.Field & Bostock; PhytoKeys 20: 44 (2013)
Distribution: Society Isl. (Tahiti)
61 SCI
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia oltmannsii (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Urostachys oltmannsii Herter ex Nessel; Bärlappgew. 231 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 168, t. 14, f. 71 (1940)
species :
Phlegmariurus ophioglossoides
(Lam.) A.R.Field & Bostock; PhytoKeys 20: 44 (2013)
Distribution: Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Annobon Isl.; Congo [Brazzaville]; D.R.Congo [Zaire]; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga); Madagascar; Comores; Mauritius; La Réunion; Seychelles
23 ANN BIO BUR CMN CON SAO ZAI 24 ETH SUD 25 KEN TAN UGA 26 MLW MOZ ZIM 27 SAF 29 COM MAU MDG REU SEY
Remarks: ophioglossoides agg.
Synonyms: hidden
= Huperzia ophioglossoides (Lam.) Rothm.; Feddes Repert. 54: 62 (1944)
= Lepidotis longifolia P.Beauv.; Prodr. Aethéogam. 109 (1805)
= Lycopodium longifolium (P.Beauv.) Sw.; Syn. Fil. 177 (1806)
= Lycopodium ophioglossoides Lam.; Encycl. 3: 646 (1789)
= Urostachys ophioglossoides (Lam.) Herter ex Nessel; Bärlappgew. 238 (1939)
= Urostachys phlegmaria var. densifolius (Spring) Herter ex Nessel; Bärlappgew. 221 (1939)
species :
Phlegmariurus orizabae
(Underw. & F.E.Lloyd) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Mexico (Guerrero, Chiapas, Veracruz, Oaxaca); Guatemala; El Salvador; Honduras
79 MEX 80 ELS GUA HON
Remarks: linifolia agg.
Synonyms: hidden
= Huperzia orizabae (Underw. & F.E.Lloyd) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium orizabae Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 110 (1906)
= Urostachys orizabae (Underw. & F.E.Lloyd) Herter ex Nessel; Bärlappgew. 175 (1939)
species :
Phlegmariurus ovatifolius
(Ching) W.M.Chu ex H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 37(1): 52 (1999)
Distribution: China (S-Yunnan)
36 CHI
Remarks: hamiltonii agg.
Synonyms: hidden
= Huperzia ovatifolia Ching; Acta Bot. Yunnan. 3: 298 (1981)
species :
Phlegmariurus pachyphyllus
(Kuhn ex Herter) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Madagascar
29 MDG
Remarks: obtusifolia agg.
Synonyms: hidden
= Huperzia obtusifolia var. pachyphylla (Kuhn ex Herter) C.Chr.; Cat. Pl. Madag. Pterid. 69 (1931)
= Huperzia pachyphylla (Kuhn ex Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium pachyphyllum Kuhn ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 51 (1909) (non Kuhn ex E. Pritz. 1901)
= Urostachys pachyphyllus (Herter) Herter ex Nessel; Bärlappgew. 190 (1939)
species :
Phlegmariurus pachyskelos
B.Øllg.; Phytotaxa 277(3): 268 (2016)
Distribution: Colombia (Boyaca, Cauca, Huila, Meta)
82 CLM
species :
Phlegmariurus parksii
(Copel.) A.R.Field & Bostock; PhytoKeys 20: 45 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea); Fiji (Viti Levu, Taveuni)
42 NWG 60 FIJ
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia parksii (Copel.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium parksii Copel.; Bull. Bernice P. Bishop Mus. 59: 8, t. 2 (1929)
= Urostachys parksii (Copel.) Herter ex Nessel; Bärlappgew. 124 (1939)
species :
Phlegmariurus patentissimus
(Alderw.) A.R.Field & Bostock; PhytoKeys 20: 45 (2013)
Distribution: Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (incl. Makira Isl.)
42 MOL NWG 43 SOL
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia patentissima (Alderw.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium patentissimum Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 4 (1917)
= Urostachys patentissimus (Alderw.) Herter ex Nessel; Bärlappgew. 226 (1939)
species :
Phlegmariurus pecten
(Baker) A.R.Field & Bostock; PhytoKeys 20: 45 (2013)
Distribution: Madagascar
29 MDG
Remarks: pecten agg.
Synonyms: hidden
= Huperzia pecten (Baker) Tardieu; Adansonia 10: 18 (1970)
= Lycopodium pecten Baker; J. Linn. Soc., Bot. 15: 421 (1876)
= Urostachys pecten (Baker) Herter ex Nessel; Bärlappgew. 67 (1939)
species :
Phlegmariurus perrerianus
(Tardieu) A.R.Field & Bostock; PhytoKeys 20: 45 (2013)
Distribution: Madagascar
29 MDG
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia perreriana Tardieu; Adansonia 10: 20, t. 2, fig. 14 (1970)
species :
Phlegmariurus pflanzii
(Nessel) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Bolivia (Cochabamba, La Paz)
83 BOL
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia pflanzii (Nessel) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100) 156 (1988)
= Lycopodium pflanzii (Nessel) Lellinger; Proc. Biol. Soc. Washington 89(61): 717 (1977)
= Urostachys pflanzii Nessel; Repert. Spec. Nov. Regni Veg. 36(12-15): 181, t. 171 (1934)
species :
Phlegmariurus phlegmaria
(L.) Holub; Preslia 36(1): 21 (1964)
Distribution: Guinea; Sierra Leone; Liberia; Ghana; Cameroon; Bioko Isl. [Fernando Poo]; Principe Isl.; São Tomé; Equatorial Guinea (Rio Muni); Gabon; Gabon; ?D.R.Congo [Zaire]; Uganda; Tanzania; Malawi; Comores (Mayotte); ?Mauritius; +Rodrigues; Seychelles; China (Guangdong, Guangxi, Hainan, Yunnan); Japan; Ryukyu Isl.; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Karnataka, Kerala, Meghalaya, Nagaland, Sikkim, Tamil Nadu, Tripura, West Bengal); Sri Lanka; Bangladesh; Andaman Isl.; Nicobar Isl.; Myanmar (Kachin, etc.); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (Perak, Pahang, Johor); Singapore; Sumatra; Borneo (Sabah, Sarawak, Mt. Kinabalu, Sabah); Sulawesi; Palawan; Palawan; Philippines; Lesser Sunda Isl.; Moluccas (Seram); Christmas Isl. [Austr.]); New Guinea (Papua New Guinea); Bismarck Arch. (Admiralty Isl.); Australia (E-Queensland); New Caledonia; Rotuma Isl.; Tonga (Late Isl., ´Eua); Niue; Fiji; Wallis & Futuna (Alofi, Futuna, ´Uvea, Wallis); American Samoa (Tutuila, Olosega, Ta´u); [Western] Samoa (Aleipata Isl., Savai´i, ´Upolu); Cook Isl. (Rarotonga); Society Isl. (Tahiti, Moorea); Fed. States of Micronesia (Pohnpei)
22 GHA GUI LBR SIE 23 BIO CMN EQG GAB PRI SAO 25 TAN UGA 26 MLW 29 COM ROD SEY 36 CHI 38 JAP NNS TAI 40 BAN BHU IND NEP SRL 41 AND CBD LAO MYA NCB THA VIE 42 BOR LSI MLY MOL NWG PHI PLW SNG SUL SUM 43 BIS 50 AUS 60 ASA FIJ NUE NWC ROT SAM TON WAL 61 COO SCI 62 MIC
Remarks: phlegmaria agg. Distribution might be overstated and some certainly refers to other species of the complex.
Conservation Status: endangered (Singapore)
Synonyms: hidden
= Huperzia feejeensis (Luerss.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Huperzia phlegmaria (L.) Rothm.; Feddes Repert. 54: 62 (1944)
= Huperzia phlegmaria var. divaricatum Blume; Enum. Pl. Javae 2: 262 (1828)
= Huperzia simonii (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lepidotis phlegmaria (L.) P.Beauv.; Prodr. Aethéogam. 109 (1805)
= Lycopodium filiforme Wall. ex Roxb.; Calcutta J. Nat. Hist. 4: 473 (1844), nom. illeg., non Sw. 1800
= Lycopodium horizontale var. submoniliforme Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 4 (1917)
= Lycopodium phlegmaria L.; Sp. Pl. 2: 1101 (1753)
= Lycopodium phlegmaria var. longibracteatum Domin; Biblioth. Bot. 20: 228 (1915)
= Lycopodium phlegmaria var. parvifolium Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 65 (1842)
= Lycopodium phlegmarium var. feejeense Luerss.; Fil. Graeff. 1: 270 (1871)
= Lycopodium transiens Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 227 (1922)
= Lycopodium vrieseanum Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 33 (1849)
= Phlegmariurus phlegmaria (L.) T.Sen & U.Sen; Fern Gaz. 11(6): 421, f. 4 a (1978), isonym
= Phlegmariurus phlegmaria var. laxosporophyllus Gias & Pasha; Indian Fern J. 14: 25 (1998), without latin descr. or type
= Urostachys borneensis Herter; Philipp. J. Sci. 22: 181 (1923)
= Urostachys feejeensis (Luerss.) Herter; Notul. Syst. (Paris) 15: 355 (1958)
= Urostachys leonis Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 67 (1949), nom. nov. for U. borneensis Herter
= Urostachys phlegmaria (L.) Herter ex Nessel; Bärlappgew. 215 (1939)
= Urostachys simonii Nessel; Revista Sudamer. Bot. 6: 167, excl. fig. (1940)
= Urostachys submoniliformis (Alderw) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 84 (1949)
= Urostachys transiens (Alderw.) Herter ex Nessel; Bärlappgew. 143 (1939)
= Urostachys vrieseanus (Spring) Herter ex Nessel; Bärlappgew. 236 (1939)
species :
Phlegmariurus phlegmarioides
(Gaudich.) A.R.Field & Bostock; PhytoKeys 20: 45 (2013)
Distribution: peninsular Malaysia (Perak, Pahang, Perlis, Kedah); Moluccas; New Guinea (Irian Jaya, Papua New Guinea, Louisiade Arch.: Woodlark Isl.); Bismarck Arch. (New Hanover); Solomon Isl. (Makira, Malaita, Rennell, Bougainville, Vangunu); Australia (NE-Queensland); Santa Cruz Isl. (Vanikoro); New Caledonia; Vanuatu; Fiji (Viti Levu, Vanua Levu, Ovalau); [Western] Samoa (´Upolu); Society Isl. (Tahiti); Austral Isl. (Raivavae)
42 MLY MOL NWG 43 BIS SOL 50 AUS 60 FIJ NWC SAM SCZ VAN 61 AUI SCI
Remarks: phlegmarioides agg.
Synonyms: hidden
= Huperzia kandavuensis (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Huperzia phlegmarioides (Gaudich.) Rothm.; Feddes Repert. 54: 62 (1944)
= Huperzia pseudophlegmaria (Kuhn) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium phlegmarioides Gaudich.; Voy. Uranie [Freycinet] 1: 281, t. 23 (1828)
= Lycopodium pseudophlegmaria Kuhn; Engl., Forschungsr. Gazelle 4: 16 (1889)
= Lycopodium tetrapterygium (Herter) F.M.Bailey; Proc. Roy. Soc. Queensl. 1: 150 (1884)
= Phlegmariurus pseudophlegmaria (Kuhn) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
= Urostachys kandavuensis Nessel; Repert. Spec. Nov. Regni Veg. 39: 67, t. 192a (1935)
= Urostachys phlegmarioides (Gaudich.) Herter ex Nessel; Bärlappgew. 247 (1939)
= Urostachys pseudophlegmaria (Kuhn) Herter ex Nessel; Bärlappgew. 237 (1939)
= Urostachys tetrapterygius (F.M.Bailey) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 86 (1949)
species :
Phlegmariurus phylicifolius
(Desv. ex Poir.) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Colombia (Antioquia); ?Venezuela; Ecuador; Galapagos; Peru; Bolivia (Cochabamba, La Paz, Santa Cruz, Tarija); Argentina (Jujuy, Salta, Tucuman); Chile
82 CLM 83 BOL ECU GAL PER 85 ARG CLE
Remarks: myrsinites agg.
Synonyms: hidden
= Huperzia phylicifolia (Desv. ex Poir.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium congestifolium Spring; Bull. Acad. Roy. Soc. Bruxelles 8: 518 (1841)
= Lycopodium erythrocaulos var. pastoense (Baker) Sodiro ex Nessel; Bärlappgew. 244 (1939)
= Lycopodium nubigenum Herzog; Meded. Rijks Herb. no. 27: 2 (1915)
= Lycopodium phylicifolium Desv. ex Poir.; Encycl. suppl. 3: 546 (1813 publ. 1814), as phylicaefolium
= Lycopodium subulatum var. pastoense Baker; Handb. Fern-Allies 21 (1887)
= Urostachys erythrocaulos var. pastoensis (Baker) Herter ex Nessel; Bärlappgew. 244 (1939)
= Urostachys nubigenus (Herzog) Herter ex Nessel; Bärlappgew. 246 (1939)
= Urostachys phylicifolius (Poir.) Herter ex Nessel; Bärlappgew. 246 (1939)
= Urostachys subulatus var. congestifolius (Spring) Nessel; Bärlappgew. 243 (1939)
species :
Phlegmariurus phyllanthus
(Hook. & Arn.) R.D.Dixit; J. Bombay Nat. Hist. Soc. 77(3): 541 (1980 publ. 1981)
Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Lanai, Hawaii Isl.)
63 HAW
Remarks: phyllanthus agg.
Synonyms: hidden
= Huperzia phyllantha (Hook. & Arn.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia phyllantha (Hook. & Arn.) Panigrahi; Higher Plants Indian Subcont., add. ser. Indian J. Forest. 7, 4: 109 (1993), isonym
= Lycopodium pachystachyon var. phyllanthum (Hook. & Arn.) Hillebr.; Fl. Hawaii Isl. 644 (1888)
= Lycopodium phyllanthum Hook. & Arn.; Bot. Beech. Voy. 102 (1832 (Oct-Nov.))
= Urostachys phyllanthum (Hook. & Arn.) Herter; Philipp. J. Sci. 22: 181 (1923)
species :
Phlegmariurus picardae
(Christ ex Krug) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Hispaniola (Haiti, Dominican Republic)
81 HIS
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia picardae (Christ ex Krug) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium picardae (Christ ex Krug) Urb.; Bot. Jahrb. Syst. 24: 148 (1897)
= Urostachys picardae (Christ ex Krug) Herter; Repert. Spec. Nov. Regni Veg. 19: 166 (1923)
species :
Phlegmariurus pichianus
(Tardieu) A.R.Field & Bostock; PhytoKeys 20: 46 (2013)
Distribution: Madagascar
29 MDG
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia pichiana Tardieu; Adansonia 10: 21, t. 2, fig. 5-8 (1970)
species :
Phlegmariurus pinifolius
(Trevis.) Kiew; PhytoKeys 96: 108 (2018)
Distribution: Sri Lanka; Andaman Isl.; Nicobar Isl.; Thailand; peninsular Malaysia (Perak, Pahang, Melaka, Johor); Sumatra; Borneo (Sabah, Sarawak, Mt. Kinabalu, Sabah); Sulawesi; Philippines; Java; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.; Santa Cruz Isl. (Vanikoro); Vanuatu
40 SRL 41 AND NCB THA 42 BOR JAW MLY MOL NWG PHI SUL SUM 43 BIS 60 SCZ VAN
Remarks: hippuris agg.
Synonyms: hidden
= Huperzia pinifolia Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1874), nom. nov. for Lycopodium pinifolium Blume
= Huperzia piscium (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium pinifolium Blume; Enum. Pl. Javae 2: 264 (1828) (non Kaulf. 1824))
= Lycopodium piscium (Herter) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 22: 103 (1967)
= Urostachys mercurii Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 69 (1949)
= Urostachys pinifolius (Blume) Herter; Bot. Arch. 3: 16 (1923)
= Urostachys piscium Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 75 (1949)
species :
Phlegmariurus pithyoides
(Schltdl. & Cham.) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Mexico (Chiapas, Colima, Guerrero, Jalisco, Oaxaca, Queretaro, San Luis Potosi, Tamaulipas, Veracruz); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Hispaniola (Haiti, Dominican Republic); Colombia (Huila); Venezuela (Aragua, Falcon)
79 MEX 80 BLZ COS ELS GUA HON NIC PAN 81 CUB HIS 82 CLM VEN
Remarks: wilsonii agg.
Synonyms: hidden
= Huperzia pithyoides (Schltdl. & Cham.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium gigas Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 50 (1909)
= Lycopodium mandiocanum var. mexicanum Spring; Mém. Acad. Roy. Sci. Bruxelles 15: 45 (1842)
= Lycopodium pithyoides Schltdl. & Cham.; Linnaea 5: 623 (1830)
= Urostachys gigas (Herter) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 164 (1923)
= Urostachys mandiocanus var. mexicanus (Spring) Herter ex Nessel; Bärlappgew. 149 (1939)
= Urostachys pithyoides (Schltdl. & Cham.) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 164 (1923)
= Urostachys pithyoides var. cubanus Herter; Repert. Spec. Nov. Regni Veg. 19: 164 (1923)
species :
Phlegmariurus pittieri
(Christ) B.Øllg.; Phytotaxa 57: 17 (2012)
Distribution: Isla del Coco
80 COC
Remarks: linifolia agg.
Synonyms: hidden
= Huperzia pittieri (Christ) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium pittieri Christ; Bull. Soc. bot. Genève 1(5): 236 (1909)
= Urostachys pittieri (Christ) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 75 (1949)
species :
Phlegmariurus podocarpensis
B.Øllg.; Phytotaxa 246(2): 100 (2016)
Distribution: Ecuador
83 ECU
species :
Phlegmariurus polycarpos
(Kunze) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Costa Rica; Panama; Colombia (Antioquia, Chocó, Quindío, Risaralda, Santander, Valle); Ecuador; Peru
80 COS PAN 82 CLM 83 ECU PER
Remarks: dichotoma agg.
Synonyms: hidden
= Huperzia polycarpos (Kunze) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium cuatrecasasii (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium polycarpos Kunze; Linnaea 9: 5 (1834)
= Urostachys cuatrecasasii Herter; Revista Sudamer. Bot. 10: 123 (1953)
= Urostachys polycarpos (Kunze) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 76 (1949)
species :
Phlegmariurus polycladus
(Sodiro) comb.ined.
Distribution: Ecuador
83 ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia polyclada (Sodiro) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Lycopodium polycladum Sodiro; Crypt. Vasc. Quit. 561 (1893) (Anales Univ. Quito 12(81): 316 (1895))
= Urostachys polycladus (Sodiro) Herter ex Nessel; Bärlappgew. 73 (1939)
species :
Phlegmariurus polydactylus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Ecuador
83 ECU
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia polydactyla B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 64, f. 11B (1988)
species :
Phlegmariurus polylepidetorum
(B.Øllg.) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Ecuador; Peru
83 ECU PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia polylepidetorum B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 42, f. 6B (1988)
species :
Phlegmariurus portoricensis
(Underw. & F.E.Lloyd) comb.ined.
Distribution: Guatemala; Nicaragua; Costa Rica; Panama; ?Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Saba; Guadeloupe; Dominica; Martinique; St. Lucia; Colombia (Cesar, Meta); Venezuela; Ecuador; ?Peru; Bolivia (Santa Cruz)
80 COS GUA NIC PAN 81 DOM GDL HIS JAM MAR PUE SAB STL 82 CLM VEN 83 BOL ECU
Remarks: verticillata agg. Contains New World populations of verticillatus, which is restricted to Western Indian Ocean. L. portoricensis seems to be the oldest available name, because setaceus type comes from La Réunion.
Synonyms: hidden
= Lycopodium portoricense Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 108 (1906)
= Lycopodium tortile Christ; Bull. Soc. bot. Genève 1(5): 235 (1909)
= Urostachys portoricensis (Underw. & F.E.Lloyd) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 163 (1923)
species :
Phlegmariurus pringlei
(Underw. & F.E.Lloyd) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Mexico (Chiapas, Colima, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Oaxaca, Puebla, Sinaloa, Veracruz); Guatemala; El Salvador; Honduras; Nicaragua
79 MEX 80 ELS GUA HON NIC
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia pringlei (Underw. & F.E.Lloyd) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Huperzia tuerckheimii (Maxon) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium fontinaloides var. mexicanum Spring; Mém. Acad. Roy. Sci. Bruxelles 15: 49 (1842)
= Lycopodium pringlei Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 109 (1906)
= Lycopodium tuerckheimii Maxon; Contr. U.S. Nat. Herb. 13(1): 23, t. 1 (1909)
= Urostachys pringlei (Underw. & F.E.Lloyd) Herter ex Nessel; Bärlappgew. 131 (1939)
= Urostachys tuerckheimii (Maxon) Herter; Repert. Spec. Nov. Regni Veg. 19: 163 (1923)
species :
Phlegmariurus proliferus
(Blume) A.R.Field & Bostock; PhytoKeys 20: 46 (2013)
Distribution: peninsular Malaysia (Perak, Pahang); Borneo (Sabah); Sulawesi; Java; Lesser Sunda Isl. (Lombok); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Guadalcanal); New Caledonia
42 BOR JAW LSI MLY MOL NWG SUL 43 SOL 60 NWC
Remarks: hippuris agg. Australian records belong to H. tetrastichoides.
Synonyms: hidden
= Huperzia prolifera (Blume) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium proliferum Blume; Enum. Pl. Javae 2: 265 (1828)
= Urostachys prolifer (Blume) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 77 (1949)
species :
Phlegmariurus pruinosus
(Hieron. & Herter) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Ecuador; Peru
83 ECU PER
Remarks: phlegmaria agg.
Conservation Status: IUCN R
Synonyms: hidden
= Huperzia durissima (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Huperzia pruinosa (Hieron. & Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium durissimum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 52 (1909)
= Lycopodium pruinosum Hieron. & Herter ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 52 (1909)
= Urostachys durissimum (Herter) Herter ex Nessel; Bärlappgew. 240 (1939)
= Urostachys pruinosus (Hieron. & Herter) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 420 (1927)
species :
Phlegmariurus pseudovarius
(Brownlie) Rouhan & A.R.Field; Phytotaxa 497(1): 55 (2021)
Distribution: New Caledonia
60 NWC
Remarks: myrtifolia agg.
Conservation Status: IUCN R
Synonyms: hidden
= Huperzia pseudovaria (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium pseudovarium Brownlie; Fl. Nouv.-Caléd. 3: 23, t. 1, f. 13-15 (1969)
species :
Phlegmariurus pulcherrimus
(Hook. & Grev.) Á.Löve & D.Löve; Taxon 26: 324 (1977)
Distribution: China (Guangxi, W-Yunnan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Manipur, Meghalaya, Sikkim, Uttarakhand, West Bengal, Mizoram, Nagaland); Sri Lanka; Myanmar (Kachin, etc.); Thailand; Vietnam; Borneo (Sabah); Sulawesi
36 CHI TIB 40 BHU IND NEP SRL 41 MYA THA VIE 42 BOR SUL
Remarks: hippuris agg.
Synonyms: hidden
= Huperzia pulcherrima (Wall. ex Hook. & Grev.) Pic.Serm.; Webbia 24: 719 (1970)
= Huperzia pulcherrima (Wall. ex Hook. & Grev.) T.Sen & U.Sen; Fern Gaz. 11(6): 419 (1978), isonym
= Huperzia setacea (Trevis.) Rothm.; Feddes Repert. Spc. Nov. Regn. Veg. 54: 61 (1961), isonym
= Huperzia setacea Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia subulifolia (Wall. ex Hook. & Grev.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia subulifolia var. assamica J.P.Mandal & U.Sen; Biol. Mem. 3(2): 225 (1979)
= Huperzia taiwanensis (Ching) Holub; Folia Geobot. Phytotax. 20(1): 77 (1985)
= Lycopodium pulcherrimum Wall. ex Hook. & Grev.; Icon. Filic. 1(2): t. 38 (1827)
= Lycopodium setaceum Buch.-Ham. ex D.Don; Prodr. Fl. Nepal. 18 (1825) (nom. illeg., non Lycopodium setaceum Lam. (1789))
= Lycopodium setaceum var. pulcherrimum (Wall. ex Hook. & Grev.) C.B.Clarke; Trans. Linn. Soc. Lond. 2, Bot. 1: 590 (1880)
= Lycopodium subulifolium Wall. ex Hook. & Grev.; Icon. Filic. 1: t. 49 (1827) (fide Fraser-Jenkins 2008)
= Phlegmariurus subulifolius (Wall. ex Hook. & Grev.) S.R.Ghosh; Pterid. Fl. E. India 66 (2004), et J. Econ. Taxon. Bot. 33(2): 364 (2009)
= Phlegmariurus taiwanensis Ching; Acta Bot. Yunnan. 4: 124 (1982 (nom. nov. superfl. for Lycopodium pulcherrimum Hook. & Grev.)
= Urostachys amphorae Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 51 (1949 (nom. nov. superfl. for Urostachys pulcherrimus (Hook. & Grev.) Nessel))
= Urostachys pulcherrimus (Wall. ex Hook. & Grev.) Herter ex Nessel; Bärlappgew. 72 (1939)
= Urostachys setaceus (Trevis.) Herter ex Nessel; Arch. Bot. São Paulo 1: 408 (1927)
= Urostachys subulifolius (Wall. ex Hook. & Grev.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 84 (1949)
species :
Phlegmariurus pungentifolius
(Silveira) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Minas Gerais, Rio de Janeiro, São Paulo)
84 BRA
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia pungentifolia (Silveira) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium ouropretanum Christ; Pl. Nov. Mineir. 2: 39 (1900)
= Lycopodium pungentifolium Silveira; Bol. Comm. Geogr. Geol. Est. Minas Gerais 2, 5: 119, t. 4 (1898)
= Urostachys hennebergorum Nessel; Repert. Spec. Nov. Regni Veg. 48: 169, t. 316, f. 1 (1940)
= Urostachys pungentifolius (Silveira) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 394 (1927)
= Urostachys reflexus var. pungentifolius (Silveira) Herter ex Nessel; Bärlappgew. 115 (1939)
species :
Phlegmariurus quadrifariatus
(Bory) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Huperzia aschersonii (Herter) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Huperzia quadrifariata (Bory ex Duperr) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 60 (1944)
= Lycopodium aschersonii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 53 (1909)
= Lycopodium fontinaloides var. coronata Fée; Crypt. Vasc. Brésil 2: 94-95 (1872 publ. 1873)
= Lycopodium quadrangulare Spring; Mart., Fl. Bras. 1(2): 112-113, t. 5, f. 1 (1840)
= Lycopodium quadrifariatum Bory; Dup., Voy. Bot. 1: 245 (1829)
= Urostachys aschersonii (Herter) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1. 422 (1927)
= Urostachys fontinaloides var. coronatus (Fée) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 403 (1927)
= Urostachys quadrangularis (Spring) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 404 (1927)
= Urostachys quadrifariatus (Bory) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 405 (1927)
species :
Phlegmariurus recurvifolius
(Rolleri) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Venezuela (Amazonas, Bolívar, Tachira); Guyana; Brazil (Bahia, Espírito Santo, Minas Gerais)
82 GUY VEN 84 BRA
Synonyms: hidden
= Huperzia recurvifolia Rolleri; Notas Mus. La Plata, Bot. 21(10, f. 1-33): 210 (1989)
species :
Phlegmariurus reflexus
(Lam.) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Mexico (Chiapas, Guerrero, Hidalgo, Jalisco, Mexico State, Oaxaca, Puebla, Veracruz); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Guadeloupe; Dominica; Martinique; St. Vincent; Trinidad; Colombia (Antioquia, Boyacá, Caldas, Caquetá, Cauca, Cesar, Chocó, Cundinamarca, Huila, Magdalena, Meta, Nariño, Norte de Santander, Putumayo, Quindío, Risaralda, Santander, Tolima, Valle); Venezuela (Amazonas, Anzoategui, Aragua, Barinas, Bolívar, Venezuela Distrito Federal, Falcon, Lara, Merida, Miranda, Monagas, Portuguesa, Sucre, Tachira, Trujillo); Guyana (Mt. Roraima); Ecuador; Galapagos; Peru; Bolivia (Beni, Chuquisaca, Cochabamba, La Paz, Santa Cruz); Brazil (Amazonas, Brasilia Distrito Federal, Goiás, Minas Gerais, Mato Grosso do Sul, Pará, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Paraguay; Uruguay (Rivera); Argentina (Salta, Jujuy)
79 MEX 80 COS ELS GUA HON NIC 81 CUB DOM GDL HIS JAM MAR PUE STV TRT 82 CLM GUY VEN 83 BOL ECU GAL PER 84 BRA 85 ARG PAR URU
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia bifida (Humb. & Bonpl. ex Willd.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Huperzia densifolia (Baker) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Huperzia leptodon (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Huperzia mexiae (Copel.) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Huperzia parvifolia (Nessel & Hoehne) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21: 156 (1988)
= Huperzia pearcei (Baker) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia reflexa (Lam.) Rothm.; Feddes Repert. 54: 60 (1944), isonym
= Huperzia reflexa (Lam.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia reflexa var. bifida (Humb. & Bonpl. ex Willd.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia reflexa var. minor (Spring) B.Øllg.; Fl. Ecuador 33: 26 (1988)
= Huperzia rigida (J.F.Gmel.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Huperzia spongiosa (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 157 (1988)
= Lycopodium bifidum Humb. & Bonpl. ex Willd.; Sp. Pl. ed. IV, 5: 53 (1810)
= Lycopodium brutum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 47 (1909)
= Lycopodium densifolium (Baker) Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 106 (1906)
= Lycopodium leptodon (Herter) Maxon ex C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium mexiae Copel.; Univ. Calif. Publ. Bot. 19: 294, t. 47 (1941)
= Lycopodium pearcei Baker; Handb. Fern-Allies 14 (1887)
= Lycopodium reflexum Lam.; Encycl. 3: 653 (1792)
= Lycopodium reflexum var. brevissimum Christ; Pittier, Prim. Fl. Costaric. 3: 56 (1901)
= Lycopodium reflexum var. caesium Vareschi; Fl. Venezuel. 1(1): 31 (1968), nom. inval. (no type indicated)
= Lycopodium reflexum var. densifolium Baker; Handb. Fern-Allies 11 (1887)
= Lycopodium reflexum var. intermedium Sodiro; Crypt. Vasc. Quit. 569 (1893)
= Lycopodium reflexum var. majus Spring; Mém. Acad. Roy. Sci. Bruxelles 15: 26 (1842)
= Lycopodium reflexum var. minus Spring; Mém. Acad. Roy. Sci. Bruxelles 15: 26 (1842)
= Lycopodium reflexum var. rigidum (J.F.Gmel.) Proctor; Rhodora 68(no. 776): 464 (1966)
= Lycopodium reflexum var. rupestre Hassl.; Trab. Inst. Bot. Farmacol. no. 45: 94 (1928)
= Lycopodium reflexum var. udum Christ; Denkschr. Kaiserl. Akad. Wiss., Wien. Math.-Naturwiss. Kl. 79: 57 (1908)
= Lycopodium reversum C.Presl; Reliq. Haenk. 1(1): 82 (1825)
= Lycopodium rigidum J.F.Gmel.; Syst. Nat. II (Syst. Veg.1(2): 128) (1791)
= Lycopodium rigidum Sw.; J. Bot. (Schrader) 1800(2): 114 (1801) (Syn. Fil. 176, 400. 1806) (non J. F. Gmel. (1791))
= Lycopodium spongiosum Rolleri; Rev. Mus. La Plata (N. S.) Bot. 14(87): 6, f. 3-4 (1985)
= Lycopodium squarrosum Sw.; Prodr. 137 (1788) (non G. Forst. 1786, nec Lam. (1792))
= Lycopodium stellae-polaris (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Phlegmariurus pearcei (Baker) B.Øllg.; Phytotaxa 57: 17 (2012)
= Phlegmariurus reflexus var. minor (Spring) B.Øllg.; Rodriguésia 63(2): 481 (2012)
= Plananthus reflexus (Lam.) P.Beauv.; Prodr. Aethéogam. 100 (1805)
= Plananthus rigidus (J.F.Gmel.) P.Beauv.; Prodr. Aethéogam. 111 (1805)
= Plananthus squarrosus (Sw., Lam. & J.F.Gmel.) P.Beauv.; Prodr. Aethéogam. 112 (1805) nom. confus., combination based on different basionyms
= Urostachys bifidus (Humb. & Bonpl. ex Willd.) Herter ex Nessel; Bärlappgew. 110, t. 24, f. 4 (1939)
= Urostachys brutus (Herter) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 162 (1923)
= Urostachys densifolius (Baker) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 58 (1949)
= Urostachys jergii Nessel; Repert. Spec. Nov. Regni Veg. 39: 70, t. 194a (1935)
= Urostachys leptodon Maxon ex Herter; Revista Sudamer. Bot. 10: 120 (1953 publ. 1954)
= Urostachys parvifolius (Raddi) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 2: 395 (1927)
= Urostachys pearcei (Baker) Herter ex Nessel; Bärlappgew. 73 (1939)
= Urostachys reflexus (Lam.) Herter; Beih. Bot. Centralbl. 39: 249 (1922)
= Urostachys reflexus var. aquaticus Nessel & Hoehne; Arch. Bot. São Paulo 1: 396 (1927)
= Urostachys reflexus var. brutus (Herter) Herter ex Nessel; Bärlappgew. 116 (1939)
= Urostachys reflexus var. densifolius (Baker) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1(4): 394 (1927)
= Urostachys reflexus var. majus (Spring) Herter ex Nessel; Bärlappgew. 115 (1939)
= Urostachys reflexus var. minor (Spring) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1(4): 393 (1927)
= Urostachys reflexus var. parvifolius (Raddi) Herter ex Nessel; Bärlappgew. 114 (1939)
= Urostachys reflexus var. schaffneri (Christ) Nessel; Bärlappgew. 115 (1939)
= Urostachys reversus (C.Presl) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 78 (1949), Alt. title: Index Lycopodiarum 78 (1949)
= Urostachys rigidus (J.F.Gmel.) Herter; Repert. Spec. Nov. Regni Veg. 19: 163 (1923)
= Urostachys rigidus var. maximus Herter ex Nessel; Bärlappgew. 108 (1939)
= Urostachys stellae-polaris Herter; Revista Sudamer. Bot. 10: 121 (1954)
species :
Phlegmariurus regnellii
(Maxon) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Minas Gerais, Rio de Janeiro)
84 BRA
Remarks: brongniartii agg.
Conservation Status: IUCN R
Synonyms: hidden
= Huperzia regnellii (Maxon) B.Øllg. & P.G.Windisch; Bradea 5: 16 (1987)
= Lycopodium regnellii Maxon; Contr. U.S. Natl. Herb. 17(4): 424, t. 23 (1914)
= Urostachys regnellii (Maxon) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 381 (1927)
species :
Phlegmariurus ribourtii
(Herter) A.R.Field & Bostock; PhytoKeys 20: 46 (2013)
Distribution: +Cook Isl. (+Rarotonga); Society Isl. (Tahiti)
61 COO SCI
Remarks: obtusifolia agg.
Synonyms: hidden
= Huperzia ribourtii (Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium ribourtii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 53 (1909)
= Urostachys ribourtii (Herter) Herter ex Nessel; Bärlappgew. 250 (1939)
species :
Phlegmariurus riobambensis
(Nessel) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cundinamarca, Norte de Santander, Santander); Venezuela (Lara, Tachira, Trujillo)
82 CLM VEN
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia riobambensis (Nessel) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium castoris (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys affinis var. riobambensis Nessel; (Bärlappgew. 98 (1939) (Revista Sudamer. Bot. 6(5-6): 162, t. 10, f. 38 (1940) (nom. inval.)
= Urostachys castoris Herter; Revista Sudamer. Bot. 10: 111 (1953 publ. 1954)
= Urostachys riobambensis (Nessel) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 79 (1949)
species :
Phlegmariurus robustus
(Klotzsch) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Venezuela (Amazonas, Bolívar)
82 VEN
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia phelpsii (Vareschi) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia robusta (Klotzsch) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium phelpsii Vareschi; Acta Bot. Venez. 1: 84 (1966)
= Lycopodium robustum Klotzsch; Linnaea 18: 518 (1844)
= Urostachys robustus (Klotzsch) Herter ex Nessel; Bärlappgew. 212 (1939)
species :
Phlegmariurus rosenstockianus
(Herter) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Colombia (Antioquia, Cauca, Nariño, Cundinamarca, Norte de Santander); Ecuador; Peru
82 CLM 83 ECU PER
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia rosenstockiana (Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Huperzia socratis (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21(100): 157 (1988)
= Lycopodium cassandrae (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium rosenstockianum Herter; Hedwigia 49: 90 (1909)
= Lycopodium socratis (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys cassandrae Herter; Revista Sudamer. Bot. 10: 116 (1953 publ. 1954)
= Urostachys rosenstockianus (Herter) Herter ex Nessel; Bärlappgew. 174 (1939)
= Urostachys socratis Herter; Revista Sudamer. Bot. 10: 117 (1953 publ. 1954)
species :
Phlegmariurus rostrifolius
(Silveira) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Minas Gerais)
84 BRA
Synonyms: hidden
= Huperzia rostrifolia (Silveira) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium rostrifolium Silveira; Bol. Comm. Geogr. Geol. Est. Minas Gerais 2, 5: 118, t. 2 (1898)
= Urostachys rostrifolius (Silveira) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 379 (1927)
species :
Phlegmariurus ruber
(Cham. & Schltdl.) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (?Bahia, Minas Gerais, ?Rio de Janeiro)
84 BRA
Synonyms: hidden
= Huperzia rubra (Cham. & Schltdl.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
= Lycopodium rubrum Cham. & Schltdl.; Linnaea 8: 389 (1833)
= Urostachys ruber (Cham. & Schltdl.) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 381 (1927)
species :
Phlegmariurus rubricus
(Herter) A.R.Field & Bostock; PhytoKeys 20: 46 (2013)
Distribution: Madagascar
29 MDG
Remarks: ophioglossoides agg.
Synonyms: hidden
= Huperzia rubrica (Herter) Tardieu; Adansonia ser. 2 10: 18 (1970)
= Lycopodium ophioglossoides var. rubricaule C.Chr.; Dansk Bot. Ark. 7: 192, t. 77, f. 9-14 (1932)
= Urostachys rubricus Herter; Revista Sudamer. Bot. 8: 25 (1949)
species :
Phlegmariurus rufescens
(Hook.) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Colombia (Arauca, Boyacá, Caldas, Cauca, Chocó, Cundinamarca, Magdalena, Nariño, Tolima); Venezuela (Merida, Trujillo); Ecuador
82 CLM VEN 83 ECU
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia rufescens (Hook.) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 60 (1944), isonym
= Huperzia rufescens (Hook.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1875)
= Lycopodium mirum Vareschi; Acta Biol. Venez. 2: 317, f. 23 (1958)
= Lycopodium rufescens Hook.; Hooker´s Icon. Pl. 1: tab 36 (1837)
= Urostachys rufescens (Hook.) Herter ex Nessel; Bärlappgew. 103 (1939)
species :
Phlegmariurus rupicola
(Alderw.) A.R.Field & Bostock; PhytoKeys 20: 46 (2013)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Remarks: squarrosa agg.
Synonyms: hidden
= Huperzia rupicola (Alderw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium rupicola Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 39 (1914)
= Urostachys rupicolus (Alderw.) Herter ex Nessel; Bärlappgew. 203 (1939)
species :
Phlegmariurus sagasteguianus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Ecuador; Peru
83 ECU PER
Synonyms: hidden
= Huperzia sagasteguiana B.Øllg.; Fieldiana, Bot. n. s. 34: 37 (1994)
species :
Phlegmariurus salvinioides
(Herter) Ching; Acta Bot. Yunnan. 4: 122 (1982)
Distribution: Japan; Taiwan; ?Vietnam; Borneo (Sabah); Palawan; Philippines
38 JAP TAI 42 BOR PHI PLW
Remarks: nummulariifolia agg.
Synonyms: hidden
= Huperzia edanoi (Copel.) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Huperzia salvinioides (Herter) Holub; Folia Geobot. Phytotax. 20. 76 (1985)
= Lycopodium edanoi Copel.; Philipp. J. Sci. 46: 209 (1931)
= Lycopodium salvinioides (Herter) Tagawa; Acta Phytotax. Geobot. 8: 20 (1939)
= Urostachys edanoi (Copel.) Nessel; Bärlappgew. 230 (1939)
= Urostachys formosanus Herter ex Nessel; Bärlappgew. 229 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 167, excl. fig. (1940)
= Urostachys salvinioides Herter; Bot. Arch. 3: 18 (1923)
species :
Phlegmariurus samoanus
(Herter ex Nessel) A.R.Field & Bostock; PhytoKeys 20: 47 (2013)
Distribution: [Western] Samoa
60 SAM
Remarks: hippuris agg.
Synonyms: hidden
= Huperzia samoana (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Urostachys samoanus Herter ex Nessel; Bärlappgew. 125, f. 29e 5 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 163 (1940)
species :
Phlegmariurus sarmentosus
(Spring) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Colombia (Antioquia, Choco); Ecuador; Peru
82 CLM 83 ECU PER
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia sarmentosa (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium sarmentosum Spring; Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 13 (1849)
= Lycopodium sarmentosum var. grenadense Hook. ex Nessel; Bärlappgew. 159 (1939)
= Urostachys sarmentosus (Spring) Herter; Repert. Spec. Nov. Regni veg. 19: 165 (1923)
species :
Phlegmariurus saururus
(Lam.) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Cameroon; D.R.Congo [Zaire]; Rwanda; Ethiopia; Uganda; Kenya; Tanzania; Malawi; E-Zimbabwe; South Africa (KwaZulu-Natal, W-Cape Prov., E-Cape Prov., Free State, Mpulamanga); Lesotho; Madagascar; Comores; Mauritius; La Réunion
23 CMN RWA ZAI 24 ETH 25 KEN TAN UGA 26 MLW ZIM 27 LES SAF 29 COM MAU MDG REU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia saururus (Lam.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1875)
= Lycopodium saururus Lam.; Encycl. 3: 653 (1789 (1792))
= Plananthus saururus (Lam.) P.Beauv.; Prodr. Aethéogam. 111 (1805)
= Urostachys saururus (Lam.) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 162 (1923)
species :
Phlegmariurus scabridus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Ecuador
83 ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia scabrida B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 60, f. 8A (1988)
species :
Phlegmariurus schlechteri
(E.Pritz.) A.R.Field & Bostock; PhytoKeys 20: 47 (2013)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Huperzia schlechteri (E.Pritz.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium schlechteri E.Pritz.; Schlechter, Bot. Jahrb. Syst. 39: 14 (1906)
= Urostachys schlechteri (E.Pritz.) Herter ex Nessel; Bärlappgew. 207 (1939)
species :
Phlegmariurus schlimii
(Herter) B.Øllg.; Phytotaxa 57: 18 (2012)
Distribution: Colombia (Boyacá, Caldas, Cauca, Magdalena, Risaralda)
82 CLM
Remarks: saururus agg.
Synonyms: hidden
= Huperzia schlimii (Herter) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium schlimii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909)
= Urostachys schlimii (Herter) Herter ex Nessel; Bärlappgew. 91 (1939)
species :
Phlegmariurus schmidtchenii
(Hieron.) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Colombia (Tolima, Caldas, ?Cauca, Cundinamarca)
82 CLM
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia goudotii (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
= Huperzia schmidtchenii (Hieron.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium goudotii Herter; Bot. Jahrb. Syst. 43. Beibl. 98: 47 (1909)
= Lycopodium schmidtchenii Hieron.; Bot. Jahrb. Syst. 34: 570 (1905)
= Phlegmariurus goudotii (Herter) B.Øllg.; Phytotaxa 57: 15 (2012)
= Urostachys goudotii (Herter) Herter ex Nessel; Bärlappgew. 106 (1939)
= Urostachys schmidtchenii (Hieron.) Herter ex Nessel; Bärlappgew. 103 (1939)
species :
Phlegmariurus sellifolius
(B.Øllg.) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Colombia (Cauca, Huila, Narino); Ecuador; Peru
82 CLM 83 ECU PER
Remarks: brevifolia agg.
Synonyms: hidden
= Urostachys compactus f. minor Herter; Revista Sudamer. Bot. 10: 119-120 (1954)
= Huperzia sellifolia B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 68, f. 12C (1988)
species :
Phlegmariurus sellowianus
(Herter) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Espírito Santo, Minas Gerais, Rio de Janeiro, Santa Catarina, São Paulo)
84 BRA
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia sellowiana (Herter) B.Øllg.; Opera Bot. 92: 169 (1987)
= Huperzia ulei (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium brasilianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 44 (1909)
= Lycopodium sellowianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 44 (1909)
= Urostachys brasilianus (Herter) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 386 (1927)
= Urostachys sellowianus (Herter) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 385 (1927)
= Urostachys ulei Herter; Repert. Spec. Nov. Regni Veg. 19: 162 (1923)
species :
Phlegmariurus serpentiformis
(Herter) B.Øllg.; Phytotaxa 277(3): 269 (2016)
Distribution: Colombia (Cundinamarca, Meta)
82 CLM
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia serpentiformis (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Lycopodium serpentiforme Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 46 (1909)
= Urostachys serpentiformis (Herter) Herter ex Nessel; Bärlappgew. 104 (1939)
species :
Phlegmariurus setifolius
(Alderw.) A.R.Field & Bostock; PhytoKeys 20: 47 (2013)
Distribution: Borneo
42 BOR
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia setifolia (Alderw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium setifolium Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 40 (1914)
= Urostachys setifolius (Alderw.) Herter ex Nessel; Bärlappgew. 233 (1939)
species :
Phlegmariurus shangsiensis
C.Y.Yang; Acta Phytotax. Sin. 22(1): 87-88, f. 1 (1984)
Distribution: China (Guangxi)
36 CHI
Synonyms: hidden
= Huperzia shangsiensis (C.Y.Yang) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
species :
Phlegmariurus shingianus
R.H.Jiang & X.C.Zhang; Turczaninowia 25(1): 131 (2022)
Distribution: China (Guangxi)
36 CHI
species :
Phlegmariurus sieberianus
(Spring) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: ?Hispaniola (?Haiti); St. Kitts; Guadeloupe; Martinique; St. Vincent
81 GDL MAR STK STV
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia sieberiana (Spring) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 60 (1944), isonym
= Huperzia sieberiana (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium sieberianum Spring; Flora 21: 153 (1838)
= Lycopodium squarrosum Lam.; Encycl. 3. 653 (1792) (non G. Forst. 1786, nec Sw. (1788))
= Urostachys sieberianus (Spring) Herter; Repert. Spec. Nov. Regni Veg. 19: 162 (1923)
species :
Phlegmariurus sieboldii
(Miq.) Ching; Acta Bot. Yunnan. 4: 121 (1982)
Distribution: South Korea (Cheju Isl.); Japan; Ryukyu Isl.; Taiwan
38 JAP KOR NNS TAI
Remarks: cancellata agg.
Synonyms: hidden
= Huperzia sieboldii (Miq.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium sieboldii Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 184 (1867)
= Urostachys sieboldii (Miq.) Herter ex Nessel; Bärlappgew. 138 (1939)
species :
Phlegmariurus silveirae
(Herter ex Nessel & Hoehne) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Minas Gerais, Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
84 BRA
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia silveirae (Herter ex Nessel & Hoehne) B.Øllg. & P.G.Windisch; Bradea 5: 17 (1987)
= Urostachys brasilianus var. schwackeanus (Herter ex Nessel & Hoehne) Herter ex Nessel; Bärlappgew. 85 (1939)
= Urostachys brasilianus var. silveirae (Nessel) Nessel; Bärlappgew. 84 (1939)
= Urostachys portoanus Nessel; Repert. Spec. Nov. Regni Veg. 39: 70, t. 194b (1935)
= Urostachys schwackeanus Herter ex Nessel & Hoehne; Arch. Bot. Est. São Paulo 1: 388 (1927)
= Urostachys silveirae Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 386 (1927)
species :
Phlegmariurus silverstonei
B.Øllg.; Phytotaxa 277(3): 269 (2016)
Distribution: Colombia (Choco)
82 CLM
Remarks: known only from type
species :
Phlegmariurus sintenisii
(Herter) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Hispaniola (Haiti, Dominican Republic); Puerto Rico
81 HIS PUE
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia sintenisii (Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium sintenisii (Herter) Maxon ex C.Chr.; Kongl. Vetensk. Acad. Handl. ser. 3, 16(2): 91 (1937)
= Urostachys sintenisii Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 161 (1923)
species :
Phlegmariurus societensis
(J.W.Moore) A.R.Field; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Society Isl. (Tahiti)
61 SCI
Synonyms: hidden
= Lycopodium societense J.W.Moore; Bull. Bernice P. Bishop Mus. 102: 9-12 (1933)
= Huperzia societensis (J.W.Moore) Govaerts; Skvortsovia 4(3): 83 (2018)
species :
Phlegmariurus sooanus
Lawalrée; Acta Bot. Acad. Sci. Hung. 19: 195, f. 1 (1973)
Distribution: D.R.Congo [Zaire]
23 ZAI
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia sooana (Lawalrée) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
species :
Phlegmariurus sphagnicola
B.Øllg.; Phytotaxa 277(3): 270 (2016)
Distribution: Colombia (Magdalena)
82 CLM
species :
Phlegmariurus squarrosus
(G.Forst.) Á.Löve & D.Löve; Taxon 26: 324 (1977)
Distribution: Madagascar; Mauritius; La Réunion; Seychelles; China (W-Guangxi, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Karnataka, Kerala, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Sikkim, Tripura, West Bengal); Sri Lanka; Bangladesh; Andaman Isl.; ?Nicobar Isl.; Myanmar (Kachin, etc.); Thailand (widespread); Laos; Cambodia; Vietnam; peninsular Malaysia (Kedah, Kelantan, Perak, Pahang); +Singapore; Sumatra; Borneo (Sabah, Sarawak, Mt. Kinabalu, Sabah); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Lombok); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea, d´Entrecasteaux Arch.: Normanby Isl., Louisiade Arch.: Woodlark Isl.); Bismarck Arch. (Admiralty Isl.); Solomon Isl. (Choiseul, Guadalcanal, Makira, New Georgia, Malaita, Rennell, Vangunu, Vella Lavella); Australia (NE-Queensland); New Caledonia; Tonga (Late Isl., Tafahi); Fiji; American Samoa (Tutuila, Ta´u); [Western] Samoa (Savai´i, ´Upolu); Cook Isl. (Rarotonga); Society Isl. (Tahiti); Austral Isl. (Rapa Iti, Tubuai); Palau (Babeldaob); Fed. States of Micronesia (Pohnpei, Kosrae)
29 MAU MDG REU SEY 36 CHI TIB 38 TAI 40 BAN BHU IND NEP SRL 41 AND CBD LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SNG SUL SUM 43 BIS SOL 50 AUS 60 ASA FIJ NWC SAM TON 61 AUI COO SCI 62 MIC PLU
Remarks: squarrosa agg. Status of P. acutifolius (Mauritius) unclear
Synonyms: hidden
= Huperzia acutifolia (Desv ex Poir.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Huperzia epiceifolia (Desv. ex Poir.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia forsteri Poir.; nom. illeg. superfl.
= Huperzia hookeri (Hook. & Grev.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Huperzia squarrosa (G.Forst.) Rothm.; Feddes Repert. 54: 62 (1944), isonym
= Huperzia squarrosa (G.Forst.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
= Huperzia ulicifolia (Sw.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium acutifolium Desv. ex Poir.; Lam., Encycl. suppl. 3: 559 (1813 publ. 1814)
= Lycopodium epiceifolium Desv. ex Poir.; Lam., Encycl. suppl. 3: 559 (1813 publ. 1814)
= Lycopodium forsteri Poir.; Encycl. suppl. 3: 545 (1813 (1814)) (nom. superfl.)
= Lycopodium hookeri Wall. ex Hook. & Grev.; Icon. Filic. 2: t. 183 (1830)
= Lycopodium lohitense D.D.Pant & P.S.Pandey; Phyta Monogr. 3: 26 (1985), as ´lohitensis´, synonym fide Fraser-Jenkins et al. (2015)
= Lycopodium madagascariense Desv.; (nom. inval.)
= Lycopodium pseudosquarrosum Pamp.; Bull. R. Soc. Tosc. Orticult. III, 13: 99, t. 2 (1908)
= Lycopodium remoganense Hayata; Icon. Pl. Formosan. 10: 72 (1921)
= Lycopodium squarrosum G.Forst.; Fl. Ins. Austr. 479 (1786)
= Lycopodium ulicifolium Vent. ex Sw.; Syn. Fil. 177 (1806)
= Phlegmariurus acutifolius (Desv. ex Poir.) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 644 (2015)
= Phlegmariurus ulicifolius (Vent. ex Sw.) S.R.Ghosh; Pterid. Fl. E. India: 64 (2004)
= Plananthus squarrosa (G.Forst.) P.Beauv.; Prodr. Aethéogam. 112 (1805)
= Urostachys epiceifolius (Desv. ex Poir.) Herter ex Nessel; Bärlappgew. 202 (1939)
= Urostachys epiceifolius var. acutifolius (Desv.) Herter ex Nessel; Bärlappgew. 292 (1939)
= Urostachys madagascariensis (Desv. ex Nessel) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 68 (1949), Alt. title: Index Lycopodiarum 68 (1949)
= Urostachys madagascariensis (Nessel) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 68 (1949)
= Urostachys squarrosus (G.Forst.) Herter; Bot. Arch. 3: 14 (1923)
= Urostachys squarrosus f. madagascariensis Desv. ex Nessel; Revista Sudamericana de Botánica 6,5/6: 165, 166, t. 12, fig. 60 (1940)
= Urostachys squarrosus var. intermedius Herter
= Urostachys squarrosus var. proliferus (Blume) Herter ex Nessel; Bärlappgew. 200 (1939)
= Urostachys ulicifolius (Sw.) Herter ex Nessel; Bärlappgew. 204 (1939)
species :
Phlegmariurus staudtii
(Nessel) A.R.Field & Bostock; PhytoKeys 20: 47 (2013)
Distribution: Ivory Coast; Ghana; Togo; Nigeria; Cameroon; Gabon; Central African Republic; D.R.Congo [Zaire] (Tshopo); Uganda; Angola
22 GHA IVO NGA TOG 23 CAF CMN GAB ZAI 25 UGA 26 ANG
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia staudtii (Nessel) Pic.Serm.; Webbia 23: 163 (1968)
= Lycopodium staudtii (Nessel) C.D.Adams & Alston; Bull. Brit. Mus. (Nat. Hist.) Bot. 1(6): 183 (1955)
= Urostachys adolfi-friedericii Herter ex Nessel; Bärlappgew. 226 (1939) (nom. inval., without Latin description)
= Urostachys staudtii Nessel; Repert. Spec. Nov. Regni Veg. 36: 189, t. 175 (1934)
species :
Phlegmariurus stemmermanniae
A.C.Medeiros & Wagner; Amer. Fern J. 86(3): 89-97 (1996)
Distribution: Hawaii
63 HAW
Synonyms: hidden
= Huperzia stemmermanniae (A.C.Medeiros & W.H.Wagner) Kartesz; Synthesis N. Amer. Fl., version 1 Nomencl. Innov.: no. 14 (1999)
species :
Phlegmariurus stephani
B.Øllg.; Phytotaxa 334(3): 283 (2018)
Distribution: Peru; Bolivia (La Paz, Cochabamba)
83 BOL PER
species :
Phlegmariurus strictus
(Baker) A.R.Field & Bostock; PhytoKeys 20: 47 (2013)
Distribution: Madagascar
29 MDG
Remarks: included by Roux (2009) in P. gnidioides
Synonyms: hidden
= Huperzia stricta (Baker) Tardieu; Adansonia ser. 2 10: 20 (1970)
= Lycopodium strictum Baker; J. Bot. 20: 271. (n. s., vol. 11) (1882)
= Urostachys strictus (Baker) Herter ex Nessel; Bärlappgew. 191, t. 50 (1939)
species :
Phlegmariurus subfalciformis
(Alderw.) A.R.Field & Bostock; PhytoKeys 20: 47 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: phlegmarioides agg.
Synonyms: hidden
= Huperzia subfalciformis (Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium subfalciforme Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 44 (1918)
= Phlegmariurus subfalciformis (Alderw.) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015), isonym
= Urostachys subfalciformis (Alderw.) Herter ex Nessel; Bärlappgew. 227 (1939)
species :
Phlegmariurus subtubulosus
(Herter) comb.ined.
Distribution: Hispaniola (Haiti, Dominican Republic)
81 HIS
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia subtubulosa (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Urostachys subtubulosus Herter; Repert. Spec. Nov. Regni Veg. 19: 166 (1923)
species :
Phlegmariurus subulatus
(Desv. ex Poir.) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Honduras; Costa Rica; Panama; Colombia (Antioquia, Caldas, Cauca, Cundinamarca, Huila, Nariño, Putumayo, Quindío, Risaralda, Tolima, Valle); Ecuador; Peru; Bolivia (Cochabamba, La Paz, Tarija); Argentina (Salta, Tucuman)
80 COS HON PAN 82 CLM 83 BOL ECU PER 85 ARG
Remarks: myrsinites agg.
Synonyms: hidden
= Huperzia subulata (Desv. ex Poir.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium ewanii (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium polymorphum Willd. ex Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 71 (1842)
= Lycopodium subulatum Desv. ex Poir.; Encycl. suppl. 3: 544. (1813 publ. 1814)
= Lycopodium subulatum var. obtusifolium Kuhn ex Urb.; Bot. Jahrb. Syst. 24(1): 149 (1897)
= Lycopodium subulatum var. rubens Spring; Mém. Acad. Roy. Sci. Belgique 24: 32 (1849)
= Urostachys ewanii Herter; Revista Sudamer. Bot. 10: 126 (1953 publ. 1954)
= Urostachys subulatus (Desv. ex Poir.) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1(4): 420 (1927)
= Urostachys subulatus var. obtusifolius (Kuhn) Nessel; Bärlappgew. 242 (1939)
= Urostachys subulatus var. paradoxus (Humb. & Bonpl.) Nessel; Bärlappgew. 243 (1939)
= Urostachys subulatus var. polymorphus (Willd. ex Spring) Nessel; Bärlappgew. 243 (1939)
species :
Phlegmariurus tafaensis
(Nessel) A.R.Field & C.W.Chen; Fern Gaz. 21(7): 307 (2022)
Distribution: New Guinea (Papua New Guinea); Solomon Isl. (Bougainville, Guadalcanal, Santa Ysabel, Malaita, Kolombangara, Makira, New Georgia, Vella Lavella)
42 NWG 43 SOL
Synonyms: hidden
= Huperzia brassii (Copel.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Lycopodium brassii (Nessel) Alston; J. Bot. 77: 290 (1939) (non Copel. (1929))
= Lycopodium brassii Copel.; J. Arnold Arbor. 10: 174 (1929)
= Phlegmariurus brassii (Copel.) A.R.Field & Bostock; PhytoKeys 20: 37 (2013)
= Urostachys arietis Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 51 (1949) (nom. nov. superfl.)
= Urostachys brassii (Copel.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 54 (1949), isonym
= Urostachys brassii (Copel.) Nessel; Repert. Spec. Nov. Regni Veg. 39: 62, t. 190a (1935)
= Urostachys tafaensis Nessel; Repert. Spec. Nov. Revni Veg. 39: 61 (1935)
species :
Phlegmariurus talamancanus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Costa Rica; Panama
80 COS PAN
Synonyms: hidden
= Huperzia talamancana B.Øllg.; Novon 3(1): 67, f. 2 (1993)
species :
Phlegmariurus talamauanus
(Alderw.) A.R.Field & Bostock; PhytoKeys 20: 48 (2013)
Distribution: Sumatra
42 SUM
Remarks: phyllantha agg.
Synonyms: hidden
= Huperzia talamauana (Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium talamauanum Alderw.; Bull. Jard. Bot. Buitenz. II,28: 45 (1918)
= Urostachys talamauanus (Alderw.) Herter ex Nessel; Bärlappgew. 212 (1939)
species :
Phlegmariurus talpiphilus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Ecuador; Bolivia (Cochabamba, La Paz, Santa Cruz)
83 BOL ECU
Remarks: saururus agg.
Synonyms: hidden
= Huperzia talpiphila B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 44, f. 9A (1988)
species :
Phlegmariurus tardieuae
(Herter) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Madagascar; ?Mauritius; La Réunion; Seychelles
29 MDG REU SEY
Synonyms: hidden
= Huperzia phlegmaria var. tardieuae (Herter) Tardieu; Adansonia ser. 2 10: 18 (1970)
= Huperzia tardieuae (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Urostachys tardieuae Herter; Notul. Syst. (Paris) 15: 355 (1958)
species :
Phlegmariurus tauri
(Herter) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Sulawesi; Java; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.)
42 JAW MOL NWG SUL 43 BIS
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia tauri (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium apiculatum Spring; Miq., Pl. Jungh. 3: 272 (1854) (non Poir. 1814)
= Lycopodium tauri (Herter) M.Kato; Acta Phytotax. Geobot. 39(4-6): 137 (1988)
= Urostachys apiculatus (Spring) Herter ex Nessel; Bärlappgew. 227 (1939)
= Urostachys tauri Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 85 (1949)
species :
Phlegmariurus taxifolius
(Sw.) Á.Löve & D.Löve; Taxon 26(2-3): 324 (1977)
Distribution: Mexico (Chiapas, Ciudad de Mexico, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Oaxaca, Puebla, San Luis Potosi, Sinaloa, Tamaulipas, Veracruz); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Saba; Nevis; St. Kitts; Montserrat; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Grenada; Trinidad; Colombia (Antioquia, Cauca, Cesar, Cundinamarca, Magdalena, Nariño, Santander, Valle, Caldas, Choco, Guajira, Norte de Santander, Risaralda); Venezuela (Amazonas, Anzoategui, Aragua, Bolívar, Carabobo, Venezuela Distrito Federal, Falcon, Lara, Merida, Miranda, Monagas, Nueva Esparta, Portuguesa, Sucre, Zulia); Guyana; Surinam; French Guiana; Ecuador; Galapagos; Peru; Bolivia (Cochabamba, La Paz); Brazil (Amapá, Ceará, Paraná, Rio de Janeiro, Roraima, Santa Catarina, São Paulo)
79 MEX 80 BLZ COS ELS GUA HON NIC PAN 81 CUB DOM GDL GRE HIS JAM MAR MTS PUE SAB STK STL STV TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU GAL PER 84 BRA
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia brauseana (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21(100): 155 (1988)
= Huperzia cubana (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Huperzia passerinoides (Kunth) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia passerinoides var. nitens (Schltdl. & Cham.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia rubiginosa Nessel
= Huperzia taxifolia (Sw.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Huperzia taxifolia var. herminieri (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17(3): 248, no. 17 (1875)
= Lycopodium americanum Desv.; (herb. Vaillant)
= Lycopodium brauseanum Herter; Symb. Ant. 7(2): 165 (1912)
= Lycopodium cubanum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 50 (1909)
= Lycopodium herminieri Spring; Bull. Acad. Roy. Soc. Bruxelles 8514 (1841)
= Lycopodium linifolium var. spongiosum Jos.Tub. ex Nessel; Bärlappgew. 168 (1939)
= Lycopodium nitens Schltdl. & Cham.; Linnaea 5: 623 (1830)
= Lycopodium passerinoides Kunth; Nov. Gen. Sp. [H. B. K.] 1: 41 (1815 publ. 1816)
= Lycopodium schwendeneri Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 50 (1909)
= Lycopodium taxifolium Sw.; Nov. Gen. Sp. seu Prodromus 138 (1788)
= Lycopodium taxifolium var. nitens (Schltdl. & Cham.) Poepp. ex Nessel & Hoehne; Arch. Bot. São Paulo 2: 388 (1927)
= Lycopodium taxifolium var. passerinoides (Kunth) Baker; Handb. Fern-Allies 16 (1887)
= Lycopodium taxifolium var. recurvatum H.Karst. ex Nessel; Bärlappgew. 94 (1939)
= Plananthus taxifolius (Sw.) P.Beauv.; Prodr. Aethéogam. 112 (1805)
= Urostachys brauseanus (Herter) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 162 (1923)
= Urostachys bruelkei Nessel; Repert. Spec. Nov. Regni Veg. 36(12-15): 184, t. 173 (1934)
= Urostachys cubanus (Herter) Herter; Repert. Spec. Nov. Regni Veg. 19: 165 (1923)
= Urostachys herminieri (Spring) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 162 (1923)
= Urostachys nitens (Schltdl. & Cham.) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 164 (1923)
= Urostachys passerinoides (Kunth) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1(4): 417 (1927)
= Urostachys passerinoides var. nitens (Schltdl. & Cham.) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1(4): 418 (1927)
= Urostachys rubiginosus Herter ex Nessel; Revista Sudamer. Bot. 6(5-6): 162, t. 8, f. 36 (1940)
= Urostachys schwendeneri (Herter) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 165 (1923)
= Urostachys schwendeneri var. nitens (Schltdl. & Cham.) Herter ex Nessel; Bärlappgew. 166 (1939)
= Urostachys schwendeneri var. spongiosus (Jos Tub) Nessel; Bärlappgew. 168 (1939)
= Urostachys spongiosus (Nessel) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 83 (1949)
= Urostachys taxifolius (Sm.) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 162 (1923)
= Urostachys taxifolius var. herminieri (Spring) Herter ex Nessel; Bärlappgew. 173 (1939)
= Urostachys taxifolius var. nitens Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 419 (1927)
= Urostachys taxifolius var. passerinoides (Kunth) Herter ex Nessel; Bärlappgew. 172 (1939)
species :
Phlegmariurus tenuicaulis
(Underw. & F.E.Lloyd) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Puerto Rico; Guadeloupe; Dominica; Guyana
81 DOM GDL PUE 82 GUY
Remarks: linifolia agg.
Synonyms: hidden
= Huperzia tenuicaulis (Underw. & F.E.Lloyd) B.Øllg.; Opera Bot. 92: 169 (1987)
= Lycopodium tenuicaule Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 113 (1906)
= Urostachys tenuicaulis (Underw. & F.E.Lloyd) Herter; Repert. Spec. Nov. Regni Veg. 19: 165 (1923)
species :
Phlegmariurus tenuis
(Humb. & Bonpl. ex Willd.) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Costa Rica; Panama; Colombia (Cauca, Narino, Norte de Santander); Venezuela (Merida); Ecuador; N-Peru; Bolivia (Cochabamba, La Paz)
80 COS PAN 82 CLM VEN 83 BOL ECU PER
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia tenuis (Humb. & Bonpl. ex Willd.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium tenue Humb. & Bonpl. ex Willd.; Sp. Pl. ed. IV, 5: 55 (1810)
= Urostachys pax-hominibus Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 74 (1949) (nom. superfl. for Lycopodium tenue Willd.)
= Urostachys tenuis (Humb. & Bonpl. ex Willd.) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1(4): 401 (1927)
species :
Phlegmariurus terrae-guilelmi
(Herter) A.R.Field & Bostock; PhytoKeys 20: 48 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia terrae-guilelmi (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium terrae-guilelmi Herter; Bot. Jahrb. Syst. 54: 229 (1916)
= Urostachys terrae-guilelmi (Herter) Herter ex Nessel; Bärlappgew. 231 (1939)
species :
Phlegmariurus tetragonus
(Hook. & Grev.) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Colombia (Caldas, Cauca, Cundinamarca, Huila, Nariño, Putumayo, Quindío, Risaralda, Tolima, Valle); Ecuador; Peru
82 CLM 83 ECU PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia tetragona (Hook. & Grev.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium catharticum Hook.; Ann. Nat. Hist. 1: 430-431, t. 14 (1838)
= Lycopodium myrsinites var. minus Spring; Mém. Acad. Roy. Sci. Bruxelles 15: 29 (1842)
= Lycopodium tetragonum Hook. & Grev.; Icon. Filic. 1: t. 109 (1829)
= Lycopodium tetragonum var. patulum Spring; Mém. Acad. Roy. Soc. Bruxelles 24: 12 (1849)
= Urostachys catharticus (Hook.) Herter ex Nessel; Bärlappgew. 135 (1939)
= Urostachys tetragonus (Hook. & Grev.) Herter ex Nessel; Bärlappgew. 135 (1939)
species :
Phlegmariurus tetrastichoides
(A.R.Field & Bostock) A.R.Field & Bostock; PhytoKeys 20: 48 (2013)
Distribution: Australia (NE-Queensland)
50 AUS
Remarks: hippuris agg. Contains Australian records for P. proliferus.
Synonyms: hidden
= Huperzia tetrastichoides A.R.Field & Bostock; Austrobaileya 7(4): 711 (2008)
species :
Phlegmariurus tetrastichus
(Kunze) A.R.Field & Bostock; PhytoKeys 20: 48 (2013)
Distribution: Vietnam; peninsular Malaysia (Kelantan, Pahang, Negeri Sembilan); Borneo (Sabah, Sarawak); Sulawesi; Java; Moluccas (Seram)
41 VIE 42 BOR JAW MLY MOL SUL
Remarks: carinata agg.
Synonyms: hidden
= Huperzia tetrasticha (Kunze ex Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium sarasinorum Christ; Verh. Naturf. Ges. Basel 11: 34, t. 2o, f. 11-12 (1894)
= Lycopodium tetrastichum Kunze; Bot. Zeitung (Berlin) 6: 99 (1848)
= Urostachys sarasinorum (Christ) Herter ex Nessel; Bärlappgew. 182 (1939)
= Urostachys tetrastichus (Kunze) Herter ex Nessel; Bärlappgew. 183 (1939, by reference to Alderw. (1915a: 35))
species :
Phlegmariurus tico
A.Rojas; Open Access Library Journal 6: e5172 (2019), nom. nov.
Distribution: Costa Rica
80 COS
Remarks: reflexus agg.
Synonyms: hidden
= Phlegmariurus nanus A.Rojas & R.Calderón; Acta Botanica Malacitana 42: 67-70 (2017), non C. Y. Ma 1990
species :
Phlegmariurus toppingii
(Herter) A.R.Field & Bostock; PhytoKeys 20: 48 (2013)
Distribution: Philippines
42 PHI
Remarks: squarrosa agg.
Synonyms: hidden
= Huperzia toppingii (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Urostachys toppingii Herter; Bot. Arch. 3: 15 (1923)
species :
Phlegmariurus tournayanus
Lawalrée; Acta Bot. Acad. Sci. Hung. 19: 197, f. 2 (1973)
Distribution: D.R.Congo [Zaire]
23 ZAI
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia tournayana (Lawalrée) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
species :
Phlegmariurus transilla
(Sodiro ex Baker) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Colombia (Chocó, Santander); Ecuador
82 CLM 83 ECU
Remarks: brevifolia agg.
Synonyms: hidden
= Huperzia transilla (Baker) Holub; Folia Geobot. Phytotax. 20: 77 (1985) (trencilla)
= Lycopodium transilla Sodiro ex Baker; J. Bot. 15: 168 (1877) (trencilla)
= Urostachys transilla (Baker) Herter ex Nessel; Bärlappgew. 106 (1939)
species :
Phlegmariurus treitubensis
(Silveira) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Brazil (Minas Gerais)
84 BRA
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia treitubensis (Silveira) B.Øllg.; Opera Bot. 92: 170 (1987)
= Lycopodium hoehnei (Nessel) Rolleri; Rev. Mus. La Plata (N. S.) Bot. 13: 77 (1981)
= Lycopodium inflexum Silveira; Bol. Comm. Geogr. Geol. Est. Minas Gerais 2,5: 118, t. 2 (1898) (non (P. Beauv.) Sw. (1806))
= Lycopodium saururus var. inflexus (Silveira) Nessel & Hoehne
= Lycopodium treitubense Silveira; Bol. Comm. Geogr. Geol. Est. Minas Gerais 2, 5: 118, t. 3 (1898)
= Urostachys capri Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 55 (1949) (nom. superfl.)
= Urostachys christii var. treitubensis (Silveira) Nessel; Bärlappgew. 46 (1939)
= Urostachys hoehnei Nessel; Arch. Bot. São Paulo 1: 383, t. 6 (1927)
= Urostachys inflexus Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 384 (1927) (nom. nov. for Lycopodium inflexum Silveira (1806) non (P. Beauv.) Sw.
= Urostachys treitubensis (Silveira) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 380 (1927)
species :
Phlegmariurus trifoliatus
(Copel.) A.R.Field & Bostock; PhytoKeys 20: 48 (2013)
Distribution: Fiji (Viti Levu, Vanua Levu, Kadavu, Ovalau, Taveuni)
60 FIJ
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia trifoliata (Copel.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium trifoliatum Copel.; Bull. Bernice P. Bishop Mus. 59: 7, t. 1 (1929)
= Urostachys trifoliatus (Copel.) Herter ex Nessel; Bärlappgew. 224 (1939)
species :
Phlegmariurus trigonus
(C.Chr.) A.R.Field & Bostock; PhytoKeys 20: 49 (2013)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Huperzia trigona (C.Chr.) Tardieu; Adansonia 10: 20 (1970)
= Urostachys trigonus (C.Chr.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 87 (1949)
= Lycopodium trigonum C.Chr.; Dansk Bot. Ark. 7:191, t. 76, f. 1-5 (1932)
species :
Phlegmariurus tryonorum
B.Øllg.; Phytotaxa 277(3): 272 (2016)
Distribution: Costa Rica
80 COS
species :
Phlegmariurus tubulosus
(Maxon) B.Øllg.; Phytotaxa 57: 19 (2012)
Distribution: Costa Rica; Panama
80 COS PAN
Remarks: taxifolia agg.
Synonyms: hidden
= Huperzia costaricensis (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Huperzia tubulosa (Maxon) B.Øllg.; Ann. Missouri Bot. Gard. 79: 697 (1992)
= Lycopodium costaricense (Herter) C.V.Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium tubulosum Maxon; Contr. U.S. Nat. Herb. 17(2): 178, t. 10 (1913)
= Urostachys costaricensis Herter; Amer. Fern J. 48: 83 (1958)
= Urostachys tubulosus (Maxon) Herter ex Nessel; Bärlappgew. 177 (1939)
species :
Phlegmariurus ulixis
(Herter) B.Øllg.; Phytotaxa 57: 20 (2012)
Distribution: Colombia (Cauca, Huila, Narino, Valle)
82 CLM
Remarks: saururus agg.
Synonyms: hidden
= Huperzia ulixis (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium ulixis (Herter) C.V.Morton; Amer. Fern J. 54: 73 (1964)
= Urostachys ulixis Herter; Revista Sudamer. Bot. 10: 115 (1953 publ. 1954)
species :
Phlegmariurus unguiculatus
(B.Øllg.) B.Øllg.; Phytotaxa 57: 20 (2012)
Distribution: Colombia (Cauca, Choco, Putumayo); Ecuador
82 CLM 83 ECU
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia unguiculata B.Øllg.; Harling & Andersson, Fl. Ecuador 33: 26, f. 2B (1988)
species :
Phlegmariurus urbanii
(Herter) B.Øllg.; Phytotaxa 57: 20 (2012)
Distribution: Colombia (Cauca, Nariño, Putumayo); Ecuador; Peru
82 CLM 83 ECU PER
Remarks: reflexa agg.
Synonyms: hidden
= Huperzia urbanii (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium urbanii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 47 (1909)
= Urostachys urbanii (Herter) Herter ex Nessel; Bärlappgew. 110 (1939)
species :
Phlegmariurus vanuatuensis
A.R.Field; PhytoKeys 109: 55 (2018)
Distribution: Vanuatu
60 VAN
species :
Phlegmariurus varius
(R.Br.) A.R.Field & Bostock; PhytoKeys 20: 49 (2013)
Distribution: Australia (Tasmania, E-Queensland, SE-New South Wales, S-Victoria); Lord Howe Isl.; New Zealand (North Isl., South Isl., Stewart Isl.); Chatham Isl.; Antipodes Isl.; Auckland Isl.; Campbell Isl.
50 AUS LHI 51 ATP AUC CTM NZL 91 CMP
Remarks: varia agg.
Synonyms: hidden
= Huperzia desvauxii Holub; Folia Geobot. Phytotax. 26(1): 92 (1991), nom. nov. for Lycopodium pachystachyum Desv. ex Poir. 1814
= Huperzia novae-zelandica (Colenso) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia polaris (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Huperzia varia (R.Br.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1875)
= Lycopodium novae-zelandicum Colenso; Trans. & Proc. New Zealand Inst. 19: 275 (1887)
= Lycopodium pachystachyum Desv.; Encyclopedie Methodique. Botanique Suppl. 3 (1813) ex Poir. (1814)
= Lycopodium phlegmaria A.Cunn.; Compan. bot. Mag. 2: 361 (non L. (1837))
= Lycopodium selago var. varium (R.Br.) C.Moore & Betche; Handbook of the Flora of New South Wales (1893)
= Lycopodium varium R.Br.; Prodr. Fl. Nov. Holland. 65 (1810)
= Lycopodium varium var. alpinum R.Br.; Prodr. Fl. Nov. Holland. 65 (1810)
= Lycopodium varium var. gracile Kirk; Trans. New Zealand Inst. 16: 377 (1884)
= Lycopodium varium var. polaris Kirk; Trans. New Zealand Inst. 16: 376 (1884)
= Lycopodium varium var. umbrosum R.Br.; Prodr. Fl. Nov. Holland. 65 (1810)
= Urostachys polaris (Nessel) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 76 (1949)
= Urostachys varius (R.Br.) Herter ex Nessel; Bärlappgew. 192 (1939)
= Urostachys varius var. polaris Herter ex Nessel; Revista Sudamer. Bot. 6: 165, t. 12, fig. 59 (1940) (non Lycopodium varium R. Br. var. polaris Kirk (1849)
species :
Phlegmariurus venezuelanicus
(Herter) B.Øllg.; Phytotaxa 57: 20 (2012)
Distribution: Venezuela (Sa. Nevada de Merida)
82 VEN
Remarks: saururus agg.
Synonyms: hidden
= Huperzia schneei (Vareschi) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Huperzia venezuelanica (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium schneei Vareschi; Acta Bot. Venez. 1: 83, f. 1 (1966)
= Lycopodium venezuelanicum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909)
= Urostachys venezuelanicus (Herter) Herter ex Nessel; Bärlappgew. 74 (1939)
species :
Phlegmariurus vernicosus
(Hook. & Grev.) Á.Löve & D.Löve; Taxon 26: 324 (1977)
Distribution: S-India (Tamil Nadu, Kerala); Sri Lanka
40 IND SRL
Conservation Status: IUCN CR (Fraser-Jenkins 2012)
Synonyms: hidden
= Huperzia vernicosa (Hook. & Grev.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
= Lycopodium vernicosum Hook. & Grev. ex Hook. & Grev.; Bot. Misc. 2: 364 (1831)
= Urostachys vernicosus (Hook. & Grev.) Herter ex Nessel; Bärlappgew. 61 (1939)
species :
Phlegmariurus verticillatus
(L.f.) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; D.R.Congo [Zaire]; Rwanda; Kenya; Tanzania; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, E-Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho; Eswatini [Swaziland]; Madagascar; Comores (Mayotte); Mauritius; La Réunion
23 BIO CMN RWA SAO ZAI 25 KEN TAN 26 MLW MOZ ZAM ZIM 27 LES SAF SWZ 29 COM MAU MDG REU
Remarks: verticillata agg. True H. verticillatus is Western Indian Ocean. Populations in Malesia, Pacific belong to P. filiformis, New World to P. portoricensis. About the continental African Populations there is still uncertainty.
Synonyms: hidden
= Huperzia verticillata (L.f.) Trevis.; Atti Soc. Critt. Ital. 17: 248 (1875)
= Lycopodium setaceum Lam.; Encycl. 3: 653 (1789 (1792))
= Lycopodium verticillatum L.f.; Suppl. Pl. 448 (1782)
= Plananthus setaceus (Lam.) P.Beauv.; Prodr. Aethéogam. 112 (1805)
= Plananthus verticillatus (L.f.) P.Beauv.; Prodr. Aethéogam. 112 (1805)
= Stachygynandrum verticillatum (L.f.) P.Beauv.; Prodr. Aethéogam. 113 (1805)
= Urostachys verticillatus (L.f.) Herter; Beih. Bot. Centralbl. 39: 249 (1922)
= Urostachys verticillatus var. gracilis Herter ex Nessel; Bärlappgew. 123 (1939)
species :
Phlegmariurus villonacensis
(B.Øllg.) B.Øllg.; Phytotaxa 57: 20 (2012)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Huperzia villonacensis B.Øllg.; Nordic J. Bot. 23(1): 33 (2004)
species :
Phlegmariurus warneckei
(Herter ex Nessel) A.R.Field & Testo; Molec. Phylogen. Evol. 94(B): 645 (2015)
Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Cameroon; São Tomé; Gabon; Congo [Brazzaville]
22 GUI IVO LBR SIE 23 CMN CON GAB SAO
Remarks: phlegmaria agg.
Synonyms: hidden
= Huperzia warneckei (Nessel) Pic.Serm.; Webbia 23: 163 (1968)
= Urostachys warneckei Herter ex Nessel; Bärlappgew. 213 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 166, t. 13, f. 64 (1940)
= Lycopodium warneckei (Nessel) Alston; Bol. Soc. Broter. Ser. 2A, 30: 20 (1956)
species :
Phlegmariurus watsonianus
(Maxon) B.Øllg.; Phytotaxa 57: 20 (2012)
Distribution: El Salvador; Costa Rica; Panama
80 COS ELS PAN
Remarks: verticillata agg.
Synonyms: hidden
= Huperzia watsoniana (Maxon) B.Øllg.; Ann. Missouri Bot. Gard. 79: 701 (1992)
= Lycopodium watsonianum Maxon; Smithsonian Misc. Collect. 56(29): 3, t. 3 (1912)
= Urostachys watsonianus (Maxon) Herter ex Nessel; Bärlappgew. 159 (1939)
species :
Phlegmariurus weberbaueri
(Nessel) B.Øllg.; Phytotaxa 57: 20 (2012)
Distribution: Ecuador; Peru
83 ECU PER
Remarks: saururus agg.
Synonyms: hidden
= Huperzia papillata (Rolleri) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Huperzia weberbaueri (Hieron. & Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Lycopodium papillatum Rolleri; Amer. Fern J. 65(1): 3 (1975)
= Urostachys weberbaueri Herter ex Nessel; Bärlappgew. 99 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 162, t. 10p, f. 41 (1940)
species :
Phlegmariurus weddellii
(Herter) B.Øllg.; Phytotaxa 57: 20 (2012)
Distribution: Ecuador; Peru; Bolivia (La Paz)
83 BOL ECU PER
Remarks: brongniartii agg.
Synonyms: hidden
= Huperzia weddellii (Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Lycopodium weddellii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909)
= Urostachys weddellii (Herter) Herter ex Nessel; Bärlappgew. 91 (1939)
species :
Phlegmariurus wilsonii
(Underw. & F.E.Lloyd) B.Øllg.; Rodriguésia 63(2): 481 (2012)
Distribution: Mexico (Oaxaca); Guatemala; Nicaragua; Costa Rica; Panama; Jamaica; Puerto Rico; Saba; Guadeloupe; Martinique; St. Lucia; St. Vincent; Trinidad; Colombia (Antioquia, Nariño, Risaralda, Santander, Valle, Meta, Choco, Norte de Santander); Venezuela (Amazonas, Aragua, Bolívar); Surinam; Ecuador; Peru; Bolivia (Cochabamba, La Paz, Santa Cruz); Brazil (Mato Grosso)
79 MEX 80 COS GUA NIC PAN 81 GDL JAM MAR PUE SAB STL STV TRT 82 CLM SUR VEN 83 BOL ECU PER 84 BRA
Remarks: wilsonii agg.
Synonyms: hidden
= Huperzia lindaviana (Herter) Holub; Folia Geobot. Phytotax. 33: 97 (1988)
= Huperzia wilsonii (Underw. & F.E.Lloyd) B.Øllg.; Opera Bot. 92: 170 (1987)
= Lycopodium andinum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 49 (1909) (non Rosenst. (1908))
= Lycopodium arcanum Maxon; Field Mus. Nat. Hist., Bot. Ser. 17(4): 310, t. 3 (1938)
= Lycopodium lindavianum Herter; Hedwigia 49(2-3): 90, as Lycopodium Lindavianum (1909)
= Lycopodium stamineum Maxon; Smithsonian Misc. Collect. 56(29): 2-3, t. 2 (1912)
= Lycopodium trichodendron Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 49 (1909)
= Lycopodium wilsonii Underw. & F.E.Lloyd; Bull. Torrey Bot. Club 33: 111 (1906)
= Urostachys lindavianus (Herter) Herter ex Nessel; Bärlappgew. 146 (1939)
= Urostachys mandiocanus var. filifolius Herter; Revista Sudamer. Bot. 10: 124 (1954)
= Urostachys stamineus (Maxon) Herter ex Nessel; Bärlappgew. 148 (1939)
= Urostachys trichodendron (Herter) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 163 (1923)
= Urostachys wilsonii (Underw. & F.E.Lloyd) Herter; Repert. Spec. Nov. Regni Veg. 19(11-13): 163 (1923)
species :
Phlegmariurus xiphophyllus
(Baker) A.R.Field & Bostock; PhytoKeys 20: 49 (2013)
Distribution: Madagascar
29 MDG
Remarks: squarrosa agg.
Synonyms: hidden
= Huperzia xiphophylla (Baker) Tardieu; Adansonia ser. 2 10: 20 (1970)
= Lycopodium xiphophyllum Baker; J. Linn. Soc., Bot. 22: 535 (1887)
= Urostachys xiphophyllus (Baker) Herter ex Nessel; Bärlappgew. 72, t. 11, f. 9 (1939)
species :
Phlegmariurus yunfengii
R.H.Jiang & X.C.Zhang; Turczaninowia 25(1): 131 (2022)
Distribution: China (Yunnan)
36 CHI
species :
Phlegmariurus yunnanensis
Ching; Acta Bot. Yunnan. 4: 121 (1982)
Distribution: China (Yunnan)
36 CHI
Remarks: cancellata agg.
Synonyms: hidden
= Huperzia yunnanensis (Ching) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
species :
Phlegmariurus ×koolauensis
W.H.Wagner; Contr. Univ. Michigan Herb. 22: 138 (1999)
Distribution: Hawaii
63 HAW
Remarks: Phlegmariurus nutans x phyllanthus
Synonyms: hidden
= Huperzia ×koolauensis (W.H.Wagner) Kartesz; Synthesis N. Amer. Fl., version 1 Nomencl. Innov.: no. 13 (1999)
