Parsley fern

Cryptogramma crispa

''Cryptogramma crispa'', the parsley fern, is an Arctic–alpine species of fern. It produces separate sterile and fertile fronds, up to 30 cm tall, and is a pioneer species on acidic screes.
Parsley Fern Growing on drystone wall on way to Place Fell. Cryptogramma crispa,Cumbria,Parsley fern

Appearance

The fronds of ''C. crispa'' are 30 centimetres long and appear in two distinct forms. Sterile leaves are 2–3-pinnate with the pinnules 5–10 mm long by 3–7 mm wide, while fertile leaves are 3–4-pinnate, and with narrower pinnules. The fertile leaves have sori scattered along the veins, each with a strongly enrolled false indusium. The sporangia are yellow and mature around midsummer.
Parsley Fern Close up of this delicate fern growing on a drystone wall Cryptogramma crispa,Cumbria,Parsley fern

Distribution

''Cryptogramma crispa'' grows among acidic rocks in areas where snow lies until late in the year. It is a pioneer species on stable scree slopes and also occurs on cliffs and dry stone walls.

In Europe, ''C. crispa'' has an Arctic–alpine distribution, growing in the mountains of Central and Southern Europe, as well as in the north of the continent, including Scandinavia and higher ground in the British Isles. In Ireland, it is rare and concentrated in the east of the country, leading Praeger to conjecture that the Irish examples are recent colonists from Great Britain, arriving as airborne spores.

Similar plants, which may belong to the same species occur in East Asia and North America, although these are usually considered a separate taxon.

Spores attributable to ''C. crispa'' have been discovered in deposits in Snowdonia from the last glacial period, as well as from more low-lying sites in Cheshire.

Habitat

''Cryptogramma crispa'' grows among acidic rocks in areas where snow lies until late in the year. It is a pioneer species on stable scree slopes and also occurs on cliffs and dry stone walls.

In Europe, ''C. crispa'' has an Arctic–alpine distribution, growing in the mountains of Central and Southern Europe, as well as in the north of the continent, including Scandinavia and higher ground in the British Isles. In Ireland, it is rare and concentrated in the east of the country, leading Praeger to conjecture that the Irish examples are recent colonists from Great Britain, arriving as airborne spores.

Similar plants, which may belong to the same species occur in East Asia and North America, although these are usually considered a separate taxon.

Spores attributable to ''C. crispa'' have been discovered in deposits in Snowdonia from the last glacial period, as well as from more low-lying sites in Cheshire.

Evolution

The parsley fern appeared in Carl Linnaeus' 1753 work ''Species Plantarum'', the starting point for botanical nomenclature, under the name ''Osmunda crispa''. The specific epithet ''crispa'' means finely waved or curled. It is placed in the family Pteridaceae, part of the order Polypodiales.

References:

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Taxonomy
KingdomPlantae
DivisionPolypodiophyta
ClassPolypodiopsida
OrderPolypodiales
FamilyPteridaceae
GenusCryptogramma
SpeciesC. crispa
Photographed in
United Kingdom