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WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
SpeciesMapsDocumentsIDAO

Rottboellia cochinchinensis (Lour.) Clayton

Accepted
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton
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🗒 Synonyms
synonymAegilops fluviatilis Blanco
synonymManisuris exaltata Kuntze
synonymManisuris exaltata var. appendiculata (Steud.) Honda
synonymManisuris exaltata var. appendiculata (Steud.) Honda, nom. superfl.
synonymOphiuros appendiculatus Steud.
synonymRottboellia arundinacea Hochst. ex A.Rich.
synonymRottboellia denudata Steud.
synonymRottboellia exaltata f. arundinacea (Hochst. ex A.Rich.) Hack.
synonymRottboellia exaltata f. glaberrima Chiov.
synonymRottboellia exaltata L.f., nom. illeg.
synonymRottboellia exaltata var. appendiculata (Steud.) Hack.
synonymRottboellia exaltata var. robusta Hook.f.
synonymRottboellia hispida Roxb. ex Hook.f. [Invalid]
synonymRottboellia hispida Roxb. ex Hook.f., pro syn.
synonymRottboellia setosa J.Presl
synonymStegosia cochinchinensis Lour.
synonymStegosia exaltata (Kuntze) Nash
🗒 Common Names
Comorian
  • Sandzemadji
Creoles and pidgins, French-based
  • Herbe fataque duvet, Fataque duvet, Herbe de riz, Herbe à poils, Herbe duvet
Creoles and pidgins;
  • Rottboellia
Creoles and pidgins; French-based
  • Zèb a diri (Antilles)
English
  • Itch grass
Malagasy
  • Kalay, Tsanganday, Paipaikatianomby
  • Fataka malailay (Moyen-Ouest)
📚 Overview
Overview
Brief

Code

ROOEX

Growth form

grass

Biological cycle

annual

Habitat

terrestrial

 

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    Diagnostic Keys
    Description
    Global description

    Rottboellia cochinchinensis is a grass in tuft, measuring up to 4 m high. The base of the stem is bristling with stiff hairs and presents harsh stilt roots. The thatch are robust, with adventitious roots at the lower nodes. The leaves have a sheath with erect stiff and prickly hairs. The blade is long, linear and hairy on the upper face. It has a very marked white midrib. The inflorescence is a fake cylindrical terminal spike, consisting of articles standing out easily.
     
    First leaves

    The seed is determinant characteristics since it remains attached for a long time to the seedling by an axis of 1 to 2 cm long. The first leaf is characteristic. The leaf blade is elliptic, 2 to 3 cm long and 5 to 6 mm wide, with wide angled or round top. This lamina is rapidly spread horizontally. The following leaves are laid obliquely. The prefoliation is rolled. The sheath becomes pubescent or hispid until it is covered with stiff hairs perpendicular to the surface implantation and very pungent. The ligule is membranous. The lamina is raised obliquely. It is linear with sharp pointed top. Some long hairs are spread in the lower part. Some ecotypes are completely glabrous.
     
    General habit

    Plant in thick tuft, high tillering and axis abundantly branched. It measures up to 4 m high.
     
    Underground system

    Roots fibrous, others hard, with stilt developing from lower nodes of the plant.
     
    Culm

    Culm cylindrical, sparsely covered with stiff hairs and with scabrous dark nodes. It is robust at the base, with a diameter reaching up to 1 cm.
     
    Leaf

    Simple, alternate leaves. The sheath is cylindrical, and slightly keeled and hispid over its entire length. It has white stiff hairs (tuberculate and perpendicular to the surface of implantation). The ligule is membranous, 1 mm high, barely cilliated at the top. The lamina is linear, flat, drawn obliquely acute apex, up to 1.5 m long and 1 to 2.5 cm wide, grooved white midrid. The margin is scabrous. The upper surface is sparsely covered with long tuberculate hairs, the underside is glabrous.
     
    Inflorescence

    Inflorescence consisting of many cylindrical spikes, 5 to 15 cm long, generated from a foliated husk and consist of articles disarticulating very easily. Each consist of a internode, 5 mm long, with a rachis flattened into a cavity in which is located the sessile spikelet. The pedicel of the pedicellate spikelet is attached to the rachis along its length. The sessile spikelet is equipped by a corneous rotula at the base that articulate on the proceeding article.

    Spikelet

    The sessile spikelet has 2 flowers, the internal is male or sterile and external fertile. The lower glume is very convex, with acuminate apex, included in the alveole of the rachis. The upper glume is oval with a wide angled tip; both are thick, leathery and glabrous. Lemma and palea are papery and translucent. The pedicellate spikelet consists of a lower oval glume and a flattened and bicarinate upper glume. Both are membranous and green.
     
    Grain

    Oblong grain, 2 to 2.5 mm long and surmounted by a short point. It remains included in spikelet, itself secured to the article of the spike that disarticulates at maturity.
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      No Data
      📚 Natural History
      Life Cycle

      Life cycle

      Annual
      Annual

      North Cameroon: Rottboellia cochinchinensis germination period begins in April with the first rains and lasts until late May. Any cultural operation (plowing, weeding, hilling) is followed by another period of germination. These new emergence can occur until September. Flowering begins in August but may be delayed until October in late emerged plants. Fruiting and dissemination take place from September to November until drying of the plant. In areas without a pronounced dry season, the species can develop at any time of year.

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        Reproduction

        Rottboellia cochinchinensis is an annual species. It multiplies only by seeds that can remain viable in the soil for 2 to 3 years. The dissemination is done by water, agricultural tools, seeds of cultivated plants and animals. An individual can produce more than 3000 seeds.

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          Morphology

          Growth form

          Tuft plant with narrow leaves
          Tuft plant with narrow leaves
          Erected
          Erected

          Leaf type

          Grass or grass-like
          Grass or grass-like

          Latex

          Without latex
          Without latex

          Root type

          Fibrous roots
          Fibrous roots

          Ligule type

          Ligule membranous large
          Ligule membranous large

          Stipule type

          No stipule
          No stipule

          Leaf attachment type

          with graminate sheathing
          with graminate sheathing

          Fruit type

          Grain of grasses
          Grain of grasses

          Lamina base

          sheathing grass-like broader
          sheathing grass-like broader

          Lamina margin

          scabrous
          scabrous
          entire
          entire

          Lamina apex

          attenuate
          attenuate

          Upperface pilosity

          Glabrous
          Glabrous
          Less hairy
          Less hairy

          Upperface hair type

          Hairs with glandulous base
          Hairs with glandulous base

          Simple leaf type

          Lamina linear
          Lamina linear

          Lamina section

          flat
          flat
          folded
          folded

          Inflorescence type

          Spike
          Spike
          Raceme with alternate sessile flowers
          Raceme with alternate sessile flowers

          Stem pilosity

          Dense hairy
          Dense hairy
          Less hairy
          Less hairy
          Look Alikes
          Hemarthria altissima (Poir.) Stapf & C.E.Hubb. can be mistaken Rottboellia cochinchinensiset is readily distinguished by the following characters: H. altissima is a perennial stoloniferous species with elliptical thatched, glabrous, with shortly membranous ligule and ciliated at the top. The cylindrical racemes laterally compressed have pedicellate spikelets that disarticulate without breaking if one stretches the raceme. Conversely, R. cochincinensis is an annual species with cylindrical thatch, often bristling with very stiff thorny bristles to shortly membranous ligule and barely ciliated. Cylindrical racemes are formed of spikelets whose pedicel is welded to the rachis, which does not allow the raceme to stretch by disarticulating without breaking if one tries to stretch.

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            Ecology

            Northern Cameroon: Rottboellia cochinchinensis is a nitrophilous species characteristic of silty clay and humid soil. It grows especially on alluvial soils along rivers, vertisols, the planosols in the shallows and on well-structured ferruginous soils.
            French Guiana: Ruderal species which appreciates heavy and humid clay soils.
            Comoros: species of medium to high altitude, as from 400 m. It is common in shaded to semi exposed areas.
            Madagascar: Species widespread in all sub-humid and semi-arid low or medium altitude area of the Island (North West, West, Midwest and Southwest). It infests recent fallow and rainfed crops in the alluvial plains, depressions and lowlands where soils (vertisols, alluvial soils, ferruginous hydromorphic soils) are relatively rich.
            Mauritius: Species present especially in the lower part of the island, it grows in abandoned fields and sugarcane fields.
            Mayotte: Rottboellia cochinchinensis is an exotic species which grows during the rainy season on heavy and wet soils. It is particularly present in the center of the island.
            Reunion: This plant is common in humid, sub-humid and warm. It appreciates the deep silty clay soil and very wet. It has now invaded all island areas below 1000 m, excluding the drylands.
            Seychelles: Absent.
            West Indies: Rottboellia cochinchinensis is an exotic species. It grows easily in untended areas and is particularly fond of clay soils in subhumid and humid areas.

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              No Data
              📚 Habitat and Distribution
              Description

              Geographical distibution

              Madagascar
              Madagascar
              Reunion Island
              Reunion Island
              Comoros
              Comoros
              Mauritius
              Mauritius
              Origin

              Rottboella conchinchinensis is native to Africa, India and Southeast Asia until Northern Australia.

              Worldwide distribution

              This species is now very common in all tropical regions of Asia, Africa, America, Caribbean, Pacific and Australia.

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                No Data
                📚 Occurrence
                No Data
                📚 Demography and Conservation
                Risk Statement

                Local harmfulness

                Benin: Rottboellia cochinchinensis is rare, but abundant when present in paddy fields.
                Burkina Faso: rare and scarce species in paddy fields.
                Northern Cameroon: This species causes extensive damage to corn crops, especially cotton and upland rice. Its rapid growth allows it to seriously compete with crops early in the crop cycle. It is slightly sensitive to commonly used pre-emergence herbicides for weed control in cotton. The manual or mechanical weeding should be done early because when the stilt roots develop, the extraction becomes very difficult, especially as the stiff hairs of the base are wounding. This species can invade a plot in 2-5 years if its development is not adequately controlled by a specific herbicide and by early and repeated weeding. However, a good control of this species in 2 or 3 years provides an almost perfect eradication because the viability of seeds in the soil is low.
                Chad: Rare and scarce in paddy fields.
                Comoros:
                major weed of banana crops and fallow.
                Ivory Coast: Frequent and usually abundant in paddy fields.
                Ghana: Frequent and scarce in paddy fields.
                French Guiana: An infrequent species and never abundant in orchards.
                Kenya: Rare but abundant when present in paddy fields.
                Madagascar: common species (43% of the plots in the Midwest and 60% in the South West) fast growing often forming a dense stand, it is especially harmful in growing cotton, maize and upland rice or peanut. Weeding is extremely difficult in rainy season or after a few days of delay because of its stiff wounding hairs and of its strong rootedness. It can quickly stifle crops thereby causing partial or total abandonment of heavily infested plots after delayed intervention.
                Mali: Frequent and scarce in paddy fields.
                Mauritius: A weed of low to medium harmfulness, hard to master.
                Mayotte: Rottboellia cochinchinensis is an infrequent weed, present in 4% of cultivated plots. It is mainly present in fruit crops. It can be found in food crops.
                Reunion: This species is present in 25% of the cultivated plots of the island. It is currently in the process of multiplying to the extent of forming real populations in sugarcane cultivation with high coverage of 30%. It also causes extensive damage to corn crops.
                Senegal: Frequent and scarce in paddy fields.
                Seychelles: Absent.
                Tanzania: Frequent and usually abundant in paddy fields.
                Uganda: Frequent and usually abundant in paddy fields.
                West Indies: Rottboellia cochinchinensis is a very damaging weed present in all crops. If it is not controlled, it forms very damaging stands, particularly in sugarcane and young fruit crops. Under these conditions, each weeding operation can be followed by a germination.

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                  No Data
                  📚 Uses and Management
                  Management

                  Global control

                  For annual grass weed control tips FOR irrigated rice and lowland in Africa, visit: http://portal.wiktrop.org/document/show/32 

                  Local control


                  Madagascar: Tillage triggers germination of Rottboellia cochinchinensis. Pulling or weeding should be done very early, before the development of the root system makes them difficult.
                  Pre-clearance, pendimethalin (or oxadiazon) are very interesting in infested plots. In post-emergence, the young roettboellia is susceptible to fluazifop-p-butyl. Glyphosate alone destroys the old seedlings.
                  Able to grow through mulching. A very thick blanket of stylosanthes however enables its control and elimination over few years.

                   La Réunion : see the site of  CaroCanne (online magasine for the experts in sugarcane in Reunionn) ; cahier technique n°34

                  Herbicide efficacy spectrum on Rottboellia cochinchinensis in sugarcane cultivation
                  Active Ingredients commercial products doses of commercial products efficiency
                  pre-emergence      
                  mésotrione +
                  S-métolachlore
                  Camix 3,75 l/ha  
                  mésotrione + S-métolachlore
                  + S-métolachlore
                  Camix + Mercantor Gold 3,75 l/ha + 0,5 l/ha  
                  mésotrione + S-métolachlore
                  + isoxaflutole
                  Camix + Merlin 3,75 l/ha + 0,1 kg/ha  
                  mésotrione + S-métolachlore
                  + pendiméthaline
                  Camix + Prowl 400 3,75 l/ha + 3,0 l/ha  
                  isoxaflutole + pendiméthaline
                  + métribuzine
                  Merlin + Prowl 400 + Sencoral 0,067 kg/ha + 1,5 l/ha + 0,625 kg/ha  
                  isoxaflutole + pendiméthaline
                  + mésotrione + S-métolachlore
                  Merlin + Prowl 400 + Camix 0,067 kg/ha + 1,5 l/ha + 2,5 l/ha  
                  post-emergence      
                  2,4-D
                  .
                  2,4-D 2,0 l/ha  
                  2,4-D
                  + mésotrione
                  2,4-D + Callisto 2,0 l/ha + 1,0 l/ha  

                  (Doses are expressed in commercial products) - 2014

                    Good efficiency
                    Medium efficiency
                    Ineffective



                  Data acquired in Reunion on the effectiveness of herbicides in the context of the sugarcane herbicide network by eRcane Network with funding from the ODEADOM and ONEMA.
                  Action led by the French Ministry of Agriculture, food and forest, with financial support from the National Agency for Water and Aquatic Environments, on the finance issued from the tax for the pollution diffused, attributed to the finance of the Ecophyto plan.

                  West Indies: Good control of Rottboellia cochinchinensis requires the use of a pre- or post-emergence herbicide when it has 4 to 5 leaves.

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                    📚 Information Listing
                    References
                    1. Johnson, D.E. 1997. Weeds of rice in West Africa. ADRAO/WARDA, Bouaké, Côte-d'Ivoire. 196p
                    2. Philips,S.,Namaganda, M., Lye, K.A.2003.Makerere University handbook no. 1.115 Ugandan Grasses. Department of Botany, Makerere University, Kampala.262p.
                    3. Akobundu O. & Agyakwa C.W.: A handbook of West African Weeds. International Institute of Tropical Agriculture, Ibadan, Nigeria (1998), 522p.
                    4. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                    1. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186.https://portal.wiktrop.org/document/show/173
                    2. https://www.feedipedia.org/node/388
                    3. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:909653-1
                    4. Zon van der A.P.M., 1992. Graminées du Cameroun, Vol. II, Flore.
                    5. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontanée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.
                    6. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                    7. Grossard, F., Le Bourgeois, T., Dumbardon-Martial, E. & Gervais, L. 2013. Adventilles - Guadeloupe & Martinique - Les adventices des Antilles françaises. Abymes, Guadeloupe, France, Les éditions du CTCS Guadeloupe. 195 p.
                    1. Vanden Berghen C., 1983. Matériaux pour une flore de la végétation herbacée de la Casamance occidentale, Sénégal, Fascicule 2, Gramineae. Jardin Botanique National de Belgique, 66p.
                    1. Holm L. G., Plucknett D. L., Pancho J. V. & Herberger J. P., 1977. The World's Worst Weeds : Distribution and Biologie. East-West Center, University Press of Hawaii, Honolulu, 609p.
                    1. Wageningen Agric. Univ. Papers 92 - 1, Wageningen, 57p.
                    1. Berhaut J., 1967. Flore du Sénégal. 2ème éd. Clairafrique éd., Dakar, Sénégal, 485p.
                    1. Grard, P., T. Le Bourgeois and H. Merlier (2010). Adventrop V.1.5 Les adventices d'Afrique soudano-sahélienne. Montpellier, France, Cirad.
                    1. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                    1. Ivens G. W., 1989. East African Weeds and Their Control. Oxford University Press, Nairobi, Kenya, 289p.
                    1. Ivens G. W., Moody K. & Egunjobi J. K., 1978. West African Weeds. Oxford University Press, Ibadan, Nigeria, 255p.
                    1. Koch W., 1981. Mauvaises herbes des cultures tropicales. In Kranz J., Schmutterer H. & Koch W. : Maladies, ravageurs et mauvaises herbes des cultures tropicales. Verlag Paul Parey, Berlin, Hambourg : 587-665.
                    1. Le Bourgeois Th., 1993. Les mauvaises herbes dans la rotation cotonnière au Nord-Cameroun (Afrique) - Amplitude d'habitat et degré d'infestation - Cycle de développement. Thèse USTL Montpellier II, Montpellier, France, 241p.
                    1. Le Bourgeois, T., Carrara, A., Dodet, M., Dogley, W., Gaungoo, A., Grard, P., Ibrahim, Y., Jeuffrault, E., Lebreton, G., Poilecot, P., Prosperi, J., Randriamampianina, J.A., Andrianaivo, A.P., Théveny, F. 2008. Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien.V.1.0. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom.Merlier H. & Montégut J., 1982. Adventices tropicales.ORSTOM-GERDAT-ENSH éd., Montpellier, France, 490p.
                    1. Stanfield D.P., 1970. The flora of Nigeria, Grasses. Stanfield and Lowe ed., Ibadan University Press, Ibadan, Nigeria, 118p.
                    1. Akobundu I.O. & Agyakwa C.W., 1989. Guide des adventices d'Afrique de l'Ouest. Institut international d'agriculture tropicale. Ibadan, Nigeria, 521p.
                    2. https://www.cabi.org/isc/datasheet/47782
                    Information Listing > References
                    1. Johnson, D.E. 1997. Weeds of rice in West Africa. ADRAO/WARDA, Bouaké, Côte-d'Ivoire. 196p
                    2. Philips,S.,Namaganda, M., Lye, K.A.2003.Makerere University handbook no. 1.115 Ugandan Grasses. Department of Botany, Makerere University, Kampala.262p.
                    3. Akobundu O. & Agyakwa C.W.: A handbook of West African Weeds. International Institute of Tropical Agriculture, Ibadan, Nigeria (1998), 522p.
                    4. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                    5. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186.https://portal.wiktrop.org/document/show/173
                    6. https://www.feedipedia.org/node/388
                    7. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:909653-1
                    8. Zon van der A.P.M., 1992. Graminées du Cameroun, Vol. II, Flore.
                    9. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontanée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.
                    10. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                    11. Grossard, F., Le Bourgeois, T., Dumbardon-Martial, E. & Gervais, L. 2013. Adventilles - Guadeloupe & Martinique - Les adventices des Antilles françaises. Abymes, Guadeloupe, France, Les éditions du CTCS Guadeloupe. 195 p.
                    12. Vanden Berghen C., 1983. Matériaux pour une flore de la végétation herbacée de la Casamance occidentale, Sénégal, Fascicule 2, Gramineae. Jardin Botanique National de Belgique, 66p.
                    13. Holm L. G., Plucknett D. L., Pancho J. V. & Herberger J. P., 1977. The World's Worst Weeds : Distribution and Biologie. East-West Center, University Press of Hawaii, Honolulu, 609p.
                    14. Wageningen Agric. Univ. Papers 92 - 1, Wageningen, 57p.
                    15. Berhaut J., 1967. Flore du Sénégal. 2ème éd. Clairafrique éd., Dakar, Sénégal, 485p.
                    16. Grard, P., T. Le Bourgeois and H. Merlier (2010). Adventrop V.1.5 Les adventices d'Afrique soudano-sahélienne. Montpellier, France, Cirad.
                    17. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                    18. Ivens G. W., 1989. East African Weeds and Their Control. Oxford University Press, Nairobi, Kenya, 289p.
                    19. Ivens G. W., Moody K. & Egunjobi J. K., 1978. West African Weeds. Oxford University Press, Ibadan, Nigeria, 255p.
                    20. Koch W., 1981. Mauvaises herbes des cultures tropicales. In Kranz J., Schmutterer H. & Koch W. : Maladies, ravageurs et mauvaises herbes des cultures tropicales. Verlag Paul Parey, Berlin, Hambourg : 587-665.
                    21. Le Bourgeois Th., 1993. Les mauvaises herbes dans la rotation cotonnière au Nord-Cameroun (Afrique) - Amplitude d'habitat et degré d'infestation - Cycle de développement. Thèse USTL Montpellier II, Montpellier, France, 241p.
                    22. Le Bourgeois, T., Carrara, A., Dodet, M., Dogley, W., Gaungoo, A., Grard, P., Ibrahim, Y., Jeuffrault, E., Lebreton, G., Poilecot, P., Prosperi, J., Randriamampianina, J.A., Andrianaivo, A.P., Théveny, F. 2008. Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien.V.1.0. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom.Merlier H. & Montégut J., 1982. Adventices tropicales.ORSTOM-GERDAT-ENSH éd., Montpellier, France, 490p.
                    23. Stanfield D.P., 1970. The flora of Nigeria, Grasses. Stanfield and Lowe ed., Ibadan University Press, Ibadan, Nigeria, 118p.
                    24. Akobundu I.O. & Agyakwa C.W., 1989. Guide des adventices d'Afrique de l'Ouest. Institut international d'agriculture tropicale. Ibadan, Nigeria, 521p.
                    25. https://www.cabi.org/isc/datasheet/47782

                    Etude floristique et phytoécologique des adventices des complexes sucriers de Ferké 1 et 2, de Borotou-Koro et de Zuenoula, en Côte d'Ivoire

                    Thomas Le Bourgeois
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                    Thomas Le Bourgeois
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                      WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areasWIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
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