Abstract
Efforts to broaden and improve the genetic stock from which horticultural material of Kalanchoe (Crassulaceae subfam. Kalanchooideae) is made available globally are on-going. Apart from making small, light-weight, highly floriferous material—both single- and double-flowered—available to the in- and outdoor plant trades, work is also underway to select Kalanchoe material that can be sold as long-lasting cut flowers. Material derived from K. lubangensis × K. sexangularis, here described as K. ×forbesiae, shows promise as a source of cut flowers.
References
<p>Adanson, M. (1763) <em>Familles des Plantes</em>.<em> Seconde Partie. Etat actuel de la Botanike</em>. Chez Vincent, Imprimeur-Libraire de M<sup>gt</sup> le Comte de Provence, Paris, 640 pp. [https://www.biodiversitylibrary.org/item/6958#page/794/mode/1up]</p>
<p>Baker, J.G. (1881 [21 February]) Notes on a collection of flowering plants made by L. Kitching, Esq., in Madagascar in 1879 (Plates VII. & VIII.). <em>Journal of the Linnean Society, Botany</em> 18: 264–281. [https://www.biodiversitylibrary.org/item/8373#page/271/mode/1up]</p>
<p>Brown, N.E. (1913) 1436. <em>Kalanchoe sexangularis</em>, N. E. Brown [Crassulaceae]. <em>In</em>: Diagnoses Africanae LIII. <em>Bulletin of Miscellaneous Information, Kew</em> 1913 (3): 118–123. [https://www.biodiversitylibrary.org/item/43726#page/125/mode/1up]</p>
<p>Fernandes, R.B. (1980) Notes sur quelques espèces du genre <em>Kalanchoe</em> Adans. <em>Boletim da Sociedade Broteriana</em> (série 2.<sup>a</sup>) 53: 325–442.</p>
<p>Forbes, H. (1941) <em>Kalanchoe albiflora</em> H. Forbes, <em>sp. nov.</em>, pp. 37–39. <em>In</em>: Anonymous, Newly described species and new combinations. <em>Bothalia</em> 4: 37–41, + plate on p. 39. [https://archive.org/details/bothaliavolume404unse/page/36/mode/2up]</p>
<p>Gunn, M. & Codd, L.E.[W.]. (1981) <em>Botanical exploration of southern Africa. An illustrated history of early botanical literature on the Cape flora. Biographical accounts of the leading plant collectors and their activities in southern Africa from the days of the East India Company until modern times</em>. A.A. Balkema, Cape Town, 400 pp.</p>
<p>Hamet, R. (1908) <em>Kalanchoe luciae</em> <em>sp. nov.</em> <em>Bulletin de l’Herbier Boissier</em>, sér. 2, 8: 254–257. [https://www.biodiversitylibrary.org/item/104945#page/290/mode/1up]</p>
<p>Harvey, W.H. (1862) Order LIII. Crassulaceae, D.C. VIII. <em>Kalanchoe</em>, Adans. <em>In</em>: W.H. Harvey & O.W. Sonder, <em>Flora capensis</em> [<em>being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal</em>] 2. L. Reeve & Co., Ltd, Kent, pp. 378–380. [https://www.biodiversitylibrary.org/item/15233#page/386/mode/1up]</p>
<p>IPNI. (2021 [continuously updated]) <em>The International Plant Names Index</em>. Available from: https://www.ipni.org/ (accessed 2021)</p>
<p>Koorders, S.H. (1919 [Mai] [24 October 1918]) Beitrag zur Kenntniss der Flora von Java No. 18. Beschreibung von <em>Kalanchoë Schumacheri</em> vom Idjen-Plateau und Revision der Crassulaceae von Java. (Mit 2 lithographischen Tafeln.). <em>Bulletin du Jardin Botanique de Buitenzorg</em> sér. 3, 1 (3): 169–180, Figures 14 and 15. [https://www.biodiversitylibrary.org/item/27173#page/205/mode/1up (text); https://www.biodiversitylibrary.org/item/27173#page/243/mode/1up (figures)]</p>
<p>Kroon, G.H., De Jong, J. & Van Raamsdonk, L.W.D. (1989) Evaluation of the genus <em>Kalanchoe</em> for breeding of new cutflowers and potplants. <em>Acta Horticulturae</em> (International Symposium on the Development of New Floricultural Crops) 252: 111–116. https://doi.org/10.17660/ActaHortic.1989.252.14</p>
<p>Raymond-Hamet. (1933) Observations sur le <em>Kalanchoe uniflora</em> (Stapf) Raymond-Hamet. <em>Bulletin de la Société Botanique de France</em> 80 (4): 542–548. https://doi.org/10.1080/00378941.1933.10833878</p>
<p>Salisbury, R.A. (1805 [1 June 1805]) <em>The Paradisus Londinensis: or coloured figures of plants cultivated in the vicinity of the metropolis</em> 1. Published by William Hooker, No. 6, Frith Street, city not stated, presumably London, 157 pp. & 117 plates. https://doi.org/10.5962/bhl.title.53520</p>
<p>Smith, G.F. (2020a) <em>Kalanchoe</em> Adans. subg. <em>Fernandesiae</em> Gideon F.Sm. (Crassulaceae subfam. Kalanchooideae), a new subgenus from southern and south-tropical Africa. <em>Haseltonia</em> 27: 2–7. https://doi.org/10.2985/026.027.0101</p>
<p>Smith, G.F. (2020b) Character inheritance in <em>Kalanchoe </em>‘Vivien’ Bischofberger (Crassulaceae subfam. Kalanchooideae), a cultivar derived from an interspecific hybrid between two southern African species (<em>K. luciae </em>Raym.-Hamet × <em>K. sexangularis </em>N.E.Br.). <em>Bradleya</em> 38: 3–13. https://doi.org/10.25223/brad.n38.2020.a1</p>
<p>Smith, G.F. (2020c) <em>Kalanchoe</em> ×<em>estrelae</em> (Crassulaceae subfam. Kalanchooideae): a new nothospecies for the hybrid between <em>K. luciae</em> and <em>K. sexangularis</em>. <em>Phytotaxa</em> 441 (2): 225–228. https://doi.org/10.11646/phytotaxa.441.2.11</p>
<p>Smith, G.F. (2021) Reinstatement of <em>Kalanchoe</em> [subg. <em>Bryophyllum</em>] sect. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae) for <em>K. porphyrocalyx</em> and <em>K. uniflora</em>, two Madagascan endemics. <em>Bradleya</em> 39: 177–181. https://doi.org/10.25223/brad.n39.2021.a17</p>
<p>Smith, G.F. & Figueiredo, E. (2018) The infrageneric classification and nomenclature of <em>Kalanchoe </em>Adans. (Crassulaceae), with special reference to the southern African species. <em>Bradleya </em>36: 162–172. https://doi.org/10.25223/brad.n36.2018.a10</p>
<p>Smith, G.F., Figueiredo, E., Loureiro, J. & Crouch, N.R. (2019b) <em>Kalanchoe</em> ×<em>gunniae</em> Gideon F.Sm & Figueiredo (Crassulaceae), a new South African nothospecies derived from <em>Kalanchoe paniculata</em> Harv. × <em>Kalanchoe sexangularis</em> N.E.Br.<em> Bradleya</em> 37: 141–150. https://doi.org/10.25223/brad.n37.2019.a9</p>
<p>Smith, G.F., Figueiredo, E. & Van Wyk, A.E. (2019a) Kalanchoe <em>(Crassulaceae) in southern Africa. Classification, biology, and cultivation</em>. Academic Press, an imprint of Elsevier, London, San Diego, Cambridge (USA), and Oxford, 328 pp.</p>
<p>Smith, G.F., Klein, D.-P. & Shtein, R. (2021b) A revision of the Malagasy <em>Kalanchoe</em> subg. <em>Kitchingia</em> (Crassulaceae subfam. Kalanchooideae): history, taxonomy, and nomenclature. <em>Phytotaxa</em> 507 (1): 67–80. https://doi.org/10.11646/phytotaxa.507.1.3</p>
<p>Smith, G.F., Shtein, R. & Klein, D.-P. (2021a) <em>Kalanchoe</em> subg. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae), a new rank for <em>K</em>. sect. <em>Alatae</em>. <em>Phytotaxa</em> 509 (2): 255–258. https://doi.org/10.11646/phytotaxa.509.2.9</p>
<p>Thiers, B. (2021 [continuously updated]) <em>Index herbariorum: a global directory of public herbaria and associated staff</em>. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (accessed 2021)</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) <em>International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. Koeltz Botanical Books, Oberreifenberg. [Regnum Vegetabile 159], 254 pp. https://doi.org/10.12705/Code.2018</p>
<p>Von Poellnitz, [J.]K.[L.A.]. (1934) XV. <em>Kalanchoe Blossfeldiana</em>. <em>Repertorium specierum novarum regni vegetabilis. Centralblatt für Sammlung und Veröffentlichung von Einzeldiagnosen neuer Pflanzen</em> 35: 159–160. [http://bibdigital.rjb.csic.es/ing/Libro.php?Libro=6015]</p>
<p>Baker, J.G. (1881 [21 February]) Notes on a collection of flowering plants made by L. Kitching, Esq., in Madagascar in 1879 (Plates VII. & VIII.). <em>Journal of the Linnean Society, Botany</em> 18: 264–281. [https://www.biodiversitylibrary.org/item/8373#page/271/mode/1up]</p>
<p>Brown, N.E. (1913) 1436. <em>Kalanchoe sexangularis</em>, N. E. Brown [Crassulaceae]. <em>In</em>: Diagnoses Africanae LIII. <em>Bulletin of Miscellaneous Information, Kew</em> 1913 (3): 118–123. [https://www.biodiversitylibrary.org/item/43726#page/125/mode/1up]</p>
<p>Fernandes, R.B. (1980) Notes sur quelques espèces du genre <em>Kalanchoe</em> Adans. <em>Boletim da Sociedade Broteriana</em> (série 2.<sup>a</sup>) 53: 325–442.</p>
<p>Forbes, H. (1941) <em>Kalanchoe albiflora</em> H. Forbes, <em>sp. nov.</em>, pp. 37–39. <em>In</em>: Anonymous, Newly described species and new combinations. <em>Bothalia</em> 4: 37–41, + plate on p. 39. [https://archive.org/details/bothaliavolume404unse/page/36/mode/2up]</p>
<p>Gunn, M. & Codd, L.E.[W.]. (1981) <em>Botanical exploration of southern Africa. An illustrated history of early botanical literature on the Cape flora. Biographical accounts of the leading plant collectors and their activities in southern Africa from the days of the East India Company until modern times</em>. A.A. Balkema, Cape Town, 400 pp.</p>
<p>Hamet, R. (1908) <em>Kalanchoe luciae</em> <em>sp. nov.</em> <em>Bulletin de l’Herbier Boissier</em>, sér. 2, 8: 254–257. [https://www.biodiversitylibrary.org/item/104945#page/290/mode/1up]</p>
<p>Harvey, W.H. (1862) Order LIII. Crassulaceae, D.C. VIII. <em>Kalanchoe</em>, Adans. <em>In</em>: W.H. Harvey & O.W. Sonder, <em>Flora capensis</em> [<em>being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal</em>] 2. L. Reeve & Co., Ltd, Kent, pp. 378–380. [https://www.biodiversitylibrary.org/item/15233#page/386/mode/1up]</p>
<p>IPNI. (2021 [continuously updated]) <em>The International Plant Names Index</em>. Available from: https://www.ipni.org/ (accessed 2021)</p>
<p>Koorders, S.H. (1919 [Mai] [24 October 1918]) Beitrag zur Kenntniss der Flora von Java No. 18. Beschreibung von <em>Kalanchoë Schumacheri</em> vom Idjen-Plateau und Revision der Crassulaceae von Java. (Mit 2 lithographischen Tafeln.). <em>Bulletin du Jardin Botanique de Buitenzorg</em> sér. 3, 1 (3): 169–180, Figures 14 and 15. [https://www.biodiversitylibrary.org/item/27173#page/205/mode/1up (text); https://www.biodiversitylibrary.org/item/27173#page/243/mode/1up (figures)]</p>
<p>Kroon, G.H., De Jong, J. & Van Raamsdonk, L.W.D. (1989) Evaluation of the genus <em>Kalanchoe</em> for breeding of new cutflowers and potplants. <em>Acta Horticulturae</em> (International Symposium on the Development of New Floricultural Crops) 252: 111–116. https://doi.org/10.17660/ActaHortic.1989.252.14</p>
<p>Raymond-Hamet. (1933) Observations sur le <em>Kalanchoe uniflora</em> (Stapf) Raymond-Hamet. <em>Bulletin de la Société Botanique de France</em> 80 (4): 542–548. https://doi.org/10.1080/00378941.1933.10833878</p>
<p>Salisbury, R.A. (1805 [1 June 1805]) <em>The Paradisus Londinensis: or coloured figures of plants cultivated in the vicinity of the metropolis</em> 1. Published by William Hooker, No. 6, Frith Street, city not stated, presumably London, 157 pp. & 117 plates. https://doi.org/10.5962/bhl.title.53520</p>
<p>Smith, G.F. (2020a) <em>Kalanchoe</em> Adans. subg. <em>Fernandesiae</em> Gideon F.Sm. (Crassulaceae subfam. Kalanchooideae), a new subgenus from southern and south-tropical Africa. <em>Haseltonia</em> 27: 2–7. https://doi.org/10.2985/026.027.0101</p>
<p>Smith, G.F. (2020b) Character inheritance in <em>Kalanchoe </em>‘Vivien’ Bischofberger (Crassulaceae subfam. Kalanchooideae), a cultivar derived from an interspecific hybrid between two southern African species (<em>K. luciae </em>Raym.-Hamet × <em>K. sexangularis </em>N.E.Br.). <em>Bradleya</em> 38: 3–13. https://doi.org/10.25223/brad.n38.2020.a1</p>
<p>Smith, G.F. (2020c) <em>Kalanchoe</em> ×<em>estrelae</em> (Crassulaceae subfam. Kalanchooideae): a new nothospecies for the hybrid between <em>K. luciae</em> and <em>K. sexangularis</em>. <em>Phytotaxa</em> 441 (2): 225–228. https://doi.org/10.11646/phytotaxa.441.2.11</p>
<p>Smith, G.F. (2021) Reinstatement of <em>Kalanchoe</em> [subg. <em>Bryophyllum</em>] sect. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae) for <em>K. porphyrocalyx</em> and <em>K. uniflora</em>, two Madagascan endemics. <em>Bradleya</em> 39: 177–181. https://doi.org/10.25223/brad.n39.2021.a17</p>
<p>Smith, G.F. & Figueiredo, E. (2018) The infrageneric classification and nomenclature of <em>Kalanchoe </em>Adans. (Crassulaceae), with special reference to the southern African species. <em>Bradleya </em>36: 162–172. https://doi.org/10.25223/brad.n36.2018.a10</p>
<p>Smith, G.F., Figueiredo, E., Loureiro, J. & Crouch, N.R. (2019b) <em>Kalanchoe</em> ×<em>gunniae</em> Gideon F.Sm & Figueiredo (Crassulaceae), a new South African nothospecies derived from <em>Kalanchoe paniculata</em> Harv. × <em>Kalanchoe sexangularis</em> N.E.Br.<em> Bradleya</em> 37: 141–150. https://doi.org/10.25223/brad.n37.2019.a9</p>
<p>Smith, G.F., Figueiredo, E. & Van Wyk, A.E. (2019a) Kalanchoe <em>(Crassulaceae) in southern Africa. Classification, biology, and cultivation</em>. Academic Press, an imprint of Elsevier, London, San Diego, Cambridge (USA), and Oxford, 328 pp.</p>
<p>Smith, G.F., Klein, D.-P. & Shtein, R. (2021b) A revision of the Malagasy <em>Kalanchoe</em> subg. <em>Kitchingia</em> (Crassulaceae subfam. Kalanchooideae): history, taxonomy, and nomenclature. <em>Phytotaxa</em> 507 (1): 67–80. https://doi.org/10.11646/phytotaxa.507.1.3</p>
<p>Smith, G.F., Shtein, R. & Klein, D.-P. (2021a) <em>Kalanchoe</em> subg. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae), a new rank for <em>K</em>. sect. <em>Alatae</em>. <em>Phytotaxa</em> 509 (2): 255–258. https://doi.org/10.11646/phytotaxa.509.2.9</p>
<p>Thiers, B. (2021 [continuously updated]) <em>Index herbariorum: a global directory of public herbaria and associated staff</em>. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (accessed 2021)</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) <em>International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. Koeltz Botanical Books, Oberreifenberg. [Regnum Vegetabile 159], 254 pp. https://doi.org/10.12705/Code.2018</p>
<p>Von Poellnitz, [J.]K.[L.A.]. (1934) XV. <em>Kalanchoe Blossfeldiana</em>. <em>Repertorium specierum novarum regni vegetabilis. Centralblatt für Sammlung und Veröffentlichung von Einzeldiagnosen neuer Pflanzen</em> 35: 159–160. [http://bibdigital.rjb.csic.es/ing/Libro.php?Libro=6015]</p>